treatments-xml/data/03/DC/87/03DC8791022FFF9B6475FCEFF19C1743.xml

<document id="49A5721B0304DB6AD5A211953B0E66F1" ID-DOI="10.37520/aemnp.2024.016" ID-ISSN="1804-6487" ID-Zenodo-Dep="14655591" ID-ZooBank="187DAF15-7C4F-4A4C-9AA3-8753B824CCCB" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1736957651372" checkinUser="felipe" docAuthor="Hájek, Jiří &amp; Shaverdo, Helena" docDate="2024" docId="03DC8791022FFF9B6475FCEFF19C1743" docLanguage="en" docName="ActaEntMusNatPra.64.2.243-247.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 64 (2)" docSource="https://doi.org/10.37520/aemnp.2024.016" docStyle="DocumentStyle:BA2C5C8540539042E893A65A182CEA85.8:ActaEntMusNatPra.2018-.journal_article" docStyleId="BA2C5C8540539042E893A65A182CEA85" docStyleName="ActaEntMusNatPra.2018-.journal_article" docStyleVersion="8" docTitle="Austrelatus mirai Hájek &amp; Shaverdo, 2024, sp. nov." docType="treatment" docVersion="2" lastPageNumber="246" masterDocId="FFE5FFE9022EFF98654FFFB5F1401616" masterDocTitle="Two new species of Austrelatus diving beetles from continental Southeast Asia, linking the distribution area of the genus (Coleoptera: Dytiscidae: Copelatinae)" masterLastPageNumber="247" masterPageNumber="243" pageNumber="244" updateTime="1736958082241" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-ND-3.0">
<mods:mods id="200776ECA98ED56E9CFBD4F56C95BB0B" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="D15BFACC2B59BE507E47C007752A478B">
<mods:title id="FE80A09762DE06B0DEA09143480D5E62">Two new species of Austrelatus diving beetles from continental Southeast Asia, linking the distribution area of the genus (Coleoptera: Dytiscidae: Copelatinae)</mods:title>
</mods:titleInfo>
<mods:name id="8CBC5C69333B9F5C31C2A74B7E539349" type="personal">
<mods:role id="83C90A9B2101C29CCC1C0FCDD7C59B29">
<mods:roleTerm id="E2D9FFA56A6FAEA6748008A775CC85E4">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="37F79C6D84E970D98601ED72104B9195">Hájek, Jiří</mods:namePart>
<mods:affiliation id="017319C71BA17A8A32CCE6E0BE8BC3D5">Department of Entomology, National Museum of the Czech Republic, Cirkusová 1740, CZ- 193 00 Praha</mods:affiliation>
</mods:name>
<mods:name id="65558D3F69B5398E011E349A5094CABE" type="personal">
<mods:role id="9701A065105257D3EB9995889F9A649B">
<mods:roleTerm id="578493355787969CA68F25A3AE65B28F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5938E95A1EE273466C1903CAECCB679E">Shaverdo, Helena</mods:namePart>
<mods:affiliation id="F3AF9156A102E4A9590B353D73A6BAD7">Naturhistorisches Museum Wien, Burgring</mods:affiliation>
</mods:name>
<mods:typeOfResource id="811D6BEA93BC896FD50F87A098E360A2">text</mods:typeOfResource>
<mods:relatedItem id="F2B2D9281C6E9100B883A5C4CD93945F" type="host">
<mods:titleInfo id="A591581E86B081D2E2EC2E86521668D2">
<mods:title id="46565505A97CC149BCFE6089EED12B23">Acta Entomologica Musei Nationalis Pragae</mods:title>
</mods:titleInfo>
<mods:part id="9D6AF9516D696A69B1ADD89D89F7A157">
<mods:date id="72833026F4B1921001B5F40E256D5292">2024</mods:date>
<mods:detail id="520E18F57C0690767581A33F89C1D489" type="series">
<mods:title id="E37CD15C29754922597DDC37A5BA4EEC">Acta. Ent. Mus. Natl. Pragae</mods:title>
</mods:detail>
<mods:detail id="AB8CD683A4DFB3A8E1286810740902B6" type="pubDate">
<mods:number id="8A213D8483C418BE856D79710B10A8AC">2024-10-01</mods:number>
</mods:detail>
<mods:detail id="FF40A9AD4865909A3325557954D3E775" type="volume">
<mods:number id="473136A6F283509926C95BCBCF7BF70F">64</mods:number>
</mods:detail>
<mods:detail id="A9F983B9E62E3F9E22A7E59CA10DC497" type="issue">
<mods:number id="EE93ACD43B7C19B64535C6FF3B3D8EA4">2</mods:number>
</mods:detail>
<mods:extent id="BEA34510601B24E22542B8BDC66AFFA2" unit="page">
<mods:start id="EB9D55B151FD1A18022B947C8D4340FA">243</mods:start>
<mods:end id="F273FC09A0B06EE6C65C181AACA15EE9">247</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="CF697FB1D1169D3C52AED8535D799A5B">
<mods:url id="2F98A1E4B58160470974263E4918F32F">https://doi.org/10.37520/aemnp.2024.016</mods:url>
</mods:location>
<mods:classification id="52297614CA811E71D84DEE2579FB4AA2">journal article</mods:classification>
