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<document id="F524234A9CBB907DCD3A8635D4A9F89B" ID-DOI="10.5281/zenodo.13799439" ID-ISSN="0068-547X" ID-Zenodo-Dep="13799439" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722538212676" checkinUser="felipe" docAuthor="Kavanaugh, David H. &amp; Rainio, Johanna" docDate="2016" docId="0A3E2657305AFFB5FF8EFA95B9710F61" docLanguage="en" docName="ProcCalifAcadSci.63.7.201-268.pdf" docOrigin="Proceedings of the California Academy of Sciences 63 (7)" docStyle="DocumentStyle:E411DA009AF5E7F9A7E023431E2B7D5B.2:ProcCalifAcadSci.2015-.journal_article" docStyleId="E411DA009AF5E7F9A7E023431E2B7D5B" docStyleName="ProcCalifAcadSci.2015-.journal_article" docStyleVersion="2" docTitle="Madecassina quadrimaculata Kavanaugh and Rainio 2016, sp. nov." docType="treatment" docVersion="2" lastPageNumber="238" masterDocId="F6075E2F3079FF90FFF5FF92BD630D1B" masterDocTitle="Twenty-six New Species of Predaceous Ground Beetles (Coleoptera: Adephaga: Carabidae) from Ranomafana National Park, Madagascar" masterLastPageNumber="268" masterPageNumber="201" pageNumber="236" updateTime="1726771873529" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="1E2A0EC3B2601D4B4359ED4806320B37" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="EDB0749890DC13BC3F6BB97A76FAD2C3">
<mods:title id="56FC5662DFE68C7AE76EEE4E7486C536">Twenty-six New Species of Predaceous Ground Beetles (Coleoptera: Adephaga: Carabidae) from Ranomafana National Park, Madagascar</mods:title>
</mods:titleInfo>
<mods:name id="7ACDE97A3F1CC3A682C6206ABCEBA762" type="personal">
<mods:role id="D47933319E15E4A83163BE88E50B0D41">
<mods:roleTerm id="1F9D7C71969D9EED52BE44B58AF32561">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="FBC7592CD851E8EB6CA31C8ABF47F7C6">Kavanaugh, David H.</mods:namePart>
<mods:affiliation id="F2894E6C864DEF01B4E9DF38B6061D15">Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA; Email: dkavanaugh @ calacademy. org; &amp; Corresponding author: David H. Kavanaugh (dkavanaugh @ calacademy. org)</mods:affiliation>
</mods:name>
<mods:name id="CBD5F80520B78BF7DC8BA886DC0516D7" type="personal">
<mods:role id="0B94ED213550D1445DC02C33E403DE2B">
<mods:roleTerm id="0AE189359B5015585A09494A3B24F235">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5E542D127042B6860855E9599A312C1B">Rainio, Johanna</mods:namePart>
<mods:affiliation id="2235822C649D4169CEA913A71601E267">Department of Ecology and Systematics, University of Helsinki, P. O. Box 65, FIN- 00014 Helsinki, Finland; Email: kjohannarainio @ gmail. com</mods:affiliation>
</mods:name>
<mods:typeOfResource id="54A9C833284CF1BC65FB3EC826CF721C">text</mods:typeOfResource>
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<mods:title id="7FCC9F4F1E3A5E62EF1EA397E25AD6C8">Proceedings of the California Academy of Sciences</mods:title>
</mods:titleInfo>
<mods:part id="F035536A2A5AEBF0FCAF910F082D5648">
<mods:date id="EC356C0DFF5D1AA0550410DEDFC531C1">2016</mods:date>
<mods:detail id="16D01B6D82C249FAC7F7CF5F5B82527B" type="pubDate">
<mods:number id="C812231B8D3C3B8CFA79E6AE6DA5CA68">2016-04-29</mods:number>
</mods:detail>
<mods:detail id="2E211C414005D46A41C2F711A3213755" type="volume">
<mods:number id="5AE1EDC6CB4D841476E4087BAF1F62CA">63</mods:number>
</mods:detail>
<mods:detail id="16A89F3ADA40125659F24DFD8B6F273B" type="issue">
