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<document id="D02926ED01975E456B2CCA5E8B19E41E" ID-DOI="10.5281/zenodo.13799439" ID-ISSN="0068-547X" ID-Zenodo-Dep="13799439" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722538212676" checkinUser="felipe" docAuthor="Kavanaugh, David H. &amp; Rainio, Johanna" docDate="2016" docId="0A3E2657306EFF87FF8EFF49BEB4082C" docLanguage="en" docName="ProcCalifAcadSci.63.7.201-268.pdf" docOrigin="Proceedings of the California Academy of Sciences 63 (7)" docStyle="DocumentStyle:E411DA009AF5E7F9A7E023431E2B7D5B.2:ProcCalifAcadSci.2015-.journal_article" docStyleId="E411DA009AF5E7F9A7E023431E2B7D5B" docStyleName="ProcCalifAcadSci.2015-.journal_article" docStyleVersion="2" docTitle="Omphreoides ranomafanae Kavanaugh and Rainio 2016, sp. nov." docType="treatment" docVersion="2" lastPageNumber="224" masterDocId="F6075E2F3079FF90FFF5FF92BD630D1B" masterDocTitle="Twenty-six New Species of Predaceous Ground Beetles (Coleoptera: Adephaga: Carabidae) from Ranomafana National Park, Madagascar" masterLastPageNumber="268" masterPageNumber="201" pageNumber="224" updateTime="1726771873529" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="6A64393F7815B8B726F542EFB44571D0" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="4FDD1B2ADAC30A7563814CFFCA371C2D">Twenty-six New Species of Predaceous Ground Beetles (Coleoptera: Adephaga: Carabidae) from Ranomafana National Park, Madagascar</mods:title>
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<mods:name id="8AD447F5F72A6FC43B8430E35E5464FB" type="personal">
<mods:role id="A312C2E8A692C7F04DFCBE7634859337">
<mods:roleTerm id="0FB4912EEA936FACC55D818C3B3F74FA">Author</mods:roleTerm>
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<mods:namePart id="1A54D1300D436942B7D285E9594DC22B">Kavanaugh, David H.</mods:namePart>
<mods:affiliation id="68A512EFF977B5E811764B15C88943D9">Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA; Email: dkavanaugh @ calacademy. org; &amp; Corresponding author: David H. Kavanaugh (dkavanaugh @ calacademy. org)</mods:affiliation>
</mods:name>
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<mods:role id="1BB26A999DFA632D8EAB97DC777280D1">
<mods:roleTerm id="D8123562AECE50C4D0C4F90F4FA1283D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="27AD843D62F4FCEDFBF34692FF81E9F7">Rainio, Johanna</mods:namePart>
<mods:affiliation id="700D020A635E47FFBD317FF8426B0BDF">Department of Ecology and Systematics, University of Helsinki, P. O. Box 65, FIN- 00014 Helsinki, Finland; Email: kjohannarainio @ gmail. com</mods:affiliation>
</mods:name>
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<mods:title id="0A5CF2EC43A70A8066EC446EF9A95EBC">Proceedings of the California Academy of Sciences</mods:title>
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<mods:part id="16868557BDD4963724A08DCDA9253E28">
<mods:date id="6371ECF0F4F91606F141BA0A3F1A18BC">2016</mods:date>
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<mods:number id="3532D49D037E70329CFEA777F742EA19">2016-04-29</mods:number>
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<mods:number id="A476FD2C067E96A9005C773108985158">63</mods:number>
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<mods:number id="2172B713BD5FCD88BDE2C2EFCF1B4E0A">7</mods:number>
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<mods:classification id="A3486B7AC430609714CB437FEE4EBFBD">journal article</mods:classification>
<mods:identifier id="A0C6D25BCC6D8C46933E612AE311AB74" type="DOI">10.5281/zenodo.13799439</mods:identifier>
