336 lines
36 KiB
XML
336 lines
36 KiB
XML
<document id="18111ACB861F16C1D49E947557330DA8" ID-CLB-Dataset="29854" ID-DOI="10.11646/zootaxa.4438.1.1" ID-GBIF-Dataset="3338d9d4-c5fa-4f77-89fb-901b1e028237" ID-ISSN="1175-5326" ID-Zenodo-Dep="1294121" ID-ZooBank="55BEB1B1-DA14-4CC4-92EB-1EA98AD6277C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1529535606996" checkinUser="plazi" docAuthor="Rakotonirina, Jean Claude & Fisher, Brian L." docDate="2018" docId="03C287BBE874FF9531A6489EFE2EF821" docLanguage="en" docName="zootaxa.4438.1.1.pdf.imf" docOrigin="Zootaxa 4438 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Camponotus lamosy Rakotonirina & Fisher 2018, sp. n." docType="treatment" docVersion="7" lastPageNumber="32" masterDocId="FFFBFFC3E86BFF8A31314822FFAAFFFA" masterDocTitle="Taxonomic revision of the Malagasy Camponotus subgenus Mayria (Hymenoptera, Formicidae) using qualitative and quantitative morphology" masterLastPageNumber="58" masterPageNumber="1" pageNumber="32" updateTime="1698550116491" updateUser="ExternalLinkService">
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<mods:title id="9F8BC35F9FDD2BCA0FBBCF51B12ECAC4">Taxonomic revision of the Malagasy Camponotus subgenus Mayria (Hymenoptera, Formicidae) using qualitative and quantitative morphology</mods:title>
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<mods:namePart id="17DA9B0012502242E989EAD29511239C">Rakotonirina, Jean Claude</mods:namePart>
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<mods:namePart id="9D3E76F7DFE19CAE62EE763855D6711E">Fisher, Brian L.</mods:namePart>
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<mods:typeOfResource id="2F2F5700EE4A9AC9A23DA7CEC337E1F8">text</mods:typeOfResource>
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<mods:title id="11234EB0AC6C0903641524F0D1168E20">Zootaxa</mods:title>
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<mods:part id="1AF487E24D2E31B38410227E18D3D7BA">
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<mods:date id="07CD7F9982D2648A41B74512A76BD494">2018</mods:date>
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<mods:number id="70E7BD94735E85DDC36D0DA696F7E526">2018-06-20</mods:number>
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<mods:number id="355F94D622C65A3B9B301307CCF672A9">4438</mods:number>
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<mods:identifier id="98A2B691E7FE6C0902C7B926F26D434D" type="DOI">10.11646/zootaxa.4438.1.1</mods:identifier>
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<treatment id="03C287BBE874FF9531A6489EFE2EF821" ID-DOI="http://doi.org/10.5281/zenodo.5987968" ID-GBIF-Taxon="146015938" ID-Zenodo-Dep="5987968" LSID="urn:lsid:plazi:treatment:03C287BBE874FF9531A6489EFE2EF821" httpUri="http://treatment.plazi.org/id/03C287BBE874FF9531A6489EFE2EF821" lastPageNumber="32" pageId="31" pageNumber="32">
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<subSubSection id="C3716526E874FF9531A6489EFE1DFF0D" pageId="31" pageNumber="32" type="nomenclature">
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<paragraph id="8BD436ADE874FF9531A6489EFD51FF2C" blockId="31.[151,763,188,247]" box="[151,763,188,214]" pageId="31" pageNumber="32">
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<heading id="D09C81C1E874FF9531A6489EFD51FF2C" bold="true" box="[151,763,188,214]" fontSize="11" level="1" pageId="31" pageNumber="32" reason="1">
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<emphasis id="B91FEABFE874FF9531A6489EFD51FF2C" bold="true" box="[151,763,188,214]" pageId="31" pageNumber="32">
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<taxonomicName id="4C6B4D2EE874FF9531A6489EFD18FF2C" authority="Rakotonirina & Fisher" authorityName="Rakotonirina & Fisher" authorityYear="2018" box="[151,690,188,214]" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="lamosy" status="sp. nov.">
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<emphasis id="B91FEABFE874FF9531A6489EFE26FF2C" bold="true" box="[151,396,188,214]" italics="true" pageId="31" pageNumber="32">Camponotus lamosy</emphasis>
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Rakotonirina & Fisher
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</taxonomicName>
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<taxonomicNameLabel id="A22C57C4E874FF953388489EFD51FF2C" box="[697,763,188,214]" pageId="31" pageNumber="32" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BD436ADE874FF9531A648FCFE1DFF0D" blockId="31.[151,763,188,247]" box="[151,439,222,247]" pageId="31" pageNumber="32">
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(
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<figureCitation id="13502A28E874FF9531AF48FCFEB4FF0D" box="[158,286,222,247]" captionStart="FIGURE 2" captionStartId="13.[151,250,1933,1955]" captionTargetBox="[151,1436,1402,1903]" captionTargetId="figure@13.[151,1436,1402,1911]" captionTargetPageId="13" captionText="FIGURE 2. Head in full-face view. A: C. lubbocki (CASENT0486998). B: C. lamosy (CASENT0408976)." httpUri="https://zenodo.org/record/1294125/files/figure.png" pageId="31" pageNumber="32">Figures 2B</figureCitation>
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,
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<figureCitation id="13502A28E874FF95301B48FDFEF6FF0D" box="[298,348,223,247]" captionStart="FIGURE 10" captionStartId="17.[151,250,657,679]" captionTargetBox="[152,1436,193,623]" captionTargetId="figure@17.[151,1436,193,635]" captionTargetPageId="17" captionText="FIGURE 10. Mesosoma in lateral view. A: C. lamosy (CASENT0408976). B: C. pulcher (CASENT0191639)." httpUri="https://zenodo.org/record/1294141/files/figure.png" pageId="31" pageNumber="32">10A</figureCitation>
