1271 lines
156 KiB
XML
1271 lines
156 KiB
XML
<document id="240067F923F5BBAE4A0EF0EB0567BB4E" ID-CLB-Dataset="299663" ID-DOI="10.11646/zootaxa.5474.4.3" ID-GBIF-Dataset="f2a05eff-9900-485d-9c8a-acdf96470c35" ID-ISSN="1175-5326" ID-Zenodo-Dep="12681166" ID-ZooBank="4F341940-B3FA-42A4-A0F5-7EF88FD5ECB9" IM.bibliography_approvedBy="julia" IM.illustrations_approvedBy="julia" IM.materialsCitations_approvedBy="julia" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="julia" IM.taxonomicNames_approvedBy="julia" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="julia" checkinTime="1720373095461" checkinUser="plazi" docAuthor="Idiiatullina, Sabira S., Nguyen, Tan Van, Bragin, Andrey M., Pawangkhanant, Parinya, Le, Dac Xuan, Vogel, Gernot, David, Patrick & Poyarkov, Nikolay A." docDate="2024" docId="CF2DB900CA01FFA5FF6AD8FAD4FAFF42" docLanguage="en" docName="zootaxa.5474.4.3.pdf" docOrigin="Zootaxa 5474 (4)" docSource="http://dx.doi.org/10.11646/zootaxa.5474.4.3" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Trimeresurus cyanolabris Idiiatullina, Nguyen, Bragin, Pawangkhanant, Le, Vogel, David & Poyarkov, 2024, sp. nov." docType="treatment" docUuid="5593ADD1-63B3-4F39-BB5B-02D400B71D5C" docUuidSource="ZooBank" docVersion="5" lastPageNumber="386" masterDocId="3314C178CA07FFA9FFFDD94DD058FFBB" masterDocTitle="A new species of green pitviper of the Trimeresurus macrops complex (Reptilia: Serpentes: Viperidae) from South Central Coastal Region of Vietnam" masterLastPageNumber="411" masterPageNumber="375" pageNumber="381" updateTime="1720647552586" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="695E2154A18C76BF8378A50C6842E7AD">A new species of green pitviper of the Trimeresurus macrops complex (Reptilia: Serpentes: Viperidae) from South Central Coastal Region of Vietnam</mods:title>
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<mods:namePart id="EF83399855B358FA334C200156EC91AD">Idiiatullina, Sabira S.</mods:namePart>
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<mods:affiliation id="F72EC3C89AF1085522158FC0D63B1676">Department of Vertebrate Zoology, Lomonosov Moscow State University, Leninskiye Gory, GSP- 1, Moscow 119991, Russia.</mods:affiliation>
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<mods:namePart id="4FD851184C0FEB0F6C1F962CA2E6281E">Nguyen, Tan Van</mods:namePart>
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<mods:affiliation id="40588DB9609E91DD02D7D4F1CBBF0DBF">Institute for Research and Training in Medicine, Biology and Pharmacy, Duy Tan University, Da Nang, 550000, Vietnam. & College of Medicine and Pharmacy, Duy Tan University, 120 Hoang Minh Thao, Lien Chieu, Da Nang, 550000, Vietnam.</mods:affiliation>
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<mods:namePart id="E35D2E4A9BE7E78EF698B00BD3400F7A">Bragin, Andrey M.</mods:namePart>
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<mods:affiliation id="1CB42922892D67BE8DF7A4D920AF8DCA">Department of Vertebrate Zoology, Lomonosov Moscow State University, Leninskiye Gory, GSP- 1, Moscow 119991, Russia. & Joint Vietnam - Russia Tropical Science and Technology Research Center, Nghia Do, Cau Giay, Hanoi, Vietnam.</mods:affiliation>
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<mods:namePart id="B6210BFCF6386EA8D945D327EA9FD8E6">Pawangkhanant, Parinya</mods:namePart>
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<mods:affiliation id="0AAA5D14FCCC718B84E9C8DB4FB88349">Rabbit in the Moon foundation, Suanphueng, Ratchaburi 70180, Thailand. & Division of Fishery, School of Agriculture and Natural Resources, University of Phayao, Phayao, Thailand.</mods:affiliation>
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<mods:namePart id="4904EC5D3FFB8B153B04092BD8C356D9">Le, Dac Xuan</mods:namePart>
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<mods:affiliation id="D6EB23F42A20E93462CED778ED1E5A1C">Joint Vietnam - Russia Tropical Science and Technology Research Center, Nghia Do, Cau Giay, Hanoi, Vietnam.</mods:affiliation>
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<mods:namePart id="007BA40FD0C4047DC8EA914E7A00AFCD">Vogel, Gernot</mods:namePart>
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<mods:affiliation id="FB2D0E545D0C01B95044AFB49CECEDFF">Society for South East Asian Herpetology, Im Sand- 3, D- 69115 Heidelberg, Germany.</mods:affiliation>
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<mods:namePart id="7F7DC3ED309A4EBB0F1D63B0E21550AE">David, Patrick</mods:namePart>
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<mods:affiliation id="7677F6A3B936CEA8C42418B006AB8C18">Reptiles & Amphibiens, UMR 7205 OSEB, Département évolution et Systématique, CP 30, Muséum National d’Histoire Naturelle. 57 rue Cuvier, F- 75231 Paris Cedex 05, France.</mods:affiliation>
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<mods:namePart id="622AC2A6700914269F5EB0749966A6E1">Poyarkov, Nikolay A.</mods:namePart>
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<mods:affiliation id="B2D8A212AD6BB7D190473E888BC04E63">Department of Vertebrate Zoology, Lomonosov Moscow State University, Leninskiye Gory, GSP- 1, Moscow 119991, Russia. & Joint Vietnam - Russia Tropical Science and Technology Research Center, Nghia Do, Cau Giay, Hanoi, Vietnam.</mods:affiliation>
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<mods:title id="3D047F952870EB59252860AC5AD28005">Zootaxa</mods:title>
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<mods:date id="108FEF434493B4B62484CFF68DD7B9B2">2024</mods:date>
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<treatment id="CF2DB900CA01FFA5FF6AD8FAD4FAFF42" ID-DOI="http://doi.org/10.5281/zenodo.12708898" ID-GBIF-Taxon="235029949" ID-Zenodo-Dep="12708898" LSID="urn:lsid:zoobank.org:act:5593ADD1-63B3-4F39-BB5B-02D400B71D5C" httpUri="http://treatment.plazi.org/id/CF2DB900CA01FFA5FF6AD8FAD4FAFF42" lastPageId="12" lastPageNumber="386" pageId="6" pageNumber="381">
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<taxonomicName id="80847395CA01FFAFFF6AD8FAD196FE69" authorityName="Idiiatullina & Nguyen & Bragin & Pawangkhanant & Le & Vogel & David & Poyarkov" authorityYear="2024" box="[151,462,439,466]" class="Squamata" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" pageId="6" pageNumber="381" phylum="Chordata" rank="species" species="cyanolabris" status="sp. nov.">
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<emphasis id="75F0D404CA01FFAFFF6AD8FAD196FE69" bold="true" box="[151,462,439,466]" italics="true" pageId="6" pageNumber="381">Trimeresurus cyanolabris</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="6EC3697FCA01FFAFFE2BD8FAD26BFE6A" box="[470,563,439,465]" pageId="6" pageNumber="381" rank="species">sp. nov.</taxonomicNameLabel>
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(
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<figureCitation id="DFBF1493CA01FFAFFDBED8FAD2F3FE6A" box="[579,683,439,465]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStartId-0="7.[152,255,1921,1945]" captionStartId-1="8.[152,255,1832,1856]" captionTargetBox-0="[165,1423,309,1897]" captionTargetBox-1="[193,1395,181,1809]" captionTargetId-0="figure-95@7.[164,1423,309,1897]" captionTargetId-1="figure-25@8.[192,1395,181,1809]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionText-0="FIGURE 3. The holotype of Trimeresurus cyanolabris sp. nov. in preservative (ZMMU Re-17679, adult male) from Nui Chua NP, Ninh Thuan Province, Vietnam. A dorsolateral and B ventral view of body; C head in dorsal, D in ventral, E in lateral right, and F in lateral left aspects. Scale bar equals 1 cm. Photographs by S.S. Idiiatullina." captionText-1="FIGURE 4. Type specimens of Trimeresurus cyanolabris sp. nov. in life from Nui Chua NP, Ninh Thuan Province, Vietnam: A holotype ZMMU Re-17679, adult male; B paratype ZMMU Re-17681, adult female. Photographs by A.M. Bragin (A) and N.A. Poyarkov (B)." figureDoi-0="http://doi.org/10.5281/zenodo.12681174" figureDoi-1="http://doi.org/10.5281/zenodo.12681176" httpUri-0="https://zenodo.org/record/12681174/files/figure.png" httpUri-1="https://zenodo.org/record/12681176/files/figure.png" pageId="6" pageNumber="381">Fig. 3–4</figureCitation>
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;
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<tableCitation id="0A063DADCA01FFAFFD45D8FAD378FE6A" box="[696,800,439,465]" captionStart="TABLE 2" captionStartId="14.[152,245,440,464]" captionTargetPageId="14" captionText="TABLE 2. Sequences and voucher specimens of the genus Trimeresurus and outgroup taxa used in this study." httpUri="http://table.plazi.org/id/13FB589ECA09FFA7FF65D8F5D568FE6B" pageId="6" pageNumber="381" tableUuid="13FB589ECA09FFA7FF65D8F5D568FE6B">Tables 2</tableCitation>
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,
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<tableCitation id="0A063DADCA01FFAFFCD0D8FAD302FE6A" box="[813,858,439,465]" captionStart-0="TABLE 4" captionStart-1="TABLE 5" captionStart-2="TABLE 6" captionStartId-0="21.[152,245,888,912]" captionStartId-1="22.[152,245,151,175]" captionStartId-2="23.[152,245,1242,1266]" captionTargetPageId-0="21" captionTargetPageId-1="22" captionTargetPageId-2="23" captionText-0="TABLE 4. Uncorrected p–distances (percentage) between the sequences of cyt b mtDNA gene of species of the subgenus Trimeresurus included in the phylogenetic analyses." captionText-1="TABLE 5. Morphological data on the type series of Trimeresurus cyanolabris sp. nov. Notes: for abbreviations see Materials and methods section; N/a: Data not available." captionText-2="TABLE 6. Comparison of morphological characters of Trimeresurus cyanolabris sp. nov. with T. cardamomensis, T. macrops, and T. rubeus. Remarks: Diagnostic differences from the new species are marked in bold. (?): requires verification." httpUri-0="http://table.plazi.org/id/13FB589ECA12FFBCFF65DA35D29AFC0F" httpUri-1="http://table.plazi.org/id/13FB589ECA11FFBFFF65D9DAD2B0FF68" httpUri-2="http://table.plazi.org/id/13FB589ECA10FFBEFF65DD97D142FA81" pageId="6" pageNumber="381" tableUuid-0="13FB589ECA12FFBCFF65DA35D29AFC0F" tableUuid-1="13FB589ECA11FFBFFF65D9DAD2B0FF68" tableUuid-2="13FB589ECA10FFBEFF65DD97D142FA81">4–6</tableCitation>
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)
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</emphasis>
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</heading>
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<subSubSection id="0F9E5B9DCA01FFAFFF6AD8B2D428FD6A" pageId="6" pageNumber="381" type="reference_group">
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<emphasis id="75F0D404CA01FFAFFF6AD8B2D118FDA2" bold="true" box="[151,320,511,537]" pageId="6" pageNumber="381">Chresonymy:</emphasis>
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</heading>
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</paragraph>
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<paragraph id="473B0816CA01FFAFFF6ADB72D0A6FDCE" blockId="6.[151,1437,574,721]" pageId="6" pageNumber="381">
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<treatmentCitationGroup id="67942F38CA01FFAFFF6ADB72D0A6FDCE" pageId="6" pageNumber="381">
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<taxonomicName id="80847395CA01FFAFFF6ADB72D1D5FDED" ID-CoL="7CTKS" authorityName="Gray" authorityYear="1842" box="[151,397,574,598]" class="Squamata" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" pageId="6" pageNumber="381" phylum="Chordata" rank="species" species="albolabris">
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<emphasis id="75F0D404CA01FFAFFF6ADB72D1D5FDED" box="[151,397,574,598]" italics="true" pageId="6" pageNumber="381">Trimeresurus albolabris</emphasis>
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</taxonomicName>
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(
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<emphasis id="75F0D404CA01FFAFFE63DB72D19CFDED" box="[414,452,575,598]" italics="true" pageId="6" pageNumber="381">non</emphasis>
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<taxonomicName id="80847395CA01FFAFFE31DB72D366FDED" ID-CoL="7CTKS" authority="Gray, 1842" authorityName="Gray" authorityYear="1842" box="[460,830,574,598]" class="Squamata" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" pageId="6" pageNumber="381" phylum="Chordata" rank="species" species="albolabris">
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<emphasis id="75F0D404CA01FFAFFE31DB72D29BFDED" box="[460,707,574,598]" italics="true" pageId="6" pageNumber="381">Trimeresurus albolabris</emphasis>
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Gray, 1842
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</taxonomicName>
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) —
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||
<treatmentCitation id="C6252E07CA01FFAFFC92DB72D42DFDED" author="Nemes, L. & Babb, R. & Devender, W. V. & Nguyen, K. V. & Le, Q. K. & Vu, T. N. & Rauhaus, A. & Nguyen, T. Q. & Ziegler, T." box="[879,1141,574,598]" page="320" pageId="6" pageNumber="381" year="2013">
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<bibRefCitation id="231575E7CA01FFAFFC92DB72D42DFDED" author="Nemes, L. & Babb, R. & Devender, W. V. & Nguyen, K. V. & Le, Q. K. & Vu, T. N. & Rauhaus, A. & Nguyen, T. Q. & Ziegler, T." box="[879,1141,574,598]" pageId="6" pageNumber="381" pagination="301 - 326" refId="ref19197" refString="Nemes, L., Babb, R., Devender, W. V., Nguyen, K. V., Le, Q. K., Vu, T. N., Rauhaus, A., Nguyen, T. Q. & Ziegler, T. (2013) First contribution to the reptile fauna of Quang Ngai province, central Vietnam. Biodiversity Journal, 4 (2), 301 - 326." type="journal article" year="2013">
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Nemes
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<emphasis id="75F0D404CA01FFAFFC40DB72D3B4FDED" box="[957,1004,574,598]" italics="true" pageId="6" pageNumber="381">et al</emphasis>
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. (2013: 320)
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</bibRefCitation>
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</treatmentCitation>
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;
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<treatmentCitation id="C6252E07CA01FFAFFB7FDB72D525FDED" author="Nguyen, T. V. & Duong, L. D. & Duong, T. T. A." box="[1154,1405,574,599]" page="40" pageId="6" pageNumber="381" year="2019">
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<bibRefCitation id="231575E7CA01FFAFFB7FDB72D525FDED" author="Nguyen, T. V. & Duong, L. D. & Duong, T. T. A." box="[1154,1405,574,599]" pageId="6" pageNumber="381" pagination="35 - 41" refId="ref19485" refString="Nguyen, T. V., Duong, L. D. & Duong, T. T. A. (2019) New records and an updated list of snakes (Squamata: Serpentes) from Binh Dinh Province, Central-Southern Vietnam. Hue University Journal of Science: Natural Science, 128 (1 B), 35 - 41. https: // doi. org / 10.26459 / hueuni-jns. v 128 i 1 B. 5203" type="journal article" year="2019">
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Nguyen
|
||
<emphasis id="75F0D404CA01FFAFFB27DB72D550FDED" box="[1242,1288,574,598]" italics="true" pageId="6" pageNumber="381">et al</emphasis>
|
||
. (2019: 40
|
||
</bibRefCitation>
|
||
</treatmentCitation>
|
||
, in part).
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="473B0816CA01FFAFFF6ADB31D414FD2F" blockId="6.[151,1437,574,721]" box="[151,1100,636,660]" pageId="6" pageNumber="381">
|
||
<treatmentCitationGroup id="67942F38CA01FFAFFF6ADB31D414FD2F" box="[151,1100,636,660]" pageId="6" pageNumber="381">
|
||
<taxonomicName id="80847395CA01FFAFFF6ADB31D122FD28" ID-CoL="7CTN5" authorityName="Kramer" authorityYear="1977" box="[151,378,636,659]" class="Squamata" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" pageId="6" pageNumber="381" phylum="Chordata" rank="species" species="macrops">
|
||
<emphasis id="75F0D404CA01FFAFFF6ADB31D122FD28" box="[151,378,636,659]" italics="true" pageId="6" pageNumber="381">Trimeresurus macrops</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<emphasis id="75F0D404CA01FFAFFE74DB31D1F7FD28" box="[393,431,636,659]" italics="true" pageId="6" pageNumber="381">non</emphasis>
|
||
<taxonomicName id="80847395CA01FFAFFE48DB31D373FD2F" ID-CoL="7CTN5" authority="Kramer, 1977" authorityName="Kramer" authorityYear="1977" box="[437,811,636,660]" class="Squamata" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" pageId="6" pageNumber="381" phylum="Chordata" rank="species" species="macrops">
|
||
<emphasis id="75F0D404CA01FFAFFE48DB31D2C0FD28" box="[437,664,636,659]" italics="true" pageId="6" pageNumber="381">Trimeresurus macrops</emphasis>
|
||
<bibRefCitation id="231575E7CA01FFAFFD63DB31D373FD2F" author="Kramer, E." box="[670,811,636,660]" pageId="6" pageNumber="381" pagination="721 - 761" refId="ref18354" refString="Kramer, E. (1977) Zur Schlangenfauna Nepals. Revue Suisse de Zoologie, 84 (3), 721 - 761, 5 figs. [in German] https: // doi. org / 10.5962 / bhl. part. 91420" type="journal article" year="1977">Kramer, 1977</bibRefCitation>
|
||
</taxonomicName>
|
||
) —
|
||
<treatmentCitation id="C6252E07CA01FFAFFCA5DB31D411FD2F" author="Hoang, M. D. & Luu, H. T. & Tran, B. V. & Le, D. & To, V. Q. & Dang, V. D. & Nguyen, H. N." box="[856,1097,636,660]" page="90" pageId="6" pageNumber="381" year="2019">
|
||
<bibRefCitation id="231575E7CA01FFAFFCA5DB31D411FD2F" author="Hoang, M. D. & Luu, H. T. & Tran, B. V. & Le, D. & To, V. Q. & Dang, V. D. & Nguyen, H. N." box="[856,1097,636,660]" pageId="6" pageNumber="381" refId="ref17529" refString="Hoang, M. D., Luu, H. T., Tran, B. V., Le, D., To, V. Q., Dang, V. D. & Nguyen, H. N. (2019) Forest biodiversity in Khanh Hoa province - Part 2 Animals. Publishing House for Science and Technology, 215 pp. [in Vietnamese]" type="book" year="2019">
|
||
Hoang
|
||
<emphasis id="75F0D404CA01FFAFFC5FDB31D397FD2F" box="[930,975,636,660]" italics="true" pageId="6" pageNumber="381">et al</emphasis>
|
||
. (2019: 90)
|
||
</bibRefCitation>
|
||
</treatmentCitation>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="473B0816CA01FFAFFF6ADBD6D3F2FD09" blockId="6.[151,1437,574,721]" box="[151,938,666,690]" pageId="6" pageNumber="381">
|
||
<treatmentCitationGroup id="67942F38CA01FFAFFF6ADBD6D3F2FD09" box="[151,938,666,690]" pageId="6" pageNumber="381">
|
||
<taxonomicName id="80847395CA01FFAFFF6ADBD6D24FFD09" ID-CoL="7CTN5" authorityName="Kramer" authorityYear="1977" box="[151,535,667,690]" class="Squamata" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" pageId="6" pageNumber="381" phylum="Chordata" rank="species" species="macrops" subGenus="Trimeresurus">
|
||
<emphasis id="75F0D404CA01FFAFFF6ADBD6D24FFD09" box="[151,535,667,690]" italics="true" pageId="6" pageNumber="381">Trimeresurus (Trimeresurus) macrops</emphasis>
|
||
</taxonomicName>
|
||
—
|
||
<treatmentCitation id="C6252E07CA01FFAFFDC0DBD7D316FD09" author="Poyarkov, N. A. & Nguyen, T. V. & Popov, E. S. & Geissler, P. & Pawangkhanant, P. & Neang, T. & Suwannapoom, C. & Ananjeva, N. B. & Orlov, N. L." box="[573,846,666,690]" page="391" pageId="6" pageNumber="381" year="2023">
|
||
<bibRefCitation id="231575E7CA01FFAFFDC0DBD7D316FD09" author="Poyarkov, N. A. & Nguyen, T. V. & Popov, E. S. & Geissler, P. & Pawangkhanant, P. & Neang, T. & Suwannapoom, C. & Ananjeva, N. B. & Orlov, N. L." box="[573,846,666,690]" pageId="6" pageNumber="381" pagination="255 - 476" refId="ref19734" refString="Poyarkov, N. A., Nguyen, T. V., Popov, E. S., Geissler, P., Pawangkhanant, P., Neang, T., Suwannapoom, C., Ananjeva, N. B. & Orlov, N. L. (2023) Recent progress in taxonomic studies, biogeographic analysis and revised checklist of Reptilians in Indochina. Russian Journal of Herpetology, 30 (5), 255 - 476. https: // doi. org / 10.30906 / 1026 - 2296 - 2023 - 30 - 5 - 255 - 476" type="journal article" year="2023">
|
||
Poyarkov
|
||
<emphasis id="75F0D404CA01FFAFFD58DBD6D289FD09" box="[677,721,666,690]" italics="true" pageId="6" pageNumber="381">et al</emphasis>
|
||
. (2023: 391
|
||
</bibRefCitation>
|
||
</treatmentCitation>
|
||
, in part).