<mods:identifier id="77744AF3AECFE1DE101301C3A5E8FAE8" type="DOI">10.37520/aemnp.2024.016</mods:identifier>
<mods:identifier id="90770A855B3052033260D2FDC035A61B" type="ISSN">1804-6487</mods:identifier>
<mods:identifier id="FD51FF00D5411A327C9A5F89FD4C4C05" type="Zenodo-Dep">14655591</mods:identifier>
<mods:identifier id="8DB91980B6750A32608AF126B2E16B80" type="ZooBank">187DAF15-7C4F-4A4C-9AA3-8753B824CCCB</mods:identifier>
</mods:mods>
<treatment id="03DC8791022FFF9B6475FCEFF19C1743" LSID="urn:lsid:plazi:treatment:03DC8791022FFF9B6475FCEFF19C1743" httpUri="http://treatment.plazi.org/id/03DC8791022FFF9B6475FCEFF19C1743" lastPageId="3" lastPageNumber="246" pageId="1" pageNumber="244">
<subSubSection id="C36F650C022FFF996475FCEFF3161564" box="[314,598,858,882]" pageId="1" pageNumber="244" type="nomenclature">
<paragraph id="8BCA3687022FFF996475FCEFF3161564" blockId="1.[314,598,858,882]" box="[314,598,858,882]" pageId="1" pageNumber="244">
<heading id="D08281EB022FFF996475FCEFF3161564" box="[314,598,858,882]" centered="true" fontSize="10" level="2" pageId="1" pageNumber="244" reason="3">
<emphasis id="B901EA95022FFF996475FCEFF3161564" bold="true" box="[314,598,858,882]" pageId="1" pageNumber="244">
<taxonomicName id="4C754D04022FFF996475FCEFF0B91564" authorityName="Hájek &amp; Shaverdo" authorityYear="2024" box="[314,505,858,882]" class="Insecta" family="Dytiscidae" genus="Austrelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="244" phylum="Arthropoda" rank="species" species="mirai" status="sp. nov.">
<emphasis id="B901EA95022FFF996475FCEFF0B91564" bold="true" box="[314,505,858,882]" italics="true" pageId="1" pageNumber="244">Austrelatus mirai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23257EE022FFF99674FFCEEF3161564" box="[512,598,859,882]" pageId="1" pageNumber="244" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36F650C022FFF9964DEFCCCF0BE159A" box="[401,510,889,908]" pageId="1" pageNumber="244" type="description">
<paragraph id="8BCA3687022FFF9964DEFCCCF0BE159A" blockId="1.[401,510,889,908]" box="[401,510,889,908]" pageId="1" pageNumber="244">
(
<figureCitation id="134E2A02022FFF9964D8FCCCF0B8159A" box="[407,504,889,908]" captionStart="Figs 1–6. 1–2" captionStartId="2.[151,188,1689,1708]" captionTargetBox="[177,1413,215,1650]" captionTargetId="figure-183@2.[151,1436,189,1673]" captionTargetPageId="2" captionText="Figs 1–6. 1–2 – habitus of Austrelatus holotypes: 1 – A. mirai sp. nov.; 2 – A. riberai sp. nov. 3–6 – male genitalia of Austrelatus: 3–4 – A. mirai sp. nov.; 5–6 – A. riberai sp. nov. 3, 5 – median lobe in lateral (a) and ventral (b) view; 4, 6 – parameres. Scale bar = 0.5 mm (Figs 3–6)." figureDoi="http://doi.org/10.5281/zenodo.14655593" httpUri="https://zenodo.org/record/14655593/files/figure.png" pageId="1" pageNumber="244">Figs 1, 3–4</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C36F650C022FFF9965D8FC16F0CE1267" pageId="1" pageNumber="244" type="materials_examined">
<paragraph id="8BCA3687022FFF9965D8FC16F34E15C7" blockId="1.[151,761,931,1137]" pageId="1" pageNumber="244">
<emphasis id="B901EA95022FFF9965D8FC16F05315A0" bold="true" box="[151,275,931,950]" pageId="1" pageNumber="244">
<typeStatus id="54CE8825022FFF9965D8FC16F18415A0" box="[151,196,931,950]" pageId="1" pageNumber="244">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F3627617022FFF996454FC16F02C15A0" box="[283,364,931,950]" name="Malaysia" pageId="1" pageNumber="244">Malaysia</collectingCountry>
, 
<collectingRegion id="49B1F865022FFF996439FC16F0B715A0" box="[374,503,931,950]" country="Malaysia" name="Selangor" pageId="1" pageNumber="244">Selangor State</collectingRegion>
, Rawang Township, Templer Park Forest Reserve, ca. 
<geoCoordinate id="EE415040022FFF996424FC0BF0F215C7" box="[363,434,958,977]" degrees="03" direction="north" minutes="17" orientation="latitude" pageId="1" pageNumber="244" precision="925" value="3.2833333">03°17′N</geoCoordinate>
<geoCoordinate id="EE415040022FFF9964F7FC0BF34915C7" box="[440,521,958,977]" degrees="103" direction="east" minutes="38" orientation="longitude" pageId="1" pageNumber="244" precision="925" value="103.63333">103°38′E</geoCoordinate>
.