<mods:identifier id="E37B1601F4D007B35523E90A23E38E0C" type="ISSN">0068-547X</mods:identifier>
<mods:identifier id="FBB54C1FC53E4CC2E414FB24F46F06DE" type="Zenodo-Dep">13799439</mods:identifier>
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<treatment id="0A3E2657306EFF87FF8EFF49BEB4082C" ID-DOI="http://doi.org/10.5281/zenodo.13799479" ID-Zenodo-Dep="13799479" LSID="urn:lsid:plazi:treatment:0A3E2657306EFF87FF8EFF49BEB4082C" httpUri="http://treatment.plazi.org/id/0A3E2657306EFF87FF8EFF49BEB4082C" lastPageNumber="224" pageId="23" pageNumber="224">
<subSubSection id="CA8DC4CA306EFF87FF8EFF49BE7D0DE8" box="[123,798,219,243]" pageId="23" pageNumber="224" type="nomenclature">
<paragraph id="82289741306EFF87FF8EFF49BE7D0DE8" blockId="23.[123,798,219,273]" box="[123,798,219,243]" pageId="23" pageNumber="224">
<heading id="D960202D306EFF87FF8EFF49BE7D0DE8" box="[123,798,219,243]" fontSize="10" level="2" pageId="23" pageNumber="224" reason="4">
<emphasis id="B0E34B53306EFF87FF8EFF49BE7D0DE8" bold="true" box="[123,798,219,243]" pageId="23" pageNumber="224">
<taxonomicName id="4597ECC2306EFF87FF8EFF49BFD90DE8" ID-CoL="49FB4" authority="Kavanaugh and Rainio" authorityName="Kavanaugh and Rainio" authorityYear="2016" box="[123,698,219,243]" class="Insecta" family="Carabidae" genus="Omphreoides" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="species" species="ranomafanae" status="sp. nov.">
<emphasis id="B0E34B53306EFF87FF8EFF49BCCA0DE8" bold="true" box="[123,425,219,243]" italics="true" pageId="23" pageNumber="224">Omphreoides ranomafanae</emphasis>
Kavanaugh and Rainio
</taxonomicName>
,
<taxonomicNameLabel id="ABD0F628306EFF87FD32FF49BE7D0DE8" box="[711,798,219,243]" pageId="23" pageNumber="224" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="CA8DC4CA306EFF87FF8EFF69BDBB0C0B" box="[123,216,251,273]" pageId="23" pageNumber="224" type="description">
<paragraph id="82289741306EFF87FF8EFF69BDBB0C0B" blockId="23.[123,798,219,273]" box="[123,216,251,273]" pageId="23" pageNumber="224">
<figureCitation id="1AAC8BC4306EFF87FF8EFF69BDBB0C0B" box="[123,216,251,273]" captionStart="FIGURE" captionStartId="24.[155,167,1561,1580]" captionTargetBox="[120,1176,216,1539]" captionTargetId="figure-53@24.[120,1176,216,1539]" captionTargetPageId="24" captionText="FIGURE 13. Digital images of holotype female of Omphreoides ranomafanae Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B. Labels; C. Forebody, dorsal aspect. Scale lines A = 1.0 mm, C–D = 0.25 mm" figureDoi="http://doi.org/10.5281/zenodo.13799471" httpUri="https://zenodo.org/record/13799471/files/figure.png" pageId="23" pageNumber="224">Figure 13</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="CA8DC4CA306EFF87FF5EFEA7BEB4082C" pageId="23" pageNumber="224" type="materials_examined">
<paragraph id="82289741306EFF87FF5EFEA7BF060CB6" blockId="23.[123,1174,309,1336]" pageId="23" pageNumber="224">
<emphasis id="B0E34B53306EFF87FF5EFEA7BC3B0C50" bold="true" box="[171,344,309,333]" pageId="23" pageNumber="224">
<smallCapsWord id="84CE019D306EFF87FF5EFEA7BD870C57" baselines="327,327" box="[171,228,309,333]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Type" pageId="23" pageNumber="224">TYPE</smallCapsWord>
<smallCapsWord id="84CE019D306EFF87FF1EFEAABC3B0C50" baselines="326" box="[235,344,312,331]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="material" pageId="23" pageNumber="224">MATERIAL</smallCapsWord>
</emphasis>
.—
