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<document id="098FBB9F641F872EE289680CECFEFF07" ID-DOI="10.5281/zenodo.13799439" ID-ISSN="0068-547X" ID-Zenodo-Dep="13799439" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722538212676" checkinUser="felipe" docAuthor="Kavanaugh, David H. &amp; Rainio, Johanna" docDate="2016" docId="0A3E26573073FF9BFF8EFD4FBCF10BA1" docLanguage="en" docName="ProcCalifAcadSci.63.7.201-268.pdf" docOrigin="Proceedings of the California Academy of Sciences 63 (7)" docStyle="DocumentStyle:E411DA009AF5E7F9A7E023431E2B7D5B.2:ProcCalifAcadSci.2015-.journal_article" docStyleId="E411DA009AF5E7F9A7E023431E2B7D5B" docStyleName="ProcCalifAcadSci.2015-.journal_article" docStyleVersion="2" docTitle="Dactyleurys minimus Kavanaugh and Rainio 2016, sp. nov." docType="treatment" docVersion="2" lastPageNumber="212" masterDocId="F6075E2F3079FF90FFF5FF92BD630D1B" masterDocTitle="Twenty-six New Species of Predaceous Ground Beetles (Coleoptera: Adephaga: Carabidae) from Ranomafana National Park, Madagascar" masterLastPageNumber="268" masterPageNumber="201" pageNumber="211" updateTime="1726771873529" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="1F3DB88A45C3450D61CA4D430E428C3C" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="DBF109AA8FF4D49FF3539A18AEFE364C">
<mods:title id="9633E7241E6C3A5668FE9F083252D3FE">Twenty-six New Species of Predaceous Ground Beetles (Coleoptera: Adephaga: Carabidae) from Ranomafana National Park, Madagascar</mods:title>
</mods:titleInfo>
<mods:name id="BFD6267A8C584B6C7C3E54E793A86605" type="personal">
<mods:role id="CCA0D2910A1A503F0D6A5BC14A3004FC">
<mods:roleTerm id="B8C891488E4DEFE22E26FE31A5B35EC2">Author</mods:roleTerm>
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<mods:namePart id="DCA21B2F59869238EED5040177BE3396">Kavanaugh, David H.</mods:namePart>
<mods:affiliation id="5A06581E9557FE1D3A4210D681581FA5">Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA; Email: dkavanaugh @ calacademy. org; &amp; Corresponding author: David H. Kavanaugh (dkavanaugh @ calacademy. org)</mods:affiliation>
</mods:name>
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<mods:role id="2A164A7BBB25D97EC02EE1518002BE84">
<mods:roleTerm id="AECE1104103060A9B2B79B2E4A0A4AE1">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="41FE4F9208150261FED79C0C6207F6A5">Rainio, Johanna</mods:namePart>
<mods:affiliation id="80AF45948D47E1B5F9B1F73408F55E04">Department of Ecology and Systematics, University of Helsinki, P. O. Box 65, FIN- 00014 Helsinki, Finland; Email: kjohannarainio @ gmail. com</mods:affiliation>
</mods:name>
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<mods:title id="4E05E7AE6C6ABFF2AAE5DF061481ABD8">Proceedings of the California Academy of Sciences</mods:title>
</mods:titleInfo>
<mods:part id="94B05819732DA00E2563425A3225277B">
<mods:date id="F45E3AB29ED7E98B10A939D7FE1D7843">2016</mods:date>
<mods:detail id="F1A63511F5C07169A9E803D1BCB664CC" type="pubDate">
<mods:number id="201FAFB43A46BDAD8E68A99E74B08237">2016-04-29</mods:number>
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<mods:detail id="9847B432EAFB61B32E16CC1CC892CD57" type="volume">
<mods:number id="6927F7AC752AC827F9CBDB9755E08CBB">63</mods:number>
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<mods:number id="7A0EEA3F21E6A3C108D84FAD4E0967A8">7</mods:number>
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<mods:classification id="D40CE7FF001238834829FFE82D0D7FBF">journal article</mods:classification>
<mods:identifier id="7AD6CF62EF09D76B2653C6E2FA29F546" type="DOI">10.5281/zenodo.13799439</mods:identifier>
<mods:identifier id="0F8B1C8819749B2370DC6DA4F1629EA2" type="ISSN">0068-547X</mods:identifier>
<mods:identifier id="E998FAAD6A8A53720543599E99C920D7" type="Zenodo-Dep">13799439</mods:identifier>
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<treatment id="0A3E26573073FF9BFF8EFD4FBCF10BA1" ID-DOI="http://doi.org/10.5281/zenodo.13799445" ID-Zenodo-Dep="13799445" LSID="urn:lsid:plazi:treatment:0A3E26573073FF9BFF8EFD4FBCF10BA1" httpUri="http://treatment.plazi.org/id/0A3E26573073FF9BFF8EFD4FBCF10BA1" lastPageId="11" lastPageNumber="212" pageId="10" pageNumber="211">
<subSubSection id="CA8DC4CA3073FF9AFF8EFD4FBFBA0FED" box="[123,729,733,758]" pageId="10" pageNumber="211" type="nomenclature">