<mods:number id="17CF0D7F12A475EA17275E515F664784">7</mods:number>
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<mods:classification id="9026F9EDDBDF24E607186056DCC9D155">journal article</mods:classification>
<mods:identifier id="C24DF76EDA71200DDC4BB44D979AEC13" type="DOI">10.5281/zenodo.13799439</mods:identifier>
<mods:identifier id="749D54AB2A2C98269B5FBDA07697D229" type="ISSN">0068-547X</mods:identifier>
<mods:identifier id="16053BC3746C7A690470991E1966DD65" type="Zenodo-Dep">13799439</mods:identifier>
</mods:mods>
<treatment id="0A3E2657305AFFB5FF8EFA95B9710F61" ID-DOI="http://doi.org/10.5281/zenodo.13799513" ID-Zenodo-Dep="13799513" LSID="urn:lsid:plazi:treatment:0A3E2657305AFFB5FF8EFA95B9710F61" httpUri="http://treatment.plazi.org/id/0A3E2657305AFFB5FF8EFA95B9710F61" lastPageId="37" lastPageNumber="238" pageId="35" pageNumber="236">
<subSubSection id="CA8DC4CA305AFFB3FF8EFA95BE56083B" box="[123,821,1287,1312]" pageId="35" pageNumber="236" type="nomenclature">
<paragraph id="82289741305AFFB3FF8EFA95BE56083B" blockId="35.[123,821,1287,1342]" box="[123,821,1287,1312]" pageId="35" pageNumber="236">
<heading id="D960202D305AFFB3FF8EFA95BE56083B" box="[123,821,1287,1312]" fontSize="10" level="2" pageId="35" pageNumber="236" reason="4">
<emphasis id="B0E34B53305AFFB3FF8EFA95BE56083B" bold="true" box="[123,821,1287,1312]" pageId="35" pageNumber="236">
<taxonomicName id="4597ECC2305AFFB3FF8EFA95BFB20804" ID-CoL="6QRM3" authority="Kavanaugh and Rainio" authorityName="Kavanaugh and Rainio" authorityYear="2016" box="[123,721,1287,1312]" class="Insecta" family="Carabidae" genus="Madecassina" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="236" phylum="Arthropoda" rank="species" species="quadrimaculata" status="sp. nov.">
<emphasis id="B0E34B53305AFFB3FF8EFA95BCA20804" bold="true" box="[123,449,1287,1311]" italics="true" pageId="35" pageNumber="236">Madecassina quadrimaculata</emphasis>
Kavanaugh and Rainio
</taxonomicName>
,
<taxonomicNameLabel id="ABD0F628305AFFB3FD2AFA9ABE56083B" box="[735,821,1288,1312]" pageId="35" pageNumber="236" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="CA8DC4CA305AFFB3FF8EFABABDBB0826" box="[123,216,1320,1342]" pageId="35" pageNumber="236" type="description">
<paragraph id="82289741305AFFB3FF8EFABABDBB0826" blockId="35.[123,821,1287,1342]" box="[123,216,1320,1342]" pageId="35" pageNumber="236">
<figureCitation id="1AAC8BC4305AFFB3FF8EFABABDBB0826" box="[123,216,1320,1342]" captionStart="FIGURE" captionStartId="36.[155,167,1570,1589]" captionTargetBox="[144,1176,216,1552]" captionTargetId="figure-53@36.[120,1176,216,1567]" captionTargetPageId="36" captionText="FIGURE 20. Digital images of holotype male of Madecassina quadrimaculata Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B. Labels C–D. Aedeagus of male genitalia; C. Left lateral aspect; D. Dorsal aspect. Scale lines A = 1.0 mm, C–D = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13799487" httpUri="https://zenodo.org/record/13799487/files/figure.png" pageId="35" pageNumber="236">Figure 20</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="CA8DC4CA305AFFB5FF5EFAF0B9710F61" lastPageId="37" lastPageNumber="238" pageId="35" pageNumber="236" type="materials_examined">
<paragraph id="82289741305AFFB3FF5EFAF0B9F508C1" blockId="35.[123,1174,1378,1725]" pageId="35" pageNumber="236">
