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<document id="ABC315F5A44CBF73B241433ABE7D3BDC" ID-CLB-Dataset="26002" ID-DOI="10.11646/zootaxa.4657.1.2" ID-GBIF-Dataset="a7c3cd9d-d4be-4686-a198-3255c948beb8" ID-ISSN="1175-5326" ID-Zenodo-Dep="3369801" ID-ZooBank="F27CB967-C001-436A-9F25-184D5484363C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1565968506341" checkinUser="plazi" docAuthor="Perkins, Philip D. &amp; Bergsten, Johannes" docDate="2019" docId="AD076747FFC1FFD4FF6DFA48B16B905F" docLanguage="en" docName="zootaxa.4657.1.2.pdf" docOrigin="Zootaxa 4657 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Incoltorrida zahamena Perkins &amp; Bergsten 2019, new species" docType="treatment" docVersion="12" lastPageNumber="80" masterDocId="513E1F3FFFD2FFC3FFFAFFB9B27A9138" masterDocTitle="New Myxophagan water beetles from Madagascar (Coleoptera: Torridincolidae, Hydroscaphidae)" masterLastPageNumber="96" masterPageNumber="57" pageNumber="76" updateTime="1698751487288" updateUser="ExternalLinkService">
<mods:mods id="F56B9F96364175B7F68B604AE8C31A34" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="4E2400F582DAD021076021964F5C9064">
<mods:title id="C206611DD48073994821743B04DA3854">New Myxophagan water beetles from Madagascar (Coleoptera: Torridincolidae, Hydroscaphidae)</mods:title>
</mods:titleInfo>
<mods:name id="9ECB91FD0689E4F5D631FF57EDA05CF4" type="personal">
<mods:role id="4CCB208610D5FDE8E3045EFFA0DAB320">
<mods:roleTerm id="F7554522DDB9E1097D60121C1AA5F295">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="3D726094D9DBB4783A9B3645BFBF6A76">Perkins, Philip D.</mods:namePart>
</mods:name>
<mods:name id="6D72619838F63498CDB7A127323EDB2A" type="personal">
<mods:role id="FED1E7D19D9D02ED9FB3A3101E249591">
<mods:roleTerm id="176638D9AD7C6F15E971C2A373D01DCA">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E9DA24811EAF5A63A4FE0DDF7D9C561C">Bergsten, Johannes</mods:namePart>
</mods:name>
<mods:typeOfResource id="A4D6C598F627C0A3E92A30E9DD2F84B6">text</mods:typeOfResource>
<mods:relatedItem id="7B78A4CB023D62F9E7AD5F53C44C5D55" type="host">
<mods:titleInfo id="08EA236E17DF0BF3D35B40CBAC2D95FC">
<mods:title id="F2383C373C0AE63C69147E6265AC7462">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="32C19856AE6C6983B37341E79BD22FB4">
<mods:date id="F0CFAF5DBD667382E670F501C982F8C6">2019</mods:date>
<mods:detail id="114428E566CC3D47DD79D1F7325BF8EA" type="pubDate">
<mods:number id="026AFEEA0C18FF1ADBB06461A6129DF3">2019-08-16</mods:number>
</mods:detail>
<mods:detail id="5F3DF048EBF33FADAB4DEC2CB4D71825" type="volume">
<mods:number id="1DD8AAD8A63DFD6A1E62B1A5827F050E">4657</mods:number>
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<mods:detail id="F4CCB507CB041B0F4074BD85958EE099" type="issue">
<mods:number id="848986FB5813BAEF3DA7141ED662FBC9">1</mods:number>
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<mods:start id="9A5DF9C4E8C6F21D14476E107B947369">57</mods:start>
<mods:end id="9FDED774BBCAC706ABF717F9F5DD2ED9">96</mods:end>
</mods:extent>
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<mods:classification id="9349BE4FC325E02839CC98F6672B80F0">journal article</mods:classification>
<mods:identifier id="A3F5DC8A04F4A02ABC06A598371B61F5" type="CLB-Dataset">26002</mods:identifier>
<mods:identifier id="13CF82B67F4F59B13FBE8E4BDEAFF764" type="DOI">10.11646/zootaxa.4657.1.2</mods:identifier>