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="473B0816CA01FFAFFF6ADBF7D428FD6A" blockId="6.[151,1437,574,721]" box="[151,1136,697,721]" pageId="6" pageNumber="381">
|
||
<treatmentCitationGroup id="67942F38CA01FFAFFF6ADBF7D428FD6A" box="[151,1136,697,721]" pageId="6" pageNumber="381">
|
||
<taxonomicName id="80847395CA01FFAFFF6ADBF7D1D0FD6A" ID-CoL="7D5PV" authorityName="Schmidt" authorityYear="1925" box="[151,392,698,721]" class="Squamata" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" pageId="6" pageNumber="381" phylum="Chordata" rank="species" species="stejnegeri">
|
||
<emphasis id="75F0D404CA01FFAFFF6ADBF7D1D0FD6A" box="[151,392,698,721]" italics="true" pageId="6" pageNumber="381">Trimeresurus stejnegeri</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<emphasis id="75F0D404CA01FFAFFE6BDBF7D1E4FD6A" box="[406,444,698,721]" italics="true" pageId="6" pageNumber="381">non</emphasis>
|
||
<taxonomicName id="80847395CA01FFAFFE3EDBF7D317FD6A" ID-CoL="7D5PV" authority="Schmidt, 1925" authorityName="Schmidt" authorityYear="1925" box="[451,847,697,721]" class="Squamata" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" pageId="6" pageNumber="381" phylum="Chordata" rank="species" species="stejnegeri">
|
||
<emphasis id="75F0D404CA01FFAFFE3EDBF7D2EBFD6A" box="[451,691,698,721]" italics="true" pageId="6" pageNumber="381">Trimeresurus stejnegeri</emphasis>
|
||
Schmidt, 1925
|
||
</taxonomicName>
|
||
) —
|
||
<treatmentCitation id="C6252E07CA01FFAFFC81DBF4D433FD6A" author="Hoang, M. D. & Luu, H. T. & Tran, B. V. & Le, D. & To, V. Q. & Dang, V. D. & Nguyen, H. N." box="[892,1131,697,721]" page="91" pageId="6" pageNumber="381" year="2019">
|
||
<bibRefCitation id="231575E7CA01FFAFFC81DBF4D433FD6A" author="Hoang, M. D. & Luu, H. T. & Tran, B. V. & Le, D. & To, V. Q. & Dang, V. D. & Nguyen, H. N." box="[892,1131,697,721]" pageId="6" pageNumber="381" refId="ref17529" refString="Hoang, M. D., Luu, H. T., Tran, B. V., Le, D., To, V. Q., Dang, V. D. & Nguyen, H. N. (2019) Forest biodiversity in Khanh Hoa province - Part 2 Animals. Publishing House for Science and Technology, 215 pp. [in Vietnamese]" type="book" year="2019">
|
||
Hoang
|
||
<emphasis id="75F0D404CA01FFAFFC3BDBF7D3AAFD6A" box="[966,1010,697,721]" italics="true" pageId="6" pageNumber="381">et al</emphasis>
|
||
. (2019: 91)
|
||
</bibRefCitation>
|
||
</treatmentCitation>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0F9E5B9DCA01FFAFFF6ADBB2D136F9EE" pageId="6" pageNumber="381" type="materials_examined">
|
||
<paragraph id="473B0816CA01FFAFFF6ADBB2D4AAFCDA" blockId="6.[151,1437,767,2017]" pageId="6" pageNumber="381">
|
||
<materialsCitation id="F7EC024BCA01FFAFFF6ADBB2D4AAFCDA" ID-GBIF-Occurrence="4904896303" collectingDate="2023-03-08" collectionCode="ZMMU" collectorName="N. A. Poyarkov & A. M. Bragin" country="Vietnam" county="Ninh Hai District" elevation="155" latitude="11.77757" location="Nui Chua National Park in Thai An Village" longLatPrecision="1" longitude="109.17603" municipality="Vinh Hai Commune" pageId="6" pageNumber="381" specimenCount="1" stateProvince="Ninh Thuan" typeStatus="holotype">
|
||
<emphasis id="75F0D404CA01FFAFFF6ADBB2D15DFCA2" bold="true" box="[151,261,767,793]" pageId="6" pageNumber="381">
|
||
<typeStatus id="983FB6B4CA01FFAFFF6ADBB2D158FCA2" box="[151,256,767,793]" pageId="6" pageNumber="381" type="holotype">Holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
<collectionCode id="219590D3CA01FFAFFEF7DA4DD107FCA1" box="[266,351,768,794]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="6" pageNumber="381" type="Museum">ZMMU</collectionCode>
|
||
Re-17679 (field ID NAP-13705), adult male, from
|
||
<location id="425B5ECDCA01FFAFFC88DBB2D56CFCA2" LSID="urn:lsid:plazi:treatment:CF2DB900CA01FFA5FF6AD8FAD4FAFF42:425B5ECDCA01FFAFFC88DBB2D56CFCA2" box="[885,1332,767,793]" country="Vietnam" county="Ninh Hai District" latitude="11.77757" longLatPrecision="1" longitude="109.17603" municipality="Vinh Hai Commune" name="Nui Chua National Park in Thai An Village" pageId="6" pageNumber="381" stateProvince="Ninh Thuan">Nui Chua National Park in Thai An Village</location>
|
||
,
|
||
<collectingMunicipality id="A75F926CCA01FFAFFAC0DBB2D151FC86" pageId="6" pageNumber="381">Vinh Hai Commune</collectingMunicipality>
|
||
,
|
||
<collectingCounty id="AE5A709ACA01FFAFFEE9DA6ED197FC86" box="[276,463,803,829]" pageId="6" pageNumber="381">Ninh Hai District</collectingCounty>
|
||
,
|
||
<collectingRegion id="8540C6F4CA01FFAFFE27DA6ED299FC86" box="[474,705,803,829]" country="Vietnam" name="Ninh Thuan" pageId="6" pageNumber="381">Ninh Thuan Province</collectingRegion>
|
||
,
|
||
<collectingCountry id="3F934886CA01FFAFFD36DA6ED37DFC86" box="[715,805,803,829]" name="Vietnam" pageId="6" pageNumber="381">Vietnam</collectingCountry>
|
||
(
|
||
<geoCoordinate id="22B06ED1CA01FFAFFCC9DA6ED3E0FC85" box="[820,952,803,830]" degrees="11.77757" direction="north" orientation="latitude" pageId="6" pageNumber="381" precision="1" value="11.77757">11.77757°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="22B06ED1CA01FFAFFC3EDA6ED40AFC86" box="[963,1106,803,829]" degrees="109.17603" direction="east" orientation="longitude" pageId="6" pageNumber="381" precision="1" value="109.17603">109.17603°E</geoCoordinate>
|
||
; altitude
|
||
<elevation id="CCA9EF25CA01FFAFFB4FDA6ED574FC86" box="[1202,1324,803,830]" metricMagnitude="2" metricUnit="m" metricValue="1.55" pageId="6" pageNumber="381" unit="m" value="155.0">
|
||
<quantity id="807CA5F3CA01FFAFFB4FDA6ED4AEFC85" box="[1202,1270,803,830]" metricMagnitude="2" metricUnit="m" metricValue="1.55" pageId="6" pageNumber="381" unit="m" value="155.0">155 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
) collected by
|
||
<collectorName id="EA716DC0CA01FFAFFF45DA0AD100FCDA" box="[184,344,839,865]" pageId="6" pageNumber="381">N.A. Poyarkov</collectorName>
|
||
and
|
||
<collectorName id="EA716DC0CA01FFAFFE74DA0AD248FCDA" box="[393,528,839,865]" pageId="6" pageNumber="381">A.M. Bragin</collectorName>
|
||
from the ground on the bank of a mountain stream on
|
||
<date id="333A2ED6CA01FFAFFBB2DA0AD4B6FCDA" box="[1103,1262,839,865]" pageId="6" pageNumber="381" value="2023-03-08">
|
||
<collectingDate id="237ED73ECA01FFAFFBB2DA0AD4B6FCDA" box="[1103,1262,839,865]" pageId="6" pageNumber="381" value="2023-03-08">March 8, 2023</collectingDate>
|
||
</date>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="473B0816CA01FFAFFF3ADA26D52DFAC6" blockId="6.[151,1437,767,2017]" pageId="6" pageNumber="381">
|
||
<materialsCitation id="F7EC024BCA01FFAFFF3ADA26D1CCFC12" ID-GBIF-Occurrence="4904896308" collectingDate="2023-03-08" collectionCode="ZMMU" collectorName="N. A. Poyarkov & A. M. Bragin" country="Vietnam" county="Ninh Hai District" elevation="155" latitude="11.77757" location="Nui Chua National Park in Thai An Village" longLatPrecision="1" longitude="109.17603" municipality="Vinh Hai Commune" pageId="6" pageNumber="381" specimenCount="1" stateProvince="Ninh Thuan" typeStatus="paratype">
|
||
<emphasis id="75F0D404CA01FFAFFF3ADA26D160FC3E" bold="true" box="[199,312,875,901]" pageId="6" pageNumber="381">
|
||
<typeStatus id="983FB6B4CA01FFAFFF3ADA26D160FC3E" box="[199,312,875,901]" pageId="6" pageNumber="381" type="paratype">Paratypes</typeStatus>
|
||
</emphasis>
|
||
(n = 9).
|
||
<collectionCode id="219590D3CA01FFAFFE68DA21D1B2FC3D" box="[405,490,876,902]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="6" pageNumber="381" type="Museum">ZMMU</collectionCode>
|
||
Re-17678 (field ID NAP-13704), adult male, collected on
|
||
<date id="333A2ED6CA01FFAFFBA6DA26D4A0FC3E" box="[1115,1272,875,901]" pageId="6" pageNumber="381" value="2023-03-08">
|
||
<collectingDate id="237ED73ECA01FFAFFBA6DA26D4A0FC3E" box="[1115,1272,875,901]" pageId="6" pageNumber="381" value="2023-03-08">March 8, 2023</collectingDate>
|
||
</date>
|
||
, from the same location as the holotype
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="F7EC024BCA01FFAFFE62DADDD2FAFBAE" ID-GBIF-Occurrence="4904896304" collectingDate="2023-01-06" collectionCode="ZMMU" collectorName="N. A. Poyarkov & D. X. Le" country="Vietnam" elevation="280" latitude="11.778" location="Nuoc Ngot River" longLatPrecision="77" longitude="109.176" municipality="Ninh Hai District" pageId="6" pageNumber="381" specimenCount="2" specimenCount-adult="2" stateProvince="Ninh Thuan" typeStatus="paratype">
|
||
<collectionCode id="219590D3CA01FFAFFE62DADDD1ACFC11" box="[415,500,912,938]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="6" pageNumber="381" type="Museum">ZMMU</collectionCode>
|
||
Re-17680–81 (field ID NAP-13229–30),
|
||
<specimenCount id="5182C39FCA01FFAFFC52DADDD449FC12" box="[943,1041,911,938]" count="2" pageId="6" pageNumber="381" type="adult">two adult</specimenCount>
|
||
females, collected by
|
||
<collectorName id="EA716DC0CA01FFAFFB00DAC2D5C5FC12" box="[1277,1437,911,937]" pageId="6" pageNumber="381">N.A. Poyarkov</collectorName>
|
||
and
|
||
<collectorName id="EA716DC0CA01FFAFFF38DAF9D17BFC75" box="[197,291,948,974]" pageId="6" pageNumber="381">D. X. Le</collectorName>
|
||
on
|
||
<date id="333A2ED6CA01FFAFFEB0DAF9D1A0FC76" box="[333,504,947,974]" pageId="6" pageNumber="381" value="2023-01-06">
|
||
<collectingDate id="237ED73ECA01FFAFFEB0DAF9D1A0FC76" box="[333,504,947,974]" pageId="6" pageNumber="381" value="2023-01-06">January 6, 2023</collectingDate>
|
||
</date>
|
||
, from the valley of
|
||
<location id="425B5ECDCA01FFAFFD37DAF9D3D9FC76" LSID="urn:lsid:plazi:treatment:CF2DB900CA01FFA5FF6AD8FAD4FAFF42:425B5ECDCA01FFAFFD37DAF9D3D9FC76" box="[714,897,947,974]" country="Vietnam" latitude="11.778" longLatPrecision="77" longitude="109.176" municipality="Ninh Hai District" name="Nuoc Ngot River" pageId="6" pageNumber="381" stateProvince="Ninh Thuan Province">Nuoc Ngot River</location>
|
||
(
|
||
<geoCoordinate id="22B06ED1CA01FFAFFC6CDAFED3A0FC76" box="[913,1016,947,973]" degrees="11.778" direction="north" orientation="latitude" pageId="6" pageNumber="381" precision="55" value="11.778">11.778°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="22B06ED1CA01FFAFFBF9DAFED42FFC76" box="[1028,1143,947,973]" degrees="109.176" direction="east" orientation="longitude" pageId="6" pageNumber="381" precision="55" value="109.176">109.176°E</geoCoordinate>
|
||
; altitude
|
||
<elevation id="CCA9EF25CA01FFAFFB25DAFED50BFC76" box="[1240,1363,947,974]" metricMagnitude="2" metricUnit="m" metricValue="2.8" pageId="6" pageNumber="381" unit="m" value="280.0">
|
||
<quantity id="807CA5F3CA01FFAFFB25DAFED544FC75" box="[1240,1308,947,974]" metricMagnitude="2" metricUnit="m" metricValue="2.8" pageId="6" pageNumber="381" unit="m" value="280.0">280 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
), ca.
|
||
<quantity id="807CA5F3CA01FFAFFA73DAFED0E1FC4A" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="6" pageNumber="381" unit="km" value="3.0">3 km</quantity>
|
||
south-west from
|
||
<collectorName id="EA716DC0CA01FFAFFE8FDA95D26AFC4A" box="[370,562,983,1010]" pageId="6" pageNumber="381">Nuoc Ngot Beach</collectorName>
|
||
, on the territory of
|
||
<collectorName id="EA716DC0CA01FFAFFD00DA9AD3C0FC49" box="[765,920,983,1010]" pageId="6" pageNumber="381">Nui Chua N.P.</collectorName>
|
||
,
|
||
<collectorName id="EA716DC0CA01FFAFFC5EDA9AD414FC4A" box="[931,1100,983,1009]" hasSafeAffix="true" pageId="6" pageNumber="381">Thai An Village</collectorName>
|
||
,
|
||
<collectorName id="EA716DC0CA01FFAFFBAADA9AD577FC4A" box="[1111,1327,983,1009]" pageId="6" pageNumber="381">Vinh Hai Commune</collectorName>
|
||
,
|
||
<collectingMunicipality id="A75F926CCA01FFAFFAC7DA9AD0BFFBAE" pageId="6" pageNumber="381">Ninh Hai District</collectingMunicipality>
|
||
,
|
||
<collectingRegion id="8540C6F4CA01FFAFFF0EDAB6D181FBAE" box="[243,473,1019,1045]" inferredBySuffix="true" pageId="6" pageNumber="381">Ninh Thuan Province</collectingRegion>
|
||
, southern
|
||
<collectingCountry id="3F934886CA01FFAFFDBBDAB6D2FAFBAE" box="[582,674,1019,1045]" name="Vietnam" pageId="6" pageNumber="381">Vietnam</collectingCountry>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="F7EC024BCA01FFAFFD50DAB1D41EFBE6" ID-GBIF-Occurrence="4904896307" collectingDate="2023-03-04" collectionCode="ZMMU" collectorName="N. A. Poyarkov & A. M. Bragin" country="Vietnam" county="Ninh Hai District" elevation="730" latitude="11.72343" location="Thai An Village" longLatPrecision="1" longitude="109.13656" municipality="Vinh Hai Commune" pageId="6" pageNumber="381" specimenCount="1" stateProvince="Ninh Thuan" typeStatus="paratype">
|
||
<collectionCode id="219590D3CA01FFAFFD50DAB1D35AFBAD" box="[685,770,1020,1046]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="6" pageNumber="381" type="Museum">ZMMU</collectionCode>
|
||
Re-17682 (field ID NAP-13643), adult female, collected from
|
||
<collectorName id="EA716DC0CA01FFAFFF6ADD52D16BFB81" box="[151,307,1055,1082]" pageId="6" pageNumber="381">Nui Chua N.P.</collectorName>
|
||
in
|
||
<location id="425B5ECDCA01FFAFFEABDD52D259FB82" LSID="urn:lsid:plazi:treatment:CF2DB900CA01FFA5FF6AD8FAD4FAFF42:425B5ECDCA01FFAFFEABDD52D259FB82" box="[342,513,1055,1081]" country="Vietnam" county="Ninh Hai District" latitude="11.72343" longLatPrecision="1" longitude="109.13656" municipality="Vinh Hai Commune" name="Thai An Village" pageId="6" pageNumber="381" stateProvince="Ninh Thuan">Thai An Village</location>
|
||
,
|
||
<collectingMunicipality id="A75F926CCA01FFAFFDF0DD52D2BFFB82" box="[525,743,1055,1081]" pageId="6" pageNumber="381">Vinh Hai Commune</collectingMunicipality>
|
||
,
|
||
<collectingCounty id="AE5A709ACA01FFAFFD0EDD52D3F6FB82" box="[755,942,1055,1081]" pageId="6" pageNumber="381">Ninh Hai District</collectingCounty>
|
||
,
|
||
<collectingRegion id="8540C6F4CA01FFAFFC47DD52D4F9FB82" box="[954,1185,1055,1081]" country="Vietnam" name="Ninh Thuan" pageId="6" pageNumber="381">Ninh Thuan Province</collectingRegion>
|
||
,
|
||
<collectingCountry id="3F934886CA01FFAFFB51DD52D55DFB82" box="[1196,1285,1055,1081]" name="Vietnam" pageId="6" pageNumber="381">Vietnam</collectingCountry>
|
||
(
|
||
<geoCoordinate id="22B06ED1CA01FFAFFAE8DD52D5C0FB82" box="[1301,1432,1055,1081]" degrees="11.72343" direction="north" orientation="latitude" pageId="6" pageNumber="381" precision="1" value="11.72343">11.72343°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="22B06ED1CA01FFAFFF6ADD0ED17DFBE5" box="[151,293,1091,1118]" degrees="109.13656" direction="east" orientation="longitude" pageId="6" pageNumber="381" precision="1" value="109.13656">109.13656°E</geoCoordinate>
|
||
; altitude
|
||
<elevation id="CCA9EF25CA01FFAFFE7EDD0ED1A3FBE6" box="[387,507,1091,1118]" metricMagnitude="2" metricUnit="m" metricValue="7.3" pageId="6" pageNumber="381" unit="m" value="730.0">
|
||
<quantity id="807CA5F3CA01FFAFFE7EDD0ED19EFBE5" box="[387,454,1091,1118]" metricMagnitude="2" metricUnit="m" metricValue="7.3" pageId="6" pageNumber="381" unit="m" value="730.0">730 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
) by
|
||
<collectorName id="EA716DC0CA01FFAFFDD1DD0ED293FBE6" box="[556,715,1091,1117]" pageId="6" pageNumber="381">N.A. Poyarkov</collectorName>
|
||
and
|
||
<collectorName id="EA716DC0CA01FFAFFD01DD0ED3D9FBE6" box="[764,897,1091,1117]" pageId="6" pageNumber="381">A.M. Bragin</collectorName>
|
||
on
|
||
<date id="333A2ED6CA01FFAFFC54DD0ED41EFBE6" box="[937,1094,1091,1117]" pageId="6" pageNumber="381" value="2023-03-04">
|
||
<collectingDate id="237ED73ECA01FFAFFC54DD0ED41EFBE6" box="[937,1094,1091,1117]" pageId="6" pageNumber="381" value="2023-03-04">March 4, 2023</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="F7EC024BCA01FFAFFBACDD09D097FB72" ID-GBIF-Occurrence="4904896301" collectingDate="2023-02-25" collectionCode="ZMMU" collectorName=" N. A. Poyarkov & A. M. Bragin" country="Vietnam" elevation="400" latitude="11.99353" location="Phuoc Binh Commune" longLatPrecision="1" longitude="108.74379" municipality="Bac Ai District" pageId="6" pageNumber="381" specimenCount="2" specimenCount-adult="2" stateProvince="Ninh Thuan" typeStatus="paratype">
|
||
<collectionCode id="219590D3CA01FFAFFBACDD09D4FEFBE5" box="[1105,1190,1092,1118]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="6" pageNumber="381" type="Museum">ZMMU</collectionCode>
|
||
Re-17683–84 (field ID NAP-13365–66),
|
||
<specimenCount id="5182C39FCA01FFAFFEA9DD25D1EEFB3A" box="[340,438,1127,1154]" count="2" pageId="6" pageNumber="381" type="adult">two adult</specimenCount>
|
||
females, collected from
|
||
<collectorName id="EA716DC0CA01FFAFFD47DD2AD332FB39" box="[698,874,1127,1154]" pageId="6" pageNumber="381">Phuoc Binh N.P.</collectorName>
|
||
,
|
||
<location id="425B5ECDCA01FFAFFC88DD2AD430FB3A" LSID="urn:lsid:plazi:treatment:CF2DB900CA01FFA5FF6AD8FAD4FAFF42:425B5ECDCA01FFAFFC88DD2AD430FB3A" box="[885,1128,1127,1153]" country="Vietnam" latitude="11.99353" longLatPrecision="1" longitude="108.74379" municipality="Bac Ai District" name="Phuoc Binh Commune" pageId="6" pageNumber="381" stateProvince="Ninh Thuan">Phuoc Binh Commune</location>
|
||
,
|
||
<collectingMunicipality id="A75F926CCA01FFAFFB8EDD25D54BFB3A" box="[1139,1299,1127,1154]" pageId="6" pageNumber="381">Bac Ai District</collectingMunicipality>
|
||
,
|
||
<collectingRegion id="8540C6F4CA01FFAFFAE0DD2AD0AEFB1E" country="Vietnam" name="Ninh Thuan" pageId="6" pageNumber="381">Ninh Thuan Province</collectingRegion>
|
||
,
|
||
<collectingCountry id="3F934886CA01FFAFFEFCDDC6D102FB1E" box="[257,346,1163,1189]" name="Vietnam" pageId="6" pageNumber="381">Vietnam</collectingCountry>
|
||
(
|
||
<geoCoordinate id="22B06ED1CA01FFAFFE94DDC6D1B4FB1D" box="[361,492,1163,1190]" degrees="11.99353" direction="north" orientation="latitude" pageId="6" pageNumber="381" precision="1" value="11.99353">11.99353°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="22B06ED1CA01FFAFFE0ADDC6D2DDFB1E" box="[503,645,1163,1189]" degrees="108.74379" direction="east" orientation="longitude" pageId="6" pageNumber="381" precision="1" value="108.74379">108.74379°E</geoCoordinate>
|
||
; altitude
|
||
<elevation id="CCA9EF25CA01FFAFFD18DDC6D307FB1E" box="[741,863,1163,1190]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="6" pageNumber="381" unit="m" value="400.0">
|
||
<quantity id="807CA5F3CA01FFAFFD18DDC6D371FB1D" box="[741,809,1163,1190]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="6" pageNumber="381" unit="m" value="400.0">400 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
) by
|
||
<collectorName id="EA716DC0CA01FFAFFC6FDDC6D469FB1E" box="[914,1073,1163,1189]" pageId="6" pageNumber="381">N.A. Poyarkov</collectorName>
|
||
and
|
||
<collectorName id="EA716DC0CA01FFAFFB98DDC6D4B3FB1E" box="[1125,1259,1163,1189]" pageId="6" pageNumber="381">A.M. Bragin</collectorName>
|
||
on
|
||
