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<document id="1F4B33482F9F9417C0752D4737AAC0AC" ID-CLB-Dataset="21992" ID-DOI="10.5852/ejt.2017.355" ID-GBIF-Dataset="d8be473a-53c9-4ce6-8bd4-ec818d87adc0" ID-ISSN="2118-9773" ID-Zenodo-Dep="3836313" ID-ZooBank="80D35410-38CD-42FB-AFAF-605B24FC0A8B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1589997846232" checkinUser="carolina" docAuthor="Jocque, Merlijn, Wellens, Siel, Andrianarivosoa, J. D., Rakotondraparany, F., Seing, Sam The &amp; Jocqué, Rudy" docDate="2017" docId="0A076352FF9EFFCB6DDF29306B32FDC9" docLanguage="en" docName="ejt-2017-355.pdf.imf" docOrigin="European Journal of Taxonomy 355" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Ocyale Audouin 1823" docType="key" docVersion="7" lastPageNumber="5" masterDocId="F63E1B2AFF9CFFCF6F4A2F006E45FFF4" masterDocTitle="A new species of Ocyale (Araneae, Lycosidae) from Madagascar, with first observations on the biology of a representative in the genus" masterLastPageNumber="13" masterPageNumber="1" pageNumber="3" updateTime="1698840500717" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="C1B7457AC954CD492BD852CEFC27E11C">A new species of Ocyale (Araneae, Lycosidae) from Madagascar, with first observations on the biology of a representative in the genus</mods:title>
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<mods:namePart id="0DF3F5371D30DED02B17D6E2FEC84520">Jocque, Merlijn</mods:namePart>
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<mods:affiliation id="0B791EB57A03C0A5CC898446A71483CE">Biodiversity Inventory for Conservation (BINCO), Walmersumstraat 44, 3380 Glabbeek, Belgium. &amp; Operation Wallacea, Wallacea House, Old Bolingbroke, Lincolnshire, PE 23 4 EX United Kingdom. &amp; Aquatic and Terrestrial Ecology (ATECO), Royal Belgian Institute of Natural Sciences (RBINS), Vautierstraat 29, 1000 Brussels, Belgium. &amp; Email: merlijnjocque @ gmail. com &amp; urn: lsid: zoobank. org: author: 6 F 4 C 1 B 5 C-BDD 9 - 4217 - 8 F 29 - 95 ADA 5 E 9 F 4 D 3</mods:affiliation>
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<mods:namePart id="DAA36336095B31558DE69F708D006484">Wellens, Siel</mods:namePart>
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<mods:affiliation id="0F37082D7BB86A0BA7BBF22553B1CAC1">Biodiversity Inventory for Conservation (BINCO), Walmersumstraat 44, 3380 Glabbeek, Belgium. &amp; Operation Wallacea, Wallacea House, Old Bolingbroke, Lincolnshire, PE 23 4 EX United Kingdom. &amp; Email: siel. wellens @ gmail. com &amp; urn: lsid: zoobank. org: author: B 549 CCD 8 - 9 A 34 - 4 FCD- 996 F-C 49633331 D 33</mods:affiliation>
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<mods:namePart id="2708C5FF8965D0D517952486D393D5DC">Andrianarivosoa, J. D.</mods:namePart>
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<mods:affiliation id="CEED5E19AD6DFD056761807EAB8E1C4F">Mention Zoologie et Biologie Animale (MZBA), Université d’Antananarivo, Madagascar. &amp; Email: davsandrianarivosoa @ gmail. com &amp; urn: lsid: zoobank. org: author: 8 F 3 B 36 C 3 - D 80 B- 455 D-B 48 E- 50 F 27491167 C</mods:affiliation>
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<mods:namePart id="CC8A2F84E426ED735802F6C00B1851BD">Rakotondraparany, F.</mods:namePart>
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<mods:affiliation id="1367ACD1F37D325E0D56DB2F12682311">Mention Zoologie et Biologie Animale (MZBA), Université d’Antananarivo, Madagascar. &amp; Email: felitenrecs @ gmail. com &amp; urn: lsid: zoobank. org: author: 487 CF 2 F 0 - E 5 AC- 4 DF 0 - A 2 B 2 - 4 BCE 6 C 7 F 8 CB 3</mods:affiliation>
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<mods:namePart id="71361F9F055765752769131F41B9697F">Seing, Sam The</mods:namePart>
<mods:affiliation id="D3184E60F4D8222CDE7665D77771562B">Development and Biodiversity Conservation Action for Madagascar (DBCAM). Lot II A 93 L, Anjanahary, Antananarivo, Madagascar. &amp; Email: sam _ theseing @ yahoo. fr &amp; urn: lsid: zoobank. org: author: 28 F 7 A 019 - 1581 - 4 C 8 B-B 0 CE- 1 C 4948287 F 65</mods:affiliation>