<mods:identifier id="64FC10B0738B3D000C730F36618541C4" type="GBIF-Dataset">a7c3cd9d-d4be-4686-a198-3255c948beb8</mods:identifier>
<mods:identifier id="214EFBA07257A051E7E995D89E806D46" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="7BFA2F6A11B20A821E6FD1CFC56BC4EA" type="Zenodo-Dep">3369801</mods:identifier>
<mods:identifier id="FF1D461FA8D6FA0EC0BAE286A66CBE21" type="ZooBank">F27CB967-C001-436A-9F25-184D5484363C</mods:identifier>
</mods:mods>
<treatment id="AD076747FFC1FFD4FF6DFA48B16B905F" ID-DOI="http://doi.org/10.5281/zenodo.5934257" ID-GBIF-Taxon="159131438" ID-Zenodo-Dep="5934257" LSID="urn:lsid:plazi:treatment:AD076747FFC1FFD4FF6DFA48B16B905F" httpUri="http://treatment.plazi.org/id/AD076747FFC1FFD4FF6DFA48B16B905F" lastPageId="23" lastPageNumber="80" pageId="19" pageNumber="76">
<subSubSection id="6DB485DAFFC1FFD0FF6DFA48B0369733" box="[151,588,1521,1548]" pageId="19" pageNumber="76" type="nomenclature">
<paragraph id="2511D651FFC1FFD0FF6DFA48B0369733" blockId="19.[151,816,1521,1584]" box="[151,588,1521,1548]" pageId="19" pageNumber="76">
<heading id="7E59613DFFC1FFD0FF6DFA48B0369733" bold="true" box="[151,588,1521,1548]" fontSize="11" level="1" pageId="19" pageNumber="76" reason="1">
<emphasis id="17DA0A43FFC1FFD0FF6DFA48B0369733" bold="true" box="[151,588,1521,1548]" pageId="19" pageNumber="76">
<taxonomicName id="E2AEADD2FFC1FFD0FF6DFA48B3D79734" authority="Perkins &amp; Bergsten, 2019" authorityName="Perkins &amp; Bergsten" authorityYear="2019" box="[151,429,1521,1548]" class="Insecta" family="Torridincolidae" genus="Incoltorrida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="76" phylum="Arthropoda" rank="species" species="zahamena" status="sp. nov.">
<emphasis id="17DA0A43FFC1FFD0FF6DFA48B3D79734" bold="true" box="[151,429,1521,1548]" italics="true" pageId="19" pageNumber="76">Incoltorrida zahamena</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="0CE9B738FFC1FFD0FE46FA4BB0369733" box="[444,588,1521,1548]" pageId="19" pageNumber="76" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="6DB485DAFFC1FFD0FF6DF9ACB14A9708" box="[151,816,1557,1584]" pageId="19" pageNumber="76" type="description">
<paragraph id="2511D651FFC1FFD0FF6DF9ACB14A9708" blockId="19.[151,816,1521,1584]" box="[151,816,1557,1584]" pageId="19" pageNumber="76">
<figureCitation id="BD95CAD4FFC1FFD0FF6DF9ACB28A9717" box="[151,240,1557,1583]" captionStart="FIGURES 11–12" captionStartId="14.[151,264,2000,2025]" captionTargetId="figure@14.[267,1320,166,1980]" captionTargetPageId="14" captionText="FIGURES 11–12. Incoltorrida zahamena, holotype, habitus and aedeagus." httpUri="https://zenodo.org/record/3387376/files/figure.png" pageId="19" pageNumber="76">Figs. 11</figureCitation>
(habitus); 12 (genitalia); 36 (map); 50, 51 (habitat)
</paragraph>
</subSubSection>
<subSubSection id="6DB485DAFFC1FFD4FF6DF9E7B16B905F" lastPageId="23" lastPageNumber="80" pageId="19" pageNumber="76" type="materials_examined">
<paragraph id="2511D651FFC1FFD0FF6DF9E7B16297DC" blockId="19.[151,1437,1629,2016]" pageId="19" pageNumber="76">
<emphasis id="17DA0A43FFC1FFD0FF6DF9E7B3339740" bold="true" box="[151,329,1629,1656]" pageId="19" pageNumber="76">Type Material.</emphasis>
<materialsCitation id="95C6DC0CFFC1FFD0FEA8F9E4B72D9787" ID-GBIF-Occurrence="2332527085" collectingDate="2018-03-10" collectionCode="NHRS" collectorName="Leg. J. Bergsten &amp; T. Ranarilalatiana" country="Madagascar" county="Alaotra-Mangoro" elevation="100" latitude="-17.5438" location="Manambato river" longLatPrecision="7" longitude="48.723" municipality="Zahamena National Park" pageId="19" pageNumber="76" specimenCode="MAD18-111" specimenCount="1" stateProvince="Toamasina" typeStatus="holotype">