<materialsCitation id="32FF9D1C306EFF87FE8AFEA7BF060CB6" collectingDate="1990-02-17" collectingDateMax="1990-02-22" collectingDateMin="1990-02-17" collectorName="W. E. Steiner" country="Madagascar" elevation="900" location="Ranomafana" pageId="23" pageNumber="224" specimenCount="2" stateProvince="Fianarantsoa" typeStatus="holotype">
<typeStatus id="5D2C29E3306EFF87FE8AFEA7BC810C56" box="[383,482,309,333]" pageId="23" pageNumber="224" type="holotype">Holotype</typeStatus>
(
<figureCitation id="1AAC8BC4306EFF87FE07FEA7BF100C57" box="[498,627,309,333]" captionStart="FIGURE" captionStartId="24.[155,167,1561,1580]" captionTargetBox="[120,1176,216,1539]" captionTargetId="figure-53@24.[120,1176,216,1539]" captionTargetPageId="24" captionText="FIGURE 13. Digital images of holotype female of Omphreoides ranomafanae Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B. Labels; C. Forebody, dorsal aspect. Scale lines A = 1.0 mm, C–D = 0.25 mm" figureDoi="http://doi.org/10.5281/zenodo.13799471" httpUri="https://zenodo.org/record/13799471/files/figure.png" pageId="23" pageNumber="224">Figs.13A–B</figureCitation>
), a female, in NMNH, labeled: “
<collectingCountry id="FA80D7D1306EFF87FC25FEA7B9ED0C56" box="[976,1166,309,333]" name="Madagascar" pageId="23" pageNumber="224">MADAGASCAR</collectingCountry>
: Prov.
<collectingRegion id="405359A3306EFF87FF4EFEC7BC200C76" box="[187,323,341,365]" country="Madagascar" name="Fianarantsoa" pageId="23" pageNumber="224">Fianarantsoa</collectingRegion>
,
<quantity id="456F3AA4306EFF87FEA7FEC4BCE80C76" box="[338,395,341,365]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="23" pageNumber="224" unit="km" value="7.0">7 km</quantity>
W
<location id="8748C19A306EFF87FE41FEC7BF580C76" LSID="urn:lsid:plazi:treatment:0A3E2657306EFF87FF8EFF49BEB4082C:8748C19A306EFF87FE41FEC7BF580C76" box="[436,571,341,365]" country="Madagascar" name="Ranomafana" pageId="23" pageNumber="224" stateProvince="Fianarantsoa">Ranomafana</location>
,
<quantity id="456F3AA4306EFF87FDBFFEC7BFF30C76" box="[586,656,341,365]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="23" pageNumber="224" unit="m" value="900.0">
<elevation id="09BA7072306EFF87FDBFFEC7BFF30C76" box="[586,656,341,365]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="23" pageNumber="224" unit="m" value="900.0">900 m</elevation>
</quantity>
<date id="F629B181306EFF87FD6CFEC7BE1D0C76" box="[665,894,341,365]" pageId="23" pageNumber="224" value="1990-02-17" valueMax="1990-02-22" valueMin="1990-02-17">
<collectingDate id="E66D4869306EFF87FD6CFEC7BE1D0C76" box="[665,894,341,365]" pageId="23" pageNumber="224" value="1990-02-17" valueMax="1990-02-22" valueMin="1990-02-17">17-22 February 1990</collectingDate>
</date>
<collectorName id="2F62F297306EFF87FC73FEC4B97B0C76" box="[902,1048,341,365]" pageId="23" pageNumber="224">W. E. Steiner</collectorName>
”/ “at black light in montane rainforest near river and stream”/ “
<typeStatus id="5D2C29E3306EFF87FD26FEE4BE030C95" box="[723,864,374,398]" pageId="23" pageNumber="224" type="holotype">HOLOTYPE</typeStatus>
<taxonomicName id="4597ECC2306EFF87FC99FEE4BC3C0CB5" authority="Kavanaugh &amp; Rainio" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" class="Insecta" family="Carabidae" genus="Omphreoides" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="species" species="ranomafanae" status="sp. nov.">
<emphasis id="B0E34B53306EFF87FC99FEE4B9F60C95" box="[876,1173,374,398]" italics="true" pageId="23" pageNumber="224">Omphreoides ranomafanae</emphasis>
Kavanaugh &amp; Rainio
</taxonomicName>
<taxonomicNameLabel id="ABD0F628306EFF87FE93FE04BCC30CB6" box="[358,416,406,429]" pageId="23" pageNumber="224" rank="species">sp. n.</taxonomicNameLabel>
2015” [red label].
