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<document id="9FFF91DDF99F131EF9E122BD7B96B5DD" ID-DOI="10.5281/zenodo.13799439" ID-ISSN="0068-547X" ID-Zenodo-Dep="13799439" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722538212676" checkinUser="felipe" docAuthor="Kavanaugh, David H. &amp; Rainio, Johanna" docDate="2016" docId="0A3E26573064FF8EFF8EFD99BEDB0B4C" docLanguage="en" docName="ProcCalifAcadSci.63.7.201-268.pdf" docOrigin="Proceedings of the California Academy of Sciences 63 (7)" docStyle="DocumentStyle:E411DA009AF5E7F9A7E023431E2B7D5B.2:ProcCalifAcadSci.2015-.journal_article" docStyleId="E411DA009AF5E7F9A7E023431E2B7D5B" docStyleName="ProcCalifAcadSci.2015-.journal_article" docStyleVersion="2" docTitle="Archicolliuris ranomafanae Kavanaugh and Rainio 2016, sp. nov." docType="treatment" docVersion="2" lastPageNumber="231" masterDocId="F6075E2F3079FF90FFF5FF92BD630D1B" masterDocTitle="Twenty-six New Species of Predaceous Ground Beetles (Coleoptera: Adephaga: Carabidae) from Ranomafana National Park, Madagascar" masterLastPageNumber="268" masterPageNumber="201" pageNumber="230" updateTime="1726771873529" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="ECCD913073B0F0EC17C3D3AE195CBF38" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="0284E82EC75D3C6D94CB7824A07700D7">Twenty-six New Species of Predaceous Ground Beetles (Coleoptera: Adephaga: Carabidae) from Ranomafana National Park, Madagascar</mods:title>
</mods:titleInfo>
<mods:name id="F7F77FAA3B4CD8C1D992F2E7B124CD2C" type="personal">
<mods:role id="FF64D3B670053493D97E2EF690FAC25B">
<mods:roleTerm id="3A7A04AC0EBEC5FEB16F41D83B771F73">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="FE8BCFF7BF8E07911336DB29FED11A9F">Kavanaugh, David H.</mods:namePart>
<mods:affiliation id="EFBF2B409C2642341E93C8476DB49558">Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA; Email: dkavanaugh @ calacademy. org; &amp; Corresponding author: David H. Kavanaugh (dkavanaugh @ calacademy. org)</mods:affiliation>
</mods:name>
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<mods:role id="ADD82E0595CFB09C46E0AF682B21E6C6">
<mods:roleTerm id="A3CE8E68FA6A47D642369664625BBD06">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="6E0DB570105427CA31A76E2E4FA01AB0">Rainio, Johanna</mods:namePart>
<mods:affiliation id="FBE0DFA5363B3798C996329B250EACC0">Department of Ecology and Systematics, University of Helsinki, P. O. Box 65, FIN- 00014 Helsinki, Finland; Email: kjohannarainio @ gmail. com</mods:affiliation>
</mods:name>
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<mods:title id="229FF82FED3E54104C5715FA2392DFA1">Proceedings of the California Academy of Sciences</mods:title>
</mods:titleInfo>
<mods:part id="7A5092C59FE86B39C1CFE414EE453707">
<mods:date id="757DA1998DB4C50A8F5BD26B54D5180B">2016</mods:date>
<mods:detail id="775D2A82197B3EB2F26F24BC29EB06FE" type="pubDate">
<mods:number id="D562613227B5AB5FD454E1F02786C557">2016-04-29</mods:number>
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<mods:detail id="906C7822F091EA21AF18633BF60668D5" type="volume">
<mods:number id="15E9210867318EEA58833321042E4437">63</mods:number>
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<mods:number id="728F4BE721B5AF200DBC0E2924685C2C">7</mods:number>
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<mods:classification id="E85036C02BD74095793EDEC3A3808774">journal article</mods:classification>
<mods:identifier id="50DF6E327210FCED3BC9C48C8DA3D0DE" type="DOI">10.5281/zenodo.13799439</mods:identifier>
<mods:identifier id="D6130496DF5881C9216D9E0A07AD7F06" type="ISSN">0068-547X</mods:identifier>
<mods:identifier id="B63B79E2220E075E0C452A6D002B8ECD" type="Zenodo-Dep">13799439</mods:identifier>
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<treatment id="0A3E26573064FF8EFF8EFD99BEDB0B4C" ID-DOI="http://doi.org/10.5281/zenodo.13799491" ID-Zenodo-Dep="13799491" LSID="urn:lsid:plazi:treatment:0A3E26573064FF8EFF8EFD99BEDB0B4C" httpUri="http://treatment.plazi.org/id/0A3E26573064FF8EFF8EFD99BEDB0B4C" lastPageId="30" lastPageNumber="231" pageId="29" pageNumber="230">
