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<document id="2E0119FDEE99CCE68942C2F000FA87F8" ID-DOI="10.5852/ejt.2024.968.2703" ID-ISSN="2118-9773" ID-Zenodo-Dep="14009186" ID-ZooBank="9864A04F-83FA-49EA-8DA3-D0BEA38160E4" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1730222331304" checkinUser="plazi" docAuthor="Gustafsson, Daniel R., Tian, Chunpo, Ren, Mengjiao, Li, Zhu, Sun, Xiuling &amp; Zou, Fasheng" docDate="2024" docId="03BD2071A848687BFDF3F4BDFE0FFE00" docLanguage="en" docName="EJT.2024.968.1-36.pdf" docOrigin="European Journal of Taxonomy 968" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2703/12467" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Goniocotes sarissa Gustafsson, Tian, Ren, Li, Sun &amp; Zou, 2024, sp. nov." docType="treatment" docUuid="CFD6B564-DC4E-44FA-B2D1-3586CCCA76E6" docUuidSource="ZooBank" docVersion="3" lastPageNumber="22" masterDocId="FF845809A859686EFF8BF72CFFACFB1E" masterDocTitle="Separation of Goniocotes Burmeister, 1838, and Dictyocotes Kéler, 1940 (Phthiraptera: Ischnocera), with descriptions of four new species" masterLastPageNumber="36" masterPageNumber="1" pageNumber="18" updateTime="1734488065995" updateUser="felipe" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="191BBD9E5D3668AD8EF3CEAB200482E9" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="CB0EB070A873A42294C08B63940046D0">
<mods:title id="F0A6251F74CD7B982FF478CE0922E1C5">Separation of Goniocotes Burmeister, 1838, and Dictyocotes Kéler, 1940 (Phthiraptera: Ischnocera), with descriptions of four new species</mods:title>
</mods:titleInfo>
<mods:name id="109DD01AA5C3A61A1B77609324F441C5" type="personal">
<mods:role id="5F9CB575C0F1DC74414B4CAC2BF119DD">
<mods:roleTerm id="1AF43EA7403E2518390536863250B8D5">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="58938AB113CE03875ECCBFF822C3BFA2">Gustafsson, Daniel R.</mods:namePart>
<mods:nameIdentifier id="D6E1F26E2AC9E303C6AD8295A0717306" type="LSID">69F60930-F3E2-467B-9403-144CE07756B4</mods:nameIdentifier>
<mods:affiliation id="3B736A1BF82D75B1385EE4E18B69992C">Guangdong Key Laboratory of Animal Conservation and Resource Utilization, Guangdong Public Library of Wild Animal Conservation and Utilization, Institute of Zoology, Guangdong Academy of Sciences, 105 Xingang West Road, Haizhu District, Guangzhou, 510260, Guangdong Province, China.</mods:affiliation>
<mods:nameIdentifier id="5BD3C06A8F370A890FBFF078B9487128" type="email">kotatsu@fripost.org</mods:nameIdentifier>
</mods:name>
<mods:name id="9FB9BA9821200D9B56A4573345F157A7" type="personal">
<mods:role id="4881B4635C2DFF9BACC5B0C6BA6A1931">
<mods:roleTerm id="6DB9B9D76A5D32BC9AE4FD772A54C7D7">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="CB0A084E142C843ABE6EA5E6528F2724">Tian, Chunpo</mods:namePart>
<mods:nameIdentifier id="0FE48321DA371B33D33E82FAB5FB66AC" type="LSID">B9545E4F-52E9-4B91-B00D-A4F54C316D8D</mods:nameIdentifier>
<mods:affiliation id="3DA3EFDCAEBCEE9A0142F478AF6AF096">Guangdong Key Laboratory of Animal Conservation and Resource Utilization, Guangdong Public Library of Wild Animal Conservation and Utilization, Institute of Zoology, Guangdong Academy of Sciences, 105 Xingang West Road, Haizhu District, Guangzhou, 510260, Guangdong Province, China. &amp; College of Life Sciences, Shaanxi Normal University, 620 West Chang’an Street, Chang’an District, Xi’an City, 710119, Shaanxi Province, China.</mods:affiliation>
<mods:nameIdentifier id="C519BCEF563A7573FC1DAFD3723186EE" type="email">tichpo0926@snnu.edu.cn</mods:nameIdentifier>
</mods:name>
<mods:name id="80C0AD445CB96D9A7319BC32FCEFC7BB" type="personal">
<mods:role id="5CEE0221FB6F16C507F203D4147674F8">
<mods:roleTerm id="7BCCA43C40CE988C8356FE2C76F38B15">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E7075BF94B408B9A462C397899B6458C">Ren, Mengjiao</mods:namePart>
<mods:nameIdentifier id="6D5860266D05C75906E4F5A56E3BAEAB" type="LSID">6C30C981-9587-4BCB-99B1-07DC9ECD8713</mods:nameIdentifier>
<mods:affiliation id="1E6E61BE9E05A6B8C27D95BCDD623CB6">Guangdong Key Laboratory of Animal Conservation and Resource Utilization, Guangdong Public Library of Wild Animal Conservation and Utilization, Institute of Zoology, Guangdong Academy of Sciences, 105 Xingang West Road, Haizhu District, Guangzhou, 510260, Guangdong Province, China.</mods:affiliation>
<mods:nameIdentifier id="F3D7C274B68BEFED68F96839E1E34BDC" type="email">renmengjiao1119@163.com</mods:nameIdentifier>
</mods:name>
<mods:name id="99ECD31E8C326C33355F29F36D73F9CA" type="personal">
<mods:role id="F7DFCE3D30880603D19EB28F8E3204BF">
<mods:roleTerm id="0A6DC2393F24F53F5ED86C8C9C48D56F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E0E3C60A5F2C1A3F3AF7967691D0C81F">Li, Zhu</mods:namePart>
<mods:nameIdentifier id="105DA26ADA9E29D5E48ACDC711731EC1" type="LSID">B79ADF17-5EC4-45F9-9F7F-B9A1D9A967D5</mods:nameIdentifier>
<mods:affiliation id="BD8190CF73CDB59BAE540F790A5CC303">Department of Life Sciences, National Natural History Museum of China, 126, Tianqiao South St. Dongcheng District, Beijing, 100050, China. &amp; Department of Life Sciences, National Natural History Museum of China, 126, Tianqiao South St. Dongcheng District, Beijing, 100050, China.</mods:affiliation>
<mods:nameIdentifier id="7117D0BA6FDD6D2E0C042D675A5E2489" type="email">bamboo_L@163.com</mods:nameIdentifier>
</mods:name>
<mods:name id="48459B24233C2E2B2203B2E7496E0CD2" type="personal">
<mods:role id="B54A6B122B5C9A0BEA5456513BDF3782">
<mods:roleTerm id="F1FBA9A5E730F233B0E5604D82F2DB02">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="796D8B17AE30EBDAE1E1E522C5A812BC">Sun, Xiuling</mods:namePart>
<mods:nameIdentifier id="5A7D5191BBDD069424CB476515F14964" type="LSID">1D6BA247-F1A6-49CF-961A-A1149FD411B9</mods:nameIdentifier>
<mods:affiliation id="F92F39C1C413B1AA87883EB868A0570C">Collections Department, National Natural History Museum of China, 126, Tianqiao South St. Dongcheng District, Beijing, 100050, China.</mods:affiliation>
<mods:nameIdentifier id="E10C0C97AAA735880C92887DD0BB03FD" type="email">sunshine-320@163.com</mods:nameIdentifier>
</mods:name>
<mods:name id="9EF5B0A1E70AE7F147FC7BB0911C068F" type="personal">
<mods:role id="35261CD6B304AAB7D708F448D16F5F77">
<mods:roleTerm id="A04BBDDF41B8162D666B6DD586002D4A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="10E6B617E27BF99BAB90B030E3F1AD26">Zou, Fasheng</mods:namePart>
<mods:nameIdentifier id="F06F5FEA214A0D522D6524888DC123D0" type="LSID">9D1BB50D-CD19-4332-8854-4D0EA411342C</mods:nameIdentifier>
<mods:affiliation id="85E72135ACC4CB3FF990C4117E3ED79E">Guangdong Key Laboratory of Animal Conservation and Resource Utilization, Guangdong Public Library of Wild Animal Conservation and Utilization, Institute of Zoology, Guangdong Academy of Sciences, 105 Xingang West Road, Haizhu District, Guangzhou, 510260, Guangdong Province, China.</mods:affiliation>
<mods:nameIdentifier id="11CCEDDC081164106B0C951A9A4C190B" type="email">zoufs@giz.gd.cn</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="85A4CEBBD514B04BF79BD66DBC2294DD">text</mods:typeOfResource>
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<mods:title id="2805987C9A3AAA2714954C83E27ABF00">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="A5B65BC211B28FED5387C03FF91384E3">
<mods:date id="F182A015582D678CDDD1821C9AE8FC17">2024</mods:date>
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<mods:number id="019F63205C2299F9EF9F6B78E9C3A5AE">2024-10-25</mods:number>
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<mods:detail id="3F59FCA72034B4ECC5D3C4E984A0B39D" type="volume">
<mods:number id="60F94C8A2551DECB90D8E125E408B26A">968</mods:number>
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<mods:start id="53CAB50C3D0FED1D99831EBC1A4C5671">1</mods:start>
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<mods:location id="4082307709DE6ED5B9DD72E710DC6ABA">
<mods:url id="21BE4AFD0ACE62F83A5587158024B78F">https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2703/12467</mods:url>
</mods:location>
<mods:classification id="BD6F0FB9CB237F8ED4AA3B5646C78370">journal article</mods:classification>
<mods:identifier id="26DEC7388B05DB9890482AB44BC5902D" type="DOI">10.5852/ejt.2024.968.2703</mods:identifier>
<mods:identifier id="8E65AC972E81EE4DC5C211B24CC6A263" type="ISSN">2118-9773</mods:identifier>
<mods:identifier id="EBCD6D6C35ED012968D84D0852AA8E06" type="Zenodo-Dep">14009186</mods:identifier>
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</mods:mods>
<treatment id="03BD2071A848687BFDF3F4BDFE0FFE00" LSID="urn:lsid:zoobank.org:act:CFD6B564-DC4E-44FA-B2D1-3586CCCA76E6" httpUri="http://treatment.plazi.org/id/03BD2071A848687BFDF3F4BDFE0FFE00" lastPageId="21" lastPageNumber="22" pageId="17" pageNumber="18">
<subSubSection id="C30EC2ECA848687FFDF3F4BDFC10F8B5" box="[632,956,913,939]" pageId="17" pageNumber="18" type="nomenclature">
<paragraph id="8BAB9167A848687FFDF3F4BDFC10F8B5" blockId="17.[355,1232,913,1009]" box="[632,956,913,939]" pageId="17" pageNumber="18">
<heading id="D0E3260BA848687FFDF3F4BDFC10F8B5" box="[632,956,913,939]" centered="true" fontSize="11" level="2" pageId="17" pageNumber="18" reason="2">
<taxonomicName id="4C14EAE4A848687FFDF3F4BDFCF6F8B5" authorityName="Gustafsson &amp; Tian &amp; Ren &amp; Li &amp; Sun &amp; Zou" authorityYear="2024" box="[632,858,913,939]" class="Insecta" family="Philopteridae" genus="Goniocotes" kingdom="Animalia" order="Psocodea" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="sarissa" status="sp. nov.">
<emphasis id="B9604D75A848687FFDF3F4BDFCF6F8B5" bold="true" box="[632,858,913,939]" italics="true" pageId="17" pageNumber="18">Goniocotes sarissa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A253F00EA848687FFCEAF4BDFC10F8B5" box="[865,956,913,939]" pageId="17" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30EC2ECA848687FFEE8F49FFCF6F8EF" pageId="17" pageNumber="18" type="description">
