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<document id="AF16173E94E0BE50699BD4CC1B0C80B7" ID-DOI="10.5281/zenodo.13799439" ID-ISSN="0068-547X" ID-Zenodo-Dep="13799439" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722538212676" checkinUser="felipe" docAuthor="Kavanaugh, David H. &amp; Rainio, Johanna" docDate="2016" docId="0A3E26573057FFA0FF8EFF49BC7D0FEE" docLanguage="en" docName="ProcCalifAcadSci.63.7.201-268.pdf" docOrigin="Proceedings of the California Academy of Sciences 63 (7)" docStyle="DocumentStyle:E411DA009AF5E7F9A7E023431E2B7D5B.2:ProcCalifAcadSci.2015-.journal_article" docStyleId="E411DA009AF5E7F9A7E023431E2B7D5B" docStyleName="ProcCalifAcadSci.2015-.journal_article" docStyleVersion="2" docTitle="Eurydera simplica Kavanaugh and Rainio 2016, sp. nov." docType="treatment" docVersion="2" lastPageNumber="249" masterDocId="F6075E2F3079FF90FFF5FF92BD630D1B" masterDocTitle="Twenty-six New Species of Predaceous Ground Beetles (Coleoptera: Adephaga: Carabidae) from Ranomafana National Park, Madagascar" masterLastPageNumber="268" masterPageNumber="201" pageNumber="247" updateTime="1726771873529" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="90485B16CECF75CC052473BA9B0AE6FB" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="13854CD14FA6BABF71A096407A7EF8E3">
<mods:title id="B9E92EB281D32A1800C80960C6E439BD">Twenty-six New Species of Predaceous Ground Beetles (Coleoptera: Adephaga: Carabidae) from Ranomafana National Park, Madagascar</mods:title>
</mods:titleInfo>
<mods:name id="1B217914FF731D55D2C8EC8426EC88B5" type="personal">
<mods:role id="FED20AF0AD1B1156C95BCFC77DBFCA31">
<mods:roleTerm id="5AD2EF4553420C8D5EC046E984B98D72">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="070E19733D454B6333EC30F1360D505C">Kavanaugh, David H.</mods:namePart>
<mods:affiliation id="871CF8B2A604C1E26CB51700F81D8FAE">Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA; Email: dkavanaugh @ calacademy. org; &amp; Corresponding author: David H. Kavanaugh (dkavanaugh @ calacademy. org)</mods:affiliation>
</mods:name>
<mods:name id="80CDC937E38A11E85BCB2EDFF66EA608" type="personal">
<mods:role id="564117CA54446D9493B5C033FA83987E">
<mods:roleTerm id="E14A02C509B57D9A2D3FE57E640453A1">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7D2661C80DAACBADF6F9E4429506883A">Rainio, Johanna</mods:namePart>
<mods:affiliation id="62E02399E86861414CB5A1F744544B19">Department of Ecology and Systematics, University of Helsinki, P. O. Box 65, FIN- 00014 Helsinki, Finland; Email: kjohannarainio @ gmail. com</mods:affiliation>
</mods:name>
<mods:typeOfResource id="DBDAE005BF46434389320DA9AD01D7BF">text</mods:typeOfResource>
<mods:relatedItem id="F552D1A5FCB633865F5BB0667FBF59C3" type="host">
<mods:titleInfo id="2EE129EFF3B6A90FEBBEF8E4CFAB6D30">
<mods:title id="154DF7F55A16407DFA3440F1F18F1A43">Proceedings of the California Academy of Sciences</mods:title>
</mods:titleInfo>
<mods:part id="DA28A8A3C624FE32CED5FD9F69D36D46">
<mods:date id="C0177D8195A09BFABCA4054792966F74">2016</mods:date>
<mods:detail id="A2B7352CC50AC455F7AC58C098A73218" type="pubDate">
<mods:number id="F87A871B47A78DA85B63677ECB5FD159">2016-04-29</mods:number>
</mods:detail>
<mods:detail id="6C08026261DABFF7518B585C008ED06D" type="volume">
<mods:number id="91D8558060A46FE2A0F47B5BFAFAF513">63</mods:number>
</mods:detail>
<mods:detail id="4E270372CA704211AED3AA8F5724D498" type="issue">
<mods:number id="367F1094CE763A51A8850AFC7F5798D4">7</mods:number>
</mods:detail>
<mods:extent id="996F400A495D51F51EFF50358D331FC9" unit="page">
<mods:start id="DE2610BC2EEF3153D8455FE1CC327CE0">201</mods:start>
<mods:end id="9D0FC72389FE506FFE94F48BC5D90EF0">268</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="29A977608CC3731C6033A95F5D9DD287">journal article</mods:classification>
<mods:identifier id="453531765D73ED332605AAB187AC541C" type="DOI">10.5281/zenodo.13799439</mods:identifier>
<mods:identifier id="58EF278B63BC5BA4425921779312AA2C" type="ISSN">0068-547X</mods:identifier>
<mods:identifier id="E5F5E491DD638783DC8D5212D5012B62" type="Zenodo-Dep">13799439</mods:identifier>
</mods:mods>
