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<document id="A57AF4D7526C958B654623989D209149" ID-CLB-Dataset="26953" ID-DOI="10.11646/zootaxa.4590.3.2" ID-GBIF-Dataset="5ed08b95-646d-41d0-bcf6-54ff44603e27" ID-ISSN="1175-5326" ID-Zenodo-Dep="2653404" ID-ZooBank="7FEE7CB2-9A74-407A-A9DB-DB714FEF1E0F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1556520044797" checkinUser="plazi" docAuthor="Branch, William R., Baptista, Ninda, Keates, Chad &amp; Edwards, Shelley" docDate="2019" docId="03BD7406FFDAFFB106ADFDD8556543F0" docLanguage="en" docName="zootaxa.4590.3.2.pdf" docOrigin="Zootaxa 4590 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Psammophylax ocellatus , Bocage 1873" docType="treatment" docVersion="9" lastPageNumber="357" masterDocId="FF840C7EFFD4FFA1063AFF9C54644324" masterDocTitle="Rediscovery, taxonomic status, and phylogenetic relationships of two rare and endemic snakes (Serpentes: Psammophiinae) from the southwestern Angolan plateau" masterLastPageNumber="366" masterPageNumber="342" pageNumber="356" updateTime="1698724313275" updateUser="ExternalLinkService">
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<mods:title id="58F1FE91498E3764AD9E73D30270EC07">Rediscovery, taxonomic status, and phylogenetic relationships of two rare and endemic snakes (Serpentes: Psammophiinae) from the southwestern Angolan plateau</mods:title>
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<mods:namePart id="754060BF6FF9B3524EA52CE1238C958D">Branch, William R.</mods:namePart>
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<mods:namePart id="91DD4DF5579285E6378584E6ED645836">Baptista, Ninda</mods:namePart>
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<mods:namePart id="3A4A5C86A7BFC7389AD8B6EA876B4F6C">Keates, Chad</mods:namePart>
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<mods:namePart id="CED4F148DE83ECF84237EE1E6D8EEC9F">Edwards, Shelley</mods:namePart>
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<mods:title id="2BC2287584B323243E009AD26E12E913">Zootaxa</mods:title>
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<mods:date id="61497E50A0124C68A23ED1B97B5FE787">2019</mods:date>
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<mods:number id="2D7DD7920739E5B94223707F9C6D88B2">2019-04-29</mods:number>
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<mods:number id="290378E7A03DAE95F879E41804A3B89B">4590</mods:number>
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<treatment id="03BD7406FFDAFFB106ADFDD8556543F0" ID-GBIF-Taxon="156872437" LSID="urn:lsid:plazi:treatment:03BD7406FFDAFFB106ADFDD8556543F0" httpUri="http://treatment.plazi.org/id/03BD7406FFDAFFB106ADFDD8556543F0" lastPageId="16" lastPageNumber="357" pageId="14" pageNumber="356">
<subSubSection id="C30E969BFFDAFFAF06ADFDD855E141A4" pageId="14" pageNumber="356" type="nomenclature">
<paragraph id="8BABC510FFDAFFAF06ADFDD85609417B" blockId="14.[151,621,580,640]" box="[151,621,580,607]" pageId="14" pageNumber="356">
<heading id="D0E3727CFFDAFFAF06ADFDD85609417B" bold="true" box="[151,621,580,607]" fontSize="11" level="1" pageId="14" pageNumber="356" reason="1">
<taxonomicName id="4C14BE93FFDAFFAF06ADFDD85609417B" authority="Bocage, 1873" authorityName=", Bocage" authorityYear="1873" box="[151,621,580,607]" class="Reptilia" family="Lamprophiidae" genus="Psammophylax" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="356" phylum="Chordata" rank="species" species="ocellatus">
<emphasis id="B9601902FFDAFFAF06ADFDD85609417B" bold="true" box="[151,621,580,607]" pageId="14" pageNumber="356">
<emphasis id="B9601902FFDAFFAF06ADFDD855A0417A" bold="true" box="[151,452,580,606]" italics="true" pageId="14" pageNumber="356">Psammophylax ocellatus</emphasis>
<bibRefCitation id="EF85B8E1FFDAFFAF07F0FDD95609417B" author="Bocage, J. V. B." box="[458,621,581,607]" pageId="14" pageNumber="356" pagination="209 - 227" refId="ref10900" refString="Bocage, J. V. B. (1873) Melanges erpetologiques. II. Sur quelques reptiles et batraciens nouveaux, rares ou peu connus d'Afrique occidentale. Jornal de Sciencias, Mathematicas, Physicas e Naturaes, Lisboa, 4, 209 - 227." type="journal article" year="1873">Bocage, 1873</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BABC510FFDAFFAF06ADFDFB55E141A4" blockId="14.[151,621,580,640]" box="[151,389,615,640]" pageId="14" pageNumber="356">Angolan Skaapsteker</paragraph>
</subSubSection>
<subSubSection id="C30E969BFFDAFFAF06ADFD3250AD41C7" pageId="14" pageNumber="356" type="reference_group">
<paragraph id="8BABC510FFDAFFAF06ADFD3250AD41C7" blockId="14.[151,1435,686,739]" pageId="14" pageNumber="356">
<taxonomicName id="4C14BE93FFDAFFAF06ADFD32565541E0" authority="Bocage 1873" authorityName=", Bocage" authorityYear="1873" box="[151,561,686,708]" class="Reptilia" family="Lamprophiidae" genus="Psammophylax" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="356" phylum="Chordata" rank="species" species="ocellatus">
<emphasis id="B9601902FFDAFFAF06ADFD3255FC41E0" box="[151,408,686,708]" italics="true" pageId="14" pageNumber="356">Psammophylax ocellatus</emphasis>
<bibRefCitation id="EF85B8E1FFDAFFAF079FFD33565541E0" author="Bocage, J. V. B." box="[421,561,686,708]" pageId="14" pageNumber="356" pagination="209 - 227" refId="ref10900" refString="Bocage, J. V. B. (1873) Melanges erpetologiques. II. Sur quelques reptiles et batraciens nouveaux, rares ou peu connus d'Afrique occidentale. Jornal de Sciencias, Mathematicas, Physicas e Naturaes, Lisboa, 4, 209 - 227." type="journal article" year="1873">Bocage 1873</bibRefCitation>
</taxonomicName>
, 221. Bocage, J.V.B. 1873. Melanges erpetologiques. II. Sur quelques reptiles et batraciens nouveaux, rares ou peu connus d‘Afrique occidentale. Jorn. Acad. Sci.
<collectingRegion id="49D00BF2FFDAFFAF023EFD51502D41C7" box="[1028,1097,717,739]" country="Portugal" name="Lisboa" pageId="14" pageNumber="356">Lisboa</collectingRegion>
4: 209–227.