</materialsCitation>
</paragraph>
<paragraph id="82289741306EFF87FF5EFE24B9500CD5" blockId="23.[123,1174,309,1336]" box="[171,1075,438,462]" pageId="23" pageNumber="224">
<materialsCitation id="32FF9D1C306EFF87FF5EFE24B94F0CD5" box="[171,1068,438,462]" country="Madagascar" location="Ranomafana National Park" pageId="23" pageNumber="224" specimenCount="1" stateProvince="Fianarantsoa" typeStatus="holotype">
<emphasis id="B0E34B53306EFF87FF5EFE24BC340CD6" bold="true" box="[171,343,438,462]" pageId="23" pageNumber="224">
<smallCapsWord id="84CE019D306EFF87FF5EFE24BD870CD6" baselines="456,456" box="[171,228,438,462]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Type" pageId="23" pageNumber="224">
<typeStatus id="5D2C29E3306EFF87FF5EFE24BD870CD6" box="[171,228,438,462]" pageId="23" pageNumber="224">TYPE</typeStatus>
</smallCapsWord>
<smallCapsWord id="84CE019D306EFF87FF19FE28BC340CD6" baselines="456" box="[236,343,442,461]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="locality" pageId="23" pageNumber="224">LOCALITY</smallCapsWord>
</emphasis>
.—
<collectingCountry id="FA80D7D1306EFF87FE8BFE24BC9C0CD5" box="[382,511,438,462]" name="Madagascar" pageId="23" pageNumber="224">Madagascar</collectingCountry>
,
<collectingRegion id="405359A3306EFF87FDF8FE24BF9F0CD5" box="[525,764,438,462]" country="Madagascar" name="Fianarantsoa" pageId="23" pageNumber="224">Fianarantsoa Province</collectingRegion>
,
<location id="8748C19A306EFF87FCFFFE24B94F0CD5" LSID="urn:lsid:plazi:treatment:0A3E2657306EFF87FF8EFF49BEB4082C:8748C19A306EFF87FCFFFE24B94F0CD5" box="[778,1068,438,462]" country="Madagascar" name="Ranomafana National Park" pageId="23" pageNumber="224" stateProvince="Fianarantsoa">Ranomafana National Park</location>
</materialsCitation>
.
</paragraph>
<paragraph id="82289741306EFF87FF5EFE45BECA0F15" blockId="23.[123,1174,309,1336]" pageId="23" pageNumber="224">
<emphasis id="B0E34B53306EFF87FF5EFE45BC910CF6" bold="true" box="[171,498,471,495]" pageId="23" pageNumber="224">
<smallCapsWord id="84CE019D306EFF87FF5EFE45BC4D0CF6" baselines="489,488" box="[171,302,471,495]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Derivation" pageId="23" pageNumber="224">DERIVATION</smallCapsWord>
<smallCapsWord id="84CE019D306EFF87FEC3FE48BC310CF6" baselines="488" box="[310,338,474,493]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="of" pageId="23" pageNumber="224">OF</smallCapsWord>
<smallCapsWord id="84CE019D306EFF87FEACFE48BCCF0CF6" baselines="488" box="[345,428,474,493]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="species" pageId="23" pageNumber="224">SPECIES</smallCapsWord>
<smallCapsWord id="84CE019D306EFF87FE40FE48BC910CF6" baselines="488" box="[437,498,474,493]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="name" pageId="23" pageNumber="224">NAME</smallCapsWord>
</emphasis>
.— The species epithet,
<taxonomicName id="4597ECC2306EFF87FD0CFE45BEE40CF4" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[761,903,471,495]" class="Insecta" family="Carabidae" genus="Omphreoides" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="species" species="ranomafanae">
<emphasis id="B0E34B53306EFF87FD0CFE45BEE40CF4" box="[761,903,471,495]" italics="true" pageId="23" pageNumber="224">ranomafanae</emphasis>
</taxonomicName>
, is a noun in apposition, derived from the name of the national park in which the
<typeStatus id="5D2C29E3306EFF87FD29FE65BE690F14" box="[732,778,503,527]" pageId="23" pageNumber="224">type</typeStatus>
was collected.