<emphasis id="B0E34B53305AFFB3FF5EFAF0BC3D0863" bold="true" box="[171,350,1378,1402]" pageId="35" pageNumber="236">
<smallCapsWord id="84CE019D305AFFB3FF5EFAF0BD870863" baselines="1396,1395" box="[171,228,1378,1402]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Type" pageId="35" pageNumber="236">TYPE</smallCapsWord>
<smallCapsWord id="84CE019D305AFFB3FF04FAF7BC3D0863" baselines="1395" box="[241,350,1381,1400]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="material" pageId="35" pageNumber="236">MATERIAL</smallCapsWord>
</emphasis>
.—
<materialsCitation id="32FF9D1C305AFFB3FE73FAF0B9F508C1" collectingDate="2005-05-19" collectorName="F. J. Ratelolahy" country="Madagascar" elevation="100" latitude="-21.09637" location="Tsinjorano village" longLatPrecision="1" longitude="47.52085" pageId="35" pageNumber="236" specimenCount="2" typeStatus="holotype">
<typeStatus id="5D2C29E3305AFFB3FE73FAF0BC8A0861" box="[390,489,1378,1402]" pageId="35" pageNumber="236" type="holotype">Holotype</typeStatus>
(
<figureCitation id="1AAC8BC4305AFFB3FE0BFAF0BFE30862" box="[510,640,1378,1402]" captionStart="FIGURE" captionStartId="36.[155,167,1570,1589]" captionTargetBox="[144,1176,216,1552]" captionTargetId="figure-53@36.[120,1176,216,1567]" captionTargetPageId="36" captionText="FIGURE 20. Digital images of holotype male of Madecassina quadrimaculata Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B. Labels C–D. Aedeagus of male genitalia; C. Left lateral aspect; D. Dorsal aspect. Scale lines A = 1.0 mm, C–D = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13799487" httpUri="https://zenodo.org/record/13799487/files/figure.png" pageId="35" pageNumber="236">Figs.20A–B</figureCitation>
), a male, in CAS, labeled: “
<collectingCountry id="FA80D7D1305AFFB3FC24FAF0B9EC0861" box="[977,1167,1378,1402]" name="Madagascar" pageId="35" pageNumber="236">MADAGASCAR</collectingCountry>
, Ranomafana NP., montane rainforest near
<location id="8748C19A305AFFB3FD91FA10BE4F0881" LSID="urn:lsid:plazi:treatment:0A3E2657305AFFB5FF8EFA95B9710F61:8748C19A305AFFB3FD91FA10BE4F0881" box="[612,812,1410,1434]" country="Madagascar" latitude="-21.09637" longLatPrecision="1" longitude="47.52085" name="Tsinjorano village" pageId="35" pageNumber="236">Tsinjorano village</location>
,
<quantity id="456F3AA4305AFFB3FCB4FA10BEEF0882" box="[833,908,1410,1434]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="35" pageNumber="236" unit="m" value="100.0">
<elevation id="09BA7072305AFFB3FCB4FA10BEEF0882" box="[833,908,1410,1434]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="35" pageNumber="236" unit="m" value="100.0">100 m</elevation>
</quantity>
in from forest edge,”/ “
<geoCoordinate id="E7A3F186305AFFB3FF73FA30BC6708A1" box="[134,260,1442,1466]" degrees="21. 09637" direction="south" orientation="latitude" pageId="35" pageNumber="236" precision="1" value="-21.09637">21.09637°S</geoCoordinate>
<geoCoordinate id="E7A3F186305AFFB3FEF9FA30BCE908A1" box="[268,394,1442,1466]" degrees="47. 52085" direction="east" orientation="longitude" pageId="35" pageNumber="236" precision="1" value="47.52085">47.52085°E</geoCoordinate>
,
<quantity id="456F3AA4305AFFB3FE6CFA30BC8808A1" box="[409,491,1442,1466]" metricMagnitude="3" metricUnit="m" metricValue="1.05" pageId="35" pageNumber="236" unit="m" value="1050.0">