<paragraph id="822897413073FF9AFF8EFD4FBFBA0FED" blockId="10.[123,729,733,788]" box="[123,729,733,758]" pageId="10" pageNumber="211">
<heading id="D960202D3073FF9AFF8EFD4FBFBA0FED" box="[123,729,733,758]" fontSize="10" level="2" pageId="10" pageNumber="211" reason="4">
<emphasis id="B0E34B533073FF9AFF8EFD4FBFBA0FED" bold="true" box="[123,729,733,758]" pageId="10" pageNumber="211">
<taxonomicName id="4597ECC23073FF9AFF8EFD4FBF150FEE" ID-CoL="6C9F6" authority="Kavanaugh and Rainio" authorityName="Kavanaugh and Rainio" authorityYear="2016" box="[123,630,733,758]" class="Insecta" family="Carabidae" genus="Dactyleurys" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="211" phylum="Arthropoda" rank="species" species="minimus" status="sp. nov.">
<emphasis id="B0E34B533073FF9AFF8EFD4FBC070FED" bold="true" box="[123,356,733,758]" italics="true" pageId="10" pageNumber="211">Dactyleurys minimus</emphasis>
Kavanaugh and Rainio
</taxonomicName>
,
<taxonomicNameLabel id="ABD0F6283073FF9AFD76FD4CBFBA0FED" box="[643,729,734,758]" pageId="10" pageNumber="211" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="CA8DC4CA3073FF9AFF8EFD6CBC300E0F" box="[123,339,766,788]" pageId="10" pageNumber="211" type="description">
<paragraph id="822897413073FF9AFF8EFD6CBC300E0F" blockId="10.[123,729,733,788]" box="[123,339,766,788]" pageId="10" pageNumber="211">
<figureCitation id="1AAC8BC43073FF9AFF8EFD6CBDB60E0F" box="[123,213,766,788]" captionStart="FIGURE" captionStartId="11.[155,167,1273,1292]" captionTargetBox="[120,1176,216,1254]" captionTargetId="figure-920@11.[120,1176,216,1255]" captionTargetPageId="11" captionText="FIGURE 6. Digital images of holotype male of Dactyleurys minimus Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B. Labels. Scale line = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13799453" httpUri="https://zenodo.org/record/13799453/files/figure.png" pageId="10" pageNumber="211">Figures 6</figureCitation>
,
<figureCitation id="1AAC8BC43073FF9AFF17FD6CBD9C0E08" box="[226,255,766,787]" captionStart="FIGURE" captionStartId="14.[587,599,1619,1638]" captionTargetBox="[123,1176,216,1715]" captionTargetId="figure-1475@14.[552,1176,216,1601]" captionTargetPageId="14" captionText="FIGURE 8. Digital images of pronota of Dactyleurys spp., dorsal aspect. A. Holotype of D. minimus Kavanaugh &amp; Rainio sp. nov.; B. Holotype of D. ranomafanae Kavanaugh &amp; Rainio sp. nov.; C. D. anomalus Tschitschérine. Scale lines = 1.0 mm" figureDoi="http://doi.org/10.5281/zenodo.13799459" httpUri="https://zenodo.org/record/13799459/files/figure.png" pageId="10" pageNumber="211">8A</figureCitation>
,
<figureCitation id="1AAC8BC43073FF9AFEF9FD6CBC300E0F" box="[268,339,766,788]" captionStart="FIGURE" captionStartId="15.[155,167,1141,1160]" captionTargetBox="[147,1159,229,1126]" captionTargetId="figure-936@15.[120,1176,216,1138]" captionTargetPageId="15" captionText="FIGURE 9. Digital images of aedeagus of male genitalia of Dactyleurys spp. A–C. left lateral aspect; D–F. dorsal aspect. A, D. Holotype of D. minimus Kavanaugh &amp; Rainio sp. nov.; B, E. Holotype of D. ranomafanae Kavanaugh &amp; Rainio sp. nov.; C, F. D. anomalus Tschitschérine. Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13799461" httpUri="https://zenodo.org/record/13799461/files/figure.png" pageId="10" pageNumber="211">9A, 9D</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="CA8DC4CA3073FF9BFF5EFCAABCF10BA1" lastPageId="11" lastPageNumber="212" pageId="10" pageNumber="211" type="materials_examined">
<paragraph id="822897413073FF9AFF5EFCAABC5A0EAA" blockId="10.[122,1174,824,1725]" pageId="10" pageNumber="211">
<emphasis id="B0E34B533073FF9AFF5EFCAABC3D0E55" bold="true" box="[171,350,824,848]" pageId="10" pageNumber="211">
<smallCapsWord id="84CE019D3073FF9AFF5EFCAABD870E54" baselines="842,842" box="[171,228,824,848]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Type" pageId="10" pageNumber="211">TYPE</smallCapsWord>
<smallCapsWord id="84CE019D3073FF9AFF04FCA9BC3D0E55" baselines="841" box="[241,350,827,846]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="material" pageId="10" pageNumber="211">MATERIAL</smallCapsWord>
</emphasis>
.—
<materialsCitation id="32FF9D1C3073FF9AFE73FCAABC5A0EAA" collectingDate="2005-04-12" collectorName="F. Ratalata" country="Madagascar" elevation="130" location="MADAGASCAR" pageId="10" pageNumber="211" specimenCount="2" typeStatus="holotype">