<subSubSection id="CA8DC4CA3064FF8DFF8EFD99BE400F38" box="[123,803,523,547]" pageId="29" pageNumber="230" type="nomenclature">
<paragraph id="822897413064FF8DFF8EFD99BE400F38" blockId="29.[123,803,523,577]" box="[123,803,523,547]" pageId="29" pageNumber="230">
<heading id="D960202D3064FF8DFF8EFD99BE400F38" box="[123,803,523,547]" fontSize="10" level="2" pageId="29" pageNumber="230" reason="4">
<emphasis id="B0E34B533064FF8DFF8EFD99BE400F38" bold="true" box="[123,803,523,547]" pageId="29" pageNumber="230">
<taxonomicName id="4597ECC23064FF8DFF8EFD99BFA30F38" ID-CoL="G8PQ" authority="Kavanaugh and Rainio" authorityName="Kavanaugh and Rainio" authorityYear="2016" box="[123,704,523,547]" class="Insecta" family="Carabidae" genus="Archicolliuris" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="230" phylum="Arthropoda" rank="species" species="ranomafanae" status="sp. nov.">
<emphasis id="B0E34B533064FF8DFF8EFD99BCCC0F38" bold="true" box="[123,431,523,547]" italics="true" pageId="29" pageNumber="230">Archicolliuris ranomafanae</emphasis>
Kavanaugh and Rainio
</taxonomicName>
,
<taxonomicNameLabel id="ABD0F6283064FF8DFD38FD99BE400F38" box="[717,803,523,547]" pageId="29" pageNumber="230" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="CA8DC4CA3064FF8DFF8EFDB9BDBB0F5A" box="[123,216,555,577]" pageId="29" pageNumber="230" type="description">
<paragraph id="822897413064FF8DFF8EFDB9BDBB0F5A" blockId="29.[123,803,523,577]" box="[123,216,555,577]" pageId="29" pageNumber="230">
<figureCitation id="1AAC8BC43064FF8DFF8EFDB9BDBB0F5A" box="[123,216,555,577]" captionStart="FIGURE" captionStartId="30.[155,167,1246,1265]" captionTargetBox="[120,1176,216,1220]" captionTargetId="figure-760@30.[120,1176,216,1239]" captionTargetPageId="30" captionText="FIGURE 16. Digital images of holotype female of Archicolliuris ranomafanae Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B. Labels. Scale line = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13799477" httpUri="https://zenodo.org/record/13799477/files/figure.png" pageId="29" pageNumber="230">Figure 16</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="CA8DC4CA3064FF8EFF5EFDF6BEDB0B4C" lastPageId="30" lastPageNumber="231" pageId="29" pageNumber="230" type="materials_examined">
<paragraph id="822897413064FF8DFF5EFDF6BF980FC7" blockId="29.[123,1174,612,1726]" pageId="29" pageNumber="230">
<emphasis id="B0E34B533064FF8DFF5EFDF6BC3B0F60" bold="true" box="[171,344,612,636]" pageId="29" pageNumber="230">
<smallCapsWord id="84CE019D3064FF8DFF5EFDF6BD870F60" baselines="630,630" box="[171,228,612,636]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Type" pageId="29" pageNumber="230">TYPE</smallCapsWord>
<smallCapsWord id="84CE019D3064FF8DFF1EFDFABC3B0F60" baselines="630" box="[235,344,616,635]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="material" pageId="29" pageNumber="230">MATERIAL</smallCapsWord>
</emphasis>
.—
<materialsCitation id="32FF9D1C3064FF8DFE8AFDF6BF980FC7" collectingDate="1988-10-22" collectingDateMax="1988-10-31" collectingDateMin="1988-10-22" collectorName="W. E. Steiner" country="Madagascar" elevation="1100" location="Ranomafana" pageId="29" pageNumber="230" specimenCount="2" stateProvince="Fianarantsoa" typeStatus="holotype">
<typeStatus id="5D2C29E33064FF8DFE8AFDF6BC810F67" box="[383,482,612,636]" pageId="29" pageNumber="230" type="holotype">Holotype</typeStatus>
(
<figureCitation id="1AAC8BC43064FF8DFE07FDF6BF100F67" box="[498,627,612,636]" captionStart="FIGURE" captionStartId="30.[155,167,1246,1265]" captionTargetBox="[120,1176,216,1220]" captionTargetId="figure-760@30.[120,1176,216,1239]" captionTargetPageId="30" captionText="FIGURE 16. Digital images of holotype female of Archicolliuris ranomafanae Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B. Labels. Scale line = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13799477" httpUri="https://zenodo.org/record/13799477/files/figure.png" pageId="29" pageNumber="230">Figs.16A–B</figureCitation>
), a female, in NMNH, labeled: “
<collectingCountry id="FA80D7D13064FF8DFC25FDF6B9ED0F67" box="[976,1166,612,636]" name="Madagascar" pageId="29" pageNumber="230">MADAGASCAR</collectingCountry>
: Prov.