<treatment id="0A3E26573057FFA0FF8EFF49BC7D0FEE" LSID="urn:lsid:plazi:treatment:0A3E26573057FFA0FF8EFF49BC7D0FEE" httpUri="http://treatment.plazi.org/id/0A3E26573057FFA0FF8EFF49BC7D0FEE" lastPageId="48" lastPageNumber="249" pageId="46" pageNumber="247">
<subSubSection id="CA8DC4CA3057FFBEFF8EFF49BFD90DE8" box="[123,698,219,243]" pageId="46" pageNumber="247" type="nomenclature">
<paragraph id="822897413057FFBEFF8EFF49BFD90DE8" blockId="46.[123,698,219,275]" box="[123,698,219,243]" pageId="46" pageNumber="247">
<heading id="D960202D3057FFBEFF8EFF49BFD90DE8" box="[123,698,219,243]" fontSize="10" level="2" pageId="46" pageNumber="247" reason="4">
<emphasis id="B0E34B533057FFBEFF8EFF49BFD90DE8" bold="true" box="[123,698,219,243]" pageId="46" pageNumber="247">
<taxonomicName id="4597ECC23057FFBEFF8EFF49BF350DE8" ID-CoL="3D44Z" authority="Kavanaugh and Rainio" authorityName="Kavanaugh and Rainio" authorityYear="2016" box="[123,598,219,243]" class="Insecta" family="Carabidae" genus="Eurydera" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="247" phylum="Arthropoda" rank="species" species="simplica" status="sp. nov.">
<emphasis id="B0E34B533057FFBEFF8EFF49BC250DE8" bold="true" box="[123,326,219,243]" italics="true" pageId="46" pageNumber="247">Eurydera simplica</emphasis>
Kavanaugh and Rainio
</taxonomicName>
,
<taxonomicNameLabel id="ABD0F6283057FFBEFD91FF49BFD90DE8" box="[612,698,219,243]" pageId="46" pageNumber="247" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="CA8DC4CA3057FFBEFF8EFF6FBE3A0C7E" pageId="46" pageNumber="247" type="description">
<paragraph id="822897413057FFBEFF8EFF6FBC660C08" blockId="46.[123,698,219,275]" box="[123,261,253,275]" pageId="46" pageNumber="247">
<figureCitation id="1AAC8BC43057FFBEFF8EFF6FBC660C08" box="[123,261,253,275]" captionStart-0="FIGURE" captionStart-1="FIGURE" captionStartId-0="46.[155,167,1667,1686]" captionStartId-1="47.[155,167,783,802]" captionTargetBox-0="[149,1176,526,1636]" captionTargetBox-1="[120,1164,230,765]" captionTargetId-0="figure-537@46.[120,1176,528,1664]" captionTargetId-1="figure-2124@47.[120,1176,216,765]" captionTargetPageId-0="46" captionTargetPageId-1="47" captionText-0="FIGURE 27. Digital images of holotype male of Eurydera simplica Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B. Labels. Scale line = 1.0 mm." captionText-1="FIGURE 28. Digital images of holotype male of Eurydera simplica Kavanaugh &amp; Rainio sp. nov. A. Head, dorsal aspect; B–C. Median lobe of aedeagus of genitalia; B. Left lateral aspect; C. Dorsal aspect. Scale lines = 0.5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.13799501" figureDoi-1="http://doi.org/10.5281/zenodo.13799503" httpUri-0="https://zenodo.org/record/13799501/files/figure.png" httpUri-1="https://zenodo.org/record/13799503/files/figure.png" pageId="46" pageNumber="247">Figures 27–28</figureCitation>
</paragraph>
<paragraph id="822897413057FFBEFF8EFEBFBE710C58" blockId="46.[123,857,301,358]" box="[123,786,301,323]" pageId="46" pageNumber="247">
<taxonomicName id="4597ECC23057FFBEFF8EFEBFBD940C58" box="[123,247,301,323]" class="Insecta" family="Carabidae" genus="Eurydera" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="247" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B0E34B533057FFBEFF8EFEBFBD940C58" box="[123,247,301,323]" italics="true" pageId="46" pageNumber="247">Eurydera sp.</emphasis>
</taxonomicName>
; Rainio and Niemelä 2006: 227 (informal designation).
</paragraph>
<paragraph id="822897413057FFBEFF8EFEC2BE3A0C7E" blockId="46.[123,857,301,358]" box="[123,857,336,358]" pageId="46" pageNumber="247">
<emphasis id="B0E34B533057FFBEFF8EFEC2BC410C7E" box="[123,290,336,357]" italics="true" pageId="46" pageNumber="247">
<taxonomicName id="4597ECC23057FFBEFF8EFEC2BDB50C7E" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[123,214,336,357]" class="Insecta" family="Carabidae" genus="Eurydera" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="247" phylum="Arthropoda" rank="genus" status="n. sp.">Eurydera</taxonomicName>
<taxonomicNameLabel id="ABD0F6283057FFBEFF29FEC2BC730C7E" box="[220,272,336,357]" pageId="46" pageNumber="247" rank="species">n. sp.</taxonomicNameLabel>
3
</emphasis>
; Rainio 2009: 31, Rainio 2012: 72 (informal designation).