<typeStatus id="5D2C29E33073FF9AFE73FCAABC8A0E4B" box="[390,489,824,848]" pageId="10" pageNumber="211" type="holotype">Holotype</typeStatus>
(
<figureCitation id="1AAC8BC43073FF9AFE0BFCAABFE30E54" box="[510,640,824,848]" captionStart="FIGURE" captionStartId="11.[155,167,1273,1292]" captionTargetBox="[120,1176,216,1254]" captionTargetId="figure-920@11.[120,1176,216,1255]" captionTargetPageId="11" captionText="FIGURE 6. Digital images of holotype male of Dactyleurys minimus Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B. Labels. Scale line = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13799453" httpUri="https://zenodo.org/record/13799453/files/figure.png" pageId="10" pageNumber="211">Figs. 6A–B</figureCitation>
), a male, in CAS, labeled: “
<collectingCountry id="FA80D7D13073FF9AFC24FCAAB9EC0E4B" box="[977,1167,824,848]" name="Madagascar" pageId="10" pageNumber="211">MADAGASCAR</collectingCountry>
, Ranomafana NP, Talatakely, X at
<quantity id="456F3AA43073FF9AFE11FCCBBF4B0E6B" box="[484,552,857,881]" metricMagnitude="2" metricUnit="m" metricValue="1.3" pageId="10" pageNumber="211" unit="m" value="130.0">
<elevation id="09BA70723073FF9AFE11FCCBBF4B0E6B" box="[484,552,857,881]" metricMagnitude="2" metricUnit="m" metricValue="1.3" pageId="10" pageNumber="211" unit="m" value="130.0">130 m</elevation>
</quantity>
mark
<date id="F629B1813073FF9AFD98FCCBBFB40E6B" box="[621,727,856,881]" pageId="10" pageNumber="211" value="2005-04-12">
<collectingDate id="E66D48693073FF9AFD98FCCBBFB40E6B" box="[621,727,856,881]" pageId="10" pageNumber="211" value="2005-04-12">12.4.2005</collectingDate>
</date>
”/ “collected by hand in daytime,
<collectorName id="2F62F2973073FF9AFBB4FCCBBDC20E8A" pageId="10" pageNumber="211">F. Ratalata</collectorName>
&amp; Johanna Rainio collectors”/ “
<typeStatus id="5D2C29E33073FF9AFE0FFCEBBFE40E8A" box="[506,647,889,913]" pageId="10" pageNumber="211" type="holotype">HOLOTYPE</typeStatus>
<taxonomicName id="4597ECC23073FF9AFD78FCEBB9350E8A" authority="Kavanaugh &amp; Rainio" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[653,1110,889,913]" class="Insecta" family="Carabidae" genus="Dactyleurys" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="211" phylum="Arthropoda" rank="species" species="minimus" status="sp. nov.">
<emphasis id="B0E34B533073FF9AFD78FCEBBE0D0E8A" box="[653,878,889,913]" italics="true" pageId="10" pageNumber="211">Dactyleurys minimus</emphasis>
Kavanaugh &amp; Rainio
</taxonomicName>
<taxonomicNameLabel id="ABD0F6283073FF9AFBA8FCEBB9F50E8B" box="[1117,1174,889,912]" pageId="10" pageNumber="211" rank="species">sp. n.</taxonomicNameLabel>
2015” [red label].
</materialsCitation>
</paragraph>
<paragraph id="822897413073FF9AFF5EFC28B9500ECA" blockId="10.[122,1174,824,1725]" box="[171,1075,954,978]" pageId="10" pageNumber="211">
<materialsCitation id="32FF9D1C3073FF9AFF5EFC28B94F0EC9" box="[171,1068,954,978]" country="Madagascar" location="Ranomafana National Park" pageId="10" pageNumber="211" specimenCount="1" stateProvince="Fianarantsoa" typeStatus="holotype">
<emphasis id="B0E34B533073FF9AFF5EFC28BC340ECB" bold="true" box="[171,343,954,978]" pageId="10" pageNumber="211">
<smallCapsWord id="84CE019D3073FF9AFF5EFC28BD870ECA" baselines="972,972" box="[171,228,954,978]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Type" pageId="10" pageNumber="211">
<typeStatus id="5D2C29E33073FF9AFF5EFC28BD870ECA" box="[171,228,954,978]" pageId="10" pageNumber="211">TYPE</typeStatus>
</smallCapsWord>
<smallCapsWord id="84CE019D3073FF9AFF19FC2FBC340ECB" baselines="971" box="[236,343,957,976]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="locality" pageId="10" pageNumber="211">LOCALITY</smallCapsWord>
</emphasis>
.—
<collectingCountry id="FA80D7D13073FF9AFE8BFC28BC9C0EC9" box="[382,511,954,978]" name="Madagascar" pageId="10" pageNumber="211">Madagascar</collectingCountry>
,
<collectingRegion id="405359A33073FF9AFDF8FC28BF9F0EC9" box="[525,764,954,978]" country="Madagascar" name="Fianarantsoa" pageId="10" pageNumber="211">Fianarantsoa Province</collectingRegion>
,
<location id="8748C19A3073FF9AFCFFFC28B94F0EC9" LSID="urn:lsid:plazi:treatment:0A3E26573073FF9BFF8EFD4FBCF10BA1:8748C19A3073FF9AFCFFFC28B94F0EC9" box="[778,1068,954,978]" country="Madagascar" name="Ranomafana National Park" pageId="10" pageNumber="211" stateProvince="Fianarantsoa">Ranomafana National Park</location>
</materialsCitation>
.