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,
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<figureCitation id="13502A28E874FF95305948FDFE2CFF0D" box="[360,390,223,247]" captionStart="FIGURE 18" captionStartId="32.[151,250,1678,1700]" captionTargetBox="[152,1434,195,1639]" captionTargetId="figure@32.[151,1436,195,1656]" captionTargetPageId="32" captionText="FIGURE 18. Camponotus lamosy minor worker CASENT0408976. A: lateral view. B: head in full-face view. C: dorsal view." httpUri="https://zenodo.org/record/1294157/files/figure.png" pageId="31" pageNumber="32">18</figureCitation>
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,
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<figureCitation id="13502A28E874FF9530A348FDFE05FF0D" box="[402,431,223,247]" captionStart="FIGURES 21–26" captionStartId="37.[151,264,1889,1911]" captionTargetBox="[151,1436,303,1863]" captionTargetId="figure@37.[151,1436,303,1867]" captionTargetPageId="37" captionText="FIGURES 21–26. Distribution maps of Camponotus subgenus Mayria in Madagascar. Fig. 21: C. christi; Fig. 22: C. dromedarius; Fig. 23: C. foersteri; Fig. 24: C. lamosy; Fig. 25: C. liandia, Fig. 26: C. lubbocki." httpUri="https://zenodo.org/record/1294163/files/figure.png" pageId="31" pageNumber="32">24</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C3716526E874FF9531A64904FEE6FDC1" pageId="31" pageNumber="32" type="materials_examined">
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||
<paragraph id="8BD436ADE874FF9531A64904FDBEFE51" blockId="31.[151,1437,294,2011]" pageId="31" pageNumber="32">
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<materialsCitation id="3B033CF0E874FF9531A64904FDBAFE51" ID-GBIF-Occurrence="1851012821" collectingDate="2001-01-20" collectingDateMax="2001-01-26" collectingDateMin="2001-01-20" collectionCode="CASC" country="Madagascar" county="Parc National Montagne d'Ambre" elevation="925" location="Fisher" municipality="Beat" pageId="31" pageNumber="32" specimenCount="1" stateProvince="Antsiranana" typeStatus="holotype">
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<emphasis id="B91FEABFE874FF9531A64904FEC4FEC5" bold="true" box="[151,366,294,319]" pageId="31" pageNumber="32">
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||
<typeStatus id="54D0880FE874FF9531A64904FEAEFEC5" box="[151,260,294,319]" pageId="31" pageNumber="32" type="holotype">Holotype</typeStatus>
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worker.
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</emphasis>
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<collectingCountry id="F37C763DE874FF95304D4904FDA2FEC5" box="[380,520,294,319]" name="Madagascar" pageId="31" pageNumber="32">Madagascar</collectingCountry>
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: Province
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<collectingRegion id="49AFF84FE874FF9533BA4904FCBFFEC5" box="[651,789,294,319]" country="Madagascar" name="Antsiranana" pageId="31" pageNumber="32">Antsiranana</collectingRegion>
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,
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<collectingCounty id="62B54E21E874FF9532164905FB10FEC5" box="[807,1210,294,319]" pageId="31" pageNumber="32">Parc National Montagne d'Ambre</collectingCounty>
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,
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<locationDeviation id="9A662F8FE874FF9535FA4904FEA7FE99" pageId="31" pageNumber="32">
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<quantity id="4C939B48E874FF9535FA4904FA8BFEC5" box="[1227,1313,294,319]" metricMagnitude="3" metricUnit="m" metricValue="3.6" pageId="31" pageNumber="32" unit="km" value="3.6">3.6 km</quantity>
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235° SW Joffreville
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</locationDeviation>
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, -12.53444, 49.1795,
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<quantity id="4C939B48E874FF9533204968FDF5FE98" box="[529,607,330,355]" metricMagnitude="2" metricUnit="m" metricValue="9.25" pageId="31" pageNumber="32" unit="m" value="925.0">
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<elevation id="0046D19EE874FF9533204968FDF5FE98" box="[529,607,330,355]" metricMagnitude="2" metricUnit="m" metricValue="9.25" pageId="31" pageNumber="32" unit="m" value="925.0">925 m</elevation>
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</quantity>
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, transect subsample #08,
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<collectingMunicipality id="6BB0ACD7E874FF9532A54969FC6DFE98" box="[916,967,331,354]" pageId="31" pageNumber="32">Beat</collectingMunicipality>
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25 sample transect,
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||
<quantity id="4C939B48E874FF9535F34969FAA8FE98" box="[1218,1282,331,355]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="31" pageNumber="32" unit="m" value="10.0">
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||
<elevation id="0046D19EE874FF9535F34969FAA8FE98" box="[1218,1282,331,355]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="31" pageNumber="32" unit="m" value="10.0">10 m</elevation>
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</quantity>
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, beating low vegetation montane rainforest,