<typeStatus id="FA1568F3FFC1FFD0FEA8F9E4B3C0974F" box="[338,442,1629,1655]" pageId="19" pageNumber="76" type="holotype">Holotype</typeStatus>
(male):
<collectingCountry id="5DB996C1FFC1FFD0FDE1F9E4B0DD974F" box="[539,679,1629,1655]" name="Madagascar" pageId="19" pageNumber="76">Madagascar</collectingCountry>
:
<collectingRegion id="E76A18B3FFC1FFD0FD48F9E4B155974F" box="[690,815,1629,1655]" country="Madagascar" name="Toamasina" pageId="19" pageNumber="76">Toamasina</collectingRegion>
:
<collectingCounty id="CC70AEDDFFC1FFD0FCC3F9E4B67B974F" box="[825,1025,1629,1655]" pageId="19" pageNumber="76">Alaotra-Mangoro</collectingCounty>
:
<collectingMunicipality id="C5754C2BFFC1FFD0FBF7F9E4B753974F" box="[1037,1321,1629,1655]" pageId="19" pageNumber="76">Zahamena National Park</collectingMunicipality>
, Antanandava Sect., midaltitude rainforest:
<location id="2071808AFFC1FFD0FDD9F938B09297A3" LSID="urn:lsid:plazi:treatment:AD076747FFC1FFD4FF6DFA48B16B905F:2071808AFFC1FFD0FDD9F938B09297A3" box="[547,744,1665,1691]" country="Madagascar" county="Alaotra-Mangoro" latitude="-17.5438" longLatPrecision="7" longitude="48.723" municipality="Zahamena National Park" name="Manambato river" pageId="19" pageNumber="76" stateProvince="Toamasina">Manambato river</location>
~
<quantity id="E2567BB4FFC1FFD0FD05F938B13A97A3" box="[767,832,1665,1691]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="19" pageNumber="76" unit="m" value="100.0">
<elevation id="AE833162FFC1FFD0FD05F938B13A97A3" box="[767,832,1665,1691]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="19" pageNumber="76" unit="m" value="100.0">100m</elevation>
</quantity>
downstream of
<location id="2071808AFFC1FFD0FC07F938B6D397A3" LSID="urn:lsid:plazi:treatment:AD076747FFC1FFD4FF6DFA48B16B905F:2071808AFFC1FFD0FC07F938B6D397A3" box="[1021,1193,1665,1691]" country="Madagascar" county="Alaotra-Mangoro" latitude="-17.5438" longLatPrecision="7" longitude="48.723" municipality="Zahamena National Park" name="Camp Cascade" pageId="19" pageNumber="76" stateProvince="Toamasina">Camp Cascade</location>
, hygropetric rocks at night,
<geoCoordinate id="409AB096FFC1FFD0FF24F91CB33697F8" box="[222,332,1701,1728]" degrees="17.5438" direction="south" orientation="latitude" pageId="19" pageNumber="76" precision="5" value="-17.5438">17.5438S</geoCoordinate>
,
<geoCoordinate id="409AB096FFC1FFD0FEACF91CB3BF9787" box="[342,453,1701,1727]" degrees="48.7230" direction="east" orientation="longitude" pageId="19" pageNumber="76" precision="5" value="48.723">48.7230E</geoCoordinate>
,
<quantity id="E2567BB4FFC1FFD0FE2AF91CB05A9787" box="[464,544,1701,1727]" metricMagnitude="3" metricUnit="m" metricValue="1.28" pageId="19" pageNumber="76" unit="m" value="1280.0">
<elevation id="AE833162FFC1FFD0FE2AF91CB05A9787" box="[464,544,1701,1727]" metricMagnitude="3" metricUnit="m" metricValue="1.28" pageId="19" pageNumber="76" unit="m" value="1280.0">1280m</elevation>
</quantity>
,
<date id="5110F091FFC1FFD0FDD1F91CB0D69787" box="[555,684,1701,1727]" pageId="19" pageNumber="76" value="2018-03-10">
<collectingDate id="41540979FFC1FFD0FDD1F91CB0D69787" box="[555,684,1701,1727]" pageId="19" pageNumber="76" value="2018-03-10">10.III.2018</collectingDate>
</date>
,