<paragraph id="8BAB9167A848687FFEE8F49FFB7CF8D0" blockId="17.[355,1232,913,1009]" box="[355,1232,947,974]" pageId="17" pageNumber="18">
<uri id="FF859D65A848687FFEE8F49FFB7CF8D0" box="[355,1232,947,974]" pageId="17" pageNumber="18">
urn:lsid:zoobank.org:act:
<uuid id="FFB2ABB2A848687FFD0BF49FFB7CF8D0" box="[640,1232,947,974]" pageId="17" pageNumber="18">CFD6B564-DC4E-44FA-B2D1-3586CCCA76E6</uuid>
</uri>
</paragraph>
<paragraph id="8BAB9167A848687FFD52F4FAFCF6F8EF" blockId="17.[355,1232,913,1009]" box="[729,858,982,1009]" pageId="17" pageNumber="18">
<figureCitation id="132F8DE2A848687FFD52F4FAFCF6F8EF" box="[729,858,982,1009]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Figs 13–15" captionStartId-0="18.[189,232,1963,1989]" captionStartId-1="19.[189,232,1998,2024]" captionStartId-2="20.[189,242,1963,1989]" captionTargetBox-0="[427,1218,685,1921]" captionTargetBox-1="[416,1161,676,1933]" captionTargetBox-2="[342,1235,650,1888]" captionTargetId-0="figure-112@18.[340,1247,671,1926]" captionTargetId-1="figure-120@19.[340,1247,670,1953]" captionTargetId-2="figure-39@20.[340,1247,636,1919]" captionTargetPageId-0="18" captionTargetPageId-1="19" captionTargetPageId-2="20" captionText-0="Fig. 11. Goniocotes sarissa sp. nov., holotype, ♂ (BMNH, slide 95, box E0026011). Habitus, dorsal and ventral views." captionText-1="Fig. 12. Goniocotes sarissa sp. nov. Female habitus, dorsal and ventral views." captionText-2="Figs 13–15. Goniocotes sarissa sp. nov. 13–14. Holotype, ♂ (BMNH, slide 95, box E0026011). 13. Head, dorsal and ventral views. 14. Genitalia, dorsal view. 15. Female genitalia, ventral view." figureDoi-0="http://doi.org/10.5281/zenodo.14009206" figureDoi-1="http://doi.org/10.5281/zenodo.14009208" figureDoi-2="http://doi.org/10.5281/zenodo.14009210" httpUri-0="https://zenodo.org/record/14009206/files/figure.png" httpUri-1="https://zenodo.org/record/14009208/files/figure.png" httpUri-2="https://zenodo.org/record/14009210/files/figure.png" pageId="17" pageNumber="18">Figs 11–15</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30EC2ECA848687CFF36F33BFCB6FA76" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18" type="diagnosis">
<paragraph id="8BAB9167A848687FFF36F33BFE9AFF2F" blockId="17.[189,310,1047,1073]" box="[189,310,1047,1073]" pageId="17" pageNumber="18">
<heading id="D0E3260BA848687FFF36F33BFE9AFF2F" bold="true" box="[189,310,1047,1073]" fontSize="11" level="3" pageId="17" pageNumber="18" reason="3">
<emphasis id="B9604D75A848687FFF36F33BFE9AFF2F" bold="true" box="[189,310,1047,1073]" pageId="17" pageNumber="18">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BAB9167A848687FFF36F369FD71FE13" blockId="17.[189,1399,1093,1293]" pageId="17" pageNumber="18">
<taxonomicName id="4C14EAE4A848687FFF36F369FE32FF41" authorityName="Gustafsson &amp; Tian &amp; Ren &amp; Li &amp; Sun &amp; Zou" authorityYear="2024" box="[189,414,1093,1119]" class="Insecta" family="Philopteridae" genus="Goniocotes" kingdom="Animalia" order="Psocodea" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="sarissa" status="sp. nov.">
<emphasis id="B9604D75A848687FFF36F369FE32FF41" box="[189,414,1093,1119]" italics="true" pageId="17" pageNumber="18">Goniocotes sarissa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A253F00EA848687FFE2DF36AFDADFF7E" box="[422,513,1094,1120]" pageId="17" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
is most similar to
<taxonomicName id="4C14EAE4A848687FFD54F369FC7FFF41" authorityName="Gustafsson, Tian &amp; Zou" authorityYear="2021" box="[735,979,1093,1119]" class="Insecta" family="Philopteridae" genus="Goniocotes" kingdom="Animalia" order="Psocodea" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="kristinae">
<emphasis id="B9604D75A848687FFD54F369FC7FFF41" box="[735,979,1093,1119]" italics="true" pageId="17" pageNumber="18">Goniocotes kristinae</emphasis>
</taxonomicName>
, with which it shares the following characters: most sclerotized plates densely reticulated (
<figureCitation id="132F8DE2A848687FFCDDF344FC4DFF9C" box="[854,993,1128,1154]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="18.[189,232,1963,1989]" captionStartId-1="19.[189,232,1998,2024]" captionTargetBox-0="[427,1218,685,1921]" captionTargetBox-1="[416,1161,676,1933]" captionTargetId-0="figure-112@18.[340,1247,671,1926]" captionTargetId-1="figure-120@19.[340,1247,670,1953]" captionTargetPageId-0="18" captionTargetPageId-1="19" captionText-0="Fig. 11. Goniocotes sarissa sp. nov., holotype, ♂ (BMNH, slide 95, box E0026011). Habitus, dorsal and ventral views." captionText-1="Fig. 12. Goniocotes sarissa sp. nov. Female habitus, dorsal and ventral views." figureDoi-0="http://doi.org/10.5281/zenodo.14009206" figureDoi-1="http://doi.org/10.5281/zenodo.14009208" httpUri-0="https://zenodo.org/record/14009206/files/figure.png" httpUri-1="https://zenodo.org/record/14009208/files/figure.png" pageId="17" pageNumber="18">Figs 11–12</figureCitation>
); male
<emphasis id="B9604D75A848687FFBB7F345FBFAFF9C" box="[1084,1110,1129,1154]" italics="true" pageId="17" pageNumber="18">os</emphasis>
macrosetae, but female
<emphasis id="B9604D75A848687FFF36F3A7FF7BFFBA" box="[189,215,1163,1188]" italics="true" pageId="17" pageNumber="18">os</emphasis>
microsetae (
<figureCitation id="132F8DE2A848687FFEFFF3A6FDACFFBB" box="[372,512,1162,1189]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="18.[189,232,1963,1989]" captionStartId-1="19.[189,232,1998,2024]" captionTargetBox-0="[427,1218,685,1921]" captionTargetBox-1="[416,1161,676,1933]" captionTargetId-0="figure-112@18.[340,1247,671,1926]" captionTargetId-1="figure-120@19.[340,1247,670,1953]" captionTargetPageId-0="18" captionTargetPageId-1="19" captionText-0="Fig. 11. Goniocotes sarissa sp. nov., holotype, ♂ (BMNH, slide 95, box E0026011). Habitus, dorsal and ventral views." captionText-1="Fig. 12. Goniocotes sarissa sp. nov. Female habitus, dorsal and ventral views." figureDoi-0="http://doi.org/10.5281/zenodo.14009206" figureDoi-1="http://doi.org/10.5281/zenodo.14009208" httpUri-0="https://zenodo.org/record/14009206/files/figure.png" httpUri-1="https://zenodo.org/record/14009208/files/figure.png" pageId="17" pageNumber="18">Figs 11–12</figureCitation>
); male pteronotum with patch of multiple
<emphasis id="B9604D75A848687FFB82F3A7FBE8FFBA" box="[1033,1092,1163,1188]" italics="true" pageId="17" pageNumber="18">smns</emphasis>
centrally (
<figureCitation id="132F8DE2A848687FFB41F3A6FA8FFFBB" box="[1226,1315,1162,1189]" captionStart="Fig" captionStartId="18.[189,232,1963,1989]" captionTargetBox="[427,1218,685,1921]" captionTargetId="figure-112@18.[340,1247,671,1926]" captionTargetPageId="18" captionText="Fig. 11. Goniocotes sarissa sp. nov., holotype, ♂ (BMNH, slide 95, box E0026011). Habitus, dorsal and ventral views." figureDoi="http://doi.org/10.5281/zenodo.14009206" httpUri="https://zenodo.org/record/14009206/files/figure.png" pageId="17" pageNumber="18">Fig. 11</figureCitation>
); male tergopleurites II–IV with
<emphasis id="B9604D75A848687FFE73F382FDB6FFD9" box="[504,538,1198,1223]" italics="true" pageId="17" pageNumber="18">tps</emphasis>
(
<figureCitation id="132F8DE2A848687FFDA4F381FD26FFD9" box="[559,650,1197,1223]" captionStart="Fig" captionStartId="18.[189,232,1963,1989]" captionTargetBox="[427,1218,685,1921]" captionTargetId="figure-112@18.[340,1247,671,1926]" captionTargetPageId="18" captionText="Fig. 11. Goniocotes sarissa sp. nov., holotype, ♂ (BMNH, slide 95, box E0026011). Habitus, dorsal and ventral views." figureDoi="http://doi.org/10.5281/zenodo.14009206" httpUri="https://zenodo.org/record/14009206/files/figure.png" pageId="17" pageNumber="18">Fig. 11</figureCitation>
);
<emphasis id="B9604D75A848687FFD2EF382FD10FFD9" box="[677,700,1198,1223]" italics="true" pageId="17" pageNumber="18">ss</emphasis>
absent on male tergopleurites VI–VIII (
<figureCitation id="132F8DE2A848687FFB3BF381FAA7FFD9" box="[1200,1291,1197,1223]" captionStart="Fig" captionStartId="18.[189,232,1963,1989]" captionTargetBox="[427,1218,685,1921]" captionTargetId="figure-112@18.[340,1247,671,1926]" captionTargetPageId="18" captionText="Fig. 11. Goniocotes sarissa sp. nov., holotype, ♂ (BMNH, slide 95, box E0026011). Habitus, dorsal and ventral views." figureDoi="http://doi.org/10.5281/zenodo.14009206" httpUri="https://zenodo.org/record/14009206/files/figure.png" pageId="17" pageNumber="18">Fig. 11</figureCitation>
); female tergopleurites II–IV with
<emphasis id="B9604D75A848687FFE65F3FDFD8EFFF4" box="[494,546,1233,1258]" italics="true" pageId="17" pageNumber="18">psps</emphasis>
(
<figureCitation id="132F8DE2A848687FFDB8F3FCFD27FFF4" box="[563,651,1232,1258]" captionStart="Fig" captionStartId="19.[189,232,1998,2024]" captionTargetBox="[416,1161,676,1933]" captionTargetId="figure-120@19.[340,1247,670,1953]" captionTargetPageId="19" captionText="Fig. 12. Goniocotes sarissa sp. nov. Female habitus, dorsal and ventral views." figureDoi="http://doi.org/10.5281/zenodo.14009208" httpUri="https://zenodo.org/record/14009208/files/figure.png" pageId="17" pageNumber="18">Fig. 12</figureCitation>
); female but not male with reticulated lateral accessory sternal plates on abdominal segments II–VI (
<figureCitation id="132F8DE2A848687FFDF3F3DEFD7CFE13" box="[632,720,1266,1293]" captionStart="Fig" captionStartId="19.[189,232,1998,2024]" captionTargetBox="[416,1161,676,1933]" captionTargetId="figure-120@19.[340,1247,670,1953]" captionTargetPageId="19" captionText="Fig. 12. Goniocotes sarissa sp. nov. Female habitus, dorsal and ventral views." figureDoi="http://doi.org/10.5281/zenodo.14009208" httpUri="https://zenodo.org/record/14009208/files/figure.png" pageId="17" pageNumber="18">Fig. 12</figureCitation>
).