</paragraph>
</subSubSection>
<subSubSection id="CA8DC4CA3057FFA0FF5EFE1CBC7D0FEE" lastPageId="48" lastPageNumber="249" pageId="46" pageNumber="247" type="materials_examined">
<paragraph id="822897413057FFBFFF5EFE1CBFAA0ECC" blockId="46.[123,1174,398,526]" lastBlockId="47.[123,1174,864,1719]" lastPageId="47" lastPageNumber="248" pageId="46" pageNumber="247">
<emphasis id="B0E34B533057FFBEFF5EFE1CBC3D0CBF" bold="true" box="[171,350,398,422]" pageId="46" pageNumber="247">
<smallCapsWord id="84CE019D3057FFBEFF5EFE1CBD870CBF" baselines="416,415" box="[171,228,398,422]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Type" pageId="46" pageNumber="247">TYPE</smallCapsWord>
<smallCapsWord id="84CE019D3057FFBEFF04FE03BC3D0CBF" baselines="415" box="[241,350,401,420]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="material" pageId="46" pageNumber="247">MATERIAL</smallCapsWord>
</emphasis>
.—
<materialsCitation id="32FF9D1C3057FFBEFE73FE1CBDB00F15" collectingDate="2004-03-08" collectorName="F. Ratalata &amp; J. Rainio" country="Madagascar" elevation="900" location="Talatakely" pageId="46" pageNumber="247" specimenCount="1" typeStatus="holotype">
<typeStatus id="5D2C29E33057FFBEFE73FE1CBC8A0CBD" box="[390,489,398,422]" pageId="46" pageNumber="247" type="holotype">Holotype</typeStatus>
(
<figureCitation id="1AAC8BC43057FFBEFE0BFE1CBFE30CBE" box="[510,640,398,422]" captionStart="FIGURE" captionStartId="46.[155,167,1667,1686]" captionTargetBox="[149,1176,526,1636]" captionTargetId="figure-537@46.[120,1176,528,1664]" captionTargetPageId="46" captionText="FIGURE 27. Digital images of holotype male of Eurydera simplica Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B. Labels. Scale line = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13799501" httpUri="https://zenodo.org/record/13799501/files/figure.png" pageId="46" pageNumber="247">Figs.27A–B</figureCitation>
), a male, in CAS, labeled: “
<collectingCountry id="FA80D7D13057FFBEFC24FE1CB9EC0CBD" box="[977,1167,398,422]" name="Madagascar" pageId="46" pageNumber="247">MADAGASCAR</collectingCountry>
, Ranomafana NP.,
<location id="8748C19A3057FFBEFEC1FE22BCC20CD3" LSID="urn:lsid:plazi:treatment:0A3E26573057FFA0FF8EFF49BC7D0FEE:8748C19A3057FFBEFEC1FE22BCC20CD3" box="[308,417,432,456]" country="Madagascar" name="Talatakely" pageId="46" pageNumber="247">Talatakely</location>
,
<quantity id="456F3AA43057FFBEFE58FE22BC930CD3" box="[429,496,432,456]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="46" pageNumber="247" unit="m" value="900.0">
<elevation id="09BA70723057FFBEFE58FE22BC930CD3" box="[429,496,432,456]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="46" pageNumber="247" unit="m" value="900.0">900 m</elevation>
</quantity>
, Trail SP at
<quantity id="456F3AA43057FFBEFD85FE22BFC60CD3" box="[624,677,432,456]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="46" pageNumber="247" unit="m" value="50.0">
<elevation id="09BA70723057FFBEFD85FE22BFC60CD3" box="[624,677,432,456]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="46" pageNumber="247" unit="m" value="50.0">50 m</elevation>
</quantity>
mark,
<date id="F629B1813057FFBEFD1BFE22BE050CD3" box="[750,870,432,456]" pageId="46" pageNumber="247" value="2004-03-08">
<collectingDate id="E66D48693057FFBEFD1BFE22BE050CD3" box="[750,870,432,456]" pageId="46" pageNumber="247" value="2004-03-08">08.03.2004</collectingDate>
</date>
”/ “collected by hand in daytime in secondary montane rainforest,
<collectorName id="2F62F2973057FFBEFDD9FE41BFC10CF0" box="[556,674,467,491]" pageId="46" pageNumber="247">F. Ratalata</collectorName>
&amp;
<collectorName id="2F62F2973057FFBEFD38FE41BE520CF0" box="[717,817,467,491]" pageId="46" pageNumber="247">J. Rainio</collectorName>
collectors”/ “106” [handwritten label]/ “
</materialsCitation>
<materialsCitation id="32FF9D1C3057FFBFFF26FE64BE120EAC" collectingDate="2000-06-15" collectionCode="MNHN" collectorName="F. Ratalata &amp; J. Rainio" country="Madagascar" elevation="900" lastPageId="47" lastPageNumber="248" location="Talatakely" pageId="46" pageNumber="247" specimenCount="2" typeStatus="holotype">
<typeStatus id="5D2C29E33057FFBEFF26FE64BC030F15" box="[211,352,502,526]" pageId="46" pageNumber="247" type="holotype">HOLOTYPE</typeStatus>
<taxonomicName id="4597ECC23057FFBEFE92FE67BE7F0F16" authority="Kavanaugh &amp; Rainio" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[359,796,501,526]" class="Insecta" family="Carabidae" genus="Eurydera" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="247" phylum="Arthropoda" rank="species" species="simplica" status="sp. nov.">
<emphasis id="B0E34B533057FFBEFE92FE67BF4D0F16" box="[359,558,501,525]" italics="true" pageId="46" pageNumber="247">Eurydera simplica</emphasis>
Kavanaugh &amp; Rainio
</taxonomicName>
<taxonomicNameLabel id="ABD0F6283057FFBEFCD1FE64BE3D0F16" box="[804,862,502,525]" pageId="46" pageNumber="247" rank="species">sp. n.</taxonomicNameLabel>
2015” [red label].