</paragraph>
</subSubSection>
<subSubSection id="C30E969BFFDAFFAF06ADFC93557C4790" pageId="14" pageNumber="356" type="materials_examined">
<paragraph id="8BABC510FFDAFFAF06ADFC93568E4724" blockId="14.[151,1436,783,1133]" pageId="14" pageNumber="356">
<emphasis id="B9601902FFDAFFAF06ADFC93555E4003" bold="true" box="[151,314,783,808]" pageId="14" pageNumber="356">Type locality:</emphasis>
“de l’interieur de Mossamedes (
<emphasis id="B9601902FFDAFFAF0494FC93576F400C" box="[686,779,783,808]" italics="true" pageId="14" pageNumber="356">Gambos</emphasis>
) par M. d’Anchieta“ (
<bibRefCitation id="EF85B8E1FFDAFFAF0230FC8C50CE400C" author="Bocage, J. V. B." box="[1034,1194,784,808]" pageId="14" pageNumber="356" pagination="209 - 227" refId="ref10900" refString="Bocage, J. V. B. (1873) Melanges erpetologiques. II. Sur quelques reptiles et batraciens nouveaux, rares ou peu connus d'Afrique occidentale. Jornal de Sciencias, Mathematicas, Physicas e Naturaes, Lisboa, 4, 209 - 227." type="journal article" year="1873">Bocage, 1873</bibRefCitation>
). Crawford-Cabral &amp; Mesquitela (1989) updated this to “Chibemba,
<collectingRegion id="49D00BF2FFDAFFAF0492FCA857364069" box="[680,850,820,845]" country="Angola" name="Huila" pageId="14" pageNumber="356">Huíla Province</collectingRegion>
(
<geoCoordinate id="EE20A3D7FFDAFFAF0558FCA857D84069" box="[866,956,820,845]" direction="south" orientation="latitude" pageId="14" pageNumber="356" precision="925" value="-15.766666">15°46'S</geoCoordinate>
, 14°06';
<quantity id="4CEC68F5FFDAFFAF021AFCA850114068" box="[1056,1141,820,845]" metricMagnitude="3" metricUnit="m" metricValue="1.316" pageId="14" pageNumber="356" unit="m" value="1316.0">1316 m</quantity>
a.s.l)”, which was refined without comment to “Chibemba,
<collectingRegion id="49D00BF2FFDAFFAF042FFCCB56A74054" box="[533,707,855,880]" country="Angola" name="Huila" pageId="14" pageNumber="356">Huíla Province</collectingRegion>
;
<geoCoordinate id="EE20A3D7FFDAFFAF04EBFCCB57104054" box="[721,884,855,880]" direction="south" orientation="latitude" pageId="14" pageNumber="356" precision="1" value="-15.753105">15°45'11.18&quot;S</geoCoordinate>
,
<geoCoordinate id="EE20A3D7FFDAFFAF05B8FCCB504E4054" box="[898,1066,855,880]" direction="east" orientation="longitude" pageId="14" pageNumber="356" precision="1" value="14.07545">14°04'31.62&quot;E</geoCoordinate>
;
<quantity id="4CEC68F5FFDAFFAF020DFCC450EA404B" box="[1079,1166,856,880]" metricMagnitude="3" metricUnit="m" metricValue="1.321" pageId="14" pageNumber="356" unit="m" value="1321.0">1321 m</quantity>
a.s.l” by
<bibRefCitation id="EF85B8E1FFDAFFAF02C4FCCB548140B0" author="Wallach, V. &amp; Williams, K. L. &amp; Boundy, J." pageId="14" pageNumber="356" refId="ref13645" refString="Wallach, V., Williams, K. L. &amp; Boundy, J. (2014) Snakes of the World: A Catalogue of Living and Extinct Species. Taylor and Francis, CRC Press, Florida, 1237 pp. https: // doi. org / 10.1201 / b 16901" type="book" year="2014">
Wallach
<emphasis id="B9601902FFDAFFAF0358FCC551F84054" box="[1378,1436,855,880]" italics="true" pageId="14" pageNumber="356">et al.</emphasis>
(2014)
</bibRefCitation>
. The use of Chibemba to anchor the ill-defined region of ‘Gambos’ by Crawford-Cabral &amp; Mesquitela (1989) is based on the region “Circunscrição dos Gambos” which in the late nineteenth century was already defined. The village Chibemba developed near a fort, &quot;Forte dos Gambos&quot; on the road between Lubango and Xangongo (L. Veríssimo pers. comm.,
<date id="FFAAE3D0FFDAFFAF0470FC7456BA4724" box="[586,734,999,1024]" pageId="14" pageNumber="356" value="2018-04-03">3 April 2018</date>
).
</paragraph>
<paragraph id="8BABC510FFDAFFAF06FDFB9057E34749" blockId="14.[151,1436,783,1133]" pageId="14" pageNumber="356">
<materialsCitation id="3B7CCF4DFFDAFFAF06FDFB9057E34749" ID-GBIF-Occurrence="2237036586" collectionCode="MBL" county="Presumed" location="Broadley" municipality="Museum" pageId="14" pageNumber="356" specimenCount="1" stateProvince="Lisboa" typeStatus="holotype">
<emphasis id="B9601902FFDAFFAF06FDFB9055594707" bold="true" box="[199,317,1036,1061]" pageId="14" pageNumber="356">
<typeStatus id="54AF7BB2FFDAFFAF06FDFB9055504701" box="[199,308,1036,1061]" pageId="14" pageNumber="356" type="holotype">Holotype</typeStatus>
:
</emphasis>
<collectionCode id="ED055DD5FFDAFFAF0773FB9055EC4700" box="[329,392,1036,1060]" country="Portugal" httpUri="http://grbio.org/cool/pyk6-nt7k" name="Museu Nacional de Historia Natural" pageId="14" pageNumber="356">MBL</collectionCode>
?
<collectingCounty id="62CABD9CFFDAFFAF07A7FB90566F4701" box="[413,523,1036,1061]" pageId="14" pageNumber="356">Presumed</collectingCounty>
lost in fire that destroyed the
<collectingRegion id="49D00BF2FFDAFFAF0547FB9057AF4701" box="[893,971,1036,1061]" country="Portugal" name="Lisboa" pageId="14" pageNumber="356">Lisbon</collectingRegion>
<collectingMunicipality id="6BCF5F6AFFDAFFAF05ECFB91505C4700" box="[982,1080,1037,1060]" pageId="14" pageNumber="356">Museum</collectingMunicipality>
collections.
<location id="8ECB93CBFFDAFFAF02EAFB9051534701" LSID="urn:lsid:plazi:treatment:03BD7406FFDAFFB106ADFDD8556543F0:8ECB93CBFFDAFFAF02EAFB9051534701" box="[1232,1335,1036,1061]" county="Presumed" municipality="Museum" name="Broadley" pageId="14" pageNumber="356" stateProvince="Lisboa">Broadley</location>
(1977b) inspected the
<location id="8ECB93CBFFDAFFAF0708FBAC55E2476C" LSID="urn:lsid:plazi:treatment:03BD7406FFDAFFB106ADFDD8556543F0:8ECB93CBFFDAFFAF0708FBAC55E2476C" box="[306,390,1072,1096]" county="Presumed" municipality="Museum" name="Bocage" pageId="14" pageNumber="356" stateProvince="Lisboa">Bocage</location>
material during a visit to the
<collectingRegion id="49D00BF2FFDAFFAF04ECFBB35740476C" box="[726,804,1071,1096]" country="Portugal" name="Lisboa" pageId="14" pageNumber="356">Lisbon</collectingRegion>
museum in 1968 but noted that
<location id="8ECB93CBFFDAFFAF02A8FBAC509C476C" LSID="urn:lsid:plazi:treatment:03BD7406FFDAFFB106ADFDD8556543F0:8ECB93CBFFDAFFAF02A8FBAC509C476C" box="[1170,1272,1072,1096]" county="Presumed" municipality="Museum" name="Bocage's" pageId="14" pageNumber="356" stateProvince="Lisboa">Bocage’s</location>
type was then absent and it is considered lost, although its fate remains unknown.