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<mods:namePart id="403048D076829D0D773C2CE78F7FBDEC">Jocqué, Rudy</mods:namePart>
<mods:affiliation id="3380541FEA3CE496EF9BF7F1FA39557B">Royal Museum for Central Africa, Leuvense Steenweg 13, 3080 Tervuren, Belgium. &amp; urn: lsid: zoobank. org: author: CF 15016 C- 8 CD 1 - 4 C 9 D- 9021 - 44 CA 7 DC 7 A 5 D 5 &amp; Corresponding author: rudy. jocque @ africamuseum. be</mods:affiliation>
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<mods:title id="C51A3FAC36079673286A77D74DA5C722">European Journal of Taxonomy</mods:title>
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<mods:date id="8635389FEE846CACE0F5E2881A0BBB39">2017</mods:date>
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<mods:number id="6BC2D6C7406BCD510BAF7BE5F3D7286F">2017-10-03</mods:number>
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<mods:identifier id="4D11AE56BF2D4130FC37212B620ED8DA" type="DOI">10.5852/ejt.2017.355</mods:identifier>
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<mods:identifier id="2393A02673CA640F61318D2C42ACC0A1" type="Zenodo-Dep">3836313</mods:identifier>
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<treatment id="0A076352FF9EFFCB6DDF29306B32FDC9" ID-DOI="http://doi.org/10.5281/zenodo.3851685" ID-GBIF-Taxon="164243579" ID-Zenodo-Dep="3851685" LSID="urn:lsid:plazi:treatment:0A076352FF9EFFCB6DDF29306B32FDC9" httpUri="http://treatment.plazi.org/id/0A076352FF9EFFCB6DDF29306B32FDC9" lastPageId="4" lastPageNumber="5" pageId="2" pageNumber="3">
<subSubSection id="CAB481CFFF9EFFCD6DDF29306CF2F978" pageId="2" pageNumber="3" type="nomenclature">
<paragraph id="8211D244FF9EFFCD6DDF29306DDBF9BF" blockId="2.[661,926,1584,1611]" box="[661,926,1584,1611]" pageId="2" pageNumber="3">
<heading id="D9596528FF9EFFCD6DDF29306DDBF9BF" box="[661,926,1584,1611]" centered="true" fontSize="11" level="2" pageId="2" pageNumber="3" reason="2">
<taxonomicName id="45AEA9C7FF9EFFCD6DDF29306DDBF9BF" ID-CoL="66R7" authority="Audouin, 1823" authorityName="Audouin" authorityYear="1823" box="[661,926,1584,1611]" class="Arachnida" family="Lycosidae" genus="Ocyale" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="B0DA0E56FF9EFFCD6DDF29306CADF9BF" bold="true" box="[661,744,1584,1611]" italics="true" pageId="2" pageNumber="3">Ocyale</emphasis>
Audouin, 1823
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<heading id="D9596528FF9EFFCD6FF729726CF2F978" bold="true" box="[189,695,1649,1676]" fontSize="11" level="3" pageId="2" pageNumber="3" reason="3">
<emphasis id="B0DA0E56FF9EFFCD6FF729726CF2F978" bold="true" box="[189,695,1649,1676]" pageId="2" pageNumber="3">
Key to the Afrotropical species of
<taxonomicName id="45AEA9C7FF9EFFCD6D2E29716CF2F978" authorityName="Audouin" authorityYear="1823" box="[612,695,1649,1676]" class="Arachnida" family="Lycosidae" genus="Ocyale" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="B0DA0E56FF9EFFCD6D2E29716CF2F978" bold="true" box="[612,695,1649,1676]" italics="true" pageId="2" pageNumber="3">Ocyale</emphasis>
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<subSubSection id="CAB481CFFF9EFFCD6FF729A06C42F81F" pageId="2" pageNumber="3" type="discussion">
<paragraph id="8211D244FF9EFFCD6FF729A06D2DF94E" blockId="2.[189,872,1696,1722]" box="[189,872,1696,1722]" pageId="2" pageNumber="3">Letters refer to characters shown by arrows in key figures.</paragraph>
<paragraph id="8211D244FF9EFFCD6FF729E16FC9F90F" blockId="2.[189,396,1761,1787]" box="[189,396,1761,1787]" pageId="2" pageNumber="3">Species included:</paragraph>
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<taxonomicName id="45AEA9C7FF9EFFCD6FF728226C10F8C8" authority="Alderweireldt, 1996" authorityName="Alderweireldt" authorityYear="1996" box="[189,597,1826,1852]" class="Arachnida" family="Lycosidae" genus="Ocyale" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="dewinterae">