<elevation id="09BA7072305AFFB3FE6CFA30BC8808A1" box="[409,491,1442,1466]" metricMagnitude="3" metricUnit="m" metricValue="1.05" pageId="35" pageNumber="236" unit="m" value="1050.0">1050 m</elevation>
</quantity>
,
<date id="F629B181305AFFB3FE0CFA30BF1308A1" box="[505,624,1442,1466]" pageId="35" pageNumber="236" value="2005-05-19">
<collectingDate id="E66D4869305AFFB3FE0CFA30BF1308A1" box="[505,624,1442,1466]" pageId="35" pageNumber="236" value="2005-05-19">19.05.2005</collectingDate>
</date>
, collected by hand in daytime,
<collectorName id="2F62F297305AFFB3FC30FA31B90608A1" box="[965,1125,1442,1466]" pageId="35" pageNumber="236">F.J. Ratelolahy</collectorName>
collector”/ “
<typeStatus id="5D2C29E3305AFFB3FF28FA51BC0908C0" box="[221,362,1475,1499]" pageId="35" pageNumber="236" type="holotype">HOLOTYPE</typeStatus>
<taxonomicName id="4597ECC2305AFFB3FE85FA51BEF708C0" authority="Kavanaugh &amp; Rainio" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[368,916,1475,1499]" class="Insecta" family="Carabidae" genus="Madecassina" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="236" phylum="Arthropoda" rank="species" species="quadrimaculata" status="sp. nov.">
<emphasis id="B0E34B53305AFFB3FE85FA51BFCF08C0" box="[368,684,1475,1499]" italics="true" pageId="35" pageNumber="236">Madecassina quadrimaculata</emphasis>
Kavanaugh &amp; Rainio
</taxonomicName>
<taxonomicNameLabel id="ABD0F628305AFFB3FC6EFA51BEB708C1" box="[923,980,1475,1498]" pageId="35" pageNumber="236" rank="species">sp. n.</taxonomicNameLabel>
2015” [red label].
</materialsCitation>
</paragraph>
<paragraph id="82289741305AFFB3FF5EFA71B95008E0" blockId="35.[123,1174,1378,1725]" box="[171,1075,1507,1531]" pageId="35" pageNumber="236">
<materialsCitation id="32FF9D1C305AFFB3FF5EFA71B94F08E0" box="[171,1068,1507,1531]" country="Madagascar" location="Ranomafana National Park" pageId="35" pageNumber="236" specimenCount="1" stateProvince="Fianarantsoa" typeStatus="holotype">
<emphasis id="B0E34B53305AFFB3FF5EFA71BC3408E1" bold="true" box="[171,343,1507,1531]" pageId="35" pageNumber="236">
<smallCapsWord id="84CE019D305AFFB3FF5EFA71BD8708E1" baselines="1525,1525" box="[171,228,1507,1531]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Type" pageId="35" pageNumber="236">
<typeStatus id="5D2C29E3305AFFB3FF5EFA71BD8708E1" box="[171,228,1507,1531]" pageId="35" pageNumber="236">TYPE</typeStatus>
</smallCapsWord>
<smallCapsWord id="84CE019D305AFFB3FF19FA75BC3408E1" baselines="1525" box="[236,343,1511,1530]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="locality" pageId="35" pageNumber="236">LOCALITY</smallCapsWord>
</emphasis>
.—
<collectingCountry id="FA80D7D1305AFFB3FE8BFA71BC9C08E0" box="[382,511,1507,1531]" name="Madagascar" pageId="35" pageNumber="236">Madagascar</collectingCountry>
,
<collectingRegion id="405359A3305AFFB3FDF8FA71BF9F08E0" box="[525,764,1507,1531]" country="Madagascar" name="Fianarantsoa" pageId="35" pageNumber="236">Fianarantsoa Province</collectingRegion>
,
<location id="8748C19A305AFFB3FCFFFA71B94F08E0" LSID="urn:lsid:plazi:treatment:0A3E2657305AFFB5FF8EFA95B9710F61:8748C19A305AFFB3FCFFFA71B94F08E0" box="[778,1068,1507,1531]" country="Madagascar" name="Ranomafana National Park" pageId="35" pageNumber="236" stateProvince="Fianarantsoa">Ranomafana National Park</location>
</materialsCitation>
.