</paragraph>
<paragraph id="8BCA3687022FFF9965D8FC6DF0CE1267" blockId="1.[151,761,931,1137]" pageId="1" pageNumber="244">
<emphasis id="B901EA95022FFF9965D8FC6DF06815FD" bold="true" box="[151,296,984,1003]" pageId="1" pageNumber="244">Type material.</emphasis>
<typeStatus id="54CE8825022FFF99647DFC6DF0D115FD" box="[306,401,984,1003]" pageId="1" pageNumber="244" type="holotype">HOLOTYPE</typeStatus>
: Ô (
<collectionCode id="ED64AE42022FFF99648DFC6DF35015FD" box="[450,528,984,1003]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="1" pageNumber="244" type="Museum">NHMW</collectionCode>
), labelled: ‘ 
<collectingCountry id="F3627617022FFF9967CFFC6DF3B715FD" box="[640,759,984,1003]" name="Malaysia" pageId="1" pageNumber="244">MALAYSIA</collectingCountry>
<date id="FFCB1047022FFF9965D8FC46F1AB1210" box="[151,235,1011,1030]" pageId="1" pageNumber="244" value="1992-01-21">21.I.1992</date>
/ 
<collectingRegion id="49B1F865022FFF99644EFC46F0381210" box="[257,376,1011,1030]" country="Malaysia" name="Selangor" pageId="1" pageNumber="244">SELANGOR</collectingRegion>
: Templer / Park N K.L. / leg. Jäch (1) [p] // 
<typeStatus id="54CE8825022FFF9965E4FBBBF05C1237" box="[171,284,1038,1057]" pageId="1" pageNumber="244" type="holotype">HOLOTYPE</typeStatus>
Ô / 
<taxonomicName id="4C754D04022FFF996406FBBBF0941237" authorityName="Shaverdo, Hajek, Hendrich, Surbakti, Panjaitan &amp; Balke" authorityYear="2023" box="[329,468,1038,1057]" class="Insecta" family="Dytiscidae" genus="Austrelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="244" phylum="Arthropoda" rank="genus">
<emphasis id="B901EA95022FFF996406FBBBF0941237" box="[329,468,1038,1057]" italics="true" pageId="1" pageNumber="244">AUSTRELATUS</emphasis>
</taxonomicName>
/ 
<taxonomicName id="4C754D04022FFF9964A5FBBBF3581237" authorityName="Hájek &amp; Shaverdo" authorityYear="2024" box="[490,536,1038,1057]" class="Insecta" family="Dytiscidae" genus="Austrelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="244" phylum="Arthropoda" rank="species" species="mirai" status="sp. nov.">
<emphasis id="B901EA95022FFF9964A5FBBBF3581237" box="[490,536,1038,1057]" italics="true" pageId="1" pageNumber="244">mirai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23257EE022FFF99676FFBBBF3261237" box="[544,614,1038,1057]" pageId="1" pageNumber="244" rank="species">sp. nov.</taxonomicNameLabel>
/ J. Hájek &amp; H. Shaverdo det. 2024 [p, red label]’. 
<collectingRegion id="49B1F865022FFF996482FB9DF366122D" box="[461,550,1064,1083]" country="Malaysia" name="PARATYPES" pageId="1" pageNumber="244">
<typeStatus id="54CE8825022FFF996482FB9DF366122D" box="[461,550,1064,1083]" pageId="1" pageNumber="244" type="paratype">PARATYPES</typeStatus>
</collectingRegion>
: 1 Ô 
<specimenCount id="9D73FD0E022FFF996713FB9DF3CE122D" box="[604,654,1064,1083]" count="2" pageId="1" pageNumber="244" type="female">2 ♀♀</specimenCount>
, same label data as 
<typeStatus id="54CE8825022FFF996598FBF6F0621240" box="[215,290,1091,1110]" pageId="1" pageNumber="244" type="holotype">holotype</typeStatus>
(
<collectionCode id="ED64AE42022FFF996462FBF6F0351240" box="[301,373,1091,1110]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="1" pageNumber="244" type="Museum">NHMW</collectionCode>
, 
<collectionCode id="ED64AE42022FFF996432FBF6F0FD1240" box="[381,445,1091,1110]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="1" pageNumber="244">NMPC</collectionCode>
). All 
<typeStatus id="54CE8825022FFF9964A4FBF6F37D1240" box="[491,573,1091,1110]" pageId="1" pageNumber="244" type="paratype">paratypes</typeStatus>
are provided with the respective red printed labels.
</paragraph>
</subSubSection>
<subSubSection id="C36F650C022FFF9965D8FB3CF4C21E0D" pageId="1" pageNumber="244" type="description">
<paragraph id="8BCA3687022FFF9965D8FB3CF38D1316" blockId="1.[151,761,1160,2080]" pageId="1" pageNumber="244">
<emphasis id="B901EA95022FFF9965D8FB3CF09512B6" bold="true" box="[151,469,1160,1184]" pageId="1" pageNumber="244">
Description. 
<emphasis id="B901EA95022FFF996462FB3DF09512B6" box="[301,469,1160,1184]" italics="true" pageId="1" pageNumber="244">
Male 
<typeStatus id="54CE8825022FFF99643FFB3DF09112B6" box="[368,465,1160,1184]" pageId="1" pageNumber="244" type="holotype">holotype</typeStatus>
.
</emphasis>
</emphasis>
Habitus (
<figureCitation id="134E2A02022FFF996709FB3CF3C812B6" box="[582,648,1161,1184]" captionStart="Figs 1–6. 1–2" captionStartId="2.[151,188,1689,1708]" captionTargetBox="[177,1413,215,1650]" captionTargetId="figure-183@2.[151,1436,189,1673]" captionTargetPageId="2" captionText="Figs 1–6. 1–2 – habitus of Austrelatus holotypes: 1 – A. mirai sp. nov.; 2 – A. riberai sp. nov. 3–6 – male genitalia of Austrelatus: 3–4 – A. mirai sp. nov.; 5–6 – A. riberai sp. nov. 3, 5 – median lobe in lateral (a) and ventral (b) view; 4, 6 – parameres. Scale bar = 0.5 mm (Figs 3–6)." figureDoi="http://doi.org/10.5281/zenodo.14655593" httpUri="https://zenodo.org/record/14655593/files/figure.png" pageId="1" pageNumber="244">Fig. 1</figureCitation>
) elongate oblong oval, broadest in one third of elytral length, dorsally convex; body outline continuous, without angle between base of pronotum and elytra. Dorsal surface submatt.