<typeStatus id="5D2C29E33057FFBEFBDBFE64B9F50F15" box="[1070,1174,502,526]" pageId="46" pageNumber="247" type="paratype">Paratypes</typeStatus>
(only 1): a female (in
<collectionCode id="E4860F843056FFBFFE98FCF3BCDD0E63" box="[365,446,865,888]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="47" pageNumber="248">MNHN</collectionCode>
) labeled ““
<collectingCountry id="FA80D7D13056FFBFFDC8FCF3BF980E62" box="[573,763,865,889]" name="Madagascar" pageId="47" pageNumber="248">MADAGASCAR</collectingCountry>
, Ranomafana NP.,
<location id="8748C19A3056FFBFFC38FCF2B9590E63" LSID="urn:lsid:plazi:treatment:0A3E26573057FFA0FF8EFF49BC7D0FEE:8748C19A3056FFBFFC38FCF2B9590E63" box="[973,1082,864,888]" country="Madagascar" name="Talatakely" pageId="47" pageNumber="248">Talatakely</location>
,
<quantity id="456F3AA43056FFBFFBBCFCF3B9EC0E63" box="[1097,1167,865,889]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="47" pageNumber="248" unit="m" value="900.0">
<elevation id="09BA70723056FFBFFBBCFCF3B9EC0E63" box="[1097,1167,865,889]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="47" pageNumber="248" unit="m" value="900.0">900 m</elevation>
</quantity>
, Trail B at
<quantity id="456F3AA43056FFBFFF1EFCEDBC5D0E8D" box="[235,318,895,919]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="47" pageNumber="248" unit="m" value="1000.0">
<elevation id="09BA70723056FFBFFF1EFCEDBC5D0E8D" box="[235,318,895,919]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="47" pageNumber="248" unit="m" value="1000.0">1000 m</elevation>
</quantity>
mark,
<date id="F629B1813056FFBFFE79FCEDBF600E8C" box="[396,515,895,919]" pageId="47" pageNumber="248" value="2000-06-15">
<collectingDate id="E66D48693056FFBFFE79FCEDBF600E8C" box="[396,515,895,919]" pageId="47" pageNumber="248" value="2000-06-15">15.06.2000</collectingDate>
</date>
”/ “collected by hand in daytime in secondary montane rainforest,
<collectorName id="2F62F2973056FFBFFF30FC32BC560EAC" box="[197,309,927,951]" pageId="47" pageNumber="248">F. Ratalata</collectorName>
&amp;
<collectorName id="2F62F2973056FFBFFEADFC32BCD50EAC" box="[344,438,927,951]" pageId="47" pageNumber="248">J. Rainio</collectorName>
collectors”/ “7/13” [handwritten label]/ “
</materialsCitation>
<typeStatus id="5D2C29E33056FFBFFC84FC0DBE960EAC" box="[881,1013,927,951]" pageId="47" pageNumber="248" type="paratype">PARATYPE</typeStatus>
<taxonomicName id="4597ECC23056FFBFFC09FC0DBCF80ECC" authority="Kavanaugh &amp; Rainio" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" class="Insecta" family="Carabidae" genus="Eurydera" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="248" phylum="Arthropoda" rank="species" species="simplica" status="sp. nov.">
<emphasis id="B0E34B533056FFBFFC09FC0DBDD30ECC" italics="true" pageId="47" pageNumber="248">Eurydera simplica</emphasis>
Kavanaugh &amp; Rainio
</taxonomicName>
<taxonomicNameLabel id="ABD0F6283056FFBFFE56FC52BCBE0ECC" box="[419,477,960,983]" pageId="47" pageNumber="248" rank="species">sp. n.</taxonomicNameLabel>
2015” [yellow label].
</paragraph>
<caption id="D6E8C7C93057FFBEFF6EF911BCA80BAB" ID-DOI="http://doi.org/10.5281/zenodo.13799501" ID-Zenodo-Dep="13799501" httpUri="https://zenodo.org/record/13799501/files/figure.png" pageId="46" pageNumber="247" startId="46.[155,167,1667,1686]" targetBox="[149,1176,526,1636]" targetPageId="46" targetType="figure">
<paragraph id="822897413057FFBEFF6EF911BCA80BAB" blockId="46.[123,1173,1666,1712]" pageId="46" pageNumber="247">
<smallCapsWord id="84CE019D3057FFBEFF6EF911BDBB0B8F" baselines="1681,1680" box="[155,216,1667,1686]" lowerCaseFontSize="6" mainFontSize="8" normCase="title" normString="Figure" pageId="46" pageNumber="247">FIGURE</smallCapsWord>
27. Digital images of holotype male of
<taxonomicName id="4597ECC23057FFBEFDB5F910BEC00B8E" authority="Kavanaugh &amp; Rainio" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[576,931,1666,1686]" class="Insecta" family="Carabidae" genus="Eurydera" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="247" phylum="Arthropoda" rank="species" species="simplica" status="sp. nov.">
<emphasis id="B0E34B533057FFBEFDB5F910BF820B8E" box="[576,737,1666,1685]" italics="true" pageId="46" pageNumber="247">Eurydera simplica</emphasis>
Kavanaugh &amp; Rainio
</taxonomicName>
<taxonomicNameLabel id="ABD0F6283057FFBEFC5EF911BE8D0B8D" box="[939,1006,1667,1686]" pageId="46" pageNumber="247" rank="species">sp. nov.</taxonomicNameLabel>