<emphasis id="B0DA0E56FF9EFFCD6FF728226F26F8C8" box="[189,355,1826,1852]" italics="true" pageId="2" pageNumber="3">O. dewinterae</emphasis>
<bibRefCitation id="E63FAFB5FF9EFFCD6E2328226C10F8C8" author="Alderweireldt M." box="[361,597,1826,1852]" pageId="2" pageNumber="3" pagination="103 - 111" refId="ref4420" refString="Alderweireldt M. 1996. A taxonomic revision of the genus Ocyale Audouin, 1826 in Africa (Araneae: Lycosidae). Journal of Natural History 30: 1349 - 1365. https: // doi. org / 10.1080 / 00222939600771261 Alderweireldt M. &amp; Jocque R. 1991. A remarkable new genus of wolf spiders from southwestern Spain (Araneae, Lycosidae). Bulletin van het Koninklijk Belgisch Instituut voor Natuurwetenschappen, Entomologie 61: 103 - 111." type="journal article" year="1996">Alderweireldt, 1996</bibRefCitation>
</taxonomicName>
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<specimenCount id="94A819CDFF9EFFCD6D3C28226CCDF8C8" box="[630,648,1826,1852]" pageId="2" pageNumber="3" type="female">♀</specimenCount>
</paragraph>
<paragraph id="8211D244FF9EFFCD6FF728456C62F8AB" blockId="2.[189,720,1826,2027]" box="[189,551,1860,1887]" pageId="2" pageNumber="3">
<taxonomicName id="45AEA9C7FF9EFFCD6FF728456C4FF8AA" authority="Roewer, 1960" authorityName="Roewer" authorityYear="1960" box="[189,522,1860,1887]" class="Arachnida" family="Lycosidae" genus="Ocyale" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="discrepans">
<emphasis id="B0DA0E56FF9EFFCD6FF728456F1AF8AB" box="[189,351,1861,1887]" italics="true" pageId="2" pageNumber="3">O. discrepans</emphasis>
Roewer, 1960
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<specimenCount id="94A819CDFF9EFFCD6D5B28456C62F8AB" box="[529,551,1861,1887]" pageId="2" pageNumber="3" type="female">♀</specimenCount>
</paragraph>
<paragraph id="8211D244FF9EFFCD6FF728686C95F876" blockId="2.[189,720,1826,2027]" box="[189,720,1895,1922]" pageId="2" pageNumber="3">
<taxonomicName id="45AEA9C7FF9EFFCD6FF728686C7EF876" authority="Jocque M. &amp; Jocque R." authorityName="Jocque M. &amp; Jocque R." authorityYear="2017" box="[189,571,1895,1922]" class="Arachnida" family="Lycosidae" genus="Ocyale" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="ghost" status="sp. nov.">
<emphasis id="B0DA0E56FF9EFFCD6FF728686F64F875" box="[189,289,1895,1922]" italics="true" pageId="2" pageNumber="3">O. ghost</emphasis>
Jocque M. &amp; Jocqué R.
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<taxonomicNameLabel id="ABE9B32DFF9EFFCD6D0828686CD8F876" box="[578,669,1896,1922]" pageId="2" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
<specimenCount id="94A819CDFF9EFFCD6DEE28686CFBF876" box="[676,702,1896,1922]" pageId="2" pageNumber="3" type="male">♂</specimenCount>
<specimenCount id="94A819CDFF9EFFCD6DF428686C95F876" box="[702,720,1896,1922]" pageId="2" pageNumber="3" type="female">♀</specimenCount>
</paragraph>
<paragraph id="8211D244FF9EFFCD6FF7288B6C25F851" blockId="2.[189,720,1826,2027]" box="[189,608,1930,1957]" pageId="2" pageNumber="3">
<taxonomicName id="45AEA9C7FF9EFFCD6FF7288B6C68F850" authority="Alderweireldt" authorityName="Alderweireldt" authorityYear="1996" box="[189,557,1930,1957]" class="Arachnida" family="Lycosidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="grandis">
<emphasis id="B0DA0E56FF9EFFCD6FF7288B6F7EF850" box="[189,315,1930,1957]" italics="true" pageId="2" pageNumber="3">O. grandis</emphasis>
<bibRefCitation id="E63FAFB5FF9EFFCD6E0A288A6C68F850" author="Alderweireldt M." box="[320,557,1930,1956]" pageId="2" pageNumber="3" pagination="103 - 111" refId="ref4420" refString="Alderweireldt M. 1996. A taxonomic revision of the genus Ocyale Audouin, 1826 in Africa (Araneae: Lycosidae). Journal of Natural History 30: 1349 - 1365. https: // doi. org / 10.1080 / 00222939600771261 Alderweireldt M. &amp; Jocque R. 1991. A remarkable new genus of wolf spiders from southwestern Spain (Araneae, Lycosidae). Bulletin van het Koninklijk Belgisch Instituut voor Natuurwetenschappen, Entomologie 61: 103 - 111." type="journal article" year="1996">Alderweireldt, 1996</bibRefCitation>