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<date id="FFD5106DE874FF9530CE494CFD12FE7D" box="[511,696,366,391]" pageId="31" pageNumber="32" value="2001-01-20" valueMax="2001-01-26" valueMin="2001-01-20">
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||
<collectingDate id="EF91E985E874FF9530CE494CFD12FE7D" box="[511,696,366,391]" pageId="31" pageNumber="32" value="2001-01-20" valueMax="2001-01-26" valueMin="2001-01-20">20–26 Jan 2001</collectingDate>
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||
</date>
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(
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<location id="8EB46076E874FF9533FA494CFCBEFE7D" LSID="urn:lsid:plazi:treatment:03C287BBE874FF9531A6489EFE2EF821:8EB46076E874FF9533FA494CFCBEFE7D" box="[715,788,366,391]" country="Madagascar" county="Parc National Montagne d'Ambre" municipality="Beat" name="Fisher" pageId="31" pageNumber="32" stateProvince="Antsiranana">Fisher</location>
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,
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<location id="8EB46076E874FF953212494CFC21FE7D" LSID="urn:lsid:plazi:treatment:03C287BBE874FF9531A6489EFE2EF821:8EB46076E874FF953212494CFC21FE7D" box="[803,907,366,391]" country="Madagascar" county="Parc National Montagne d'Ambre" municipality="Beat" name="Griswold" pageId="31" pageNumber="32" stateProvince="Antsiranana">Griswold</location>
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<emphasis id="B91FEABFE874FF9532A7494DFC78FE7D" box="[918,978,366,391]" italics="true" pageId="31" pageNumber="32">et al.</emphasis>
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) collection code: BLF02566, specimen code: CASENT0408976 (
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<collectionCode id="ED7AAE68E874FF95308949B1FDADFE51" box="[440,519,403,427]" pageId="31" pageNumber="32">CASC</collectionCode>
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)
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="8BD436ADE874FF9531F64994FEE6FDC1" blockId="31.[151,1437,294,2011]" pageId="31" pageNumber="32">
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<materialsCitation id="3B033CF0E874FF9531F64994FEE6FDC1" ID-GBIF-Occurrence="1851012827" collectingDate="2001-01-20" collectingDateMax="2001-01-26" collectingDateMin="2001-01-20" collectionCode="CASC, BMNH, MHNG, PBZT" country="Madagascar" county="Parc National Montagne d'Ambre" elevation="925" location="Fisher" municipality="Beat" pageId="31" pageNumber="32" specimenCount="11" specimenCount-worker="11" stateProvince="Antsiranana" typeStatus="paratype">
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<emphasis id="B91FEABFE874FF9531F64994FEECFE35" bold="true" box="[199,326,438,463]" pageId="31" pageNumber="32">
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<typeStatus id="54D0880FE874FF9531F64994FEEBFE35" box="[199,321,438,463]" pageId="31" pageNumber="32" type="paratype">Paratypes</typeStatus>
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.
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</emphasis>
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<specimenCount id="9D6DFD24E874FF95306B4995FE49FE35" box="[346,483,438,463]" pageId="31" pageNumber="32" type="worker">11 workers</specimenCount>
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with same data as holotype but with specimen codes: CASENT0408963, CASENT0408965, CASENT0408966, CASENT0408968, CASENT0408970, CASENT0408971, CASENT0408972, CASENT0408973, CASENT0408974, CASENT0408975, CASENT0408977 (
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<collectionCode id="ED7AAE68E874FF9535D949DDFA9FFDED" box="[1256,1333,511,535]" pageId="31" pageNumber="32">CASC</collectionCode>
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,
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<collectionCode id="ED7AAE68E874FF95347049DDFA32FDEC" box="[1345,1432,511,534]" pageId="31" pageNumber="32">BMNH</collectionCode>
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,
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<collectionCode id="ED7AAE68E874FF9531A64A01FF46FDC1" box="[151,236,547,571]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="31" pageNumber="32">MHNG</collectionCode>
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,
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<collectionCode id="ED7AAE68E874FF9531C64A01FEEAFDC1" box="[247,320,547,571]" country="Madagascar" httpUri="http://grbio.org/cool/a6tz-5rcd" name="Parc Botanique et Zoologique de Tsimbazaza" pageId="31" pageNumber="32">PBZT</collectionCode>
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).
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</materialsCitation>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C3716526E874FF9531F64A64FE5AFD31" pageId="31" pageNumber="32" type="diagnosis">
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||
<paragraph id="8BD436ADE874FF9531F64A64FE5AFD31" blockId="31.[151,1437,294,2011]" pageId="31" pageNumber="32">
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||
<emphasis id="B91FEABFE874FF9531F64A64FEEBFDA5" bold="true" box="[199,321,582,607]" pageId="31" pageNumber="32">Diagnosis.</emphasis>
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Clypeus without median longitudinal carina; mandible with six teeth; anteromedian margin of clypeus not lamellate; in lateral view, mesosoma short and high, its dorsal outline continuously arched in a domelike structure; propodeal declivity inclined posteriorly; in lateral view, propodeal dorsum slightly excised medially, petiolar node longer than high.