<specimenCode id="75087E2AFFC1FFD0FD4DF91CB1329787" box="[695,840,1701,1727]" collectionCode="MAD" country="India" httpUri="http://grbio.org/cool/dqz8-sfkc" name="Madras Museum" pageId="19" pageNumber="76">
<collectionCode id="43BF4E94FFC1FFD0FD4DF91CB0849787" box="[695,766,1701,1727]" country="India" httpUri="http://grbio.org/cool/dqz8-sfkc" name="Madras Museum" pageId="19" pageNumber="76">MAD</collectionCode>
18-111
</specimenCode>
,
<collectorName id="885BB387FFC1FFD0FCA9F91FB67D97F8" box="[851,1031,1702,1728]" pageId="19" pageNumber="76">Leg. J. Bergsten</collectorName>
&amp;
<collectorName id="885BB387FFC1FFD0FBD0F91FB68A9787" box="[1066,1264,1701,1728]" pageId="19" pageNumber="76">T. Ranarilalatiana</collectorName>
(
<collectionCode id="43BF4E94FFC1FFD0FB04F91CB7359787" box="[1278,1359,1701,1727]" country="Sweden" httpUri="http://grbio.org/cool/rsbi-vwmq" name="Swedish Museum of Natural History, Entomology Collections" pageId="19" pageNumber="76">NHRS</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="95C6DC0CFFC1FFD0FA9BF91FB16E97DC" ID-GBIF-Occurrence="2332527095" collectingDate="2018-03-10" collectionCode="NHRS, BMNH" collectorName="Leg. J. Bergsten &amp; T. Ranarilalatiana" country="Madagascar" county="Alaotra-Mangoro" elevation="100" latitude="-17.5438" location="Manambato river" longLatPrecision="7" longitude="48.723" municipality="Zahamena National Park" pageId="19" pageNumber="76" specimenCount="2" specimenCount-female="2" stateProvince="Toamasina" typeStatus="paratype">
<typeStatus id="FA1568F3FFC1FFD0FA9BF91FB2AF97DC" pageId="19" pageNumber="76" type="paratype">Paratypes</typeStatus>
, same data as
<typeStatus id="FA1568F3FFC1FFD0FE8FF970B3AD97DB" box="[373,471,1737,1763]" pageId="19" pageNumber="76" type="holotype">holotype</typeStatus>
(
<specimenCount id="33A81DD8FFC1FFD0FE1CF970B02997DB" box="[486,595,1737,1763]" pageId="19" pageNumber="76" type="female">2 females</specimenCount>
<collectionCode id="43BF4E94FFC1FFD0FDA0F970B0DD97DB" box="[602,679,1737,1763]" country="Sweden" httpUri="http://grbio.org/cool/rsbi-vwmq" name="Swedish Museum of Natural History, Entomology Collections" pageId="19" pageNumber="76">NHRS</collectionCode>
,
<collectionCode id="43BF4E94FFC1FFD0FD49F973B17697DC" box="[691,780,1738,1764]" pageId="19" pageNumber="76">BMNH</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="2511D651FFC1FFD0FF3DF954B1FE9668" blockId="19.[151,1437,1629,2016]" pageId="19" pageNumber="76">
<emphasis id="17DA0A43FFC1FFD0FF3DF954B3A99630" bold="true" box="[199,467,1773,1800]" pageId="19" pageNumber="76">Differential Diagnosis.</emphasis>
Differentiated from other members of the genus by the combination of small size (ca.
<quantity id="E2567BB4FFC1FFD0FF6DF8A8B37A9614" box="[151,256,1809,1836]" metricMagnitude="-3" metricUnit="m" metricValue="1.97" pageId="19" pageNumber="76" unit="mm" value="1.97">1.97 mm</quantity>
), oval habitus (
<figureCitation id="BD95CAD4FFC1FFD0FE57F8A8B07A9613" box="[429,512,1809,1835]" captionStart="FIGURES 11–12" captionStartId="14.[151,264,2000,2025]" captionTargetId="figure@14.[267,1320,166,1980]" captionTargetPageId="14" captionText="FIGURES 11–12. Incoltorrida zahamena, holotype, habitus and aedeagus." httpUri="https://zenodo.org/record/3387376/files/figure.png" pageId="19" pageNumber="76">Fig. 11</figureCitation>
), pronotum without median longitudinal ridge, and elytron with eight costae. The elytral series are irregular, giving the elytra a scabrous appearance.
