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<document id="158A04B1308FB717AA86122A8D954F15" ID-DOI="10.5281/zenodo.13799439" ID-ISSN="0068-547X" ID-Zenodo-Dep="13799439" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722538212676" checkinUser="felipe" docAuthor="Kavanaugh, David H. &amp; Rainio, Johanna" docDate="2016" docId="0A3E26573042FFADFF8EFA33BF1E0823" docLanguage="en" docName="ProcCalifAcadSci.63.7.201-268.pdf" docOrigin="Proceedings of the California Academy of Sciences 63 (7)" docStyle="DocumentStyle:E411DA009AF5E7F9A7E023431E2B7D5B.2:ProcCalifAcadSci.2015-.journal_article" docStyleId="E411DA009AF5E7F9A7E023431E2B7D5B" docStyleName="ProcCalifAcadSci.2015-.journal_article" docStyleVersion="2" docTitle="Eunostus minimus Kavanaugh and Rainio 2016, sp. nov." docType="treatment" docVersion="2" lastPageNumber="262" masterDocId="F6075E2F3079FF90FFF5FF92BD630D1B" masterDocTitle="Twenty-six New Species of Predaceous Ground Beetles (Coleoptera: Adephaga: Carabidae) from Ranomafana National Park, Madagascar" masterLastPageNumber="268" masterPageNumber="201" pageNumber="260" updateTime="1726771873529" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="7B46DE7A6F8EC050E06DB403DB9E3258" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="416977A94864D2BBDF6C1B44236984C3">
<mods:title id="6F2B3E184AF42378F786195D52FB738E">Twenty-six New Species of Predaceous Ground Beetles (Coleoptera: Adephaga: Carabidae) from Ranomafana National Park, Madagascar</mods:title>
</mods:titleInfo>
<mods:name id="7216BD3CFBE6BA8C0E94F00714987E52" type="personal">
<mods:role id="DA76847867354BA1BE4D26246F1CE860">
<mods:roleTerm id="0977C3BBA795E6941E185F50D30FCC72">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A9F703B0CBBD9BF3585A6D7635245DED">Kavanaugh, David H.</mods:namePart>
<mods:affiliation id="24680503CC01E1B9B6229E0D742D5716">Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA; Email: dkavanaugh @ calacademy. org; &amp; Corresponding author: David H. Kavanaugh (dkavanaugh @ calacademy. org)</mods:affiliation>
</mods:name>
<mods:name id="3D30007B73C8FCDA7F5A16F04DAF5007" type="personal">
<mods:role id="B46A37E33BC1F469E8F4F0D1919BBE34">
<mods:roleTerm id="0D03061712E89B794DAA9FD41AD9A8D7">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="988162A2FDC20F02195424CC4FB6B290">Rainio, Johanna</mods:namePart>
<mods:affiliation id="84051BBD445B9D5821E37417924FF50A">Department of Ecology and Systematics, University of Helsinki, P. O. Box 65, FIN- 00014 Helsinki, Finland; Email: kjohannarainio @ gmail. com</mods:affiliation>
</mods:name>
<mods:typeOfResource id="EA43DFACA0EB6D0FA47B792C16924F84">text</mods:typeOfResource>
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<mods:titleInfo id="F4212561FE11AA331468D9D9CA8BBED6">
<mods:title id="206E343D817526EF179D1D7259C0B767">Proceedings of the California Academy of Sciences</mods:title>
</mods:titleInfo>
<mods:part id="85E1620BA54841EDDFB68C9E916EAF85">
<mods:date id="8A797DAECFB3349E4CD077BE1ECF9C21">2016</mods:date>
<mods:detail id="9C4D3CF5737AB312FBDCB1B0713368A3" type="pubDate">
<mods:number id="01978AC2CFF0D6B8F516D121D90BF94F">2016-04-29</mods:number>
</mods:detail>
<mods:detail id="91124DA60D5FC00FC6E5389D55518130" type="volume">
<mods:number id="5FAC302972FDCF65A0D980D24402BAA1">63</mods:number>
</mods:detail>
<mods:detail id="47DB1A443CC384F30D9C2A99479E85FD" type="issue">
<mods:number id="AE26B9DA48D110E71E17D0AE38FAAE19">7</mods:number>
</mods:detail>
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<mods:start id="A643A80FC7B0DB2C16969983D19963CE">201</mods:start>
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</mods:extent>
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<mods:classification id="0B07D5DCE621959AEF15BF33B691CA34">journal article</mods:classification>
<mods:identifier id="0B376A2095FAEFE4D72DB381007B2CD1" type="DOI">10.5281/zenodo.13799439</mods:identifier>
<mods:identifier id="B1E9BA6F64BAFE250C684DCCC773AFEC" type="ISSN">0068-547X</mods:identifier>
<mods:identifier id="67CCF32929DD272E68C259FB40E8493E" type="Zenodo-Dep">13799439</mods:identifier>
</mods:mods>
<treatment id="0A3E26573042FFADFF8EFA33BF1E0823" ID-DOI="http://doi.org/10.5281/zenodo.13799531" ID-Zenodo-Dep="13799531" LSID="urn:lsid:plazi:treatment:0A3E26573042FFADFF8EFA33BF1E0823" httpUri="http://treatment.plazi.org/id/0A3E26573042FFADFF8EFA33BF1E0823" lastPageId="61" lastPageNumber="262" pageId="59" pageNumber="260">
<subSubSection id="CA8DC4CA3042FFABFF8EFA33BFDC08A2" box="[123,703,1441,1465]" pageId="59" pageNumber="260" type="nomenclature">
<paragraph id="822897413042FFABFF8EFA33BFDC08A2" blockId="59.[123,703,1441,1495]" box="[123,703,1441,1465]" pageId="59" pageNumber="260">
<heading id="D960202D3042FFABFF8EFA33BFDC08A2" box="[123,703,1441,1465]" fontSize="10" level="2" pageId="59" pageNumber="260" reason="4">
<emphasis id="B0E34B533042FFABFF8EFA33BFDC08A2" bold="true" box="[123,703,1441,1465]" pageId="59" pageNumber="260">
<taxonomicName id="4597ECC23042FFABFF8EFA33BF3808A2" ID-CoL="3CG64" authority="Kavanaugh and Rainio" authorityName="Kavanaugh and Rainio" authorityYear="2016" box="[123,603,1441,1465]" class="Insecta" family="Carabidae" genus="Eunostus" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="260" phylum="Arthropoda" rank="species" species="minimus" status="sp. nov.">
<emphasis id="B0E34B533042FFABFF8EFA33BC2908A2" bold="true" box="[123,330,1441,1465]" italics="true" pageId="59" pageNumber="260">Eunostus minimus</emphasis>
Kavanaugh and Rainio
</taxonomicName>
,
<taxonomicNameLabel id="ABD0F6283042FFABFD9CFA33BFDC08A2" box="[617,703,1441,1465]" pageId="59" pageNumber="260" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="CA8DC4CA3042FFABFF8EFA53BE350B1A" pageId="59" pageNumber="260" type="description">
<paragraph id="822897413042FFABFF8EFA53BC6608CD" blockId="59.[123,703,1441,1495]" box="[123,261,1473,1495]" pageId="59" pageNumber="260">
<figureCitation id="1AAC8BC43042FFABFF8EFA53BC6608CD" box="[123,261,1473,1495]" captionStart-0="FIGURE" captionStart-1="FIGURE" captionStartId-0="60.[155,167,1162,1181]" captionStartId-1="61.[155,167,825,844]" captionTargetBox-0="[130,1176,216,1147]" captionTargetBox-1="[120,1176,216,803]" captionTargetId-0="figure-1103@60.[120,1176,216,1159]" captionTargetId-1="figure-1453@61.[120,1176,216,803]" captionTargetPageId-0="60" captionTargetPageId-1="61" captionText-0="FIGURE 36. Digital images of holotype male of Eunostus minimus Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B. Labels. Scale line A = 1.0 mm." captionText-1="FIGURE 37. Digital images of Eunostus minimus Kavanaugh &amp; Rainio sp. nov. A–B. Apical part of left hind femur, oblique medial aspect; A. Holotype male; B. Female; C–D. Aedeagus of holotype male genitalia; C. Left lateral aspect; D. Dorsal aspect. Scale lines = 0.5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.13799521" figureDoi-1="http://doi.org/10.5281/zenodo.13799523" httpUri-0="https://zenodo.org/record/13799521/files/figure.png" httpUri-1="https://zenodo.org/record/13799523/files/figure.png" pageId="59" pageNumber="260">Figures 36–37</figureCitation>
</paragraph>
<paragraph id="822897413042FFABFF8EFA7FBE350B1A" blockId="59.[123,854,1516,1538]" box="[123,854,1516,1538]" pageId="59" pageNumber="260">
<emphasis id="B0E34B533042FFABFF8EFA7FBC430B1A" box="[123,288,1516,1538]" italics="true" pageId="59" pageNumber="260">
<taxonomicName id="4597ECC23042FFABFF8EFA7FBDB00B19" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[123,211,1517,1538]" class="Insecta" family="Carabidae" genus="Eunostus" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="260" phylum="Arthropoda" rank="genus" status="n. sp.">Eunostus</taxonomicName>
<taxonomicNameLabel id="ABD0F6283042FFABFF2CFA7FBC6E0B19" box="[217,269,1517,1538]" pageId="59" pageNumber="260" rank="species">n. sp.</taxonomicNameLabel>
1
</emphasis>
; Rainio 2009: 31, Rainio 2012: 72 (informal designation).
</paragraph>
</subSubSection>
<subSubSection id="CA8DC4CA3042FFADFF5EF9B7BF1E0823" lastPageId="61" lastPageNumber="262" pageId="59" pageNumber="260" type="materials_examined">
<paragraph id="822897413042FFACFF5EF9B7BC1808A3" blockId="59.[123,1174,1573,1726]" lastBlockId="60.[123,1174,1246,1725]" lastPageId="60" lastPageNumber="261" pageId="59" pageNumber="260">
<emphasis id="B0E34B533042FFABFF5EF9B7BC390B20" bold="true" box="[171,346,1573,1597]" pageId="59" pageNumber="260">
<smallCapsWord id="84CE019D3042FFABFF5EF9B7BD870B27" baselines="1591,1591" box="[171,228,1573,1597]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Type" pageId="59" pageNumber="260">TYPE</smallCapsWord>
<smallCapsWord id="84CE019D3042FFABFF18F9BABC390B20" baselines="1590" box="[237,346,1576,1595]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="material" pageId="59" pageNumber="260">MATERIAL</smallCapsWord>
</emphasis>
.—
<materialsCitation id="32FF9D1C3042FFABFE74F9B7BEA70BA6" collectingDate="2003-03-27" collectingDateMax="2003-03-31" collectingDateMin="2003-03-27" collectorName="Fisher" country="Madagascar" county="Parc National de Ranomafana" elevation="1100" latitude="-21.289999" location="California Acad. of Sciences" longLatPrecision="21" longitude="47.433334" municipality="Vatoharanana River" pageId="59" pageNumber="260" specimenCount="2" stateProvince="Fianarantsoa" typeStatus="holotype">
<typeStatus id="5D2C29E33042FFABFE74F9B7BC870B26" box="[385,484,1573,1597]" pageId="59" pageNumber="260" type="holotype">Holotype</typeStatus>
(
<figureCitation id="1AAC8BC43042FFABFE00F9B7BF1C0B27" box="[501,639,1573,1597]" captionStart="FIGURE" captionStartId="60.[155,167,1162,1181]" captionTargetBox="[130,1176,216,1147]" captionTargetId="figure-1103@60.[120,1176,216,1159]" captionTargetPageId="60" captionText="FIGURE 36. Digital images of holotype male of Eunostus minimus Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B. Labels. Scale line A = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13799521" httpUri="https://zenodo.org/record/13799521/files/figure.png" pageId="59" pageNumber="260">Figs. 36A–B</figureCitation>
), a male, in CAS, labeled: “CASENT 8069195”/ “
<collectingCountry id="FA80D7D13042FFABFF73F9D7BC270B46" box="[134,324,1605,1629]" name="Madagascar" pageId="59" pageNumber="260">MADAGASCAR</collectingCountry>
:
<collectingRegion id="405359A33042FFABFEA6F9D7BCB80B46" box="[339,475,1605,1629]" country="Madagascar" name="Fianarantsoa" pageId="59" pageNumber="260">Fianarantsoa</collectingRegion>
<collectingCounty id="6B49EFCD3042FFABFE17F9D4BE400B46" box="[482,803,1605,1629]" pageId="59" pageNumber="260">Parc National de Ranomafana</collectingCounty>
,
<collectingMunicipality id="624C0D3B3042FFABFCC5F9D7B9600B46" box="[816,1027,1605,1629]" pageId="59" pageNumber="260">Vatoharanana River</collectingMunicipality>
,
<locationDeviation id="939A8E633042FFABFBE4F9D7BC500B66" location="Ranomafana" pageId="59" pageNumber="260">
<quantity id="456F3AA43042FFABFBE4F9D7B9380B46" box="[1041,1115,1605,1629]" metricMagnitude="3" metricUnit="m" metricValue="4.1" pageId="59" pageNumber="260" unit="km" value="4.1">4.1 km</quantity>
231° SW Ranomafana
</locationDeviation>
elev
<quantity id="456F3AA43042FFABFE86F9F7BCDF0B66" box="[371,444,1637,1661]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="59" pageNumber="260" unit="m" value="1100.0">
<elevation id="09BA70723042FFABFE86F9F7BCDF0B66" box="[371,444,1637,1661]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="59" pageNumber="260" unit="m" value="1100.0">1100m</elevation>
</quantity>
<date id="F629B1813042FFABFE33F9F7BF1A0B66" box="[454,633,1637,1661]" pageId="59" pageNumber="260" value="2003-03-27" valueMax="2003-03-31" valueMin="2003-03-27">
<collectingDate id="E66D48693042FFABFE33F9F7BF1A0B66" box="[454,633,1637,1661]" pageId="59" pageNumber="260" value="2003-03-27" valueMax="2003-03-31" valueMin="2003-03-27">27-31 Mar 2003</collectingDate>
</date>
”/ “
<geoCoordinate id="E7A3F1863042FFABFD56F9F7BE420B66" box="[675,801,1637,1661]" degrees="21" direction="south" minutes="17" orientation="latitude" pageId="59" pageNumber="260" precision="15" seconds="24" value="-21.289999">21°17’24”S</geoCoordinate>
<geoCoordinate id="E7A3F1863042FFABFCDEF9F7BEDB0B66" box="[811,952,1637,1661]" degrees="047" direction="east" minutes="26" orientation="longitude" pageId="59" pageNumber="260" precision="15" seconds="00" value="47.433334">047°26’00”E</geoCoordinate>
<location id="8748C19A3042FFABFC37F9F7BDBB0B86" LSID="urn:lsid:plazi:treatment:0A3E26573042FFADFF8EFA33BF1E0823:8748C19A3042FFABFC37F9F7BDBB0B86" country="Madagascar" county="Parc National de Ranomafana" latitude="-21.289999" longLatPrecision="21" longitude="47.433334" municipality="Vatoharanana River" name="California Acad. of Sciences" pageId="59" pageNumber="260" stateProvince="Fianarantsoa">California Acad. of Sciences</location>
coll.
<collectorName id="2F62F2973042FFABFEE2F917BC3A0B86" box="[279,345,1669,1693]" pageId="59" pageNumber="260">Fisher</collectorName>
, Griswold et al. sifted litter, montane rainforest collection code: BLF8400”/ “
<typeStatus id="5D2C29E33042FFABFF72F934BC770BA5" box="[135,276,1702,1726]" pageId="59" pageNumber="260" type="holotype">HOLOTYPE</typeStatus>
<taxonomicName id="4597ECC23042FFABFEEFF934BFA40BA6" authority="Kavanaugh &amp; Rainio" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[282,711,1701,1726]" class="Insecta" family="Carabidae" genus="Eunostus" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="260" phylum="Arthropoda" rank="species" species="minimus" status="sp. nov.">
<emphasis id="B0E34B533042FFABFEEFF934BCBE0BA6" box="[282,477,1702,1725]" italics="true" pageId="59" pageNumber="260">Eunostus minimus</emphasis>
Kavanaugh &amp; Rainio
</taxonomicName>
<taxonomicNameLabel id="ABD0F6283042FFABFD3BF934BE640BA6" box="[718,775,1702,1725]" pageId="59" pageNumber="260" rank="species">sp. n.</taxonomicNameLabel>
2015” [red label]
</materialsCitation>
.
<materialsCitation id="32FF9D1C3042FFACFC27F934BC1708A3" collectingDate="2001-12-21" collectingDateMax="2004-08-24" collectingDateMin="2001-12-21" collectionCode="MNHN, CAS" collectorName="R. Harin'Hala California Acad &amp; F. Ratalata &amp; J. Rainio" country="Madagascar" county="Ranomafana" elevation="690" lastPageId="60" lastPageNumber="261" latitude="-21.2485" location="Ranomafana" longLatPrecision="12" longitude="47.452168" municipality="Harin'Hala California Acad of Sciences" pageId="59" pageNumber="260" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Fianarantsoa" typeStatus="paratype">
<typeStatus id="5D2C29E33042FFABFC27F934B9590BA5" box="[978,1082,1702,1726]" pageId="59" pageNumber="260" type="paratype">Paratypes</typeStatus>
(total of 2):
<specimenCount id="94915CC83045FFACFF5FFB4CBD9D09ED" box="[170,254,1246,1270]" count="1" pageId="60" pageNumber="261" type="male">1 male</specimenCount>
(in
<collectionCode id="E4860F843045FFACFECAFB4CBCF109EE" box="[319,402,1246,1269]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="60" pageNumber="261">MNHN</collectionCode>
) labeled “CASENT 8069129”/ “
<collectingCountry id="FA80D7D13045FFACFCD3FB4CBE8B09ED" box="[806,1000,1246,1270]" name="Madagascar" pageId="60" pageNumber="261">MADAGASCAR</collectingCountry>
:
<collectingRegion id="405359A33045FFACFBF4FB4CB9ED09ED" box="[1025,1166,1246,1270]" country="Madagascar" name="Fianarantsoa" pageId="60" pageNumber="261">Fianarantsoa</collectingRegion>
,
<collectingCounty id="6B49EFCD3045FFACFF8EFB6CBC61080D" box="[123,258,1278,1302]" pageId="60" pageNumber="261">Ranomafana</collectingCounty>
JIRAMA water works,
<geoCoordinate id="E7A3F1863045FFACFDF1FB6CBF1E080D" box="[516,637,1278,1302]" degrees="21" direction="south" minutes="14. 91" orientation="latitude" pageId="60" pageNumber="261" precision="9" value="-21.2485">21°14.91’S</geoCoordinate>
,
<geoCoordinate id="E7A3F1863045FFACFD7EFB6CBE70080D" box="[651,787,1278,1302]" degrees="047" direction="east" minutes="27. 13" orientation="longitude" pageId="60" pageNumber="261" precision="9" value="47.452168">047°27.13’E</geoCoordinate>
”/ “
<date id="F629B1813045FFACFCCCFB6CB94A080D" box="[825,1065,1278,1302]" pageId="60" pageNumber="261" value="2001-12-21" valueMax="2001-12-24" valueMin="2001-12-21">
<collectingDate id="E66D48693045FFACFCCCFB6CB94A080D" box="[825,1065,1278,1302]" pageId="60" pageNumber="261" value="2001-12-21" valueMax="2001-12-24" valueMin="2001-12-21">21-24 December 2001</collectingDate>
</date>
collector:
<collectorName id="2F62F2973045FFACFF8EFA8DBCA5082C" box="[123,454,1311,1335]" pageId="60" pageNumber="261">
R.
<collectingMunicipality id="624C0D3B3045FFACFF6EFA8DBCA5082C" box="[155,454,1311,1335]" pageId="60" pageNumber="261">Harin’Hala California Acad</collectingMunicipality>
</collectorName>
of Sciences malaise trap near river elev.
<quantity id="456F3AA43045FFACFC73FA8DBEA8082D" box="[902,971,1311,1335]" metricMagnitude="2" metricUnit="m" metricValue="6.9" pageId="60" pageNumber="261" unit="m" value="690.0">
<elevation id="09BA70723045FFACFC73FA8DBEA8082D" box="[902,971,1311,1335]" metricMagnitude="2" metricUnit="m" metricValue="6.9" pageId="60" pageNumber="261" unit="m" value="690.0">690 m</elevation>
</quantity>
, MA-02-09D-08”;
<specimenCount id="94915CC83045FFACFF8EFAADBDBA084C" box="[123,217,1343,1367]" count="1" pageId="60" pageNumber="261" type="female">1 female</specimenCount>
(in
<collectionCode id="E4860F843045FFACFEF0FAADBC5A084C" box="[261,313,1343,1367]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="60" pageNumber="261" type="Museum">CAS</collectionCode>
) labeled “CASENT 1051146”/ “
<collectingCountry id="FA80D7D13045FFACFD69FAADBE39084C" box="[668,858,1343,1367]" name="Madagascar" pageId="60" pageNumber="261">MADAGASCAR</collectingCountry>
<location id="8748C19A3045FFACFC97FAADBE8A084C" LSID="urn:lsid:plazi:treatment:0A3E26573042FFADFF8EFA33BF1E0823:8748C19A3045FFACFC97FAADBE8A084C" box="[866,1001,1343,1367]" country="Madagascar" county="Ranomafana" latitude="-21.2485" longLatPrecision="12" longitude="47.452168" municipality="Harin'Hala California Acad of Sciences" name="Ranomafana" pageId="60" pageNumber="261" stateProvince="Fianarantsoa">Ranomafana</location>
NP, Vatoharana trail S 600
<date id="F629B1813045FFACFF0DFAF2BC010863" box="[248,354,1376,1400]" pageId="60" pageNumber="261" value="2004-08-24">
<collectingDate id="E66D48693045FFACFF0DFAF2BC010863" box="[248,354,1376,1400]" pageId="60" pageNumber="261" value="2004-08-24">24.8.2004</collectingDate>
</date>
”/ “collected by hand in daytime,
<collectorName id="2F62F2973045FFACFD2CFAF2BE2F0863" box="[729,844,1376,1400]" pageId="60" pageNumber="261">F. Ratalata</collectorName>
&amp;
<collectorName id="2F62F2973045FFACFC81FAF2BEB60863" box="[884,981,1376,1400]" pageId="60" pageNumber="261">J. Rainio</collectorName>
collectors”. Both
<typeStatus id="5D2C29E33045FFACFF8EFA13BD820882" box="[123,225,1409,1433]" pageId="60" pageNumber="261" type="paratype">paratypes</typeStatus>
also bear the following label: “
<typeStatus id="5D2C29E33045FFACFDC0FA13BFDA0882" box="[565,697,1409,1433]" pageId="60" pageNumber="261" type="paratype">PARATYPE</typeStatus>
<taxonomicName id="4597ECC23045FFACFD34FA13B9130883" authority="Kavanaugh &amp; Rainio" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[705,1136,1408,1433]" class="Insecta" family="Carabidae" genus="Eunostus" kingdom="Animalia" order="Coleoptera" pageId="60" pageNumber="261" phylum="Arthropoda" rank="species" species="minimus" status="sp. nov.">
<emphasis id="B0E34B533045FFACFD34FA13BEE70883" box="[705,900,1409,1432]" italics="true" pageId="60" pageNumber="261">Eunostus minimus</emphasis>
Kavanaugh &amp; Rainio
</taxonomicName>
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2015” [yellow label]
</materialsCitation>
.
</paragraph>
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36. Digital images of holotype male of
<taxonomicName id="4597ECC23045FFACFDB4FB18BEC10986" authority="Kavanaugh &amp; Rainio" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[577,930,1162,1181]" class="Insecta" family="Carabidae" genus="Eunostus" kingdom="Animalia" order="Coleoptera" pageId="60" pageNumber="261" phylum="Arthropoda" rank="species" species="minimus" status="sp. nov.">
<emphasis id="B0E34B533045FFACFDB4FB18BFBC0987" box="[577,735,1162,1180]" italics="true" pageId="60" pageNumber="261">Eunostus minimus</emphasis>
Kavanaugh &amp; Rainio
</taxonomicName>
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A. Habitus, dorsal aspect; B. Labels. Scale line A = 1.0 mm.
</paragraph>
</caption>
<paragraph id="822897413045FFACFF5EFA50B95008C2" blockId="60.[123,1174,1246,1725]" box="[171,1075,1473,1498]" pageId="60" pageNumber="261">
<materialsCitation id="32FF9D1C3045FFACFF5EFA50B94F08C2" box="[171,1068,1473,1498]" country="Madagascar" location="Ranomafana National Park" pageId="60" pageNumber="261" specimenCount="1" stateProvince="Fianarantsoa" typeStatus="holotype">
<emphasis id="B0E34B533045FFACFF5EFA50BC3408C3" bold="true" box="[171,343,1474,1498]" pageId="60" pageNumber="261">
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<typeStatus id="5D2C29E33045FFACFF5EFA50BD8708C3" box="[171,228,1474,1498]" pageId="60" pageNumber="261">TYPE</typeStatus>
</smallCapsWord>
<smallCapsWord id="84CE019D3045FFACFF19FA57BC3408C3" baselines="1491" box="[236,343,1477,1496]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="locality" pageId="60" pageNumber="261">LOCALITY</smallCapsWord>
</emphasis>
.—
<collectingCountry id="FA80D7D13045FFACFE8BFA53BC9C08C2" box="[382,511,1473,1497]" name="Madagascar" pageId="60" pageNumber="261">Madagascar</collectingCountry>
,
<collectingRegion id="405359A33045FFACFDF8FA53BF9F08C2" box="[525,764,1473,1497]" country="Madagascar" name="Fianarantsoa" pageId="60" pageNumber="261">Fianarantsoa Province</collectingRegion>
,
<location id="8748C19A3045FFACFCFFFA53B94F08C2" LSID="urn:lsid:plazi:treatment:0A3E26573042FFADFF8EFA33BF1E0823:8748C19A3045FFACFCFFFA53B94F08C2" box="[778,1068,1473,1497]" country="Madagascar" name="Ranomafana National Park" pageId="60" pageNumber="261" stateProvince="Fianarantsoa">Ranomafana National Park</location>
</materialsCitation>
.
</paragraph>
<paragraph id="822897413045FFACFF5EFA70BFC20B20" blockId="60.[123,1174,1246,1725]" pageId="60" pageNumber="261">
<emphasis id="B0E34B533045FFACFF5EFA70BC8F08E2" bold="true" box="[171,492,1506,1530]" pageId="60" pageNumber="261">
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<smallCapsWord id="84CE019D3045FFACFEC1FA74BC3308E2" baselines="1524" box="[308,336,1510,1529]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="of" pageId="60" pageNumber="261">OF</smallCapsWord>
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<smallCapsWord id="84CE019D3045FFACFE5AFA74BC8F08E2" baselines="1524" box="[431,492,1510,1529]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="name" pageId="60" pageNumber="261">NAME</smallCapsWord>
</emphasis>
.— The species epithet,
<taxonomicName id="4597ECC23045FFACFD18FA71BE2408E1" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[749,839,1507,1530]" class="Insecta" family="Carabidae" genus="Eunostus" kingdom="Animalia" order="Coleoptera" pageId="60" pageNumber="261" phylum="Arthropoda" rank="species" species="minimus">
<emphasis id="B0E34B533045FFACFD18FA71BE2408E1" box="[749,839,1507,1530]" italics="true" pageId="60" pageNumber="261">minimus</emphasis>
</taxonomicName>
, is the Latin adjective meaning smallest, a reference to fact that the known specimens of this species are smaller than those of any of the other known species that we have examined.
</paragraph>
<paragraph id="822897413045FFADFF5EF9D6B9F50963" blockId="60.[123,1174,1246,1725]" lastBlockId="61.[123,1174,928,1336]" lastPageId="61" lastPageNumber="262" pageId="60" pageNumber="261">
<smallCapsWord id="84CE019D3045FFACFF5EF9D6BC210B41" baselines="1622,1621" box="[171,322,1604,1628]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Recognition" pageId="60" pageNumber="261">
<emphasis id="B0E34B533045FFACFF5EF9D6BC210B41" bold="true" box="[171,322,1604,1628]" pageId="60" pageNumber="261">RECOGNITION</emphasis>
</smallCapsWord>
.— Small for genus, SBL of male =
<quantity id="456F3AA43045FFACFD38F9D6BE430B40" box="[717,800,1604,1628]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="60" pageNumber="261" unit="mm" value="5.5">5.5 mm</quantity>
, of female 5.0
<emphasis id="B0E34B533045FFACFC34F9D6BEAD0B47" bold="true" box="[961,974,1604,1628]" pageId="60" pageNumber="261">–</emphasis>
5.
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. Members of this species (
<figureCitation id="1AAC8BC43045FFACFED2F9F6BCEB0B66" box="[295,392,1636,1661]" captionStart="FIGURE" captionStartId="60.[155,167,1162,1181]" captionTargetBox="[130,1176,216,1147]" captionTargetId="figure-1103@60.[120,1176,216,1159]" captionTargetPageId="60" captionText="FIGURE 36. Digital images of holotype male of Eunostus minimus Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B. Labels. Scale line A = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13799521" httpUri="https://zenodo.org/record/13799521/files/figure.png" pageId="60" pageNumber="261">Fig. 36A</figureCitation>
) do not key out in Jeannel’s (1949) key for
<taxonomicName id="4597ECC23045FFACFC9AF9F7B94D0B67" box="[879,1070,1636,1660]" class="Insecta" family="Carabidae" genus="Eunostus" kingdom="Animalia" order="Coleoptera" pageId="60" pageNumber="261" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B0E34B533045FFACFC9AF9F7BEB20B67" box="[879,977,1637,1660]" italics="true" pageId="60" pageNumber="261">Eunostus</emphasis>
species.
</taxonomicName>
They are much smaller and shinier than members of any other described species. Their heads are slightly narrower, with the tempora more evenly rounded to the back of the head, their antennal scapes shorter and thicker than in other species, thickest slightly distal to mid-length. Lateral margins of the pronotum are smoothly rounded in anterior half with a moderately short and deep subbasal sinuation anterior to slightly obtuse to rectangular hind angles. The elytral setae are generally sparser than in other species and arranged in 2 more or less distinct longitudinal rows on each interval. Male (
<figureCitation id="1AAC8BC43044FFADFF31FBB2BC450922" box="[196,294,1056,1081]" captionStart="FIGURE" captionStartId="61.[155,167,825,844]" captionTargetBox="[120,1176,216,803]" captionTargetId="figure-1453@61.[120,1176,216,803]" captionTargetPageId="61" captionText="FIGURE 37. Digital images of Eunostus minimus Kavanaugh &amp; Rainio sp. nov. A–B. Apical part of left hind femur, oblique medial aspect; A. Holotype male; B. Female; C–D. Aedeagus of holotype male genitalia; C. Left lateral aspect; D. Dorsal aspect. Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13799523" httpUri="https://zenodo.org/record/13799523/files/figure.png" pageId="61" pageNumber="262">Fig. 37A</figureCitation>
) and female (
<figureCitation id="1AAC8BC43044FFADFE33FBB2BF450923" box="[454,550,1056,1080]" captionStart="FIGURE" captionStartId="61.[155,167,825,844]" captionTargetBox="[120,1176,216,803]" captionTargetId="figure-1453@61.[120,1176,216,803]" captionTargetPageId="61" captionText="FIGURE 37. Digital images of Eunostus minimus Kavanaugh &amp; Rainio sp. nov. A–B. Apical part of left hind femur, oblique medial aspect; A. Holotype male; B. Female; C–D. Aedeagus of holotype male genitalia; C. Left lateral aspect; D. Dorsal aspect. Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13799523" httpUri="https://zenodo.org/record/13799523/files/figure.png" pageId="61" pageNumber="262">Fig. 37B</figureCitation>
) hind femora are sexual dimorphic, each unique among Malagasy species for each sex. Median lobe of male genitalia
<figureCitation id="1AAC8BC43044FFADFCEAFBD2BECB0943" box="[799,936,1088,1112]" captionStart="FIGURE" captionStartId="61.[155,167,825,844]" captionTargetBox="[120,1176,216,803]" captionTargetId="figure-1453@61.[120,1176,216,803]" captionTargetPageId="61" captionText="FIGURE 37. Digital images of Eunostus minimus Kavanaugh &amp; Rainio sp. nov. A–B. Apical part of left hind femur, oblique medial aspect; A. Holotype male; B. Female; C–D. Aedeagus of holotype male genitalia; C. Left lateral aspect; D. Dorsal aspect. Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13799523" httpUri="https://zenodo.org/record/13799523/files/figure.png" pageId="61" pageNumber="262">Figs. 37C–D</figureCitation>
markedly different in shape from those of the other species for which males are known (see Jeannel 1949, Fig. 513d–e).
</paragraph>
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37. Digital images of
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<emphasis id="B0E34B533044FFADFE56FCABBF220E50" box="[419,577,825,843]" italics="true" pageId="61" pageNumber="262">Eunostus minimus</emphasis>
Kavanaugh &amp; Rainio
</taxonomicName>
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A–B. Apical part of left hind femur, oblique medial aspect; A. Holotype male; B. Female; C–D. Aedeagus of holotype male genitalia; C. Left lateral aspect; D. Dorsal aspect. Scale lines = 0.5 mm.
</paragraph>
</caption>
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<emphasis id="B0E34B533044FFADFF5EFB12BC92098C" bold="true" box="[171,497,1152,1176]" pageId="61" pageNumber="262">
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</emphasis>
.— At present, known only from the
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locality.
</paragraph>
<paragraph id="822897413044FFADFF5EFB32BF1E0823" blockId="61.[123,1174,928,1336]" pageId="61" pageNumber="262">
<emphasis id="B0E34B533044FFADFF5EFB32BCC609AC" bold="true" box="[171,421,1184,1208]" pageId="61" pageNumber="262">
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</emphasis>
.— Specimens of the
<typeStatus id="5D2C29E33044FFADFD7AFB33BFDE09A2" box="[655,701,1185,1209]" pageId="61" pageNumber="262">type</typeStatus>
series were collected in both primary montane rainforest in the Vatoharanana area (at elevations ranging from of
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) and in secondary montane rainforest
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north of Ranomafana village at an elevation of
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<quantity id="456F3AA43044FFADFC2FFB72B97C09E3" box="[986,1055,1248,1272]" metricMagnitude="2" metricUnit="m" metricValue="6.9" pageId="61" pageNumber="262" unit="m" value="690.0">690 m</quantity>
.
</specimenCount>
The specimens were collected in a malaise trap and sifted from forest litter, respectively, at the primary forest site and by hand at the secondary forest site.
</paragraph>
</subSubSection>
</treatment>
</document>
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