</paragraph>
<paragraph id="8BCA3687022FFF9965FAFABCF3AF1396" blockId="1.[151,761,1160,2080]" pageId="1" pageNumber="244">
<emphasis id="B901EA95022FFF9965FAFABCF07F1336" box="[181,319,1289,1312]" italics="true" pageId="1" pageNumber="244">Colouration.</emphasis>
Head ferruginous with darker circumocular band; pronotum ferruginous, somewhat darkened along anterior and posterior margin; elytra brown-blackish with paler base; appendages orange; ventral side ferruginous.
</paragraph>
<paragraph id="8BCA3687022FFF9965FAFA3CF3B910F6" blockId="1.[151,761,1160,2080]" pageId="1" pageNumber="244">
<emphasis id="B901EA95022FFF9965FAFA3CF1AF13B6" box="[181,239,1417,1440]" italics="true" pageId="1" pageNumber="244">Head</emphasis>
moderately broad, ca. 0.66× width of pronotum, semicircular. Anterior margin of clypeus indistinctly concave. Antenna with antennomeres long and slender. Reticulation consisting of very weakly impressed, hardly perceptible polygonal isodiametric meshes. Punctation double, consisting of coarse setigerous punctures and fine punctures spread sparsely on surface; row of coarse punctures present alongside inner margin of eyes, several punctures present also in fronto-clypeal depressions. Surface between eyes with several short longitudinal strioles; a few short transverse strioles present also in centre of head.
</paragraph>
<paragraph id="8BCA3687022FFF9965FAF95CF37A1E16" blockId="1.[151,761,1160,2080]" pageId="1" pageNumber="244">
<emphasis id="B901EA95022FFF9965FAF95CF0611116" box="[181,289,1769,1792]" italics="true" pageId="1" pageNumber="244">Pronotum</emphasis>
strongly transverse (width/length ratio = 2.46), broadest between posterior angles, lateral margins moderately curved. Lateral sides with thin beading except for anterior angles. Reticulation similar to that of head, but even less perceptible. Punctation similar to that of head; rows of coarse setigerous punctures present along anterior margin, close to lateral sides and in shallow basolateral depressions along basal margin. Whole surface of pronotum covered with long longitudinal strioles.
</paragraph>
<paragraph id="8BCA3687022FFF9965FAF7BCF35B1E36" blockId="1.[151,761,1160,2080]" box="[181,539,2057,2080]" pageId="1" pageNumber="244">
<taxonomicName id="4C754D04022FFF9965FAF7BCF05E1E36" box="[181,286,2057,2080]" class="Trilobita" family="Styginidae" genus="Scutellum" kingdom="Animalia" order="Corynexochida" pageId="1" pageNumber="244" phylum="Arthropoda" rank="genus">
<emphasis id="B901EA95022FFF9965FAF7BCF05E1E36" box="[181,286,2057,2080]" italics="true" pageId="1" pageNumber="244">Scutellum</emphasis>
</taxonomicName>
transversely triangular.
</paragraph>
<paragraph id="8BCA3687022FFF996615FF71F5F0144D" blockId="1.[828,1437,196,2075]" pageId="1" pageNumber="244">
<emphasis id="B901EA95022FFF996615FF71F2E416CD" box="[858,932,196,219]" italics="true" pageId="1" pageNumber="244">Elytra.</emphasis>
Base of elytra as broad as pronotal base; lateral margins of elytra slightly diverging in basal third, subparallel in middle third, distinctly narrowing to apex in apical third. Eleven dorsal striae and one submarginal longitudinal stria present on each elytron: all striae beginning at base; striae 1, 3, 5–6, 8 and 11 ending subapically; striae 2 and 4 longest, ending close to apex; striae 7, 9–10 progressively shorter, the latter ending at apical fourth; submarginal stria long, beginning before elytral mid-length and ending subapically. Reticulation similar to that of head. Punctation consisting of coarse setigerous punctures and very fine sparse punctures; coarse punctures present along elytral striae and lateral margins of elytra.
</paragraph>
<paragraph id="8BCA3687022FFF996615FDD1F28D14ED" blockId="1.[828,1437,196,2075]" pageId="1" pageNumber="244">
<emphasis id="B901EA95022FFF996615FDD1F2D4146D" box="[858,916,612,635]" italics="true" pageId="1" pageNumber="244">Legs.</emphasis>
Protibia broadened anteriorly, club shaped. Pro- and mesotarsomeres 1–3 distinctly broadened, with rows of adhesive setae on their ventral side; posterior (outer) protarsal claw slightly broader and more curved than anterior claw.
</paragraph>
<paragraph id="8BCA3687022FFF996615FCB1F437138D" blockId="1.[828,1437,196,2075]" pageId="1" pageNumber="244">
<emphasis id="B901EA95022FFF996615FCB1F540150D" box="[858,1024,772,795]" italics="true" pageId="1" pageNumber="244">Ventral surface.</emphasis>
Prosternum sinuate anteriorly, obtusely keeled medially; rather coarsely punctate; reticulation consisting of fine transverse meshes. Prosternal process shortly lanceolate, in cross-section convex, apex obtuse; process distinctly bordered laterally; reticulation consisting of shallow, hardly perceptible polygonal meshes. Metaventrite with reticulation consisting of transverse polygonal meshes; lateral parts of metaventrite (“metasternal wings”) tongue-shaped, slender. Metacoxal lines nearly complete, absent only close to metaventrite. Metacoxal plates covered with rather long, oblique strioles; reticulation consisting of elongate, longitudinal polygonal meshes. Metacoxal processes rounded at posterior margin.Abdominal ventrites I–II with longitudinal strioles; ventrites III–IV with oblique strioles laterally; ventrite V with a few transverse striolae laterally. Tuft of setae present antero-medially on ventrites III–V; ventrite VI with setigerous punctures laterally on either side. Abdominal reticulation consisting of elongate polygonal meshes, longitudinal on ventrites I–II, oblique on ventrite III and transverse on ventrites IV–VI. Punctation consisting of fine, sparsely distributed punctures.