</materialsCitation>
</paragraph>
<paragraph id="8BABC510FFDAFFAF06ADFB00557C4790" blockId="14.[151,805,1180,1384]" box="[151,280,1180,1204]" pageId="14" pageNumber="356">Synonymy:</paragraph>
</subSubSection>
<subSubSection id="C30E969BFFDAFFAF06ADFB235741464C" pageId="14" pageNumber="356" type="reference_group">
<paragraph id="8BABC510FFDAFFAF06ADFB2356F147FC" blockId="14.[151,805,1180,1384]" box="[151,661,1215,1240]" pageId="14" pageNumber="356">
<taxonomicName id="4C14BE93FFDAFFAF06ADFB2356F547FC" authority="Bocage 1873, p 221" authorityName=", Bocage" authorityPageNumber="221" authorityYear="1873" box="[151,657,1215,1240]" class="Reptilia" family="Lamprophiidae" genus="Psammophylax" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="356" phylum="Chordata" rank="species" species="ocellatus">
<emphasis id="B9601902FFDAFFAF06ADFB2355CA47FC" box="[151,430,1215,1240]" italics="true" pageId="14" pageNumber="356">Psammophylax ocellatus</emphasis>
<bibRefCitation id="EF85B8E1FFDAFFAF078FFB5C562F47FC" author="Bocage, J. V. B." box="[437,587,1216,1240]" pageId="14" pageNumber="356" pagination="209 - 227" refId="ref10900" refString="Bocage, J. V. B. (1873) Melanges erpetologiques. II. Sur quelques reptiles et batraciens nouveaux, rares ou peu connus d'Afrique occidentale. Jornal de Sciencias, Mathematicas, Physicas e Naturaes, Lisboa, 4, 209 - 227." type="journal article" year="1873">Bocage 1873</bibRefCitation>
, p221
</taxonomicName>
.
</paragraph>
<paragraph id="8BABC510FFDAFFAF06ADFB7856A647D8" blockId="14.[151,805,1180,1384]" box="[151,706,1252,1277]" pageId="14" pageNumber="356">
<taxonomicName id="4C14BE93FFDAFFAF06ADFB78560C47D9" authority="Bocage 1895" authorityName="Bocage" authorityYear="1895" box="[151,616,1252,1277]" class="Reptilia" family="Lamprophiidae" genus="Psammophylax" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="356" phylum="Chordata" rank="species" species="rhombeatus">
<emphasis id="B9601902FFDAFFAF06ADFB7855A947D9" box="[151,461,1252,1277]" italics="true" pageId="14" pageNumber="356">Psammophylax rhombeatus</emphasis>
<bibRefCitation id="EF85B8E1FFDAFFAF07EEFB79560C47D9" author="Bocage, J. V. B." box="[468,616,1252,1277]" pageId="14" pageNumber="356" refId="ref10948" refString="Bocage, J. V. B. (1895) Herpetologie d'Angola et du Congo. Imprimerie Nationale, Lisbonne, 203 pp., 20 pls." type="book" year="1895">Bocage 1895</bibRefCitation>
</taxonomicName>
n, p108.
</paragraph>
<paragraph id="8BABC510FFDAFFAF06ADFA9B576B4604" blockId="14.[151,805,1180,1384]" box="[151,783,1287,1312]" pageId="14" pageNumber="356">
<taxonomicName id="4C14BE93FFDAFFAF06ADFA9B55A64604" box="[151,450,1287,1312]" class="Reptilia" family="Colubridae" genus="Trimerorhinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="356" phylum="Chordata" rank="species" species="rhombeatus">
<emphasis id="B9601902FFDAFFAF06ADFA9B55A64604" box="[151,450,1287,1312]" italics="true" pageId="14" pageNumber="356">Trimerorhinus rhombeatus</emphasis>
</taxonomicName>
(part) Boulenger 1896, p138.
</paragraph>
<paragraph id="8BABC510FFDAFFAF06ADFAB056834660" blockId="14.[151,805,1180,1384]" box="[151,743,1324,1349]" pageId="14" pageNumber="356">
<taxonomicName id="4C14BE93FFDAFFAF06ADFAB056874660" authority="FitzSimons, 1957, p 239" authorityName="FitzSimons" authorityPageNumber="239" authorityYear="1957" box="[151,739,1324,1349]" class="Reptilia" family="Lamprophiidae" genus="Psammophylax" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="356" phylum="Chordata" rank="species" species="rhombeatus">
<emphasis id="B9601902FFDAFFAF06ADFAB055A94661" box="[151,461,1324,1349]" italics="true" pageId="14" pageNumber="356">Psammophylax rhombeatus</emphasis>
FitzSimons, 1957, p239
</taxonomicName>
.