<date id="333A2ED6CA01FFAFFAE8DDC6D097FB72" pageId="6" pageNumber="381" value="2023-02-25">
|
||
<collectingDate id="237ED73ECA01FFAFFAE8DDC6D097FB72" pageId="6" pageNumber="381" value="2023-02-25">February 25, 2023</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="F7EC024BCA01FFAFFF27DDFDD1FAFAAA" ID-GBIF-Occurrence="4904896302" collectingDate="2023-05-15" collectionCode="DTU" collectorName="T. V. Phan & T. D. Nguyen & T. V. Nguyen" country="Vietnam" county="Cam Lam District" elevation="130" latitude="12.134169" location="Hon Ba" longLatPrecision="1" longitude="109.034454" municipality="Suoi Cat Commune" pageId="6" pageNumber="381" specimenCode="DTU 605" specimenCount="1" stateProvince="Khanh Hoa" typeStatus="paratype">
|
||
<specimenCode id="1722A06DCA01FFAFFF27DDFDD119FB71" box="[218,321,1199,1226]" collectionCode="DTU" pageId="6" pageNumber="381">DTU 605</specimenCode>
|
||
(adult female) from a secondary forest near
|
||
<location id="425B5ECDCA01FFAFFCE7DDFDD334FB71" LSID="urn:lsid:plazi:treatment:CF2DB900CA01FFA5FF6AD8FAD4FAFF42:425B5ECDCA01FFAFFCE7DDFDD334FB71" box="[794,876,1200,1226]" country="Vietnam" county="Cam Lam District" latitude="12.134169" longLatPrecision="1" longitude="109.034454" municipality="Suoi Cat Commune" name="Hon Ba" pageId="6" pageNumber="381" stateProvince="Khanh Hoa">Hon Ba</location>
|
||
NR,
|
||
<collectingMunicipality id="A75F926CCA01FFAFFC5ADDE2D424FB72" box="[935,1148,1199,1225]" pageId="6" pageNumber="381">Suoi Cat Commune</collectingMunicipality>
|
||
,
|
||
<collectingCounty id="AE5A709ACA01FFAFFB7ADDE2D512FB72" box="[1159,1354,1199,1226]" pageId="6" pageNumber="381">Cam Lam District</collectingCounty>
|
||
,
|
||
<collectingRegion id="8540C6F4CA01FFAFFAA8DDE2D172FB56" country="Vietnam" name="Khanh Hoa" pageId="6" pageNumber="381">Khanh Hoa Province</collectingRegion>
|
||
,
|
||
<collectingCountry id="3F934886CA01FFAFFEC9DD9ED1D5FB56" box="[308,397,1235,1261]" name="Vietnam" pageId="6" pageNumber="381">Vietnam</collectingCountry>
|
||
(
|
||
<geoCoordinate id="22B06ED1CA01FFAFFE61DD9ED275FB56" box="[412,557,1235,1261]" degrees="12.134169" direction="north" orientation="latitude" pageId="6" pageNumber="381" precision="1" value="12.134169">12.134169°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="22B06ED1CA01FFAFFDC5DD9ED28AFB55" box="[568,722,1235,1262]" degrees="109.034452" direction="east" orientation="longitude" pageId="6" pageNumber="381" precision="1" value="109.034454">109.034452°E</geoCoordinate>
|
||
; altitude
|
||
<elevation id="CCA9EF25CA01FFAFFCCFDD9ED3F3FB56" box="[818,939,1235,1262]" metricMagnitude="2" metricUnit="m" metricValue="1.3" pageId="6" pageNumber="381" unit="m" value="130.0">
|
||
<quantity id="807CA5F3CA01FFAFFCCFDD9ED32EFB55" box="[818,886,1235,1262]" metricMagnitude="2" metricUnit="m" metricValue="1.3" pageId="6" pageNumber="381" unit="m" value="130.0">130 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
) collected by
|
||
<collectorName id="EA716DC0CA01FFAFFBBCDD99D4F2FB56" box="[1089,1194,1235,1262]" pageId="6" pageNumber="381">T.V. Phan</collectorName>
|
||
,
|
||
<collectorName id="EA716DC0CA01FFAFFB49DD99D566FB55" box="[1204,1342,1236,1262]" pageId="6" pageNumber="381">T.D. Nguyen</collectorName>
|
||
and
|
||
<collectorName id="EA716DC0CA01FFAFFA8CDD99D0B3FAA9" pageId="6" pageNumber="381">T.V. Nguyen</collectorName>
|
||
on
|
||
<date id="333A2ED6CA01FFAFFEF2DDB5D1FAFAAA" box="[271,418,1271,1298]" pageId="6" pageNumber="381" value="2023-05-15">
|
||
<collectingDate id="237ED73ECA01FFAFFEF2DDB5D1FAFAAA" box="[271,418,1271,1298]" pageId="6" pageNumber="381" value="2023-05-15">May 15, 2023</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="F7EC024BCA01FFAFFE57DDB5D5C0FA8E" ID-GBIF-Occurrence="4904896309" collectingDate="2023-05-20" collectionCode="DTU" collectorName="T. V. Phan & T. D. Nguyen & T. V. Nguyen" country="Vietnam" elevation="90" latitude="12.327065" location="An Chua Area" longLatPrecision="1" longitude="109.098045" municipality="Diem Khanh Distrist" pageId="6" pageNumber="381" specimenCode="DTU 606" specimenCount="1" stateProvince="Khanh Hoa" typeStatus="paratype">
|
||
<specimenCode id="1722A06DCA01FFAFFE57DDB5D256FAAA" box="[426,526,1271,1298]" collectionCode="DTU" pageId="6" pageNumber="381">DTU 606</specimenCode>
|
||
(adult female) from
|
||
<location id="425B5ECDCA01FFAFFD23DDBAD32CFAAA" LSID="urn:lsid:plazi:treatment:CF2DB900CA01FFA5FF6AD8FAD4FAFF42:425B5ECDCA01FFAFFD23DDBAD32CFAAA" box="[734,884,1271,1297]" country="Vietnam" latitude="12.327065" longLatPrecision="1" longitude="109.098045" municipality="Diem Khanh Distrist" name="An Chua Area" pageId="6" pageNumber="381" stateProvince="Khanh Hoa">An Chua Area</location>
|
||
,
|
||
<collectingMunicipality id="A75F926CCA01FFAFFC80DDBAD40EFAAA" box="[893,1110,1271,1297]" pageId="6" pageNumber="381">Diem Khanh Distrist</collectingMunicipality>
|
||
,
|
||
<collectingRegion id="8540C6F4CA01FFAFFBA2DDBAD562FAAA" box="[1119,1338,1271,1298]" country="Vietnam" name="Khanh Hoa" pageId="6" pageNumber="381">Khanh Hoa Province</collectingRegion>
|
||
,
|
||
<collectingCountry id="3F934886CA01FFAFFABFDDBAD5C4FAAA" box="[1346,1436,1271,1297]" name="Vietnam" pageId="6" pageNumber="381">Vietnam</collectingCountry>
|
||
(
|
||
<geoCoordinate id="22B06ED1CA01FFAFFF62DC56D168FA8D" box="[159,304,1307,1334]" degrees="12.327065" direction="north" orientation="latitude" pageId="6" pageNumber="381" precision="1" value="12.327065">12.327065°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="22B06ED1CA01FFAFFEC4DC56D18CFA8D" box="[313,468,1307,1334]" degrees="109.098045" direction="east" orientation="longitude" pageId="6" pageNumber="381" precision="1" value="109.098045">109.098045°E</geoCoordinate>
|
||
; altitude
|
||
<elevation id="CCA9EF25CA01FFAFFDCDDC56D2C1FA8E" box="[560,665,1307,1334]" metricMagnitude="1" metricUnit="m" metricValue="9.0" pageId="6" pageNumber="381" unit="m" value="90.0">
|
||
<quantity id="807CA5F3CA01FFAFFDCDDC56D23DFA8D" box="[560,613,1307,1334]" metricMagnitude="1" metricUnit="m" metricValue="9.0" pageId="6" pageNumber="381" unit="m" value="90.0">90 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
) collected by
|
||
<collectorName id="EA716DC0CA01FFAFFCD6DC51D3CAFA8E" box="[811,914,1307,1334]" pageId="6" pageNumber="381">T.V. Phan</collectorName>
|
||
,
|
||
<collectorName id="EA716DC0CA01FFAFFC66DC51D47DFA8D" box="[923,1061,1308,1334]" pageId="6" pageNumber="381">T.D. Nguyen</collectorName>
|
||
, and
|
||
<collectorName id="EA716DC0CA01FFAFFBA7DC51D486FA8D" box="[1114,1246,1308,1334]" pageId="6" pageNumber="381">T.V. Nguyen</collectorName>
|
||
on
|
||
<date id="333A2ED6CA01FFAFFAFEDC51D5C0FA8E" box="[1283,1432,1307,1334]" pageId="6" pageNumber="381" value="2023-05-20">
|
||
<collectingDate id="237ED73ECA01FFAFFAFEDC51D5C0FA8E" box="[1283,1432,1307,1334]" pageId="6" pageNumber="381" value="2023-05-20">May 20, 2023</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="F7EC024BCA01FFAFFF6ADC0DD529FAC6" ID-GBIF-Occurrence="4904896311" collectingDate="2023-05-19" collectionCode="DTU" collectorName="T. V. Phan & T. D. Nguyen & T. V. Nguyen" country="Vietnam" elevation="45" latitude="12.397168" location="Hon Heo Island" longLatPrecision="1" longitude="109.294044" pageId="6" pageNumber="381" specimenCode="DTU 650" specimenCount="1" stateProvince="Khanh Hoa" typeStatus="paratype">
|
||
<specimenCode id="1722A06DCA01FFAFFF6ADC0DD0A5FAE1" box="[151,253,1343,1370]" collectionCode="DTU" pageId="6" pageNumber="381">DTU 650</specimenCode>
|
||
(adult female) collected by
|
||
<collectorName id="EA716DC0CA01FFAFFDE2DC0DD2DEFAE2" box="[543,646,1343,1370]" pageId="6" pageNumber="381">T.V. Phan</collectorName>
|
||
,
|
||
<collectorName id="EA716DC0CA01FFAFFD6DDC0DD342FAE1" box="[656,794,1344,1370]" pageId="6" pageNumber="381">T.D. Nguyen</collectorName>
|
||
, and
|
||
<collectorName id="EA716DC0CA01FFAFFCADDC0DD38DFAE1" box="[848,981,1344,1370]" pageId="6" pageNumber="381">T.V. Nguyen</collectorName>
|
||
from
|
||
<location id="425B5ECDCA01FFAFFBE9DC0DD4E4FAE2" LSID="urn:lsid:plazi:treatment:CF2DB900CA01FFA5FF6AD8FAD4FAFF42:425B5ECDCA01FFAFFBE9DC0DD4E4FAE2" box="[1044,1212,1343,1370]" country="Vietnam" latitude="12.397168" longLatPrecision="1" longitude="109.294044" name="Hon Heo Island" pageId="6" pageNumber="381" stateProvince="Khanh Hoa">Hon Heo Island</location>
|
||
, Ninh Da Commune, Ninh Hoa District,
|
||
<collectingRegion id="8540C6F4CA01FFAFFEA3DC2ED265FAC6" box="[350,573,1379,1406]" country="Vietnam" name="Khanh Hoa" pageId="6" pageNumber="381">Khanh Hoa Province</collectingRegion>
|
||
,
|
||
<collectingCountry id="3F934886CA01FFAFFDBADC2ED2F8FAC6" box="[583,672,1379,1405]" name="Vietnam" pageId="6" pageNumber="381">Vietnam</collectingCountry>
|
||
(
|
||
<geoCoordinate id="22B06ED1CA01FFAFFD50DC2ED366FAC6" box="[685,830,1379,1405]" degrees="12.397168" direction="north" orientation="latitude" pageId="6" pageNumber="381" precision="1" value="12.397168">12.397168°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="22B06ED1CA01FFAFFCB5DC2ED3BBFAC6" box="[840,995,1379,1405]" degrees="109.294047" direction="east" orientation="longitude" pageId="6" pageNumber="381" precision="1" value="109.294044">109.294047°E</geoCoordinate>
|
||
; altitude
|
||
<elevation id="CCA9EF25CA01FFAFFBBCDC2ED4F3FAC6" box="[1089,1195,1379,1406]" metricMagnitude="1" metricUnit="m" metricValue="4.5" pageId="6" pageNumber="381" unit="m" value="45.0">
|
||
<quantity id="807CA5F3CA01FFAFFBBCDC2ED42EFAC5" box="[1089,1142,1379,1406]" metricMagnitude="1" metricUnit="m" metricValue="4.5" pageId="6" pageNumber="381" unit="m" value="45.0">45 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
) on
|
||
<date id="333A2ED6CA01FFAFFB26DC29D529FAC6" box="[1243,1393,1379,1406]" pageId="6" pageNumber="381" value="2023-05-19">
|
||
<collectingDate id="237ED73ECA01FFAFFB26DC29D529FAC6" box="[1243,1393,1379,1406]" pageId="6" pageNumber="381" value="2023-05-19">May 19, 2023</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="473B0816CA01FFAFFF3ADCCAD136F9EE" blockId="6.[151,1437,767,2017]" pageId="6" pageNumber="381">
|
||
<emphasis id="75F0D404CA01FFAFFF3ADCCAD1E6FA1A" bold="true" box="[199,446,1415,1441]" pageId="6" pageNumber="381">Additional materials</emphasis>
|
||
(n = 3).
|
||
<materialsCitation id="F7EC024BCA01FFAFFDDFDCC5D2EDFA7E" ID-GBIF-Occurrence="4904896310" collectionCode="ZFMK" country="Vietnam" elevation="140" latitude="14.707444" location="Ba To District" longLatPrecision="1" longitude="108.997215" pageId="6" pageNumber="381" specimenCode="ZFMK 94670" specimenCount="1" stateProvince="Quang Ngai">
|
||
<specimenCode id="1722A06DCA01FFAFFDDFDCC5D2EFFA1A" box="[546,695,1415,1442]" collectionCode="ZFMK" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="6" pageNumber="381" type="Museum">ZFMK 94670</specimenCode>
|
||
(adult male) from
|
||
<location id="425B5ECDCA01FFAFFC82DCC5D44FFA1A" LSID="urn:lsid:plazi:treatment:CF2DB900CA01FFA5FF6AD8FAD4FAFF42:425B5ECDCA01FFAFFC82DCC5D44FFA1A" box="[895,1047,1415,1442]" country="Vietnam" latitude="14.707444" longLatPrecision="1" longitude="108.997215" name="Ba To District" pageId="6" pageNumber="381" stateProvince="Quang Ngai">Ba To District</location>
|
||
,
|
||
<collectingRegion id="8540C6F4CA01FFAFFBDCDCCAD552FA1A" box="[1057,1290,1415,1441]" country="Vietnam" name="Quang Ngai" pageId="6" pageNumber="381">Quang Ngai Province</collectingRegion>
|
||
,
|
||
<collectingCountry id="3F934886CA01FFAFFAE9DCCAD536FA1A" box="[1300,1390,1415,1441]" name="Vietnam" pageId="6" pageNumber="381">Vietnam</collectingCountry>
|
||
(ca.
|
||
<geoCoordinate id="22B06ED1CA01FFAFFF6ADCE6D170FA7E" box="[151,296,1451,1477]" degrees="14.707444" direction="north" orientation="latitude" pageId="6" pageNumber="381" precision="1" value="14.707444">14.707444°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="22B06ED1CA01FFAFFECEDCE6D197FA7D" box="[307,463,1451,1478]" degrees="108.997215" direction="east" orientation="longitude" pageId="6" pageNumber="381" precision="1" value="108.997215">108.997215°E</geoCoordinate>
|
||
; altitude
|
||
<elevation id="CCA9EF25CA01FFAFFDD2DCE6D2F3FA7E" box="[559,683,1451,1478]" metricMagnitude="2" metricUnit="m" metricValue="1.4" pageId="6" pageNumber="381" unit="m" value="140.0">
|
||
<quantity id="807CA5F3CA01FFAFFDD2DCE6D22BFA7D" box="[559,627,1451,1478]" metricMagnitude="2" metricUnit="m" metricValue="1.4" pageId="6" pageNumber="381" unit="m" value="140.0">140 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="F7EC024BCA01FFAFFD3DDCE1D202F9B6" ID-GBIF-Occurrence="4904896306" collectingDate="2023-08-03" collectionCode="DTU" collectorName="T. V. Phan & T. D. Nguyen" country="Vietnam" county="Quy Nhon City" elevation="280" latitude="13.695361" location="Suoi Mo Area" longLatPrecision="1" longitude="109.17055" municipality="Bui Thi Xuan Town" pageId="6" pageNumber="381" specimenCode="DTU 653" specimenCount="1" stateProvince="Binh Dinh">
|
||
<specimenCode id="1722A06DCA01FFAFFD3DDCE1D37EFA7D" box="[704,806,1451,1478]" collectionCode="DTU" pageId="6" pageNumber="381">DTU 653</specimenCode>
|
||
(adult female, released) from
|
||
<location id="425B5ECDCA01FFAFFB9BDCE6D4A6FA7E" LSID="urn:lsid:plazi:treatment:CF2DB900CA01FFA5FF6AD8FAD4FAFF42:425B5ECDCA01FFAFFB9BDCE6D4A6FA7E" box="[1126,1278,1451,1478]" country="Vietnam" county="Quy Nhon City" latitude="13.695361" longLatPrecision="1" longitude="109.17055" municipality="Bui Thi Xuan Town" name="Suoi Mo Area" pageId="6" pageNumber="381" stateProvince="Binh Dinh">Suoi Mo Area</location>
|
||
,
|
||
<collectingMunicipality id="A75F926CCA01FFAFFAF4DCE6D08EFA51" pageId="6" pageNumber="381">Bui Thi Xuan Town</collectingMunicipality>
|
||
,
|
||
<collectingCounty id="AE5A709ACA01FFAFFF1DDC82D1DBFA52" box="[224,387,1487,1513]" pageId="6" pageNumber="381">Quy Nhon City</collectingCounty>
|
||
,
|
||
<collectingRegion id="8540C6F4CA01FFAFFE71DC82D238FA52" box="[396,608,1487,1513]" country="Vietnam" name="Binh Dinh" pageId="6" pageNumber="381">Binh Dinh Province</collectingRegion>
|
||
,
|
||
<collectingCountry id="3F934886CA01FFAFFD97DC82D29CFA52" box="[618,708,1487,1513]" name="Vietnam" pageId="6" pageNumber="381">Vietnam</collectingCountry>
|
||
(
|
||
<geoCoordinate id="22B06ED1CA01FFAFFD2CDC82D33BFA51" box="[721,867,1487,1514]" degrees="13.695361" direction="north" orientation="latitude" pageId="6" pageNumber="381" precision="1" value="13.695361">13.695361°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="22B06ED1CA01FFAFFC90DC82D451FA51" box="[877,1033,1487,1514]" degrees="109.170546" direction="east" orientation="longitude" pageId="6" pageNumber="381" precision="1" value="109.17055">109.170546°E</geoCoordinate>
|
||
; altitude
|
||
<elevation id="CCA9EF25CA01FFAFFB9BDC82D486FA52" box="[1126,1246,1487,1514]" metricMagnitude="2" metricUnit="m" metricValue="2.8" pageId="6" pageNumber="381" unit="m" value="280.0">
|
||
<quantity id="807CA5F3CA01FFAFFB9BDC82D4F1FA51" box="[1126,1193,1487,1514]" metricMagnitude="2" metricUnit="m" metricValue="2.8" pageId="6" pageNumber="381" unit="m" value="280.0">280 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
) collected by
|
||
<collectorName id="EA716DC0CA01FFAFFA8CDC9DD094F9B6" pageId="6" pageNumber="381">T.V. Phan</collectorName>
|
||
and
|
||
<collectorName id="EA716DC0CA01FFAFFF00DCB9D1DDF9B5" box="[253,389,1524,1550]" pageId="6" pageNumber="381">T.D. Nguyen</collectorName>
|
||
on
|
||
<date id="333A2ED6CA01FFAFFE54DCBED202F9B6" box="[425,602,1523,1549]" pageId="6" pageNumber="381" value="2023-08-03">
|
||
<collectingDate id="237ED73ECA01FFAFFE54DCBED202F9B6" box="[425,602,1523,1549]" pageId="6" pageNumber="381" value="2023-08-03">August 03, 2023</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="F7EC024BCA01FFAFFD99DCB9D132F9EE" ID-GBIF-Occurrence="4904896305" collectingDate="2023-08-10" collectionCode="DTU" collectorName="T. V. Phan & T. D. Nguyen" country="Vietnam" elevation="250" latitude="13.6643" location="Hiep Ha Commune" longLatPrecision="1" longitude="108.97258" municipality="Van Canh District" pageId="6" pageNumber="381" specimenCode="DTU 636" specimenCount="1" stateProvince="Binh Dinh">
|
||
<specimenCode id="1722A06DCA01FFAFFD99DCB9D291F9B6" box="[612,713,1523,1550]" collectionCode="DTU" pageId="6" pageNumber="381">DTU 636</specimenCode>
|
||
(adult female, released) from
|
||
<location id="425B5ECDCA01FFAFFBFCDCBED489F9B6" LSID="urn:lsid:plazi:treatment:CF2DB900CA01FFA5FF6AD8FAD4FAFF42:425B5ECDCA01FFAFFBFCDCBED489F9B6" box="[1025,1233,1523,1550]" country="Vietnam" latitude="13.6643" longLatPrecision="1" longitude="108.97258" municipality="Van Canh District" name="Hiep Ha Commune" pageId="6" pageNumber="381" stateProvince="Binh Dinh">Hiep Ha Commune</location>
|
||
,
|
||
<collectingMunicipality id="A75F926CCA01FFAFFB27DCB9D5C1F9B6" box="[1242,1433,1523,1550]" pageId="6" pageNumber="381">Van Canh District</collectingMunicipality>
|
||
,
|
||
<collectingRegion id="8540C6F4CA01FFAFFF6ADF5AD133F98A" box="[151,363,1559,1585]" country="Vietnam" name="Binh Dinh" pageId="6" pageNumber="381">Binh Dinh Province</collectingRegion>
|
||
,
|
||
<collectingCountry id="3F934886CA01FFAFFE88DF5AD197F98A" box="[373,463,1559,1585]" name="Vietnam" pageId="6" pageNumber="381">Vietnam</collectingCountry>
|
||
(
|
||
<geoCoordinate id="22B06ED1CA01FFAFFE21DF5AD236F98A" box="[476,622,1559,1585]" degrees="13.664300" direction="north" orientation="latitude" pageId="6" pageNumber="381" precision="1" value="13.6643">13.664300°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="22B06ED1CA01FFAFFD85DF5AD34CF989" box="[632,788,1559,1586]" degrees="108.972583" direction="east" orientation="longitude" pageId="6" pageNumber="381" precision="1" value="108.97258">108.972583°E</geoCoordinate>
|
||
; altitude
|
||
<elevation id="CCA9EF25CA01FFAFFC8CDF5AD3B1F98A" box="[881,1001,1559,1586]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="6" pageNumber="381" unit="m" value="250.0">
|
||
<quantity id="807CA5F3CA01FFAFFC8CDF5AD3ECF989" box="[881,948,1559,1586]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="6" pageNumber="381" unit="m" value="250.0">250 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
) collected by
|
||
<collectorName id="EA716DC0CA01FFAFFB81DF55D4BAF98A" box="[1148,1250,1559,1586]" pageId="6" pageNumber="381">T.V. Phan</collectorName>
|
||
and
|
||
<collectorName id="EA716DC0CA01FFAFFAEEDF55D5C4F989" box="[1299,1436,1560,1586]" pageId="6" pageNumber="381">T.D. Nguyen</collectorName>
|
||
on
|
||
<date id="333A2ED6CA01FFAFFF4ADF76D132F9EE" box="[183,362,1595,1621]" pageId="6" pageNumber="381" value="2023-08-10">
|
||
<collectingDate id="237ED73ECA01FFAFFF4ADF76D132F9EE" box="[183,362,1595,1621]" pageId="6" pageNumber="381" value="2023-08-10">August 10, 2023</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0F9E5B9DCA01FFAFFF3ADF12D404F8EF" pageId="6" pageNumber="381" type="etymology">