</paragraph>
<paragraph id="8BCA3687022FFF996615FA11F50F10CD" blockId="1.[828,1437,196,2075]" pageId="1" pageNumber="244">
<emphasis id="B901EA95022FFF996615FA11F54613AD" box="[858,1030,1444,1467]" italics="true" pageId="1" pageNumber="244">Male genitalia.</emphasis>
Median lobe of aedeagus in lateral view sickle-shaped, widest after mid-length, then rather subparallel and in apical eighth tapering to pointed apex (
<figureCitation id="134E2A02022FFF99660BF9B1F2D3100D" box="[836,915,1540,1563]" captionStart="Figs 1–6. 1–2" captionStartId="2.[151,188,1689,1708]" captionTargetBox="[177,1413,215,1650]" captionTargetId="figure-183@2.[151,1436,189,1673]" captionTargetPageId="2" captionText="Figs 1–6. 1–2 – habitus of Austrelatus holotypes: 1 – A. mirai sp. nov.; 2 – A. riberai sp. nov. 3–6 – male genitalia of Austrelatus: 3–4 – A. mirai sp. nov.; 5–6 – A. riberai sp. nov. 3, 5 – median lobe in lateral (a) and ventral (b) view; 4, 6 – parameres. Scale bar = 0.5 mm (Figs 3–6)." figureDoi="http://doi.org/10.5281/zenodo.14655593" httpUri="https://zenodo.org/record/14655593/files/figure.png" pageId="1" pageNumber="244">Fig. 3a</figureCitation>
); in ventral view, indistinctly constricted in two thirds of its length, then tapering to pointed apex (
<figureCitation id="134E2A02022FFF99600EF991F4CF102D" box="[1345,1423,1572,1595]" captionStart="Figs 1–6. 1–2" captionStartId="2.[151,188,1689,1708]" captionTargetBox="[177,1413,215,1650]" captionTargetId="figure-183@2.[151,1436,189,1673]" captionTargetPageId="2" captionText="Figs 1–6. 1–2 – habitus of Austrelatus holotypes: 1 – A. mirai sp. nov.; 2 – A. riberai sp. nov. 3–6 – male genitalia of Austrelatus: 3–4 – A. mirai sp. nov.; 5–6 – A. riberai sp. nov. 3, 5 – median lobe in lateral (a) and ventral (b) view; 4, 6 – parameres. Scale bar = 0.5 mm (Figs 3–6)." figureDoi="http://doi.org/10.5281/zenodo.14655593" httpUri="https://zenodo.org/record/14655593/files/figure.png" pageId="1" pageNumber="244">Fig. 3b</figureCitation>
); ventral lobe apically pointed, closely pressed from left side to dorsal lobe in ventral view (
<figureCitation id="134E2A02022FFF99613AF9D1F582106D" box="[1141,1218,1636,1659]" captionStart="Figs 1–6. 1–2" captionStartId="2.[151,188,1689,1708]" captionTargetBox="[177,1413,215,1650]" captionTargetId="figure-183@2.[151,1436,189,1673]" captionTargetPageId="2" captionText="Figs 1–6. 1–2 – habitus of Austrelatus holotypes: 1 – A. mirai sp. nov.; 2 – A. riberai sp. nov. 3–6 – male genitalia of Austrelatus: 3–4 – A. mirai sp. nov.; 5–6 – A. riberai sp. nov. 3, 5 – median lobe in lateral (a) and ventral (b) view; 4, 6 – parameres. Scale bar = 0.5 mm (Figs 3–6)." figureDoi="http://doi.org/10.5281/zenodo.14655593" httpUri="https://zenodo.org/record/14655593/files/figure.png" pageId="1" pageNumber="244">Fig. 3b</figureCitation>
). Paramere narrowly triangular, with basal part broad, laterally rounded; distal part narrow, subparallel, apically with long setae; apical lobe club-shaped (
<figureCitation id="134E2A02022FFF99614FF971F50210CD" box="[1024,1090,1732,1755]" captionStart="Figs 1–6. 1–2" captionStartId="2.[151,188,1689,1708]" captionTargetBox="[177,1413,215,1650]" captionTargetId="figure-183@2.[151,1436,189,1673]" captionTargetPageId="2" captionText="Figs 1–6. 1–2 – habitus of Austrelatus holotypes: 1 – A. mirai sp. nov.; 2 – A. riberai sp. nov. 3–6 – male genitalia of Austrelatus: 3–4 – A. mirai sp. nov.; 5–6 – A. riberai sp. nov. 3, 5 – median lobe in lateral (a) and ventral (b) view; 4, 6 – parameres. Scale bar = 0.5 mm (Figs 3–6)." figureDoi="http://doi.org/10.5281/zenodo.14655593" httpUri="https://zenodo.org/record/14655593/files/figure.png" pageId="1" pageNumber="244">Fig. 4</figureCitation>
).
</paragraph>
<paragraph id="8BCA3687022FFF996615F956F2CF112D" blockId="1.[828,1437,196,2075]" pageId="1" pageNumber="244">
<emphasis id="B901EA95022FFF996615F956F2F410ED" bold="true" box="[858,948,1763,1787]" pageId="1" pageNumber="244">
<emphasis id="B901EA95022FFF996615F956F2ED10ED" bold="true" box="[858,941,1763,1787]" italics="true" pageId="1" pageNumber="244">Female</emphasis>
.