</paragraph>
<paragraph id="8BABC510FFDAFFAF06ADFAD35741464C" blockId="14.[151,805,1180,1384]" box="[151,805,1359,1384]" pageId="14" pageNumber="356">
<taxonomicName id="4C14BE93FFDAFFAF06ADFAD35744464C" authority="Broadley 1977, p 20" authorityName="Broadley" authorityPageNumber="20" authorityYear="1977" box="[151,800,1359,1384]" class="Reptilia" family="Lamprophiidae" genus="Psammophylax" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="356" phylum="Chordata" rank="subSpecies" species="rhombeatus" subSpecies="ocellatus">
<emphasis id="B9601902FFDAFFAF06ADFAD3565D464C" box="[151,569,1359,1384]" italics="true" pageId="14" pageNumber="356">Psammophylax rhombeatus ocellatus</emphasis>
Broadley 1977, p20
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C30E969BFFDAFFAF06ADFA0B5685443C" pageId="14" pageNumber="356" type="materials_examined">
<paragraph id="8BABC510FFDAFFAF06ADFA0B5685443C" blockId="14.[151,1436,1431,2033]" pageId="14" pageNumber="356">
<emphasis id="B9601902FFDAFFAF06ADFA0B55E04694" bold="true" box="[151,388,1431,1456]" pageId="14" pageNumber="356">Material examined.</emphasis>
<taxonomicName id="4C14BE93FFDAFFAF07B4FA0B56CC4694" authorityName=", Bocage" authorityYear="1873" box="[398,680,1431,1456]" class="Reptilia" family="Lamprophiidae" genus="Psammophylax" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="356" phylum="Chordata" rank="species" species="ocellatus">
<emphasis id="B9601902FFDAFFAF07B4FA0B56CC4694" box="[398,680,1431,1456]" italics="true" pageId="14" pageNumber="356">Psammophylax ocellatus</emphasis>
</taxonomicName>
:
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a.s.l),
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,
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,
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<date id="FFAAE3D0FFDAFFAF0451FA20575546F0" box="[619,817,1468,1493]" pageId="14" pageNumber="356" value="2009-01-14">14 January 2009</date>
</collectingDate>
,
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&amp;
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;
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<collectionCode id="ED055DD5FFDAFFAF02B2FA2150A646F0" box="[1160,1218,1469,1492]" country="South Africa" httpUri="http://grbio.org/cool/xqn4-34sv" name="Port Elizabeth Museum" pageId="14" pageNumber="356" type="Museum">PEM</collectionCode>
<specimenCode id="DBB26D6BFFDAFFAF02F7FA20514246F1" box="[1229,1318,1468,1493]" pageId="14" pageNumber="356">R16270</specimenCode>
(
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),
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,
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, (
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,
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<quantity id="4CEC68F5FFDAFFAF05F8FA43507C46DC" box="[962,1048,1503,1528]" metricMagnitude="3" metricUnit="m" metricValue="2.275" pageId="14" pageNumber="356" unit="m" value="2275.0">2275 m</quantity>
</elevation>
a.s.l),
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,
<collectingCountry id="F3038580FFDAFFAF0325FA43511046DC" box="[1311,1396,1503,1528]" name="Angola" pageId="14" pageNumber="356">Angola</collectingCountry>
,
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<date id="FFAAE3D0FFDAFFAF03BAFA7C555B4538" pageId="14" pageNumber="356" value="2017-02-19">19 February 2017</date>
</collectingDate>
</materialsCitation>
;
<materialsCitation id="3B7CCF4DFFDAFFAF0776F998557C4564" ID-GBIF-Occurrence="2338497915" collectingDate="2016-04-13" collectionCode="NB" country="Angola" elevation="2263" latitude="-14.819722" location="Tundavala dirt road" longLatPrecision="21" longitude="13.396944" pageId="14" pageNumber="356" specimenCode="NB171" specimenCount="1" stateProvince="Huila Province">
<specimenCode id="DBB26D6BFFDAFFAF0776F99855FB4538" box="[332,415,1540,1564]" pageId="14" pageNumber="356">NB171</specimenCode>
,
<location id="8ECB93CBFFDAFFAF0797F99856E94539" LSID="urn:lsid:plazi:treatment:03BD7406FFDAFFB106ADFDD8556543F0:8ECB93CBFFDAFFAF0797F99856E94539" box="[429,653,1540,1565]" country="Angola" latitude="-14.819722" longLatPrecision="21" longitude="13.396944" municipality="Tundavala" name="Tundavala dirt road" pageId="14" pageNumber="356" stateProvince="Huila Province">Tundavala dirt road</location>
, (
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,
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,
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<quantity id="4CEC68F5FFDAFFAF05FBF99850734538" box="[961,1047,1540,1565]" metricMagnitude="3" metricUnit="m" metricValue="2.263" pageId="14" pageNumber="356" unit="m" value="2263.0">2263 m</quantity>
</elevation>
a.s.l),
<collectingRegion id="49D00BF2FFDAFFAF025FF99851764539" box="[1125,1298,1540,1565]" country="Angola" name="Huila" pageId="14" pageNumber="356">Huíla Province</collectingRegion>
,
<collectingCountry id="F3038580FFDAFFAF0324F99851174539" box="[1310,1395,1540,1565]" name="Angola" pageId="14" pageNumber="356">Angola</collectingCountry>
,
<collectingDate id="EFEE1A38FFDAFFAF03BAF998557C4564" pageId="14" pageNumber="356" value="2016-04-13">
<date id="FFAAE3D0FFDAFFAF03BAF998557C4564" pageId="14" pageNumber="356" value="2016-04-13">13 April 2016</date>
</collectingDate>
</materialsCitation>
;
<materialsCitation id="3B7CCF4DFFDAFFAF0713F9BB55DB4541" ID-GBIF-Occurrence="2338497278" collectingDate="2016-11-21" collectionCode="NB" country="Angola" elevation="2275" latitude="-14.808111" location="Tundavala grassland near rocks, south of observatory" longLatPrecision="1" longitude="13.401444" municipality="Tundavala grassland near rocks, south of observatory" pageId="14" pageNumber="356" specimenCode="NB491" specimenCount="1" stateProvince="Huila Province">
<specimenCode id="DBB26D6BFFDAFFAF0713F9BB551C4564" box="[297,376,1575,1600]" pageId="14" pageNumber="356">NB491</specimenCode>
,
<collectingMunicipality id="6BCF5F6AFFDAFFAF0746F9BB57944564" box="[380,1008,1575,1600]" country="Angola" latitude="-14.808001" longLatPrecision="1" longitude="13.401999" municipality="Tundavala" name="Tundavala grassland near rocks, south of observatory" pageId="14" pageNumber="356" stateProvince="Huila Province">Tundavala grassland near rocks, south of observatory</collectingMunicipality>
(
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,
<geoCoordinate id="EE20A3D7FFDAFFAF0290F9BB51574564" box="[1194,1331,1575,1600]" direction="east" orientation="longitude" pageId="14" pageNumber="356" precision="1" value="13.401444">13°24'5.2''E</geoCoordinate>
,
<elevation id="00392223FFDAFFAF0379F9BB51F84564" box="[1347,1436,1575,1600]" metricMagnitude="3" metricUnit="m" metricValue="2.275" pageId="14" pageNumber="356" unit="m" value="2275.0">
<quantity id="4CEC68F5FFDAFFAF0379F9BB51F84564" box="[1347,1436,1575,1600]" metricMagnitude="3" metricUnit="m" metricValue="2.275" pageId="14" pageNumber="356" unit="m" value="2275.0">2275 m</quantity>