</paragraph>
<paragraph id="2511D651FFC1FFD0FF3DF8E0B0949683" blockId="19.[151,1437,1629,2016]" pageId="19" pageNumber="76">
<emphasis id="17DA0A43FFC1FFD0FF3DF8E0B32B964C" bold="true" box="[199,337,1881,1908]" pageId="19" pageNumber="76">Description</emphasis>
. Size:
<typeStatus id="FA1568F3FFC1FFD0FE5EF8E0B07C964B" box="[420,518,1881,1907]" pageId="19" pageNumber="76" type="holotype">holotype</typeStatus>
(length/width, mm): body (length to elytral apices) 1.97/1.19; head width 0.39; pronotum 0.43/0.92; elytra 1.21/1.19. Dorsum dark brown to black, venter reddish brown, legs reddish brown except femoral-tibial articulations and tarsi dark brown.
</paragraph>
<paragraph id="2511D651FFC1FFD0FF3DF87CB65296E7" blockId="19.[151,1437,1629,2016]" box="[199,1064,1989,2016]" pageId="19" pageNumber="76">Sides of frontoclypeal shield slightly arcuate. Clypeus about as long as frons.</paragraph>
<caption id="71D186D9FFC6FFD7FF6DF8FEB72E9658" box="[151,1364,1863,1889]" httpUri="https://zenodo.org/record/3369833/files/figure.png" pageId="20" pageNumber="77" startId="20.[151,250,1863,1888]" targetBox="[175,1442,198,1825]" targetPageId="20">
<paragraph id="2511D651FFC6FFD7FF6DF8FEB72E9658" blockId="20.[151,1364,1863,1889]" box="[151,1364,1863,1889]" pageId="20" pageNumber="77">
<emphasis id="17DA0A43FFC6FFD7FF6DF8FEB33C9658" bold="true" box="[151,326,1863,1888]" pageId="20" pageNumber="77">FIGURE 23–24.</emphasis>
(23) Last instar larva of
<taxonomicName id="E2AEADD2FFC6FFD7FDB8F8F1B1349658" authorityName="Perkins &amp; Bergsten" authorityYear="2019" box="[578,846,1864,1888]" class="Insecta" family="Torridincolidae" genus="Incoltorrida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="77" phylum="Arthropoda" rank="species" species="quintacostata">
<emphasis id="17DA0A43FFC6FFD7FDB8F8F1B1349658" box="[578,846,1864,1888]" italics="true" pageId="20" pageNumber="77">Incoltorrida quintacostata</emphasis>
</taxonomicName>
; (24) transparency mount of elytron of
<taxonomicName id="E2AEADD2FFC6FFD7FB25F8F0B7379658" authorityName="Perkins &amp; Bergsten" authorityYear="2019" box="[1247,1357,1865,1888]" class="Insecta" family="Torridincolidae" genus="Incoltorrida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="77" phylum="Arthropoda" rank="species" species="marojejy">
<emphasis id="17DA0A43FFC6FFD7FB25F8F0B7379658" box="[1247,1357,1865,1888]" italics="true" pageId="20" pageNumber="77">I. marojejy</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<caption id="71D186D9FFC7FFD6FF6DF8FDB1139665" box="[151,873,1860,1885]" httpUri="https://zenodo.org/record/3369835/files/figure.png" pageId="21" pageNumber="78" startId="21.[151,250,1860,1885]" targetBox="[220,1376,233,1809]" targetPageId="21">
<paragraph id="2511D651FFC7FFD6FF6DF8FDB1139665" blockId="21.[151,873,1860,1885]" box="[151,873,1860,1885]" pageId="21" pageNumber="78">
<emphasis id="17DA0A43FFC7FFD6FF6DF8FDB35A9665" bold="true" box="[151,288,1860,1885]" pageId="21" pageNumber="78">FIGURE 25.</emphasis>
<taxonomicName id="E2AEADD2FFC7FFD6FEDCF8FDB05F9664" authorityName="Paulian" authorityYear="1949" box="[294,549,1860,1884]" class="Insecta" family="Hydroscaphidae" genus="Hydroscapha" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="78" phylum="Arthropoda" rank="species" species="saboureaui">
<emphasis id="17DA0A43FFC7FFD6FEDCF8FDB05F9664" box="[294,549,1860,1884]" italics="true" pageId="21" pageNumber="78">Hydroscapha saboureaui</emphasis>
</taxonomicName>
, habitus and labels of holotype.