<collectingRegion id="405359A33064FF8DFF4DFD16BC230F87" box="[184,320,644,668]" country="Madagascar" name="Fianarantsoa" pageId="29" pageNumber="230">Fianarantsoa</collectingRegion>
,
<quantity id="456F3AA43064FF8DFEB8FD17BCE10F87" box="[333,386,644,668]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="29" pageNumber="230" unit="km" value="7.0">7 km</quantity>
W
<location id="8748C19A3064FF8DFE52FD16BF4D0F87" LSID="urn:lsid:plazi:treatment:0A3E26573064FF8EFF8EFD99BEDB0B4C:8748C19A3064FF8DFE52FD16BF4D0F87" box="[423,558,644,668]" country="Madagascar" name="Ranomafana" pageId="29" pageNumber="230" stateProvince="Fianarantsoa">Ranomafana</location>
,
<quantity id="456F3AA43064FF8DFDCFFD16BFE70F87" box="[570,644,644,668]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="29" pageNumber="230" unit="m" value="1100.0">
<elevation id="09BA70723064FF8DFDCFFD16BFE70F87" box="[570,644,644,668]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="29" pageNumber="230" unit="m" value="1100.0">1100m</elevation>
</quantity>
<date id="F629B1813064FF8DFD7CFD16BE3C0F87" box="[649,863,644,668]" pageId="29" pageNumber="230" value="1988-10-22" valueMax="1988-10-31" valueMin="1988-10-22">
<collectingDate id="E66D48693064FF8DFD7CFD16BE3C0F87" box="[649,863,644,668]" pageId="29" pageNumber="230" value="1988-10-22" valueMax="1988-10-31" valueMin="1988-10-22">22-31 October 1988</collectingDate>
</date>
<collectorName id="2F62F2973064FF8DFC90FD17BE920F87" box="[869,1009,644,668]" pageId="29" pageNumber="230">W. E. Steiner</collectorName>
”/ “
<collectingMethod id="5BD6EF563064FF8DFBE3FD16BC0E0FA7" pageId="29" pageNumber="230">Flight intercept-yellow pan trap</collectingMethod>
, island in stream, montane rainforest”/ “
<typeStatus id="5D2C29E33064FF8DFC94FD36BE8C0FA7" box="[865,1007,676,700]" pageId="29" pageNumber="230" type="holotype">HOLOTYPE</typeStatus>
<taxonomicName id="4597ECC23064FF8DFBF5FD36BC970FC7" authority="Kavanaugh &amp; Rainio" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" class="Insecta" family="Carabidae" genus="Archicolliuris" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="230" phylum="Arthropoda" rank="species" species="ranomafanae" status="sp. nov.">
<emphasis id="B0E34B533064FF8DFBF5FD36BC6A0FC7" italics="true" pageId="29" pageNumber="230">Archicolliuris ranomafanae</emphasis>
Kavanaugh &amp; Rainio
</taxonomicName>
<taxonomicNameLabel id="ABD0F6283064FF8DFE0EFD57BF550FC7" box="[507,566,709,732]" pageId="29" pageNumber="230" rank="species">sp. n.</taxonomicNameLabel>
2015” [red label].
</materialsCitation>
</paragraph>
<paragraph id="822897413064FF8DFF5EFD77B9500FE7" blockId="29.[123,1174,612,1726]" box="[171,1075,740,765]" pageId="29" pageNumber="230">
<materialsCitation id="32FF9D1C3064FF8DFF5EFD77B94F0FE7" box="[171,1068,740,765]" country="Madagascar" location="Ranomafana National Park" pageId="29" pageNumber="230" specimenCount="1" stateProvince="Fianarantsoa" typeStatus="holotype">
<emphasis id="B0E34B533064FF8DFF5EFD77BC340FE0" bold="true" box="[171,343,741,765]" pageId="29" pageNumber="230">
<smallCapsWord id="84CE019D3064FF8DFF5EFD77BD870FE0" baselines="759,758" box="[171,228,741,765]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Type" pageId="29" pageNumber="230">
<typeStatus id="5D2C29E33064FF8DFF5EFD77BD870FE0" box="[171,228,741,765]" pageId="29" pageNumber="230">TYPE</typeStatus>
</smallCapsWord>
<smallCapsWord id="84CE019D3064FF8DFF19FD7ABC340FE0" baselines="758" box="[236,343,744,763]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="locality" pageId="29" pageNumber="230">LOCALITY</smallCapsWord>
</emphasis>
.—
<collectingCountry id="FA80D7D13064FF8DFE8BFD76BC9C0FE7" box="[382,511,740,764]" name="Madagascar" pageId="29" pageNumber="230">Madagascar</collectingCountry>
,
<collectingRegion id="405359A33064FF8DFDF8FD76BF9F0FE7" box="[525,764,740,764]" country="Madagascar" name="Fianarantsoa" pageId="29" pageNumber="230">Fianarantsoa Province</collectingRegion>
,
<location id="8748C19A3064FF8DFCFFFD76B94F0FE7" LSID="urn:lsid:plazi:treatment:0A3E26573064FF8EFF8EFD99BEDB0B4C:8748C19A3064FF8DFCFFFD76B94F0FE7" box="[778,1068,740,764]" country="Madagascar" name="Ranomafana National Park" pageId="29" pageNumber="230" stateProvince="Fianarantsoa">Ranomafana National Park</location>
</materialsCitation>
.