</elevation>
a.s.l),
<collectingDate id="EFEE1A38FFDAFFAF06E4F9D055DB4541" box="[222,447,1612,1637]" pageId="14" pageNumber="356" value="2016-11-21">
<date id="FFAAE3D0FFDAFFAF06E4F9D055DB4541" box="[222,447,1612,1637]" pageId="14" pageNumber="356" value="2016-11-21">21 November 2016</date>
</collectingDate>
</materialsCitation>
;
<materialsCitation id="3B7CCF4DFFDAFFAF07F7F9D0576845AC" ID-GBIF-Occurrence="2338498300" collectingDate="2016-11-16" collectingMethod="Tundavala rock trap" collectionCode="NB" country="Angola" elevation="2275" latitude="-14.808056" location="south of observatory" longLatPrecision="21" longitude="13.401111" pageId="14" pageNumber="356" specimenCode="NB492" specimenCount="1" stateProvince="Huila Province">
<specimenCode id="DBB26D6BFFDAFFAF07F7F9D056444540" box="[461,544,1612,1636]" pageId="14" pageNumber="356">NB492</specimenCode>
,
<collectingMethod id="5255BD07FFDAFFAF0415F9D057734540" box="[559,791,1612,1637]" pageId="14" pageNumber="356">Tundavala rock trap</collectingMethod>
,
<location id="8ECB93CBFFDAFFAF051FF9D050744541" LSID="urn:lsid:plazi:treatment:03BD7406FFDAFFB106ADFDD8556543F0:8ECB93CBFFDAFFAF051FF9D050744541" box="[805,1040,1612,1637]" country="Angola" latitude="-14.808056" longLatPrecision="21" longitude="13.401111" municipality="Tundavala" name="south of observatory" pageId="14" pageNumber="356" stateProvince="Huila Province">south of observatory</location>
(
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,
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,
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<quantity id="4CEC68F5FFDAFFAF037FF9D051F84540" box="[1349,1436,1612,1637]" metricMagnitude="3" metricUnit="m" metricValue="2.275" pageId="14" pageNumber="356" unit="m" value="2275.0">2275 m</quantity>
</elevation>
a.s.l),
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,
<collectingCountry id="F3038580FFDAFFAF078BF9F3566245AC" box="[433,518,1647,1672]" name="Angola" pageId="14" pageNumber="356">Angola</collectingCountry>
,
<collectingDate id="EFEE1A38FFDAFFAF0426F9F3576845AC" box="[540,780,1647,1672]" pageId="14" pageNumber="356" value="2016-11-16">
<date id="FFAAE3D0FFDAFFAF0426F9F3576845AC" box="[540,780,1647,1672]" pageId="14" pageNumber="356" value="2016-11-16">16 November 2016</date>
</collectingDate>
</materialsCitation>
;
<materialsCitation id="3B7CCF4DFFDAFFAF0518F9F350704588" ID-GBIF-Occurrence="2338496938" collectingDate="2017-04-21" collectingMethod="Tundavala grass trap" collectionCode="NB" country="Angola" elevation="2286" latitude="-14.810278" location="south of observatory" longLatPrecision="21" longitude="13.403611" pageId="14" pageNumber="356" specimenCode="NB585" specimenCount="1" stateProvince="Huila Province">
<specimenCode id="DBB26D6BFFDAFFAF0518F9F3571145AC" box="[802,885,1647,1672]" pageId="14" pageNumber="356">NB585</specimenCode>
,
<collectingMethod id="5255BD07FFDAFFAF05B6F9F350EF45AC" box="[908,1163,1647,1673]" pageId="14" pageNumber="356">Tundavala grass trap</collectingMethod>
,
<location id="8ECB93CBFFDAFFAF029BF9F351F845AC" LSID="urn:lsid:plazi:treatment:03BD7406FFDAFFB106ADFDD8556543F0:8ECB93CBFFDAFFAF029BF9F351F845AC" box="[1185,1436,1647,1672]" country="Angola" latitude="-14.810278" longLatPrecision="21" longitude="13.403611" municipality="Tundavala" name="south of observatory" pageId="14" pageNumber="356" stateProvince="Huila Province">south of observatory</location>
(
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,
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,
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<quantity id="4CEC68F5FFDAFFAF0786F90856754588" box="[444,529,1684,1709]" metricMagnitude="3" metricUnit="m" metricValue="2.286" pageId="14" pageNumber="356" unit="m" value="2286.0">2286 m</quantity>
</elevation>
a.s.l),
<collectingRegion id="49D00BF2FFDAFFAF0466F908576D4589" box="[604,777,1684,1709]" country="Angola" name="Huila" pageId="14" pageNumber="356">Huíla Province</collectingRegion>
,
<collectingCountry id="F3038580FFDAFFAF0529F908570C4589" box="[787,872,1684,1709]" name="Angola" pageId="14" pageNumber="356">Angola</collectingCountry>
,
<collectingDate id="EFEE1A38FFDAFFAF054EF90850704588" box="[884,1044,1684,1709]" pageId="14" pageNumber="356" value="2017-04-21">
<date id="FFAAE3D0FFDAFFAF054EF90850704588" box="[884,1044,1684,1709]" pageId="14" pageNumber="356" value="2017-04-21">21 April 2017</date>
</collectingDate>
</materialsCitation>
;
<materialsCitation id="3B7CCF4DFFDAFFAF021AF90855D945D1" ID-GBIF-Occurrence="2338496375" collectingDate="2017-04-20" collectingMethod="Tundavala grassland near grass trap" collectionCode="NB" collectorName="N. Baptista" country="Angola" elevation="2276" latitude="-14.808001" location="south of observatory" longLatPrecision="1" longitude="13.401999" pageId="14" pageNumber="356" specimenCode="NB586" specimenCount="1" stateProvince="Huila Province">
<specimenCode id="DBB26D6BFFDAFFAF021AF90850104589" box="[1056,1140,1684,1709]" pageId="14" pageNumber="356">NB586</specimenCode>
,
<collectingMethod id="5255BD07FFDAFFAF0245F908556245F4" pageId="14" pageNumber="356">Tundavala grassland near grass trap</collectingMethod>
,
<location id="8ECB93CBFFDAFFAF0729F92B559F45F4" LSID="urn:lsid:plazi:treatment:03BD7406FFDAFFB106ADFDD8556543F0:8ECB93CBFFDAFFAF0729F92B559F45F4" box="[275,507,1719,1744]" country="Angola" latitude="-14.808001" longLatPrecision="1" longitude="13.401999" municipality="Tundavala" name="south of observatory" pageId="14" pageNumber="356" stateProvince="Huila Province">south of observatory</location>
(
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,
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,
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<quantity id="4CEC68F5FFDAFFAF0576F92B57C645F4" box="[844,930,1719,1744]" metricMagnitude="3" metricUnit="m" metricValue="2.276" pageId="14" pageNumber="356" unit="m" value="2276.0">2276 m</quantity>
</elevation>
a.s.l),
<collectingDate id="EFEE1A38FFDAFFAF05D5F92450EA45F4" box="[1007,1166,1719,1744]" pageId="14" pageNumber="356" value="2017-04-20">
<date id="FFAAE3D0FFDAFFAF05D5F92450EA45F4" box="[1007,1166,1719,1744]" pageId="14" pageNumber="356" value="2017-04-20">20 April 2017</date>
</collectingDate>
(all
<location id="8ECB93CBFFDAFFAF02F9F92B515C45F4" LSID="urn:lsid:plazi:treatment:03BD7406FFDAFFB106ADFDD8556543F0:8ECB93CBFFDAFFAF02F9F92B515C45F4" box="[1219,1336,1719,1744]" country="Angola" latitude="-14.808001" longLatPrecision="1" longitude="13.401999" name="Tundavala" pageId="14" pageNumber="356" stateProvince="Huila Province">Tundavala</location>
material collected by
<collectorName id="26E1A0C6FFDAFFAF0714F94155D145D1" box="[302,437,1756,1781]" pageId="14" pageNumber="356">N. Baptista</collectorName>
)
</materialsCitation>
.