</emphasis>
Identical to male in habitus. Pro- and mesotarsomeres 1–3 not dilated, without adhesive setae; proclaws slender.
</paragraph>
<paragraph id="8BCA3687022FFF996673F8F1F562118D" blockId="1.[828,1437,196,2075]" pageId="1" pageNumber="244">
<emphasis id="B901EA95022FFF996673F8F1F2FA114D" bold="true" box="[828,954,1860,1883]" pageId="1" pageNumber="244">Variability.</emphasis>
All specimens of the 
<typeStatus id="54CE8825022FFF9961D1F8F1F58C114D" box="[1182,1228,1860,1883]" pageId="1" pageNumber="244">type</typeStatus>
series are identical. Small variability can only be seen in extent of longitudinal striolae on pronotum.
</paragraph>
<paragraph id="8BCA3687022FFF996615F816F4C21E0D" blockId="1.[828,1437,196,2075]" pageId="1" pageNumber="244">
<emphasis id="B901EA95022FFF996615F816F54211AC" bold="true" box="[858,1026,1955,1978]" italics="true" pageId="1" pageNumber="244">Measurements.</emphasis>
TL: 5.0– 
<quantity id="4C8D9B62022FFF996126F811F5FB11AD" box="[1129,1211,1956,1979]" metricMagnitude="-3" metricUnit="m" metricValue="5.6" pageId="1" pageNumber="244" unit="mm" value="5.6">5.6 mm</quantity>
(mean value: 5.35 ± 
<quantity id="4C8D9B62022FFF996673F871F2E111CD" box="[828,929,1988,2011]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="244" unit="mm" value="0.2">0.20 mm</quantity>
); 
<typeStatus id="54CE8825022FFF9966F9F871F55811CD" box="[950,1048,1988,2011]" pageId="1" pageNumber="244" type="holotype">holotype</typeStatus>
: 
<quantity id="4C8D9B62022FFF99616BF871F53B11CD" box="[1060,1147,1988,2011]" metricMagnitude="-3" metricUnit="m" metricValue="5.6" pageId="1" pageNumber="244" unit="mm" value="5.6">5.6 mm</quantity>
. TL-h: 
<quantity id="4C8D9B62022FFF996181F871F41011CD" box="[1230,1360,1988,2011]" metricMagnitude="-3" metricUnit="m" metricValue="4.7" metricValueMax="4.9" metricValueMin="4.5" pageId="1" pageNumber="244" unit="mm" value="4.7" valueMax="4.9" valueMin="4.5">4.5–4.9 mm</quantity>
(mean value: 4.75 ± 
<quantity id="4C8D9B62022FFF996681F851F57211ED" box="[974,1074,2020,2043]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="244" unit="mm" value="0.15">0.15 mm</quantity>
); 
<typeStatus id="54CE8825022FFF99610BF851F5E611ED" box="[1092,1190,2020,2043]" pageId="1" pageNumber="244" type="holotype">holotype</typeStatus>
: 
<quantity id="4C8D9B62022FFF9961E0F851F44411ED" box="[1199,1284,2020,2043]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" pageId="1" pageNumber="244" unit="mm" value="4.9">4.9 mm</quantity>
. MW: 
<quantity id="4C8D9B62022FFF996003F851F2251E0D" metricMagnitude="-3" metricUnit="m" metricValue="2.4" metricValueMax="2.5" metricValueMin="2.3" pageId="1" pageNumber="244" unit="mm" value="2.4" valueMax="2.5" valueMin="2.3">2.3–2.5 mm</quantity>
(mean value: 2.45 ± 
<quantity id="4C8D9B62022FFF996107F7B1F5EB1E0D" box="[1096,1195,2052,2075]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="244" unit="mm" value="0.1">0.10 mm</quantity>
); 
<typeStatus id="54CE8825022FFF9961F2F7B1F45F1E0D" box="[1213,1311,2052,2075]" pageId="1" pageNumber="244" type="holotype">holotype</typeStatus>
: 
<quantity id="4C8D9B62022FFF996066F7B1F43E1E0D" box="[1321,1406,2052,2075]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="1" pageNumber="244" unit="mm" value="2.5">2.5 mm</quantity>
.