</paragraph>
<paragraph id="82289741305AFFB3FF5EF996B91D0B47" blockId="35.[123,1174,1378,1725]" pageId="35" pageNumber="236">
<emphasis id="B0E34B53305AFFB3FF5EF996BC8E0B01" bold="true" box="[171,493,1540,1564]" pageId="35" pageNumber="236">
<smallCapsWord id="84CE019D305AFFB3FF5EF996BC4D0B01" baselines="1558,1557" box="[171,302,1540,1564]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Derivation" pageId="35" pageNumber="236">DERIVATION</smallCapsWord>
<smallCapsWord id="84CE019D305AFFB3FEC1F995BC330B01" baselines="1557" box="[308,336,1543,1562]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="of" pageId="35" pageNumber="236">OF</smallCapsWord>
<smallCapsWord id="84CE019D305AFFB3FEA3F995BCCA0B01" baselines="1557" box="[342,425,1543,1562]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="species" pageId="35" pageNumber="236">SPECIES</smallCapsWord>
<smallCapsWord id="84CE019D305AFFB3FE45F995BC8E0B01" baselines="1557" box="[432,493,1543,1562]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="name" pageId="35" pageNumber="236">NAME</smallCapsWord>
</emphasis>
.— The species epithet,
<taxonomicName id="4597ECC2305AFFB3FD1AF991BEFA0B00" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[751,921,1539,1563]" class="Insecta" family="Carabidae" genus="Madecassina" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="236" phylum="Arthropoda" rank="species" species="quadrimaculata">
<emphasis id="B0E34B53305AFFB3FD1AF991BEFA0B00" box="[751,921,1539,1563]" italics="true" pageId="35" pageNumber="236">quadrimaculata</emphasis>
</taxonomicName>
, is an adjective derived from the Latin words,
<emphasis id="B0E34B53305AFFB3FE80F9B7BCB20B27" box="[373,465,1573,1596]" italics="true" pageId="35" pageNumber="236">quattuor</emphasis>
, meaning four, and
<emphasis id="B0E34B53305AFFB3FD47F9B6BE620B27" box="[690,769,1572,1596]" italics="true" pageId="35" pageNumber="236">macula</emphasis>
, meaning spot or spotted. The name refers to the two (subbasal and subapical) pairs of elytral spots seen in members of this species.
</paragraph>
<paragraph id="82289741305AFFB5FF5EF9F7BE840F03" blockId="35.[123,1174,1378,1725]" lastBlockId="37.[123,1174,219,634]" lastPageId="37" lastPageNumber="238" pageId="35" pageNumber="236">
<smallCapsWord id="84CE019D305AFFB3FF5EF9F7BC210B60" baselines="1655,1654" box="[171,322,1637,1661]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Recognition" pageId="35" pageNumber="236">
<emphasis id="B0E34B53305AFFB3FF5EF9F7BC210B60" bold="true" box="[171,322,1637,1661]" pageId="35" pageNumber="236">RECOGNITION</emphasis>
</smallCapsWord>
.— Size average for genus, SBL =
<quantity id="456F3AA4305AFFB3FD34F9F6BE770B67" box="[705,788,1636,1661]" metricMagnitude="-3" metricUnit="m" metricValue="5.2" pageId="35" pageNumber="236" unit="mm" value="5.2">5.2 mm</quantity>
. The
<typeStatus id="5D2C29E3305AFFB3FCA3F9F7BED00B66" box="[854,947,1637,1661]" pageId="35" pageNumber="236" type="holotype">holotype</typeStatus>
male of this species (
<figureCitation id="1AAC8BC4305AFFB3FF71F917BD870B86" box="[132,228,1669,1693]" captionStart="FIGURE" captionStartId="36.[155,167,1570,1589]" captionTargetBox="[144,1176,216,1552]" captionTargetId="figure-53@36.[120,1176,216,1567]" captionTargetPageId="36" captionText="FIGURE 20. Digital images of holotype male of Madecassina quadrimaculata Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B. Labels C–D. Aedeagus of male genitalia; C. Left lateral aspect; D. Dorsal aspect. Scale lines A = 1.0 mm, C–D = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13799487" httpUri="https://zenodo.org/record/13799487/files/figure.png" pageId="35" pageNumber="236">Fig. 20A</figureCitation>
) keys to
<taxonomicName id="4597ECC2305AFFB3FEB9F917BF9C0B86" authority="Alluaud (1899 b)" authorityName="Alluaud" authorityYear="1899" box="[332,767,1669,1693]" class="Insecta" family="Carabidae" genus="Madecassina" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="236" phylum="Arthropoda" rank="species" species="maculata">