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C3716526E874FF9531F64AF4FDDFFAF1" pageId="31" pageNumber="32" type="description">
|
||
<paragraph id="8BD436ADE874FF9531F64AF4FB13FC11" blockId="31.[151,1437,294,2011]" pageId="31" pageNumber="32">
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<emphasis id="B91FEABFE874FF9531F64AF4FDB3FD15" bold="true" box="[199,537,726,751]" pageId="31" pageNumber="32">Description. Minor worker.</emphasis>
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In full-face view head longer than broad (CWb/CL: 0.77±0.03; 0.73–0.82), lateral margins roughly straight and slightly converging anteriorly and posteriorly; posterior margin more or less straight. Clypeus with straight anteromedian margin and rounded anterolateral angle. Eyes sometimes breaking the lateral outlines of the head, their posterior margins located well behind the mid-length of the head (PoOc/CL: 0.34±0.01; 0.31–0.37). Mandible triangular, apical margin with six sharp teeth. Antennal scape long, roughly its apical third surpassing posterior margin of head. In lateral view, outline of pronotum more or less straight and mesonotum forming a separate convexity; propodeum slightly concave anteriorly and weakly convex posteriorly; its junction to declivity angulate; declivity inclined posteriorly. Petiolar node longer than high.
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</paragraph>
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<paragraph id="8BD436ADE874FF9531F64BD5FEC3FB65" blockId="31.[151,1437,294,2011]" pageId="31" pageNumber="32">Dorsum of head and gastral segments with abundant erect filiform hairs; pronotum with 2 pairs, mesonotum, propodeum and petiolar node with one pair of erect hairs. Pubescence short and sparse on gastral segments. Head black, mesosoma, petiolar node, gaster and legs yellow-orange with black to dark brown spot on lower level of lateral surface of pronotum, propodeum and petiole. Coxa and trochanter whitish-yellow, base and apical portion of femur dark brown.</paragraph>
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<paragraph id="8BD436ADE874FF9531F64C88FDDFFAF1" blockId="31.[151,1437,294,2011]" pageId="31" pageNumber="32">
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||
<emphasis id="B91FEABFE874FF9531F64C88FED3FB39" bold="true" box="[199,377,1194,1219]" pageId="31" pageNumber="32">Major worker.</emphasis>
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Differing from minor worker in having larger heart-shaped head, apical fourth of antennal scape surpassing posterior cephalic margin; robust mesosoma with distinct metanotum, and as high as long petiolar node. Other characters as in minor worker.
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C3716526E874FF9531F64D34FCC6FA44" pageId="31" pageNumber="32" type="discussion">
|
||
<paragraph id="8BD436ADE874FF9531F64D34FD8AFAA9" blockId="31.[151,1437,294,2011]" pageId="31" pageNumber="32">
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<emphasis id="B91FEABFE874FF9531F64D34FEE6FAD5" bold="true" box="[199,332,1302,1327]" pageId="31" pageNumber="32">Discussion.</emphasis>
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<taxonomicName id="4C6B4D2EE874FF9530654D35FD93FAD5" authorityName="Rakotonirina & Fisher" authorityYear="2018" box="[340,569,1302,1327]" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="lamosy">
|
||
<emphasis id="B91FEABFE874FF9530654D35FD93FAD5" box="[340,569,1302,1327]" italics="true" pageId="31" pageNumber="32">Camponotus lamosy</emphasis>
|
||
</taxonomicName>
|
||
is similar to
|
||
<taxonomicName id="4C6B4D2EE874FF9533E04D35FC2BFAD5" box="[721,897,1302,1327]" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="dromedarius">
|
||
<emphasis id="B91FEABFE874FF9533E04D35FC2BFAD5" box="[721,897,1302,1327]" italics="true" pageId="31" pageNumber="32">C. dromedarius</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C6B4D2EE874FF95328A4D35FB9BFAD5" box="[955,1073,1302,1327]" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="pulcher">
|
||
<emphasis id="B91FEABFE874FF95328A4D35FB9BFAD5" box="[955,1073,1302,1327]" italics="true" pageId="31" pageNumber="32">C. pulcher</emphasis>
|
||
</taxonomicName>
|
||
, but can be distinguished by its slightly excised propodeal dorsum.