</paragraph>
<paragraph id="822897413064FF8DFF5EFC97BECA0E27" blockId="29.[123,1174,612,1726]" pageId="29" pageNumber="230">
<emphasis id="B0E34B533064FF8DFF5EFC97BC910E00" bold="true" box="[171,498,773,797]" pageId="29" pageNumber="230">
<smallCapsWord id="84CE019D3064FF8DFF5EFC97BC4D0E00" baselines="791,790" box="[171,302,773,797]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Derivation" pageId="29" pageNumber="230">DERIVATION</smallCapsWord>
<smallCapsWord id="84CE019D3064FF8DFEC3FC9ABC310E00" baselines="790" box="[310,338,776,795]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="of" pageId="29" pageNumber="230">OF</smallCapsWord>
<smallCapsWord id="84CE019D3064FF8DFEACFC9ABCCF0E00" baselines="790" box="[345,428,776,795]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="species" pageId="29" pageNumber="230">SPECIES</smallCapsWord>
<smallCapsWord id="84CE019D3064FF8DFE40FC9ABC910E00" baselines="790" box="[437,498,776,795]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="name" pageId="29" pageNumber="230">NAME</smallCapsWord>
</emphasis>
.— The species epithet,
<taxonomicName id="4597ECC23064FF8DFD0CFC97BEE40E06" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[761,903,773,797]" class="Insecta" family="Carabidae" genus="Archicolliuris" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="230" phylum="Arthropoda" rank="species" species="ranomafanae">
<emphasis id="B0E34B533064FF8DFD0CFC97BEE40E06" box="[761,903,773,797]" italics="true" pageId="29" pageNumber="230">ranomafanae</emphasis>
</taxonomicName>
, is a noun in apposition, derived from the name of the national park in which the
<typeStatus id="5D2C29E33064FF8DFD29FCB7BE690E26" box="[732,778,805,829]" pageId="29" pageNumber="230">type</typeStatus>
was collected.
</paragraph>
<paragraph id="822897413064FF8DFF5EFCD7BFD10B86" blockId="29.[123,1174,612,1726]" pageId="29" pageNumber="230">
<smallCapsWord id="84CE019D3064FF8DFF5EFCD7BC210E40" baselines="855,854" box="[171,322,837,861]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Recognition" pageId="29" pageNumber="230">
<emphasis id="B0E34B533064FF8DFF5EFCD7BC210E40" bold="true" box="[171,322,837,861]" pageId="29" pageNumber="230">RECOGNITION</emphasis>
</smallCapsWord>
.— Size large for genus, SBL =
<quantity id="456F3AA43064FF8DFD53FCD7BF990E47" box="[678,762,837,861]" metricMagnitude="-3" metricUnit="m" metricValue="8.2" pageId="29" pageNumber="230" unit="mm" value="8.2">8.2 mm</quantity>
. The
<typeStatus id="5D2C29E33064FF8DFCCBFCD6BEF80E47" box="[830,923,836,860]" pageId="29" pageNumber="230" type="holotype">holotype</typeStatus>
female of this species (
<figureCitation id="1AAC8BC43064FF8DFF71FCF6BD800E66" box="[132,227,868,893]" captionStart="FIGURE" captionStartId="30.[155,167,1246,1265]" captionTargetBox="[120,1176,216,1220]" captionTargetId="figure-760@30.[120,1176,216,1239]" captionTargetPageId="30" captionText="FIGURE 16. Digital images of holotype female of Archicolliuris ranomafanae Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B. Labels. Scale line = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13799477" httpUri="https://zenodo.org/record/13799477/files/figure.png" pageId="29" pageNumber="230">Fig. 16A</figureCitation>
) is easily distinguished from members of all other Malagasy odacanthine species based on numerous features. The following features used by Jeannel (1948) in his key to genera are shared with the three species that he included in “
<taxonomicName id="4597ECC23064FF8DFD79FC37BF910EA7" authorityName="Latreille &amp; Dejean" authorityYear="1822" box="[652,754,933,956]" class="Insecta" family="Carabidae" genus="Casnonia" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="230" phylum="Arthropoda" rank="genus">
<emphasis id="B0E34B533064FF8DFD79FC37BF910EA7" box="[652,754,933,956]" italics="true" pageId="29" pageNumber="230">Casnonia</emphasis>
</taxonomicName>
”: integument glabrous (without pubescence), pronotum long, narrow and tubular anteriorly, impunctate, with one or more setae present along each lateral margin. The
<typeStatus id="5D2C29E33064FF8DFE34FC77BF7D0EE6" box="[449,542,997,1021]" pageId="29" pageNumber="230" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4597ECC23064FF8DFDB4FC77BF880EE6" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[577,747,997,1021]" class="Insecta" family="Carabidae" genus="Archicolliuris" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="230" phylum="Arthropoda" rank="species" species="ranomafanae">
<emphasis id="B0E34B533064FF8DFDB4FC77BF880EE6" box="[577,747,997,1021]" italics="true" pageId="29" pageNumber="230">A. ranomafanae</emphasis>
</taxonomicName>