</paragraph>
<paragraph id="82289741306EFF87FF5EFD85B94909CD" blockId="23.[123,1174,309,1336]" pageId="23" pageNumber="224">
<smallCapsWord id="84CE019D306EFF87FF5EFD85BC210F35" baselines="553,553" box="[171,322,535,559]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Recognition" pageId="23" pageNumber="224">
<emphasis id="B0E34B53306EFF87FF5EFD85BC210F35" bold="true" box="[171,322,535,559]" pageId="23" pageNumber="224">RECOGNITION</emphasis>
</smallCapsWord>
.— Size larger than average for genus, SBL =
<quantity id="456F3AA4306EFF87FCB4FD85BEC00F34" box="[833,931,535,559]" metricMagnitude="-2" metricUnit="m" metricValue="1.34" pageId="23" pageNumber="224" unit="mm" value="13.4">13.4 mm</quantity>
. The unique
<typeStatus id="5D2C29E3306EFF87FBCDFD85B9F60F34" box="[1080,1173,535,559]" pageId="23" pageNumber="224" type="holotype">holotype</typeStatus>
female of
<taxonomicName id="4597ECC2306EFF87FF04FDAABCC60F4B" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[241,421,568,592]" class="Insecta" family="Carabidae" genus="Omphreoides" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="species" species="ranomafanae">
<emphasis id="B0E34B53306EFF87FF04FDAABCC60F4B" box="[241,421,568,592]" italics="true" pageId="23" pageNumber="224">O. ranomafanae</emphasis>
</taxonomicName>
(
<figureCitation id="1AAC8BC4306EFF87FE4FFDAABF7E0F4B" box="[442,541,568,592]" captionStart="FIGURE" captionStartId="24.[155,167,1561,1580]" captionTargetBox="[120,1176,216,1539]" captionTargetId="figure-53@24.[120,1176,216,1539]" captionTargetPageId="24" captionText="FIGURE 13. Digital images of holotype female of Omphreoides ranomafanae Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B. Labels; C. Forebody, dorsal aspect. Scale lines A = 1.0 mm, C–D = 0.25 mm" figureDoi="http://doi.org/10.5281/zenodo.13799471" httpUri="https://zenodo.org/record/13799471/files/figure.png" pageId="23" pageNumber="224">Fig. 13A</figureCitation>
) is clearly unlike members of any described species of
<taxonomicName id="4597ECC2306EFF87FF8EFDCABC690F6B" baseAuthorityName="Lorenz" baseAuthorityYear="2005" box="[123,266,600,624]" class="Insecta" family="Carabidae" genus="Omphreoides" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="genus">
<emphasis id="B0E34B53306EFF87FF8EFDCABC690F6B" box="[123,266,600,624]" italics="true" pageId="23" pageNumber="224">Omphreoides</emphasis>
</taxonomicName>
, with several features intermediate between the four typical species and
<taxonomicName id="4597ECC2306EFF87FBDAFDCAB9EC0F6B" authorityName="Alluaud" authorityYear="1910" box="[1071,1167,600,624]" class="Insecta" family="Carabidae" genus="Omphreoides" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="species" species="quodi">
<emphasis id="B0E34B53306EFF87FBDAFDCAB9EC0F6B" box="[1071,1167,600,624]" italics="true" pageId="23" pageNumber="224">O. quodi</emphasis>
</taxonomicName>
. The head of the
<typeStatus id="5D2C29E3306EFF87FED8FDEABCE90F8B" box="[301,394,632,656]" pageId="23" pageNumber="224" type="holotype">holotype</typeStatus>
(
<figureCitation id="1AAC8BC4306EFF87FE6EFDEABC990F8A" box="[411,506,632,657]" captionStart="FIGURE" captionStartId="24.[155,167,1561,1580]" captionTargetBox="[120,1176,216,1539]" captionTargetId="figure-53@24.[120,1176,216,1539]" captionTargetPageId="24" captionText="FIGURE 13. Digital images of holotype female of Omphreoides ranomafanae Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B. Labels; C. Forebody, dorsal aspect. Scale lines A = 1.0 mm, C–D = 0.25 mm" figureDoi="http://doi.org/10.5281/zenodo.13799471" httpUri="https://zenodo.org/record/13799471/files/figure.png" pageId="23" pageNumber="224">Fig. 13C</figureCitation>