</paragraph>
<paragraph id="8BAB9167A848687FFF36F218FDD4FEE5" blockId="17.[186,1399,1331,1531]" pageId="17" pageNumber="18">
<taxonomicName id="4C14EAE4A848687FFF36F218FE0EFE53" authorityName="Gustafsson &amp; Tian &amp; Ren &amp; Li &amp; Sun &amp; Zou" authorityYear="2024" box="[189,418,1332,1358]" class="Insecta" family="Philopteridae" genus="Goniocotes" kingdom="Animalia" order="Psocodea" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="sarissa" status="sp. nov.">
<emphasis id="B9604D75A848687FFF36F218FE0EFE53" box="[189,418,1332,1358]" italics="true" pageId="17" pageNumber="18">Goniocotes sarissa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A253F00EA848687FFE24F218FDA3FE50" box="[431,527,1332,1358]" pageId="17" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
can be separated from
<taxonomicName id="4C14EAE4A848687FFCB0F218FC67FE53" authorityName="Gustafsson, Tian &amp; Zou" authorityYear="2021" box="[827,971,1331,1358]" class="Insecta" family="Philopteridae" genus="Goniocotes" kingdom="Animalia" order="Psocodea" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="kristinae">
<emphasis id="B9604D75A848687FFCB0F218FC67FE53" box="[827,971,1331,1358]" italics="true" pageId="17" pageNumber="18">G. kristinae</emphasis>
</taxonomicName>
by the following characters: male tergopleurites II–III with more numerous
<emphasis id="B9604D75A848687FFD19F27BFD18FE6E" box="[658,692,1367,1392]" italics="true" pageId="17" pageNumber="18">tps</emphasis>
in
<taxonomicName id="4C14EAE4A848687FFD58F27AFCEAFE6E" authorityName="Gustafsson &amp; Tian &amp; Ren &amp; Li &amp; Sun &amp; Zou" authorityYear="2024" box="[723,838,1366,1392]" class="Insecta" family="Philopteridae" genus="Goniocotes" kingdom="Animalia" order="Psocodea" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="sarissa">
<emphasis id="B9604D75A848687FFD58F27AFCEAFE6E" box="[723,838,1366,1392]" italics="true" pageId="17" pageNumber="18">G. sarissa</emphasis>
</taxonomicName>
(
<figureCitation id="132F8DE2A848687FFCDCF27AFC0BFE6E" box="[855,935,1366,1392]" captionStart="Fig" captionStartId="18.[189,232,1963,1989]" captionTargetBox="[427,1218,685,1921]" captionTargetId="figure-112@18.[340,1247,671,1926]" captionTargetPageId="18" captionText="Fig. 11. Goniocotes sarissa sp. nov., holotype, ♂ (BMNH, slide 95, box E0026011). Habitus, dorsal and ventral views." figureDoi="http://doi.org/10.5281/zenodo.14009206" httpUri="https://zenodo.org/record/14009206/files/figure.png" pageId="17" pageNumber="18">Fig. 11</figureCitation>
) than in
<taxonomicName id="4C14EAE4A848687FFB89F27AFB26FE6E" authorityName="Gustafsson, Tian &amp; Zou" authorityYear="2021" box="[1026,1162,1366,1392]" class="Insecta" family="Philopteridae" genus="Goniocotes" kingdom="Animalia" order="Psocodea" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="kristinae">
<emphasis id="B9604D75A848687FFB89F27AFB26FE6E" box="[1026,1162,1366,1392]" italics="true" pageId="17" pageNumber="18">G. kristinae</emphasis>
</taxonomicName>
; male tergopleurites VI with
<emphasis id="B9604D75A848687FFE90F255FE9EFE8C" box="[283,306,1401,1426]" italics="true" pageId="17" pageNumber="18">ss</emphasis>
in
<taxonomicName id="4C14EAE4A848687FFEDDF255FE67FE8C" authorityName="Gustafsson &amp; Tian &amp; Ren &amp; Li &amp; Sun &amp; Zou" authorityYear="2024" box="[342,459,1401,1427]" class="Insecta" family="Philopteridae" genus="Goniocotes" kingdom="Animalia" order="Psocodea" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="sarissa">
<emphasis id="B9604D75A848687FFEDDF255FE67FE8C" box="[342,459,1401,1427]" italics="true" pageId="17" pageNumber="18">G. sarissa</emphasis>
</taxonomicName>
(
<figureCitation id="132F8DE2A848687FFE51F255FD82FE8D" box="[474,558,1401,1427]" captionStart="Fig" captionStartId="18.[189,232,1963,1989]" captionTargetBox="[427,1218,685,1921]" captionTargetId="figure-112@18.[340,1247,671,1926]" captionTargetPageId="18" captionText="Fig. 11. Goniocotes sarissa sp. nov., holotype, ♂ (BMNH, slide 95, box E0026011). Habitus, dorsal and ventral views." figureDoi="http://doi.org/10.5281/zenodo.14009206" httpUri="https://zenodo.org/record/14009206/files/figure.png" pageId="17" pageNumber="18">Fig. 11</figureCitation>
), but without
<emphasis id="B9604D75A848687FFD46F255FD48FE8C" box="[717,740,1401,1426]" italics="true" pageId="17" pageNumber="18">ss</emphasis>
in
<taxonomicName id="4C14EAE4A848687FFC8CF255FC3CFE8D" authorityName="Gustafsson, Tian &amp; Zou" authorityYear="2021" box="[775,912,1401,1427]" class="Insecta" family="Philopteridae" genus="Goniocotes" kingdom="Animalia" order="Psocodea" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="kristinae">
<emphasis id="B9604D75A848687FFC8CF255FC3CFE8D" box="[775,912,1401,1427]" italics="true" pageId="17" pageNumber="18">G. kristinae</emphasis>
</taxonomicName>
; proepimera fused medianly in
<taxonomicName id="4C14EAE4A848687FFB74F255FAD9FE8C" authorityName="Gustafsson &amp; Tian &amp; Ren &amp; Li &amp; Sun &amp; Zou" authorityYear="2024" box="[1279,1397,1401,1427]" class="Insecta" family="Philopteridae" genus="Goniocotes" kingdom="Animalia" order="Psocodea" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="sarissa">
<emphasis id="B9604D75A848687FFB74F255FAD9FE8C" box="[1279,1397,1401,1427]" italics="true" pageId="17" pageNumber="18">G. sarissa</emphasis>
</taxonomicName>
(
<figureCitation id="132F8DE2A848687FFF4EF2B7FEE3FEAB" box="[197,335,1435,1461]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="18.[189,232,1963,1989]" captionStartId-1="19.[189,232,1998,2024]" captionTargetBox-0="[427,1218,685,1921]" captionTargetBox-1="[416,1161,676,1933]" captionTargetId-0="figure-112@18.[340,1247,671,1926]" captionTargetId-1="figure-120@19.[340,1247,670,1953]" captionTargetPageId-0="18" captionTargetPageId-1="19" captionText-0="Fig. 11. Goniocotes sarissa sp. nov., holotype, ♂ (BMNH, slide 95, box E0026011). Habitus, dorsal and ventral views." captionText-1="Fig. 12. Goniocotes sarissa sp. nov. Female habitus, dorsal and ventral views." figureDoi-0="http://doi.org/10.5281/zenodo.14009206" figureDoi-1="http://doi.org/10.5281/zenodo.14009208" httpUri-0="https://zenodo.org/record/14009206/files/figure.png" httpUri-1="https://zenodo.org/record/14009208/files/figure.png" pageId="17" pageNumber="18">Figs 11–12</figureCitation>
), but medianly separated in
<taxonomicName id="4C14EAE4A848687FFD20F2B0FC96FEAB" authorityName="Gustafsson, Tian &amp; Zou" authorityYear="2021" box="[683,826,1435,1462]" class="Insecta" family="Philopteridae" genus="Goniocotes" kingdom="Animalia" order="Psocodea" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="kristinae">
<emphasis id="B9604D75A848687FFD20F2B0FC96FEAB" box="[683,826,1435,1462]" italics="true" pageId="17" pageNumber="18">G. kristinae</emphasis>
</taxonomicName>
; temples more flaring posteriorly in
<taxonomicName id="4C14EAE4A848687FFB77F2B0FADAFEAB" authorityName="Gustafsson &amp; Tian &amp; Ren &amp; Li &amp; Sun &amp; Zou" authorityYear="2024" box="[1276,1398,1436,1462]" class="Insecta" family="Philopteridae" genus="Goniocotes" kingdom="Animalia" order="Psocodea" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="sarissa">
<emphasis id="B9604D75A848687FFB77F2B0FADAFEAB" box="[1276,1398,1436,1462]" italics="true" pageId="17" pageNumber="18">G. sarissa</emphasis>
</taxonomicName>
(