</paragraph>
</subSubSection>
<caption id="DF0A660F022CFF9A65D8F92CF59710D0" ID-DOI="http://doi.org/10.5281/zenodo.14655593" ID-Zenodo-Dep="14655593" httpUri="https://zenodo.org/record/14655593/files/figure.png" pageId="2" pageNumber="245" startId="2.[151,188,1689,1708]" targetBox="[177,1413,215,1650]" targetPageId="2" targetType="figure">
<paragraph id="8BCA3687022CFF9A65D8F92CF59710D0" blockId="2.[151,1437,1689,1734]" pageId="2" pageNumber="245">
Figs 1–6. 1–2 – habitus of 
<taxonomicName id="4C754D04022CFF9A6438F92CF09710BA" authorityName="Shaverdo, Hajek, Hendrich, Surbakti, Panjaitan &amp; Balke" authorityYear="2023" box="[375,471,1689,1708]" class="Insecta" family="Dytiscidae" genus="Austrelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="245" phylum="Arthropoda" rank="genus">
<emphasis id="B901EA95022CFF9A6438F92CF09710BA" box="[375,471,1689,1708]" italics="true" pageId="2" pageNumber="245">Austrelatus</emphasis>
</taxonomicName>
holotypes: 1 – 
<taxonomicName id="4C754D04022CFF9A6718F92CF3DC10BA" authorityName="Hájek &amp; Shaverdo" authorityYear="2024" box="[599,668,1689,1708]" class="Insecta" family="Dytiscidae" genus="Austrelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="245" phylum="Arthropoda" rank="species" species="mirai" status="sp. nov.">
<emphasis id="B901EA95022CFF9A6718F92CF3DC10BA" box="[599,668,1689,1708]" italics="true" pageId="2" pageNumber="245">A. mirai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23257EE022CFF9A67EEF92CF3A610BA" box="[673,742,1689,1708]" pageId="2" pageNumber="245" rank="species">sp. nov.</taxonomicNameLabel>
; 2 – 
<taxonomicName id="4C754D04022CFF9A6644F92CF21D10BA" authorityName="Hájek &amp; Shaverdo" authorityYear="2024" box="[779,861,1689,1708]" class="Insecta" family="Dytiscidae" genus="Austrelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="245" phylum="Arthropoda" rank="species" species="riberai" status="sp. nov.">
<emphasis id="B901EA95022CFF9A6644F92CF21D10BA" box="[779,861,1689,1708]" italics="true" pageId="2" pageNumber="245">A. riberai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23257EE022CFF9A662DF92CF2E410BA" box="[866,932,1689,1708]" pageId="2" pageNumber="245" rank="species">sp. nov.</taxonomicNameLabel>
3–6 – male genitalia of 
<taxonomicName id="4C754D04022CFF9A6122F92CF58D10BA" authorityName="Shaverdo, Hajek, Hendrich, Surbakti, Panjaitan &amp; Balke" authorityYear="2023" box="[1133,1229,1689,1708]" class="Insecta" family="Dytiscidae" genus="Austrelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="245" phylum="Arthropoda" rank="genus">
<emphasis id="B901EA95022CFF9A6122F92CF58D10BA" box="[1133,1229,1689,1708]" italics="true" pageId="2" pageNumber="245">Austrelatus</emphasis>
</taxonomicName>
: 3–4 – 
<taxonomicName id="4C754D04022CFF9A6043F92CF41110BA" authorityName="Hájek &amp; Shaverdo" authorityYear="2024" box="[1292,1361,1689,1708]" class="Insecta" family="Dytiscidae" genus="Austrelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="245" phylum="Arthropoda" rank="species" species="mirai" status="sp. nov.">
<emphasis id="B901EA95022CFF9A6043F92CF41110BA" box="[1292,1361,1689,1708]" italics="true" pageId="2" pageNumber="245">A. mirai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23257EE022CFF9A601AF92CF4DA10BA" box="[1365,1434,1689,1708]" pageId="2" pageNumber="245" rank="species">sp. nov.</taxonomicNameLabel>
; 5–6 – 
<taxonomicName id="4C754D04022CFF9A6582F906F05F10D0" authorityName="Hájek &amp; Shaverdo" authorityYear="2024" box="[205,287,1715,1734]" class="Insecta" family="Dytiscidae" genus="Austrelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="245" phylum="Arthropoda" rank="species" species="riberai" status="sp. nov.">
<emphasis id="B901EA95022CFF9A6582F906F05F10D0" box="[205,287,1715,1734]" italics="true" pageId="2" pageNumber="245">A. riberai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23257EE022CFF9A646AF906F02810D0" box="[293,360,1715,1734]" pageId="2" pageNumber="245" rank="species">sp. nov.</taxonomicNameLabel>
3, 5 – median lobe in lateral (a) and ventral (b) view; 4, 6 – parameres. Scale bar = 0.5 mm (Figs 3–6).
</paragraph>
</caption>
<subSubSection id="C36F650C022CFF9A65D8F94BF4C11121" pageId="2" pageNumber="245" type="diagnosis">
<paragraph id="8BCA3687022CFF9A65D8F94BF4C11121" blockId="2.[151,760,1790,2069]" lastBlockId="2.[828,1437,1792,2071]" pageId="2" pageNumber="245">
<emphasis id="B901EA95022CFF9A65D8F94BF0C51103" bold="true" box="[151,389,1790,1814]" pageId="2" pageNumber="245">Differential diagnosis.</emphasis>
With rather small body length, more or less unicolour ferruginous colouration and presence of 11 dorsal striae + a submarginal stria on each elytron, the new species is most similar to South Indian 
<taxonomicName id="4C754D04022CFF9A6730F8EBF0B81183" authority="(Hendrich &amp; Balke, 1998)" baseAuthorityName="Hendrich &amp; Balke" baseAuthorityYear="1998" class="Insecta" family="Dytiscidae" genus="Austrelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="245" phylum="Arthropoda" rank="species" species="boukali">
<emphasis id="B901EA95022CFF9A6730F8EBF1A61183" italics="true" pageId="2" pageNumber="245">Austrelatus boukali</emphasis>
(Hendrich &amp; Balke, 1998)
</taxonomicName>
and 
<taxonomicName id="4C754D04022CFF9A6769F8CBF19911A3" authority="(Wewalka, 2017)" baseAuthorityName="Wewalka" baseAuthorityYear="2017" class="Insecta" family="Dytiscidae" genus="Austrelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="245" phylum="Arthropoda" rank="species" species="davidi">
<emphasis id="B901EA95022CFF9A6769F8CBF3C31183" box="[550,643,1918,1941]" italics="true" pageId="2" pageNumber="245">A. davidi</emphasis>
(Wewalka, 2017)
</taxonomicName>
. However, 
<taxonomicName id="4C754D04022CFF9A6400F82BF0E511A3" authorityName="Hájek &amp; Shaverdo" authorityYear="2024" box="[335,421,1950,1973]" class="Insecta" family="Dytiscidae" genus="Austrelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="245" phylum="Arthropoda" rank="species" species="mirai" status="sp. nov.">
<emphasis id="B901EA95022CFF9A6400F82BF0E511A3" box="[335,421,1950,1973]" italics="true" pageId="2" pageNumber="245">A. mirai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23257EE022CFF9A64E4F82BF0BC11A3" box="[427,508,1950,1973]" pageId="2" pageNumber="245" rank="species">sp. nov.</taxonomicNameLabel>
can be recognised from those species based on more subparallel habitus, presence of long striolae on pronotum, and different shape of male genitalia. We assume, that the new species is most closely related to several, so far undescribed, 
<taxonomicName id="4C754D04022CFF9A619FF8B5F4DC1101" box="[1232,1436,1792,1815]" pageId="2" pageNumber="245">
<emphasis id="B901EA95022CFF9A619FF8B5F4091101" box="[1232,1353,1792,1815]" italics="true" pageId="2" pageNumber="245">Austrelatus</emphasis>
species
</taxonomicName>
from Borneo Island (J. Hájek et al., unpublished data).