) is longer and more slender than that of females of any other species except
<taxonomicName id="4597ECC2306EFF87FEEFFD0BBC140FAA" authorityName="Alluaud" authorityYear="1910" box="[282,375,665,689]" class="Insecta" family="Carabidae" genus="Omphreoides" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="species" species="quodi">
<emphasis id="B0E34B53306EFF87FEEFFD0BBC140FAA" box="[282,375,665,689]" italics="true" pageId="23" pageNumber="224">O. quodi</emphasis>
</taxonomicName>
, which has an even longer and narrower head (see Jeannel 1949, Fig. 364). In
<taxonomicName id="4597ECC2306EFF87FF62FD28BC260FCA" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[151,325,697,721]" class="Insecta" family="Carabidae" genus="Omphreoides" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="species" species="ranomafanae">
<emphasis id="B0E34B53306EFF87FF62FD28BC260FCA" box="[151,325,697,721]" italics="true" pageId="23" pageNumber="224">O. ranomafanae</emphasis>
</taxonomicName>
, the tempora are straight, parallel in anterior their two-thirds, slightly more than twice as long as the diameter of eye, distinctly delimited posteriorly by narrowly round angles. The dorsal longitudinal grooves typical of all members of this genus, except those of
<taxonomicName id="4597ECC2306EFF87FBF2FD68BCF00E29" authority="Fairmaire (1896)" authorityName="Fairmaire" authorityYear="1896" class="Insecta" family="Carabidae" genus="Omphreoides" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="species" species="bispinus">
<emphasis id="B0E34B53306EFF87FBF2FD68BDB70E29" italics="true" pageId="23" pageNumber="224">Omphreoides bispinus</emphasis>
<bibRefCitation id="E606EAB0306EFF87FF29FC88BCF00E29" author="FAIRMAIRE, L." box="[220,403,794,818]" pageId="23" pageNumber="224" pagination="48 - 50" refId="ref30287" refString="FAIRMAIRE, L. 1896. Description de six Carabiques de la region Malgache. Bulletin de la Societe Entomologique de France 65: 48 - 50." type="journal article" year="1896">Fairmaire (1896)</bibRefCitation>
</taxonomicName>
, are extended as sharply-defined grooves only about halfway from the point of insertion of the anterior supraorbital setae to that of posterior supraorbital setae and not continued on to base of the head as they are in
<taxonomicName id="4597ECC2306EFF87FD67FCC9B9060E68" authority="Alluaud (1899 a)" authorityName="Alluaud" authorityYear="1899" box="[658,1125,859,883]" class="Insecta" family="Carabidae" genus="Omphreoides" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="species" species="bucculentus">
<emphasis id="B0E34B53306EFF87FD67FCC9BECA0E68" box="[658,937,859,883]" italics="true" pageId="23" pageNumber="224">Omphreoides bucculentus</emphasis>
<bibRefCitation id="E606EAB0306EFF87FC47FCC9B9060E68" author="ALLUAUD, C." box="[946,1125,859,883]" pageId="23" pageNumber="224" pagination="410 - 413" refId="ref28748" refString="ALLUAUD, C. 1899 a. Sur quelques Coleoptera de Madagascar de famille Carabiques, recueillis par M. Mocquerys et acquis par le Museum. Bulletin du Museum National d'Histoire Naturelle 5: 410 - 413." type="journal article" year="1899">Alluaud (1899a)</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4597ECC2306EFF87FF8EFCE9BF7D0E88" authority="Alluaud (1936)" authorityName="Alluaud" authorityYear="1936" box="[123,542,891,915]" class="Insecta" family="Carabidae" genus="Omphreoides" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="species" species="distinctus">