<figureCitation id="132F8DE2A848687FFF4EF292FEE6FEC6" box="[197,330,1470,1496]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="18.[189,232,1963,1989]" captionStartId-1="19.[189,232,1998,2024]" captionTargetBox-0="[427,1218,685,1921]" captionTargetBox-1="[416,1161,676,1933]" captionTargetId-0="figure-112@18.[340,1247,671,1926]" captionTargetId-1="figure-120@19.[340,1247,670,1953]" captionTargetPageId-0="18" captionTargetPageId-1="19" captionText-0="Fig. 11. Goniocotes sarissa sp. nov., holotype, ♂ (BMNH, slide 95, box E0026011). Habitus, dorsal and ventral views." captionText-1="Fig. 12. Goniocotes sarissa sp. nov. Female habitus, dorsal and ventral views." figureDoi-0="http://doi.org/10.5281/zenodo.14009206" figureDoi-1="http://doi.org/10.5281/zenodo.14009208" httpUri-0="https://zenodo.org/record/14009206/files/figure.png" httpUri-1="https://zenodo.org/record/14009208/files/figure.png" pageId="17" pageNumber="18">Figs 11–12</figureCitation>
) than in
<taxonomicName id="4C14EAE4A848687FFE3DF292FDECFEC6" authorityName="Gustafsson, Tian &amp; Zou" authorityYear="2021" box="[438,576,1470,1496]" class="Insecta" family="Philopteridae" genus="Goniocotes" kingdom="Animalia" order="Psocodea" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="kristinae">
<emphasis id="B9604D75A848687FFE3DF292FDECFEC6" box="[438,576,1470,1496]" italics="true" pageId="17" pageNumber="18">G. kristinae</emphasis>
</taxonomicName>
; rhombic sclerite separate from pronotum in
<taxonomicName id="4C14EAE4A848687FFBD1F292FB7DFEC6" authorityName="Gustafsson &amp; Tian &amp; Ren &amp; Li &amp; Sun &amp; Zou" authorityYear="2024" box="[1114,1233,1470,1496]" class="Insecta" family="Philopteridae" genus="Goniocotes" kingdom="Animalia" order="Psocodea" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="sarissa">
<emphasis id="B9604D75A848687FFBD1F292FB7DFEC6" box="[1114,1233,1470,1496]" italics="true" pageId="17" pageNumber="18">G. sarissa</emphasis>
</taxonomicName>
(
<figureCitation id="132F8DE2A848687FFB6AF292FAC4FEC6" box="[1249,1384,1470,1496]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="18.[189,232,1963,1989]" captionStartId-1="19.[189,232,1998,2024]" captionTargetBox-0="[427,1218,685,1921]" captionTargetBox-1="[416,1161,676,1933]" captionTargetId-0="figure-112@18.[340,1247,671,1926]" captionTargetId-1="figure-120@19.[340,1247,670,1953]" captionTargetPageId-0="18" captionTargetPageId-1="19" captionText-0="Fig. 11. Goniocotes sarissa sp. nov., holotype, ♂ (BMNH, slide 95, box E0026011). Habitus, dorsal and ventral views." captionText-1="Fig. 12. Goniocotes sarissa sp. nov. Female habitus, dorsal and ventral views." figureDoi-0="http://doi.org/10.5281/zenodo.14009206" figureDoi-1="http://doi.org/10.5281/zenodo.14009208" httpUri-0="https://zenodo.org/record/14009206/files/figure.png" httpUri-1="https://zenodo.org/record/14009208/files/figure.png" pageId="17" pageNumber="18">Figs 11–12</figureCitation>
), but fused to pronotum in
<taxonomicName id="4C14EAE4A848687FFE6DF2CDFDDDFEE5" authorityName="Gustafsson, Tian &amp; Zou" authorityYear="2021" box="[486,625,1505,1531]" class="Insecta" family="Philopteridae" genus="Goniocotes" kingdom="Animalia" order="Psocodea" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="kristinae">
<emphasis id="B9604D75A848687FFE6DF2CDFDDDFEE5" box="[486,625,1505,1531]" italics="true" pageId="17" pageNumber="18">G. kristinae</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BAB9167A848687FFF36F10DFD0BFDD9" blockId="17.[189,1399,1569,1735]" pageId="17" pageNumber="18">
Reticulated lateral accessory sternal plates are also known in
<taxonomicName id="4C14EAE4A848687FFC07F10EFBC3FD25" authorityName="Giebel" authorityYear="1874" box="[908,1135,1569,1596]" class="Insecta" family="Philopteridae" genus="Goniocotes" kingdom="Animalia" order="Psocodea" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="albidus">
<emphasis id="B9604D75A848687FFC07F10EFBC3FD25" box="[908,1135,1569,1596]" italics="true" pageId="17" pageNumber="18">Goniocotes albidus</emphasis>
</taxonomicName>
, where they are found in both male and female. However, in
<taxonomicName id="4C14EAE4A848687FFD03F169FCAFFD40" authorityName="Giebel" authorityYear="1874" box="[648,771,1604,1631]" class="Insecta" family="Philopteridae" genus="Goniocotes" kingdom="Animalia" order="Psocodea" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="albidus">
<emphasis id="B9604D75A848687FFD03F169FCAFFD40" box="[648,771,1604,1631]" italics="true" pageId="17" pageNumber="18">G. albidus</emphasis>
</taxonomicName>
the plates are small and rounded or oval, whereas in
<taxonomicName id="4C14EAE4A848687FFF35F14BFE98FD9F" authorityName="Gustafsson &amp; Tian &amp; Ren &amp; Li &amp; Sun &amp; Zou" authorityYear="2024" box="[190,308,1639,1665]" class="Insecta" family="Philopteridae" genus="Goniocotes" kingdom="Animalia" order="Psocodea" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="sarissa" status="sp. nov.">
<emphasis id="B9604D75A848687FFF35F14BFE98FD9F" box="[190,308,1639,1665]" italics="true" pageId="17" pageNumber="18">G. sarissa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A253F00EA848687FFEB1F14BFE38FD9F" box="[314,404,1639,1665]" pageId="17" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
they are elongated. Moreover,
<taxonomicName id="4C14EAE4A848687FFD74F14BFCD4FD9F" authorityName="Giebel" authorityYear="1874" box="[767,888,1639,1665]" class="Insecta" family="Philopteridae" genus="Goniocotes" kingdom="Animalia" order="Psocodea" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="albidus">
<emphasis id="B9604D75A848687FFD74F14BFCD4FD9F" box="[767,888,1639,1665]" italics="true" pageId="17" pageNumber="18">G. albidus</emphasis>
</taxonomicName>
has a rounded, not flaring, temporal corner, males have dense rows of
<emphasis id="B9604D75A848687FFE75F1A6FD8CFDBD" box="[510,544,1674,1699]" italics="true" pageId="17" pageNumber="18">tps</emphasis>
on all of tergopleurites II–VI and only a single
<emphasis id="B9604D75A848687FFBEFF1A6FB33FDBD" box="[1124,1183,1674,1699]" italics="true" pageId="17" pageNumber="18">smns</emphasis>
on each side, and females lack
<emphasis id="B9604D75A848687FFED3F181FE20FDD8" box="[344,396,1709,1734]" italics="true" pageId="17" pageNumber="18">psps</emphasis>
on tergopleurites II–III.
</paragraph>
<paragraph id="8BAB9167A848687CFF36F1C2FCB6FA76" blockId="17.[187,1399,1773,2008]" lastBlockId="18.[189,1398,265,361]" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18">
<taxonomicName id="4C14EAE4A848687FFF36F1C2FE60FC19" authorityName="Nitzsch" authorityYear="1866" box="[189,460,1773,1800]" class="Insecta" family="Philopteridae" genus="Goniocotes" kingdom="Animalia" order="Psocodea" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="diplogonus">
<emphasis id="B9604D75A848687FFF36F1C2FE60FC19" box="[189,460,1773,1800]" italics="true" pageId="17" pageNumber="18">Goniocotes diplogonus</emphasis>
</taxonomicName>
is probably also closely related to
<taxonomicName id="4C14EAE4A848687FFCE1F1C2FC4DFC19" authorityName="Gustafsson &amp; Tian &amp; Ren &amp; Li &amp; Sun &amp; Zou" authorityYear="2024" box="[874,993,1774,1800]" class="Insecta" family="Philopteridae" genus="Goniocotes" kingdom="Animalia" order="Psocodea" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="sarissa" status="sp. nov.">
<emphasis id="B9604D75A848687FFCE1F1C2FC4DFC19" box="[874,993,1774,1800]" italics="true" pageId="17" pageNumber="18">G. sarissa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A253F00EA848687FFC63F1C2FBE8FC16" box="[1000,1092,1774,1800]" pageId="17" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="4C14EAE4A848687FFBF5F1C2FB5BFC19" box="[1150,1271,1773,1800]" class="Insecta" family="Philopteridae" genus="Dictyocotes" kingdom="Animalia" order="Psocodea" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="rolandi">
<emphasis id="B9604D75A848687FFBF5F1C2FB5BFC19" box="[1150,1271,1773,1800]" italics="true" pageId="17" pageNumber="18">G. rolandi</emphasis>
</taxonomicName>
, and share the reticulated lateral accessory sternal plates. No specimens of this species were examined.