|
||
</paragraph>
|
||
<paragraph id="8BD436ADE874FF9531F64D7CFCC6FA44" blockId="31.[151,1437,294,2011]" pageId="31" pageNumber="32">
|
||
The definition of
|
||
<taxonomicName id="4C6B4D2EE874FF95309C4D7DFD36FA8D" authorityName="Rakotonirina & Fisher" authorityYear="2018" box="[429,668,1374,1399]" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="lamosy">
|
||
<emphasis id="B91FEABFE874FF95309C4D7DFD36FA8D" box="[429,668,1374,1399]" italics="true" pageId="31" pageNumber="32">Camponotus lamosy</emphasis>
|
||
</taxonomicName>
|
||
based on qualitative morphological analysis and multivariate morphometry is congruent. Recognition of the species is supported by the clustering shown by the dendrogram (
|
||
<figureCitation id="13502A28E874FF9531AE4D84FF4AFA45" box="[159,224,1446,1471]" captionStart="FIGURE 1" captionStartId="12.[151,250,1952,1974]" captionTargetBox="[503,1067,196,1920]" captionTargetId="figure@12.[501,1070,193,1921]" captionTargetPageId="12" captionText="FIGURE 1. Dendrogram of NC-PART-clustering of the 14 species recognized in the Camponotus subgenus Mayria. Color bars represent the species hypothesis generated by PART using “hclust” and “kmeans” clustering methods. The corresponding label on the tip of the branch (the species name followed by the collection code) is given the same color code." httpUri="https://zenodo.org/record/1294123/files/figure.png" pageId="31" pageNumber="32">Fig. 1</figureCitation>
|
||
) and confirmed by LDA at a 100% classification success.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3716526E874FF9531F64DE8FBE2F9B5" pageId="31" pageNumber="32" type="distribution">
|
||
<paragraph id="8BD436ADE874FF9531F64DE8FBE2F9B5" blockId="31.[151,1437,294,2011]" pageId="31" pageNumber="32">
|
||
<emphasis id="B91FEABFE874FF9531F64DE8FE5BFA19" bold="true" box="[199,497,1482,1507]" pageId="31" pageNumber="32">Distribution and biology.</emphasis>
|
||
<taxonomicName id="4C6B4D2EE874FF9530C94DE9FD71FA19" authorityName="Rakotonirina & Fisher" authorityYear="2018" box="[504,731,1482,1507]" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="lamosy">
|
||
<emphasis id="B91FEABFE874FF9530C94DE9FD71FA19" box="[504,731,1482,1507]" italics="true" pageId="31" pageNumber="32">Camponotus lamosy</emphasis>
|
||
</taxonomicName>
|
||
occupies the littoral forest habitats of the north, the rainforests of the northeast, and the montane rainforests of the central high plateau and the southeast of
|
||
<collectingCountry id="F37C763DE874FF9535944DCCFA85F9FD" box="[1189,1327,1518,1543]" name="Madagascar" pageId="31" pageNumber="32">Madagascar</collectingCountry>
|
||
. Endemic to the island, the species is found foraging both on the ground and through the leaf litter or on the lower portions of vegetation, and its nests are located in rotten logs or in dead branches above ground.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3716526E874FF9531F64E78FE2EF821" pageId="31" pageNumber="32" type="materials_examined">
|
||
<paragraph id="8BD436ADE874FF9531F64E78FE2EF821" blockId="31.[151,1437,294,2011]" pageId="31" pageNumber="32">
|
||
<materialsCitation id="3B033CF0E874FF9531F64E78FD57F96D" ID-GBIF-Occurrence="1851012766" collectionCode="CASC" collectorName="B. L. Fisher" elevation="875" location="Reserve Anjanaharibe-Sud" pageId="31" pageNumber="32" specimenCount="1" stateProvince="Antsiranana">
|
||
<emphasis id="B91FEABFE874FF9531F64E78FD9EF989" bold="true" box="[199,564,1626,1651]" pageId="31" pageNumber="32">Additional material examined.</emphasis>
|
||
Province
|
||
<emphasis id="B91FEABFE874FF9533974E78FCEEF989" bold="true" box="[678,836,1626,1651]" pageId="31" pageNumber="32">
|
||
<collectingRegion id="49AFF84FE874FF9533974E78FC95F989" box="[678,831,1626,1651]" country="Madagascar" name="Antsiranana" pageId="31" pageNumber="32">Antsiranana</collectingRegion>
|
||
:
|
||
</emphasis>
|
||
<locationDeviation id="9A662F8FE874FF95327D4E78FBF7F989" box="[844,1117,1626,1651]" pageId="31" pageNumber="32">
|
||
<quantity id="4C939B48E874FF95327D4E78FC30F989" box="[844,922,1626,1651]" metricMagnitude="3" metricUnit="m" metricValue="6.5" pageId="31" pageNumber="32" unit="km" value="6.5">6.5 km</quantity>
|
||
SSW Befingotra
|
||
</locationDeviation>
|
||
,
|
||