</paragraph>
</caption>
<paragraph id="2511D651FFC7FFD6FF3DF832B35496D5" blockId="21.[151,1436,1931,2029]" pageId="21" pageNumber="78">Pronotum with short oblique carinae on anterior 1/3 moderately developed, no indication of midlongitudinal carina, area posterior to carinae transversely rounded, with no indication of ridges or depressions. Cuticle rough, subgranulate.</paragraph>
<caption id="71D186D9FFC4FFD5FF6DF8FEB1749658" box="[151,782,1863,1888]" httpUri="https://zenodo.org/record/3369837/files/figure.png" pageId="22" pageNumber="79" startId="22.[151,264,1863,1888]" targetBox="[272,1310,217,1827]" targetPageId="22">
<paragraph id="2511D651FFC4FFD5FF6DF8FEB1749658" blockId="22.[151,782,1863,1888]" box="[151,782,1863,1888]" pageId="22" pageNumber="79">
<emphasis id="17DA0A43FFC4FFD5FF6DF8FEB3549658" bold="true" box="[151,302,1863,1888]" pageId="22" pageNumber="79">FIGURES 26.</emphasis>
<taxonomicName id="E2AEADD2FFC4FFD5FECEF8F1B04D9658" authorityName="Perkins &amp; Bergsten" authorityYear="2019" box="[308,567,1864,1888]" class="Insecta" family="Hydroscaphidae" genus="Hydroscapha" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="79" phylum="Arthropoda" rank="species" species="andringitra">
<emphasis id="17DA0A43FFC4FFD5FECEF8F1B04D9658" box="[308,567,1864,1888]" italics="true" pageId="22" pageNumber="79">Hydroscapha andringitra</emphasis>
</taxonomicName>
, habitus of holotype.
</paragraph>
</caption>
<paragraph id="2511D651FFC4FFD5FF3DF837B3A396C9" blockId="22.[151,1437,1934,2033]" pageId="22" pageNumber="79">Elytron with eight costae of varying height; costa #6 very low and short, costa #8 generally higher and stronger than other costae; transverse ridges between costae absent or very indistinct. Serial punctures large and coarse, producing scabrous appearance.</paragraph>
<paragraph id="2511D651FFC5FFD4FF3DFF20B0E491EF" blockId="23.[151,1436,153,359]" pageId="23" pageNumber="80">Metaventral tabella with very faint, indistinct transverse grooves; midlongitudinal groove in posterior 1/2 of metaventral tabella widest at about midlength.</paragraph>
<paragraph id="2511D651FFC5FFD4FF3DFF58B6E791C3" blockId="23.[151,1436,153,359]" box="[199,1181,225,251]" pageId="23" pageNumber="80">Midlongitudinal carina of first abdominal ventrite strong and extends length of ventrite.</paragraph>
<paragraph id="2511D651FFC5FFD4FF3DFEBCB3BC907B" blockId="23.[151,1436,153,359]" pageId="23" pageNumber="80">
Male genitalia in lateral view distinctively thickened in basal 1/2, apical 1/2 narrowing, with tip slightly widened and spinose (
<figureCitation id="BD95CAD4FFC5FFD4FE9CFE90B3C3907B" box="[358,441,297,323]" captionStart="FIGURES 11–12" captionStartId="14.[151,264,2000,2025]" captionTargetId="figure@14.[267,1320,166,1980]" captionTargetPageId="14" captionText="FIGURES 11–12. Incoltorrida zahamena, holotype, habitus and aedeagus." httpUri="https://zenodo.org/record/3387376/files/figure.png" pageId="23" pageNumber="80">Fig. 12</figureCitation>
).
</paragraph>
<paragraph id="2511D651FFC5FFD4FF3DFEF4B16B905F" blockId="23.[151,1436,153,359]" box="[199,785,333,359]" pageId="23" pageNumber="80">
<emphasis id="17DA0A43FFC5FFD4FF3DFEF4B332905F" bold="true" box="[199,328,333,359]" pageId="23" pageNumber="80">Etymology</emphasis>
. Named in reference to the
<typeStatus id="FA1568F3FFC5FFD4FD7AFEF4B0CA905F" box="[640,688,333,359]" pageId="23" pageNumber="80">type</typeStatus>
locality.
</paragraph>
</subSubSection>
</treatment>
</document>
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