<bibRefCitation id="EF85EC96A848687FFB6FF03CFADFFC34" author="Keler" box="[1252,1395,1808,1834]" firstAuthor="Keler" pageId="17" pageNumber="18" pagination="1 - 254" refId="ref14956" refString="Keler S. von 1940. [1939] Baustoff zu einer Monographie der Mallophagen. II. Teil: Uberfamilie der Nirmoidea (1) Die Familien Trichophilopteridae, Goniodidae, Heptapsogastridae. Nova Acta Leopoldiana (N. F.) 8: 1 - 254." type="journal article" year="1940">Kéler (1940</bibRefCitation>
: fig. 88) illustrated the male and female, but noted that his male was morphologically aberrant. Based on Kéler’s illustrations,
<taxonomicName id="4C14EAE4A848687FFE6AF07AFDF7FC71" authorityName="Gustafsson &amp; Tian &amp; Ren &amp; Li &amp; Sun &amp; Zou" authorityYear="2024" box="[481,603,1878,1904]" class="Insecta" family="Philopteridae" genus="Goniocotes" kingdom="Animalia" order="Psocodea" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="sarissa">
<emphasis id="B9604D75A848687FFE6AF07AFDF7FC71" box="[481,603,1878,1904]" italics="true" pageId="17" pageNumber="18">G. sarissa</emphasis>
</taxonomicName>
can be separated from
<taxonomicName id="4C14EAE4A848687FFCFCF07AFBB3FC71" authorityName="Nitzsch" authorityYear="1866" box="[887,1055,1877,1904]" class="Insecta" family="Philopteridae" genus="Goniocotes" kingdom="Animalia" order="Psocodea" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="diplogonus">
<emphasis id="B9604D75A848687FFCFCF07AFBB3FC71" box="[887,1055,1877,1904]" italics="true" pageId="17" pageNumber="18">G. diplogonus</emphasis>
</taxonomicName>
by the following characters: temples more flaring and frons more flattened in
<taxonomicName id="4C14EAE4A848687FFD7CF054FCC1FC8C" authorityName="Gustafsson &amp; Tian &amp; Ren &amp; Li &amp; Sun &amp; Zou" authorityYear="2024" box="[759,877,1912,1938]" class="Insecta" family="Philopteridae" genus="Goniocotes" kingdom="Animalia" order="Psocodea" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="sarissa">
<emphasis id="B9604D75A848687FFD7CF054FCC1FC8C" box="[759,877,1912,1938]" italics="true" pageId="17" pageNumber="18">G. sarissa</emphasis>
</taxonomicName>
(
<figureCitation id="132F8DE2A848687FFCF6F054FC7DFC8C" box="[893,977,1912,1938]" captionStart="Figs 13–15" captionStartId="20.[189,242,1963,1989]" captionTargetBox="[342,1235,650,1888]" captionTargetId="figure-39@20.[340,1247,636,1919]" captionTargetPageId="20" captionText="Figs 13–15. Goniocotes sarissa sp. nov. 13–14. Holotype, ♂ (BMNH, slide 95, box E0026011). 13. Head, dorsal and ventral views. 14. Genitalia, dorsal view. 15. Female genitalia, ventral view." figureDoi="http://doi.org/10.5281/zenodo.14009210" httpUri="https://zenodo.org/record/14009210/files/figure.png" pageId="17" pageNumber="18">Fig. 13</figureCitation>
) than in
<taxonomicName id="4C14EAE4A848687FFBB3F054FB71FC8C" authorityName="Nitzsch" authorityYear="1866" box="[1080,1245,1912,1938]" class="Insecta" family="Philopteridae" genus="Goniocotes" kingdom="Animalia" order="Psocodea" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="diplogonus">
<emphasis id="B9604D75A848687FFBB3F054FB71FC8C" box="[1080,1245,1912,1938]" italics="true" pageId="17" pageNumber="18">G. diplogonus</emphasis>
</taxonomicName>
(but temples drawn differently on each side of the head of the male by
<bibRefCitation id="EF85EC96A848687FFCE0F0B6FC58FCAB" author="Keler" box="[875,1012,1946,1973]" firstAuthor="Keler" pageId="17" pageNumber="18" pagination="1 - 254" refId="ref14956" refString="Keler S. von 1940. [1939] Baustoff zu einer Monographie der Mallophagen. II. Teil: Uberfamilie der Nirmoidea (1) Die Familien Trichophilopteridae, Goniodidae, Heptapsogastridae. Nova Acta Leopoldiana (N. F.) 8: 1 - 254." type="journal article" year="1940">Kéler 1940</bibRefCitation>
); male pteronotum with patch of
<emphasis id="B9604D75A848687FFF35F092FF55FCC9" box="[190,249,1982,2007]" italics="true" pageId="17" pageNumber="18">smns</emphasis>
in
<taxonomicName id="4C14EAE4A848687FFEABF092FE3BFCC9" authorityName="Gustafsson &amp; Tian &amp; Ren &amp; Li &amp; Sun &amp; Zou" authorityYear="2024" box="[288,407,1982,2008]" class="Insecta" family="Philopteridae" genus="Goniocotes" kingdom="Animalia" order="Psocodea" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="sarissa">
<emphasis id="B9604D75A848687FFEABF092FE3BFCC9" box="[288,407,1982,2008]" italics="true" pageId="17" pageNumber="18">G. sarissa</emphasis>
</taxonomicName>
(
<figureCitation id="132F8DE2A848687FFE23F091FE51FCC9" box="[424,509,1981,2007]" captionStart="Fig" captionStartId="18.[189,232,1963,1989]" captionTargetBox="[427,1218,685,1921]" captionTargetId="figure-112@18.[340,1247,671,1926]" captionTargetPageId="18" captionText="Fig. 11. Goniocotes sarissa sp. nov., holotype, ♂ (BMNH, slide 95, box E0026011). Habitus, dorsal and ventral views." figureDoi="http://doi.org/10.5281/zenodo.14009206" httpUri="https://zenodo.org/record/14009206/files/figure.png" pageId="17" pageNumber="18">Fig. 11</figureCitation>
), but this is apparently lacking in
<taxonomicName id="4C14EAE4A848687FFC18F092FB95FCC9" authorityName="Nitzsch" authorityYear="1866" box="[915,1081,1981,2008]" class="Insecta" family="Philopteridae" genus="Goniocotes" kingdom="Animalia" order="Psocodea" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="diplogonus">
<emphasis id="B9604D75A848687FFC18F092FB95FCC9" box="[915,1081,1981,2008]" italics="true" pageId="17" pageNumber="18">G. diplogonus</emphasis>
</taxonomicName>
; male tergopleurites V–VI with
<emphasis id="B9604D75A84B687CFF76F626FEB8FA3D" box="[253,276,266,291]" italics="true" pageId="18" pageNumber="19">ss</emphasis>
in
<taxonomicName id="4C14EAE4A84B687CFECAF625FE10FA3D" authorityName="Gustafsson &amp; Tian &amp; Ren &amp; Li &amp; Sun &amp; Zou" authorityYear="2024" box="[321,444,265,291]" class="Insecta" family="Philopteridae" genus="Goniocotes" kingdom="Animalia" order="Psocodea" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="sarissa">
<emphasis id="B9604D75A84B687CFECAF625FE10FA3D" box="[321,444,265,291]" italics="true" pageId="18" pageNumber="19">G. sarissa</emphasis>
</taxonomicName>
(
<figureCitation id="132F8DE2A84B687CFE5AF625FD86FA3D" box="[465,554,265,291]" captionStart="Fig" captionStartId="18.[189,232,1963,1989]" captionTargetBox="[427,1218,685,1921]" captionTargetId="figure-112@18.[340,1247,671,1926]" captionTargetPageId="18" captionText="Fig. 11. Goniocotes sarissa sp. nov., holotype, ♂ (BMNH, slide 95, box E0026011). Habitus, dorsal and ventral views." figureDoi="http://doi.org/10.5281/zenodo.14009206" httpUri="https://zenodo.org/record/14009206/files/figure.png" pageId="18" pageNumber="19">Fig. 11</figureCitation>
), but these apparently lacking in
<taxonomicName id="4C14EAE4A84B687CFC42F625FBDFFA3D" authorityName="Nitzsch" authorityYear="1866" box="[969,1139,265,291]" class="Insecta" family="Philopteridae" genus="Goniocotes" kingdom="Animalia" order="Psocodea" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="diplogonus">
<emphasis id="B9604D75A84B687CFC42F625FBDFFA3D" box="[969,1139,265,291]" italics="true" pageId="18" pageNumber="19">G. diplogonus</emphasis>
</taxonomicName>
;
<emphasis id="B9604D75A84B687CFB0CF626FB05FA3D" box="[1159,1193,266,291]" italics="true" pageId="18" pageNumber="19">tps</emphasis>
present on male tergopleurite IV in
<taxonomicName id="4C14EAE4A84B687CFE11F600FDBCFA5B" authorityName="Gustafsson &amp; Tian &amp; Ren &amp; Li &amp; Sun &amp; Zou" authorityYear="2024" box="[410,528,300,326]" class="Insecta" family="Philopteridae" genus="Goniocotes" kingdom="Animalia" order="Psocodea" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="sarissa">
<emphasis id="B9604D75A84B687CFE11F600FDBCFA5B" box="[410,528,300,326]" italics="true" pageId="18" pageNumber="19">G. sarissa</emphasis>
</taxonomicName>
(
<figureCitation id="132F8DE2A84B687CFDABF600FDD8FA58" box="[544,628,300,326]" captionStart="Fig" captionStartId="18.[189,232,1963,1989]" captionTargetBox="[427,1218,685,1921]" captionTargetId="figure-112@18.[340,1247,671,1926]" captionTargetPageId="18" captionText="Fig. 11. Goniocotes sarissa sp. nov., holotype, ♂ (BMNH, slide 95, box E0026011). Habitus, dorsal and ventral views." figureDoi="http://doi.org/10.5281/zenodo.14009206" httpUri="https://zenodo.org/record/14009206/files/figure.png" pageId="18" pageNumber="19">Fig. 11</figureCitation>
), but apparently absent in
<taxonomicName id="4C14EAE4A84B687CFC2EF600FBE6FA58" authorityName="Nitzsch" authorityYear="1866" box="[933,1098,300,326]" class="Insecta" family="Philopteridae" genus="Goniocotes" kingdom="Animalia" order="Psocodea" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="diplogonus">
<emphasis id="B9604D75A84B687CFC2EF600FBE6FA58" box="[933,1098,300,326]" italics="true" pageId="18" pageNumber="19">G. diplogonus</emphasis>
</taxonomicName>
. However,
<taxonomicName id="4C14EAE4A84B687CFB5BF600FAD9FA58" authorityName="Nitzsch" authorityYear="1866" box="[1232,1397,300,326]" class="Insecta" family="Philopteridae" genus="Goniocotes" kingdom="Animalia" order="Psocodea" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="diplogonus">
<emphasis id="B9604D75A84B687CFB5BF600FAD9FA58" box="[1232,1397,300,326]" italics="true" pageId="18" pageNumber="19">G. diplogonus</emphasis>
</taxonomicName>
needs to be redescribed to confirm these characters.
</paragraph>
</subSubSection>
<subSubSection id="C30EC2ECA84B687CFF36F6A3FB85F902" pageId="18" pageNumber="19" type="etymology">
<paragraph id="8BAB9167A84B687CFF36F6A3FEE8FAB7" blockId="18.[189,324,399,425]" box="[189,324,399,425]" pageId="18" pageNumber="19">
<heading id="D0E3260BA84B687CFF36F6A3FEE8FAB7" bold="true" box="[189,324,399,425]" fontSize="11" level="3" pageId="18" pageNumber="19" reason="3">
<emphasis id="B9604D75A84B687CFF36F6A3FEE8FAB7" bold="true" box="[189,324,399,425]" pageId="18" pageNumber="19">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BAB9167A84B687CFF36F691FB85F902" blockId="18.[189,1399,445,541]" pageId="18" pageNumber="19">
The specific name is derived from
<taxonomicName id="4C14EAE4A84B687CFDFFF691FD7DFAC6" authority="Greek" authorityName="Gustafsson &amp; Tian &amp; Ren &amp; Li &amp; Sun &amp; Zou" authorityYear="2024" box="[628,721,445,472]" class="Insecta" family="Philopteridae" genus="Goniocotes" kingdom="Animalia" order="Psocodea" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="sarissa">
‘
<emphasis id="B9604D75A84B687CFDF5F692FD7DFAC6" box="[638,721,446,472]" italics="true" pageId="18" pageNumber="19">sárissa</emphasis>
</taxonomicName>
’, Greek for the long pikes used in traditional phalanx formations. This refers to the patch of
<emphasis id="B9604D75A84B687CFD0DF6CDFD6DFAE4" box="[646,705,481,506]" italics="true" pageId="18" pageNumber="19">smns</emphasis>
on the male pteronotum combined with the rows of
<emphasis id="B9604D75A84B687CFABBF6CDFAFEFAE4" box="[1328,1362,481,506]" italics="true" pageId="18" pageNumber="19">tps</emphasis>
on male tergopleurites II–IV, which are reminiscent of a loose, small phalanx.