<materialsCitation id="3B7CCF4DFFDAFFAF07F1F94056B9443C" ID-GBIF-Occurrence="2237036585" collectingDate="1983-07" collectionCode="SAM" collectorName="John Visser Coll." country="Angola" elevation="1108" latitude="-17.375" location="Calueque" pageId="14" pageNumber="356" specimenCode="ZR 46424" specimenCount="1" stateProvince="Cunene Province">
<collectionCode id="ED055DD5FFDAFFAF07F1F940566C45D0" box="[459,520,1756,1780]" country="South Africa" httpUri="http://grbio.org/cool/mhew-9czz" name="South African Museum" pageId="14" pageNumber="356">SAM</collectionCode>
<specimenCode id="DBB26D6BFFDAFFAF0429F94156EE45D1" box="[531,650,1756,1781]" collectionCode="SAM" country="South Africa" httpUri="http://grbio.org/cool/mhew-9czz" name="South African Museum" pageId="14" pageNumber="356">ZR 46424</specimenCode>
,
<location id="8ECB93CBFFDAFFAF04A3F940576245D1" LSID="urn:lsid:plazi:treatment:03BD7406FFDAFFB106ADFDD8556543F0:8ECB93CBFFDAFFAF04A3F940576245D1" box="[665,774,1756,1781]" country="Angola" latitude="-17.375" name="Calueque" pageId="14" pageNumber="356" stateProvince="Cunene Province">Calueque</location>
,
<collectingRegion id="49D00BF2FFDAFFAF052FF94057B845D1" box="[789,988,1756,1781]" country="Angola" name="Cunene" pageId="14" pageNumber="356">Cunene Province</collectingRegion>
,
<collectingCountry id="F3038580FFDAFFAF05D1F940505945D1" box="[1003,1085,1756,1781]" name="Angola" pageId="14" pageNumber="356">Angola</collectingCountry>
(
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,
<geoCoordinate id="EE20A3D7FFDAFFAF02E2F940515245D1" box="[1240,1334,1756,1781]" pageId="14" pageNumber="356">14.6250</geoCoordinate>
,
<elevation id="00392223FFDAFFAF037FF94051F845D0" box="[1349,1436,1756,1780]" metricMagnitude="3" metricUnit="m" metricValue="1.108" pageId="14" pageNumber="356" unit="m" value="1108.0">
<quantity id="4CEC68F5FFDAFFAF037FF94051F845D0" box="[1349,1436,1756,1780]" metricMagnitude="3" metricUnit="m" metricValue="1.108" pageId="14" pageNumber="356" unit="m" value="1108.0">1108 m</quantity>
</elevation>
a.s.l),
<collectingDate id="EFEE1A38FFDAFFAF06E1F963552D443C" box="[219,329,1791,1816]" pageId="14" pageNumber="356" value="1983-07">
<date id="FFAAE3D0FFDAFFAF06E1F963552D443C" box="[219,329,1791,1816]" pageId="14" pageNumber="356" value="1983-07">July 1983</date>
</collectingDate>
(2930—part of
<collectorName id="26E1A0C6FFDAFFAF07C4F96356DB443C" box="[510,703,1791,1816]" pageId="14" pageNumber="356">John Visser Coll.</collectorName>
?)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30E969BFFDAFFAE06FDF8B85103401E" lastPageId="15" lastPageNumber="357" pageId="14" pageNumber="356" type="description">
<paragraph id="8BABC510FFDAFFAE06FDF8B850014289" blockId="14.[151,1436,1431,2033]" lastBlockId="15.[151,1437,151,1043]" lastPageId="15" lastPageNumber="357" pageId="14" pageNumber="356">
<emphasis id="B9601902FFDAFFAF06FDF8B8553C441F" bold="true" box="[199,344,1828,1853]" pageId="14" pageNumber="356">Description.</emphasis>
The smallest skaapsteker (maximum
<quantity id="4CEC68F5FFDAFFAF052BF8B857114418" box="[785,885,1828,1853]" metricMagnitude="-1" metricUnit="m" metricValue="8.75" pageId="14" pageNumber="356" unit="mm" value="875.0">875 mm</quantity>
; see Broadley 1977b for other species). Nostril pierced between single pre- and post-nasals; preocular 1, separated from frontal; postoculars 2 (sometimes 3 on one side); temporals usually 2+3 (sometimes 2+2 or 2+4); supralabials 8, the fourth &amp; fifth entering orbit (sometimes 3–4); infralabials 10–12, the first 5 usually in contact with anterior sublinguals (rarely 4–6); dorsal scales in 17-17- 13 rows; ventrals 156–183 (see discussion above); cloacal shield divided; subcaudals 57–68. Tail length 21.5–29.1% of total length (see discussion above). Dorsum grey, heavily patterned with longitudinal series of paravertebral and lateral ocelli; paravertebral ocelli comprise 1–2 central reddish brown scales irregularly encircled in black and then with a thin yellow-orange ring, and separated in the midline by a thin white-yellowish line that runs from the neck to the base of the tail; the lateral ocelli are larger and have 1–3 greyish scales and a narrow black edge. Head uniform brown on top with a narrow elongate extension onto the neck; iris of eye brilliant red; the supra- and infralabials are white with scattered black blotches that form more regular bars on the latter; two large, solid back blotches occur on each side of the head, the largest on the temporal region, the other on the side of the neck; ventrum white with 2–5 subtriangular black spots that arise at the base and extend halfway to the free edge of most ventral scales; subcaudals grey-white with scattered blotches on the lateral edges.
</paragraph>
<paragraph id="8BABC510FFDBFFAE06ADFE2B577142F4" blockId="15.[151,1437,151,1043]" box="[151,789,439,464]" pageId="15" pageNumber="357">
Scalation details of specimen are summarized in
<tableCitation id="C696F0ABFFDBFFAE0481FE2B577542F4" box="[699,785,439,464]" captionStart="TABLE 2" captionStartId="9.[147,229,593,615]" captionTargetPageId="9" captionText="TABLE 2. Measurements and Scalation of Psammophylax ocellatus" httpUri="http://table.plazi.org/id/DF6B9598FFDDFFA806A9FDCD57484143" pageId="15" pageNumber="357" tableUuid="DF6B9598FFDDFFA806A9FDCD57484143">Table 2</tableCitation>
.