<emphasis id="B0E34B53305AFFB3FEB9F917BF270B86" box="[332,580,1669,1693]" italics="true" pageId="35" pageNumber="236">Madecassina maculata</emphasis>
<bibRefCitation id="E606EAB0305AFFB3FDB9F917BF9C0B86" author="ALLUAUD, C." box="[588,767,1669,1693]" pageId="35" pageNumber="236" pagination="341 - 344" refId="ref28787" refString="ALLUAUD, C. 1899 b. Contributions a la faune entomologique de la Region malagache. 6 e Note. Bulletin de la Societe Entomologique de France 1899: 341 - 344." type="journal article" year="1899">Alluaud (1899b)</bibRefCitation>
</taxonomicName>
in Jeannel’s (1949) key based on the following combination of features: body black, forebody without metallic reflection, pronotum punctate laterally, and elytra shiny with four pale spots. It differs from members of
<taxonomicName id="4597ECC2305CFFB5FC06FF4EB9190DE8" authorityName="Alluaud" authorityYear="1899" box="[1011,1146,219,243]" class="Insecta" family="Carabidae" genus="Madecassina" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="238" phylum="Arthropoda" rank="species" species="maculata">
<emphasis id="B0E34B53305CFFB5FC06FF4EB9190DE8" box="[1011,1146,219,243]" italics="true" pageId="37" pageNumber="238">M. maculata</emphasis>
</taxonomicName>
in having a smaller body size (SBL = 5.5–6.0 mm in
<taxonomicName id="4597ECC2305CFFB5FD56FF6EBE480C0F" authorityName="Alluaud" authorityYear="1899" box="[675,811,252,276]" class="Insecta" family="Carabidae" genus="Madecassina" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="238" phylum="Arthropoda" rank="species" species="maculata">
<emphasis id="B0E34B53305CFFB5FD56FF6EBE480C0F" box="[675,811,252,276]" italics="true" pageId="37" pageNumber="238">M. maculata</emphasis>
</taxonomicName>
), the pronotum punctate laterally and laterobasally only, with conspicuous transverse grooves and ridges on disc (in
<taxonomicName id="4597ECC2305CFFB5FC1DFE8FB90D0C2F" authorityName="Alluaud" authorityYear="1899" box="[1000,1134,284,308]" class="Insecta" family="Carabidae" genus="Madecassina" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="238" phylum="Arthropoda" rank="species" species="maculata">
<emphasis id="B0E34B53305CFFB5FC1DFE8FB90D0C2F" box="[1000,1134,284,308]" italics="true" pageId="37" pageNumber="238">M. maculata</emphasis>
</taxonomicName>
the pronotum is punctate on the disc also, between the ridges, in contrast to what Jeannel said in his key), elytral intervals shinier due to more shallowly impressed isodiametric microsculpture (elytral intervals duller due to more deeply impressed isodiametric microsculpture in
<taxonomicName id="4597ECC2305CFFB5FC27FEEDB9380C8D" authorityName="Alluaud" authorityYear="1899" box="[978,1115,382,406]" class="Insecta" family="Carabidae" genus="Madecassina" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="238" phylum="Arthropoda" rank="species" species="maculata">
<emphasis id="B0E34B53305CFFB5FC27FEEDB9380C8D" box="[978,1115,382,406]" italics="true" pageId="37" pageNumber="238">M. maculata</emphasis>
</taxonomicName>
), the elytral spots much more distinct (much less so in
<taxonomicName id="4597ECC2305CFFB5FD65FE0DBE7B0CAC" authorityName="Alluaud" authorityYear="1899" box="[656,792,415,439]" class="Insecta" family="Carabidae" genus="Madecassina" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="238" phylum="Arthropoda" rank="species" species="maculata">
<emphasis id="B0E34B53305CFFB5FD65FE0DBE7B0CAC" box="[656,792,415,439]" italics="true" pageId="37" pageNumber="238">M. maculata</emphasis>
</taxonomicName>
) and median lobe of male genitalia with apex hooked dorsally and thicker in lateral aspect (
<figureCitation id="1AAC8BC4305CFFB5FD16FE2DBE200CCC" box="[739,835,447,471]" captionStart="FIGURE" captionStartId="36.[155,167,1570,1589]" captionTargetBox="[144,1176,216,1552]" captionTargetId="figure-53@36.[120,1176,216,1567]" captionTargetPageId="36" captionText="FIGURE 20. Digital images of holotype male of Madecassina quadrimaculata Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B. Labels C–D. Aedeagus of male genitalia; C. Left lateral aspect; D. Dorsal aspect. Scale lines A = 1.0 mm, C–D = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13799487" httpUri="https://zenodo.org/record/13799487/files/figure.png" pageId="37" pageNumber="238">Fig. 20C</figureCitation>