<location id="8EB46076E874FF9535594E79FA32F989" LSID="urn:lsid:plazi:treatment:03C287BBE874FF9531A6489EFE2EF821:8EB46076E874FF9535594E79FA32F989" box="[1128,1432,1626,1651]" name="Reserve Anjanaharibe-Sud" pageId="31" pageNumber="32" stateProvince="Antsiranana">Réserve Anjanaharibe-Sud</location>
|
||
,
|
||
<quantity id="4C939B48E874FF9531A64E5CFF48F96C" box="[151,226,1662,1687]" metricMagnitude="2" metricUnit="m" metricValue="8.75" pageId="31" pageNumber="32" unit="m" value="875.0">
|
||
<elevation id="0046D19EE874FF9531A64E5CFF48F96C" box="[151,226,1662,1687]" metricMagnitude="2" metricUnit="m" metricValue="8.75" pageId="31" pageNumber="32" unit="m" value="875.0">875 m</elevation>
|
||
</quantity>
|
||
, -14.75, 49.5, rainforest, (
|
||
<collectorName id="269E537BE874FF95333D4E5DFD24F96D" box="[524,654,1662,1687]" pageId="31" pageNumber="32">B.L. Fisher</collectorName>
|
||
) (
|
||
<collectionCode id="ED7AAE68E874FF9533974E5DFD5FF96D" box="[678,757,1663,1687]" pageId="31" pageNumber="32">CASC</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B033CF0E874FF95323A4E5CFD15F941" ID-GBIF-Occurrence="1851012776" collectionCode="CASC" collectorName="B. L. Fisher" elevation="880" location="Bekona River" municipality="Betaolana Forest" pageId="31" pageNumber="32" specimenCount="1" stateProvince="Antsiranana">
|
||
<collectingMunicipality id="6BB0ACD7E874FF95323A4E5CFC66F96C" box="[779,972,1662,1687]" pageId="31" pageNumber="32">Betaolana Forest</collectingMunicipality>
|
||
, along
|
||
<location id="8EB46076E874FF9535114E5CFB14F96D" LSID="urn:lsid:plazi:treatment:03C287BBE874FF9531A6489EFE2EF821:8EB46076E874FF9535114E5CFB14F96D" box="[1056,1214,1662,1687]" municipality="Betaolana Forest" name="Bekona River" pageId="31" pageNumber="32" stateProvince="Antsiranana">Bekona River</location>
|
||
,
|
||
<quantity id="4C939B48E874FF9535FA4E5DFABFF96C" box="[1227,1301,1663,1687]" metricMagnitude="2" metricUnit="m" metricValue="8.8" pageId="31" pageNumber="32" unit="m" value="880.0">
|
||
<elevation id="0046D19EE874FF9535FA4E5DFABFF96C" box="[1227,1301,1663,1687]" metricMagnitude="2" metricUnit="m" metricValue="8.8" pageId="31" pageNumber="32" unit="m" value="880.0">880 m</elevation>
|
||
</quantity>
|
||
, -14.52996, 49.44039, rainforest, (
|
||
<collectorName id="269E537BE874FF9530A04E81FDBAF941" box="[401,528,1698,1723]" pageId="31" pageNumber="32">B.L. Fisher</collectorName>
|
||
<emphasis id="B91FEABFE874FF9533264E86FDFAF941" box="[535,592,1698,1723]" italics="true" pageId="31" pageNumber="32">et al.</emphasis>
|
||
) (
|
||
<collectionCode id="ED7AAE68E874FF9533594E81FD1DF941" box="[616,695,1699,1723]" pageId="31" pageNumber="32">CASC</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B033CF0E874FF9533FA4E81FB5CF925" ID-GBIF-Occurrence="1851012757" collectionCode="CASC" collectorName="B. L. Fisher" elevation="775" location="Manantenina River" municipality="Parc National de Marojejy" pageId="31" pageNumber="32" specimenCount="1" stateProvince="Antsiranana">
|
||
<collectingMunicipality id="6BB0ACD7E874FF9533FA4E81FC5FF941" box="[715,1013,1698,1723]" pageId="31" pageNumber="32">Parc National de Marojejy</collectingMunicipality>
|
||
,
|
||
<location id="8EB46076E874FF9535314E80FB70F941" LSID="urn:lsid:plazi:treatment:03C287BBE874FF9531A6489EFE2EF821:8EB46076E874FF9535314E80FB70F941" box="[1024,1242,1698,1723]" municipality="Parc National de Marojejy" name="Manantenina River" pageId="31" pageNumber="32" stateProvince="Antsiranana">Manantenina River</location>
|
||
,
|
||
<quantity id="4C939B48E874FF9535D44E80FAEBF941" box="[1253,1345,1698,1723]" metricMagnitude="4" metricUnit="m" metricValue="2.76" pageId="31" pageNumber="32" unit="km" value="27.6">27.6 km</quantity>
|
||
35° NE Andapa,
|
||
<quantity id="4C939B48E874FF9531CC4EE4FEE1F925" box="[253,331,1734,1759]" metricMagnitude="3" metricUnit="m" metricValue="9.6" pageId="31" pageNumber="32" unit="km" value="9.6">9.6 km</quantity>
|
||
327° NNW Manantenina,
|
||
<quantity id="4C939B48E874FF95334A4EE4FD6EF924" box="[635,708,1734,1759]" metricMagnitude="2" metricUnit="m" metricValue="7.75" pageId="31" pageNumber="32" unit="m" value="775.0">
|
||
<elevation id="0046D19EE874FF95334A4EE4FD6EF924" box="[635,708,1734,1759]" metricMagnitude="2" metricUnit="m" metricValue="7.75" pageId="31" pageNumber="32" unit="m" value="775.0">775 m</elevation>
|