A. Habitus, dorsal aspect; B. Labels. Scale line = 1.0 mm.
</paragraph>
</caption>
<caption id="D6E8C7C93056FFBFFF6EFC9DB96A0E27" ID-DOI="http://doi.org/10.5281/zenodo.13799503" ID-Zenodo-Dep="13799503" httpUri="https://zenodo.org/record/13799503/files/figure.png" pageId="47" pageNumber="248" startId="47.[155,167,783,802]" targetBox="[120,1164,230,765]" targetPageId="47" targetType="figure">
<paragraph id="822897413056FFBFFF6EFC9DB9F60E39" blockId="47.[123,1173,783,828]" box="[155,1173,783,802]" pageId="47" pageNumber="248">
<smallCapsWord id="84CE019D3056FFBFFF6EFC9DBDBB0E3A" baselines="797,797" box="[155,216,783,802]" lowerCaseFontSize="6" mainFontSize="8" normCase="title" normString="Figure" pageId="47" pageNumber="248">FIGURE</smallCapsWord>
28. Digital images of holotype male of
<taxonomicName id="4597ECC23056FFBFFDD9FC9DBEE60E39" authority="Kavanaugh &amp; Rainio" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[556,901,783,802]" class="Insecta" family="Carabidae" genus="Eurydera" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="248" phylum="Arthropoda" rank="species" species="simplica" status="sp. nov.">
<emphasis id="B0E34B533056FFBFFDD9FC9DBFA90E39" box="[556,714,783,802]" italics="true" pageId="47" pageNumber="248">Eurydera simplica</emphasis>
Kavanaugh &amp; Rainio
</taxonomicName>
<taxonomicNameLabel id="ABD0F6283056FFBFFC7FFC9DBEA80E39" box="[906,971,783,802]" pageId="47" pageNumber="248" rank="species">sp. nov.</taxonomicNameLabel>
A. Head, dorsal aspect;
</paragraph>
<paragraph id="822897413056FFBFFF8EFCBBB96A0E27" blockId="47.[123,1173,783,828]" box="[123,1033,808,828]" pageId="47" pageNumber="248">B–C. Median lobe of aedeagus of genitalia; B. Left lateral aspect; C. Dorsal aspect. Scale lines = 0.5 mm.</paragraph>
</caption>
<paragraph id="822897413056FFBFFF5EFC4DB9500EED" blockId="47.[123,1174,864,1719]" box="[171,1075,991,1015]" pageId="47" pageNumber="248">
<materialsCitation id="32FF9D1C3056FFBFFF5EFC4DB94F0EEC" box="[171,1068,991,1015]" country="Madagascar" location="Ranomafana National Park" pageId="47" pageNumber="248" specimenCount="1" stateProvince="Fianarantsoa" typeStatus="holotype">
<emphasis id="B0E34B533056FFBFFF5EFC4DBC340EEE" bold="true" box="[171,343,991,1015]" pageId="47" pageNumber="248">
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<typeStatus id="5D2C29E33056FFBFFF5EFC4DBD870EED" box="[171,228,991,1015]" pageId="47" pageNumber="248">TYPE</typeStatus>
</smallCapsWord>
<smallCapsWord id="84CE019D3056FFBFFF19FC70BC340EEE" baselines="1008" box="[236,343,994,1013]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="locality" pageId="47" pageNumber="248">LOCALITY</smallCapsWord>
</emphasis>
.—
<collectingCountry id="FA80D7D13056FFBFFE8BFC4DBC9C0EEC" box="[382,511,991,1015]" name="Madagascar" pageId="47" pageNumber="248">Madagascar</collectingCountry>
,
<collectingRegion id="405359A33056FFBFFDF8FC4DBF9F0EEC" box="[525,764,991,1015]" country="Madagascar" name="Fianarantsoa" pageId="47" pageNumber="248">Fianarantsoa Province</collectingRegion>
,
<location id="8748C19A3056FFBFFCFFFC4DB94F0EEC" LSID="urn:lsid:plazi:treatment:0A3E26573057FFA0FF8EFF49BC7D0FEE:8748C19A3056FFBFFCFFFC4DB94F0EEC" box="[778,1068,991,1015]" country="Madagascar" name="Ranomafana National Park" pageId="47" pageNumber="248" stateProvince="Fianarantsoa">Ranomafana National Park</location>
</materialsCitation>
.
</paragraph>
<paragraph id="822897413056FFBFFF5EFC6DBDD0096C" blockId="47.[123,1174,864,1719]" pageId="47" pageNumber="248">
<emphasis id="B0E34B533056FFBFFF5EFC6DBC90090E" bold="true" box="[171,499,1023,1047]" pageId="47" pageNumber="248">
<smallCapsWord id="84CE019D3056FFBFFF5EFC6DBC4D090E" baselines="1041,1040" box="[171,302,1023,1047]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Derivation" pageId="47" pageNumber="248">DERIVATION</smallCapsWord>
<smallCapsWord id="84CE019D3056FFBFFEC3FB90BC31090E" baselines="1040" box="[310,338,1026,1045]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="of" pageId="47" pageNumber="248">OF</smallCapsWord>
<smallCapsWord id="84CE019D3056FFBFFEAFFB90BCCE090E" baselines="1040" box="[346,429,1026,1045]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="species" pageId="47" pageNumber="248">SPECIES</smallCapsWord>
<smallCapsWord id="84CE019D3056FFBFFE43FB90BC90090E" baselines="1040" box="[438,499,1026,1045]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="name" pageId="47" pageNumber="248">NAME</smallCapsWord>
</emphasis>
.— The species epithet,
<taxonomicName id="4597ECC23056FFBFFD0FFC6DBE37090C" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[762,852,1023,1047]" class="Insecta" family="Carabidae" genus="Eurydera" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="248" phylum="Arthropoda" rank="species" species="simplica">
<emphasis id="B0E34B533056FFBFFD0FFC6DBE37090C" box="[762,852,1023,1047]" italics="true" pageId="47" pageNumber="248">simplica</emphasis>
</taxonomicName>
, is an adjective derived from the Latin word,
<emphasis id="B0E34B533056FFBFFEDEFB8DBC1F092C" box="[299,380,1055,1079]" italics="true" pageId="47" pageNumber="248">simplex</emphasis>
, meaning simple. The name refers to the relative simplicity of form and surface macrosculpture of member of this species compared with those of other species of
<taxonomicName id="4597ECC23056FFBFFBACFBD2BDCF096C" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" class="Insecta" family="Carabidae" genus="Eurydera" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="248" phylum="Arthropoda" rank="genus">