</taxonomicName>
<specimenCount id="94A819CDFF9EFFCD6D7E288B6C0BF851" box="[564,590,1931,1957]" pageId="2" pageNumber="3" type="male">♂</specimenCount>
<specimenCount id="94A819CDFF9EFFCD6D04288B6C25F851" box="[590,608,1931,1957]" pageId="2" pageNumber="3" type="female">♀</specimenCount>
</paragraph>
<paragraph id="8211D244FF9EFFCD6FF728AE6C4EF83C" blockId="2.[189,720,1826,2027]" box="[189,523,1965,1992]" pageId="2" pageNumber="3">
<taxonomicName id="45AEA9C7FF9EFFCD6FF728AE6F9DF83C" authority="Karsch, 1878" authorityName="Karsch" authorityYear="1878" box="[189,472,1965,1992]" class="Arachnida" family="Lycosidae" genus="Ocyale" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="guttata">
<emphasis id="B0DA0E56FF9EFFCD6FF728AE6F71F833" box="[189,308,1966,1992]" italics="true" pageId="2" pageNumber="3">O. guttata</emphasis>
Karsch, 1878
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<specimenCount id="94A819CDFF9EFFCD6E9528AE6FBCF83C" box="[479,505,1966,1992]" pageId="2" pageNumber="3" type="male">♂</specimenCount>
<specimenCount id="94A819CDFF9EFFCD6EB328AE6C4EF83C" box="[505,523,1966,1992]" pageId="2" pageNumber="3" type="female">♀</specimenCount>
</paragraph>
<paragraph id="8211D244FF9EFFCD6FF728D16C42F81F" blockId="2.[189,720,1826,2027]" box="[189,519,2001,2027]" pageId="2" pageNumber="3">
<taxonomicName id="45AEA9C7FF9EFFCD6FF728D16F91F81F" authority="Roewer, 1960" authorityName="Roewer" authorityYear="1960" box="[189,468,2001,2027]" class="Arachnida" family="Lycosidae" genus="Ocyale" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="pilosa">
<emphasis id="B0DA0E56FF9EFFCD6FF728D16F6CF81F" box="[189,297,2001,2027]" italics="true" pageId="2" pageNumber="3">O. pilosa</emphasis>
Roewer, 1960
</taxonomicName>
<specimenCount id="94A819CDFF9EFFCD6E9128D16FB0F81F" box="[475,501,2001,2027]" pageId="2" pageNumber="3" type="male">♂</specimenCount>
<specimenCount id="94A819CDFF9EFFCD6EBF28D16C42F81F" box="[501,519,2001,2027]" pageId="2" pageNumber="3" type="female">♀</specimenCount>
</paragraph>
</subSubSection>
<subSubSection id="CAB481CFFF9FFFCB6FF72E0B6B32FDC9" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4" type="key">
<key id="F63F8575FF9FFFCB6FF72E0B6B32FDC9" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4">
<keyStep id="395ACAE1FF9FFFCC6FF72E0B6B32FEBC" pageId="3" pageNumber="4">
<paragraph id="8211D244FF9FFFCC6FF72E0B6B32FED1" blockId="3.[189,1399,267,329]" box="[189,1399,267,294]" pageId="3" pageNumber="4">
<keyLead id="395F7171FF9FFFCC6FF72E0B6B32FED1" box="[189,1399,267,294]" pageId="3" pageNumber="4">1. Males ................................................................................................................................................. 2</keyLead>
</paragraph>
<paragraph id="8211D244FF9FFFCC6F8B2E2F6B32FEBC" blockId="3.[189,1399,267,329]" box="[193,1399,302,329]" pageId="3" pageNumber="4">
<keyLead id="395F7171FF9FFFCC6F8B2E2F6B32FEBC" box="[193,1399,302,329]" pageId="3" pageNumber="4">– Females ............................................................................................................................................. 6</keyLead>
</paragraph>
</keyStep>
<keyStep id="395ACAE1FF9FFFCC6FF72E706B32FE00" pageId="3" pageNumber="4">
<paragraph id="8211D244FF9FFFCC6FF72E706B32FE59" blockId="3.[189,1399,368,500]" pageId="3" pageNumber="4">
<keyLead id="395F7171FF9FFFCC6FF72E706B32FE59" pageId="3" pageNumber="4">
2. Small species: TL
<quantity id="45567FA1FF9FFFCC6E982E706C7EFE7E" box="[466,571,368,394]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" metricValueMax="8.0" metricValueMin="6.0" pageId="3" pageNumber="4" unit="mm" value="7.0" valueMax="8.0" valueMin="6.0">6–8 mm</quantity>
; distal part of cymbium shorter than bulbus (b) and bulbus clearly broadened (a) ..............................................................................
<taxonomicName id="45AEA9C7FF9FFFCC6C912E936B32FE59" ID-CoL="48L6Y" authority="Alderweireldt, 1996" authorityName="Alderweireldt" authorityYear="1996" box="[987,1399,403,430]" class="Arachnida" family="Lycosidae" genus="Ocyale" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="dewinterae">
<emphasis id="B0DA0E56FF9FFFCC6C912E936AC6FE5A" bold="true" box="[987,1155,403,430]" italics="true" pageId="3" pageNumber="4">O. dewinterae</emphasis>
<bibRefCitation id="E63FAFB5FF9FFFCC6BC02E936B32FE59" author="Alderweireldt M." box="[1162,1399,403,429]" pageId="3" pageNumber="4" pagination="103 - 111" refId="ref4420" refString="Alderweireldt M. 1996. A taxonomic revision of the genus Ocyale Audouin, 1826 in Africa (Araneae: Lycosidae). Journal of Natural History 30: 1349 - 1365. https: // doi. org / 10.1080 / 00222939600771261 Alderweireldt M. &amp; Jocque R. 1991. A remarkable new genus of wolf spiders from southwestern Spain (Araneae, Lycosidae). Bulletin van het Koninklijk Belgisch Instituut voor Natuurwetenschappen, Entomologie 61: 103 - 111." type="journal article" year="1996">Alderweireldt, 1996</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8211D244FF9FFFCC6F8B2EB76B32FE00" blockId="3.[189,1399,368,500]" pageId="3" pageNumber="4">
<keyLead id="395F7171FF9FFFCC6F8B2EB76B32FE00" pageId="3" pageNumber="4">
– Larger species, TL&gt;
<quantity id="45567FA1FF9FFFCC6EA82EB66C7CFE25" box="[482,569,438,465]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" pageId="3" pageNumber="4" unit="mm" value="16.0">16 mm</quantity>
; distal part of cymbium longer than bulbus (c) and bulbus clearly not broadened (d) .................................................................................................................................... 3
</keyLead>
</paragraph>
</keyStep>
<keyStep id="395ACAE1FF9FFFCC6FF72D1B6B32FD88" pageId="3" pageNumber="4">
<paragraph id="8211D244FF9FFFCC6FF72D1B6B32FDAD" blockId="3.[189,1399,539,636]" pageId="3" pageNumber="4">
<keyLead id="395F7171FF9FFFCC6FF72D1B6B32FDAD" pageId="3" pageNumber="4">
3. Distal prong of palea much narrower than proximal prong (e); ventral process of MA subrectangular with long parallel margins (f) ............................................
<taxonomicName id="45AEA9C7FF9FFFCC6CDF2D3E6B50FDAD" ID-CoL="48L73" authority="Jocque M. &amp; Jocque R." authorityName="Jocque M. &amp; Jocque R." authorityYear="2017" box="[917,1301,574,601]" class="Arachnida" family="Lycosidae" genus="Ocyale" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="ghost" status="sp. nov.">
<emphasis id="B0DA0E56FF9FFFCC6CDF2D3E6DBFFDAD" bold="true" box="[917,1018,574,601]" italics="true" pageId="3" pageNumber="4">O. ghost</emphasis>
Jocque M. &amp; Jocqué R.
</taxonomicName>
<taxonomicNameLabel id="ABE9B32DFF9FFFCC6A562D3F6B32FDAD" box="[1308,1399,575,601]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8211D244FF9FFFCC6F8B2D626B32FD88" blockId="3.[189,1399,539,636]" box="[193,1399,609,636]" pageId="3" pageNumber="4">
<keyLead id="395F7171FF9FFFCC6F8B2D626B32FD88" box="[193,1399,609,636]" pageId="3" pageNumber="4">– Prongs of palea of similar size (g); long margin of ventral MA process with bulge (h) .................. 4</keyLead>
</paragraph>
</keyStep>
<keyStep id="395ACAE1FF9FFFCC6FF72B4E6B32FB5B" pageId="3" pageNumber="4">
<paragraph id="8211D244FF9FFFCC6FF72B4E6B32FB7F" blockId="3.[189,1399,1102,1200]" pageId="3" pageNumber="4">
<keyLead id="395F7171FF9FFFCC6FF72B4E6B32FB7F" pageId="3" pageNumber="4">
4. Lateral process of MA slender, tapered to sharp tip (i); TL&gt;
<quantity id="45567FA1FF9FFFCC6CFB2B4E6A46FB9D" box="[945,1027,1102,1129]" metricMagnitude="-2" metricUnit="m" metricValue="2.3" pageId="3" pageNumber="4" unit="mm" value="23.0">23 mm</quantity>
................................................ ...........................................................................................................
<taxonomicName id="45AEA9C7FF9FFFCC6B4E2B716B32FB7F" ID-CoL="48L74" authority="Alderweireldt, 1996" authorityName="Alderweireldt" authorityYear="1996" box="[1028,1399,1137,1164]" class="Arachnida" family="Lycosidae" genus="Ocyale" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="grandis">
<emphasis id="B0DA0E56FF9FFFCC6B4E2B716AC6FB78" bold="true" box="[1028,1155,1137,1164]" italics="true" pageId="3" pageNumber="4">O. grandis</emphasis>
<bibRefCitation id="E63FAFB5FF9FFFCC6BC02B716B32FB7F" author="Alderweireldt M." box="[1162,1399,1137,1163]" pageId="3" pageNumber="4" pagination="103 - 111" refId="ref4420" refString="Alderweireldt M. 1996. A taxonomic revision of the genus Ocyale Audouin, 1826 in Africa (Araneae: Lycosidae). Journal of Natural History 30: 1349 - 1365. https: // doi. org / 10.1080 / 00222939600771261 Alderweireldt M. &amp; Jocque R. 1991. A remarkable new genus of wolf spiders from southwestern Spain (Araneae, Lycosidae). Bulletin van het Koninklijk Belgisch Instituut voor Natuurwetenschappen, Entomologie 61: 103 - 111." type="journal article" year="1996">Alderweireldt, 1996</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8211D244FF9FFFCC6F8B2B956B32FB5B" blockId="3.[189,1399,1102,1200]" box="[193,1399,1172,1200]" pageId="3" pageNumber="4">
<keyLead id="395F7171FF9FFFCC6F8B2B956B32FB5B" box="[193,1399,1172,1200]" pageId="3" pageNumber="4">
– Lateral process of MA, shorter and broader, blunt (j); TL &lt;
<quantity id="45567FA1FF9FFFCC6CE92B956DB3FB5B" box="[931,1014,1173,1199]" metricMagnitude="-2" metricUnit="m" metricValue="2.3" pageId="3" pageNumber="4" unit="mm" value="23.0">23 mm</quantity>
................................................ 5
</keyLead>
</paragraph>
</keyStep>
<keyStep id="395ACAE1FF9FFFCC6FF72BD66B32FAAD" pageId="3" pageNumber="4">
<paragraph id="8211D244FF9FFFCC6FF72BD66B32FAE7" blockId="3.[189,1399,1238,1370]" pageId="3" pageNumber="4">
<keyLead id="395F7171FF9FFFCC6FF72BD66B32FAE7" pageId="3" pageNumber="4">
5. Sperm duct near proximal margin of tegulum, partly hidden by subtegulum (k); long margin of ventral MA process with deep long incision before bulge ...........................
<taxonomicName id="45AEA9C7FF9FFFCC6B102BF96B32FAE7" ID-CoL="48L75" authority="Karsch, 1878" authorityName="Karsch" authorityYear="1878" box="[1114,1399,1273,1300]" class="Arachnida" family="Lycosidae" genus="Ocyale" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="guttata">
<emphasis id="B0DA0E56FF9FFFCC6B102BF96A96FAE0" bold="true" box="[1114,1235,1273,1300]" italics="true" pageId="3" pageNumber="4">O. guttata</emphasis>
Karsch, 1878
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8211D244FF9FFFCC6F8B2A1D6B32FAAD" blockId="3.[189,1399,1238,1370]" pageId="3" pageNumber="4">
<keyLead id="395F7171FF9FFFCC6F8B2A1D6B32FAAD" pageId="3" pageNumber="4">
– Sperm duct further from proximal margin of tegulum, not hidden by subtegulum (l); long margin of ventral MA process with shallow, short incision before bulge (m) ...............
<taxonomicName id="45AEA9C7FF9FFFCC6B2A2A406B32FAAD" ID-CoL="48L7D" authority="Roewer, 1960" authorityName="Roewer" authorityYear="1960" box="[1120,1399,1343,1370]" class="Arachnida" family="Lycosidae" genus="Ocyale" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="pilosa">
<emphasis id="B0DA0E56FF9FFFCC6B2A2A406A89FAAE" bold="true" box="[1120,1228,1343,1370]" italics="true" pageId="3" pageNumber="4">O. pilosa</emphasis>
Roewer, 1960
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="395ACAE1FF9FFFCC6FF72A816B32FA4B" pageId="3" pageNumber="4">
<paragraph id="8211D244FF9FFFCC6FF72A816B32FA68" blockId="3.[189,1399,1409,1471]" box="[189,1399,1409,1436]" pageId="3" pageNumber="4">
<keyLead id="395F7171FF9FFFCC6FF72A816B32FA68" box="[189,1399,1409,1436]" pageId="3" pageNumber="4">
6. Epigyne surrounded but not covered by white setae (n) ....
<taxonomicName id="45AEA9C7FF9FFFCC6CDF2A816B50FA68" ID-CoL="48L73" authority="Jocque M. &amp; Jocque R." authorityName="Jocque M. &amp; Jocque R." authorityYear="2017" box="[917,1301,1409,1436]" class="Arachnida" family="Lycosidae" genus="Ocyale" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="ghost" status="sp. nov.">
<emphasis id="B0DA0E56FF9FFFCC6CDF2A816DBFFA68" bold="true" box="[917,1018,1409,1436]" italics="true" pageId="3" pageNumber="4">O. ghost</emphasis>
Jocque M. &amp; Jocqué R.
</taxonomicName>
<taxonomicNameLabel id="ABE9B32DFF9FFFCC6A562A826B32FA68" box="[1308,1399,1410,1436]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8211D244FF9FFFCC6F8B2AA56B32FA4B" blockId="3.[189,1399,1409,1471]" box="[193,1399,1444,1471]" pageId="3" pageNumber="4">
<keyLead id="395F7171FF9FFFCC6F8B2AA56B32FA4B" box="[193,1399,1444,1471]" pageId="3" pageNumber="4">– Epigyne almost completely covered by white setae (o) ................................................................... 7</keyLead>
</paragraph>
</keyStep>
<keyStep id="395ACAE1FF9FFFCC6FF72AE66B32F9B2" pageId="3" pageNumber="4">
<paragraph id="8211D244FF9FFFCC6FF72AE66B32F9D7" blockId="3.[189,1399,1510,1607]" pageId="3" pageNumber="4">
<keyLead id="395F7171FF9FFFCC6FF72AE66B32F9D7" pageId="3" pageNumber="4">
7. Large species, TL&gt;
<quantity id="45567FA1FF9FFFCC6E902AE66C75F9F4" box="[474,560,1510,1536]" metricMagnitude="-2" metricUnit="m" metricValue="2.3" pageId="3" pageNumber="4" unit="mm" value="23.0">23 mm</quantity>
; epigyne with small and short septum (p)............................................... ...........................................................................................................
<taxonomicName id="45AEA9C7FF9FFFCC6B4E29096B32F9D7" ID-CoL="48L74" authority="Alderweireldt, 1996" authorityName="Alderweireldt" authorityYear="1996" box="[1028,1399,1544,1571]" class="Arachnida" family="Lycosidae" genus="Ocyale" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="grandis">
<emphasis id="B0DA0E56FF9FFFCC6B4E29096AC6F9D7" bold="true" box="[1028,1155,1544,1571]" italics="true" pageId="3" pageNumber="4">O. grandis</emphasis>
<bibRefCitation id="E63FAFB5FF9FFFCC6BC029096B32F9D7" author="Alderweireldt M." box="[1162,1399,1545,1571]" pageId="3" pageNumber="4" pagination="103 - 111" refId="ref4420" refString="Alderweireldt M. 1996. A taxonomic revision of the genus Ocyale Audouin, 1826 in Africa (Araneae: Lycosidae). Journal of Natural History 30: 1349 - 1365. https: // doi. org / 10.1080 / 00222939600771261 Alderweireldt M. &amp; Jocque R. 1991. A remarkable new genus of wolf spiders from southwestern Spain (Araneae, Lycosidae). Bulletin van het Koninklijk Belgisch Instituut voor Natuurwetenschappen, Entomologie 61: 103 - 111." type="journal article" year="1996">Alderweireldt, 1996</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8211D244FF9FFFCC6F8B292D6B32F9B2" blockId="3.[189,1399,1510,1607]" box="[193,1399,1580,1607]" pageId="3" pageNumber="4">
<keyLead id="395F7171FF9FFFCC6F8B292D6B32F9B2" box="[193,1399,1580,1607]" pageId="3" pageNumber="4">
– Smaller species, TL &lt;
<quantity id="45567FA1FF9FFFCC6EB8292C6C02F9B3" box="[498,583,1580,1607]" metricMagnitude="-2" metricUnit="m" metricValue="2.3" pageId="3" pageNumber="4" unit="mm" value="23.0">23 mm</quantity>
; epigyne with well-developed inverted T-shaped septum (q, r) ........ 8
</keyLead>
</paragraph>
</keyStep>
<keyStep id="395ACAE1FF98FFCB6FF72E0B6B32FE64" pageId="4" pageNumber="5">
<paragraph id="8211D244FF98FFCB6FF72E0B6B32FEBD" blockId="4.[189,1399,267,401]" pageId="4" pageNumber="5">
<keyLead id="395F7171FF98FFCB6FF72E0B6B32FEBD" pageId="4" pageNumber="5">
8. Longitudinal part of inverted T longer than transverse part (q); colour pattern uniform .................... .....................................................................................................
<taxonomicName id="45AEA9C7FF98FFCB6C912E2F6B32FEBD" ID-CoL="48L6Y" authority="Alderweireldt" authorityName="Alderweireldt" authorityYear="1996" box="[987,1399,302,329]" class="Arachnida" family="Lycosidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="dewinterae">
<emphasis id="B0DA0E56FF98FFCB6C912E2F6AC6FEBD" bold="true" box="[987,1155,302,329]" italics="true" pageId="4" pageNumber="5">O. dewinterae</emphasis>
<bibRefCitation id="E63FAFB5FF98FFCB6BC02E2F6B32FEBD" author="Alderweireldt M." box="[1162,1399,303,329]" pageId="4" pageNumber="5" pagination="103 - 111" refId="ref4420" refString="Alderweireldt M. 1996. A taxonomic revision of the genus Ocyale Audouin, 1826 in Africa (Araneae: Lycosidae). Journal of Natural History 30: 1349 - 1365. https: // doi. org / 10.1080 / 00222939600771261 Alderweireldt M. &amp; Jocque R. 1991. A remarkable new genus of wolf spiders from southwestern Spain (Araneae, Lycosidae). Bulletin van het Koninklijk Belgisch Instituut voor Natuurwetenschappen, Entomologie 61: 103 - 111." type="journal article" year="1996">Alderweireldt, 1996</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8211D244FF98FFCB6F8B2E536B32FE64" blockId="4.[189,1399,267,401]" pageId="4" pageNumber="5">
<keyLead id="395F7171FF98FFCB6F8B2E536B32FE64" pageId="4" pageNumber="5">– Longitudinal part of inverted T as long as or shorter than transverse part (r); brown to yellow carapace pattern (in ethanol) ............................................................................................................................ 9</keyLead>
</paragraph>
</keyStep>
<keyStep id="395ACAE1FF98FFCB6FF72EB86B32FDC9" pageId="4" pageNumber="5">
<paragraph id="8211D244FF98FFCB6FF72EB86B32FE01" blockId="4.[189,1399,440,573]" pageId="4" pageNumber="5">
<keyLead id="395F7171FF98FFCB6FF72EB86B32FE01" pageId="4" pageNumber="5">
9. Lateral epigyne margins along median septum swollen and shiny; longitudinal part of inverted T slender, longer than transverse part ..............
<taxonomicName id="45AEA9C7FF98FFCB6DB62EDB6A5EFE02" ID-CoL="48L75" authority="Karsch, 1878" authorityName="Karsch" authorityYear="1878" box="[764,1051,475,502]" class="Arachnida" family="Lycosidae" genus="Ocyale" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="guttata">
<emphasis id="B0DA0E56FF98FFCB6DB62EDB6D30FE02" bold="true" box="[764,885,475,502]" italics="true" pageId="4" pageNumber="5">O. guttata</emphasis>
Karsch, 1878
</taxonomicName>
,
<taxonomicName id="45AEA9C7FF98FFCB6B6D2EDB6B32FE01" ID-CoL="48L6Z" authority="Roewer, 1960" authorityName="Roewer" authorityYear="1960" box="[1063,1399,475,502]" class="Arachnida" family="Lycosidae" genus="Ocyale" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="discrepans">
<emphasis id="B0DA0E56FF98FFCB6B6D2EDB6A89FE02" bold="true" box="[1063,1228,475,502]" italics="true" pageId="4" pageNumber="5">O. discrepans</emphasis>
Roewer, 1960
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8211D244FF98FFCB6F8B2D006B32FDC9" blockId="4.[189,1399,440,573]" pageId="4" pageNumber="5">
<keyLead id="395F7171FF98FFCB6F8B2D006B32FDC9" pageId="4" pageNumber="5">
– Lateral epigyne margins along median septum not swollen and shiny; longitudinal part of inverted T broad, only half as long as transverse part (r) ................................................
<taxonomicName id="45AEA9C7FF98FFCB6B2A2D236B32FDC9" ID-CoL="48L7D" authority="Roewer, 1960" authorityName="Roewer" authorityYear="1960" box="[1120,1399,546,573]" class="Arachnida" family="Lycosidae" genus="Ocyale" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="pilosa">
<emphasis id="B0DA0E56FF98FFCB6B2A2D236A89FDC9" bold="true" box="[1120,1228,546,573]" italics="true" pageId="4" pageNumber="5">O. pilosa</emphasis>
Roewer, 1960
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
</treatment>
</document>
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