</paragraph>
</subSubSection>
<caption id="DF6BC1EFA84B687CFF36F087FECDFCF6" ID-DOI="http://doi.org/10.5281/zenodo.14009206" ID-Zenodo-Dep="14009206" httpUri="https://zenodo.org/record/14009206/files/figure.png" pageId="18" pageNumber="19" startId="18.[189,232,1963,1989]" targetBox="[427,1218,685,1921]" targetPageId="18">
<paragraph id="8BAB9167A84B687CFF36F087FECDFCF6" blockId="18.[188,1397,1963,2024]" pageId="18" pageNumber="19">
<emphasis id="B9604D75A84B687CFF36F087FEB4FCDB" bold="true" box="[189,280,1963,1989]" pageId="18" pageNumber="19">Fig. 11.</emphasis>
<taxonomicName id="4C14EAE4A84B687CFE95F080FE51FCDB" authorityName="Gustafsson &amp; Tian &amp; Ren &amp; Li &amp; Sun &amp; Zou" authorityYear="2024" box="[286,509,1964,1990]" class="Insecta" family="Philopteridae" genus="Goniocotes" kingdom="Animalia" order="Psocodea" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="sarissa" status="sp. nov.">
<emphasis id="B9604D75A84B687CFE95F080FE51FCDB" box="[286,509,1964,1990]" italics="true" pageId="18" pageNumber="19">Goniocotes sarissa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A253F00EA84B687CFD8FF080FDCCFCD8" box="[516,608,1964,1990]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, ♂ (BMNH, slide 95, box E0026011). Habitus, dorsal and ventral views.
</paragraph>
</caption>
<subSubSection id="C30EC2ECA84A687DFF36F625FBA6F978" pageId="19" pageNumber="20" type="materials_examined">
<paragraph id="8BAB9167A84A687DFF36F625FEC0FA3C" blockId="19.[189,364,264,291]" box="[189,364,264,291]" pageId="19" pageNumber="20">
<heading id="D0E3260BA84A687DFF36F625FEC0FA3C" bold="true" box="[189,364,264,291]" fontSize="11" level="3" pageId="19" pageNumber="20" reason="3">
<emphasis id="B9604D75A84A687DFF36F625FEC0FA3C" bold="true" box="[189,364,264,291]" pageId="19" pageNumber="20">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="8BAB9167A84A687DFF61F61BFCD0FA89" blockId="19.[187,1399,311,408]" pageId="19" pageNumber="20">
<materialsCitation id="3B7C9B3AA84A687DFF61F61BFCD0FA89" collectingDate="1990-10-30" collectionCode="BMNH" country="China" location="Fujian Province" pageId="19" pageNumber="20" specimenCount="1" specimenCount-male="1" stateProvince="Fujian Province" typeStatus="holotype">
<emphasis id="B9604D75A84A687DFF61F61BFEF0FA4F" bold="true" box="[234,348,311,337]" pageId="19" pageNumber="20">
<typeStatus id="54AF2FC5A84A687DFF61F61BFEF0FA4F" box="[234,348,311,337]" pageId="19" pageNumber="20" type="holotype">Holotype</typeStatus>
</emphasis>
(ex
<taxonomicName id="4C14EAE4A84A687DFE1AF614FDFFFA4F" baseAuthorityName="Gould" baseAuthorityYear="1857" box="[401,595,311,337]" class="Aves" family="Phasianidae" genus="Tragopan" kingdom="Animalia" order="Galliformes" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="caboti">
<emphasis id="B9604D75A84A687DFE1AF614FDFFFA4F" box="[401,595,311,337]" italics="true" pageId="19" pageNumber="20">Tragopan caboti</emphasis>
</taxonomicName>
)
<collectingCountry id="F303D1F7A84A687DFF36F676FEB4FA6A" box="[189,280,346,372]" name="China" pageId="19" pageNumber="20">CHINA</collectingCountry>
•
<specimenCount id="9D125AEEA84A687DFEA6F672FEEFFA6A" box="[301,323,350,372]" count="1" pageId="19" pageNumber="20" type="male">♂</specimenCount>
;
<collectingRegion id="49D05F85A84A687DFEDAF676FDA1FA6A" box="[337,525,346,372]" country="China" name="Fujian" pageId="19" pageNumber="20">Fujian Province</collectingRegion>
;
<date id="FFAAB7A7A84A687DFD9CF676FD03FA6A" box="[535,687,346,372]" pageId="19" pageNumber="20" value="1990-10-30">
<collectingDate id="EFEE4E4FA84A687DFD9CF676FD03FA6A" box="[535,687,346,372]" pageId="19" pageNumber="20" value="1990-10-30">30 Oct. 1990</collectingDate>
</date>
; collector unknown; box E0026011, slide 95;
<collectionCode id="ED0509A2A84A687DFB35F677FABAFA6B" box="[1214,1302,347,373]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="19" pageNumber="20">BMNH</collectionCode>
[second male from right in top row, marked with black dot on slide].
</materialsCitation>
</paragraph>
<paragraph id="8BAB9167A84A687DFF61F693FBA6F978" blockId="19.[189,1399,447,614]" pageId="19" pageNumber="20">
<materialsCitation id="3B7C9B3AA84A687DFF61F693FC3CFAE3" collectingDate="1990-10-30" collectionCode="BMNH" country="China" location="CHINA" pageId="19" pageNumber="20" specimenCount="38" specimenCount-female="21" specimenCount-male="17" stateProvince="Fujian Province" typeStatus="paratype">
<emphasis id="B9604D75A84A687DFF61F693FECBFAC7" bold="true" box="[234,359,447,473]" pageId="19" pageNumber="20">
<typeStatus id="54AF2FC5A84A687DFF61F693FECBFAC7" box="[234,359,447,473]" pageId="19" pageNumber="20" type="paratype">Paratypes</typeStatus>
</emphasis>
(ex
<taxonomicName id="4C14EAE4A84A687DFE17F6ECFDF2FAC7" baseAuthorityName="Gould" baseAuthorityYear="1857" box="[412,606,447,473]" class="Aves" family="Phasianidae" genus="Tragopan" kingdom="Animalia" order="Galliformes" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="caboti">
<emphasis id="B9604D75A84A687DFE17F6ECFDF2FAC7" box="[412,606,447,473]" italics="true" pageId="19" pageNumber="20">Tragopan caboti</emphasis>
</taxonomicName>
)
<collectingCountry id="F303D1F7A84A687DFF36F6CEFEB4FAE2" box="[189,280,482,508]" name="China" pageId="19" pageNumber="20">CHINA</collectingCountry>
•
<specimenCount id="9D125AEEA84A687DFEB8F6CEFE29FAE2" box="[307,389,482,508]" count="17" pageId="19" pageNumber="20" type="male">17 ♂♂</specimenCount>
,
<specimenCount id="9D125AEEA84A687DFE1EF6CEFE44FAE3" box="[405,488,482,509]" count="21" pageId="19" pageNumber="20" type="female">21 ♀♀</specimenCount>
; same data as for holotype;
<collectionCode id="ED0509A2A84A687DFCB3F6CFFC3CFAE3" box="[824,912,483,509]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="19" pageNumber="20">BMNH</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B7C9B3AA84A687DFC20F6CEFC0FF93E" collectingDate="1990-09-29" collectionCode="BMNH" location="Fujian Province" pageId="19" pageNumber="20" specimenCount="9" specimenCount-female="4" specimenCount-male="5" stateProvince="Fujian Province" typeStatus="paratype">
<specimenCount id="9D125AEEA84A687DFC20F6CEFC42FAE2" box="[939,1006,482,509]" count="5" pageId="19" pageNumber="20" type="male">5 ♂♂</specimenCount>
,
<specimenCount id="9D125AEEA84A687DFC75F6CEFBEEFAE3" box="[1022,1090,482,509]" count="4" pageId="19" pageNumber="20" type="female">4 ♀♀</specimenCount>
;
<collectingRegion id="49D05F85A84A687DFBDAF6CEFABDFAE2" box="[1105,1297,482,508]" country="China" name="Fujian" pageId="19" pageNumber="20">Fujian Province</collectingRegion>
;
<date id="FFAAB7A7A84A687DFA96F6CEFF57F901" pageId="19" pageNumber="20" value="1990-09-29">
<collectingDate id="EFEE4E4FA84A687DFA96F6CEFF57F901" pageId="19" pageNumber="20" value="1990-09-29">29 Sep. 1990</collectingDate>
</date>
; collector unknown; box E0026136, slides 24–25;
<collectionCode id="ED0509A2A84A687DFCC0F52AFC0FF93E" box="[843,931,518,544]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="19" pageNumber="20">BMNH</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B7C9B3AA84A687DFC31F529FC65F95D" collectingDate="1997-01-07" collectionCode="BMNH" location="Jianou" municipality="Jianou" pageId="19" pageNumber="20" specimenCount="4" specimenCount-female="2" specimenCount-male="2" stateProvince="Fujian Province" typeStatus="paratype">
<specimenCount id="9D125AEEA84A687DFC31F529FC50F901" box="[954,1020,517,543]" count="2" pageId="19" pageNumber="20" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="9D125AEEA84A687DFB81F529FBE0F93E" box="[1034,1100,517,544]" count="2" pageId="19" pageNumber="20" type="female">2 ♀♀</specimenCount>
;
<collectingRegion id="49D05F85A84A687DFBD1F529FABAF901" box="[1114,1302,517,543]" country="China" name="Fujian" pageId="19" pageNumber="20">Fujian Province</collectingRegion>
,
<collectingMunicipality id="6BCF0B1DA84A687DFAA9F529FADEF901" box="[1314,1394,517,543]" country="China" name="Jianou" pageId="19" pageNumber="20" stateProvince="Fujian">Jianou</collectingMunicipality>
;
<date id="FFAAB7A7A84A687DFF36F505FEE1F95D" box="[189,333,553,579]" pageId="19" pageNumber="20" value="1997-01-07">
<collectingDate id="EFEE4E4FA84A687DFF36F505FEE1F95D" box="[189,333,553,579]" pageId="19" pageNumber="20" value="1997-01-07">7 Jan. 1997</collectingDate>
</date>
; collector unknown; box E0026195, slide 4;
<collectionCode id="ED0509A2A84A687DFCFAF505FC65F95D" box="[881,969,553,579]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="19" pageNumber="20">BMNH</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B7C9B3AA84A687DFC63F504FBAAF978" collectingDate="1995-01-10" collectionCode="BMNH" location="Guangxi Province" pageId="19" pageNumber="20" specimenCount="12" specimenCount-female="7" specimenCount-male="5" stateProvince="Guangxi Province" typeStatus="paratype">
<specimenCount id="9D125AEEA84A687DFC63F504FB81F95C" box="[1000,1069,552,579]" count="5" pageId="19" pageNumber="20" type="male">5 ♂♂</specimenCount>