|
||
<paragraph id="473B0816CA01FFAFFF3ADF12D404F8EF" blockId="6.[151,1437,767,2017]" pageId="6" pageNumber="381">
|
||
<emphasis id="75F0D404CA01FFAFFF3ADF12D11AF9C2" bold="true" box="[199,322,1631,1657]" pageId="6" pageNumber="381">Etymology</emphasis>
|
||
. The specific name
|
||
<taxonomicName id="80847395CA01FFAFFDE2DF12D2FFF9C2" authorityName="Idiiatullina & Nguyen & Bragin & Pawangkhanant & Le & Vogel & David & Poyarkov" authorityYear="2024" box="[543,679,1631,1657]" class="Squamata" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" pageId="6" pageNumber="381" phylum="Chordata" rank="species" species="cyanolabris">
|
||
“
|
||
<emphasis id="75F0D404CA01FFAFFDD7DF12D2FFF9C2" box="[554,679,1631,1657]" italics="true" pageId="6" pageNumber="381">cyanolabris</emphasis>
|
||
</taxonomicName>
|
||
” is formed of the Latin adjective “
|
||
<emphasis id="75F0D404CA01FFAFFBDCDF2DD42EF9C2" box="[1057,1142,1632,1657]" italics="true" pageId="6" pageNumber="381">cyaneus</emphasis>
|
||
,” derived from a Latizined form of the classical Greek adjective
|
||
<emphasis id="75F0D404CA01FFAFFDE5DFC9D23BF925" box="[536,611,1668,1694]" italics="true" pageId="6" pageNumber="381">κυαΝΟΣ</emphasis>
|
||
(
|
||
<emphasis id="75F0D404CA01FFAFFD8DDFCED2E2F926" box="[624,698,1667,1693]" italics="true" pageId="6" pageNumber="381">kyanos</emphasis>
|
||
), meaning “dark blue” or “deep blue,” and of the Latin noun
|
||
<emphasis id="75F0D404CA01FFAFFACCDFCED5C4F926" box="[1329,1436,1667,1693]" italics="true" pageId="6" pageNumber="381">“labrum”</emphasis>
|
||
(-
|
||
<emphasis id="75F0D404CA01FFAFFF55DFE5D0F7F97A" box="[168,175,1704,1729]" italics="true" pageId="6" pageNumber="381">i</emphasis>
|
||
), the lip. This specific name means “deep blue-lipped,” and refers to the distinctive blue color found on the throat, chin, and infralabials of the new species. Though traditionally it is recommended that the Greek and Latin words not be combined in one species nomen, we would like to underline that the adjective “
|
||
<emphasis id="75F0D404CA01FFAFFCA3DFBDD3EBF8B2" box="[862,947,1776,1801]" italics="true" pageId="6" pageNumber="381">cyaneus</emphasis>
|
||
” is common in Latin texts and can be regarded as a proper Latin adjective. We suggest the following common English name: “Blue-lipped Green Pitviper”, Russian name: “Golubogubaya bambukovaya kufiya”, and Vietnamese name: “Rắn lục mép xanh dƯƠng”.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0F9E5B9DCA01FFAEFF3ADE16D579FEA6" lastPageId="7" lastPageNumber="382" pageId="6" pageNumber="381" type="diagnosis">
|
||
<paragraph id="473B0816CA01FFAEFF3ADE16D579FEA6" blockId="6.[151,1437,767,2017]" lastBlockId="7.[151,1436,150,285]" lastPageId="7" lastPageNumber="382" pageId="6" pageNumber="381">
|
||
<emphasis id="75F0D404CA01FFAFFF3ADE16D16CF8CE" bold="true" box="[199,308,1883,1909]" pageId="6" pageNumber="381">Diagnosis</emphasis>
|
||
. The new species is assigned to the subgenus
|
||
<taxonomicName id="80847395CA01FFAFFCABDE11D3BBF8CE" box="[854,995,1884,1909]" pageId="6" pageNumber="381" rank="subGenus" subGenus="Trimeresurus">
|
||
<emphasis id="75F0D404CA01FFAFFCABDE11D3BBF8CE" box="[854,995,1884,1909]" italics="true" pageId="6" pageNumber="381">Trimeresurus</emphasis>
|
||
</taxonomicName>
|
||
based on the following morphological attributes: a long calyculate hemipenis and a partially fused first supralabial and nasal scales (
|
||
<bibRefCitation id="231575E7CA01FFAFFB74DE32D5C4F822" author="Malhotra, A. & Thorpe, R. S." box="[1161,1436,1919,1945]" pageId="6" pageNumber="381" pagination="83 - 100" refId="ref18749" refString="Malhotra, A. & Thorpe, R. S. (2004) A phylogeny of four mitochondrial gene regions suggests a revised taxonomy for Asian pitvipers (Trimeresurus and Ovophis). Molecular Phylogenetics and Evolution, 32, 83 - 100. https: // doi. org / 10.1016 / j. ympev. 2004.02.008" type="journal article" year="2004">Malhotra & Thorpe 2004</bibRefCitation>
|
||
[as
|
||
<taxonomicName id="80847395CA01FFAFFF41DEEED114F806" authorityName="Cope" authorityYear="1859" box="[188,332,1955,1981]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="381" phylum="Chordata" rank="genus">
|
||
<emphasis id="75F0D404CA01FFAFFF41DEEED114F806" box="[188,332,1955,1981]" italics="true" pageId="6" pageNumber="381">Cryptelytrops</emphasis>
|
||
</taxonomicName>
|
||
];
|
||
<bibRefCitation id="231575E7CA01FFAFFE9FDEEED243F806" author="David, P. & Vogel, G. & Dubois, A." box="[354,539,1955,1982]" pageId="6" pageNumber="381" pagination="1 - 51" refId="ref16865" refString="David, P., Vogel, G. & Dubois, A. (2011) On the need to follow rigorously the rules of the code for the subsequent designation of a nucleospecies (type species) for a nominal genus which lacked one: the case of the nominal genus Trimeresurus Lacepede, 1804 (Reptilia: Squamata: Viperidae). Zootaxa, 2992 (1), 1 - 51. https: // doi. org / 10.11646 / zootaxa. 2992.1.1" type="journal article" year="2011">
|
||
David
|
||
<emphasis id="75F0D404CA01FFAFFE54DEE9D18EF806" box="[425,470,1955,1981]" italics="true" pageId="6" pageNumber="381">et al</emphasis>
|
||
. 2011
|
||
</bibRefCitation>
|
||
). The new species,
|
||
<taxonomicName id="80847395CA01FFAFFD1ADEE9D3AEF806" authorityName="Idiiatullina & Nguyen & Bragin & Pawangkhanant & Le & Vogel & David & Poyarkov" authorityYear="2024" box="[743,1014,1955,1981]" class="Squamata" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" pageId="6" pageNumber="381" phylum="Chordata" rank="species" species="cyanolabris" status="sp. nov.">
|
||
<emphasis id="75F0D404CA01FFAFFD1ADEE9D3AEF806" box="[743,1014,1955,1981]" italics="true" pageId="6" pageNumber="381">Trimeresurus cyanolabris</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="6EC3697FCA01FFAFFC01DEEED409F805" box="[1020,1105,1955,1982]" pageId="6" pageNumber="381" rank="species">
|
||
<emphasis id="75F0D404CA01FFAFFC01DEEED412F806" bold="true" box="[1020,1098,1955,1981]" pageId="6" pageNumber="381">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
, differs from other members of the subgenus
|
||
<taxonomicName id="80847395CA01FFAFFED7DE85D1EFF85A" box="[298,439,1992,2017]" pageId="6" pageNumber="381" rank="subGenus" subGenus="Trimeresurus">
|
||
<emphasis id="75F0D404CA01FFAFFED7DE85D1EFF85A" box="[298,439,1992,2017]" italics="true" pageId="6" pageNumber="381">Trimeresurus</emphasis>
|
||
</taxonomicName>
|
||
by the combination of the following morphological characters: small size, maximum known SVL of
|
||
<quantity id="807CA5F3CA00FFAEFF11D9DAD110FF0A" box="[236,328,151,177]" metricMagnitude="-1" metricUnit="m" metricValue="6.38" pageId="7" pageNumber="382" unit="mm" value="638.0">638 mm</quantity>
|
||
; dorsal scales in 21 (rarely 23)–21–15 rows, moderately keeled except the outermost rows; ventral scales 166–178; subcaudal scales 52–75, all paired; hemipenis forked, calyculate, reaching the 8
|
||
<superScript id="B0F1A55ECA00FFAEFBBCD9F7D415FF73" attach="left" box="[1089,1101,186,200]" fontSize="6" pageId="7" pageNumber="382">th</superScript>
|
||
subcaudal; eye bright yellow in both sexes; dorsal surfaces deep green lacking cross-bands; postocular streak white in both sexes; ventrolateral stripe, faint, present on the first few dorsal scale rows in males, absent in females; throat, chin, and lower labials blue color.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="13FB589ECA00FFAEFF65DECCD3B1F85A" ID-DOI="http://doi.org/10.5281/zenodo.12681174" ID-Zenodo-Dep="12681174" httpUri="https://zenodo.org/record/12681174/files/figure.png" pageId="7" pageNumber="382" startId="7.[152,255,1921,1945]" targetBox="[165,1423,309,1897]" targetPageId="7" targetType="figure">
|
||
<paragraph id="473B0816CA00FFAEFF65DECCD3B1F85A" blockId="7.[151,1436,1921,2017]" pageId="7" pageNumber="382">
|
||
<emphasis id="75F0D404CA00FFAEFF65DECCD140F822" bold="true" box="[152,280,1921,1945]" pageId="7" pageNumber="382">FIGURE 3.</emphasis>
|
||
The holotype of
|
||
<taxonomicName id="80847395CA00FFAEFE39DECFD290F822" authorityName="Idiiatullina & Nguyen & Bragin & Pawangkhanant & Le & Vogel & David & Poyarkov" authorityYear="2024" box="[452,712,1921,1945]" class="Squamata" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" pageId="7" pageNumber="382" phylum="Chordata" rank="species" species="cyanolabris" status="sp. nov.">
|
||
<emphasis id="75F0D404CA00FFAEFE39DECFD290F822" box="[452,712,1921,1945]" italics="true" pageId="7" pageNumber="382">Trimeresurus cyanolabris</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="75F0D404CA00FFAEFD33DECCD346F822" bold="true" box="[718,798,1921,1945]" pageId="7" pageNumber="382">
|
||
<taxonomicNameLabel id="6EC3697FCA00FFAEFD33DECCD346F822" box="[718,798,1921,1945]" pageId="7" pageNumber="382" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in preservative (ZMMU Re-17679, adult male) from Nui Chua NP, Ninh Thuan Province, Vietnam.
|
||
<emphasis id="75F0D404CA00FFAEFDFADEE8D241F806" bold="true" box="[519,537,1957,1981]" pageId="7" pageNumber="382">A</emphasis>
|
||
dorsolateral and
|
||
<emphasis id="75F0D404CA00FFAEFD3ADEE8D280F806" bold="true" box="[711,728,1957,1981]" pageId="7" pageNumber="382">B</emphasis>
|
||
ventral view of body;
|
||
<emphasis id="75F0D404CA00FFAEFC46DEE8D395F806" bold="true" box="[955,973,1957,1981]" pageId="7" pageNumber="382">C</emphasis>
|
||
head in dorsal,
|
||
<emphasis id="75F0D404CA00FFAEFB90DEE8D427F806" bold="true" box="[1133,1151,1957,1981]" pageId="7" pageNumber="382">D</emphasis>
|
||
in ventral,
|
||
<emphasis id="75F0D404CA00FFAEFB0CDEE8D55AF806" bold="true" box="[1265,1282,1957,1981]" pageId="7" pageNumber="382">E</emphasis>
|
||
in lateral right, and
|
||
<emphasis id="75F0D404CA00FFAEFF3FDE84D089F85A" bold="true" box="[194,209,1993,2017]" pageId="7" pageNumber="382">F</emphasis>
|
||
in lateral left aspects. Scale bar equals 1 cm. Photographs by S.S. Idiiatullina.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="13FB589ECA0FFFA1FF65DE65D107F832" ID-DOI="http://doi.org/10.5281/zenodo.12681176" ID-Zenodo-Dep="12681176" httpUri="https://zenodo.org/record/12681176/files/figure.png" pageId="8" pageNumber="383" startId="8.[152,255,1832,1856]" targetBox="[193,1395,181,1809]" targetPageId="8" targetType="figure">
|
||
<paragraph id="473B0816CA0FFFA1FF65DE65D107F832" blockId="8.[151,1437,1832,1929]" pageId="8" pageNumber="383">
|
||
<emphasis id="75F0D404CA0FFFA1FF65DE65D141F8FB" bold="true" box="[152,281,1832,1856]" pageId="8" pageNumber="383">FIGURE 4.</emphasis>
|
||
Type specimens of
|
||
<taxonomicName id="80847395CA0FFFA1FE18DE64D2B2F8FB" authorityName="Idiiatullina & Nguyen & Bragin & Pawangkhanant & Le & Vogel & David & Poyarkov" authorityYear="2024" box="[485,746,1832,1856]" class="Squamata" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" pageId="8" pageNumber="383" phylum="Chordata" rank="species" species="cyanolabris" status="sp. nov.">
|
||
<emphasis id="75F0D404CA0FFFA1FE18DE64D2B2F8FB" box="[485,746,1832,1856]" italics="true" pageId="8" pageNumber="383">Trimeresurus cyanolabris</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="75F0D404CA0FFFA1FD0CDE64D31AF8FA" bold="true" box="[753,834,1833,1857]" pageId="8" pageNumber="383">
|
||
<taxonomicNameLabel id="6EC3697FCA0FFFA1FD0CDE64D31AF8FA" box="[753,834,1833,1857]" pageId="8" pageNumber="383" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in life from Nui Chua NP, Ninh Thuan Province, Vietnam: A holotype ZMMU Re-17679, adult male; B paratype ZMMU Re-17681, adult female. Photographs by A.M. Bragin (A) and N.A. Poyarkov (B).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="0F9E5B9DCA0EFFA3FF3AD9DBD1DBF8CA" lastPageId="10" lastPageNumber="385" pageId="9" pageNumber="384" type="description">
|
||
<paragraph id="473B0816CA0EFFA0FF3AD9DBD4E3FCE7" blockId="9.[151,1437,150,2013]" pageId="9" pageNumber="384">
|
||
<emphasis id="75F0D404CA0EFFA0FF3AD9DBD24BFF0B" bold="true" box="[199,531,150,176]" pageId="9" pageNumber="384">
|
||
Description of the
|
||
<typeStatus id="983FB6B4CA0EFFA0FE58D9DBD256FF0B" box="[421,526,150,176]" pageId="9" pageNumber="384" type="holotype">holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
Adult male (
|
||
<figureCitation id="DFBF1493CA0EFFA0FD55D9DBD2B4FF0B" box="[680,748,150,176]" captionStart="FIGURE 3" captionStartId="7.[152,255,1921,1945]" captionTargetBox="[165,1423,309,1897]" captionTargetId="figure-95@7.[164,1423,309,1897]" captionTargetPageId="7" captionText="FIGURE 3. The holotype of Trimeresurus cyanolabris sp. nov. in preservative (ZMMU Re-17679, adult male) from Nui Chua NP, Ninh Thuan Province, Vietnam. A dorsolateral and B ventral view of body; C head in dorsal, D in ventral, E in lateral right, and F in lateral left aspects. Scale bar equals 1 cm. Photographs by S.S. Idiiatullina." figureDoi="http://doi.org/10.5281/zenodo.12681174" httpUri="https://zenodo.org/record/12681174/files/figure.png" pageId="9" pageNumber="384">Fig. 3</figureCitation>
|
||
), specimen in a good state of preservation. Body cylindrical, long, and thin (SVL
|
||
<quantity id="807CA5F3CA0EFFA0FE87D9F7D180FF6E" box="[378,472,186,213]" metricMagnitude="-1" metricUnit="m" metricValue="4.64" pageId="9" pageNumber="384" unit="mm" value="464.0">464 mm</quantity>
|
||
, TaL
|
||
<quantity id="807CA5F3CA0EFFA0FDE9D9F7D22AFF6E" box="[532,626,186,213]" metricMagnitude="-1" metricUnit="m" metricValue="1.06" pageId="9" pageNumber="384" unit="mm" value="106.0">106 mm</quantity>
|
||
, TL
|
||
<quantity id="807CA5F3CA0EFFA0FD5ED9F7D35AFF6E" box="[675,770,186,213]" metricMagnitude="-1" metricUnit="m" metricValue="5.7" pageId="9" pageNumber="384" unit="mm" value="570.0">570 mm</quantity>
|
||
, TaL/TL 0.186) (
|
||
<figureCitation id="DFBF1493CA0EFFA0FC43D9F7D461FF6F" box="[958,1081,186,212]" captionStart="FIGURE 3" captionStartId="7.[152,255,1921,1945]" captionTargetBox="[165,1423,309,1897]" captionTargetId="figure-95@7.[164,1423,309,1897]" captionTargetPageId="7" captionText="FIGURE 3. The holotype of Trimeresurus cyanolabris sp. nov. in preservative (ZMMU Re-17679, adult male) from Nui Chua NP, Ninh Thuan Province, Vietnam. A dorsolateral and B ventral view of body; C head in dorsal, D in ventral, E in lateral right, and F in lateral left aspects. Scale bar equals 1 cm. Photographs by S.S. Idiiatullina." figureDoi="http://doi.org/10.5281/zenodo.12681174" httpUri="https://zenodo.org/record/12681174/files/figure.png" pageId="9" pageNumber="384">Fig. 3A–B</figureCitation>
|
||
). Head triangular in dorsal view (
|
||
<figureCitation id="DFBF1493CA0EFFA0FF5DD993D0AEFF43" box="[160,246,222,248]" captionStart="FIGURE 3" captionStartId="7.[152,255,1921,1945]" captionTargetBox="[165,1423,309,1897]" captionTargetId="figure-95@7.[164,1423,309,1897]" captionTargetPageId="7" captionText="FIGURE 3. The holotype of Trimeresurus cyanolabris sp. nov. in preservative (ZMMU Re-17679, adult male) from Nui Chua NP, Ninh Thuan Province, Vietnam. A dorsolateral and B ventral view of body; C head in dorsal, D in ventral, E in lateral right, and F in lateral left aspects. Scale bar equals 1 cm. Photographs by S.S. Idiiatullina." figureDoi="http://doi.org/10.5281/zenodo.12681174" httpUri="https://zenodo.org/record/12681174/files/figure.png" pageId="9" pageNumber="384">Fig. 3C</figureCitation>
|
||
), elongate, clearly distinct from the neck (HL
|
||
<quantity id="807CA5F3CA0EFFA0FD0BD993D304FF42" box="[758,860,222,249]" metricMagnitude="-2" metricUnit="m" metricValue="2.12" pageId="9" pageNumber="384" unit="mm" value="21.2">21.2 mm</quantity>
|
||
; HW 13.0 mm; HW/HL ratio 0.61). Snout elongate, flattened, and rounded in dorsal view (
|
||
<figureCitation id="DFBF1493CA0EFFA0FDB6D84FD2FBFEA7" box="[587,675,258,284]" captionStart="FIGURE 3" captionStartId="7.[152,255,1921,1945]" captionTargetBox="[165,1423,309,1897]" captionTargetId="figure-95@7.[164,1423,309,1897]" captionTargetPageId="7" captionText="FIGURE 3. The holotype of Trimeresurus cyanolabris sp. nov. in preservative (ZMMU Re-17679, adult male) from Nui Chua NP, Ninh Thuan Province, Vietnam. A dorsolateral and B ventral view of body; C head in dorsal, D in ventral, E in lateral right, and F in lateral left aspects. Scale bar equals 1 cm. Photographs by S.S. Idiiatullina." figureDoi="http://doi.org/10.5281/zenodo.12681174" httpUri="https://zenodo.org/record/12681174/files/figure.png" pageId="9" pageNumber="384">Fig. 3C</figureCitation>
|
||
), rather rectangular in lateral view (
|
||
<figureCitation id="DFBF1493CA0EFFA0FBC4D84FD4F6FEA7" box="[1081,1198,258,284]" captionStart="FIGURE 3" captionStartId="7.[152,255,1921,1945]" captionTargetBox="[165,1423,309,1897]" captionTargetId="figure-95@7.[164,1423,309,1897]" captionTargetPageId="7" captionText="FIGURE 3. The holotype of Trimeresurus cyanolabris sp. nov. in preservative (ZMMU Re-17679, adult male) from Nui Chua NP, Ninh Thuan Province, Vietnam. A dorsolateral and B ventral view of body; C head in dorsal, D in ventral, E in lateral right, and F in lateral left aspects. Scale bar equals 1 cm. Photographs by S.S. Idiiatullina." figureDoi="http://doi.org/10.5281/zenodo.12681174" httpUri="https://zenodo.org/record/12681174/files/figure.png" pageId="9" pageNumber="384">Fig. 3E–F</figureCitation>
|
||
), with a very distinct and sharp canthus rostralis. Eyes large (ED
|
||
<quantity id="807CA5F3CA0EFFA0FD8AD86BD296FEFA" box="[631,718,294,321]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="9" pageNumber="384" unit="mm" value="3.5">3.5 mm</quantity>
|
||
; EN
|
||
<quantity id="807CA5F3CA0EFFA0FCFED86BD303FEFA" box="[771,859,294,321]" metricMagnitude="-3" metricUnit="m" metricValue="5.2" pageId="9" pageNumber="384" unit="mm" value="5.2">5.2 mm</quantity>
|
||
; ED/EN ratio 0.67). Rostral slightly visible in dorsal aspect, triangular (
|
||
<figureCitation id="DFBF1493CA0EFFA0FE91D807D19FFEDF" box="[364,455,330,356]" captionStart="FIGURE 3" captionStartId="7.[152,255,1921,1945]" captionTargetBox="[165,1423,309,1897]" captionTargetId="figure-95@7.[164,1423,309,1897]" captionTargetPageId="7" captionText="FIGURE 3. The holotype of Trimeresurus cyanolabris sp. nov. in preservative (ZMMU Re-17679, adult male) from Nui Chua NP, Ninh Thuan Province, Vietnam. A dorsolateral and B ventral view of body; C head in dorsal, D in ventral, E in lateral right, and F in lateral left aspects. Scale bar equals 1 cm. Photographs by S.S. Idiiatullina." figureDoi="http://doi.org/10.5281/zenodo.12681174" httpUri="https://zenodo.org/record/12681174/files/figure.png" pageId="9" pageNumber="384">Fig. 3C</figureCitation>
|
||
). Pupil vertically elliptical, loreal pit present, triangular in shape (
|
||