||
</quantity>
|
||
, -14.435, 49.76, rainforest, (
|
||
<collectorName id="269E537BE874FF9535364EE5FB22F925" box="[1031,1160,1734,1759]" pageId="31" pageNumber="32">B.L. Fisher</collectorName>
|
||
) (
|
||
<collectionCode id="ED7AAE68E874FF9535AE4EE5FB44F925" box="[1183,1262,1735,1759]" pageId="31" pageNumber="32">CASC</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B033CF0E874FF9534334EE5FB5DF8F9" ID-GBIF-Occurrence="1851012677" collectionCode="CASC" collectorName="Joy Boutin" elevation="925" location="Parc National Montagne d'Ambre" pageId="31" pageNumber="32" specimenCount="1" stateProvince="Antsiranana">
|
||
<location id="8EB46076E874FF9534334EE5FEDEF8F9" LSID="urn:lsid:plazi:treatment:03C287BBE874FF9531A6489EFE2EF821:8EB46076E874FF9534334EE5FEDEF8F9" name="Parc National Montagne d'Ambre" pageId="31" pageNumber="32" stateProvince="Antsiranana">Parc National Montagne d'Ambre</location>
|
||
,
|
||
<locationDeviation id="9A662F8FE874FF95304E4EC8FD12F8F9" box="[383,696,1770,1795]" pageId="31" pageNumber="32">
|
||
<quantity id="4C939B48E874FF95304E4EC8FE67F8F9" box="[383,461,1770,1795]" metricMagnitude="3" metricUnit="m" metricValue="3.6" pageId="31" pageNumber="32" unit="km" value="3.6">3.6 km</quantity>
|
||
235° SW Joffreville
|
||
</locationDeviation>
|
||
,
|
||
<quantity id="4C939B48E874FF9533F24EC8FCA7F8F8" box="[707,781,1770,1795]" metricMagnitude="2" metricUnit="m" metricValue="9.25" pageId="31" pageNumber="32" unit="m" value="925.0">
|
||
<elevation id="0046D19EE874FF9533F24EC8FCA7F8F8" box="[707,781,1770,1795]" metricMagnitude="2" metricUnit="m" metricValue="9.25" pageId="31" pageNumber="32" unit="m" value="925.0">925 m</elevation>
|
||
</quantity>
|
||
, -12.53444, 49.1795, (
|
||
<collectorName id="269E537BE874FF95353A4EC9FB23F8F9" box="[1035,1161,1770,1795]" pageId="31" pageNumber="32">Joy Boutin</collectorName>
|
||
) (
|
||
<collectionCode id="ED7AAE68E874FF9535914EC9FB45F8F9" box="[1184,1263,1771,1795]" pageId="31" pageNumber="32">CASC</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
; Parc National Montagne d'Ambre, Crête,
|
||
<quantity id="4C939B48E874FF9530F94F2DFDB7F8DC" box="[456,541,1807,1831]" metricMagnitude="3" metricUnit="m" metricValue="1.1099999999999999" pageId="31" pageNumber="32" unit="m" value="1110.0">1110 m</quantity>
|
||
, -12.58132, 49.13368, montane rainforest, (B.L. Fisher
|
||
<emphasis id="B91FEABFE874FF9535BE4F2DFB6DF8DD" box="[1167,1223,1806,1831]" italics="true" pageId="31" pageNumber="32">et al.</emphasis>
|
||
) (CASC);
|
||
<materialsCitation id="3B033CF0E874FF9534734F2DFED8F895" ID-GBIF-Occurrence="1851012756" collectionCode="CASC" elevation="40" location="3 km W Sakalava Beach" pageId="31" pageNumber="32" specimenCount="1" stateProvince="Antsiranana">
|
||
<location id="8EB46076E874FF9534734F2DFEE1F8B1" LSID="urn:lsid:plazi:treatment:03C287BBE874FF9531A6489EFE2EF821:8EB46076E874FF9534734F2DFEE1F8B1" name="3 km W Sakalava Beach" pageId="31" pageNumber="32" stateProvince="Antsiranana">
|
||
<quantity id="4C939B48E874FF9534734F2DFAD0F8DD" box="[1346,1402,1806,1831]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="31" pageNumber="32" unit="km" value="3.0">3 km</quantity>
|
||
W Sakalava Beach
|
||
</location>
|
||
[white dunes site],
|
||
<quantity id="4C939B48E874FF95331F4F11FDC1F8B0" box="[558,619,1843,1867]" metricMagnitude="1" metricUnit="m" metricValue="4.0" pageId="31" pageNumber="32" unit="m" value="40.0">
|
||
<elevation id="0046D19EE874FF95331F4F11FDC1F8B0" box="[558,619,1843,1867]" metricMagnitude="1" metricUnit="m" metricValue="4.0" pageId="31" pageNumber="32" unit="m" value="40.0">40 m</elevation>
|
||
</quantity>
|
||
, -12.28617, 49.36667, white dunes in littoral forest, (Harin'Hala, Irwin, Schlinger) (
|
||
<collectionCode id="ED7AAE68E874FF95302A4F75FEC0F895" box="[283,362,1879,1903]" pageId="31" pageNumber="32">CASC</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B033CF0E874FF95304F4F75FF5CF869" ID-GBIF-Occurrence="1851012768" collectionCode="CASC" collectorName="R. Harin'Hala" elevation="360" location="Fisher" pageId="31" pageNumber="32" specimenCount="1" stateProvince="Antsiranana">
|
||
<locationDeviation id="9A662F8FE874FF95304F4F75FDE7F895" box="[382,589,1878,1903]" pageId="31" pageNumber="32">
|
||