,
<specimenCount id="9D125AEEA84A687DFBB4F505FB29F95D" box="[1087,1157,553,579]" count="7" pageId="19" pageNumber="20" type="female">7 ♀♀</specimenCount>
;
<collectingRegion id="49D05F85A84A687DFB1CF504FADEF95C" box="[1175,1394,552,578]" country="China" name="Guangxi" pageId="19" pageNumber="20">Guangxi Province</collectingRegion>
;
<date id="FFAAB7A7A84A687DFF36F560FEF9F978" box="[189,341,587,614]" pageId="19" pageNumber="20" value="1995-01-10">
<collectingDate id="EFEE4E4FA84A687DFF36F560FEF9F978" box="[189,341,587,614]" pageId="19" pageNumber="20" value="1995-01-10">10 Jan. 1995</collectingDate>
</date>
; collector unknown; box E0026136, slides 20–22;
<collectionCode id="ED0509A2A84A687DFC21F560FBAAF978" box="[938,1030,588,614]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="19" pageNumber="20">BMNH</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="DF6BC1EFA84A687DFF36F0E2FBFDFCF6" ID-DOI="http://doi.org/10.5281/zenodo.14009208" ID-Zenodo-Dep="14009208" box="[189,1105,1998,2025]" httpUri="https://zenodo.org/record/14009208/files/figure.png" pageId="19" pageNumber="20" startId="19.[189,232,1998,2024]" targetBox="[416,1161,676,1933]" targetPageId="19">
<paragraph id="8BAB9167A84A687DFF36F0E2FBFDFCF6" blockId="19.[189,1105,1998,2025]" box="[189,1105,1998,2025]" pageId="19" pageNumber="20">
<emphasis id="B9604D75A84A687DFF36F0E2FEB6FCF6" bold="true" box="[189,282,1998,2024]" pageId="19" pageNumber="20">Fig. 12.</emphasis>
<taxonomicName id="4C14EAE4A84A687DFEA9F0E3FDADFCF6" authorityName="Gustafsson &amp; Tian &amp; Ren &amp; Li &amp; Sun &amp; Zou" authorityYear="2024" box="[290,513,1999,2025]" class="Insecta" family="Philopteridae" genus="Goniocotes" kingdom="Animalia" order="Psocodea" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="sarissa" status="sp. nov.">
<emphasis id="B9604D75A84A687DFEA9F0E3FDADFCF6" box="[290,513,1999,2025]" italics="true" pageId="19" pageNumber="20">Goniocotes sarissa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A253F00EA84A687DFD82F0E3FDC8FCF7" box="[521,612,1999,2025]" pageId="19" pageNumber="20" rank="species">sp. nov.</taxonomicNameLabel>
Female habitus, dorsal and ventral views.
</paragraph>
</caption>
<subSubSection id="C30EC2ECA84D687AFF36F625FCB6FA4C" pageId="20" pageNumber="21" type="biology_ecology">
<paragraph id="8BAB9167A84D687AFF36F625FE9AFA3D" blockId="20.[189,310,265,291]" box="[189,310,265,291]" pageId="20" pageNumber="21">
<heading id="D0E3260BA84D687AFF36F625FE9AFA3D" bold="true" box="[189,310,265,291]" fontSize="11" level="3" pageId="20" pageNumber="21" reason="3">
<emphasis id="B9604D75A84D687AFF36F625FE9AFA3D" bold="true" box="[189,310,265,291]" pageId="20" pageNumber="21">Type host</emphasis>
</heading>
</paragraph>
<paragraph id="8BAB9167A84D687AFF36F614FCB6FA4C" blockId="20.[189,794,311,338]" box="[189,794,311,338]" pageId="20" pageNumber="21">
<taxonomicName id="4C14EAE4A84D687AFF36F614FD81FA4C" authority="(Gould, 1857)" baseAuthorityName="Gould" baseAuthorityYear="1857" box="[189,557,311,338]" class="Aves" family="Phasianidae" genus="Tragopan" kingdom="Animalia" order="Galliformes" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="caboti">
<emphasis id="B9604D75A84D687AFF36F614FED3FA4F" box="[189,383,311,337]" italics="true" pageId="20" pageNumber="21">Tragopan caboti</emphasis>
(
<bibRefCitation id="EF85EC96A84D687AFE04F61BFD89FA4C" author="Gould" box="[399,549,311,338]" firstAuthor="Gould" pageId="20" pageNumber="21" pagination="161 - 162" refId="ref14013" refString="Gould J. 1857. [Untitled]. Proceedings of the Zoological Society of London 25: 161 - 162. Available from https: // www. biodiversitylibrary. org / page / 32451306 [accessed 22 Jan. 2024]." type="book chapter" year="1857">Gould, 1857</bibRefCitation>
)
</taxonomicName>
– Cabot’s tragopan.
</paragraph>
</subSubSection>
<subSubSection id="C30EC2ECA84D687AFF36F655FC13FADF" pageId="20" pageNumber="21" type="distribution">
<paragraph id="8BAB9167A84D687AFF36F655FEF1FA8C" blockId="20.[189,349,376,403]" box="[189,349,376,403]" pageId="20" pageNumber="21">
<heading id="D0E3260BA84D687AFF36F655FEF1FA8C" bold="true" box="[189,349,376,403]" fontSize="11" level="3" pageId="20" pageNumber="21" reason="3">
<emphasis id="B9604D75A84D687AFF36F655FEF1FA8C" bold="true" box="[189,349,376,403]" pageId="20" pageNumber="21">Type locality</emphasis>
</heading>
</paragraph>
<paragraph id="8BAB9167A84D687AFF36F68BFC13FADF" blockId="20.[189,959,423,450]" box="[189,959,423,450]" pageId="20" pageNumber="21">
<collectingRegion id="49D05F85A84D687AFF36F68BFED6FADF" box="[189,378,423,449]" country="China" name="Fujian" pageId="20" pageNumber="21">Fujian Province</collectingRegion>
,
<collectingCountry id="F303D1F7A84D687AFE0DF68BFE7CFADF" box="[390,464,423,449]" name="China" pageId="20" pageNumber="21">China</collectingCountry>
; no more detailed locality given on slides.
</paragraph>
</subSubSection>
<caption id="DF6BC1EFA84D687AFF36F087FA9EFCF6" ID-DOI="http://doi.org/10.5281/zenodo.14009210" ID-Zenodo-Dep="14009210" httpUri="https://zenodo.org/record/14009210/files/figure.png" pageId="20" pageNumber="21" startId="20.[189,242,1963,1989]" targetBox="[342,1235,650,1888]" targetPageId="20">
<paragraph id="8BAB9167A84D687AFF36F087FA9EFCF6" blockId="20.[189,1399,1963,2025]" pageId="20" pageNumber="21">
<emphasis id="B9604D75A84D687AFF36F087FEFDFCDB" bold="true" box="[189,337,1963,1989]" pageId="20" pageNumber="21">Figs 13–15.</emphasis>
<taxonomicName id="4C14EAE4A84D687AFED5F080FDEBFCDB" authorityName="Gustafsson &amp; Tian &amp; Ren &amp; Li &amp; Sun &amp; Zou" authorityYear="2024" box="[350,583,1964,1990]" class="Insecta" family="Philopteridae" genus="Goniocotes" kingdom="Animalia" order="Psocodea" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="sarissa" status="sp. nov.">
<emphasis id="B9604D75A84D687AFED5F080FDEBFCDB" box="[350,583,1964,1990]" italics="true" pageId="20" pageNumber="21">Goniocotes sarissa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A253F00EA84D687AFDDEF080FD1AFCD8" box="[597,694,1964,1990]" pageId="20" pageNumber="21" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B9604D75A84D687AFD48F087FCA1FCDB" bold="true" box="[707,781,1963,1989]" pageId="20" pageNumber="21">13–14</emphasis>
. Holotype, ♂ (BMNH, slide 95, box E0026011).
<emphasis id="B9604D75A84D687AFF36F0E2FF76FCF6" bold="true" box="[189,218,1998,2024]" pageId="20" pageNumber="21">13</emphasis>
. Head, dorsal and ventral views.
<emphasis id="B9604D75A84D687AFDE9F0E2FDD3FCF6" bold="true" box="[610,639,1998,2024]" pageId="20" pageNumber="21">14</emphasis>
. Genitalia, dorsal view.
<emphasis id="B9604D75A84D687AFC16F0E2FC16FCF6" bold="true" box="[925,954,1998,2024]" pageId="20" pageNumber="21">15</emphasis>
. Female genitalia, ventral view.
</paragraph>
</caption>
<subSubSection id="C30EC2ECA84C687BFF36F624FDA7FF79" pageId="21" pageNumber="22" type="description">
<paragraph id="8BAB9167A84C687BFF36F624FEE2FA3C" blockId="21.[189,334,264,290]" box="[189,334,264,290]" pageId="21" pageNumber="22">
<heading id="D0E3260BA84C687BFF36F624FEE2FA3C" bold="true" box="[189,334,264,290]" fontSize="11" level="3" pageId="21" pageNumber="22" reason="3">
<emphasis id="B9604D75A84C687BFF36F624FEE2FA3C" bold="true" box="[189,334,264,290]" pageId="21" pageNumber="22">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BAB9167A84C687BFF36F614FB47FAA5" blockId="21.[189,1399,311,443]" pageId="21" pageNumber="22">
Frons slightly flattened (
<figureCitation id="132F8DE2A84C687BFE6CF61BFDEEFA4F" box="[487,578,311,337]" captionStart="Figs 13–15" captionStartId="20.[189,242,1963,1989]" captionTargetBox="[342,1235,650,1888]" captionTargetId="figure-39@20.[340,1247,636,1919]" captionTargetPageId="20" captionText="Figs 13–15. Goniocotes sarissa sp. nov. 13–14. Holotype, ♂ (BMNH, slide 95, box E0026011). 13. Head, dorsal and ventral views. 14. Genitalia, dorsal view. 15. Female genitalia, ventral view." figureDoi="http://doi.org/10.5281/zenodo.14009210" httpUri="https://zenodo.org/record/14009210/files/figure.png" pageId="21" pageNumber="22">Fig. 13</figureCitation>
). Marginal carina broad, with clear attendant canals of preantennal setae. No internal line present. Preantennal nodi elongated, narrowed at base, broader in female than in male. Male
<emphasis id="B9604D75A84C687BFEC1F652FEC8FA89" box="[330,356,382,407]" italics="true" pageId="21" pageNumber="22">os</emphasis>
macrosetae, female
<emphasis id="B9604D75A84C687BFDD3F652FDDEFA89" box="[600,626,382,407]" italics="true" pageId="21" pageNumber="22">os</emphasis>
microsetae. Lateral temporal margins slightly concave, divergent posteriorly. Occipital bulge present, prominent;
<emphasis id="B9604D75A84C687BFD7BF68DFC8BFAA5" box="[752,807,417,443]" italics="true" pageId="21" pageNumber="22">mts5</emphasis>
situated apically. Occipital nodi large.