<figureCitation id="DFBF1493CA0EFFA0FB3CD807D561FEDF" box="[1217,1337,330,356]" captionStart="FIGURE 3" captionStartId="7.[152,255,1921,1945]" captionTargetBox="[165,1423,309,1897]" captionTargetId="figure-95@7.[164,1423,309,1897]" captionTargetPageId="7" captionText="FIGURE 3. The holotype of Trimeresurus cyanolabris sp. nov. in preservative (ZMMU Re-17679, adult male) from Nui Chua NP, Ninh Thuan Province, Vietnam. A dorsolateral and B ventral view of body; C head in dorsal, D in ventral, E in lateral right, and F in lateral left aspects. Scale bar equals 1 cm. Photographs by S.S. Idiiatullina." figureDoi="http://doi.org/10.5281/zenodo.12681174" httpUri="https://zenodo.org/record/12681174/files/figure.png" pageId="9" pageNumber="384">Fig. 3E–F</figureCitation>
|
||
). Nostril completely enclosed in nasal scale; nasal scale partially fused with first supralabial (
|
||
<figureCitation id="DFBF1493CA0EFFA0FBA8D823D493FE33" box="[1109,1227,366,392]" captionStart="FIGURE 3" captionStartId="7.[152,255,1921,1945]" captionTargetBox="[165,1423,309,1897]" captionTargetId="figure-95@7.[164,1423,309,1897]" captionTargetPageId="7" captionText="FIGURE 3. The holotype of Trimeresurus cyanolabris sp. nov. in preservative (ZMMU Re-17679, adult male) from Nui Chua NP, Ninh Thuan Province, Vietnam. A dorsolateral and B ventral view of body; C head in dorsal, D in ventral, E in lateral right, and F in lateral left aspects. Scale bar equals 1 cm. Photographs by S.S. Idiiatullina." figureDoi="http://doi.org/10.5281/zenodo.12681174" httpUri="https://zenodo.org/record/12681174/files/figure.png" pageId="9" pageNumber="384">Fig. 3E–F</figureCitation>
|
||
). Shield bordering the anterior edge of the loreal pit fused with second supralabial, which is tall. Anterior subocular long, thin, crescentlike, separated from the 4
|
||
<superScript id="B0F1A55ECA0EFFA0FE4BD8FBD19BFE7F" attach="left" box="[438,451,438,452]" fontSize="6" pageId="9" pageNumber="384">th</superScript>
|
||
and 5
|
||
<superScript id="B0F1A55ECA0EFFA0FDFAD8FBD24CFE7F" attach="left" box="[519,532,438,452]" fontSize="6" pageId="9" pageNumber="384">th</superScript>
|
||
supralabials by one row of scales; posterior subocular ovoid, separated from the 6
|
||
<superScript id="B0F1A55ECA0EFFA0FF58D897D0EAFE53" attach="left" box="[165,178,474,488]" fontSize="6" pageId="9" pageNumber="384">th</superScript>
|
||
supralabial by 2/1 scales (
|
||
<figureCitation id="DFBF1493CA0EFFA0FE1CD897D20FFE4F" box="[481,599,474,500]" captionStart="FIGURE 3" captionStartId="7.[152,255,1921,1945]" captionTargetBox="[165,1423,309,1897]" captionTargetId="figure-95@7.[164,1423,309,1897]" captionTargetPageId="7" captionText="FIGURE 3. The holotype of Trimeresurus cyanolabris sp. nov. in preservative (ZMMU Re-17679, adult male) from Nui Chua NP, Ninh Thuan Province, Vietnam. A dorsolateral and B ventral view of body; C head in dorsal, D in ventral, E in lateral right, and F in lateral left aspects. Scale bar equals 1 cm. Photographs by S.S. Idiiatullina." figureDoi="http://doi.org/10.5281/zenodo.12681174" httpUri="https://zenodo.org/record/12681174/files/figure.png" pageId="9" pageNumber="384">Fig. 3E–F</figureCitation>
|
||
). Three preoculars on each side of the head; two upper preoculars located above the loreal pit, elongate, in contact with the single loreal, which separates them from the nasal; lower preocular forming the lower margin of the loreal pit, lower preocular in contact with third supralabial (
|
||
<figureCitation id="DFBF1493CA0EFFA0FB47DB6FD569FD87" box="[1210,1329,546,572]" captionStart="FIGURE 3" captionStartId="7.[152,255,1921,1945]" captionTargetBox="[165,1423,309,1897]" captionTargetId="figure-95@7.[164,1423,309,1897]" captionTargetPageId="7" captionText="FIGURE 3. The holotype of Trimeresurus cyanolabris sp. nov. in preservative (ZMMU Re-17679, adult male) from Nui Chua NP, Ninh Thuan Province, Vietnam. A dorsolateral and B ventral view of body; C head in dorsal, D in ventral, E in lateral right, and F in lateral left aspects. Scale bar equals 1 cm. Photographs by S.S. Idiiatullina." figureDoi="http://doi.org/10.5281/zenodo.12681174" httpUri="https://zenodo.org/record/12681174/files/figure.png" pageId="9" pageNumber="384">Fig. 3E–F</figureCitation>
|
||
). A small scale between nasal and second supralabial; 2/2 postoculars; 10/10 supralabials, third the largest (
|
||
<figureCitation id="DFBF1493CA0EFFA0FB29DB0BD511FDDB" box="[1236,1353,582,608]" captionStart="FIGURE 3" captionStartId="7.[152,255,1921,1945]" captionTargetBox="[165,1423,309,1897]" captionTargetId="figure-95@7.[164,1423,309,1897]" captionTargetPageId="7" captionText="FIGURE 3. The holotype of Trimeresurus cyanolabris sp. nov. in preservative (ZMMU Re-17679, adult male) from Nui Chua NP, Ninh Thuan Province, Vietnam. A dorsolateral and B ventral view of body; C head in dorsal, D in ventral, E in lateral right, and F in lateral left aspects. Scale bar equals 1 cm. Photographs by S.S. Idiiatullina." figureDoi="http://doi.org/10.5281/zenodo.12681174" httpUri="https://zenodo.org/record/12681174/files/figure.png" pageId="9" pageNumber="384">Fig. 3E–F</figureCitation>
|
||
); 12/12 infralabials, those of the first pair in contact with each other behind the mental; the first three pairs of infralabials in contact with the single pair of chin shields (
|
||
<figureCitation id="DFBF1493CA0EFFA0FD83DBC3D2ABFD13" box="[638,755,654,680]" captionStart="FIGURE 3" captionStartId="7.[152,255,1921,1945]" captionTargetBox="[165,1423,309,1897]" captionTargetId="figure-95@7.[164,1423,309,1897]" captionTargetPageId="7" captionText="FIGURE 3. The holotype of Trimeresurus cyanolabris sp. nov. in preservative (ZMMU Re-17679, adult male) from Nui Chua NP, Ninh Thuan Province, Vietnam. A dorsolateral and B ventral view of body; C head in dorsal, D in ventral, E in lateral right, and F in lateral left aspects. Scale bar equals 1 cm. Photographs by S.S. Idiiatullina." figureDoi="http://doi.org/10.5281/zenodo.12681174" httpUri="https://zenodo.org/record/12681174/files/figure.png" pageId="9" pageNumber="384">Fig. 3E–F</figureCitation>
|
||
). Six pairs of gulars aligned between the chin shields and the first preventral (
|
||
<figureCitation id="DFBF1493CA0EFFA0FEB0DBFFD1F0FD77" box="[333,424,690,716]" captionStart="FIGURE 3" captionStartId="7.[152,255,1921,1945]" captionTargetBox="[165,1423,309,1897]" captionTargetId="figure-95@7.[164,1423,309,1897]" captionTargetPageId="7" captionText="FIGURE 3. The holotype of Trimeresurus cyanolabris sp. nov. in preservative (ZMMU Re-17679, adult male) from Nui Chua NP, Ninh Thuan Province, Vietnam. A dorsolateral and B ventral view of body; C head in dorsal, D in ventral, E in lateral right, and F in lateral left aspects. Scale bar equals 1 cm. Photographs by S.S. Idiiatullina." figureDoi="http://doi.org/10.5281/zenodo.12681174" httpUri="https://zenodo.org/record/12681174/files/figure.png" pageId="9" pageNumber="384">Fig. 3D</figureCitation>
|
||
). One large pair of enlarged internasals, separated by three small scales. One supraocular, large (
|
||
<figureCitation id="DFBF1493CA0EFFA0FF20DB9BD16CFD4B" box="[221,308,726,752]" captionStart="FIGURE 3" captionStartId="7.[152,255,1921,1945]" captionTargetBox="[165,1423,309,1897]" captionTargetId="figure-95@7.[164,1423,309,1897]" captionTargetPageId="7" captionText="FIGURE 3. The holotype of Trimeresurus cyanolabris sp. nov. in preservative (ZMMU Re-17679, adult male) from Nui Chua NP, Ninh Thuan Province, Vietnam. A dorsolateral and B ventral view of body; C head in dorsal, D in ventral, E in lateral right, and F in lateral left aspects. Scale bar equals 1 cm. Photographs by S.S. Idiiatullina." figureDoi="http://doi.org/10.5281/zenodo.12681174" httpUri="https://zenodo.org/record/12681174/files/figure.png" pageId="9" pageNumber="384">Fig. 3C</figureCitation>
|
||
). Scales on snout and in the interorbital region smooth, irregular, subimbricate; temporal and occipital scales feebly keeled (
|
||
<figureCitation id="DFBF1493CA0EFFA0FE7BDBB7D190FCAF" box="[390,456,762,788]" captionStart="FIGURE 3" captionStartId="7.[152,255,1921,1945]" captionTargetBox="[165,1423,309,1897]" captionTargetId="figure-95@7.[164,1423,309,1897]" captionTargetPageId="7" captionText="FIGURE 3. The holotype of Trimeresurus cyanolabris sp. nov. in preservative (ZMMU Re-17679, adult male) from Nui Chua NP, Ninh Thuan Province, Vietnam. A dorsolateral and B ventral view of body; C head in dorsal, D in ventral, E in lateral right, and F in lateral left aspects. Scale bar equals 1 cm. Photographs by S.S. Idiiatullina." figureDoi="http://doi.org/10.5281/zenodo.12681174" httpUri="https://zenodo.org/record/12681174/files/figure.png" pageId="9" pageNumber="384">Fig. 3</figureCitation>
|
||
E-F). Dorsal scales in 21–21–15 rows, moderately keeled, except the first row, which is smooth (
|
||
<figureCitation id="DFBF1493CA0EFFA0FF04DA53D10CFC83" box="[249,340,798,824]" captionStart="FIGURE 3" captionStartId="7.[152,255,1921,1945]" captionTargetBox="[165,1423,309,1897]" captionTargetId="figure-95@7.[164,1423,309,1897]" captionTargetPageId="7" captionText="FIGURE 3. The holotype of Trimeresurus cyanolabris sp. nov. in preservative (ZMMU Re-17679, adult male) from Nui Chua NP, Ninh Thuan Province, Vietnam. A dorsolateral and B ventral view of body; C head in dorsal, D in ventral, E in lateral right, and F in lateral left aspects. Scale bar equals 1 cm. Photographs by S.S. Idiiatullina." figureDoi="http://doi.org/10.5281/zenodo.12681174" httpUri="https://zenodo.org/record/12681174/files/figure.png" pageId="9" pageNumber="384">Fig. 3A</figureCitation>
|
||
). One preventral and 175 ventrals. Anal plate single; 72 subcaudals, all divided. Hemipenes long, papillose, and deeply forked with small, soft basal spines, reaching the 8
|
||
<superScript id="B0F1A55ECA0EFFA0FC3EDA0FD388FCEB" attach="left" box="[963,976,834,848]" fontSize="6" pageId="9" pageNumber="384">th</superScript>
|
||
subcaudal (
|
||
<figureCitation id="DFBF1493CA0EFFA0FBABDA0FD4F6FCE7" box="[1110,1198,834,860]" captionStart="FIGURE 3" captionStartId="7.[152,255,1921,1945]" captionTargetBox="[165,1423,309,1897]" captionTargetId="figure-95@7.[164,1423,309,1897]" captionTargetPageId="7" captionText="FIGURE 3. The holotype of Trimeresurus cyanolabris sp. nov. in preservative (ZMMU Re-17679, adult male) from Nui Chua NP, Ninh Thuan Province, Vietnam. A dorsolateral and B ventral view of body; C head in dorsal, D in ventral, E in lateral right, and F in lateral left aspects. Scale bar equals 1 cm. Photographs by S.S. Idiiatullina." figureDoi="http://doi.org/10.5281/zenodo.12681174" httpUri="https://zenodo.org/record/12681174/files/figure.png" pageId="9" pageNumber="384">Fig. 3B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="473B0816CA0EFFA0FF3ADA2BD3A7FB8F" blockId="9.[151,1437,150,2013]" pageId="9" pageNumber="384">
|
||
<emphasis id="75F0D404CA0EFFA0FF3ADA2BD110FC3B" bold="true" box="[199,328,870,896]" pageId="9" pageNumber="384">Coloration</emphasis>
|
||
. In life (
|
||
<figureCitation id="DFBF1493CA0EFFA0FE54DA2BD25BFC3B" box="[425,515,870,896]" captionStart="FIGURE 4" captionStartId="8.[152,255,1832,1856]" captionTargetBox="[193,1395,181,1809]" captionTargetId="figure-25@8.[192,1395,181,1809]" captionTargetPageId="8" captionText="FIGURE 4. Type specimens of Trimeresurus cyanolabris sp. nov. in life from Nui Chua NP, Ninh Thuan Province, Vietnam: A holotype ZMMU Re-17679, adult male; B paratype ZMMU Re-17681, adult female. Photographs by A.M. Bragin (A) and N.A. Poyarkov (B)." figureDoi="http://doi.org/10.5281/zenodo.12681176" httpUri="https://zenodo.org/record/12681176/files/figure.png" pageId="9" pageNumber="384">Fig. 4A</figureCitation>
|
||
), the body is uniformly bright grass-green above and on the upper part of the sides and gradually turns yellow on the lower sides, without a ventrolateral stripe. Tail dull brick red, dorsally extending as far as vent, with a faint margin on lateral side of tail. Venter light greenish-yellow, paler in its anterior part, and light green in its posterior part. The dorsal surface of the head and the temporal region are green without a postocular streak; supralabial, infralabial, chin and throat regions are blue. Iris bright yellow. In preservative (
|
||
<figureCitation id="DFBF1493CA0EFFA0FAE4DABBD538FBAB" box="[1305,1376,1014,1040]" captionStart="FIGURE 3" captionStartId="7.[152,255,1921,1945]" captionTargetBox="[165,1423,309,1897]" captionTargetId="figure-95@7.[164,1423,309,1897]" captionTargetPageId="7" captionText="FIGURE 3. The holotype of Trimeresurus cyanolabris sp. nov. in preservative (ZMMU Re-17679, adult male) from Nui Chua NP, Ninh Thuan Province, Vietnam. A dorsolateral and B ventral view of body; C head in dorsal, D in ventral, E in lateral right, and F in lateral left aspects. Scale bar equals 1 cm. Photographs by S.S. Idiiatullina." figureDoi="http://doi.org/10.5281/zenodo.12681174" httpUri="https://zenodo.org/record/12681174/files/figure.png" pageId="9" pageNumber="384">Fig. 3</figureCitation>
|
||
), the background dorsal color faded to a light gray; the ventral color became white.
|
||
</paragraph>
|
||
<paragraph id="473B0816CA0EFFA0FF3ADD73D47EFB1A" blockId="9.[151,1437,150,2013]" pageId="9" pageNumber="384">
|
||
<emphasis id="75F0D404CA0EFFA0FF3ADD73D160FBE3" bold="true" box="[199,312,1086,1112]" pageId="9" pageNumber="384">Variation</emphasis>
|
||
(for detailed information, see
|
||
<tableCitation id="0A063DADCA0EFFA0FD77DD73D287FBE2" box="[650,735,1086,1113]" captionStart="TABLE 5" captionStartId="22.[152,245,151,175]" captionTargetPageId="22" captionText="TABLE 5. Morphological data on the type series of Trimeresurus cyanolabris sp. nov. Notes: for abbreviations see Materials and methods section; N/a: Data not available." httpUri="http://table.plazi.org/id/13FB589ECA11FFBFFF65D9DAD2B0FF68" pageId="9" pageNumber="384" tableUuid="13FB589ECA11FFBFFF65D9DAD2B0FF68">Table 5</tableCitation>
|
||
). The longest known specimen is
|
||
<quantity id="807CA5F3CA0EFFA0FBA7DD73D4EEFBE2" box="[1114,1206,1086,1113]" metricMagnitude="-1" metricUnit="m" metricValue="7.43" pageId="9" pageNumber="384" unit="mm" value="743.0">743 mm</quantity>
|
||
long (SVL
|
||
<quantity id="807CA5F3CA0EFFA0FAC4DD73D5C0FBE2" box="[1337,1432,1086,1113]" metricMagnitude="-1" metricUnit="m" metricValue="6.38" pageId="9" pageNumber="384" unit="mm" value="638.0">638 mm</quantity>
|
||
, TaL
|
||
<quantity id="807CA5F3CA0EFFA0FF31DD2FD176FBC6" box="[204,302,1122,1149]" metricMagnitude="-1" metricUnit="m" metricValue="1.05" pageId="9" pageNumber="384" unit="mm" value="105.0">105 mm</quantity>
|
||
; female, DTU 635); the longest known male is
|
||
<quantity id="807CA5F3CA0EFFA0FCA9DD2FD3EBFBC6" box="[852,947,1122,1149]" metricMagnitude="-1" metricUnit="m" metricValue="6.07" pageId="9" pageNumber="384" unit="mm" value="607.0">607 mm</quantity>
|
||
long (SVL
|
||
<quantity id="807CA5F3CA0EFFA0FBC1DD2FD4C5FBC6" box="[1084,1181,1122,1149]" metricMagnitude="-1" metricUnit="m" metricValue="4.779999999999999" pageId="9" pageNumber="384" unit="mm" value="478.0">478 mm</quantity>
|
||
, TaL
|
||
<quantity id="807CA5F3CA0EFFA0FB22DD2FD519FBC6" box="[1247,1345,1122,1149]" metricMagnitude="-1" metricUnit="m" metricValue="1.29" pageId="9" pageNumber="384" unit="mm" value="129.0">129 mm</quantity>
|
||
; ZFMK 94670). Ratio TaL/TL:
|
||
<geoCoordinate id="22B06ED1CA0EFFA0FE60DDCBD184FB1B" box="[413,476,1158,1184]" degrees="0.129" direction="north" orientation="latitude" pageId="9" pageNumber="384" precision="55" value="0.129">0.129</geoCoordinate>
|
||
<geoCoordinate id="22B06ED1CA0EFFA0FE21DDCBD271FB1B" box="[476,553,1158,1184]" degrees="0.213" direction="west" orientation="longitude" pageId="9" pageNumber="384" precision="55" value="-0.213">–0.213</geoCoordinate>
|
||
(males:
|
||
<geoCoordinate id="22B06ED1CA0EFFA0FD77DDCBD291FB1B" box="[650,713,1158,1184]" degrees="0.181" direction="north" orientation="latitude" pageId="9" pageNumber="384" precision="55" value="0.181">0.181</geoCoordinate>
|
||
<geoCoordinate id="22B06ED1CA0EFFA0FD34DDCBD341FB1B" box="[713,793,1158,1184]" degrees="0.213" direction="west" orientation="longitude" pageId="9" pageNumber="384" precision="55" value="-0.213">–0.213</geoCoordinate>
|
||
, females:
|
||
<geoCoordinate id="22B06ED1CA0EFFA0FC74DDCBD392FB1A" box="[905,970,1158,1185]" degrees="0.129" direction="north" orientation="latitude" pageId="9" pageNumber="384" precision="55" value="0.129">0.129</geoCoordinate>
|
||
<geoCoordinate id="22B06ED1CA0EFFA0FC36DDCBD442FB1A" box="[971,1050,1158,1185]" degrees="0.154" direction="west" orientation="longitude" pageId="9" pageNumber="384" precision="55" value="-0.154">–0.154</geoCoordinate>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="473B0816CA0EFFA0FF3ADDE7D128FAB7" blockId="9.[151,1437,150,2013]" pageId="9" pageNumber="384">
|
||
<emphasis id="75F0D404CA0EFFA0FF3ADDE7D12AFB7F" box="[199,370,1194,1220]" italics="true" pageId="9" pageNumber="384">Body scalation</emphasis>
|
||
. 21 (rarely
|
||
<quantity id="807CA5F3CA0EFFA0FE00DDE7D261FB7F" box="[509,569,1194,1220]" metricMagnitude="-1" metricUnit="m" metricValue="5.842" pageId="9" pageNumber="384" unit="in" value="23.0">23 in</quantity>
|
||
the ZMMU Re-
|
||
<specimenCount id="5182C39FCA0EFFA0FD07DDE7D3CCFB7E" box="[762,916,1194,1221]" count="176" pageId="9" pageNumber="384" type="generic" typeStatus="paratype">176 Paratype</specimenCount>
|
||
)–21–15 DSR; 166–178
|
||
<collectingRegion id="8540C6F4CA0EFFA0FB57DDE6D4BEFB7E" box="[1194,1254,1195,1221]" country="Austria" name="Wien" pageId="9" pageNumber="384">VEN</collectingRegion>
|
||
, without sexual dimorphism; 52–75 SC (males: 71–75, females: 52–62); total number of
|
||
<collectingRegion id="8540C6F4CA0EFFA0FC11DD83D472FB53" box="[1004,1066,1230,1256]" country="Austria" name="Wien" pageId="9" pageNumber="384">VEN</collectingRegion>
|
||
+SC: 223–247 (males: 241–247, females: 223–237).
|
||
</paragraph>
|
||
<paragraph id="473B0816CA0EFFA0FF3ADC5BD12CFAEF" blockId="9.[151,1437,150,2013]" pageId="9" pageNumber="384">
|
||
<emphasis id="75F0D404CA0EFFA0FF3ADC5BD12AFA8B" box="[199,370,1302,1328]" italics="true" pageId="9" pageNumber="384">Head scalation</emphasis>
|
||
. InSep: none, internasals in contact in all known specimens; Cep: 9–10; SL: 10 (exceptionally 9 or 11); IL: 10–13.
|
||
</paragraph>
|
||
<paragraph id="473B0816CA0EFFA0FF3ADC12D460FA26" blockId="9.[151,1437,150,2013]" pageId="9" pageNumber="384">
|
||
<emphasis id="75F0D404CA0EFFA0FF3ADC12D241FAC3" box="[199,537,1374,1400]" italics="true" pageId="9" pageNumber="384">Main characters of the pattern</emphasis>
|
||
. The postocular streak is always absent in both sexes; ventrolateral stripe present, white, faint, and thin in males; absent in females. Eyes bright yellow in both sexes.