<emphasis id="B0E34B53306EFF87FF8EFCE9BC150E88" box="[123,374,891,915]" italics="true" pageId="23" pageNumber="224">Omphreoides distinctus</emphasis>
Alluaud (1936)
</taxonomicName>
. Posterior to the sharply-defined segments, broad and shallow depressions extend posteriorly, arcuately convergent and nearly joined in the midline near the back of the head. The pronotum (
<figureCitation id="1AAC8BC4306EFF87FE2BFC2EBF5F0ECF" box="[478,572,956,980]" captionStart="FIGURE" captionStartId="24.[155,167,1561,1580]" captionTargetBox="[120,1176,216,1539]" captionTargetId="figure-53@24.[120,1176,216,1539]" captionTargetPageId="24" captionText="FIGURE 13. Digital images of holotype female of Omphreoides ranomafanae Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B. Labels; C. Forebody, dorsal aspect. Scale lines A = 1.0 mm, C–D = 0.25 mm" figureDoi="http://doi.org/10.5281/zenodo.13799471" httpUri="https://zenodo.org/record/13799471/files/figure.png" pageId="23" pageNumber="224">Fig. 13C</figureCitation>
) is subquadrate, longer than wide (ratio PW/PL = 0.75), with a wavy diagonal row of sparse, coarse punctures (seen also in
<taxonomicName id="4597ECC2306EFF87FCC8FC4FBEA00EEF" authorityName="Alluaud" authorityYear="1936" box="[829,963,988,1012]" class="Insecta" family="Carabidae" genus="Omphreoides" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="species" species="distinctus">
<emphasis id="B0E34B53306EFF87FCC8FC4FBEA00EEF" box="[829,963,988,1012]" italics="true" pageId="23" pageNumber="224">O. distinctus</emphasis>
</taxonomicName>
and
<taxonomicName id="4597ECC2306EFF87FC00FC4FB9310EEF" authorityName="Alluaud" authorityYear="1910" box="[1013,1106,988,1012]" class="Insecta" family="Carabidae" genus="Omphreoides" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="species" species="quodi">
<emphasis id="B0E34B53306EFF87FC00FC4FB9310EEF" box="[1013,1106,988,1012]" italics="true" pageId="23" pageNumber="224">O. quodi</emphasis>
</taxonomicName>
members but absent from those of
<taxonomicName id="4597ECC2306EFF87FE3EFC6FBF0D090E" authorityName="Alluaud" authorityYear="1899" box="[459,622,1021,1045]" class="Insecta" family="Carabidae" genus="Omphreoides" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="species" species="bucculentus">
<emphasis id="B0E34B53306EFF87FE3EFC6FBF0D090E" box="[459,622,1021,1045]" italics="true" pageId="23" pageNumber="224">O. bucculentus</emphasis>
</taxonomicName>
and
<taxonomicName id="4597ECC2306EFF87FD5CFC6FBE45090E" authorityName="Fairmaire" authorityYear="1896" box="[681,806,1021,1045]" class="Insecta" family="Carabidae" genus="Omphreoides" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="species" species="bispinus">
<emphasis id="B0E34B53306EFF87FD5CFC6FBE45090E" box="[681,806,1021,1045]" italics="true" pageId="23" pageNumber="224">O. bispinus</emphasis>
</taxonomicName>
) extended from near the anterior transverse impression paramedially to the basal foveae posteriorly; lateral pronotal margins with short but distinct sinuations anterior to slightly obtuse hind angles; median longitudinal impression deep, wider than in all other
<taxonomicName id="4597ECC2306EFF87FE5EFBCCBF59096D" baseAuthorityName="Lorenz" baseAuthorityYear="2005" box="[427,570,1118,1142]" class="Insecta" family="Carabidae" genus="Omphreoides" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="genus">
<emphasis id="B0E34B53306EFF87FE5EFBCCBF59096D" box="[427,570,1118,1142]" italics="true" pageId="23" pageNumber="224">Omphreoides</emphasis>