</paragraph>
<paragraph id="822897413073FF9AFF5EFC49BFC20928" blockId="10.[122,1174,824,1725]" pageId="10" pageNumber="211">
<emphasis id="B0E34B533073FF9AFF5EFC49BC8F0EEA" bold="true" box="[171,492,987,1011]" pageId="10" pageNumber="211">
<smallCapsWord id="84CE019D3073FF9AFF5EFC49BC4D0EEA" baselines="1005,1004" box="[171,302,987,1011]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Derivation" pageId="10" pageNumber="211">DERIVATION</smallCapsWord>
<smallCapsWord id="84CE019D3073FF9AFEC1FC4CBC330EEA" baselines="1004" box="[308,336,990,1009]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="of" pageId="10" pageNumber="211">OF</smallCapsWord>
<smallCapsWord id="84CE019D3073FF9AFEA0FC4CBCCB0EEA" baselines="1004" box="[341,424,990,1009]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="species" pageId="10" pageNumber="211">SPECIES</smallCapsWord>
<smallCapsWord id="84CE019D3073FF9AFE5AFC4CBC8F0EEA" baselines="1004" box="[431,492,990,1009]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="name" pageId="10" pageNumber="211">NAME</smallCapsWord>
</emphasis>
.— The species epithet,
<taxonomicName id="4597ECC23073FF9AFD18FC49BE240EE9" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[749,839,987,1010]" class="Insecta" family="Carabidae" genus="Dactyleurys" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="211" phylum="Arthropoda" rank="species" species="minimus">
<emphasis id="B0E34B533073FF9AFD18FC49BE240EE9" box="[749,839,987,1010]" italics="true" pageId="10" pageNumber="211">minimus</emphasis>
</taxonomicName>
, is the Latin adjective meaning smallest, a reference to fact that the known adult male of this species is smaller than males of any of the other known species that we have examined.
</paragraph>
<paragraph id="822897413073FF9BFF5EFBAEBF0D08E1" blockId="10.[122,1174,824,1725]" lastBlockId="11.[123,1174,1347,1723]" lastPageId="11" lastPageNumber="212" pageId="10" pageNumber="211">
<smallCapsWord id="84CE019D3073FF9AFF5EFBAEBC210949" baselines="1102,1101" box="[171,322,1084,1108]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Recognition" pageId="10" pageNumber="211">
<emphasis id="B0E34B533073FF9AFF5EFBAEBC210949" bold="true" box="[171,322,1084,1108]" pageId="10" pageNumber="211">RECOGNITION</emphasis>
</smallCapsWord>
.— Smallest adult known for the genus, SBL =
<quantity id="456F3AA43073FF9AFCB2FBAEBEFA0948" box="[839,921,1084,1108]" metricMagnitude="-3" metricUnit="m" metricValue="6.9" pageId="10" pageNumber="211" unit="mm" value="6.9">6.9 mm</quantity>
. The following combination of features identify members of this species as coelostomine pterostichines of the genus
<taxonomicName id="4597ECC23073FF9AFF8EFBEFBD9F098E" authorityName="Tschitscherine" authorityYear="1899" box="[123,252,1149,1173]" class="Insecta" family="Carabidae" genus="Dactyleurys" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="211" phylum="Arthropoda" rank="genus">
<emphasis id="B0E34B533073FF9AFF8EFBEFBD9F098E" box="[123,252,1149,1173]" italics="true" pageId="10" pageNumber="211">Dactyleurys</emphasis>
</taxonomicName>
: antennae with antennomere 3 attached symmetrically or nearly symmetrically to antennomere 2; mentum with deep emargination, apex of median mental tooth distinctly posterior to apices of epilobes; elytron with epipleuron interrupted by internal plica subapically; parascutellar stria not evident; basal setiferious puncture situated at base of stria 3; tarsomeres 1 to 4 of all legs with dense pads of setae ventrally; front and middle tarsomeres 1 to 3 smoothly convex dorsally; male front tarsomeres 1 to 4 wide, dilated, not latero- or medioapically toothed; aedeagus of male with right face dorsal in repose, right paramere conchoid, left paramere reduced, short, apically digitiform. Members of
<taxonomicName id="4597ECC23073FF9AFE35FAF3BF5F0863" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[448,572,1377,1400]" class="Insecta" family="Carabidae" genus="Dactyleurys" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="211" phylum="Arthropoda" rank="species" species="minimus">
<emphasis id="B0E34B533073FF9AFE35FAF3BF5F0863" box="[448,572,1377,1400]" italics="true" pageId="10" pageNumber="211">D. minimus</emphasis>
</taxonomicName>
(
<figureCitation id="1AAC8BC43073FF9AFDBBFAF2BFC10862" box="[590,674,1376,1401]" captionStart="FIGURE" captionStartId="11.[155,167,1273,1292]" captionTargetBox="[120,1176,216,1254]" captionTargetId="figure-920@11.[120,1176,216,1255]" captionTargetPageId="11" captionText="FIGURE 6. Digital images of holotype male of Dactyleurys minimus Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B. Labels. Scale line = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13799453" httpUri="https://zenodo.org/record/13799453/files/figure.png" pageId="10" pageNumber="211">Fig. 6A</figureCitation>
) differ from those of
<taxonomicName id="4597ECC23073FF9AFC61FAF3B97C0863" authorityName="Tschitscherine. Scale" authorityYear="1899" box="[916,1055,1376,1400]" class="Insecta" family="Carabidae" genus="Dactyleurys" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="211" phylum="Arthropoda" rank="species" species="anomalus">
<emphasis id="B0E34B533073FF9AFC61FAF3B97C0863" box="[916,1055,1376,1400]" italics="true" pageId="10" pageNumber="211">D. anomalus</emphasis>
</taxonomicName>
in the following features: smaller size (SBL =
<quantity id="456F3AA43073FF9AFDFEFA13BF3F0883" box="[523,604,1409,1433]" metricMagnitude="-3" metricUnit="m" metricValue="6.9" pageId="10" pageNumber="211" unit="mm" value="6.9">6.9 mm</quantity>
compared with 7.7–9.