</paragraph>
<paragraph id="8BAB9167A84C687BFF36F6CEFE12F843" blockId="21.[186,1400,482,861]" pageId="21" pageNumber="22">
Thoracic and abdominal segments and chaetotaxy as in
<figureCitation id="132F8DE2A84C687BFCFAF6CEFC56FAE2" box="[881,1018,482,508]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="18.[189,232,1963,1989]" captionStartId-1="19.[189,232,1998,2024]" captionTargetBox-0="[427,1218,685,1921]" captionTargetBox-1="[416,1161,676,1933]" captionTargetId-0="figure-112@18.[340,1247,671,1926]" captionTargetId-1="figure-120@19.[340,1247,670,1953]" captionTargetPageId-0="18" captionTargetPageId-1="19" captionText-0="Fig. 11. Goniocotes sarissa sp. nov., holotype, ♂ (BMNH, slide 95, box E0026011). Habitus, dorsal and ventral views." captionText-1="Fig. 12. Goniocotes sarissa sp. nov. Female habitus, dorsal and ventral views." figureDoi-0="http://doi.org/10.5281/zenodo.14009206" figureDoi-1="http://doi.org/10.5281/zenodo.14009208" httpUri-0="https://zenodo.org/record/14009206/files/figure.png" httpUri-1="https://zenodo.org/record/14009208/files/figure.png" pageId="21" pageNumber="22">Figs 11–12</figureCitation>
. Rhombic sclerite not fused to pronotum. Postero-lateral corners of pronotum slightly extended laterally. Proepimera fused medianly. Postero-lateral corner of pteronotum not extended laterally; posterior margin gently rounded;
<emphasis id="B9604D75A84C687BFA97F505FAFBF95C" box="[1308,1367,553,578]" italics="true" pageId="21" pageNumber="22">smns</emphasis>
as patch of 4–5 setae on each side in male, but as single seta on each side in female. Male tergopleurites extensive on segments II–VI, but much reduced on segments VII–VIII, with that of segments VIII lacking median extension; lateral tergopleurite IX+X small, arched, not reaching median sclerite IX+X. Lateral accessory sternal plates present in female, as large reticulated plates on segments II–VI. Male abdominal chaetotaxy:
<emphasis id="B9604D75A84C687BFE4CF5F5FE72F9EC" box="[455,478,729,754]" italics="true" pageId="21" pageNumber="22">ss</emphasis>
present on tergopleurites II–VI;
<emphasis id="B9604D75A84C687BFCD9F5F5FCD8F9EC" box="[850,884,729,754]" italics="true" pageId="21" pageNumber="22">tps</emphasis>
present on tergopleurites II–IV;
<emphasis id="B9604D75A84C687BFB6DF5F5FAB6F9EC" box="[1254,1306,729,754]" italics="true" pageId="21" pageNumber="22">psps</emphasis>
present on tergopleurites II–VII;
<emphasis id="B9604D75A84C687BFE68F5D0FDA0F80B" box="[483,524,764,789]" italics="true" pageId="21" pageNumber="22">aps</emphasis>
absent;
<emphasis id="B9604D75A84C687BFDE5F5D0FD24F80B" box="[622,648,764,789]" italics="true" pageId="21" pageNumber="22">ps</emphasis>
present on segments III–VIII. Female abdominal chaetotaxy:
<emphasis id="B9604D75A84C687BFAD4F5D0FADAF80B" box="[1375,1398,764,789]" italics="true" pageId="21" pageNumber="22">ss</emphasis>
present on tergopleurites II–VIII;
<emphasis id="B9604D75A84C687BFDDBF40CFDDEF827" box="[592,626,800,825]" italics="true" pageId="21" pageNumber="22">tps</emphasis>
and
<emphasis id="B9604D75A84C687BFD25F40CFD7BF827" box="[686,727,800,825]" italics="true" pageId="21" pageNumber="22">aps</emphasis>
absent;
<emphasis id="B9604D75A84C687BFCB0F40CFCC3F827" box="[827,879,800,825]" italics="true" pageId="21" pageNumber="22">psps</emphasis>
present on tergopleurites II–VII;
<emphasis id="B9604D75A84C687BFA8BF40CFAB6F827" box="[1280,1306,800,825]" italics="true" pageId="21" pageNumber="22">ps</emphasis>
present on segments III–VIII.
</paragraph>
<paragraph id="8BAB9167A84C687BFF36F4AFFBD9F883" blockId="21.[189,1141,899,926]" box="[189,1141,899,926]" pageId="21" pageNumber="22">
Male genitalia as in
<figureCitation id="132F8DE2A84C687BFE20F4AFFDADF880" box="[427,513,899,926]" captionStart="Figs 13–15" captionStartId="20.[189,242,1963,1989]" captionTargetBox="[342,1235,650,1888]" captionTargetId="figure-39@20.[340,1247,636,1919]" captionTargetPageId="20" captionText="Figs 13–15. Goniocotes sarissa sp. nov. 13–14. Holotype, ♂ (BMNH, slide 95, box E0026011). 13. Head, dorsal and ventral views. 14. Genitalia, dorsal view. 15. Female genitalia, ventral view." figureDoi="http://doi.org/10.5281/zenodo.14009210" httpUri="https://zenodo.org/record/14009210/files/figure.png" pageId="21" pageNumber="22">Fig. 14</figureCitation>
; solenoid, with no clear distal margin;
<emphasis id="B9604D75A84C687BFC42F4A8FC47F883" box="[969,1003,900,925]" italics="true" pageId="21" pageNumber="22">pst</emphasis>
not visible.
</paragraph>
<paragraph id="8BAB9167A84C687BFF36F4E9FE77FF3B" blockId="21.[189,1399,965,1061]" pageId="21" pageNumber="22">
Female genitalia as in
<figureCitation id="132F8DE2A84C687BFE4BF4E9FDB8F8FE" box="[448,532,965,992]" captionStart="Figs 13–15" captionStartId="20.[189,242,1963,1989]" captionTargetBox="[342,1235,650,1888]" captionTargetId="figure-39@20.[340,1247,636,1919]" captionTargetPageId="20" captionText="Figs 13–15. Goniocotes sarissa sp. nov. 13–14. Holotype, ♂ (BMNH, slide 95, box E0026011). 13. Head, dorsal and ventral views. 14. Genitalia, dorsal view. 15. Female genitalia, ventral view." figureDoi="http://doi.org/10.5281/zenodo.14009210" httpUri="https://zenodo.org/record/14009210/files/figure.png" pageId="21" pageNumber="22">Fig. 15</figureCitation>
; subgenital plate absent. Vulval margin somewhat flattened medianly, with 8–11 short, slender
<emphasis id="B9604D75A84C687BFE22F4C5FE7BFF1C" box="[425,471,1001,1026]" italics="true" pageId="21" pageNumber="22">vms</emphasis>
and 3 longer, thorn-like
<emphasis id="B9604D75A84C687BFD74F4C5FC8FFF1C" box="[767,803,1001,1026]" italics="true" pageId="21" pageNumber="22">vss</emphasis>
on each side; 3–6 short, slender
<emphasis id="B9604D75A84C687BFB26F4C5FB78FF1C" box="[1197,1236,1001,1026]" italics="true" pageId="21" pageNumber="22">vos</emphasis>
on each side. Subvulval plates absent.
</paragraph>
<paragraph id="8BAB9167A84C687BFF36F361FDA7FF79" blockId="21.[189,523,1101,1127]" box="[189,523,1101,1127]" pageId="21" pageNumber="22">
Measurements as in
<tableCitation id="C696A4DCA84C687BFE26F361FDA9FF79" box="[429,517,1101,1127]" captionStart="Table 1" captionStartId="3.[186,256,660,688]" captionTargetPageId="3" captionText="Table 1. Measurements of the species described herein. Measurements were taken from live images of the specimens, for the following dimensions: AW = abdominal width (at segment V); HL = head length (at midline); HW = head width (at widest point of temples); PRW = prothoracic width; PTW= pterothoracic width; TL = total length (at midline)." pageId="21" pageNumber="22">Table 1</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30EC2ECA84C687BFF36F3A2FE0FFE00" pageId="21" pageNumber="22" type="discussion">
<paragraph id="8BAB9167A84C687BFF36F3A2FE83FFB6" blockId="21.[189,303,1166,1192]" box="[189,303,1166,1192]" pageId="21" pageNumber="22">
<heading id="D0E3260BA84C687BFF36F3A2FE83FFB6" bold="true" box="[189,303,1166,1192]" fontSize="11" level="3" pageId="21" pageNumber="22" reason="3">
<emphasis id="B9604D75A84C687BFF36F3A2FE83FFB6" bold="true" box="[189,303,1166,1192]" pageId="21" pageNumber="22">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BAB9167A84C687BFF36F391FE0FFE00" blockId="21.[189,1399,1213,1310]" pageId="21" pageNumber="22">
A single female of a second, larger species of
<taxonomicName id="4C14EAE4A84C687BFD6FF391FCC6FFC9" authorityName="Nitzsch" authorityYear="1818" box="[740,874,1213,1239]" class="Insecta" family="Philopteridae" genus="Goniocotes" kingdom="Animalia" order="Psocodea" pageId="21" pageNumber="22" phylum="Arthropoda" rank="genus">
<emphasis id="B9604D75A84C687BFD6FF391FCC6FFC9" box="[740,874,1213,1239]" italics="true" pageId="21" pageNumber="22">Goniocotes</emphasis>
</taxonomicName>
has also been seen from the same host (box E0026011, slide 95 at BMNH). More specimens are needed to identify this species, and verify that it occurs on
<taxonomicName id="4C14EAE4A84C687BFEBDF228FE37FE03" baseAuthorityName="Gould" baseAuthorityYear="1857" box="[310,411,1283,1309]" class="Aves" family="Phasianidae" genus="Tragopan" kingdom="Animalia" order="Galliformes" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="caboti">
<emphasis id="B9604D75A84C687BFEBDF228FE37FE03" box="[310,411,1283,1309]" italics="true" pageId="21" pageNumber="22">T. caboti</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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