</paragraph>
</subSubSection>
<subSubSection id="C36F650C022CFF9B6673F8F5F06A16E3" lastPageId="3" lastPageNumber="246" pageId="2" pageNumber="245" type="etymology">
<paragraph id="8BCA3687022CFF9A6673F8F5F5501181" blockId="2.[828,1437,1792,2071]" pageId="2" pageNumber="245">
<emphasis id="B901EA95022CFF9A6673F8F5F2FC1141" bold="true" box="[828,956,1856,1879]" pageId="2" pageNumber="245">Etymology.</emphasis>
The new species is dedicated to Miroslav Vít (Chyňava, 
<collectingCountry id="F3627617022CFF9A66FFF8D5F51E1161" box="[944,1118,1888,1911]" name="Czech Republic" pageId="2" pageNumber="245">Czech Republic</collectingCountry>
); the specific epithet is a noun in the genitive case.
</paragraph>
<paragraph id="8BCA3687022CFF9B6673F815F06A16E3" blockId="2.[828,1437,1792,2071]" lastBlockId="3.[150,759,190,341]" lastPageId="3" lastPageNumber="246" pageId="2" pageNumber="245">
<emphasis id="B901EA95022CFF9A6673F815F51F11A1" bold="true" box="[828,1119,1952,1975]" pageId="2" pageNumber="245">Collecting circumstances.</emphasis>
The species was collected in the ca. 
<quantity id="4C8D9B62022CFF9A66C4F875F29011C1" box="[907,976,1984,2007]" metricMagnitude="0" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="2" pageNumber="245" unit="m" value="2.5" valueMax="3.0" valueMin="2.0">2–3 m</quantity>
wide forest stream with bottom of granite stones and much sand between them and some leaf litter (
<figureCitation id="134E2A02022CFF9A660BF7B5F2C91E01" box="[836,905,2048,2071]" captionStart="Fig" captionStartId="3.[828,859,1092,1111]" captionTargetBox="[828,1436,189,1074]" captionTargetId="figure-691@3.[827,1436,189,1074]" captionTargetPageId="3" captionText="Fig. 7. Habitat of Austrelatus mirai sp. nov. in Templer Park Forest Reserve, Selangor, Malaysia." figureDoi="http://doi.org/10.5281/zenodo.14655595" httpUri="https://zenodo.org/record/14655595/files/figure.png" pageId="2" pageNumber="245">Fig. 7</figureCitation>
). The altitude was ca. 
<quantity id="4C8D9B62022CFF9A61C6F7B5F5901E01" box="[1161,1232,2048,2071]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="245" unit="m" value="200.0">200 m</quantity>
a.s.l.; there was a primary forest in the stream’s upper reaches (M. A. Jäch, pers. comm.).
</paragraph>
</subSubSection>
<subSubSection id="C36F650C022DFF9B65D9FF4BF19C1743" pageId="3" pageNumber="246" type="distribution">
<paragraph id="8BCA3687022DFF9B65D9FF4BF19C1743" blockId="3.[150,759,190,341]" pageId="3" pageNumber="246">
<emphasis id="B901EA95022DFF9B65D9FF4BF0691703" bold="true" box="[150,297,254,277]" pageId="3" pageNumber="246">Distribution.</emphasis>
<taxonomicName id="4C754D04022DFF9B647CFF4BF0AF1703" authorityName="Hájek &amp; Shaverdo" authorityYear="2024" box="[307,495,254,277]" class="Insecta" family="Dytiscidae" genus="Austrelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="246" phylum="Arthropoda" rank="species" species="mirai" status="sp. nov.">
<emphasis id="B901EA95022DFF9B647CFF4BF0AF1703" box="[307,495,254,277]" italics="true" pageId="3" pageNumber="246">Austrelatus mirai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23257EE022DFF9B64B6FF4BF3101703" box="[505,592,254,277]" pageId="3" pageNumber="246" rank="species">sp. nov.</taxonomicNameLabel>
is known only from the 
<typeStatus id="54CE8825022DFF9B65B4FEABF0691723" box="[251,297,286,309]" pageId="3" pageNumber="246">type</typeStatus>
locality in 
<collectingRegion id="49B1F865022DFF9B64E8FEABF3061723" box="[423,582,286,309]" country="Malaysia" name="Selangor" pageId="3" pageNumber="246">Selangor State</collectingRegion>
, Peninsular 
<collectingCountry id="F3627617022DFF9B6784FEABF1981743" name="Malaysia" pageId="3" pageNumber="246">Malaysia</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>