</taxonomicName>
except
<taxonomicName id="4597ECC2306EFF87FD78FBCCBF89096D" authorityName="Alluaud" authorityYear="1910" box="[653,746,1118,1142]" class="Insecta" family="Carabidae" genus="Omphreoides" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="species" species="quodi">
<emphasis id="B0E34B53306EFF87FD78FBCCBF89096D" box="[653,746,1118,1142]" italics="true" pageId="23" pageNumber="224">O. quodi</emphasis>
</taxonomicName>
(in which the impression is even wider) and sparsely but coarsely punctate; lateral explanation narrow (narrower at middle than that in
<taxonomicName id="4597ECC2306EFF87FF8EFB0DBC7909AD" authorityName="Alluaud" authorityYear="1899" box="[123,282,1182,1206]" class="Insecta" family="Carabidae" genus="Omphreoides" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="species" species="bucculentus">
<emphasis id="B0E34B53306EFF87FF8EFB0DBC7909AD" box="[123,282,1182,1206]" italics="true" pageId="23" pageNumber="224">O. bucculentus</emphasis>
</taxonomicName>
members and similar to that in
<taxonomicName id="4597ECC2306EFF87FD92FB0DBF8E09AD" authorityName="Alluaud" authorityYear="1936" box="[615,749,1182,1206]" class="Insecta" family="Carabidae" genus="Omphreoides" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="species" species="distinctus">
<emphasis id="B0E34B53306EFF87FD92FB0DBF8E09AD" box="[615,749,1182,1206]" italics="true" pageId="23" pageNumber="224">O. distinctus</emphasis>
</taxonomicName>
member. The elytra are relatively wider than in all other species; each elytron with a straight apical spine in line with interval 3.
</paragraph>
<paragraph id="82289741306EFF87FF5EFB4DB96909EC" blockId="23.[123,1174,309,1336]" box="[171,1034,1247,1272]" pageId="23" pageNumber="224">
<emphasis id="B0E34B53306EFF87FF5EFB4DBC9209EE" bold="true" box="[171,497,1247,1271]" pageId="23" pageNumber="224">
<smallCapsWord id="84CE019D306EFF87FF5EFB4DBC3509EE" baselines="1265,1264" box="[171,342,1247,1271]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Geographical" pageId="23" pageNumber="224">GEOGRAPHICAL</smallCapsWord>
<smallCapsWord id="84CE019D306EFF87FEA8FB70BC9209EE" baselines="1264" box="[349,497,1250,1269]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="distribution" pageId="23" pageNumber="224">DISTRIBUTION</smallCapsWord>
</emphasis>
.— At present, known only from the
<typeStatus id="5D2C29E3306EFF87FC8AFB72BECE09E3" box="[895,941,1248,1272]" pageId="23" pageNumber="224">type</typeStatus>
locality.
</paragraph>
<paragraph id="82289741306EFF87FF5EFA92BEB4082C" blockId="23.[123,1174,309,1336]" pageId="23" pageNumber="224">
<emphasis id="B0E34B53306EFF87FF5EFA92BCC6080D" bold="true" box="[171,421,1280,1304]" pageId="23" pageNumber="224">
<smallCapsWord id="84CE019D306EFF87FF5EFA92BC6A080D" baselines="1298,1297" box="[171,265,1280,1304]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Habitat" pageId="23" pageNumber="224">HABITAT</smallCapsWord>
<smallCapsWord id="84CE019D306EFF87FEE4FA91BCC6080D" baselines="1297" box="[273,421,1283,1302]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="distribution" pageId="23" pageNumber="224">DISTRIBUTION</smallCapsWord>
</emphasis>
.— The unique
<typeStatus id="5D2C29E3306EFF87FDA5FB6DBFCE080C" box="[592,685,1279,1303]" pageId="23" pageNumber="224" type="holotype">holotype</typeStatus>
was collected at ultraviolet light in montane rainforest near the junction of the Namorona River and a small tributary stream.
</paragraph>
</subSubSection>
</treatment>
</document>
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