) and slightly swollen and bent right in dorsal aspect (
<figureCitation id="1AAC8BC4305CFFB5FE9FFE72BCAB0CEC" box="[362,456,480,504]" captionStart="FIGURE" captionStartId="36.[155,167,1570,1589]" captionTargetBox="[144,1176,216,1552]" captionTargetId="figure-53@36.[120,1176,216,1567]" captionTargetPageId="36" captionText="FIGURE 20. Digital images of holotype male of Madecassina quadrimaculata Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B. Labels C–D. Aedeagus of male genitalia; C. Left lateral aspect; D. Dorsal aspect. Scale lines A = 1.0 mm, C–D = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13799487" httpUri="https://zenodo.org/record/13799487/files/figure.png" pageId="37" pageNumber="238">Fig. 20D</figureCitation>
) (apex hooked but thinner in lateral aspect and more parallel-sided and straight in dorsal aspect in
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<emphasis id="B0E34B53305CFFB5FE3FFD93BF310F03" box="[458,594,512,536]" italics="true" pageId="37" pageNumber="238">M. maculata</emphasis>
</taxonomicName>
males) (see Jeannel 1949, Fig. 474f).
</paragraph>
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<paragraph id="82289741305DFFB4FF6EF9B0BC630B73" blockId="36.[123,1173,1570,1640]" pageId="36" pageNumber="237">
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20. Digital images of holotype male of
<taxonomicName id="4597ECC2305DFFB4FDDBF9B0BE8B0B2E" authority="Kavanaugh &amp; Rainio" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[558,1000,1570,1589]" class="Insecta" family="Carabidae" genus="Madecassina" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="237" phylum="Arthropoda" rank="species" species="quadrimaculata" status="sp. nov.">
<emphasis id="B0E34B53305DFFB4FDDBF9B0BE4F0B2E" box="[558,812,1570,1589]" italics="true" pageId="36" pageNumber="237">Madecassina quadrimaculata</emphasis>
Kavanaugh &amp; Rainio
</taxonomicName>
<taxonomicNameLabel id="ABD0F628305DFFB4FC18F9B0B94C0B2E" box="[1005,1071,1570,1589]" pageId="36" pageNumber="237" rank="species">sp. nov.</taxonomicNameLabel>
A. Habitus, dorsal aspect; B. Labels C–D. Aedeagus of male genitalia; C. Left lateral aspect; D. Dorsal aspect. Scale lines A = 1.0 mm, C–D = 0.5 mm.
</paragraph>
</caption>
<paragraph id="82289741305CFFB5FF5EFDB3B9690F22" blockId="37.[123,1174,219,634]" box="[171,1034,545,570]" pageId="37" pageNumber="238">
<emphasis id="B0E34B53305CFFB5FF5EFDB3BC920F2C" bold="true" box="[171,497,545,569]" pageId="37" pageNumber="238">
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</emphasis>
.— At present, known only from the
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locality.
</paragraph>
<paragraph id="82289741305CFFB5FF5EFDD0B9710F61" blockId="37.[123,1174,219,634]" pageId="37" pageNumber="238">
<emphasis id="B0E34B53305CFFB5FF5EFDD0BCC00F43" bold="true" box="[171,419,578,602]" pageId="37" pageNumber="238">
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<smallCapsWord id="84CE019D305CFFB5FEFAFDD7BCC00F43" baselines="595" box="[271,419,581,600]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="distribution" pageId="37" pageNumber="238">DISTRIBUTION</smallCapsWord>
</emphasis>
.— The unique
<typeStatus id="5D2C29E3305CFFB5FDBCFDD0BFC50F41" box="[585,678,578,602]" pageId="37" pageNumber="238" type="holotype">holotype</typeStatus>
was collected in montane rainforest near Tsinjorano village at a distance of
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inside the forest edge at an elevation of
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.
</paragraph>
</subSubSection>
</treatment>
</document>
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