differs from members of
<taxonomicName id="4597ECC23064FF8DFC0FFC77BC060906" authority="Gestro (1895)" authorityName="Gestro" authorityYear="1895" class="Insecta" family="Carabidae" genus="Casnonia" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="230" phylum="Arthropoda" rank="species" species="fairmairei">
<emphasis id="B0E34B533064FF8DFC0FFC77BDA30907" italics="true" pageId="29" pageNumber="230">Casnonia fairmairei</emphasis>
<bibRefCitation id="E606EAB03064FF8DFF3EFB97BC060906" author="GESTRO, R." box="[203,357,1029,1053]" pageId="29" pageNumber="230" pagination="247 - 478" refId="ref30448" refString="GESTRO, R. 1895. Esplorazione del Giuba e dei suoi affluienti compiuta dal Cap. V. Bottego durante gli anni 1892 - 93 sotto gli auspicii della Societa Geografica Italiana. Risultati zoologici. Coleotteri. Annali del Museo Civico di Storia Naturale di Genova 35: 247 - 478." type="journal article" year="1895">Gestro (1895)</bibRefCitation>
</taxonomicName>
[currently included in genus
<taxonomicName id="4597ECC23064FF8DFD4DFB97B9630906" authority="Liebke (1931)" authorityName="Liebke" authorityYear="1931" box="[696,1024,1029,1053]" class="Insecta" family="Carabidae" genus="Erectocolliuris" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="230" phylum="Arthropoda" rank="genus">
<emphasis id="B0E34B533064FF8DFD4DFB97BE340906" box="[696,855,1029,1053]" italics="true" pageId="29" pageNumber="230">Erectocolliuris</emphasis>
<bibRefCitation id="E606EAB03064FF8DFC96FB97B9630906" author="LIEBKE, M." box="[867,1024,1029,1053]" pageId="29" pageNumber="230" pagination="280 - 301" refId="ref31583" refString="LIEBKE, M. 1931. Die Africanischen Arten der Gattung Colliuris Degeer (Col. Car.). Revue de Zoologie et de Botanique Africaines 20 (1930 - 31): 280 - 301." type="journal article" year="1931">Liebke (1931)</bibRefCitation>
</taxonomicName>
] in having a pronotum black in color and with only a single pair of midlateral setae (pronotum rufotestaceous in color and with 5 or 6 pairs of lateral setae present in
<taxonomicName id="4597ECC23064FF8DFD1FFBD4BE160946" authorityName="Gestro" authorityYear="1895" box="[746,885,1093,1117]" class="Insecta" family="Carabidae" genus="Casnonia" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="230" phylum="Arthropoda" rank="species" species="fairmairei">
<emphasis id="B0E34B533064FF8DFD1FFBD4BE160946" box="[746,885,1093,1117]" italics="true" pageId="29" pageNumber="230">E. fairmairei</emphasis>
</taxonomicName>
members). It differs from members
<taxonomicName id="4597ECC23064FF8DFF11FBF7BF920966" authority="Kunckel d'Herculais (1887)" authorityName="Kunckel d'Herculais" authorityYear="1887" box="[228,753,1125,1149]" class="Insecta" family="Carabidae" genus="Casnonia" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="230" phylum="Arthropoda" rank="species" species="coerulans">
<emphasis id="B0E34B533064FF8DFF11FBF7BCD80966" box="[228,443,1125,1149]" italics="true" pageId="29" pageNumber="230">Casnonia coerulans</emphasis>
Künckel d’Herculais (1887)
</taxonomicName>
[currently included in genus
<taxonomicName id="4597ECC23064FF8DFBC5FBF7BC300986" authority="Liebke (1931)" authorityName="Liebke" authorityYear="1931" class="Insecta" family="Carabidae" genus="Protocolliuris" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="230" phylum="Arthropoda" rank="genus">
<emphasis id="B0E34B533064FF8DFBC5FBF7BDD00986" italics="true" pageId="29" pageNumber="230">Protocolliuris</emphasis>
<bibRefCitation id="E606EAB03064FF8DFF4FFB17BC300986" author="LIEBKE, M." box="[186,339,1157,1181]" pageId="29" pageNumber="230" pagination="280 - 301" refId="ref31583" refString="LIEBKE, M. 1931. Die Africanischen Arten der Gattung Colliuris Degeer (Col. Car.). Revue de Zoologie et de Botanique Africaines 20 (1930 - 31): 280 - 301." type="journal article" year="1931">Liebke (1931)</bibRefCitation>
</taxonomicName>
in having the pronotum only about twice as long as wide (three times as long as wide in
<taxonomicName id="4597ECC23064FF8DFF03FB34BC1E09A6" authorityName="Kunckel d'Herculais" authorityYear="1887" box="[246,381,1189,1213]" class="Insecta" family="Carabidae" genus="Casnonia" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="230" phylum="Arthropoda" rank="species" species="coerulans">
<emphasis id="B0E34B533064FF8DFF03FB34BC1E09A6" box="[246,381,1189,1213]" italics="true" pageId="29" pageNumber="230">P. coerulans</emphasis>
</taxonomicName>
), the elytra without metallic reflection (elytra with a dark metallic blue reflection), and legs dark black to piceous, except trochanthers rufous and basal parts of all femora pale (legs pale in