|
||
</paragraph>
|
||
<paragraph id="473B0816CA0EFFA0FF3ADCEBD43AF9EB" blockId="9.[151,1437,150,2013]" pageId="9" pageNumber="384">
|
||
<emphasis id="75F0D404CA0EFFA0FF3ADCEBD13EFA7B" bold="true" box="[199,358,1446,1472]" pageId="9" pageNumber="384">Comparisons</emphasis>
|
||
. The new species is phylogenetically placed within the subgenus
|
||
<taxonomicName id="80847395CA0EFFA0FB91DCEAD558FA7B" box="[1132,1280,1447,1472]" pageId="9" pageNumber="384" rank="subGenus" subGenus="Trimeresurus">
|
||
<emphasis id="75F0D404CA0EFFA0FB91DCEAD558FA7B" box="[1132,1280,1447,1472]" italics="true" pageId="9" pageNumber="384">Trimeresurus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="231575E7CA0EFFA0FAEEDCEBD172FA5F" author="Malhotra, A. & Thorpe, R. S." pageId="9" pageNumber="384" pagination="83 - 100" refId="ref18749" refString="Malhotra, A. & Thorpe, R. S. (2004) A phylogeny of four mitochondrial gene regions suggests a revised taxonomy for Asian pitvipers (Trimeresurus and Ovophis). Molecular Phylogenetics and Evolution, 32, 83 - 100. https: // doi. org / 10.1016 / j. ympev. 2004.02.008" type="journal article" year="2004">Malhotra & Thorpe 2004</bibRefCitation>
|
||
[as
|
||
<taxonomicName id="80847395CA0EFFA0FEA3DC87D1AFFA5F" authorityName="Cope" authorityYear="1859" box="[350,503,1482,1508]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="384" phylum="Chordata" rank="genus">
|
||
<emphasis id="75F0D404CA0EFFA0FEA3DC87D1AFFA5F" box="[350,503,1482,1508]" italics="true" pageId="9" pageNumber="384">Cryptelytrops</emphasis>
|
||
</taxonomicName>
|
||
];
|
||
<bibRefCitation id="231575E7CA0EFFA0FDEFDC87D287FA5F" author="David, P. & Vogel, G. & Dubois, A." box="[530,735,1482,1509]" pageId="9" pageNumber="384" pagination="1 - 51" refId="ref16865" refString="David, P., Vogel, G. & Dubois, A. (2011) On the need to follow rigorously the rules of the code for the subsequent designation of a nucleospecies (type species) for a nominal genus which lacked one: the case of the nominal genus Trimeresurus Lacepede, 1804 (Reptilia: Squamata: Viperidae). Zootaxa, 2992 (1), 1 - 51. https: // doi. org / 10.11646 / zootaxa. 2992.1.1" type="journal article" year="2011">
|
||
David
|
||
<emphasis id="75F0D404CA0EFFA0FD9CDC86D2CDFA5F" box="[609,661,1482,1508]" italics="true" pageId="9" pageNumber="384">et al</emphasis>
|
||
. 2011
|
||
</bibRefCitation>
|
||
) and is morphologically most similar to other green pitviper species from the
|
||
<taxonomicName id="80847395CA0EFFA0FE99DCA2D1BCF9B3" authorityName="Kramer" authorityYear="1977" box="[356,484,1519,1544]" class="Squamata" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" pageId="9" pageNumber="384" phylum="Chordata" rank="species" species="macrops">
|
||
<emphasis id="75F0D404CA0EFFA0FE99DCA2D1BCF9B3" box="[356,484,1519,1544]" italics="true" pageId="9" pageNumber="384">T. macrops</emphasis>
|
||
</taxonomicName>
|
||
complex, including
|
||
<taxonomicName id="80847395CA0EFFA0FD24DCA2D31DF9B3" box="[729,837,1518,1544]" class="Squamata" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" pageId="9" pageNumber="384" phylum="Chordata" rank="species" species="rubeus">
|
||
<emphasis id="75F0D404CA0EFFA0FD24DCA2D31DF9B3" box="[729,837,1518,1544]" italics="true" pageId="9" pageNumber="384">T. rubeus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="80847395CA0EFFA0FCA5DCA2D470F9B3" baseAuthorityName="Malhotra, Thorpe, Mrinalini & Stuart" baseAuthorityYear="2011" box="[856,1064,1518,1544]" class="Squamata" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" pageId="9" pageNumber="384" phylum="Chordata" rank="species" species="cardamomensis">
|
||
<emphasis id="75F0D404CA0EFFA0FCA5DCA2D470F9B3" box="[856,1064,1518,1544]" italics="true" pageId="9" pageNumber="384">T. cardamomensis</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="80847395CA0EFFA0FB92DCA2D4B6F9B3" authorityName="Kramer" authorityYear="1977" box="[1135,1262,1519,1544]" class="Squamata" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" pageId="9" pageNumber="384" phylum="Chordata" rank="species" species="macrops">
|
||
<emphasis id="75F0D404CA0EFFA0FB92DCA2D4B6F9B3" box="[1135,1262,1519,1544]" italics="true" pageId="9" pageNumber="384">T. macrops</emphasis>
|
||
</taxonomicName>
|
||
; therefore, the comparisons with these three species appear to be the most pertinent. The main diagnostic characters separating
|
||
<taxonomicName id="80847395CA0EFFA0FF6ADF7AD1EFF9EB" authorityName="Idiiatullina & Nguyen & Bragin & Pawangkhanant & Le & Vogel & David & Poyarkov" authorityYear="2024" box="[151,439,1590,1616]" class="Squamata" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" pageId="9" pageNumber="384" phylum="Chordata" rank="species" species="cyanolabris" status="sp. nov.">
|
||
<emphasis id="75F0D404CA0EFFA0FF6ADF7AD1EFF9EB" box="[151,439,1590,1616]" italics="true" pageId="9" pageNumber="384">Trimeresurus cyanolabris</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="75F0D404CA0EFFA0FE43DF7BD24FF9EB" bold="true" box="[446,535,1590,1616]" pageId="9" pageNumber="384">
|
||
<taxonomicNameLabel id="6EC3697FCA0EFFA0FE43DF7BD24FF9EB" box="[446,535,1590,1616]" pageId="9" pageNumber="384" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
from these three species are summarized in
|
||
<tableCitation id="0A063DADCA0EFFA0FBF5DF7BD406F9EB" box="[1032,1118,1590,1616]" captionStart="TABLE 6" captionStartId="23.[152,245,1242,1266]" captionTargetPageId="23" captionText="TABLE 6. Comparison of morphological characters of Trimeresurus cyanolabris sp. nov. with T. cardamomensis, T. macrops, and T. rubeus. Remarks: Diagnostic differences from the new species are marked in bold. (?): requires verification." httpUri="http://table.plazi.org/id/13FB589ECA10FFBEFF65DD97D142FA81" pageId="9" pageNumber="384" tableUuid="13FB589ECA10FFBEFF65DD97D142FA81">Table 6</tableCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="473B0816CA0EFFA3FF3ADF16D4C6FF42" blockId="9.[151,1437,150,2013]" lastBlockId="10.[151,1437,151,2013]" lastPageId="10" lastPageNumber="385" pageId="9" pageNumber="384">
|
||
<taxonomicName id="80847395CA0EFFA0FF3ADF16D1BFF9CF" authorityName="Idiiatullina & Nguyen & Bragin & Pawangkhanant & Le & Vogel & David & Poyarkov" authorityYear="2024" box="[199,487,1626,1652]" class="Squamata" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" pageId="9" pageNumber="384" phylum="Chordata" rank="species" species="cyanolabris" status="sp. nov.">
|
||
<emphasis id="75F0D404CA0EFFA0FF3ADF16D1BFF9CF" box="[199,487,1626,1652]" italics="true" pageId="9" pageNumber="384">Trimeresurus cyanolabris</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="75F0D404CA0EFFA0FE12DF17D212F9CF" bold="true" box="[495,586,1626,1652]" pageId="9" pageNumber="384">
|
||
<taxonomicNameLabel id="6EC3697FCA0EFFA0FE12DF17D212F9CF" box="[495,586,1626,1652]" pageId="9" pageNumber="384" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is distinguished from
|
||
<taxonomicName id="80847395CA0EFFA0FCBADF16D3E8F9CF" box="[839,944,1626,1652]" class="Squamata" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" pageId="9" pageNumber="384" phylum="Chordata" rank="species" species="rubeus">
|
||
<emphasis id="75F0D404CA0EFFA0FCBADF16D3E8F9CF" box="[839,944,1626,1652]" italics="true" pageId="9" pageNumber="384">T. rubeus</emphasis>
|
||
</taxonomicName>
|
||
(restricted to southern
|
||
<collectingCountry id="3F934886CA0EFFA0FB4BDF17D54DF9CF" box="[1206,1301,1626,1652]" name="Vietnam" pageId="9" pageNumber="384">Vietnam</collectingCountry>
|
||
and eastern
|
||
<collectingCountry id="3F934886CA0EFFA0FF6ADF33D154F923" box="[151,268,1662,1688]" name="Cambodia" pageId="9" pageNumber="384">Cambodia</collectingCountry>
|
||
) by having: (1) lower max SVL in males (
|
||
<quantity id="807CA5F3CA0EFFA0FD20DF33D362F922" box="[733,826,1662,1689]" metricMagnitude="-1" metricUnit="m" metricValue="4.779999999999999" pageId="9" pageNumber="384" unit="mm" value="478.0">478 mm</quantity>
|
||
vs.
|
||
<quantity id="807CA5F3CA0EFFA0FC9BDF33D39DF923" box="[870,965,1662,1689]" metricMagnitude="-1" metricUnit="m" metricValue="5.86" pageId="9" pageNumber="384" unit="mm" value="586.0">586 mm</quantity>
|
||
), but higher max SVL in females (
|
||
<quantity id="807CA5F3CA0EFFA0FAC2DF33D5C5F922" box="[1343,1437,1662,1689]" metricMagnitude="-1" metricUnit="m" metricValue="6.38" pageId="9" pageNumber="384" unit="mm" value="638.0">638 mm</quantity>
|
||
vs.
|
||
<quantity id="807CA5F3CA0EFFA0FF42DFEFD179F907" box="[191,289,1698,1725]" metricMagnitude="-1" metricUnit="m" metricValue="5.029999999999999" pageId="9" pageNumber="384" unit="mm" value="503.0">503 mm</quantity>
|
||
); (2) slightly lower ratio TaL/TL in both sexes (0.18–0.21 [X = 0.19] in males, 0.13–0.15 [X = 0.14] in females vs. 0.21–0.22 [X = 0.22] in males, 0.15–0.19 [X = 0.17] in females); (3) slightly higher ratio HW/HL in both sexes (0.55–0.68 [X = 0.61] in males, 0.65–0.70 [X = 0.66] in females vs. 0.53–0.60 [X = 0.56] in males, 0.46–0.72 [X = 0.61] in females); (4) slightly higher ratio ED/EN in males (0.67–0.78 [X = 0.74] vs. 0.66–0.74 [X = 0.70], but lower ratio ED/EN in females (0.62–0.88 [X = 0.71] vs. 0.79–0.95 [X = 0.85]; (5) slightly higher number VEN in both sexes (166–175 [X = 171.0] in males, 169–178 [X = 174.1] in females vs. 164–171 [X = 167.5] in males, 158–163 [X = 159.8] in females); (6) slightly lower number SC in males (71–75 [͞x =72.7] in males, 52–62 [X = 57.7] in females vs. 78–80 [X = 79.0] in males, 52–69 [X = 61.6] in females; (7) slightly lower total number of VEN+SC in females (223–237 [X = 231.8] vs. 214–228 [X = 221.4]; (8) without postocular white stripe in males (vs. present); (9) ventrolateral body stripe present, faint, poorly contrasted in males (vs. well developed, highly contrasted with reddish and white); (10) ventral surface of body greenish-blue (vs. greenish-yellow); (11) eye color bright yellow in both sexes (vs. bright red, blood-red or deep reddish–orange in both sexes).
|
||
</paragraph>
|
||
<paragraph id="473B0816CA0DFFA3FF3AD84ED300FD12" blockId="10.[151,1437,151,2013]" pageId="10" pageNumber="385">
|
||
<taxonomicName id="80847395CA0DFFA3FF3AD84ED1BDFEA6" authorityName="Idiiatullina & Nguyen & Bragin & Pawangkhanant & Le & Vogel & David & Poyarkov" authorityYear="2024" box="[199,485,259,285]" class="Squamata" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" pageId="10" pageNumber="385" phylum="Chordata" rank="species" species="cyanolabris" status="sp. nov.">
|
||
<emphasis id="75F0D404CA0DFFA3FF3AD84ED1BDFEA6" box="[199,485,259,285]" italics="true" pageId="10" pageNumber="385">Trimeresurus cyanolabris</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="75F0D404CA0DFFA3FE16D84ED21CFEA6" bold="true" box="[491,580,259,285]" pageId="10" pageNumber="385">
|
||
<taxonomicNameLabel id="6EC3697FCA0DFFA3FE16D84ED21CFEA6" box="[491,580,259,285]" pageId="10" pageNumber="385" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
further differs from
|
||
<taxonomicName id="80847395CA0DFFA3FCDAD84ED3A8FEA6" baseAuthorityName="Malhotra, Thorpe, Mrinalini & Stuart" baseAuthorityYear="2011" box="[807,1008,259,285]" class="Squamata" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" pageId="10" pageNumber="385" phylum="Chordata" rank="species" species="cardamomensis">
|
||
<emphasis id="75F0D404CA0DFFA3FCDAD84ED3A8FEA6" box="[807,1008,259,285]" italics="true" pageId="10" pageNumber="385">T. cardamomensis</emphasis>
|
||
</taxonomicName>
|
||
(restricted to southwestern
|
||
<collectingCountry id="3F934886CA0DFFA3FADFD84ED5C0FEA6" box="[1314,1432,259,285]" name="Cambodia" pageId="10" pageNumber="385">Cambodia</collectingCountry>
|
||
, southeastern
|
||
<collectingCountry id="3F934886CA0DFFA3FED7D86AD1C8FEFA" box="[298,400,295,321]" name="Thailand" pageId="10" pageNumber="385">Thailand</collectingCountry>
|
||
, and the southermost part of
|
||
<collectingCountry id="3F934886CA0DFFA3FD2AD86AD360FEFA" box="[727,824,295,321]" name="Vietnam" pageId="10" pageNumber="385">Vietnam</collectingCountry>
|
||
) by having: (1) lower max SVL in males (
|
||
<quantity id="807CA5F3CA0DFFA3FAEBD86AD52DFEFA" box="[1302,1397,295,321]" metricMagnitude="-1" metricUnit="m" metricValue="4.779999999999999" pageId="10" pageNumber="385" unit="mm" value="478.0">478 mm</quantity>
|
||
vs.
|
||
<quantity id="807CA5F3CA0DFFA3FF6AD807D0A3FEDE" box="[151,251,330,357]" metricMagnitude="-1" metricUnit="m" metricValue="5.35" pageId="10" pageNumber="385" unit="mm" value="535.0">535 mm</quantity>
|
||
), but higher max SVL in females (
|
||
<quantity id="807CA5F3CA0DFFA3FD73D806D2B7FEDE" box="[654,751,331,357]" metricMagnitude="-1" metricUnit="m" metricValue="6.38" pageId="10" pageNumber="385" unit="mm" value="638.0">638 mm</quantity>
|
||
vs.
|
||
<quantity id="807CA5F3CA0DFFA3FCDCD807D3DDFEDE" box="[801,901,330,357]" metricMagnitude="-1" metricUnit="m" metricValue="5.36" pageId="10" pageNumber="385" unit="mm" value="536.0">536 mm</quantity>
|
||
); (2) slightly lower ratio ED/EN in both sexes (0.67–0.78 [X = 0.74] in males, 0.62–0.88 [X = 0.71] in females vs. 0.66–0.91 [X = 0.78] in males, 0.51–0.97 [X = 0.74] in females); (3) slightly higher number of SC in males (71–75 [X = 72.7] vs. 60–75 [X = 67.0]); (4) higher total number of VEN+SC in males (241–247 [X = 243.7] vs. 226–243 [X = 234.0]); (5) postocular stripe absent in males (vs. present, white or blue); (6) ventrolateral body stripe present, faint, poorly contrasted in male (vs. well developed, highly contrasted, white); (7) ventral surface of body greenish-blue (vs. greenish-yellow); (8) head shape more elongate-oval (vs. more triangular); (9) small size scales on head behind internasal scales (vs. lager). Furthermore,
|
||
<taxonomicName id="80847395CA0DFFA3FECEDB0AD18DFDDA" authorityName="Idiiatullina & Nguyen & Bragin & Pawangkhanant & Le & Vogel & David & Poyarkov" authorityYear="2024" box="[307,469,583,609]" class="Squamata" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" pageId="10" pageNumber="385" phylum="Chordata" rank="species" species="cyanolabris" status="sp. nov.">
|
||
<emphasis id="75F0D404CA0DFFA3FECEDB0AD18DFDDA" box="[307,469,583,609]" italics="true" pageId="10" pageNumber="385">T. cyanolabris</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="75F0D404CA0DFFA3FE23DB0AD262FDDA" bold="true" box="[478,570,583,609]" pageId="10" pageNumber="385">
|
||
<taxonomicNameLabel id="6EC3697FCA0DFFA3FE23DB0AD262FDDA" box="[478,570,583,609]" pageId="10" pageNumber="385" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is widely separated from
|
||
<taxonomicName id="80847395CA0DFFA3FC9ADB0AD46CFDDA" baseAuthorityName="Malhotra, Thorpe, Mrinalini & Stuart" baseAuthorityYear="2011" box="[871,1076,583,609]" class="Squamata" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" pageId="10" pageNumber="385" phylum="Chordata" rank="species" species="cardamomensis">
|
||
<emphasis id="75F0D404CA0DFFA3FC9ADB0AD46CFDDA" box="[871,1076,583,609]" italics="true" pageId="10" pageNumber="385">T. cardamomensis</emphasis>
|
||
</taxonomicName>
|
||
on a geographical basis, as the latter species inhabits only southeastern
|
||
<collectingCountry id="3F934886CA0DFFA3FDA8DB26D2E3FD3E" box="[597,699,619,645]" name="Thailand" pageId="10" pageNumber="385">Thailand</collectingCountry>
|
||
, southerwestern
|
||
<collectingCountry id="3F934886CA0DFFA3FC88DB26D3B3FD3E" box="[885,1003,619,645]" name="Cambodia" pageId="10" pageNumber="385">Cambodia</collectingCountry>
|
||
, and southernmost
|
||
<collectingCountry id="3F934886CA0DFFA3FB3DDB26D547FD3E" box="[1216,1311,619,645]" name="Vietnam" pageId="10" pageNumber="385">Vietnam</collectingCountry>
|
||
(Phu Quoc Isl.). Moreover,
|
||
<taxonomicName id="80847395CA0DFFA3FEB0DBC2D1ECFD12" box="[333,436,655,681]" class="Squamata" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" pageId="10" pageNumber="385" phylum="Chordata" rank="species" species="rubeus">
|
||
<emphasis id="75F0D404CA0DFFA3FEB0DBC2D1ECFD12" box="[333,436,655,681]" italics="true" pageId="10" pageNumber="385">T. rubeus</emphasis>
|
||
</taxonomicName>
|
||
separates the ranges of both species..
|
||
</paragraph>
|
||
<paragraph id="473B0816CA0DFFA3FF3ADBFED105FBAA" blockId="10.[151,1437,151,2013]" pageId="10" pageNumber="385">
|
||
In particular,
|
||
<taxonomicName id="80847395CA0DFFA3FEA6DBFED221FD76" authorityName="Idiiatullina & Nguyen & Bragin & Pawangkhanant & Le & Vogel & David & Poyarkov" authorityYear="2024" box="[347,633,691,717]" class="Squamata" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" pageId="10" pageNumber="385" phylum="Chordata" rank="species" species="cyanolabris" status="sp. nov.">
|
||
<emphasis id="75F0D404CA0DFFA3FEA6DBFED221FD76" box="[347,633,691,717]" italics="true" pageId="10" pageNumber="385">Trimeresurus cyanolabris</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="75F0D404CA0DFFA3FD82DBFED28FFD76" bold="true" box="[639,727,691,717]" pageId="10" pageNumber="385">
|
||
<taxonomicNameLabel id="6EC3697FCA0DFFA3FD82DBFED28FFD76" box="[639,727,691,717]" pageId="10" pageNumber="385" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is distinguished from
|
||
<taxonomicName id="80847395CA0DFFA3FC31DBFED41DFD77" authorityName="Kramer" authorityYear="1977" box="[972,1093,691,716]" class="Squamata" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" pageId="10" pageNumber="385" phylum="Chordata" rank="species" species="macrops">
|
||
<emphasis id="75F0D404CA0DFFA3FC31DBFED41DFD77" box="[972,1093,691,716]" italics="true" pageId="10" pageNumber="385">T. macrops</emphasis>
|
||
</taxonomicName>
|
||
by having: (1) lower max SVL in males (
|
||
<quantity id="807CA5F3CA0DFFA3FEFEDB9AD138FD4A" box="[259,352,727,753]" metricMagnitude="-1" metricUnit="m" metricValue="4.779999999999999" pageId="10" pageNumber="385" unit="mm" value="478.0">478 mm</quantity>
|
||
vs.
|
||
<quantity id="807CA5F3CA0DFFA3FE71DB9BD1B5FD4A" box="[396,493,726,753]" metricMagnitude="-1" metricUnit="m" metricValue="5.57" pageId="10" pageNumber="385" unit="mm" value="557.0">557 mm</quantity>
|
||
), but higher max SVL in females (
|
||
<quantity id="807CA5F3CA0DFFA3FC95DB9AD39EFD4A" box="[872,966,727,753]" metricMagnitude="-1" metricUnit="m" metricValue="6.38" pageId="10" pageNumber="385" unit="mm" value="638.0">638 mm</quantity>
|
||
vs.
|
||
<quantity id="807CA5F3CA0DFFA3FC0FDB9AD40AFD4A" box="[1010,1106,727,753]" metricMagnitude="-1" metricUnit="m" metricValue="6.29" pageId="10" pageNumber="385" unit="mm" value="629.0">629 mm</quantity>
|
||
); (2) slightly lower ratio HW/ HL in males (0.55–0.68 [X = 0.61] vs. 0.56–0.82 [X = 0.68]); (3) lower ratio ED/EN in both sexes (0.67–0.78 [X = 0.74] in males, 0.62–0.88 [X = 0.71] in females vs. 0.62–1.12 [X = 0.89] in males, 0.64–0.97 [X = 0.83] in females); (4) slightly higher number
|
||
<collectingRegion id="8540C6F4CA0DFFA3FE38DA0ED1A7FCE6" box="[453,511,835,861]" country="Austria" name="Wien" pageId="10" pageNumber="385">VEN</collectingRegion>
|
||
in both sexes (166–175 [X = 171.0] in males, 169–178 [X = 174.1] in females vs. 158–173 [X = 165.0] in males, 159–173 [X = 166.4] in females); (5) higher total number of
|
||
<collectingRegion id="8540C6F4CA0DFFA3FB64DA2AD48CFC3A" box="[1177,1236,871,897]" country="Austria" name="Wien" pageId="10" pageNumber="385">VEN</collectingRegion>
|
||
+SC in both sexes (241–247 [X = 243.7] in males, 223–237 [X = 231.8] in females vs. 225–240 [X = 233.0] in males, 214–235 [X = 222.1] in females); (6) postocular stripe absent in males (vs. present); (7) ventrolateral body stripe present, faint, poorly contrasted in male (vs. well developed, highly contrasted); (8) ventral surface of body greenish-blue (vs. greenish-yellow).