<quantity id="456F3AA43073FF9AFCB8FA13BEE80883" box="[845,907,1409,1433]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="10" pageNumber="211" unit="mm" value="4.0">4 mm</quantity>
in
<taxonomicName id="4597ECC23073FF9AFC5BFA10B9540882" authorityName="Tschitscherine. Scale" authorityYear="1899" box="[942,1079,1409,1433]" class="Insecta" family="Carabidae" genus="Dactyleurys" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="211" phylum="Arthropoda" rank="species" species="anomalus">
<emphasis id="B0E34B533073FF9AFC5BFA10B9540882" box="[942,1079,1409,1433]" italics="true" pageId="10" pageNumber="211">D. anomalus</emphasis>
</taxonomicName>
); pronotum (
<figureCitation id="1AAC8BC43073FF9AFF40FA33BC6408A1" box="[181,263,1441,1466]" captionStart="FIGURE" captionStartId="14.[587,599,1619,1638]" captionTargetBox="[123,1176,216,1715]" captionTargetId="figure-1475@14.[552,1176,216,1601]" captionTargetPageId="14" captionText="FIGURE 8. Digital images of pronota of Dactyleurys spp., dorsal aspect. A. Holotype of D. minimus Kavanaugh &amp; Rainio sp. nov.; B. Holotype of D. ranomafanae Kavanaugh &amp; Rainio sp. nov.; C. D. anomalus Tschitschérine. Scale lines = 1.0 mm" figureDoi="http://doi.org/10.5281/zenodo.13799459" httpUri="https://zenodo.org/record/13799459/files/figure.png" pageId="10" pageNumber="211">Fig. 8A</figureCitation>
) relatively narrower, widest at middle (relatively wider (
<figureCitation id="1AAC8BC43073FF9AFC9AFA33BEA208A1" box="[879,961,1441,1466]" captionStart="FIGURE" captionStartId="14.[587,599,1619,1638]" captionTargetBox="[123,1176,216,1715]" captionTargetId="figure-1475@14.[552,1176,216,1601]" captionTargetPageId="14" captionText="FIGURE 8. Digital images of pronota of Dactyleurys spp., dorsal aspect. A. Holotype of D. minimus Kavanaugh &amp; Rainio sp. nov.; B. Holotype of D. ranomafanae Kavanaugh &amp; Rainio sp. nov.; C. D. anomalus Tschitschérine. Scale lines = 1.0 mm" figureDoi="http://doi.org/10.5281/zenodo.13799459" httpUri="https://zenodo.org/record/13799459/files/figure.png" pageId="10" pageNumber="211">Fig. 8C</figureCitation>
) and clearly widest anterior to middle in
<taxonomicName id="4597ECC23073FF9AFEA9FA51BC8608C1" authorityName="Tschitscherine. Scale" authorityYear="1899" box="[348,485,1474,1498]" class="Insecta" family="Carabidae" genus="Dactyleurys" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="211" phylum="Arthropoda" rank="species" species="anomalus">
<emphasis id="B0E34B533073FF9AFEA9FA51BC8608C1" box="[348,485,1474,1498]" italics="true" pageId="10" pageNumber="211">D. anomalus</emphasis>
</taxonomicName>
); elytra faintly but evidently iridescence and elytral microsculpture markedly transverse with sculpticells great than 2.5 times as wide as long (elytra not iridescent and elytral microsculpture moderately transverse with sculpticells less than 2.5 times as wide as long in
<taxonomicName id="4597ECC23073FF9AFF1CF9B6BC120B20" authorityName="Tschitscherine. Scale" authorityYear="1899" box="[233,369,1571,1595]" class="Insecta" family="Carabidae" genus="Dactyleurys" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="211" phylum="Arthropoda" rank="species" species="anomalus">
<emphasis id="B0E34B533073FF9AFF1CF9B6BC120B20" box="[233,369,1571,1595]" italics="true" pageId="10" pageNumber="211">D. anomalus</emphasis>
</taxonomicName>
); and median lobe of male genitalia (
<figureCitation id="1AAC8BC43073FF9AFCF4F9B1BE310B27" box="[769,850,1571,1596]" captionStart="FIGURE" captionStartId="15.[155,167,1141,1160]" captionTargetBox="[147,1159,229,1126]" captionTargetId="figure-936@15.[120,1176,216,1138]" captionTargetPageId="15" captionText="FIGURE 9. Digital images of aedeagus of male genitalia of Dactyleurys spp. A–C. left lateral aspect; D–F. dorsal aspect. A, D. Holotype of D. minimus Kavanaugh &amp; Rainio sp. nov.; B, E. Holotype of D. ranomafanae Kavanaugh &amp; Rainio sp. nov.; C, F. D. anomalus Tschitschérine. Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13799461" httpUri="https://zenodo.org/record/13799461/files/figure.png" pageId="10" pageNumber="211">Fig. 9A</figureCitation>
) with shaft abruptly bent ventrally in apical fourth with apex nearly straight in lateral aspect (shaft nearly straight ventrally (