</paragraph>
<paragraph id="8BABC510FFDBFFAE06FDFE41571842D0" blockId="15.[151,1437,151,1043]" box="[199,892,476,501]" pageId="15" pageNumber="357">
<emphasis id="B9601902FFDBFFAE06FDFE41549A42D0" box="[199,254,477,500]" italics="true" pageId="15" pageNumber="357">Size:</emphasis>
Maximum size: Male (MBL 1724) 720 + 155 =
<quantity id="4CEC68F5FFDBFFAE051AFE40571842D0" box="[800,892,476,501]" metricMagnitude="-1" metricUnit="m" metricValue="8.75" pageId="15" pageNumber="357" unit="mm" value="875.0">875 mm</quantity>
</paragraph>
<paragraph id="8BABC510FFDBFFAE06FDFE635103401E" blockId="15.[151,1437,151,1043]" pageId="15" pageNumber="357">
<emphasis id="B9601902FFDBFFAE06FDFE63554E4133" bold="true" box="[199,298,511,536]" pageId="15" pageNumber="357">Habitat.</emphasis>
Recent material was collected in montane grassland on the edge of the Humpata
<collectingRegion id="49D00BF2FFDBFFAE02CEFE63512D413C" box="[1268,1353,511,536]" country="Nigeria" name="Plateau" pageId="15" pageNumber="357">Plateau</collectingRegion>
, in the vicinity of the Chela Escarpment at an altitude of
<quantity id="4CEC68F5FFDBFFAE04EEFDB857144118" box="[724,880,548,572]" metricMagnitude="3" metricUnit="m" metricValue="2.2" metricValueMax="2.3" metricValueMin="2.1" pageId="15" pageNumber="357" unit="m" value="2200.0" valueMax="2300.0" valueMin="2100.0">2100–2300 m</quantity>
a.s.l, with scattered sandstone outcrops, seepage lines, and patches of
<taxonomicName id="4C14BE93FFDBFFAE07B1FDD555B24144" box="[395,470,585,608]" class="Magnoliopsida" family="Proteaceae" genus="Protea" kingdom="Plantae" order="Proteales" pageId="15" pageNumber="357" phylum="Tracheophyta" rank="genus">
<emphasis id="B9601902FFDBFFAE07B1FDD555B24144" box="[395,470,585,608]" italics="true" pageId="15" pageNumber="357">Protea</emphasis>
</taxonomicName>
sp. bush and dwarf miombo woodland dominated by
<taxonomicName id="4C14BE93FFDBFFAE027EFDDB513A4144" box="[1092,1374,583,608]" class="Magnoliopsida" family="Fabaceae" genus="Brachystegia" kingdom="Plantae" order="Fabales" pageId="15" pageNumber="357" phylum="Tracheophyta" rank="species" species="spiciformis">
<emphasis id="B9601902FFDBFFAE027EFDDB513A4144" box="[1092,1374,583,608]" italics="true" pageId="15" pageNumber="357">Brachystegia spiciformis</emphasis>
</taxonomicName>
(
<figureCitation id="132FD995FFDBFFAE0354FDDB54DE41A0" captionStart="FIGURE 1" captionStartId="7.[151,250,1602,1624]" captionTargetBox="[150,1436,733,1581]" captionTargetId="figure@7.[150,1436,733,1581]" captionTargetPageId="7" captionText="FIGURE 1. Psammophylax ocellatus PEM R17986, Estação Zootécnica, montane grasslands, Huíla Province. A: dorsum showing the characteristic boldly blotched body, a thin vertebral line separating the paravertebral ocelli, and the absence of a dark nape collar; B: ventrum showing the white belly with irregular rows of dark spots, that are absent below the tail; C: Live specimen showing general colouration and striking red eye; D: Habitat—montane grasslands, Estação Zootécnica, Huíla Province, Angola. All photos by W. R. Branch." httpUri="https://zenodo.org/record/2653406/files/figure.png" pageId="15" pageNumber="357">Fig. 1C</figureCitation>
). The climate, pedology, geomorphology and woody species composition were characterised by
<bibRefCitation id="EF85B8E1FFDBFFAE02D5FDF051F841A0" author="Barbosa, L. A. G." box="[1263,1436,620,645]" pageId="15" pageNumber="357" refId="ref10872" refString="Barbosa, L. A. G. (1970) Carta fitogeografica de Angola. Instituto de Investigacao Cientifica de Angola, Luanda, 323 pp." type="book" year="1970">Barbosa (1970)</bibRefCitation>
for Angolan montane grasslands in the Humpata and Tundavala regions. The Tundavala collecting sites were dominated by the grasses
<emphasis id="B9601902FFDBFFAE07F2FD28556541D4" italics="true" pageId="15" pageNumber="357">
<taxonomicName id="4C14BE93FFDBFFAE07F2FD28572741E9" authority=", Schizachyrium" authorityName="Schizachyrium" box="[456,835,692,717]" class="Liliopsida" family="Poaceae" genus="Loudetia" kingdom="Plantae" order="Poales" pageId="15" pageNumber="357" phylum="Tracheophyta" rank="species" species="simplex">Loudetia simplex, Schizachyrium</taxonomicName>
sanguineum, Diheteropogom filifolius, Hyparrhenia cymbaria
</emphasis>
and
<taxonomicName id="4C14BE93FFDBFFAE0703FD4B566241D4" box="[313,518,727,752]" class="Liliopsida" family="Poaceae" genus="Elionurus" kingdom="Plantae" order="Poales" pageId="15" pageNumber="357" phylum="Tracheophyta" rank="species" species="muticus">
<emphasis id="B9601902FFDBFFAE0703FD4B566241D4" box="[313,518,727,752]" italics="true" pageId="15" pageNumber="357">Elionurus muticus</emphasis>
</taxonomicName>
. All Tundavala snakes, except for NB171, were collected within close proximity (
<quantity id="4CEC68F5FFDBFFAE06A5FD6354954033" box="[159,241,766,791]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="15" pageNumber="357" unit="km" value="0.1">0.1 km</quantity>
<superScript id="7C616858FFDBFFAE06CBFD67549D402D" attach="left" box="[241,249,763,777]" fontSize="6" pageId="15" pageNumber="357">2</superScript>
) and around
<quantity id="4CEC68F5FFDBFFAE07ACFD6355B74033" box="[406,467,766,791]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="15" pageNumber="357" unit="km" value="1.0">1 km</quantity>
from the edge of the plateau in the south of the Tundavala SASSCAL biodiversity observatory in habitat consisting of montane grasslands on sandy soil and with weathered sandstone outcrops.
</paragraph>
</subSubSection>
<subSubSection id="C30E969BFFDBFFAE06FDFCDA554B4737" pageId="15" pageNumber="357" type="distribution">
<paragraph id="8BABC510FFDBFFAE06FDFCDA554B4737" blockId="15.[151,1437,151,1043]" pageId="15" pageNumber="357">
<emphasis id="B9601902FFDBFFAE06FDFCDA5504407B" bold="true" box="[199,352,838,863]" pageId="15" pageNumber="357">Distribution.</emphasis>
Currently restricted to isolated populations, with recent material inhabiting montane grasslands at the edge of the Humpata
<collectingRegion id="49D00BF2FFDBFFAE07F7FCF6564440A7" box="[461,544,874,899]" country="Nigeria" name="Plateau" pageId="15" pageNumber="357">Plateau</collectingRegion>
in the vicinity of the Chela Escarpment (
<figureCitation id="132FD995FFDBFFAE05D7FCF6505640A7" box="[1005,1074,874,899]" captionStart="FIGURE 6" captionStartId="15.[151,250,1724,1746]" captionTargetBox="[164,1400,1104,1674]" captionTargetId="figure@15.[150,1436,1095,1702]" captionTargetPageId="15" captionText="FIGURE 6. Distribution of Psammophylax ocellatus (green) and Psammophis ansorgii (blue). Right (A): Africa with location of Angola in blue. Middle (B): Topographic map of Angola, showing relief and central localities, and red square outlining the location of the Humpata Plateau, adjacent to the Chela Escarpment. Localities: 1–2 Catchiungo—Chinguar (blue star) type locality of Psammophis ansorgii; 3, Cunhangamua; 4, Tundavala; 5, Estação Zootécnica; 6, Chibemba (green star) type locality of Psammophylax ocellatus; 7, Humbe; 8, Calueque, near Ruacaná. Left (C): zoomed in region of Humpata Plateau." httpUri="https://zenodo.org/record/2653416/files/figure.png" pageId="15" pageNumber="357">Fig. 6</figureCitation>
). It represents a range extension of about
<quantity id="4CEC68F5FFDBFFAE0738FC13553E4083" box="[258,346,910,935]" metricMagnitude="5" metricUnit="m" metricValue="1.22" pageId="15" pageNumber="357" unit="km" value="122.0">122 km</quantity>
to the northwest from historical material from the Main
<collectingRegion id="49D00BF2FFDBFFAE05C3FC12502F4083" box="[1017,1099,910,935]" country="Nigeria" name="Plateau" pageId="15" pageNumber="357">Plateau</collectingRegion>
region of
<collectingRegion id="49D00BF2FFDBFFAE02F0FC12516C4083" box="[1226,1288,910,935]" country="Angola" name="Huila" pageId="15" pageNumber="357">Huíla</collectingRegion>
and
<collectingRegion id="49D00BF2FFDBFFAE037DFC1351F84083" box="[1351,1436,911,935]" country="Angola" name="Cunene" pageId="15" pageNumber="357">Cunene</collectingRegion>
provinces. It also more than doubles the altitudinal range of the species (
<quantity id="4CEC68F5FFDBFFAE05F1FC2F504540EE" box="[971,1057,947,971]" metricMagnitude="3" metricUnit="m" metricValue="1.108" pageId="15" pageNumber="357" unit="m" value="1108.0">1108 m</quantity>
to
<quantity id="4CEC68F5FFDBFFAE027CFC2E50FF40EE" box="[1094,1179,946,971]" metricMagnitude="3" metricUnit="m" metricValue="2.286" pageId="15" pageNumber="357" unit="m" value="2286.0">2286 m</quantity>
a.s.l). The presence of another putatively isolated population in the Angolan section of the Cuvelai-Etosha Basin from Calueque requires confirmation.