|
||
</paragraph>
|
||
<paragraph id="473B0816CA0DFFA3FF3ADD56D39AFB52" blockId="10.[151,1437,151,2013]" pageId="10" pageNumber="385">
|
||
Among the other species in the subgenus
|
||
<taxonomicName id="80847395CA0DFFA3FD50DD56D319FB8F" box="[685,833,1051,1076]" pageId="10" pageNumber="385" rank="subGenus" subGenus="Trimeresurus">
|
||
<emphasis id="75F0D404CA0DFFA3FD50DD56D319FB8F" box="[685,833,1051,1076]" italics="true" pageId="10" pageNumber="385">Trimeresurus</emphasis>
|
||
</taxonomicName>
|
||
, the new species can be readily distinguished from:
|
||
<taxonomicName id="80847395CA0DFFA3FF6ADD72D15BFBC6" authority="Idiiatullina, Pawangkhanant, Tawan, Worranuch, Dechochai, Suwannapoom, Nguyen, Chanhome & Poyarkov" authorityName="Idiiatullina, Pawangkhanant, Tawan, Worranuch, Dechochai, Suwannapoom, Nguyen, Chanhome & Poyarkov" class="Squamata" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" pageId="10" pageNumber="385" phylum="Chordata" rank="species" species="ciliaris">
|
||
<emphasis id="75F0D404CA0DFFA3FF6ADD72D15FFBE2" box="[151,263,1087,1113]" italics="true" pageId="10" pageNumber="385">T. ciliaris</emphasis>
|
||
Idiiatullina, Pawangkhanant, Tawan, Worranuch, Dechochai, Suwannapoom, Nguyen, Chanhome & Poyarkov
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="80847395CA0DFFA3FEEFDD2ED270FBC6" authority="(Boulenger)" baseAuthorityName="Boulenger" baseAuthorityYear="1896" box="[274,552,1123,1149]" class="Squamata" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" pageId="10" pageNumber="385" phylum="Chordata" rank="species" species="fasciatus">
|
||
<emphasis id="75F0D404CA0DFFA3FEEFDD2ED1CCFBC6" box="[274,404,1123,1149]" italics="true" pageId="10" pageNumber="385">T. fasciatus</emphasis>
|
||
(Boulenger)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="80847395CA0DFFA3FDCBDD2ED340FBC6" authority="Fitzinger" authorityName="Fitzinger" box="[566,792,1123,1149]" class="Squamata" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" pageId="10" pageNumber="385" phylum="Chordata" rank="species" species="labialis">
|
||
<emphasis id="75F0D404CA0DFFA3FDCBDD2ED2F1FBC6" box="[566,681,1123,1149]" italics="true" pageId="10" pageNumber="385">T. labialis</emphasis>
|
||
Fitzinger
|
||
</taxonomicName>
|
||
<emphasis id="75F0D404CA0DFFA3FCDFDD2ED360FBC7" box="[802,824,1123,1148]" italics="true" pageId="10" pageNumber="385">in</emphasis>
|
||
Steindachner,
|
||
<taxonomicName id="80847395CA0DFFA3FC19DD2ED4B8FBC6" authority="Stoliczka" authorityName="Stoliczka" authorityYear="1870" box="[996,1248,1123,1149]" class="Squamata" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" pageId="10" pageNumber="385" phylum="Chordata" rank="species" species="mutabilis">
|
||
<emphasis id="75F0D404CA0DFFA3FC19DD2ED432FBC6" box="[996,1130,1123,1149]" italics="true" pageId="10" pageNumber="385">T. mutabilis</emphasis>
|
||
Stoliczka
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="80847395CA0DFFA3FB13DD2ED5CDFBC6" baseAuthorityName="Grismer, Ngo & Grismer" baseAuthorityYear="2008" box="[1262,1429,1123,1149]" class="Squamata" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" pageId="10" pageNumber="385" phylum="Chordata" rank="species" species="honsonensis">
|
||
<emphasis id="75F0D404CA0DFFA3FB13DD2ED5CDFBC6" box="[1262,1429,1123,1149]" italics="true" pageId="10" pageNumber="385">T. honsonensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="80847395CA0DFFA3FF6ADDCAD52CFB1A" authority="Idiiatullina, Pawangkhanant, Suwannapoom, Chanhome, Nguyen, David, Vogel & Poyarkov" authorityName="Idiiatullina, Pawangkhanant, Suwannapoom, Chanhome, Nguyen, David, Vogel & Poyarkov" box="[151,1396,1159,1185]" class="Squamata" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" pageId="10" pageNumber="385" phylum="Chordata" rank="species" species="kraensis">
|
||
<emphasis id="75F0D404CA0DFFA3FF6ADDCAD140FB1A" box="[151,280,1159,1185]" italics="true" pageId="10" pageNumber="385">T. kraensis</emphasis>
|
||
Idiiatullina, Pawangkhanant, Suwannapoom, Chanhome, Nguyen, David, Vogel & Poyarkov
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="80847395CA0DFFA3FA75DDCAD17EFB7E" authorityName="Smith" authorityYear="1943" class="Squamata" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" pageId="10" pageNumber="385" phylum="Chordata" rank="species" species="kanburiensis">
|
||
<emphasis id="75F0D404CA0DFFA3FA75DDCAD17EFB7E" italics="true" pageId="10" pageNumber="385">T. kanburiensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="80847395CA0DFFA3FECBDDE6D1FCFB7E" box="[310,420,1195,1221]" class="Squamata" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" pageId="10" pageNumber="385" phylum="Chordata" rank="species" species="kuiburi">
|
||
<emphasis id="75F0D404CA0DFFA3FECBDDE6D1FCFB7E" box="[310,420,1195,1221]" italics="true" pageId="10" pageNumber="385">T. kuiburi</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="80847395CA0DFFA3FE19DDE6D239FB7F" authorityName="Vogel" authorityYear="1991" box="[484,609,1195,1220]" class="Squamata" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" pageId="10" pageNumber="385" phylum="Chordata" rank="species" species="venustus">
|
||
<emphasis id="75F0D404CA0DFFA3FE19DDE6D239FB7F" box="[484,609,1195,1220]" italics="true" pageId="10" pageNumber="385">T. venustus</emphasis>
|
||
</taxonomicName>
|
||
by coloration and pattern (uniform green vs. reddish-brown or purple or dark olive-brown bands on emerald-green or olive-greyish background).
|
||
</paragraph>
|
||
<paragraph id="473B0816CA0DFFA3FF3ADDBED437F9CE" blockId="10.[151,1437,151,2013]" pageId="10" pageNumber="385">
|
||
<taxonomicName id="80847395CA0DFFA3FF3ADDBED1ABFAB6" authorityName="Idiiatullina & Nguyen & Bragin & Pawangkhanant & Le & Vogel & David & Poyarkov" authorityYear="2024" box="[199,499,1267,1293]" class="Squamata" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" pageId="10" pageNumber="385" phylum="Chordata" rank="species" species="cyanolabris" status="sp. nov.">
|
||
<emphasis id="75F0D404CA0DFFA3FF3ADDBED1ABFAB6" box="[199,499,1267,1293]" italics="true" pageId="10" pageNumber="385">Trimeresurus cyanolabris</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="75F0D404CA0DFFA3FDFBDDBED234FAB6" bold="true" box="[518,620,1267,1293]" pageId="10" pageNumber="385">
|
||
<taxonomicNameLabel id="6EC3697FCA0DFFA3FDFBDDBED234FAB6" box="[518,620,1267,1293]" pageId="10" pageNumber="385" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
can be distinguished from
|
||
<taxonomicName id="80847395CA0DFFA3FC24DDBED529FAB6" authority="Theobald" authorityName="Theobald" authorityYear="1868" box="[985,1393,1267,1293]" class="Squamata" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" pageId="10" pageNumber="385" phylum="Chordata" rank="species" species="andersoni">
|
||
<emphasis id="75F0D404CA0DFFA3FC24DDBED4A9FAB6" box="[985,1265,1267,1293]" italics="true" pageId="10" pageNumber="385">Trimeresurus andersoni</emphasis>
|
||
Theobald
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="80847395CA0DFFA3FA75DDBED10EFA8A" authorityName="Chan, Anuar, Sankar, Law, Law, Shivaram, Christian, Mulcahy & Malhotra" authorityYear="2023" class="Squamata" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" pageId="10" pageNumber="385" phylum="Chordata" rank="species" species="ayeyarwadyensis">
|
||
<emphasis id="75F0D404CA0DFFA3FA75DDBED10EFA8A" italics="true" pageId="10" pageNumber="385">T. ayeyarwadyensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="80847395CA0DFFA3FE94DC5AD261FA8A" authority="(Blyth)" baseAuthorityName="Blyth" baseAuthorityYear="1846" box="[361,569,1303,1329]" class="Squamata" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" pageId="10" pageNumber="385" phylum="Chordata" rank="species" species="cantori">
|
||
<emphasis id="75F0D404CA0DFFA3FE94DC5AD182FA8B" box="[361,474,1303,1328]" italics="true" pageId="10" pageNumber="385">T. cantori</emphasis>
|
||
(Blyth)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="80847395CA0DFFA3FDB4DC5AD287FA8A" box="[585,735,1303,1329]" class="Squamata" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" pageId="10" pageNumber="385" phylum="Chordata" rank="species" species="erythrurus">
|
||
<emphasis id="75F0D404CA0DFFA3FDB4DC5AD287FA8A" box="[585,735,1303,1329]" italics="true" pageId="10" pageNumber="385">T. erythrurus</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="80847395CA0DFFA3FCDBDC5AD420FA8A" authority="(Gray)" baseAuthorityName="Gray" baseAuthorityYear="1832" box="[806,1144,1303,1329]" class="Squamata" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" pageId="10" pageNumber="385" phylum="Chordata" rank="species" species="purpureomaculatus">
|
||
<emphasis id="75F0D404CA0DFFA3FCDBDC5AD479FA8A" box="[806,1057,1303,1329]" italics="true" pageId="10" pageNumber="385">T. purpureomaculatus</emphasis>
|
||
(Gray)
|
||
</taxonomicName>
|
||
by the lower number of midbody dorsal scale rows (21 vs.
|
||
<quantity id="807CA5F3CA0DFFA3FDD8DC77D2D2FAEE" box="[549,650,1338,1365]" metricMagnitude="-1" metricUnit="m" metricValue="6.096" metricValueMax="6.35" metricValueMin="5.842" pageId="10" pageNumber="385" unit="in" value="24.0" valueMax="25.0" valueMin="23.0">23–25 in</quantity>
|
||
<taxonomicName id="80847395CA0DFFA3FD6FDC76D347FAEE" authorityName="Theobald" authorityYear="1868" box="[658,799,1339,1365]" class="Squamata" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" pageId="10" pageNumber="385" phylum="Chordata" rank="species" species="andersoni">
|
||
<emphasis id="75F0D404CA0DFFA3FD6FDC76D347FAEE" box="[658,799,1339,1365]" italics="true" pageId="10" pageNumber="385">T. andersoni</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="80847395CA0DFFA3FCA5DC76D46CFAEE" authorityName="Chan, Anuar, Sankar, Law, Law, Shivaram, Christian, Mulcahy & Malhotra" authorityYear="2023" box="[856,1076,1339,1365]" class="Squamata" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" pageId="10" pageNumber="385" phylum="Chordata" rank="species" species="ayeyarwadyensis">
|
||
<emphasis id="75F0D404CA0DFFA3FCA5DC76D46CFAEE" box="[856,1076,1339,1365]" italics="true" pageId="10" pageNumber="385">T. ayeyarwadyensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<quantity id="807CA5F3CA0DFFA3FBBEDC77D4F0FAEE" box="[1091,1192,1338,1365]" metricMagnitude="-1" metricUnit="m" metricValue="6.858" metricValueMax="7.366" metricValueMin="6.35" pageId="10" pageNumber="385" unit="in" value="27.0" valueMax="29.0" valueMin="25.0">25–29 in</quantity>
|
||
<taxonomicName id="80847395CA0DFFA3FB4DDC76D546FAEF" baseAuthorityName="Blyth" baseAuthorityYear="1846" box="[1200,1310,1339,1364]" class="Squamata" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" pageId="10" pageNumber="385" phylum="Chordata" rank="species" species="cantori">
|
||
<emphasis id="75F0D404CA0DFFA3FB4DDC76D546FAEF" box="[1200,1310,1339,1364]" italics="true" pageId="10" pageNumber="385">T. cantori</emphasis>
|
||
</taxonomicName>
|
||
, 23 (rarely 24, 25) in
|
||
<taxonomicName id="80847395CA0DFFA3FEF4DC12D1C2FAC2" box="[265,410,1375,1401]" class="Squamata" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" pageId="10" pageNumber="385" phylum="Chordata" rank="species" species="erythrurus">
|
||
<emphasis id="75F0D404CA0DFFA3FEF4DC12D1C2FAC2" box="[265,410,1375,1401]" italics="true" pageId="10" pageNumber="385">T. erythrurus</emphasis>
|
||
</taxonomicName>
|
||
, 25 (rarely 27, 29) in
|
||
<taxonomicName id="80847395CA0DFFA3FD73DC12D3DBFAC2" baseAuthorityName="Gray" baseAuthorityYear="1832" box="[654,899,1375,1401]" class="Squamata" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" pageId="10" pageNumber="385" phylum="Chordata" rank="species" species="purpureomaculatus">
|
||
<emphasis id="75F0D404CA0DFFA3FD73DC12D3DBFAC2" box="[654,899,1375,1401]" italics="true" pageId="10" pageNumber="385">T. purpureomaculatus</emphasis>
|
||
</taxonomicName>
|
||
). The new species can be distinguished from:
|
||
<taxonomicName id="80847395CA0DFFA3FA75DC12D265FA26" authority="Chen, Ding, Vogel & Shi" authorityName="Chen, Ding, Vogel & Shi" authorityYear="2020" class="Squamata" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" pageId="10" pageNumber="385" phylum="Chordata" rank="species" species="caudornatus">
|
||
<emphasis id="75F0D404CA0DFFA3FA75DC12D17BFA26" italics="true" pageId="10" pageNumber="385">T. caudornatus</emphasis>
|
||
Chen, Ding, Vogel & Shi
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="80847395CA0DFFA3FDBADCCED472FA26" authority="Chandramouli, Campbell & Vogel" authorityName="Chandramouli, Campbell & Vogel" authorityYear="2020" box="[583,1066,1411,1437]" class="Squamata" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" pageId="10" pageNumber="385" phylum="Chordata" rank="species" species="davidi">
|
||
<emphasis id="75F0D404CA0DFFA3FDBADCCED2FEFA26" box="[583,678,1411,1437]" italics="true" pageId="10" pageNumber="385">T. davidi</emphasis>
|
||
Chandramouli, Campbell & Vogel
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="80847395CA0DFFA3FBCFDCCED5CFFA26" authority="Chen, Shi, Vogel & Ding" authorityName="Chen, Shi, Vogel & Ding" authorityYear="2021" box="[1074,1431,1411,1437]" class="Squamata" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" pageId="10" pageNumber="385" phylum="Chordata" rank="species" species="guoi">
|
||
<emphasis id="75F0D404CA0DFFA3FBCFDCCED426FA27" box="[1074,1150,1411,1436]" italics="true" pageId="10" pageNumber="385">T. guoi</emphasis>
|
||
Chen, Shi, Vogel & Ding
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="80847395CA0DFFA3FF6ADCEAD12BFA7A" authority="Kramer" authorityName="Kramer" authorityYear="1977" box="[151,371,1447,1473]" class="Squamata" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" pageId="10" pageNumber="385" phylum="Chordata" rank="species" species="insularis">
|
||
<emphasis id="75F0D404CA0DFFA3FF6ADCEAD14CFA7A" box="[151,276,1447,1473]" italics="true" pageId="10" pageNumber="385">T. insularis</emphasis>
|
||
Kramer
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="80847395CA0DFFA3FE80DCEAD40EFA7A" authority="Mirza, Bhosale, Phansalkar, Sawant, Gowande & Patel" authorityName="Mirza, Bhosale, Phansalkar, Sawant, Gowande & Patel" authorityYear="2020" box="[381,1110,1447,1473]" class="Squamata" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" pageId="10" pageNumber="385" phylum="Chordata" rank="species" species="salazar">
|
||
<emphasis id="75F0D404CA0DFFA3FE80DCEAD1B1FA7A" box="[381,489,1447,1473]" italics="true" pageId="10" pageNumber="385">T. salazar</emphasis>
|
||
Mirza, Bhosale, Phansalkar, Sawant, Gowande & Patel
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="80847395CA0DFFA3FB9DDCEAD525FA7A" authority="Kramer" authorityName="Kramer" authorityYear="1977" box="[1120,1405,1447,1473]" class="Squamata" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" pageId="10" pageNumber="385" phylum="Chordata" rank="species" species="septentrionalis">
|
||
<emphasis id="75F0D404CA0DFFA3FB9DDCEAD578FA7A" box="[1120,1312,1447,1473]" italics="true" pageId="10" pageNumber="385">T. septentrionalis</emphasis>
|
||
Kramer
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="80847395CA0DFFA3FA75DCEAD1BBFA5E" authority="Vogel, Nguyen & David" authorityName="Vogel, Nguyen & David" authorityYear="2023" class="Squamata" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" pageId="10" pageNumber="385" phylum="Chordata" rank="species" species="uetzi">
|
||
<emphasis id="75F0D404CA0DFFA3FA75DCEAD094FA5F" italics="true" pageId="10" pageNumber="385">T. uetzi</emphasis>
|
||
Vogel, Nguyen & David
|
||
</taxonomicName>
|
||
by having: body size smaller (max SVL
|
||
<quantity id="807CA5F3CA0DFFA3FC4DDC86D455FA5E" box="[944,1037,1483,1509]" metricMagnitude="-1" metricUnit="m" metricValue="6.38" pageId="10" pageNumber="385" unit="mm" value="638.0">638 mm</quantity>
|
||
vs.
|
||
<quantity id="807CA5F3CA0DFFA3FBC6DC86D4C1FA5E" box="[1083,1177,1483,1509]" metricMagnitude="-1" metricUnit="m" metricValue="8.04" pageId="10" pageNumber="385" unit="mm" value="804.0">804 mm</quantity>
|
||
in
|
||
<taxonomicName id="80847395CA0DFFA3FB41DC86D53CFA5E" authorityName="Chen, Ding, Vogel & Shi" authorityYear="2020" box="[1212,1380,1483,1509]" class="Squamata" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" pageId="10" pageNumber="385" phylum="Chordata" rank="species" species="caudornatus">
|
||
<emphasis id="75F0D404CA0DFFA3FB41DC86D53CFA5E" box="[1212,1380,1483,1509]" italics="true" pageId="10" pageNumber="385">T. caudornatus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<quantity id="807CA5F3CA0DFFA3FA8FDC87D09BF9B2" metricMagnitude="-1" metricUnit="m" metricValue="8.35" pageId="10" pageNumber="385" unit="mm" value="835.0">835 mm</quantity>
|
||
in
|
||
<taxonomicName id="80847395CA0DFFA3FF15DCA2D112F9B2" authorityName="Chandramouli, Campbell & Vogel" authorityYear="2020" box="[232,330,1519,1545]" class="Squamata" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" pageId="10" pageNumber="385" phylum="Chordata" rank="species" species="davidi">
|
||
<emphasis id="75F0D404CA0DFFA3FF15DCA2D112F9B2" box="[232,330,1519,1545]" italics="true" pageId="10" pageNumber="385">T. davidi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<quantity id="807CA5F3CA0DFFA3FEA4DCA2D1EEF9B2" box="[345,438,1519,1545]" metricMagnitude="-1" metricUnit="m" metricValue="8.04" pageId="10" pageNumber="385" unit="mm" value="804.0">804 mm</quantity>
|
||
in
|
||
<taxonomicName id="80847395CA0DFFA3FE26DCA2D271F9B3" authorityName="Chen, Shi, Vogel & Ding" authorityYear="2021" box="[475,553,1519,1544]" class="Squamata" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" pageId="10" pageNumber="385" phylum="Chordata" rank="species" species="guoi">
|
||
<emphasis id="75F0D404CA0DFFA3FE26DCA2D271F9B3" box="[475,553,1519,1544]" italics="true" pageId="10" pageNumber="385">T. guoi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<quantity id="807CA5F3CA0DFFA3FDC5DCA3D2CDF9B2" box="[568,661,1518,1545]" metricMagnitude="-1" metricUnit="m" metricValue="7.35" pageId="10" pageNumber="385" unit="mm" value="735.0">735 mm</quantity>
|
||
in
|
||
<taxonomicName id="80847395CA0DFFA3FD47DCA2D360F9B2" authorityName="Kramer" authorityYear="1977" box="[698,824,1519,1545]" class="Squamata" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" pageId="10" pageNumber="385" phylum="Chordata" rank="species" species="insularis">
|
||
<emphasis id="75F0D404CA0DFFA3FD47DCA2D360F9B2" box="[698,824,1519,1545]" italics="true" pageId="10" pageNumber="385">T. insularis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<quantity id="807CA5F3CA0DFFA3FCBADCA2D3FCF9B2" box="[839,932,1519,1545]" metricMagnitude="-1" metricUnit="m" metricValue="8.19" pageId="10" pageNumber="385" unit="mm" value="819.0">819 mm</quantity>
|
||
in
|
||
<taxonomicName id="80847395CA0DFFA3FC34DCA2D4D3F9B2" authorityName="Kramer" authorityYear="1977" box="[969,1163,1519,1545]" class="Squamata" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" pageId="10" pageNumber="385" phylum="Chordata" rank="species" species="septentrionalis">
|
||
<emphasis id="75F0D404CA0DFFA3FC34DCA2D4D3F9B2" box="[969,1163,1519,1545]" italics="true" pageId="10" pageNumber="385">T. septentrionalis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<quantity id="807CA5F3CA0DFFA3FB67DCA2D4A0F9B2" box="[1178,1272,1519,1545]" metricMagnitude="-1" metricUnit="m" metricValue="6.89" pageId="10" pageNumber="385" unit="mm" value="689.0">689 mm</quantity>
|
||
in
|
||
<taxonomicName id="80847395CA0DFFA3FAE0DCA2D536F9B3" authorityName="Vogel, Nguyen & David" authorityYear="2023" box="[1309,1390,1519,1544]" class="Squamata" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" pageId="10" pageNumber="385" phylum="Chordata" rank="species" species="uetzi">
|
||
<emphasis id="75F0D404CA0DFFA3FAE0DCA2D536F9B3" box="[1309,1390,1519,1544]" italics="true" pageId="10" pageNumber="385">T. uetzi</emphasis>
|
||
</taxonomicName>
|
||
); in contrast, body size larger (max SVL
|
||
<quantity id="807CA5F3CA0DFFA3FDCFDF5ED2D7F996" box="[562,655,1555,1581]" metricMagnitude="-1" metricUnit="m" metricValue="6.38" pageId="10" pageNumber="385" unit="mm" value="638.0">638 mm</quantity>
|
||
vs.
|
||
<quantity id="807CA5F3CA0DFFA3FD40DF5FD342F996" box="[701,794,1554,1581]" metricMagnitude="-1" metricUnit="m" metricValue="5.65" pageId="10" pageNumber="385" unit="mm" value="565.0">565 mm</quantity>
|
||
in
|
||
<taxonomicName id="80847395CA0DFFA3FCC0DF5ED3F2F996" authorityName="Mirza, Bhosale, Phansalkar, Sawant, Gowande & Patel" authorityYear="2020" box="[829,938,1555,1581]" class="Squamata" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" pageId="10" pageNumber="385" phylum="Chordata" rank="species" species="salazar">
|
||
<emphasis id="75F0D404CA0DFFA3FCC0DF5ED3F2F996" box="[829,938,1555,1581]" italics="true" pageId="10" pageNumber="385">T. salazar</emphasis>
|
||
</taxonomicName>
|
||
); throat, chin, and lower labials in shades of blue (vs. creamy or white); relatively larger size of the eye (most obvious in adults); relatively wider supraoculars; the shape of the head (widens quite abruptly behind the eyes vs. more elongate or oval).
|
||
</paragraph>
|
||
<paragraph id="473B0816CA0DFFA3FF3ADF32D1DBF8CA" blockId="10.[151,1437,151,2013]" pageId="10" pageNumber="385">
|
||
Lastly,
|
||
<taxonomicName id="80847395CA0DFFA3FEE0DF32D267F922" authorityName="Idiiatullina & Nguyen & Bragin & Pawangkhanant & Le & Vogel & David & Poyarkov" authorityYear="2024" box="[285,575,1663,1689]" class="Squamata" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" pageId="10" pageNumber="385" phylum="Chordata" rank="species" species="cyanolabris" status="sp. nov.">
|
||
<emphasis id="75F0D404CA0DFFA3FEE0DF32D267F922" box="[285,575,1663,1689]" italics="true" pageId="10" pageNumber="385">Trimeresurus cyanolabris</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="75F0D404CA0DFFA3FDB4DF32D2FEF922" bold="true" box="[585,678,1663,1689]" pageId="10" pageNumber="385">
|
||
<taxonomicNameLabel id="6EC3697FCA0DFFA3FDB4DF32D2FEF922" box="[585,678,1663,1689]" pageId="10" pageNumber="385" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
was previously misidentified with
|
||
<taxonomicName id="80847395CA0DFFA3FBBDDF32D48AF922" authorityName="Gray" authorityYear="1842" box="[1088,1234,1663,1689]" class="Squamata" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" pageId="10" pageNumber="385" phylum="Chordata" rank="species" species="albolabris">
|
||
<emphasis id="75F0D404CA0DFFA3FBBDDF32D48AF922" box="[1088,1234,1663,1689]" italics="true" pageId="10" pageNumber="385">T. albolabris</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="80847395CA0DFFA3FAF2DF32D0ACF906" authority="Schmidt" authorityName="Schmidt" authorityYear="1925" class="Squamata" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" pageId="10" pageNumber="385" phylum="Chordata" rank="species" species="stejnegeri">
|
||
<emphasis id="75F0D404CA0DFFA3FAF2DF32D5C4F923" box="[1295,1436,1663,1688]" italics="true" pageId="10" pageNumber="385">T. stejnegeri</emphasis>
|
||
Schmidt
|
||
</taxonomicName>
|
||
(see
|
||
<bibRefCitation id="231575E7CA0DFFA3FECDDFEED253F906" author="Nguyen, S. V. & Ho, C. T. & Nguyen, T. Q." box="[304,523,1699,1725]" pageId="10" pageNumber="385" refId="ref19451" refString="Nguyen, S. V., Ho, C. T. & Nguyen, T. Q. (2009) Herpetofauna of Vietnam. Chimaira, Frankfurt, 768 pp." type="book" year="2009">
|
||
Nguyen
|
||
<emphasis id="75F0D404CA0DFFA3FE6DDFEED19AF906" box="[400,450,1699,1725]" italics="true" pageId="10" pageNumber="385">et al</emphasis>
|
||
. 2009
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="231575E7CA0DFFA3FDEADFEED2BDF906" author="Hoang, M. D. & Luu, H. T. & Tran, B. V. & Le, D. & To, V. Q. & Dang, V. D. & Nguyen, H. N." box="[535,741,1699,1725]" pageId="10" pageNumber="385" refId="ref17529" refString="Hoang, M. D., Luu, H. T., Tran, B. V., Le, D., To, V. Q., Dang, V. D. & Nguyen, H. N. (2019) Forest biodiversity in Khanh Hoa province - Part 2 Animals. Publishing House for Science and Technology, 215 pp. [in Vietnamese]" type="book" year="2019">
|
||
Hoang
|
||
<emphasis id="75F0D404CA0DFFA3FD94DFEED2C2F906" box="[617,666,1699,1725]" italics="true" pageId="10" pageNumber="385">et al</emphasis>
|
||
. 2019
|
||
</bibRefCitation>
|
||
). The new species can be distinguished from
|
||
<taxonomicName id="80847395CA0DFFA3FB1FDFEED529F906" authorityName="Gray" authorityYear="1842" box="[1250,1393,1699,1725]" class="Squamata" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" pageId="10" pageNumber="385" phylum="Chordata" rank="species" species="albolabris">
|
||
<emphasis id="75F0D404CA0DFFA3FB1FDFEED529F906" box="[1250,1393,1699,1725]" italics="true" pageId="10" pageNumber="385">T. albolabris</emphasis>
|
||
</taxonomicName>
|
||
by: body size smaller (max SVL
|
||
<quantity id="807CA5F3CA0DFFA3FE2BDF8AD269F95A" box="[470,561,1735,1761]" metricMagnitude="-1" metricUnit="m" metricValue="6.38" pageId="10" pageNumber="385" unit="mm" value="638.0">638 mm</quantity>
|
||
vs.