<taxonomicName id="4597ECC23064FF8DFEADFB74BCBE09E6" authorityName="Kunckel d'Herculais" authorityYear="1887" box="[344,477,1253,1277]" class="Insecta" family="Carabidae" genus="Casnonia" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="230" phylum="Arthropoda" rank="species" species="coerulans">
<emphasis id="B0E34B533064FF8DFEADFB74BCBE09E6" box="[344,477,1253,1277]" italics="true" pageId="29" pageNumber="230">P. coerulans</emphasis>
</taxonomicName>
, except apical parts of femora dark). It is similar to members of the last of Jeannel’s “
<taxonomicName id="4597ECC23064FF8DFEABFA97BCA70807" authorityName="Latreille &amp; Dejean" authorityYear="1822" box="[350,452,1285,1308]" class="Insecta" family="Carabidae" genus="Casnonia" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="230" phylum="Arthropoda" rank="genus">
<emphasis id="B0E34B533064FF8DFEABFA97BCA70807" box="[350,452,1285,1308]" italics="true" pageId="29" pageNumber="230">Casnonia</emphasis>
</taxonomicName>
” species,
<taxonomicName id="4597ECC23064FF8DFDC5FA97BECA0806" authority="Alluaud (1935)" authorityName="Alluaud" authorityYear="1935" box="[560,937,1285,1309]" class="Insecta" family="Carabidae" genus="Colliuris" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="230" phylum="Arthropoda" rank="species" species="olsoufieffi">
<emphasis id="B0E34B533064FF8DFDC5FA97BE630806" box="[560,768,1285,1309]" italics="true" pageId="29" pageNumber="230">Colliuris olsoufieffi</emphasis>
<bibRefCitation id="E606EAB03064FF8DFCF0FA97BECA0806" author="ALLUAUD, C." box="[773,937,1285,1309]" pageId="29" pageNumber="230" pagination="1 - 27" refId="ref28945" refString="ALLUAUD, C. 1935. Carabidae recueillis a Madagascar par MM. A. Syrig et G. Olsoufieff de 1933 a 1935. Afra, Cahiers d'Entomologie 10: 1 - 27." type="journal article" year="1935">Alluaud (1935)</bibRefCitation>
</taxonomicName>
[currently included in
<taxonomicName id="4597ECC23064FF8DFF8EFAB7BC6D0826" authorityName="Liebke" authorityYear="1931" box="[123,270,1317,1341]" class="Insecta" family="Carabidae" genus="Archicolliuris" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="230" phylum="Arthropoda" rank="genus">
<emphasis id="B0E34B533064FF8DFF8EFAB7BC6D0826" box="[123,270,1317,1341]" italics="true" pageId="29" pageNumber="230">Archicolliuris</emphasis>
</taxonomicName>
] in having a shiny black dorsum (with the only pale areas present as elytral pale spots), the pronotum with transverse grooves and ridges in the basal half and anterior angles projected laterally, and the elytra with a deep transverse depression at the basal one-fourth and with three or four discal setiferous pores on elytral interval 3. However, it differs with members of
<taxonomicName id="4597ECC23064FF8DFF8EFA37BC6B08A6" authorityName="Alluaud" authorityYear="1935" box="[123,264,1445,1469]" class="Insecta" family="Carabidae" genus="Colliuris" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="230" phylum="Arthropoda" rank="species" species="olsoufieffi">
<emphasis id="B0E34B533064FF8DFF8EFA37BC6B08A6" box="[123,264,1445,1469]" italics="true" pageId="29" pageNumber="230">A. olsoufieffi</emphasis>
</taxonomicName>
in several features: body size larger (SBL =
<quantity id="456F3AA43064FF8DFCF1FA37BE3908A6" box="[772,858,1445,1469]" metricMagnitude="-3" metricUnit="m" metricValue="6.8" pageId="29" pageNumber="230" unit="mm" value="6.8">6.8 mm</quantity>
in
<taxonomicName id="4597ECC23064FF8DFC70FA37B97108A6" authorityName="Alluaud" authorityYear="1935" box="[901,1042,1445,1469]" class="Insecta" family="Carabidae" genus="Colliuris" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="230" phylum="Arthropoda" rank="species" species="olsoufieffi">
<emphasis id="B0E34B533064FF8DFC70FA37B97108A6" box="[901,1042,1445,1469]" italics="true" pageId="29" pageNumber="230">A. olsoufieffi</emphasis>
</taxonomicName>
); pronotum impunctate (coarsely punctate laterally and basally in
<taxonomicName id="4597ECC23064FF8DFD40FA54BE5E08C6" authorityName="Alluaud" authorityYear="1935" box="[693,829,1477,1501]" class="Insecta" family="Carabidae" genus="Colliuris" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="230" phylum="Arthropoda" rank="species" species="olsoufieffi">
<emphasis id="B0E34B533064FF8DFD40FA54BE5E08C6" box="[693,829,1477,1501]" italics="true" pageId="29" pageNumber="230">A. olsoufieffi</emphasis>
</taxonomicName>
), slightly less than twice as long as wide (2.5 times as long as wide in
<taxonomicName id="4597ECC23064FF8DFDE7FA74BFFF08E6" authorityName="Alluaud" authorityYear="1935" box="[530,668,1509,1533]" class="Insecta" family="Carabidae" genus="Colliuris" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="230" phylum="Arthropoda" rank="species" species="olsoufieffi">