</paragraph>
</subSubSection>
<caption id="DF6B9598FFDBFFAE06ADF92051544469" httpUri="https://zenodo.org/record/2653416/files/figure.png" pageId="15" pageNumber="357" startId="15.[151,250,1724,1746]" targetBox="[164,1400,1104,1674]" targetPageId="15">
<paragraph id="8BABC510FFDBFFAE06ADF92051544469" blockId="15.[151,1436,1724,1869]" pageId="15" pageNumber="357">
<emphasis id="B9601902FFDBFFAE06ADF920557745F6" bold="true" box="[151,275,1724,1746]" pageId="15" pageNumber="357">FIGURE 6.</emphasis>
Distribution of
<taxonomicName id="4C14BE93FFDBFFAE078CF92056D745F6" authorityName=", Bocage" authorityYear="1873" box="[438,691,1724,1746]" class="Reptilia" family="Lamprophiidae" genus="Psammophylax" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="357" phylum="Chordata" rank="species" species="ocellatus">
<emphasis id="B9601902FFDBFFAE078CF92056D745F6" box="[438,691,1724,1746]" italics="true" pageId="15" pageNumber="357">Psammophylax ocellatus</emphasis>
</taxonomicName>
(green) and
<taxonomicName id="4C14BE93FFDBFFAE050EF920506B45F6" authorityName=", Boulenger" authorityYear="1905" box="[820,1039,1724,1746]" class="Reptilia" family="Lamprophiidae" genus="Psammophis" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="357" phylum="Chordata" rank="species" species="ansorgii">
<emphasis id="B9601902FFDBFFAE050EF920506B45F6" box="[820,1039,1724,1746]" italics="true" pageId="15" pageNumber="357">Psammophis ansorgii</emphasis>
</taxonomicName>
(blue). Right (A): Africa with location of Angola in blue. Middle (B): Topographic map of Angola, showing relief and central localities, and red square outlining the location of the Humpata Plateau, adjacent to the Chela Escarpment. Localities: 1–2 Catchiungo—Chinguar (blue star) type locality of
<taxonomicName id="4C14BE93FFDBFFAE073EF88455BB440A" authorityName=", Boulenger" authorityYear="1905" box="[260,479,1816,1838]" class="Reptilia" family="Lamprophiidae" genus="Psammophis" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="357" phylum="Chordata" rank="species" species="ansorgii">
<emphasis id="B9601902FFDBFFAE073EF88455BB440A" box="[260,479,1816,1838]" italics="true" pageId="15" pageNumber="357">Psammophis ansorgii</emphasis>
</taxonomicName>
; 3, Cunhangamua; 4, Tundavala; 5, Estação Zootécnica; 6, Chibemba (green star) type locality of
<taxonomicName id="4C14BE93FFDBFFAE0688F8AB55CA4469" authorityName=", Bocage" authorityYear="1873" box="[178,430,1847,1869]" class="Reptilia" family="Lamprophiidae" genus="Psammophylax" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="357" phylum="Chordata" rank="species" species="ocellatus">
<emphasis id="B9601902FFDBFFAE0688F8AB55CA4469" box="[178,430,1847,1869]" italics="true" pageId="15" pageNumber="357">Psammophylax ocellatus</emphasis>
</taxonomicName>
; 7, Humbe; 8, Calueque, near Ruacaná. Left (C): zoomed in region of Humpata Plateau.
</paragraph>
</caption>
<subSubSection id="C30E969BFFDBFFB106FDF8E5556543F0" lastPageId="16" lastPageNumber="358" pageId="15" pageNumber="357" type="biology_ecology">
<paragraph id="8BABC510FFDBFFB106FDF8E5556543F0" blockId="15.[151,1436,1913,2010]" lastBlockId="16.[151,1436,151,212]" lastPageId="16" lastPageNumber="358" pageId="15" pageNumber="357">
<emphasis id="B9601902FFDBFFAE06FDF8E5554F44B5" bold="true" box="[199,299,1913,1938]" pageId="15" pageNumber="357">Ecology.</emphasis>
NB586 and NB491 were collected while active during the day. PEM R17986 was collected during the day under a small slab within
<quantity id="4CEC68F5FFDBFFAE0437F80256354492" box="[525,593,1950,1974]" metricMagnitude="-1" metricUnit="m" metricValue="3.0" pageId="15" pageNumber="357" unit="cm" value="30.0">30 cm</quantity>
of the edge of a small stream. It had an adult
<taxonomicName id="4C14BE93FFDBFFAE0276F803511B4492" baseAuthorityName="Boulenger" baseAuthorityYear="1882" box="[1100,1407,1949,1974]" class="Amphibia" family="Pyxicephalidae" genus="Tomopterna" kingdom="Animalia" order="Anura" pageId="15" pageNumber="357" phylum="Chordata" rank="species" species="tuberculosa">
<emphasis id="B9601902FFDBFFAE0276F80350B64492" box="[1100,1234,1951,1974]" italics="true" pageId="15" pageNumber="357">Tomopterna</emphasis>
cf.
<emphasis id="B9601902FFDBFFAE02C6F801511B4492" box="[1276,1407,1949,1974]" italics="true" pageId="15" pageNumber="357">tuberculosa</emphasis>
</taxonomicName>
in its stomach, that was mainly undigested and probably collected during the previous night. This was the most frequently seen species of snake in the scope of the Tundavala monitoring events, indicating that it is locally common.
</paragraph>
</subSubSection>
</treatment>
</document>
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