<emphasis id="B0E34B533056FFBFFBACFBD2BDCF096C" italics="true" pageId="47" pageNumber="248">Eurydera</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="822897413056FFA0FF5EFBEDB9F50F6F" blockId="47.[123,1174,864,1719]" lastBlockId="48.[123,1174,219,758]" lastPageId="48" lastPageNumber="249" pageId="47" pageNumber="248">
<smallCapsWord id="84CE019D3056FFBFFF5EFBEDBC21098E" baselines="1169,1168" box="[171,322,1151,1175]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Recognition" pageId="47" pageNumber="248">
<emphasis id="B0E34B533056FFBFFF5EFBEDBC21098E" bold="true" box="[171,322,1151,1175]" pageId="47" pageNumber="248">RECOGNITION</emphasis>
</smallCapsWord>
.— Size average for genus, SBL of male =
<quantity id="456F3AA43056FFBFFCFCFBEDBE04098D" box="[777,871,1151,1175]" metricMagnitude="-2" metricUnit="m" metricValue="1.31" pageId="47" pageNumber="248" unit="mm" value="13.1">13.1 mm</quantity>
, of female = 12.4. Members of this species (
<figureCitation id="1AAC8BC43056FFBFFED0FB0DBCEC09AC" box="[293,399,1183,1207]" captionStart="FIGURE" captionStartId="46.[155,167,1667,1686]" captionTargetBox="[149,1176,526,1636]" captionTargetId="figure-537@46.[120,1176,528,1664]" captionTargetPageId="46" captionText="FIGURE 27. Digital images of holotype male of Eurydera simplica Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B. Labels. Scale line = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13799501" httpUri="https://zenodo.org/record/13799501/files/figure.png" pageId="47" pageNumber="248">Figs. 27A</figureCitation>
) are unlike those of any other species in the genus. In Jeannel’s (1949) key species of
<taxonomicName id="4597ECC23056FFBFFEE9FB2DBCE209CC" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[284,385,1215,1239]" class="Insecta" family="Carabidae" genus="Eurydera" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="248" phylum="Arthropoda" rank="genus">
<emphasis id="B0E34B533056FFBFFEE9FB2DBCE209CC" box="[284,385,1215,1239]" italics="true" pageId="47" pageNumber="248">Eurydera</emphasis>
</taxonomicName>
, they key to his
<taxonomicName id="4597ECC23056FFBFFDC2FB2DBFF209CC" baseAuthorityName="Klug" baseAuthorityYear="1833" box="[567,657,1215,1239]" class="Insecta" family="Carabidae" genus="Eurydera" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="248" phylum="Arthropoda" rank="species" species="unicolor">
<emphasis id="B0E34B533056FFBFFDC2FB2DBFF209CC" box="[567,657,1215,1239]" italics="true" pageId="47" pageNumber="248">unicolor</emphasis>
</taxonomicName>
species group, based on the following features: size medium; head (
<figureCitation id="1AAC8BC43056FFBFFEA1FB4DBCD709EC" box="[340,436,1247,1271]" captionStart="FIGURE" captionStartId="47.[155,167,783,802]" captionTargetBox="[120,1164,230,765]" captionTargetId="figure-2124@47.[120,1176,216,765]" captionTargetPageId="47" captionText="FIGURE 28. Digital images of holotype male of Eurydera simplica Kavanaugh &amp; Rainio sp. nov. A. Head, dorsal aspect; B–C. Median lobe of aedeagus of genitalia; B. Left lateral aspect; C. Dorsal aspect. Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13799503" httpUri="https://zenodo.org/record/13799503/files/figure.png" pageId="47" pageNumber="248">Fig. 28A</figureCitation>
) with posterior part of frons formed as a broad, flat, smooth, elevat- ed plateau, laterally delimited by deep furrows extended posteriorly beyond the level of the posterior margin of the eye then slightly and arcuately convergent posteriorly and terminated abruptly at the level of the posterior margin of the ocular swelling and just less than halfway from the midline to the lateral margin of the head, area between the furrow smooth and only slightly elevated; mandibles moderately short and broad; median lobe of male genitalia without a large, angulate projection on the ventral margin of the shaft near the base. Jeannel included only
<taxonomicName id="4597ECC23056FFBFFC3BFA0DBC6208CC" authority="(Klug 1833)" baseAuthorityName="Klug" baseAuthorityYear="1833" class="Insecta" family="Carabidae" genus="Eurydera" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="248" phylum="Arthropoda" rank="species" species="unicolor">
<emphasis id="B0E34B533056FFBFFC3BFA0DB9F608AC" box="[974,1173,1439,1463]" italics="true" pageId="47" pageNumber="248">Eurydera unicolor</emphasis>
(
<bibRefCitation id="E606EAB03056FFBFFF71FA2DBD9B08CC" author="KLUG, J. C. F." box="[132,248,1471,1495]" pageId="47" pageNumber="248" pagination="91 - 223" refId="ref30967" refString="KLUG, J. C. F. 1833. Bericht uber eine auf Madagascar veranstaltete. Sammlung von Insecten aus der Ordnung Coleoptera. Abhandlungen der Koniglichen Preussischen Akademie der Wissenschaften 1832 - 33: 91 - 223 + 5 pls." type="book chapter" year="1833">Klug 1833</bibRefCitation>
)
</taxonomicName>
in this species group, but
<bibRefCitation id="E606EAB03056FFBFFDDBFA52BFA708CC" author="MATEU, J." box="[558,708,1471,1495]" pageId="47" pageNumber="248" pagination="845 - 855" refId="ref31792" refString="MATEU, J. 1973. Notes sur les Lebiens malagaches (Col. Caraboidea) (3 e note). Le genre Eurydera Castelnau. Bulletin du Museum National d'Histoire Naturelle (Ser. 3) No. 139, Zoologie 103: 845 - 855." type="journal article" year="1973">Mateu (1973)</bibRefCitation>
added another species,
<taxonomicName id="4597ECC23056FFBFFC33FA2DBDDC08ED" authority="Mateu" authorityName="Mateu" authorityYear="1973" class="Insecta" family="Carabidae" genus="Eurydera" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="248" phylum="Arthropoda" rank="species" species="fossulata">