<emphasis id="B0E34B533064FF8DFDE7FA74BFFF08E6" box="[530,668,1509,1533]" italics="true" pageId="29" pageNumber="230">A. olsoufieffi</emphasis>
</taxonomicName>
), with lateral borders vaguely present but only as very faintly impressed lines (lateral borders absent from
<taxonomicName id="4597ECC23064FF8DFCFAF994BEF90B06" authorityName="Alluaud" authorityYear="1935" box="[783,922,1541,1565]" class="Insecta" family="Carabidae" genus="Colliuris" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="230" phylum="Arthropoda" rank="species" species="olsoufieffi">
<emphasis id="B0E34B533064FF8DFCFAF994BEF90B06" box="[783,922,1541,1565]" italics="true" pageId="29" pageNumber="230">A. olsoufieffi</emphasis>
</taxonomicName>
members); elytra with only one pair of small pale spots, located at apical one-third on interval 4 (both subapical and subbasal pairs of spots present in
<taxonomicName id="4597ECC23064FF8DFE36F9D4BF2F0B46" authorityName="Alluaud" authorityYear="1935" box="[451,588,1605,1629]" class="Insecta" family="Carabidae" genus="Colliuris" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="230" phylum="Arthropoda" rank="species" species="olsoufieffi">
<emphasis id="B0E34B533064FF8DFE36F9D4BF2F0B46" box="[451,588,1605,1629]" italics="true" pageId="29" pageNumber="230">A. olsoufieffi</emphasis>
</taxonomicName>
) and with three or four discal setiferous pores also on interval 5 (absent from
<taxonomicName id="4597ECC23064FF8DFE8EF9F4BF660B66" authorityName="Alluaud" authorityYear="1935" box="[379,517,1637,1661]" class="Insecta" family="Carabidae" genus="Colliuris" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="230" phylum="Arthropoda" rank="species" species="olsoufieffi">
<emphasis id="B0E34B533064FF8DFE8EF9F4BF660B66" box="[379,517,1637,1661]" italics="true" pageId="29" pageNumber="230">A. olsoufieffi</emphasis>
</taxonomicName>
members); and legs dark, except pale at base of femora and on trochanters (legs pale thoughout in
<taxonomicName id="4597ECC23064FF8DFDECF914BFC10B86" authorityName="Alluaud" authorityYear="1935" box="[537,674,1669,1693]" class="Insecta" family="Carabidae" genus="Colliuris" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="230" phylum="Arthropoda" rank="species" species="olsoufieffi">
<emphasis id="B0E34B533064FF8DFDECF914BFC10B86" box="[537,674,1669,1693]" italics="true" pageId="29" pageNumber="230">A. olsoufieffi</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="822897413064FF8EFF5EF934B9EA08EA" blockId="29.[123,1174,612,1726]" lastBlockId="30.[123,1174,1328,1624]" lastPageId="30" lastPageNumber="231" pageId="29" pageNumber="230">
In addition, the
<typeStatus id="5D2C29E33064FF8DFE91F937BCA20BA6" box="[356,449,1701,1725]" pageId="29" pageNumber="230" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4597ECC23064FF8DFE04F934BFC10BA6" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[497,674,1701,1725]" class="Insecta" family="Carabidae" genus="Archicolliuris" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="230" phylum="Arthropoda" rank="species" species="ranomafanae">
<emphasis id="B0E34B533064FF8DFE04F934BFC10BA6" box="[497,674,1701,1725]" italics="true" pageId="29" pageNumber="230">A. ranomafanae</emphasis>
</taxonomicName>
is unique among members of all Malagasy odcanthine species in having the elytra distinctly convex throughout, except in the area of the transverse depression, elytral intervals flat, and elytral striae 3 to 7 effaced except in the transverse depression and apically, where they are evident. Striae 1 and 2 are very shallowly impressed but evident (stria 2 less so) in and posterior to the transverse depression. The form and location of the transverse depression of the elytra and the nearly effaced striae are similar to these features of members and genus
<taxonomicName id="4597ECC23067FF8EFEA2FA48BFED08E9" authority="Liebke (1933)" authorityName="Liebke" authorityYear="1933" box="[343,654,1498,1522]" class="Insecta" family="Carabidae" genus="Mimocolliuris" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="231" phylum="Arthropoda" rank="genus">
<emphasis id="B0E34B533067FF8EFEA2FA48BC8D08E9" box="[343,494,1498,1522]" italics="true" pageId="30" pageNumber="231">Mimocolliuris</emphasis>
<bibRefCitation id="E606EAB03067FF8EFE03FA48BFED08E9" author="LIEBKE, M." box="[502,654,1498,1522]" pageId="30" pageNumber="231" pagination="202 - 210" refId="ref31621" refString="LIEBKE, M. 1933. Neue Colliurinen. Stylops 2: 202 - 210." type="journal article" year="1933">Liebke (1933)</bibRefCitation>
</taxonomicName>
of the eastern Palearctic and Oriental Regions.