<figureCitation id="1AAC8BC43073FF9AFB9EF9D6BDF90B66" captionStart="FIGURE" captionStartId="14.[587,599,1619,1638]" captionTargetBox="[123,1176,216,1715]" captionTargetId="figure-1475@14.[552,1176,216,1601]" captionTargetPageId="14" captionText="FIGURE 8. Digital images of pronota of Dactyleurys spp., dorsal aspect. A. Holotype of D. minimus Kavanaugh &amp; Rainio sp. nov.; B. Holotype of D. ranomafanae Kavanaugh &amp; Rainio sp. nov.; C. D. anomalus Tschitschérine. Scale lines = 1.0 mm" figureDoi="http://doi.org/10.5281/zenodo.13799459" httpUri="https://zenodo.org/record/13799459/files/figure.png" pageId="10" pageNumber="211">Fig. 8C</figureCitation>
) in apical fourth with apex slightly recurved dorsally in
<taxonomicName id="4597ECC23073FF9AFD09F9F7BEE50B67" authorityName="Tschitscherine. Scale" authorityYear="1899" box="[764,902,1636,1660]" class="Insecta" family="Carabidae" genus="Dactyleurys" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="211" phylum="Arthropoda" rank="species" species="anomalus">
<emphasis id="B0E34B533073FF9AFD09F9F7BEE50B67" box="[764,902,1636,1660]" italics="true" pageId="10" pageNumber="211">D. anomalus</emphasis>
</taxonomicName>
), apex narrowly rounded in dorsal aspect (
<figureCitation id="1AAC8BC43073FF9AFEC6F917BCE60B87" box="[307,389,1669,1693]" captionStart="FIGURE" captionStartId="15.[155,167,1141,1160]" captionTargetBox="[147,1159,229,1126]" captionTargetId="figure-936@15.[120,1176,216,1138]" captionTargetPageId="15" captionText="FIGURE 9. Digital images of aedeagus of male genitalia of Dactyleurys spp. A–C. left lateral aspect; D–F. dorsal aspect. A, D. Holotype of D. minimus Kavanaugh &amp; Rainio sp. nov.; B, E. Holotype of D. ranomafanae Kavanaugh &amp; Rainio sp. nov.; C, F. D. anomalus Tschitschérine. Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13799461" httpUri="https://zenodo.org/record/13799461/files/figure.png" pageId="10" pageNumber="211">Fig. 9D</figureCitation>
) (more broadly rounded (
<figureCitation id="1AAC8BC43073FF9AFD6BF917BF8F0B87" box="[670,748,1669,1693]" captionStart="FIGURE" captionStartId="15.[155,167,1141,1160]" captionTargetBox="[147,1159,229,1126]" captionTargetId="figure-936@15.[120,1176,216,1138]" captionTargetPageId="15" captionText="FIGURE 9. Digital images of aedeagus of male genitalia of Dactyleurys spp. A–C. left lateral aspect; D–F. dorsal aspect. A, D. Holotype of D. minimus Kavanaugh &amp; Rainio sp. nov.; B, E. Holotype of D. ranomafanae Kavanaugh &amp; Rainio sp. nov.; C, F. D. anomalus Tschitschérine. Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13799461" httpUri="https://zenodo.org/record/13799461/files/figure.png" pageId="10" pageNumber="211">Fig. 9F</figureCitation>
) in
<taxonomicName id="4597ECC23073FF9AFCEDF914BEC10B86" authorityName="Tschitscherine. Scale" authorityYear="1899" box="[792,930,1669,1693]" class="Insecta" family="Carabidae" genus="Dactyleurys" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="211" phylum="Arthropoda" rank="species" species="anomalus">
<emphasis id="B0E34B533073FF9AFCEDF914BEC10B86" box="[792,930,1669,1693]" italics="true" pageId="10" pageNumber="211">D. anomalus</emphasis>
</taxonomicName>
). Members of
<taxonomicName id="4597ECC23073FF9AFBB6F914BDCF0BA6" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" class="Insecta" family="Carabidae" genus="Dactyleurys" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="211" phylum="Arthropoda" rank="species" species="minimus">
<emphasis id="B0E34B533073FF9AFBB6F914BDCF0BA6" italics="true" pageId="10" pageNumber="211">D. minimus</emphasis>
</taxonomicName>
differ from those of the other new species,
<taxonomicName id="4597ECC23073FF9AFD7BF937BEC50BA6" authorityName="Kavanaugh and Rainio" authorityYear="2016" box="[654,934,1701,1725]" class="Insecta" family="Carabidae" genus="Dactyleurys" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="211" phylum="Arthropoda" rank="species" species="ranomafanae">
<emphasis id="B0E34B533073FF9AFD7BF937BEC50BA6" box="[654,934,1701,1725]" italics="true" pageId="10" pageNumber="211">Dactyleurys ranomafanae</emphasis>
</taxonomicName>
, in the following features: smaller size (SBL = 7.7–8.