<emphasis id="B0E34B533056FFBFFC33FA2DB9F608CC" box="[966,1173,1471,1495]" italics="true" pageId="47" pageNumber="248">Eurydera fossulata</emphasis>
Mateu
</taxonomicName>
to the group, in part based on the shared lack of sexual dimorphism in the front tarsi in members of these two species. Unlike males of most
<taxonomicName id="4597ECC23056FFBFFD75FA6DBE5B0B0C" box="[640,824,1535,1559]" pageId="47" pageNumber="248">
<emphasis id="B0E34B533056FFBFFD75FA6DBF860B0C" box="[640,741,1535,1559]" italics="true" pageId="47" pageNumber="248">Eurydera</emphasis>
species
</taxonomicName>
, those of
<taxonomicName id="4597ECC23056FFBFFC56F992B9790B0C" baseAuthorityName="Klug" baseAuthorityYear="1833" box="[931,1050,1535,1559]" class="Insecta" family="Carabidae" genus="Eurydera" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="248" phylum="Arthropoda" rank="species" species="unicolor">
<emphasis id="B0E34B533056FFBFFC56F992B9790B0C" box="[931,1050,1535,1559]" italics="true" pageId="47" pageNumber="248">E. unicolor</emphasis>
</taxonomicName>
and
<taxonomicName id="4597ECC23056FFBFFBBAF992BDDF0B2C" authorityName="Mateu" authorityYear="1973" class="Insecta" family="Carabidae" genus="Eurydera" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="248" phylum="Arthropoda" rank="species" species="fossulata">
<emphasis id="B0E34B533056FFBFFBBAF992BDDF0B2C" italics="true" pageId="47" pageNumber="248">E. fossulata</emphasis>
</taxonomicName>
have tarsomeres 1 to 3 no wider than in females and without ventral adhesive setae. Members of
<taxonomicName id="4597ECC23056FFBFFF25F9D2BC290B4C" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[208,330,1599,1623]" class="Insecta" family="Carabidae" genus="Eurydera" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="248" phylum="Arthropoda" rank="species" species="simplica">
<emphasis id="B0E34B533056FFBFFF25F9D2BC290B4C" box="[208,330,1599,1623]" italics="true" pageId="47" pageNumber="248">E. simplica</emphasis>
</taxonomicName>
are easily distinguished from those of these two species, as well as from all other described
<taxonomicName id="4597ECC23056FFBFFED3F9CDBCBC0B6C" box="[294,479,1631,1655]" pageId="47" pageNumber="248">
<emphasis id="B0E34B533056FFBFFED3F9CDBCE80B6C" box="[294,395,1631,1655]" italics="true" pageId="47" pageNumber="248">Eurydera</emphasis>
species
</taxonomicName>
, by the uniquely shaped pronotum (
<figureCitation id="1AAC8BC43056FFBFFC90F9CDBEA70B6C" box="[869,964,1631,1655]" captionStart="FIGURE" captionStartId="46.[155,167,1667,1686]" captionTargetBox="[149,1176,526,1636]" captionTargetId="figure-537@46.[120,1176,528,1664]" captionTargetPageId="46" captionText="FIGURE 27. Digital images of holotype male of Eurydera simplica Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B. Labels. Scale line = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13799501" httpUri="https://zenodo.org/record/13799501/files/figure.png" pageId="47" pageNumber="248">Fig. 27A</figureCitation>
), which is relatively small and short for the genus, with the disc smooth, lateral explanation very narrow, lateral margins with very short and shallow sinuation anterior to markedly obtuse hind angles, and basal margin obliquely angulate laterally. Unlike males of
<taxonomicName id="4597ECC23049FFA0FD72FF4EBF9C0DE8" baseAuthorityName="Klug" baseAuthorityYear="1833" box="[647,767,219,243]" class="Insecta" family="Carabidae" genus="Eurydera" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="249" phylum="Arthropoda" rank="species" species="unicolor">
<emphasis id="B0E34B533049FFA0FD72FF4EBF9C0DE8" box="[647,767,219,243]" italics="true" pageId="48" pageNumber="249">E. unicolor</emphasis>
</taxonomicName>
and
<taxonomicName id="4597ECC23049FFA0FCC1FF4EBED10DE8" authorityName="Mateu" authorityYear="1973" box="[820,946,219,243]" class="Insecta" family="Carabidae" genus="Eurydera" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="249" phylum="Arthropoda" rank="species" species="fossulata">
<emphasis id="B0E34B533049FFA0FCC1FF4EBED10DE8" box="[820,946,219,243]" italics="true" pageId="48" pageNumber="249">E. fossulata</emphasis>
</taxonomicName>
, those of
<taxonomicName id="4597ECC23049FFA0FBE8FF4EB9F60DE8" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[1053,1173,219,243]" class="Insecta" family="Carabidae" genus="Eurydera" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="249" phylum="Arthropoda" rank="species" species="simplica">
<emphasis id="B0E34B533049FFA0FBE8FF4EB9F60DE8" box="[1053,1173,219,243]" italics="true" pageId="48" pageNumber="249">E. simplica</emphasis>
</taxonomicName>
have front tarsi with tarsomeres 1 to 3 wider than in females and with at least a few adhesive setae ventrally, which may call into question a close relationship with these two these species. Members of
<taxonomicName id="4597ECC23049FFA0FF6FFEAEBC770C48" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[154,276,315,339]" class="Insecta" family="Carabidae" genus="Eurydera" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="249" phylum="Arthropoda" rank="species" species="simplica">
<emphasis id="B0E34B533049FFA0FF6FFEAEBC770C48" box="[154,276,315,339]" italics="true" pageId="48" pageNumber="249">E. simplica</emphasis>
</taxonomicName>
also differ from those of
<taxonomicName id="4597ECC23049FFA0FDC4FEAEBFC80C48" baseAuthorityName="Klug" baseAuthorityYear="1833" box="[561,683,315,339]" class="Insecta" family="Carabidae" genus="Eurydera" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="249" phylum="Arthropoda" rank="species" species="unicolor">