</paragraph>
<caption id="D6E8C7C93067FF8EFF6EFB4CBF660810" ID-DOI="http://doi.org/10.5281/zenodo.13799477" ID-Zenodo-Dep="13799477" httpUri="https://zenodo.org/record/13799477/files/figure.png" pageId="30" pageNumber="231" startId="30.[155,167,1246,1265]" targetBox="[120,1176,216,1220]" targetPageId="30" targetType="figure">
<paragraph id="822897413067FF8EFF6EFB4CBF660810" blockId="30.[123,1173,1246,1291]" pageId="30" pageNumber="231">
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16. Digital images of holotype female of
<taxonomicName id="4597ECC23067FF8EFDCAFB4CBE8B09EA" authority="Kavanaugh &amp; Rainio" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[575,1000,1246,1265]" class="Insecta" family="Carabidae" genus="Archicolliuris" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="231" phylum="Arthropoda" rank="species" species="ranomafanae" status="sp. nov.">
<emphasis id="B0E34B533067FF8EFDCAFB4CBE4F09EA" box="[575,812,1246,1265]" italics="true" pageId="30" pageNumber="231">Archicolliuris ranomafanae</emphasis>
Kavanaugh &amp; Rainio
</taxonomicName>
<taxonomicNameLabel id="ABD0F6283067FF8EFC18FB4CB94C09EA" box="[1005,1071,1246,1265]" pageId="30" pageNumber="231" rank="species">sp. nov.</taxonomicNameLabel>
A. Habitus, dorsal aspect; B. Labels. Scale line = 1.0 mm.
</paragraph>
</caption>
<paragraph id="822897413067FF8EFF5EFA69B9690B08" blockId="30.[123,1174,1328,1624]" box="[171,1034,1531,1556]" pageId="30" pageNumber="231">
<emphasis id="B0E34B533067FF8EFF5EFA69BC920B09" bold="true" box="[171,497,1531,1555]" pageId="30" pageNumber="231">
<smallCapsWord id="84CE019D3067FF8EFF5EFA69BC350B09" baselines="1549,1549" box="[171,342,1531,1555]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Geographical" pageId="30" pageNumber="231">GEOGRAPHICAL</smallCapsWord>
<smallCapsWord id="84CE019D3067FF8EFEA8FA6DBC920B09" baselines="1549" box="[349,497,1535,1554]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="distribution" pageId="30" pageNumber="231">DISTRIBUTION</smallCapsWord>
</emphasis>
.— At present, known only from the
<typeStatus id="5D2C29E33067FF8EFC8AFA6EBECE0B0F" box="[895,941,1532,1556]" pageId="30" pageNumber="231">type</typeStatus>
locality.
</paragraph>
<paragraph id="822897413067FF8EFF5EF98CBEDB0B4C" blockId="30.[123,1174,1328,1624]" pageId="30" pageNumber="231">
<emphasis id="B0E34B533067FF8EFF5EF98CBCC70B2F" bold="true" box="[171,420,1566,1590]" pageId="30" pageNumber="231">
<smallCapsWord id="84CE019D3067FF8EFF5EF98CBC6A0B2F" baselines="1584,1583" box="[171,265,1566,1590]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Habitat" pageId="30" pageNumber="231">HABITAT</smallCapsWord>
<smallCapsWord id="84CE019D3067FF8EFEE5F9B3BCC70B2F" baselines="1583" box="[272,420,1569,1588]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="distribution" pageId="30" pageNumber="231">DISTRIBUTION</smallCapsWord>
</emphasis>
.— The unique
<typeStatus id="5D2C29E33067FF8EFDB9F98FBFCA0B2E" box="[588,681,1565,1589]" pageId="30" pageNumber="231" type="holotype">holotype</typeStatus>
was collected in a flight intercept-yellow pan trap on an island in a stream in montane rainforest at an elevation of
<quantity id="456F3AA43067FF8EFC94F9D2BED10B4C" box="[865,946,1600,1624]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="30" pageNumber="231" unit="m" value="1100.0">1100 m</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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