<quantity id="456F3AA43072FF9BFE11FAD1BF460841" box="[484,549,1347,1371]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="11" pageNumber="212" unit="mm" value="3.0">3 mm</quantity>
in
<taxonomicName id="4597ECC23072FF9BFDBBFAD6BF9C0840" authorityName="Kavanaugh and Rainio" authorityYear="2016" box="[590,767,1347,1371]" class="Insecta" family="Carabidae" genus="Dactyleurys" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="212" phylum="Arthropoda" rank="species" species="ranomafanae">
<emphasis id="B0E34B533072FF9BFDBBFAD6BF9C0840" box="[590,767,1347,1371]" italics="true" pageId="11" pageNumber="212">D. ranomafanae</emphasis>
</taxonomicName>
); pronotum (
<figureCitation id="1AAC8BC43072FF9BFC61FAD1BE8A0840" box="[916,1001,1347,1371]" captionStart="FIGURE" captionStartId="14.[587,599,1619,1638]" captionTargetBox="[123,1176,216,1715]" captionTargetId="figure-1475@14.[552,1176,216,1601]" captionTargetPageId="14" captionText="FIGURE 8. Digital images of pronota of Dactyleurys spp., dorsal aspect. A. Holotype of D. minimus Kavanaugh &amp; Rainio sp. nov.; B. Holotype of D. ranomafanae Kavanaugh &amp; Rainio sp. nov.; C. D. anomalus Tschitschérine. Scale lines = 1.0 mm" figureDoi="http://doi.org/10.5281/zenodo.13799459" httpUri="https://zenodo.org/record/13799459/files/figure.png" pageId="11" pageNumber="212">Fig. 8A</figureCitation>
) relatively narrower and with evenly rounded lateral margin (slightly wider (
<figureCitation id="1AAC8BC43072FF9BFCC5FAF1BEE00861" box="[816,899,1379,1403]" captionStart="FIGURE" captionStartId="14.[587,599,1619,1638]" captionTargetBox="[123,1176,216,1715]" captionTargetId="figure-1475@14.[552,1176,216,1601]" captionTargetPageId="14" captionText="FIGURE 8. Digital images of pronota of Dactyleurys spp., dorsal aspect. A. Holotype of D. minimus Kavanaugh &amp; Rainio sp. nov.; B. Holotype of D. ranomafanae Kavanaugh &amp; Rainio sp. nov.; C. D. anomalus Tschitschérine. Scale lines = 1.0 mm" figureDoi="http://doi.org/10.5281/zenodo.13799459" httpUri="https://zenodo.org/record/13799459/files/figure.png" pageId="11" pageNumber="212">Fig. 8B</figureCitation>
) and with lateral margin slightly straighter in basal one-third in
<taxonomicName id="4597ECC23072FF9BFDD4FA16BFB20880" authorityName="Kavanaugh and Rainio" authorityYear="2016" box="[545,721,1411,1435]" class="Insecta" family="Carabidae" genus="Dactyleurys" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="212" phylum="Arthropoda" rank="species" species="ranomafanae">
<emphasis id="B0E34B533072FF9BFDD4FA16BFB20880" box="[545,721,1411,1435]" italics="true" pageId="11" pageNumber="212">D. ranomafanae</emphasis>
</taxonomicName>
); and median lobe of male genitalia (
<figureCitation id="1AAC8BC43072FF9BFB9EFA11BDF808A0" captionStart="FIGURE" captionStartId="15.[155,167,1141,1160]" captionTargetBox="[147,1159,229,1126]" captionTargetId="figure-936@15.[120,1176,216,1138]" captionTargetPageId="15" captionText="FIGURE 9. Digital images of aedeagus of male genitalia of Dactyleurys spp. A–C. left lateral aspect; D–F. dorsal aspect. A, D. Holotype of D. minimus Kavanaugh &amp; Rainio sp. nov.; B, E. Holotype of D. ranomafanae Kavanaugh &amp; Rainio sp. nov.; C, F. D. anomalus Tschitschérine. Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13799461" httpUri="https://zenodo.org/record/13799461/files/figure.png" pageId="11" pageNumber="212">Fig. 9A</figureCitation>
) with apex nearly straight in lateral aspect (apex slightly recurved dorsally (
<figureCitation id="1AAC8BC43072FF9BFBE0FA31B90808A0" box="[1045,1131,1443,1467]" captionStart="FIGURE" captionStartId="15.[155,167,1141,1160]" captionTargetBox="[147,1159,229,1126]" captionTargetId="figure-936@15.[120,1176,216,1138]" captionTargetPageId="15" captionText="FIGURE 9. Digital images of aedeagus of male genitalia of Dactyleurys spp. A–C. left lateral aspect; D–F. dorsal aspect. A, D. Holotype of D. minimus Kavanaugh &amp; Rainio sp. nov.; B, E. Holotype of D. ranomafanae Kavanaugh &amp; Rainio sp. nov.; C, F. D. anomalus Tschitschérine. Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13799461" httpUri="https://zenodo.org/record/13799461/files/figure.png" pageId="11" pageNumber="212">Fig. 9C</figureCitation>
) in
<taxonomicName id="4597ECC23072FF9BFF8EFA56BC4A08C0" authorityName="Kavanaugh and Rainio" authorityYear="2016" box="[123,297,1475,1499]" class="Insecta" family="Carabidae" genus="Dactyleurys" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="212" phylum="Arthropoda" rank="species" species="ranomafanae">
<emphasis id="B0E34B533072FF9BFF8EFA56BC4A08C0" box="[123,297,1475,1499]" italics="true" pageId="11" pageNumber="212">D. ranomafanae</emphasis>
</taxonomicName>
), apical part of shaft (