|
||
<quantity id="807CA5F3CA0DFFA3FDA1DF8AD2E3F95A" box="[604,699,1735,1761]" metricMagnitude="-1" metricUnit="m" metricValue="7.49" pageId="10" pageNumber="385" unit="mm" value="749.0">749 mm</quantity>
|
||
); relatively larger size of the eye, the relatively wider supraoculars, and the shape of the head (widens quite abruptly behind vs. more elongate or oval); it is distinguished from
|
||
<taxonomicName id="80847395CA0DFFA3FA75DFA6D15DF893" authorityName="Schmidt" authorityYear="1925" class="Squamata" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" pageId="10" pageNumber="385" phylum="Chordata" rank="species" species="stejnegeri">
|
||
<emphasis id="75F0D404CA0DFFA3FA75DFA6D15DF893" italics="true" pageId="10" pageNumber="385">T. stejnegeri</emphasis>
|
||
</taxonomicName>
|
||
by the 1
|
||
<superScript id="B0F1A55ECA0DFFA3FE97DE42D12DF8A6" attach="left" box="[362,373,1807,1821]" fontSize="6" pageId="10" pageNumber="385">st</superScript>
|
||
supralabial fused with nasal (vs. entirely separated by a suture); body size smaller (max SVL
|
||
<quantity id="807CA5F3CA0DFFA3FF6ADE7ED0ABF8F6" box="[151,243,1843,1869]" metricMagnitude="-1" metricUnit="m" metricValue="6.38" pageId="10" pageNumber="385" unit="mm" value="638.0">638 mm</quantity>
|
||
vs.
|
||
<quantity id="807CA5F3CA0DFFA3FEE3DE7ED126F8F6" box="[286,382,1843,1869]" metricMagnitude="-1" metricUnit="m" metricValue="7.2" pageId="10" pageNumber="385" unit="mm" value="720.0">720 mm</quantity>
|
||
); no postocular stripe in males (vs. present); eye color bright yellow in both sexes (vs. in shades of red in both sexes).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0F9E5B9DCA0DFFA2FF3ADE37D395FF42" lastPageId="11" lastPageNumber="386" pageId="10" pageNumber="385" type="distribution">
|
||
<paragraph id="473B0816CA0DFFA2FF3ADE37D395FF42" blockId="10.[151,1437,151,2013]" lastBlockId="11.[151,1437,151,753]" lastPageId="11" lastPageNumber="386" pageId="10" pageNumber="385">
|
||
<emphasis id="75F0D404CA0DFFA3FF3ADE37D101F82F" bold="true" box="[199,345,1914,1940]" pageId="10" pageNumber="385">Distribution</emphasis>
|
||
. Currently,
|
||
<taxonomicName id="80847395CA0DFFA3FE19DE36D35EF82E" authorityName="Idiiatullina & Nguyen & Bragin & Pawangkhanant & Le & Vogel & David & Poyarkov" authorityYear="2024" box="[484,774,1915,1941]" class="Squamata" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" pageId="10" pageNumber="385" phylum="Chordata" rank="species" species="cyanolabris" status="sp. nov.">
|
||
<emphasis id="75F0D404CA0DFFA3FE19DE36D35EF82E" box="[484,774,1915,1941]" italics="true" pageId="10" pageNumber="385">Trimeresurus cyanolabris</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="75F0D404CA0DFFA3FCEDDE36D334F82E" bold="true" box="[784,876,1915,1941]" pageId="10" pageNumber="385">
|
||
<taxonomicNameLabel id="6EC3697FCA0DFFA3FCEDDE36D334F82E" box="[784,876,1915,1941]" pageId="10" pageNumber="385" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is known from the southern and central parts of coastal
|
||
<collectingCountry id="3F934886CA0DFFA3FF16DED2D112F802" box="[235,330,1951,1977]" name="Vietnam" pageId="10" pageNumber="385">Vietnam</collectingCountry>
|
||
(from Ba To District in
|
||
<collectingRegion id="8540C6F4CA0DFFA3FDA4DED2D317F802" box="[601,847,1951,1977]" country="Vietnam" name="Quang Ngai" pageId="10" pageNumber="385">Quang Ngai Province</collectingRegion>
|
||
; from Quy Nhon City and Van Canh District in
|
||
<collectingRegion id="8540C6F4CA0DFFA3FA9BDED2D166F866" country="Vietnam" name="Binh Dinh" pageId="10" pageNumber="385">Binh Dinh Province</collectingRegion>
|
||
; from Diem Khanh District, Ninh Hoa Town, and Hon Ba NR in Cam Lam District in
|
||
<collectingRegion id="8540C6F4CA0DFFA2FAE6DE8ED0A6FF0A" country="Vietnam" lastPageId="11" lastPageNumber="386" name="Khanh Hoa" pageId="10" pageNumber="385">Khanh Hoa Province</collectingRegion>
|
||
; from Nui Chua NP in Ninh Hai District and Phuoc Binh NP., Bac Ai District in
|
||
<collectingRegion id="8540C6F4CA0CFFA2FB62D9DAD5CCFF0A" box="[1183,1428,151,177]" country="Vietnam" name="Ninh Thuan" pageId="11" pageNumber="386">Ninh Thuan Province</collectingRegion>
|
||
) (
|
||
<figureCitation id="DFBF1493CA0CFFA2FF62D9F6D0BCFF6E" box="[159,228,187,213]" captionStart="FIGURE 1" captionStartId="2.[152,255,1655,1679]" captionTargetBox="[154,1433,378,1629]" captionTargetId="figure-158@2.[154,1433,378,1629]" captionTargetPageId="2" captionText="FIGURE 1. Distribution of the green-colored members of the Trimeresurus macrops species complex in Indochina. Colors of icons and locality numbers correspond to those of Fig. 2 and Appendix III. Stars indicate the type localities; circles indicate the localities of specimens that were used in phylogenetic analysis; (for the Trimeresurus cyanolabris sp. nov. specimens from the type locality were included in the phylogenetic analysis) and triangles indicate distribution records (see Appendix III)." figureDoi="http://doi.org/10.5281/zenodo.12681168" httpUri="https://zenodo.org/record/12681168/files/figure.png" pageId="11" pageNumber="386">Fig. 1</figureCitation>
|
||
, Appendix III). The occurrence of the new species in the
|
||
<collectingRegion id="8540C6F4CA0CFFA2FC7ED9F6D3BFFF6E" box="[899,999,187,213]" country="Vietnam" name="Dak Lak" pageId="11" pageNumber="386">Dak Lak</collectingRegion>
|
||
and
|
||
<collectingRegion id="8540C6F4CA0CFFA2FBDFD9F6D4DAFF6E" box="[1058,1154,187,213]" country="Vietnam" name="Phu Yen" pageId="11" pageNumber="386">Phu Yen</collectingRegion>
|
||
provinces of
|
||
<collectingCountry id="3F934886CA0CFFA2FADCD9F6D5D8FF6E" box="[1313,1408,187,213]" name="Vietnam" pageId="11" pageNumber="386">Vietnam</collectingCountry>
|
||
is strongly anticipated. The actual extent of its distribution is still unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0F9E5B9DCA0CFFA2FF3AD84ED364FD4A" pageId="11" pageNumber="386" type="biology_ecology">
|
||
<paragraph id="473B0816CA0CFFA2FF3AD84ED364FD4A" blockId="11.[151,1437,151,753]" pageId="11" pageNumber="386">
|
||
<emphasis id="75F0D404CA0CFFA2FF3AD84ED1D9FEA7" bold="true" box="[199,385,258,285]" pageId="11" pageNumber="386">Natural history</emphasis>
|
||
.
|
||
<taxonomicName id="80847395CA0CFFA2FE6DD84ED2E9FEA6" authorityName="Idiiatullina & Nguyen & Bragin & Pawangkhanant & Le & Vogel & David & Poyarkov" authorityYear="2024" box="[400,689,259,285]" class="Squamata" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" pageId="11" pageNumber="386" phylum="Chordata" rank="species" species="cyanolabris" status="sp. nov.">
|
||
<emphasis id="75F0D404CA0CFFA2FE6DD84ED2E9FEA6" box="[400,689,259,285]" italics="true" pageId="11" pageNumber="386">Trimeresurus cyanolabris</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="75F0D404CA0CFFA2FD44D84ED34CFEA6" bold="true" box="[697,788,259,285]" pageId="11" pageNumber="386">
|
||
<taxonomicNameLabel id="6EC3697FCA0CFFA2FD44D84ED34CFEA6" box="[697,788,259,285]" pageId="11" pageNumber="386" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
was recorded from relatively low- to mid-elevations (ca.
|
||
<quantity id="807CA5F3CA0CFFA2FF6AD86AD151FEFA" box="[151,265,295,321]" metricMagnitude="2" metricUnit="m" metricValue="2.45" metricValueMax="4.0" metricValueMin="0.9" pageId="11" pageNumber="386" unit="m" value="245.0" valueMax="400.0" valueMin="90.0">90–400 m</quantity>
|
||
a.s.l.) in tropical dry forests of the central southern region of
|
||
<collectingCountry id="3F934886CA0CFFA2FC3CD86AD478FEFA" box="[961,1056,295,321]" name="Vietnam" pageId="11" pageNumber="386">Vietnam</collectingCountry>
|
||
(
|
||
<bibRefCitation id="231575E7CA0CFFA2FBD3D86AD578FEFA" author="Poyarkov, N. A. & Nguyen, T. V. & Popov, E. S. & Geissler, P. & Pawangkhanant, P. & Neang, T. & Suwannapoom, C. & Ananjeva, N. B. & Orlov, N. L." box="[1070,1312,295,321]" pageId="11" pageNumber="386" pagination="255 - 476" refId="ref19734" refString="Poyarkov, N. A., Nguyen, T. V., Popov, E. S., Geissler, P., Pawangkhanant, P., Neang, T., Suwannapoom, C., Ananjeva, N. B. & Orlov, N. L. (2023) Recent progress in taxonomic studies, biogeographic analysis and revised checklist of Reptilians in Indochina. Russian Journal of Herpetology, 30 (5), 255 - 476. https: // doi. org / 10.30906 / 1026 - 2296 - 2023 - 30 - 5 - 255 - 476" type="journal article" year="2023">
|
||
Poyarkov
|
||
<emphasis id="75F0D404CA0CFFA2FB5ED86AD484FEFA" box="[1187,1244,295,321]" italics="true" pageId="11" pageNumber="386">et al.</emphasis>
|
||
2023
|
||
</bibRefCitation>
|
||
). The male
|
||
<typeStatus id="983FB6B4CA0CFFA2FF6AD806D0A1FEDE" box="[151,249,331,357]" pageId="11" pageNumber="386" type="holotype">holotype</typeStatus>
|
||
ZMMU Re-17679 was collected from the ground near a small mountain stream; the male
|
||
<typeStatus id="983FB6B4CA0CFFA2FB22D806D567FEDE" box="[1247,1343,331,357]" pageId="11" pageNumber="386" type="paratype">paratype</typeStatus>
|
||
ZMMU Re-17678 while perching on a bush ca.
|
||
<quantity id="807CA5F3CA0CFFA2FDA9D822D2C1FE32" box="[596,665,367,393]" metricMagnitude="-1" metricUnit="m" metricValue="4.0" pageId="11" pageNumber="386" unit="cm" value="40.0">40 cm</quantity>
|
||
above the ground;
|
||
<specimenCount id="5182C39FCA0CFFA2FC8FD822D3B7FE32" box="[882,1007,367,393]" count="2" pageId="11" pageNumber="386" type="female">two female</specimenCount>
|
||
<typeStatus id="983FB6B4CA0CFFA2FC0BD822D439FE32" box="[1014,1121,367,393]" pageId="11" pageNumber="386" type="paratype">paratypes</typeStatus>
|
||
ZMMU Re-17680–81 were collected on the ground (large flat stones) on the bank of a small river (see
|
||
<figureCitation id="DFBF1493CA0CFFA2FC03D8DED41BFE16" box="[1022,1091,402,429]" captionStart="FIGURE 5" captionStartId="11.[152,255,1764,1788]" captionTargetBox="[151,1435,777,1741]" captionTargetId="figure-358@11.[151,1435,777,1741]" captionTargetPageId="11" captionText="FIGURE 5. Habitat of Trimeresurus cyanolabris sp. nov. in Nui Chua NP, Ninh Thuan Province, Vietnam. Photograph by A. M. Bragin." figureDoi="http://doi.org/10.5281/zenodo.12681180" httpUri="https://zenodo.org/record/12681180/files/figure.png" pageId="11" pageNumber="386">Fig. 5</figureCitation>
|
||
); the female
|
||
<typeStatus id="983FB6B4CA0CFFA2FB21D8DED564FE16" box="[1244,1340,403,429]" pageId="11" pageNumber="386" type="paratype">paratype</typeStatus>
|
||
ZMMU Re-17682 was collected while perching on a tree branch ca.
|
||
<quantity id="807CA5F3CA0CFFA2FCC4D8FAD33CFE6A" box="[825,868,439,465]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="11" pageNumber="386" unit="m" value="1.0">1 m</quantity>
|
||
above the ground, while female
|
||
<typeStatus id="983FB6B4CA0CFFA2FB2ED8FAD566FE6A" box="[1235,1342,439,465]" pageId="11" pageNumber="386" type="paratype">paratypes</typeStatus>
|
||
ZMMU Re-17683–84 were collected from the ground in the tropical montane forests on the leaf litter; all specimens were recorded between 1700 h and 2300 h. Therefore, the new species is semi-arboreal with crepuscular and nocturnal activity. During the day, it often perches on the branches or in the hollows of trees
|
||
<quantity id="807CA5F3CA0CFFA2FBC5DB6FD4F2FD86" box="[1080,1194,546,573]" metricMagnitude="0" metricUnit="m" metricValue="1.0" metricValueMax="1.5" metricValueMin="0.5" pageId="11" pageNumber="386" unit="m" value="1.0" valueMax="1.5" valueMin="0.5">0.5–1.5 m</quantity>
|
||
high near streams. In
|
||
<collectingRegion id="8540C6F4CA0CFFA2FF6ADB0AD1DFFDDA" box="[151,391,583,609]" country="Vietnam" name="Khanh Hoa" pageId="11" pageNumber="386">Khanh Hoa Province</collectingRegion>
|
||
, it is common in the beginning (August) and at the end of the rainy season (October), often in the lower layers of trees or on dry branches under the trees, in contrast to
|
||
<taxonomicName id="80847395CA0CFFA2FC06DB26D4D2FD3E" authorityName="Gray" authorityYear="1842" box="[1019,1162,619,645]" class="Squamata" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" pageId="11" pageNumber="386" phylum="Chordata" rank="species" species="albolabris">
|
||
<emphasis id="75F0D404CA0CFFA2FC06DB26D4D2FD3E" box="[1019,1162,619,645]" italics="true" pageId="11" pageNumber="386">T. albolabris</emphasis>
|
||
</taxonomicName>
|
||
which usually occurs at higher tree levels. There may be an ecological stratification between these two species in the
|
||
<collectingRegion id="8540C6F4CA0CFFA2FB62DBC2D547FD12" box="[1183,1311,655,681]" country="Vietnam" name="Khanh Hoa" pageId="11" pageNumber="386">Khanh Hoa</collectingRegion>
|
||
population (T.V. Phan pers. comm.). The new species feeds on small frogs and small lizards (such as
|
||
<taxonomicName id="80847395CA0CFFA2FB7ADBFED4ADFD76" box="[1159,1269,691,717]" class="Squamata" family="Gekkonidae" genus="Gekko" kingdom="Animalia" pageId="11" pageNumber="386" phylum="Chordata" rank="species" species="undetermined">
|
||
<emphasis id="75F0D404CA0CFFA2FB7ADBFED497FD76" box="[1159,1231,691,717]" italics="true" pageId="11" pageNumber="386">Gekko</emphasis>
|
||
sp.
|
||
</taxonomicName>
|
||
) (T.V. Nguyen pers. obser.). Nothing is known about its reproduction cycle.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="13FB589ECA0CFFA2FF65DFA9D15FF89B" ID-DOI="http://doi.org/10.5281/zenodo.12681180" ID-Zenodo-Dep="12681180" httpUri="https://zenodo.org/record/12681180/files/figure.png" pageId="11" pageNumber="386" startId="11.[152,255,1764,1788]" targetBox="[151,1435,777,1741]" targetPageId="11" targetType="figure">
|
||
<paragraph id="473B0816CA0CFFA2FF65DFA9D15FF89B" blockId="11.[151,1437,1764,1824]" pageId="11" pageNumber="386">
|
||
<emphasis id="75F0D404CA0CFFA2FF65DFA9D141F947" bold="true" box="[152,281,1764,1788]" pageId="11" pageNumber="386">FIGURE 5.</emphasis>
|
||
Habitat of
|
||
<taxonomicName id="80847395CA0CFFA2FE70DFA8D2CAF947" authorityName="Idiiatullina & Nguyen & Bragin & Pawangkhanant & Le & Vogel & David & Poyarkov" authorityYear="2024" box="[397,658,1764,1788]" class="Squamata" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" pageId="11" pageNumber="386" phylum="Chordata" rank="species" species="cyanolabris" status="sp. nov.">
|
||
<emphasis id="75F0D404CA0CFFA2FE70DFA8D2CAF947" box="[397,658,1764,1788]" italics="true" pageId="11" pageNumber="386">Trimeresurus cyanolabris</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="75F0D404CA0CFFA2FD65DFA9D2B1F947" bold="true" box="[664,745,1764,1788]" pageId="11" pageNumber="386">
|
||
<taxonomicNameLabel id="6EC3697FCA0CFFA2FD65DFA9D2B1F947" box="[664,745,1764,1788]" pageId="11" pageNumber="386" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in Nui Chua NP, Ninh Thuan Province, Vietnam. Photograph by A. M. Bragin.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="0F9E5B9DCA0CFFA5FF3ADE3FD4FAFF42" lastPageId="12" lastPageNumber="387" pageId="11" pageNumber="386" type="conservation">
|
||
<paragraph id="473B0816CA0CFFA5FF3ADE3FD4FAFF42" blockId="11.[151,1437,1906,2041]" lastBlockId="12.[151,1437,151,249]" lastPageId="12" lastPageNumber="387" pageId="11" pageNumber="386">
|
||
<emphasis id="75F0D404CA0CFFA2FF3ADE3FD1E7F837" bold="true" box="[199,447,1906,1932]" pageId="11" pageNumber="386">Conservation status.</emphasis>
|
||
Further research is required to clarify the extent of the distribution, population size, trends, and conservation status of the new species.
|
||
<taxonomicName id="80847395CA0CFFA2FD35DEDAD3BEF80B" authorityName="Idiiatullina & Nguyen & Bragin & Pawangkhanant & Le & Vogel & David & Poyarkov" authorityYear="2024" box="[712,998,1942,1968]" class="Squamata" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" pageId="11" pageNumber="386" phylum="Chordata" rank="species" species="cyanolabris" status="sp. nov.">
|
||
<emphasis id="75F0D404CA0CFFA2FD35DEDAD3BEF80B" box="[712,998,1942,1968]" italics="true" pageId="11" pageNumber="386">Trimeresurus cyanolabris</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="75F0D404CA0CFFA2FC11DEDBD41CF80B" bold="true" box="[1004,1092,1942,1968]" pageId="11" pageNumber="386">
|
||
<taxonomicNameLabel id="6EC3697FCA0CFFA2FC11DEDBD41CF80B" box="[1004,1092,1942,1968]" pageId="11" pageNumber="386" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is distributed over a large area, including several protected zones. Across its range, the new species seems to be quite common, especially in
|
||
<collectingRegion id="8540C6F4CA0CFFA2FAAFDEF7D16CF843" country="Vietnam" name="Khanh Hoa" pageId="11" pageNumber="386">Khanh Hoa Province</collectingRegion>
|
||
, where many local people and tourists have observed and reported it through the social networking group “Snake Identification and First Aid for Snakebites in
|
||
<collectingCountry id="3F934886CA0BFFA5FCCCD9DAD3CBFF0A" box="[817,915,151,177]" name="Vietnam" pageId="12" pageNumber="387">Vietnam</collectingCountry>
|
||
” (SIFASV,
|
||
<collectingCountry id="3F934886CA0BFFA5FBE9D9DAD42EFF0A" box="[1044,1142,151,177]" name="Vietnam" pageId="12" pageNumber="387">Vietnam</collectingCountry>
|
||
, T.V. Nguyen pers. obser.) Thus, we tentatively suggest that
|
||
<taxonomicName id="80847395CA0BFFA5FDE4D9F6D362FF6E" authorityName="Idiiatullina & Nguyen & Bragin & Pawangkhanant & Le & Vogel & David & Poyarkov" authorityYear="2024" box="[537,826,187,213]" class="Squamata" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" pageId="12" pageNumber="387" phylum="Chordata" rank="species" species="cyanolabris" status="sp. nov.">
|
||
<emphasis id="75F0D404CA0BFFA5FDE4D9F6D362FF6E" box="[537,826,187,213]" italics="true" pageId="12" pageNumber="387">Trimeresurus cyanolabris</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="75F0D404CA0BFFA5FCB9D9F6D3C7FF6E" bold="true" box="[836,927,187,213]" pageId="12" pageNumber="387">
|
||
<taxonomicNameLabel id="6EC3697FCA0BFFA5FCB9D9F6D3C7FF6E" box="[836,927,187,213]" pageId="12" pageNumber="387" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
should be considered a Least Concern (LC) following the IUCN‘s Red List categories (IUCN Standards and Petitions Committee 2019).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |