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<document id="CF08F22FB4C17A296D79E02DA5C7C113" ID-DOI="10.3897/zookeys.1216.132561" ID-ZooBank="0E173F72-36D3-4B97-BC43-F096067A3D1E" ID-publisher-id="132561" URI-arpha="D43572E8-1035-521E-817F-77B73A520DC2" URI-zoobank="http://zoobank.org/0E173F72-36D3-4B97-BC43-F096067A3D1E" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.treatments_approvedBy="admin" article-type="research-article" checkinTime="1729868652905" checkinUser="pensoft" docAuthor="Wang, Bing, Li, Jinglin, Li, Shuqiang &amp; Yao, Zhiyuan" docDate="2024" docId="0A3BBAAB9E695516BEF769327FD252EF" docLanguage="en" docName="ZooKeys 1216: 265-302" docOrigin="ZooKeys 1216" docSource="https://zookeys.pensoft.net/article/132561/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Belisana yongcong Wang, S. Li &amp; Yao 2024, sp. nov." docType="treatment" docUuid="1E1B4058-113E-48F6-9734-39D486FF0FB9" docUuidSource="ZooBank" docVersion="3" dtd-version="3.0" lastPageNumber="302" masterDocId="D43572E81035521E817F77B73A520DC2" masterDocTitle="Nine new spider species of Belisana Thorell, 1898 (Araneae, Pholcidae) from karst caves, with a list of species of the genus from Guizhou, southwestern China" masterLastPageNumber="302" masterPageNumber="265" pageNumber="265" updateTime="1729876391189" updateUser="admin">
<mods:mods id="C2D1479EE1DA19DADE5DF4AB635D97A9" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="9498DE60D5FA5EE7B3888F4E315EBCDB">
<mods:title id="B405112DABF1D0BBB373C081107C8345">Nine new spider species of Belisana Thorell, 1898 (Araneae, Pholcidae) from karst caves, with a list of species of the genus from Guizhou, southwestern China</mods:title>
</mods:titleInfo>
<mods:name id="7C5636FB559B87878E9CE8335CF8D4EA" type="personal">
<mods:role id="FE7DE60597C9EB78A10CB73626F2B879">
<mods:roleTerm id="0B990DE7ED18D6A70DFF320508CD6B46">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="632CF900BA420BBAA5B76534D3B8EF07">Wang, Bing</mods:namePart>
<mods:nameIdentifier id="2BCB677353947F6C3CD8820B93B1FA9D" type="ORCID">https://orcid.org/0009-0009-7047-4395</mods:nameIdentifier>
<mods:affiliation id="48172067D08CC769FBB6A80B7DA71592">College of Life Science, Shenyang Normal University, Shenyang 110034, Liaoning, China</mods:affiliation>
</mods:name>
<mods:name id="AD4688738FCE1D36B387B667538D4602" type="personal">
<mods:role id="9B8D06529AFCC80C765A1EA4AB76211C">
<mods:roleTerm id="7C25413ACF156CE58640D7F650233E75">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="33D28E85CB6BB8252F16410EA28506D3">Li, Jinglin</mods:namePart>
<mods:nameIdentifier id="EC0B36822E57109345D1F4577D289730" type="ORCID">https://orcid.org/0009-0008-3932-309X</mods:nameIdentifier>
<mods:affiliation id="B040929D94318795D74050F99B86F91C">College of Life Science, Shenyang Normal University, Shenyang 110034, Liaoning, China</mods:affiliation>
</mods:name>
<mods:name id="BFD86930224306DF07A2C10CF0B641B5" type="personal">
<mods:role id="20D955AD24AFD006438C93CDA9F640E5">
<mods:roleTerm id="D25C7DFE6B82BEB3A9EB98CED30A4A9E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F94C5039A072625406378768B3684C37">Li, Shuqiang</mods:namePart>
<mods:nameIdentifier id="84F4860FC3FCF095C913A3F604A2D034" type="ORCID">0000-0002-3290-5416</mods:nameIdentifier>
<mods:affiliation id="147535995DE145FD2CE9028313A5CB10">Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China</mods:affiliation>
</mods:name>
<mods:name id="5921495EB238A6751B503D16B79E0BCF" type="personal">
<mods:role id="CD4953BD798CB396774471B3CAA5F380">
<mods:roleTerm id="5A1154152FCD1B052773E2B17989A6EB">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="723CAE550B00D3409B35A21951544608">Yao, Zhiyuan</mods:namePart>
<mods:nameIdentifier id="3B1D47CB0FEB59938BED47146A64D513" type="ORCID">0000-0002-1631-0949</mods:nameIdentifier>
<mods:affiliation id="98137661883E48276BE8CAAA722F6B43">College of Life Science, Shenyang Normal University, Shenyang 110034, Liaoning, China</mods:affiliation>
</mods:name>
<mods:typeOfResource id="503831A052EFC7634AFF4F579E12EB85">text</mods:typeOfResource>
<mods:relatedItem id="1135BF1F24BBD1FFF87EC528C2B803C6" type="host">
<mods:titleInfo id="C559E641D417FF95D12D1282E50E6F99">
<mods:title id="8E6A916138F88D87F76EA7B09AB78356">ZooKeys</mods:title>
</mods:titleInfo>
<mods:part id="B9CFA4D0FE3FAA87AAD429D67980D4F6">
<mods:date id="2E09BE47FFAF9D14139F2FFCA6547052">2024</mods:date>
<mods:detail id="249281AF74BA39C11E03E3384D9A647E" type="pubDate">
<mods:number id="7EA5125D35914218F033B13B7006A06B">2024-10-25</mods:number>
</mods:detail>
<mods:detail id="A6825A2DDD39B1611D7D26BEF850050A" type="volume">
<mods:number id="BA26CA17689CB0B5F80875957F504B9B">1216</mods:number>
</mods:detail>
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<mods:start id="3E5768BC040126514DEFC455E95A0E63">265</mods:start>
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<mods:classification id="87ED52AD65A20C29728EFDAB70553448">journal article</mods:classification>
<mods:identifier id="AED48EC721BF4F1B652458E71D5CFEF5" type="DOI">10.3897/zookeys.1216.132561</mods:identifier>
<mods:identifier id="34FACC108A1F2EF375037D218EBDE39D" type="ZooBank">0E173F72-36D3-4B97-BC43-F096067A3D1E</mods:identifier>
</mods:mods>
<treatment id="0A3BBAAB9E695516BEF769327FD252EF" ID-DOI="http://doi.org/10.5281/zenodo.13992768" ID-Zenodo-Dep="13992768" ID-arpha="0A3BBAAB-9E69-5516-BEF7-69327FD252EF" ID-zoobank="https://zoobank.org/1E1B4058-113E-48F6-9734-39D486FF0FB9" LSID="urn:lsid:zoobank.org:act:1E1B4058-113E-48F6-9734-39D486FF0FB9" LSID-ZBK="urn:lsid:zoobank.org:act:1E1B4058-113E-48F6-9734-39D486FF0FB9" httpUri="http://treatment.plazi.org/id/0A3BBAAB9E695516BEF769327FD252EF">
<subSubSection id="EC44C1510DC78300FE096E619BD98307" type="nomenclature">
<paragraph id="10B3CEB996F609267F2C9B15D9C093AD">
<taxonomicName id="AD35EF2090B505422092699F332A251F" authority="Wang, S. Li &amp; Yao" authorityName="Wang, S. Li &amp; Yao" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="yongcong" status="sp. nov.">
<emphasis id="19BB750051F2787B8241C1D54B72FE82" italics="true">Belisana yongcong</emphasis>
Wang, S. Li &amp; Yao
</taxonomicName>
<taxonomicNameLabel id="2F77201A2CD2C9DAF7D623D4205E50CF" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="8C82330A88A974FB1DBFBEB87A15CB52" type="description">
<paragraph id="C6B0B038F7BDF96EDC4CB18DEC7BFF4F">
<figureCitation id="FB66DB2DE27896DDDB90F987FB144474" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Belisana yongcong sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at ventro-subdistal membranous lamella, arrow 2 points at prolatero-distal membranous lamella, arrow 3 points at sclerotized dorso-subdistal apophysis D retrolateral view). Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.10 (A, B); 0.02 (C, D)." figureDoi="10.3897/zookeys.1216.132561.figure14" httpUri="https://binary.pensoft.net/fig/1161059">Figs 14</figureCitation>
,
<figureCitation id="85EFE8915E35B05F86B683888F89205E" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Belisana yongcong sp. nov., holotype male (C – F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E – H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (A – D); 0.40 (E – H)." figureDoi="10.3897/zookeys.1216.132561.figure15" httpUri="https://binary.pensoft.net/fig/1161060">15</figureCitation>
,
<figureCitation id="FF798F433EB20134093C44E729BD6529" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Procursus in prolateral and retrolateral views (The arrows point at the same structures as those shown in the photos of each species) A, B Belisana yongcong sp. nov. C, D B. zhangi Tong &amp; Li, 2007 E, F B. zhouxi sp. nov. G, H B. zunyi sp. nov. Abbreviation: f = flap. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure23" httpUri="https://binary.pensoft.net/fig/1161068">23 A, B</figureCitation>
,
<figureCitation id="7AB73F6F7FE1CB6BEEDBA3C9483B343E" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Female genitalia in ventral and dorsal views A, B Belisana xiuwen sp. nov. C, D B. yongcong sp. nov. E, F B. zhangi Tong &amp; Li, 2007 G, H B. zhouxi sp. nov. I, J B. zunyi sp. nov. Abbreviations: aa = anterior arch, ep = epigynal pocket, pp = pore plate. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure25" httpUri="https://binary.pensoft.net/fig/1161070">25 C, D</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EIUAI" type="type material">
<paragraph id="2852AC0BDA970E99EBD3A48C1D72AF45">
<heading id="E7FF8DAEFC7A298BBD08C720D330B10D" reason="title">Type material.</heading>
</paragraph>
<paragraph id="4758EE97AB6B21E475D506E73B440741">
<materialsCitation id="F45C9D06D45587356B4B54B4EBEAE47D" collectingDate="2007-05-23" collectionCode="IZCAS" collectorName="J. Liu &amp; Y. Lin" country="China" county="Qiandongnan" elevation="544" latitude="25.9838" location="Guantuan Cave" longLatPrecision="1" longitude="109.11632" municipality="Yongcong Town" specimenCode="Ar 45218" specimenCount="1" specimenCount-male="1" stateProvince="Guizhou" typeStatus="holotype">
<emphasis id="DA1AE2D9F5C47576E92E16794F4E3E4F" bold="true">
<emphasis id="789EE1AA866047525F61295B42E07356" bold="true" italics="true">
<typeStatus id="4A32BB7F200C0ADCD7FCDAE057C2FEFE" type="holotype">Holotype</typeStatus>
</emphasis>
:
<collectingCountry id="C5BAD92999ED82D53554177305C46306" name="China">China</collectingCountry>
</emphasis>
•
<specimenCount id="02A841C688F6884B75BDA97C0BA43399" count="1" type="male">♂</specimenCount>
;
<collectingRegion id="E7A415FBD59AE649FE901C537883C5CA" country="China" name="Guizhou">Guizhou</collectingRegion>
,
<collectingCounty id="B22F319B42549230D8C7D2B998EB3813">Qiandongnan</collectingCounty>
,
<collectingCounty id="7A242C435108A7C03CCC78A37D5BE0AA">Liping County</collectingCounty>
,
<collectingMunicipality id="BC1D62063D80156A31E8D29B2B7ACF9A">Yongcong Town</collectingMunicipality>
,
<collectingMunicipality id="0C93078F5DF306CFC89DF357A929BB81">Guantuan Village</collectingMunicipality>
,
<location id="BF8C054371C50E66C6369D2FCD072114" LSID="urn:lsid:plazi:treatment:0A3BBAAB9E695516BEF769327FD252EF:BF8C054371C50E66C6369D2FCD072114" country="China" county="Qiandongnan" latitude="25.9838" longLatPrecision="1" longitude="109.11632" municipality="Yongcong Town" name="Guantuan Cave" stateProvince="Guizhou">Guantuan Cave</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EWUAI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[109.116317,25.983800]}">
<geoCoordinate id="B86A2E2008464241EB7DA67097B7BEE7" degrees="25" direction="north" minutes="59.028" orientation="latitude" precision="1" value="25.9838">25 ° 59.028 ' N</geoCoordinate>
,
<geoCoordinate id="48A2C2B0417111DA6187A45B3EEA0F73" degrees="109" direction="east" minutes="06.979" orientation="longitude" precision="1" value="109.11632">109 ° 06.979 ' E</geoCoordinate>
</named-content>
; alt.
<quantity id="C49EF18FBA2208B71701CF7BF32AF52A" metricMagnitude="2" metricUnit="m" metricValue="5.4399999999999995" unit="m" value="544.0">
<elevation id="C21C47D780BC360D6B4642E96C21B9FB" metricMagnitude="2" metricUnit="m" metricValue="5.4399999999999995" unit="m" value="544.0">544 m</elevation>
</quantity>
;
<date id="306F01121AF2CB9037808461EAE42D36" value="2007-05-23">
<collectingDate id="5FF1B5AF758B52B6D171B30232CC9162" value="2007-05-23">23 May 2007</collectingDate>
</date>
;
<collectorName id="DEC8AA90D68552DB74EEF9CA54B8D379">J. Liu</collectorName>
&amp;
<collectorName id="F2E25EFFD31DC0F3E294C1B502CEAB20">Y. Lin</collectorName>
leg.;
<named-content id="CB00DD673C18A0F47B468E078B1C9723" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/institute-zoology-chinese-academy-sciences" xlink_title="Institute of Zoology, Chinese Academy of Sciences">
<collectionCode id="2AD9B85A9AB47E195BB37E248731BD27" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" type="">IZCAS</collectionCode>
</named-content>
-
<specimenCode id="636F73C7BF3117CB783F39081A02D7DB">Ar 45218</specimenCode>
</materialsCitation>
.
<materialsCitation id="2B6091AC6E11E02201EFA3F4193B8F12" collectingDate="2007-05-23" collectionCode="IZCAS" collectorName="J. Liu &amp; Y. Lin" country="China" county="Liping County" elevation="544" latitude="25.9838" location="China" longLatPrecision="1" longitude="109.11632" municipality="Qiandongnan" specimenCode="Ar 45219" specimenCount="2" specimenCount-male="2" stateProvince="Guizhou" typeStatus="paratype">
<emphasis id="7510BD6BDB755EDA44E2AD0548AD8561" bold="true">
<emphasis id="8B2E86802D988A119B8E85C365620EF7" bold="true" italics="true">
<typeStatus id="91D205DFDE6E9F2448014A3AB8429CF5" type="paratype">Paratypes</typeStatus>
</emphasis>
:
<collectingCountry id="FF0A5CDE61F5661EA546D1502609902D" name="China">China</collectingCountry>
</emphasis>
•
<specimenCount id="F49EE154DF5CDEA6FFDFBB1D44949FE7" count="2" type="male">2 ♂</specimenCount>
; same data as for holotype;
<named-content id="0F1CE3D3E6C546DA57DDB370BF2573DD" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/institute-zoology-chinese-academy-sciences" xlink_title="Institute of Zoology, Chinese Academy of Sciences">
<collectionCode id="812EA57D8F8C79D252050EFAAB5F46BD" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" type="">IZCAS</collectionCode>
</named-content>
-
<specimenCode id="CF83204640C68F79DA488195EE95E32B">Ar 45219</specimenCode>
–20
</materialsCitation>
•
<materialsCitation id="2F067B092FA1149DDB020120AFDEDA63" collectingDate="2007-05-23" collectionCode="IZCAS" collectorName="J. Liu &amp; Y. Lin" country="China" county="Liping County" elevation="544" latitude="25.9838" location="Yongcong Town" longLatPrecision="1" longitude="109.11632" municipality="Qiandongnan" specimenCode="Ar 45221" specimenCount="5" specimenCount-female="5" stateProvince="Guizhou" typeStatus="paratype">
<specimenCount id="C6DB5BB0B6C7F5E54802BAB3CD86A0C4" count="5" type="female">5 ♀</specimenCount>
; same data as for holotype;
<named-content id="E76714BD7A29F3DA7CF7F5120F4122E3" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/institute-zoology-chinese-academy-sciences" xlink_title="Institute of Zoology, Chinese Academy of Sciences">
<collectionCode id="C4D81800021B380120F6DE07E3972388" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" type="">IZCAS</collectionCode>
</named-content>
-
<specimenCode id="0FDF0E2A217F64E258E8DEF38F64E2BB">Ar 45221</specimenCode>
–25
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ELVAI" type="diagnosis">
<paragraph id="2795AAE6A9C75A6DAD9B328A29587844">
<heading id="46461715F0470051C71609C0F407FA48" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="1CFD74FD1D9C2068E2D8D6BFC6ECC7FF">
The new species resembles
<taxonomicName id="119DB947A3C309EC2BE0C6376338C3C9" authorityName="Wang &amp; Li &amp; Li &amp; Yao" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="nayong" status="sp. nov.">
<emphasis id="8CE892AD515DC0DD6481B8EF6BF7C783" italics="true">B. nayong</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F2E04392E5C5A21F3EBB0381855EA557" rank="species">sp. nov.</taxonomicNameLabel>
(Figs
<figureCitation id="1E4E5F4D178CEEED3BF48E7B533FA19C" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Belisana nayong sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at ventro-subdistal membranous process, arrow 2 points at prolatero-distal membranous lamella, arrow 3 points at dorso-distal spine D retrolateral view). Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.20 (A, B); 0.05 (C, D)." figureDoi="10.3897/zookeys.1216.132561.figure8" httpUri="https://binary.pensoft.net/fig/1161053">8</figureCitation>
,
<figureCitation id="622E8C4924F57F1F2FEBA42E81FFF31E" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Belisana nayong sp. nov., holotype male (C – F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E – H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A – D); 0.40 (E – H)." figureDoi="10.3897/zookeys.1216.132561.figure9" httpUri="https://binary.pensoft.net/fig/1161054">9</figureCitation>
,
<figureCitation id="882F96A6636A30D0825B55571B56B964" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Procursus in prolateral and retrolateral views (The arrows point at the same structures as those shown in the photos of each species) A, B Belisana bijie sp. nov. C, D B. liupanshui sp. nov. E, F B. majiang sp. nov. G, H B. nayong sp. nov. I, J B. qixingguan sp. nov. K, L B. xiuwen sp. nov. Abbreviation: f = flap. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure22" httpUri="https://binary.pensoft.net/fig/1161067">22 G, H</figureCitation>
,
<figureCitation id="9BE98CBDF53663D5472270A074CF9E2B" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Female genitalia in ventral and dorsal views A, B Belisana bijie sp. nov., arrow points at lateral sclerite C, D B. liupanshui sp. nov., arrow points at lateral sclerite E, F B. majiang sp. nov. G, H B. nayong sp. nov I, J B. qixingguan sp. nov., arrow points at sclerotized protrusion. Abbreviations: aa = anterior arch, ep = epigynal pocket, pp = pore plate. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure24" httpUri="https://binary.pensoft.net/fig/1161069">24 G, H</figureCitation>
) by having similar bulbal apophysis (hooked; Fig.
<figureCitation id="CD13860494FC9DE9E9BA303F206C7449" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Belisana yongcong sp. nov., holotype male (C – F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E – H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (A – D); 0.40 (E – H)." figureDoi="10.3897/zookeys.1216.132561.figure15" httpUri="https://binary.pensoft.net/fig/1161060">15 C</figureCitation>
) and vulva (anterior arch ridge-shaped, pore plates curved, long elliptic and 8 × longer than wide; Figs
<figureCitation id="2843106302A76D2509D6A8683A91BB3E" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Belisana yongcong sp. nov., holotype male (C – F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E – H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (A – D); 0.40 (E – H)." figureDoi="10.3897/zookeys.1216.132561.figure15" httpUri="https://binary.pensoft.net/fig/1161060">15 B</figureCitation>
,
<figureCitation id="E37200AAE4079D63407FF236AF767967" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Female genitalia in ventral and dorsal views A, B Belisana xiuwen sp. nov. C, D B. yongcong sp. nov. E, F B. zhangi Tong &amp; Li, 2007 G, H B. zhouxi sp. nov. I, J B. zunyi sp. nov. Abbreviations: aa = anterior arch, ep = epigynal pocket, pp = pore plate. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure25" httpUri="https://binary.pensoft.net/fig/1161070">25 D</figureCitation>
), but can be distinguished by procursus with blunt ventro-subdistal membranous process (arrow
<quantity id="FC26D145C072BC099FAB6894D94BCECF" metricMagnitude="-2" metricUnit="m" metricValue="2.54" unit="in" value="1.0">1 in</quantity>
Figs
<figureCitation id="B504101C84DF2279B4E2E219D224FD8A" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Belisana yongcong sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at ventro-subdistal membranous lamella, arrow 2 points at prolatero-distal membranous lamella, arrow 3 points at sclerotized dorso-subdistal apophysis D retrolateral view). Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.10 (A, B); 0.02 (C, D)." figureDoi="10.3897/zookeys.1216.132561.figure14" httpUri="https://binary.pensoft.net/fig/1161059">14 C</figureCitation>
,
<figureCitation id="1048AE44E8C248B3E147744C5FA9C013" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Procursus in prolateral and retrolateral views (The arrows point at the same structures as those shown in the photos of each species) A, B Belisana yongcong sp. nov. C, D B. zhangi Tong &amp; Li, 2007 E, F B. zhouxi sp. nov. G, H B. zunyi sp. nov. Abbreviation: f = flap. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure23" httpUri="https://binary.pensoft.net/fig/1161068">23 A</figureCitation>
vs pointed), by male cheliceral distal apophyses pointing outwards (
<abbrev id="ABBRID0EAXAI" xlink_title="distal apophysis">da</abbrev>
in Fig.
<figureCitation id="985480961DECC2C058705294C8C462F2" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Belisana yongcong sp. nov., holotype male (C – F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E – H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (A – D); 0.40 (E – H)." figureDoi="10.3897/zookeys.1216.132561.figure15" httpUri="https://binary.pensoft.net/fig/1161060">15 D</figureCitation>
vs inwards), and by epigyne with lateral pockets (
<abbrev id="ABBRID0EIXAI" xlink_title="epigynal pocket">ep</abbrev>
in Figs
<figureCitation id="12F7DED755AB284476EEC0F42E0E86E5" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Belisana yongcong sp. nov., holotype male (C – F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E – H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (A – D); 0.40 (E – H)." figureDoi="10.3897/zookeys.1216.132561.figure15" httpUri="https://binary.pensoft.net/fig/1161060">15 A</figureCitation>
,
<figureCitation id="99D785FC437874EB4789144A48EBB7FA" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Female genitalia in ventral and dorsal views A, B Belisana xiuwen sp. nov. C, D B. yongcong sp. nov. E, F B. zhangi Tong &amp; Li, 2007 G, H B. zhouxi sp. nov. I, J B. zunyi sp. nov. Abbreviations: aa = anterior arch, ep = epigynal pocket, pp = pore plate. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure25" httpUri="https://binary.pensoft.net/fig/1161070">25 C</figureCitation>
vs postero-median).
</paragraph>
<caption id="28AA904B691A0D2EE386C1AF712DC176" ID-DOI="10.3897/zookeys.1216.132561.figure14" ID-arpha="5F656357-BA10-542A-A335-57B3191C9436" httpUri="https://binary.pensoft.net/fig/1161059" startId="F14">
<paragraph id="1000EBE2F9E6775235D01D47D53E983F">
<label id="3B89632B84BA412D5D18A1B9509517FC">Figure 14.</label>
</paragraph>
<paragraph id="119691F24EF2F295C83927554B3F91DB">
<taxonomicName id="6251AB9AD8060D9770E5C751DA2DC841" authorityName="Wang &amp; Li &amp; Li &amp; Yao" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="yongcong" status="sp. nov.">
<emphasis id="BC795235696A52F38F63B839D693D40F" italics="true">Belisana yongcong</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5263C2A0D46BE346B4A5664D768FD913" rank="species">sp. nov.</taxonomicNameLabel>
, holotype male
<emphasis id="3A78C3A7EF0C0122E49050D13A5AF9B7" bold="true">A, B</emphasis>
palp (
<emphasis id="2C195939DEBE43C6239FAA68D591F92F" bold="true">A</emphasis>
prolateral view
<emphasis id="256038764CA43EEE36C9327C574F5A53" bold="true">B</emphasis>
retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion)
<emphasis id="B87181E1868D6F5BE58EBD6FF5095234" bold="true">C, D</emphasis>
distal part of procursus (
<emphasis id="5E24673EDCA1E140F592EF9E28714883" bold="true">C</emphasis>
prolateral view, arrow 1 points at ventro-subdistal membranous lamella, arrow 2 points at prolatero-distal membranous lamella, arrow 3 points at sclerotized dorso-subdistal apophysis
<emphasis id="628ED961383680AA06F76F74113ACBFD" bold="true">D</emphasis>
retrolateral view). Abbreviations: b = bulb,
<abbrev id="ABBRID0ETYAI" xlink_title="bulbal apophysis">ba</abbrev>
= bulbal apophysis, e = embolus, f = flap,
<abbrev id="ABBRID0EXYAI" xlink_title="procursus">pr</abbrev>
= procursus. Scale bars: 0.10 (
<emphasis id="B9EAEFC6EE36273436A9A18B9EC74B3B" bold="true">A, B</emphasis>
); 0.02 (
<emphasis id="9FD3B9ADEDF68CD8F9B26BDD1DF16BFF" bold="true">C, D</emphasis>
).
</paragraph>
</caption>
<caption id="7651A7CFFDBC6874C1FC2E457EFAD1F5" ID-DOI="10.3897/zookeys.1216.132561.figure15" ID-arpha="13B34B84-B654-5554-97FE-D90C46E63039" httpUri="https://binary.pensoft.net/fig/1161060" startId="F15">
<paragraph id="50CA1F7E56785CAF3A0888BA33615A91">
<label id="898016993234B2ADC3E9471297C68BBB">Figure 15.</label>
</paragraph>
<paragraph id="231D4D7E57CCA6DAB44E7239BABB5DDE">
<taxonomicName id="F858B08340DE99DD97D3262095ED53E2" authorityName="Wang &amp; Li &amp; Li &amp; Yao" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="yongcong" status="sp. nov.">
<emphasis id="759FE0722EAD5701D79221F189FECCDF" italics="true">Belisana yongcong</emphasis>
</taxonomicName>
<taxonomicNameLabel id="13F136D813C64C52063452865CF4618F" rank="species">sp. nov.</taxonomicNameLabel>
, holotype male (
<emphasis id="AEB485E1234EC40535987DD72D332449" bold="true">C – F</emphasis>
) and paratype female (
<emphasis id="0B8C49BFC86698285101FDD2E7BDBE63" bold="true">A, B, G, H</emphasis>
)
<emphasis id="7977D7ECDE80976D9FE01F22699C6CE7" bold="true">A</emphasis>
epigyne, ventral view
<emphasis id="80D0921F800C9D5C5D1FCAD2291403FA" bold="true">B</emphasis>
vulva, dorsal view
<emphasis id="5132D8632D8539333A5FDB635DB1CBD7" bold="true">C</emphasis>
bulb, prolateral view
<emphasis id="BA36CE362838B858F7896EA09FE92DC0" bold="true">D</emphasis>
chelicerae, frontal view
<emphasis id="70D98B52164825BE43BAE5EB152813A2" bold="true">E – H</emphasis>
habitus (
<emphasis id="64AA3482369D6CB93768104216572FFB" bold="true">E, G</emphasis>
dorsal view
<emphasis id="92BC6861DDDC6FF65339E9BFE9D1E836" bold="true">F</emphasis>
lateral view
<emphasis id="D5646FE0BA9695ADEBC4D934AC6F5AD2" bold="true">H</emphasis>
ventral view). Abbreviations:
<abbrev id="ABBRID0EL1AI" xlink_title="anterior arch">aa</abbrev>
= anterior arch, b = bulb,
<abbrev id="ABBRID0EP1AI" xlink_title="bulbal apophysis">ba</abbrev>
= bulbal apophysis,
<abbrev id="ABBRID0ET1AI" xlink_title="distal apophysis">da</abbrev>
= distal apophysis, e = embolus,
<abbrev id="ABBRID0EX1AI" xlink_title="epigynal pocket">ep</abbrev>
= epigynal pocket,
<abbrev id="ABBRID0E21AI" xlink_title="proximo-lateral apophysis">pa</abbrev>
= proximo-lateral apophysis,
<abbrev id="ABBRID0E61AI" xlink_title="pore plate">pp</abbrev>
= pore plate. Scale bars: 0.10 (
<emphasis id="34C6D7C15A5A44C4B3B096A4FD4FB9E7" bold="true">A – D</emphasis>
); 0.40 (
<emphasis id="35E54E177E58ED3BEC13A5D17B05822C" bold="true">E – H</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EM2AI" type="description">
<paragraph id="C384EF4ECEC16358488FAF32C0F4C13B">
<heading id="8A10232D2C633604F7D2BF36883F1883" reason="title">Description.</heading>
</paragraph>
<paragraph id="DE5B6C3F53F45E32DE9CEEEA548329B3">
<emphasis id="D728FB96CF1C79A88577399BAB555851" bold="true">Male</emphasis>
(
<emphasis id="4DA5FEF8BB3FC7D6FC823153943788FF" bold="true">
<typeStatus id="5F8A5E677AA8C481B863FE6CDD74E923" type="holotype">holotype</typeStatus>
</emphasis>
): Total length 1.73 (1.82 with clypeus), prosoma 0.83 long, 0.67 wide, opisthosoma 0.90 long, 0.66 wide. Leg I: 14.76 (3.60, 0.34, 4.05, 5.58, 1.56), leg II: 12.45 (3.36, 0.32, 3.08, 4.60, 1.09), leg III: 8.21 (2.38, 0.26, 1.98, 2.84, 0.75), leg IV: 10.64 (3.20, 0.26, 2.66, 3.72, 0.80); tibia I
<abbrev id="ABBRID0EW2AI" xlink_title="length/diameter">L / d</abbrev>
: 46. Eye interdistances and diameters:
<abbrev id="ABBRID0E12AI" xlink_title="posterior median eye">PME</abbrev>
–
<abbrev id="ABBRID0E52AI" xlink_title="posterior median eye">PME</abbrev>
0.06,
<abbrev id="ABBRID0EC3AI" xlink_title="posterior median eye">PME</abbrev>
0.09,
<abbrev id="ABBRID0EG3AI" xlink_title="posterior median eye">PME</abbrev>
–
<abbrev id="ABBRID0EK3AI" xlink_title="anterior lateral eye">ALE</abbrev>
0.02. Sternum width / length: 0.54 / 0.49. Habitus as in Fig.
<figureCitation id="9C6755D0F305F25A70374BECBF8653D1" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Belisana yongcong sp. nov., holotype male (C – F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E – H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (A – D); 0.40 (E – H)." figureDoi="10.3897/zookeys.1216.132561.figure15" httpUri="https://binary.pensoft.net/fig/1161060">15 E, F</figureCitation>
. Dorsal shield of prosoma yellowish, with brownish radiating marks; clypeus and sternum yellowish. Legs whitish, without darker rings. Opisthosoma yellowish, without spots. Thoracic furrow absent. Clypeus unmodified. Chelicerae with pair of proximo-lateral apophyses (
<abbrev id="ABBRID0ES3AI" xlink_title="proximo-lateral apophysis">pa</abbrev>
in Fig.
<figureCitation id="A05E4F44D4B9288EF0C454F6A6B95C5F" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Belisana yongcong sp. nov., holotype male (C – F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E – H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (A – D); 0.40 (E – H)." figureDoi="10.3897/zookeys.1216.132561.figure15" httpUri="https://binary.pensoft.net/fig/1161060">15 D</figureCitation>
) and pair of curved distal apophyses (distance between tips: 0.32;
<abbrev id="ABBRID0E13AI" xlink_title="distal apophysis">da</abbrev>
in Fig.
<figureCitation id="9C7C5A2D0F97175A200EBAA24975AD18" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Belisana yongcong sp. nov., holotype male (C – F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E – H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (A – D); 0.40 (E – H)." figureDoi="10.3897/zookeys.1216.132561.figure15" httpUri="https://binary.pensoft.net/fig/1161060">15 D</figureCitation>
). Palp as in Fig.
<figureCitation id="2A58551CC2EAB6FE9AB573CA332553D5" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Belisana yongcong sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at ventro-subdistal membranous lamella, arrow 2 points at prolatero-distal membranous lamella, arrow 3 points at sclerotized dorso-subdistal apophysis D retrolateral view). Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.10 (A, B); 0.02 (C, D)." figureDoi="10.3897/zookeys.1216.132561.figure14" httpUri="https://binary.pensoft.net/fig/1161059">14 A, B</figureCitation>
; trochanter with ventral apophysis (as long as wide, arrow
<quantity id="0ECB97B87BE3C5414E71E7AA29963AA8" metricMagnitude="-2" metricUnit="m" metricValue="2.54" unit="in" value="1.0">1 in</quantity>
Fig.
<figureCitation id="8BFB77AA75AB499CD665B24B2EA1E911" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Belisana yongcong sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at ventro-subdistal membranous lamella, arrow 2 points at prolatero-distal membranous lamella, arrow 3 points at sclerotized dorso-subdistal apophysis D retrolateral view). Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.10 (A, B); 0.02 (C, D)." figureDoi="10.3897/zookeys.1216.132561.figure14" httpUri="https://binary.pensoft.net/fig/1161059">14 B</figureCitation>
); femur with retrolatero-proximal protrusion (arrow
<quantity id="AA74845FFA4F8E06B46F0A67D0774BAD" metricMagnitude="-2" metricUnit="m" metricValue="5.08" unit="in" value="2.0">2 in</quantity>
Fig.
<figureCitation id="F30DC805ACC49C5DC440F2A555DE929F" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Belisana yongcong sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at ventro-subdistal membranous lamella, arrow 2 points at prolatero-distal membranous lamella, arrow 3 points at sclerotized dorso-subdistal apophysis D retrolateral view). Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.10 (A, B); 0.02 (C, D)." figureDoi="10.3897/zookeys.1216.132561.figure14" httpUri="https://binary.pensoft.net/fig/1161059">14 B</figureCitation>
); procursus simple proximally but complex distally, with ventro-subdistal membranous lamella (arrow
<quantity id="7A835CC1489810D1E06F6F93FEA47A94" metricMagnitude="-2" metricUnit="m" metricValue="2.54" unit="in" value="1.0">1 in</quantity>
Figs
<figureCitation id="61058A9F1C98FB55DDD07EB362C6A8C5" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Belisana yongcong sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at ventro-subdistal membranous lamella, arrow 2 points at prolatero-distal membranous lamella, arrow 3 points at sclerotized dorso-subdistal apophysis D retrolateral view). Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.10 (A, B); 0.02 (C, D)." figureDoi="10.3897/zookeys.1216.132561.figure14" httpUri="https://binary.pensoft.net/fig/1161059">14 C</figureCitation>
,
<figureCitation id="4C6D044C21FE8D3ABE024E6E55FFA5FF" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Procursus in prolateral and retrolateral views (The arrows point at the same structures as those shown in the photos of each species) A, B Belisana yongcong sp. nov. C, D B. zhangi Tong &amp; Li, 2007 E, F B. zhouxi sp. nov. G, H B. zunyi sp. nov. Abbreviation: f = flap. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure23" httpUri="https://binary.pensoft.net/fig/1161068">23 A</figureCitation>
) bearing proximally sclerotized part, prolatero-distal membranous lamella (arrow
<quantity id="7A4CC4CD8249149FE8EBAD45F7B8DE93" metricMagnitude="-2" metricUnit="m" metricValue="5.08" unit="in" value="2.0">2 in</quantity>
Figs
<figureCitation id="0A279E94052738304385CC4611B9EF4A" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Belisana yongcong sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at ventro-subdistal membranous lamella, arrow 2 points at prolatero-distal membranous lamella, arrow 3 points at sclerotized dorso-subdistal apophysis D retrolateral view). Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.10 (A, B); 0.02 (C, D)." figureDoi="10.3897/zookeys.1216.132561.figure14" httpUri="https://binary.pensoft.net/fig/1161059">14 C</figureCitation>
,
<figureCitation id="4E03A66A1D2063D9BA459669FB11CCB7" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Procursus in prolateral and retrolateral views (The arrows point at the same structures as those shown in the photos of each species) A, B Belisana yongcong sp. nov. C, D B. zhangi Tong &amp; Li, 2007 E, F B. zhouxi sp. nov. G, H B. zunyi sp. nov. Abbreviation: f = flap. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure23" httpUri="https://binary.pensoft.net/fig/1161068">23 A</figureCitation>
) bearing proximally sclerotized part, sclerotized dorso-subdistal apophysis (arrow
<quantity id="6E83B98BE943B5E3B1013656D2902C6B" metricMagnitude="-2" metricUnit="m" metricValue="7.62" unit="in" value="3.0">3 in</quantity>
Figs
<figureCitation id="68B863C592A29446A24CE71B0EFDB0BA" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Belisana yongcong sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at ventro-subdistal membranous lamella, arrow 2 points at prolatero-distal membranous lamella, arrow 3 points at sclerotized dorso-subdistal apophysis D retrolateral view). Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.10 (A, B); 0.02 (C, D)." figureDoi="10.3897/zookeys.1216.132561.figure14" httpUri="https://binary.pensoft.net/fig/1161059">14 C</figureCitation>
,
<figureCitation id="5B8D912C0A1C514C5510FAC81E62C922" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Procursus in prolateral and retrolateral views (The arrows point at the same structures as those shown in the photos of each species) A, B Belisana yongcong sp. nov. C, D B. zhangi Tong &amp; Li, 2007 E, F B. zhouxi sp. nov. G, H B. zunyi sp. nov. Abbreviation: f = flap. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure23" httpUri="https://binary.pensoft.net/fig/1161068">23 A</figureCitation>
), and retrolateral membranous flap (f in Figs
<figureCitation id="9C88EB8227F36B0FD8333870C2A81287" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Belisana yongcong sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at ventro-subdistal membranous lamella, arrow 2 points at prolatero-distal membranous lamella, arrow 3 points at sclerotized dorso-subdistal apophysis D retrolateral view). Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.10 (A, B); 0.02 (C, D)." figureDoi="10.3897/zookeys.1216.132561.figure14" httpUri="https://binary.pensoft.net/fig/1161059">14 D</figureCitation>
,
<figureCitation id="A5C22BA9DB46BF04B5ED50138E4113B3" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Procursus in prolateral and retrolateral views (The arrows point at the same structures as those shown in the photos of each species) A, B Belisana yongcong sp. nov. C, D B. zhangi Tong &amp; Li, 2007 E, F B. zhouxi sp. nov. G, H B. zunyi sp. nov. Abbreviation: f = flap. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure23" httpUri="https://binary.pensoft.net/fig/1161068">23 B</figureCitation>
); bulb with hooked apophysis (
<abbrev id="ABBRID0EO5AI" xlink_title="bulbal apophysis">ba</abbrev>
in Fig.
<figureCitation id="D033D622A7CEF254B7B33DA5C740207C" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Belisana yongcong sp. nov., holotype male (C – F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E – H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (A – D); 0.40 (E – H)." figureDoi="10.3897/zookeys.1216.132561.figure15" httpUri="https://binary.pensoft.net/fig/1161060">15 C</figureCitation>
) and simple embolus (e in Fig.
<figureCitation id="446A9E10F5859E521C49293E8897C421" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Belisana yongcong sp. nov., holotype male (C – F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E – H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (A – D); 0.40 (E – H)." figureDoi="10.3897/zookeys.1216.132561.figure15" httpUri="https://binary.pensoft.net/fig/1161060">15 C</figureCitation>
). Retrolateral trichobothria on tibia I at 10 % proximally; legs with short vertical setae on metatarsi; tarsus I with 20 distinct pseudosegments.
</paragraph>
<paragraph id="AC16706B4577CEA243C59A093FE2F7D3">
<emphasis id="2BE1311248F5C67F2C62B9108F06706B" bold="true">Female</emphasis>
(
<emphasis id="27FFAB6D56C589F6A701092DA847D6F0" bold="true">
<typeStatus id="8C7A2ABD09A330240C16D10E37753BA3" type="paratype">paratype</typeStatus>
</emphasis>
,
<named-content id="E4D38F220800F03040D157A45E2F215E" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/institute-zoology-chinese-academy-sciences" xlink_title="Institute of Zoology, Chinese Academy of Sciences">
<collectionCode id="A8AC9C5B037B390B1C9951C5F9263804" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" type="">IZCAS</collectionCode>
</named-content>
-
<specimenCode id="4BDB0D49907D78EF00D1472BFA81A8C4">Ar 45221</specimenCode>
): Similar to male, habitus as in Fig.
<figureCitation id="F5458D05EE807C2C01B66EDB139C47E3" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Belisana yongcong sp. nov., holotype male (C – F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E – H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (A – D); 0.40 (E – H)." figureDoi="10.3897/zookeys.1216.132561.figure15" httpUri="https://binary.pensoft.net/fig/1161060">15 G, H</figureCitation>
. Total length 2.17 (2.29 with clypeus), prosoma 0.87 long, 0.76 wide, opisthosoma 1.30 long, 1.08 wide; tibia I: 3.65; tibia I
<abbrev id="ABBRID0EI6AI" xlink_title="length/diameter">L / d</abbrev>
: 47. Eye interdistances and diameters:
<abbrev id="ABBRID0EM6AI" xlink_title="posterior median eye">PME</abbrev>
–
<abbrev id="ABBRID0EQ6AI" xlink_title="posterior median eye">PME</abbrev>
0.09,
<abbrev id="ABBRID0EU6AI" xlink_title="posterior median eye">PME</abbrev>
0.08,
<abbrev id="ABBRID0EY6AI" xlink_title="posterior median eye">PME</abbrev>
–
<abbrev id="ABBRID0E36AI" xlink_title="anterior lateral eye">ALE</abbrev>
0.02. Sternum width / length: 0.59 / 0.45. Epigyne simple and flat, posteriorly curved, with pair of lateral pockets 0.35 apart (
<abbrev id="ABBRID0EAABI" xlink_title="epigynal pocket">ep</abbrev>
in Figs
<figureCitation id="17649A7F9536611652566ADD793577F5" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Belisana yongcong sp. nov., holotype male (C – F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E – H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (A – D); 0.40 (E – H)." figureDoi="10.3897/zookeys.1216.132561.figure15" httpUri="https://binary.pensoft.net/fig/1161060">15 A</figureCitation>
,
<figureCitation id="AC8C6306480D20D1B34225E3D211E1D3" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Female genitalia in ventral and dorsal views A, B Belisana xiuwen sp. nov. C, D B. yongcong sp. nov. E, F B. zhangi Tong &amp; Li, 2007 G, H B. zhouxi sp. nov. I, J B. zunyi sp. nov. Abbreviations: aa = anterior arch, ep = epigynal pocket, pp = pore plate. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure25" httpUri="https://binary.pensoft.net/fig/1161070">25 C</figureCitation>
). Vulva with ridge-shaped anterior arch (
<abbrev id="ABBRID0EMABI" xlink_title="anterior arch">aa</abbrev>
in Figs
<figureCitation id="F4835899F898D9E9E32B8B0FE54CF921" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Belisana yongcong sp. nov., holotype male (C – F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E – H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (A – D); 0.40 (E – H)." figureDoi="10.3897/zookeys.1216.132561.figure15" httpUri="https://binary.pensoft.net/fig/1161060">15 B</figureCitation>
,
<figureCitation id="B211FB56E6AEDA995D5467F3D70587B2" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Female genitalia in ventral and dorsal views A, B Belisana xiuwen sp. nov. C, D B. yongcong sp. nov. E, F B. zhangi Tong &amp; Li, 2007 G, H B. zhouxi sp. nov. I, J B. zunyi sp. nov. Abbreviations: aa = anterior arch, ep = epigynal pocket, pp = pore plate. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure25" httpUri="https://binary.pensoft.net/fig/1161070">25 D</figureCitation>
) and pair of curved, long elliptic pore plates (9 × longer than wide,
<abbrev id="ABBRID0EYABI" xlink_title="pore plate">pp</abbrev>
in Figs
<figureCitation id="D05F7DBE1FF48939432F919E1CC4C87B" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Belisana yongcong sp. nov., holotype male (C – F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E – H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (A – D); 0.40 (E – H)." figureDoi="10.3897/zookeys.1216.132561.figure15" httpUri="https://binary.pensoft.net/fig/1161060">15 B</figureCitation>
,
<figureCitation id="6A0F1794CEA46D96A4FBDB3C4910EE81" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Female genitalia in ventral and dorsal views A, B Belisana xiuwen sp. nov. C, D B. yongcong sp. nov. E, F B. zhangi Tong &amp; Li, 2007 G, H B. zhouxi sp. nov. I, J B. zunyi sp. nov. Abbreviations: aa = anterior arch, ep = epigynal pocket, pp = pore plate. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure25" httpUri="https://binary.pensoft.net/fig/1161070">25 D</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EEBBI" type="variation">
<paragraph id="3ED59D51287080444CCCC47E38A02939">
<heading id="4B6FE540057CD67AA10B95901B9E9777" reason="title">Variation.</heading>
</paragraph>
<paragraph id="DD48DE64B5AA070513ADB2802248652E">
Tibia I in
<specimenCount id="BAF72977FCD93D3C11CFC5174AD92088" count="1" type="male">one male</specimenCount>
<typeStatus id="1D395A079AC9B42EF93E8FBB32C37232" type="paratype">paratype</typeStatus>
(
<named-content id="5188D2E6711D25D6BA8750183D97C887" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/institute-zoology-chinese-academy-sciences" xlink_title="Institute of Zoology, Chinese Academy of Sciences">
<collectionCode id="110D1581519D6F7623C58BFFD36E5EDA" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" type="">IZCAS</collectionCode>
</named-content>
-
<specimenCode id="3DDB7B9EAE84A7ED27587F97912B2A33">Ar 45219</specimenCode>
): 5.32 (leg I missing in
<named-content id="8B41E652133493D584A9A3BBF8347965" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/institute-zoology-chinese-academy-sciences" xlink_title="Institute of Zoology, Chinese Academy of Sciences">
<collectionCode id="691B5AD71C96E69C3C7ED0A82A64C3A8" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" type="">IZCAS</collectionCode>
</named-content>
-
<specimenCode id="736E97FDC6360708BA035C3FDB097AD8">Ar 45220</specimenCode>
). Tibia I in the other
<specimenCount id="CA90D9E641578E9A0EAE977C327A6EF1" count="4" type="female">four female</specimenCount>
<typeStatus id="03873553D8179C5B55E895771A25A974" type="paratype">paratypes</typeStatus>
(
<named-content id="7F0E175122BBA16B3EDFC35CDE602FD9" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/institute-zoology-chinese-academy-sciences" xlink_title="Institute of Zoology, Chinese Academy of Sciences">
<collectionCode id="4E4A0D877A15273E27D838A23D63598D" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" type="">IZCAS</collectionCode>
</named-content>
-
<specimenCode id="DEA93A5D0F545B07A9221FD49A3B02A5">Ar 45222</specimenCode>
–25): 3.76, 3.85, 3.96, 4.16.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EWBBI" type="habitat">
<paragraph id="70C6274EE2EED3908D97A32E1B6E5D48">
<heading id="7FCD3065B55F97AEE261C2B1DD0342C0" reason="title">Habitat.</heading>
</paragraph>
<paragraph id="D7B707704703EF5EC72D9FB3F9E5F16B">The species was found in the dark zone inside the cave.</paragraph>
</subSubSection>
<subSubSection id="SECID0E2BBI" type="distribution">
<paragraph id="66124BB45953CF2D50BC2B418CAD4473">
<heading id="0D694D3CA342C1858376AE16463395CE" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="7D59A0F4AA036FF30BE19D0E7F770F10">
<collectingCountry id="7F7BC44E4B77FB932647FEB139E81B79" name="China">China</collectingCountry>
(
<collectingRegion id="BD0C17938584A71879BFE3D3A228FB45" country="China" name="Guizhou">Guizhou</collectingRegion>
, type locality; Fig.
<figureCitation id="1E7C05C3E9872644E583E2786270CD14" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Distribution records of the Belisana species from Guizhou, China 1 Belisana bijie sp. nov. 2 B. daji Chen, Zhang &amp; Zhu, 2009 3 B. douqing Chen, Zhang &amp; Zhu, 2009 4 B. galeiformis Zhang &amp; Peng, 2011 5 B. lii Chen, Yu &amp; Guo, 2016 6 B. liupanshui sp. nov. 7 B. majiang sp. nov. 8 B. nayong sp. nov. 9 B. qixingguan sp. nov. 10 B. wangchengi Wang, Yao &amp; Zhang, 2024 11 B. xishui Chen, Zhang &amp; Zhu, 2009 12 B. xiuwen sp. nov. 13 B. yanhe Chen, Zhang &amp; Zhu, 2009 14 B. yongcong sp. nov. 15 B. yuhaoi Yang &amp; Yao, 2023 16 B. zhangi Tong &amp; Li, 2007 17 B. zhouxi sp. nov. 18 B. zunyi sp. nov." figureDoi="10.3897/zookeys.1216.132561.figure1" httpUri="https://binary.pensoft.net/fig/1161046">1</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EFCBI" type="etymology">
<paragraph id="19F10BA43FD171EB838E08F45EFD5D03">
<heading id="73EC9B34C7D3EB9704339566F8457FFC" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="64EAD874C0B143B813A4687881A7F143">The specific name refers to the type locality; noun in apposition.</paragraph>
</subSubSection>
</treatment>
</document>
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