<emphasis id="B0E34B533049FFA0FDC4FEAEBFC80C48" box="[561,683,315,339]" italics="true" pageId="48" pageNumber="249">E. unicolor</emphasis>
</taxonomicName>
in having the pronotum with anterior angles much less projected anteriorly. They differ from members of
<taxonomicName id="4597ECC23049FFA0FCE2FECEBEF50C6F" authorityName="Mateu" authorityYear="1973" box="[791,918,348,372]" class="Insecta" family="Carabidae" genus="Eurydera" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="249" phylum="Arthropoda" rank="species" species="fossulata">
<emphasis id="B0E34B533049FFA0FCE2FECEBEF50C6F" box="[791,918,348,372]" italics="true" pageId="48" pageNumber="249">E. fossulata</emphasis>
</taxonomicName>
in having the posterior part of the head, between the lateral furrows, smooth and slightly convex, whereas
<taxonomicName id="4597ECC23049FFA0FBE0FEEFB9F60C8F" authorityName="Mateu" authorityYear="1973" box="[1045,1173,380,404]" class="Insecta" family="Carabidae" genus="Eurydera" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="249" phylum="Arthropoda" rank="species" species="fossulata">
<emphasis id="B0E34B533049FFA0FBE0FEEFB9F60C8F" box="[1045,1173,380,404]" italics="true" pageId="48" pageNumber="249">E. fossulata</emphasis>
</taxonomicName>
members have a deep, transverse medial depression, the bottom of which is flat and covered with deep transverse creases, on that part of the head. They also have the elytral intervals without the roughened surfaces typical of members of several species in genus, including
<taxonomicName id="4597ECC23049FFA0FC49FE4FB9590CEF" authorityName="Mateu" authorityYear="1973" box="[956,1082,476,500]" class="Insecta" family="Carabidae" genus="Eurydera" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="249" phylum="Arthropoda" rank="species" species="fossulata">
<emphasis id="B0E34B533049FFA0FC49FE4FB9590CEF" box="[956,1082,476,500]" italics="true" pageId="48" pageNumber="249">E. fossulata</emphasis>
</taxonomicName>
(at least subapically), and the sutural spines are very long and straight (similar to those in
<taxonomicName id="4597ECC23049FFA0FC1BFE6FB9040F0F" baseAuthorityName="Klug" baseAuthorityYear="1833" box="[1006,1127,508,532]" class="Insecta" family="Carabidae" genus="Eurydera" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="249" phylum="Arthropoda" rank="species" species="unicolor">
<emphasis id="B0E34B533049FFA0FC1BFE6FB9040F0F" box="[1006,1127,508,532]" italics="true" pageId="48" pageNumber="249">E. unicolor</emphasis>
</taxonomicName>
and
<taxonomicName id="4597ECC23049FFA0FF8EFD8FBD990F2E" authorityName="Mateu" authorityYear="1973" box="[123,250,541,565]" class="Insecta" family="Carabidae" genus="Eurydera" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="249" phylum="Arthropoda" rank="species" species="fossulata">
<emphasis id="B0E34B533049FFA0FF8EFD8FBD990F2E" box="[123,250,541,565]" italics="true" pageId="48" pageNumber="249">E. fossulata</emphasis>
</taxonomicName>
members). Finally, the median lobe of male genitalia of the
<typeStatus id="5D2C29E33049FFA0FC63FD8FBE900F2E" box="[918,1011,541,565]" pageId="48" pageNumber="249" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4597ECC23049FFA0FBEEFD8FB9F60F2E" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[1051,1173,541,565]" class="Insecta" family="Carabidae" genus="Eurydera" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="249" phylum="Arthropoda" rank="species" species="simplica">
<emphasis id="B0E34B533049FFA0FBEEFD8FB9F60F2E" box="[1051,1173,541,565]" italics="true" pageId="48" pageNumber="249">E. simplica</emphasis>
</taxonomicName>
(
<figureCitation id="1AAC8BC43049FFA0FF71FDAFBC620F4E" box="[132,257,573,597]" captionStart="FIGURE" captionStartId="47.[155,167,783,802]" captionTargetBox="[120,1164,230,765]" captionTargetId="figure-2124@47.[120,1176,216,765]" captionTargetPageId="47" captionText="FIGURE 28. Digital images of holotype male of Eurydera simplica Kavanaugh &amp; Rainio sp. nov. A. Head, dorsal aspect; B–C. Median lobe of aedeagus of genitalia; B. Left lateral aspect; C. Dorsal aspect. Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13799503" httpUri="https://zenodo.org/record/13799503/files/figure.png" pageId="48" pageNumber="249">Fig. 28B–C</figureCitation>
) has the shaft much narrower basally and the apical orifice shorter than in
<emphasis id="B0E34B533049FFA0FBDEFDAFBDF00F6E" italics="true" pageId="48" pageNumber="249">E. unicol- or</emphasis>
males (see Jeannel 1949, Fig. 495). No male of
<taxonomicName id="4597ECC23049FFA0FD6EFDCCBE7A0F6E" authorityName="Mateu" authorityYear="1973" box="[667,793,605,629]" class="Insecta" family="Carabidae" genus="Eurydera" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="249" phylum="Arthropoda" rank="species" species="fossulata">
<emphasis id="B0E34B533049FFA0FD6EFDCCBE7A0F6E" box="[667,793,605,629]" italics="true" pageId="48" pageNumber="249">E. fossulata</emphasis>
</taxonomicName>
has been available for comparison.
</paragraph>
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.— At present, known only from the
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locality.
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.— The two known specimens of this species both were collected by hand in daytime in secondary montane rainforest in the Talatakely area at elevations ranging from
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.
</paragraph>
</subSubSection>
</treatment>
</document>
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