<figureCitation id="1AAC8BC43072FF9BFDE2FA51BF0B08C1" box="[535,616,1475,1499]" captionStart="FIGURE" captionStartId="15.[155,167,1141,1160]" captionTargetBox="[147,1159,229,1126]" captionTargetId="figure-936@15.[120,1176,216,1138]" captionTargetPageId="15" captionText="FIGURE 9. Digital images of aedeagus of male genitalia of Dactyleurys spp. A–C. left lateral aspect; D–F. dorsal aspect. A, D. Holotype of D. minimus Kavanaugh &amp; Rainio sp. nov.; B, E. Holotype of D. ranomafanae Kavanaugh &amp; Rainio sp. nov.; C, F. D. anomalus Tschitschérine. Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13799461" httpUri="https://zenodo.org/record/13799461/files/figure.png" pageId="11" pageNumber="212">Fig. 9D</figureCitation>
) slightly deflected right in dorsal aspect (apical part of shaft (
<figureCitation id="1AAC8BC43072FF9BFF29FA71BC4808E1" box="[220,299,1507,1531]" captionStart="FIGURE" captionStartId="15.[155,167,1141,1160]" captionTargetBox="[147,1159,229,1126]" captionTargetId="figure-936@15.[120,1176,216,1138]" captionTargetPageId="15" captionText="FIGURE 9. Digital images of aedeagus of male genitalia of Dactyleurys spp. A–C. left lateral aspect; D–F. dorsal aspect. A, D. Holotype of D. minimus Kavanaugh &amp; Rainio sp. nov.; B, E. Holotype of D. ranomafanae Kavanaugh &amp; Rainio sp. nov.; C, F. D. anomalus Tschitschérine. Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13799461" httpUri="https://zenodo.org/record/13799461/files/figure.png" pageId="11" pageNumber="212">Fig. 9E</figureCitation>
) straight in
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<emphasis id="B0E34B533072FF9BFE5AFA76BF3D08E0" box="[431,606,1507,1531]" italics="true" pageId="11" pageNumber="212">D. ranomafanae</emphasis>
</taxonomicName>
).
</paragraph>
<caption id="D6E8C7C93072FF9BFF6EFB6BBCA8083D" ID-DOI="http://doi.org/10.5281/zenodo.13799453" ID-Zenodo-Dep="13799453" httpUri="https://zenodo.org/record/13799453/files/figure.png" pageId="11" pageNumber="212" startId="11.[155,167,1273,1292]" targetBox="[120,1176,216,1254]" targetPageId="11" targetType="figure">
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6. Digital images of holotype male of
<taxonomicName id="4597ECC23072FF9BFDC5FB6BBEC50817" authority="Kavanaugh &amp; Rainio" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[560,934,1273,1292]" class="Insecta" family="Carabidae" genus="Dactyleurys" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="212" phylum="Arthropoda" rank="species" species="minimus" status="sp. nov.">
<emphasis id="B0E34B533072FF9BFDC5FB6BBF850810" box="[560,742,1273,1292]" italics="true" pageId="11" pageNumber="212">Dactyleurys minimus</emphasis>
Kavanaugh &amp; Rainio
</taxonomicName>
<taxonomicNameLabel id="ABD0F6283072FF9BFC58FB6BBE930817" box="[941,1008,1273,1292]" pageId="11" pageNumber="212" rank="species">sp. nov.</taxonomicNameLabel>
A. Habitus, dorsal aspect; B. Labels. Scale line = 1.0 mm.
</paragraph>
</caption>
<paragraph id="822897413072FF9BFF5EF990B9690B01" blockId="11.[123,1174,1347,1723]" box="[171,1034,1538,1563]" pageId="11" pageNumber="212">
<emphasis id="B0E34B533072FF9BFF5EF990BC920B02" bold="true" box="[171,497,1538,1562]" pageId="11" pageNumber="212">
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</emphasis>
.— At present, known only from the
<typeStatus id="5D2C29E33072FF9BFC8AF991BECE0B00" box="[895,941,1539,1563]" pageId="11" pageNumber="212">type</typeStatus>
locality.
</paragraph>
<paragraph id="822897413072FF9BFF5EF9B1BCF10BA1" blockId="11.[123,1174,1347,1723]" pageId="11" pageNumber="212">
<emphasis id="B0E34B533072FF9BFF5EF9B1BCC60B22" bold="true" box="[171,421,1571,1595]" pageId="11" pageNumber="212">
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<smallCapsWord id="84CE019D3072FF9BFEE4F9B4BCC60B22" baselines="1588" box="[273,421,1574,1593]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="distribution" pageId="11" pageNumber="212">DISTRIBUTION</smallCapsWord>
</emphasis>
.— The unique
<typeStatus id="5D2C29E33072FF9BFDBBF9B1BFC80B20" box="[590,683,1571,1595]" pageId="11" pageNumber="212" type="holotype">holotype</typeStatus>
was collected by hand in daytime in secondary montane rain forest in the Talatakely area, at an elevation of
<quantity id="456F3AA43072FF9BFCEAF9D1BE010B41" box="[799,866,1603,1627]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="11" pageNumber="212" unit="m" value="900.0">900 m</quantity>
, at the
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mark along Trail “X”. Vegetation in that area consisted of bamboo, small young trees and some larger trees with diameters at breast-height (DBH) of
<quantity id="456F3AA43072FF9BFDC8F911BFCB0B81" box="[573,680,1667,1691]" metricMagnitude="-1" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="11" pageNumber="212" unit="cm" value="25.0" valueMax="30.0" valueMin="20.0">20–30 cm</quantity>
. Tree trunks and branches were heavily laden with epiphytes and lianas.
</paragraph>
</subSubSection>
</treatment>
</document>
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