<quantity id="4C939B48E874FF95304F4F75FE1DF895" box="[382,439,1878,1903]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="31" pageNumber="32" unit="km" value="7.0">7 km</quantity>
|
||
N Joffreville
|
||
</locationDeviation>
|
||
[camp 2 of
|
||
<location id="8EB46076E874FF9533E44F74FC8BF895" LSID="urn:lsid:plazi:treatment:03C287BBE874FF9531A6489EFE2EF821:8EB46076E874FF9533E44F74FC8BF895" box="[725,801,1878,1903]" name="Fisher" pageId="31" pageNumber="32" stateProvince="Antsiranana">Fisher</location>
|
||
],
|
||
<quantity id="4C939B48E874FF9532024F74FCD6F894" box="[819,892,1878,1903]" metricMagnitude="2" metricUnit="m" metricValue="3.6" pageId="31" pageNumber="32" unit="m" value="360.0">
|
||
<elevation id="0046D19EE874FF9532024F74FCD6F894" box="[819,892,1878,1903]" metricMagnitude="2" metricUnit="m" metricValue="3.6" pageId="31" pageNumber="32" unit="m" value="360.0">360 m</elevation>
|
||
</quantity>
|
||
, -12.33333, 49.25, in dry forest, (
|
||
<collectorName id="269E537BE874FF9535C64F75FA3FF895" box="[1271,1429,1878,1903]" pageId="31" pageNumber="32">R. Harin'Hala</collectorName>
|
||
) (
|
||
<collectionCode id="ED7AAE68E874FF9531AE4F59FF44F869" box="[159,238,1915,1939]" pageId="31" pageNumber="32">CASC</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B033CF0E874FF9530304F59FF5CF84D" ID-GBIF-Occurrence="1851012674" collectionCode="CASC" collectorName="R. Harin'Hala" elevation="960" location="Parc National Montagne d'Ambre" pageId="31" pageNumber="32" specimenCount="1" stateProvince="Antsiranana">
|
||
<location id="8EB46076E874FF9530304F59FDD0F869" LSID="urn:lsid:plazi:treatment:03C287BBE874FF9531A6489EFE2EF821:8EB46076E874FF9530304F59FDD0F869" box="[257,634,1914,1939]" name="Parc National Montagne d'Ambre" pageId="31" pageNumber="32" stateProvince="Antsiranana">Parc National Montagne d'Ambre</location>
|
||
[1
|
||
<taxonomicName id="4C6B4D2EE874FF9533A24F59FCB0F869" box="[659,794,1914,1939]" pageId="31" pageNumber="32" rank="subSpecies" subSpecies="campsite">st campsite</taxonomicName>
|
||
],
|
||
<quantity id="4C939B48E874FF95321D4F58FCDEF868" box="[812,884,1914,1939]" metricMagnitude="2" metricUnit="m" metricValue="9.6" pageId="31" pageNumber="32" unit="m" value="960.0">
|
||
<elevation id="0046D19EE874FF95321D4F58FCDEF868" box="[812,884,1914,1939]" metricMagnitude="2" metricUnit="m" metricValue="9.6" pageId="31" pageNumber="32" unit="m" value="960.0">960 m</elevation>
|
||
</quantity>
|
||
, -12.51444, 49.18139, rainforest, (
|
||
<collectorName id="269E537BE874FF9535C94F59FA3FF869" box="[1272,1429,1914,1939]" pageId="31" pageNumber="32">R. Harin'Hala</collectorName>
|
||
) (
|
||
<collectionCode id="ED7AAE68E874FF9531AE4FBDFF44F84D" box="[159,238,1951,1975]" pageId="31" pageNumber="32">CASC</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B033CF0E874FF9530384FBDFE2AF821" ID-GBIF-Occurrence="1851012767" collectionCode="CASC" collectorName="R. Harin'Hala" elevation="1125" location="Petit Lac" municipality="Parc National Montagne d'Ambre" pageId="31" pageNumber="32" specimenCount="1" stateProvince="Antsiranana">
|
||
<collectingMunicipality id="6BB0ACD7E874FF9530384FBDFD31F84D" box="[265,667,1950,1975]" pageId="31" pageNumber="32">Parc National Montagne d'Ambre</collectingMunicipality>
|
||
[
|
||
<location id="8EB46076E874FF9533814FBCFCB5F84C" LSID="urn:lsid:plazi:treatment:03C287BBE874FF9531A6489EFE2EF821:8EB46076E874FF9533814FBCFCB5F84C" box="[688,799,1950,1975]" municipality="Parc National Montagne d'Ambre" name="Petit Lac" pageId="31" pageNumber="32" stateProvince="Antsiranana">Petit Lac</location>
|
||
road],
|
||
<quantity id="4C939B48E874FF95324E4FBCFC77F84C" box="[895,989,1950,1975]" metricMagnitude="3" metricUnit="m" metricValue="1.125" pageId="31" pageNumber="32" unit="m" value="1125.0">
|
||
<elevation id="0046D19EE874FF95324E4FBCFC77F84C" box="[895,989,1950,1975]" metricMagnitude="3" metricUnit="m" metricValue="1.125" pageId="31" pageNumber="32" unit="m" value="1125.0">1125 m</elevation>
|
||
</quantity>
|
||
, -12.52028, 49.17917, rainforest, (
|
||
<collectorName id="269E537BE874FF9534B34FBDFEB9F821" pageId="31" pageNumber="32">R. Harin'Hala</collectorName>
|
||
) (
|
||
<collectionCode id="ED7AAE68E874FF9530184FE1FED2F821" box="[297,376,1987,2011]" pageId="31" pageNumber="32">CASC</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |