treatments-xml/data/0A/05/78/0A057876FFAF2B2A35939FCAFA92F871.xml

<document ID-DOI="10.11646/zootaxa.5249.2.6" ID-ISSN="1175-5326" ID-Zenodo-Dep="7687382" ID-ZooBank="051A50BF-8ABF-43EB-AE41-EE9CFA06AFAD" approvalRequired="1" approvalRequired_for_textStreams="1" checkinTime="1677655999792" checkinUser="plazi" docAuthor="Yang, Qiao-Zhi, Geshev, Georgi, Pathomwattananurak, Wuttipon &amp; Zhao, Ming-Zhi" docDate="2023" docId="0A057876FFAF2B2A35939FCAFA92F871" docLanguage="en" docName="zootaxa.5249.2.6.pdf" docOrigin="Zootaxa 5249 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5249.2.6" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Eophileurus varipunctatus Yang, Pathomwattananurak &amp; Zhao 2023, new species" docType="treatment" docVersion="2" lastPageNumber="293" masterDocId="F63C000EFFAE2B2935049D01FFCDFFC8" masterDocTitle="Two new species of the genus Eophileurus Arrow, 1908 (Coleoptera: Scarabaeidae Dynastinae) from the Philippines and Vietnam, with first description of E quadratifovealis Yang &amp; Pathomwattananurak, 2022 female" masterLastPageNumber="300" masterPageNumber="290" pageNumber="291" updateTime="1677656497655" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo>
<mods:title>Two new species of the genus Eophileurus Arrow, 1908 (Coleoptera: Scarabaeidae Dynastinae) from the Philippines and Vietnam, with first description of E quadratifovealis Yang &amp; Pathomwattananurak, 2022 female</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Yang, Qiao-Zhi</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Geshev, Georgi</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Pathomwattananurak, Wuttipon</mods:namePart>
<mods:affiliation>1051 / 38 Wiang, Mueang Dist., Chiang Rai Province, Thailand 57000.</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Zhao, Ming-Zhi</mods:namePart>
<mods:affiliation>College of Plant Protection, South China Agricultural University, Guangzhou, 510642, P. R. China.</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>Zootaxa</mods:title>
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<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-03-01</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>5249</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>2</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>290</mods:start>
<mods:end>300</mods:end>
</mods:extent>
</mods:part>
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<mods:location>
<mods:url>http://dx.doi.org/10.11646/zootaxa.5249.2.6</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.11646/zootaxa.5249.2.6</mods:identifier>
<mods:identifier type="ISSN">1175-5326</mods:identifier>
<mods:identifier type="Zenodo-Dep">7687382</mods:identifier>
<mods:identifier type="ZooBank">051A50BF-8ABF-43EB-AE41-EE9CFA06AFAD</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.7687408" ID-Zenodo-Dep="7687408" LSID="urn:lsid:plazi:treatment:0A057876FFAF2B2A35939FCAFA92F871" httpUri="http://treatment.plazi.org/id/0A057876FFAF2B2A35939FCAFA92F871" lastPageId="3" lastPageNumber="293" pageId="1" pageNumber="291">
<subSubSection box="[151,1099,715,742]" pageId="1" pageNumber="291" type="nomenclature">
<paragraph blockId="1.[151,1099,715,778]" box="[151,1099,715,742]" pageId="1" pageNumber="291">
<heading bold="true" box="[151,1099,715,742]" fontSize="11" level="1" pageId="1" pageNumber="291" reason="1">
<emphasis bold="true" box="[151,1099,715,742]" pageId="1" pageNumber="291">
<taxonomicName authority="Yang, Pathomwattananurak &amp; Zhao" authorityName="Yang, Pathomwattananurak &amp; Zhao" authorityYear="2023" box="[151,942,715,742]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="291" phylum="Arthropoda" rank="species" species="varipunctatus" status="new species">
<emphasis bold="true" box="[151,474,715,742]" italics="true" pageId="1" pageNumber="291">Eophileurus varipunctatus</emphasis>
Yang, Pathomwattananurak &amp; Zhao
</taxonomicName>
, 
<taxonomicNameLabel box="[955,1099,716,742]" pageId="1" pageNumber="291" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[151,402,752,778]" pageId="1" pageNumber="291" type="description">
<paragraph blockId="1.[151,1099,715,778]" box="[151,402,752,778]" pageId="1" pageNumber="291">
(
<figureCitation box="[159,300,752,778]" captionStart="FIGURE 1" captionStartId="2.[151,250,1702,1726]" captionTargetBox="[179,1410,187,1675]" captionTargetId="figure-18@2.[177,1411,179,1676]" captionTargetPageId="2" captionText="FIGURE 1. Habitus of Eophileurus varipunctatus Yang, Pathomwattananurak &amp; Zhao, new species. A–C, holotype; D–F, female; A, D, dorsal view; B, E, ventral view; C, F, oblique view." figureDoi="http://doi.org/10.5281/zenodo.7687384" httpUri="https://zenodo.org/record/7687384/files/figure.png" pageId="1" pageNumber="291">Figs. 1A–1F</figureCitation>
, 
<figureCitation box="[312,395,752,778]" captionStart="FIGURE 3" captionStartId="5.[151,250,1251,1275]" captionTargetBox="[179,1409,187,1219]" captionTargetId="figure-20@5.[151,1435,181,1227]" captionTargetPageId="5" captionText="FIGURE 3. Male genitalia of Eophileurus species. A–B, E. varipunctatus Yang, Pathomwattananurak &amp; Zhao, new species, holotype; C–D, E. iwasei Muramoto, 1995; A, C, frontal view of parameres; B, D, lateral view of aedeagus." figureDoi="http://doi.org/10.5281/zenodo.7687388" httpUri="https://zenodo.org/record/7687388/files/figure.png" pageId="1" pageNumber="291">3A–3B</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection lastPageId="2" lastPageNumber="292" pageId="1" pageNumber="291" type="materials_examined">
<paragraph blockId="1.[151,1437,824,2039]" pageId="1" pageNumber="291">
<emphasis bold="true" box="[151,316,824,850]" pageId="1" pageNumber="291">Type material</emphasis>
(3♁♁). 
<materialsCitation collectingDate="2021-02" collectionCode="CAU" collectorName="Noel Mohagan" country="Philippines" county="The" elevation="1500" location="Mt. Halcon" municipality="Naujuan Municipality" pageId="1" pageNumber="291" specimenCount="1" stateProvince="Mindoro Oriental" typeStatus="holotype">
<emphasis bold="true" box="[407,525,824,850]" pageId="1" pageNumber="291">
<typeStatus box="[407,520,824,850]" pageId="1" pageNumber="291">Holotype</typeStatus>
:
</emphasis>
♁ (
<collectionCode box="[564,628,824,850]" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33074" lsid="urn:lsid:biocol.org:col:33074" name="China Agricultural University" pageId="1" pageNumber="291">CAU</collectionCode>
), “ 
<location LSID="urn:lsid:plazi:treatment:0A057876FFAF2B2A35939FCAFA92F871:87739FBBFFAF2B2837949E39FCDFFC9A" box="[656,786,824,850]" country="Philippines" county="The" municipality="Naujuan Municipality" name="Mt. Halcon" pageId="1" pageNumber="291" stateProvince="Mindoro Oriental">Mt. Halcon</location>
, 
<location LSID="urn:lsid:plazi:treatment:0A057876FFAF2B2A35939FCAFA92F871:87739FBBFFAF2B28361F9E39FC1FFC9B" box="[795,978,824,851]" country="Philippines" county="The" municipality="Naujuan Municipality" name="Paitan Barangay" pageId="1" pageNumber="291" stateProvince="Mindoro Oriental">Paitan Barangay</location>
, 
<collectingMunicipality box="[987,1233,824,850]" pageId="1" pageNumber="291">Naujuan Municipality</collectingMunicipality>
, 
<collectingRegion country="Philippines" name="Mindoro Oriental" pageId="1" pageNumber="291">Oriental Mindoro Prov.</collectingRegion>
, Mindoro Is., 
<collectingCounty box="[366,410,860,886]" pageId="1" pageNumber="291">The</collectingCounty>
<collectingCountry box="[416,545,860,886]" name="Philippines" pageId="1" pageNumber="291">Philippines</collectingCountry>
, alt. 
<quantity box="[596,683,860,887]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="291" unit="m" value="1500.0">
<elevation box="[596,683,860,887]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="291" unit="m" value="1500.0">1500 m</elevation>
</quantity>
, 
<date box="[693,777,860,886]" pageId="1" pageNumber="291" value="2021-02">
<collectingDate box="[693,777,860,886]" pageId="1" pageNumber="291" value="2021-02">II.2021</collectingDate>
</date>
, 
<collectorName box="[788,955,860,886]" pageId="1" pageNumber="291">Noel Mohagan</collectorName>
leg.”
</materialsCitation>
<materialsCitation collectingDate="2021-02" collectionCode="CQZY" collectorName="Noel Mohagan" country="Philippines" county="The" elevation="1500" location="Mt. Halcon" municipality="Naujuan Municipality" pageId="1" pageNumber="291" specimenCount="1" stateProvince="Mindoro Oriental" typeStatus="paratype">
<emphasis bold="true" box="[1021,1150,860,886]" pageId="1" pageNumber="291">
<typeStatus box="[1021,1145,860,886]" pageId="1" pageNumber="291">Paratypes</typeStatus>
:
</emphasis>
2♁♁ (
<collectionCode box="[1226,1308,860,886]" pageId="1" pageNumber="291">CQZY</collectionCode>
), same data as the holotype
</materialsCitation>
.
</paragraph>
<paragraph blockId="1.[151,1437,824,2039]" pageId="1" pageNumber="291">
<emphasis bold="true" box="[199,568,932,958]" pageId="1" pageNumber="291">Additional material examined</emphasis>
(
<specimenCount box="[592,625,932,958]" pageId="1" pageNumber="291" type="female">1♀</specimenCount>
). 
<materialsCitation collectingDate="2019-04" collectionCode="CQZY" collectorName="Noel Mohagan" country="Philippines" county="The" elevation="800" location="Brooke's Point Municipality" municipality="Mt. Mantalingajan" pageId="1" pageNumber="291" specimenCount="1" specimenCount-female="1" stateProvince="Palawan">
<emphasis bold="true" box="[654,856,932,958]" pageId="1" pageNumber="291">
<collectingCounty box="[654,701,932,958]" pageId="1" pageNumber="291">The</collectingCounty>
<collectingCountry box="[713,851,932,958]" name="Philippines" pageId="1" pageNumber="291">Philippines</collectingCountry>
:
</emphasis>
<specimenCount box="[868,903,932,958]" pageId="1" pageNumber="291" type="female">1♀</specimenCount>
(
<collectionCode box="[924,1006,932,958]" pageId="1" pageNumber="291">CQZY</collectionCode>
), 
<collectingMunicipality box="[1030,1247,932,959]" pageId="1" pageNumber="291">Mt. Mantalingajan</collectingMunicipality>
, 
<location LSID="urn:lsid:plazi:treatment:0A057876FFAF2B2A35939FCAFA92F871:87739FBBFFAF2B2831F49EA5FEE4FC2A" country="Philippines" county="The" municipality="Mt. Mantalingajan" name="Brooke's Point Municipality" pageId="1" pageNumber="291" stateProvince="Palawan">Brooke’s Point Municipality</location>
, 
<collectingRegion box="[310,475,968,995]" country="Philippines" name="Palawan" pageId="1" pageNumber="291">Palawan Prov.</collectingRegion>
, 
<collectingRegion box="[488,583,968,994]" country="Philippines" name="Palawan" pageId="1" pageNumber="291">Palawan</collectingRegion>
Is., 
<location LSID="urn:lsid:plazi:treatment:0A057876FFAF2B2A35939FCAFA92F871:87739FBBFFAF2B28377E9EC9FCE2FC2A" box="[634,815,968,994]" country="Philippines" county="The" municipality="Mt. Mantalingajan" name="The Philippines" pageId="1" pageNumber="291" stateProvince="Palawan">The Philippines</location>
, alt. 
<quantity box="[872,947,968,995]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="1" pageNumber="291" unit="m" value="800.0">
<elevation box="[872,947,968,995]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="1" pageNumber="291" unit="m" value="800.0">800 m</elevation>
</quantity>
, 
<date box="[961,1052,968,994]" pageId="1" pageNumber="291" value="2019-04">
<collectingDate box="[961,1052,968,994]" pageId="1" pageNumber="291" value="2019-04">IV.2019</collectingDate>
</date>
, 
<collectorName box="[1066,1235,968,994]" pageId="1" pageNumber="291">Noel Mohagan</collectorName>
leg.; see remarks below
</materialsCitation>
.
</paragraph>
<paragraph blockId="1.[151,1437,824,2039]" pageId="1" pageNumber="291">
<emphasis bold="true" box="[199,308,1040,1066]" pageId="1" pageNumber="291">
<typeStatus box="[199,308,1040,1066]" pageId="1" pageNumber="291">Holotype</typeStatus>
</emphasis>
(male). 
<emphasis bold="true" box="[408,504,1040,1066]" pageId="1" pageNumber="291">General</emphasis>
(
<figureCitation box="[522,670,1040,1066]" captionStart="FIGURE 1" captionStartId="2.[151,250,1702,1726]" captionTargetBox="[179,1410,187,1675]" captionTargetId="figure-18@2.[177,1411,179,1676]" captionTargetPageId="2" captionText="FIGURE 1. Habitus of Eophileurus varipunctatus Yang, Pathomwattananurak &amp; Zhao, new species. A–C, holotype; D–F, female; A, D, dorsal view; B, E, ventral view; C, F, oblique view." figureDoi="http://doi.org/10.5281/zenodo.7687384" httpUri="https://zenodo.org/record/7687384/files/figure.png" pageId="1" pageNumber="291">Figs. 1A–1C</figureCitation>
): Body black, oblong, flattened, surface moderately shiny; ventral surface with dense, short reddish-yellow setae. 
<emphasis bold="true" box="[672,745,1076,1102]" pageId="1" pageNumber="291">Head:</emphasis>
Clypeus with dense, small punctures, denser and rugopunctate on sides, with a short and slightly backward, obtuse horn at center, without any ridge from horn to apex; clypeal apex slightly acute, reflexed. Frons slightly depressed, disk with dense, large punctures. Mandible sinuate on outer margin. 
<emphasis bold="true" box="[247,378,1184,1210]" pageId="1" pageNumber="291">Pronotum:</emphasis>
Surface densely punctate, punctures mostly small, becoming denser on lateral and anterior margins, and rugopunctate on anterior one-fourth of pronotal surface and cluster of dense punctures on each side close to disc, punctures slightly sparser near posterior margin. Anterior region with a small, deep, hexagonal fovea; fovea strongly and densely rugopunctate, margin with 2 weak protuberances on each side posteriorly and close to middle. Lateral margin roundly curved, subparallel at middle; all margins beaded except before fovea, more strongly beaded on sides. Anterior angle protruding, apex slightly rounded, posterior angle obtusely angulate. 
<emphasis bold="true" box="[151,280,1400,1426]" pageId="1" pageNumber="291">Scutellum:</emphasis>
Surface with sparse punctures near base and on disc. Sides concave near base and convex posteriorly. 
<emphasis bold="true" box="[151,237,1436,1462]" pageId="1" pageNumber="291">Elytra:</emphasis>
Surface with dense micropunctures, and dense, round or semicircular, large punctures. Interstice 1 with punctures in 2 irregular rows combined to 1 irregular row close to center, interstice 2 with 1 irregular row. Primary costae A and B indistinct, inner punctate row of primary costa B slightly irregular. Punctures in primary costae C, D, E and interstices 3, 4 distinctly larger and deeper than those in more inner primary costae and interstices. Subapical umbone slightly prominent. Margin of elytron weakly and horizontally dilated from base to posterior two-fifth. 
<emphasis bold="true" box="[268,393,1616,1642]" pageId="1" pageNumber="291">Pygidium:</emphasis>
Slightly convex, with dense, small punctures throughout, very densely rugopunctate near base, punctures slightly sparser on sides and sparser near apex. 
<emphasis bold="true" box="[862,1033,1652,1678]" pageId="1" pageNumber="291">Metasternum:</emphasis>
Punctures dense and large on sides; middle with sparser, slightly smaller punctures; setae on anterior region and each side yellow, short. 
<emphasis bold="true" pageId="1" pageNumber="291">Abdominal ventrites:</emphasis>
Surfaces with sparse, small punctures throughout, punctures becoming slightly denser and larger on each side, some bearing setae; ventrite 6 rugopunctate on anterior region. 
<emphasis bold="true" box="[939,1004,1760,1786]" pageId="1" pageNumber="291">Legs:</emphasis>
Protibia tridentate, protarsus strongly thickened, inner protibial claw strongly enlarged, elongated. Inner metatibial spur long, slightly curved outward, outer metatibial spur shorter, slightly more strongly curved outward. 
<emphasis bold="true" box="[945,1074,1832,1858]" pageId="1" pageNumber="291">Parameres</emphasis>
(
<figureCitation box="[1091,1239,1832,1858]" captionStart="FIGURE 3" captionStartId="5.[151,250,1251,1275]" captionTargetBox="[179,1409,187,1219]" captionTargetId="figure-20@5.[151,1435,181,1227]" captionTargetPageId="5" captionText="FIGURE 3. Male genitalia of Eophileurus species. A–B, E. varipunctatus Yang, Pathomwattananurak &amp; Zhao, new species, holotype; C–D, E. iwasei Muramoto, 1995; A, C, frontal view of parameres; B, D, lateral view of aedeagus." figureDoi="http://doi.org/10.5281/zenodo.7687388" httpUri="https://zenodo.org/record/7687388/files/figure.png" pageId="1" pageNumber="291">Figs. 3A–3B</figureCitation>
): In frontal view (
<figureCitation box="[159,251,1868,1894]" captionStart="FIGURE 3" captionStartId="5.[151,250,1251,1275]" captionTargetBox="[179,1409,187,1219]" captionTargetId="figure-20@5.[151,1435,181,1227]" captionTargetPageId="5" captionText="FIGURE 3. Male genitalia of Eophileurus species. A–B, E. varipunctatus Yang, Pathomwattananurak &amp; Zhao, new species, holotype; C–D, E. iwasei Muramoto, 1995; A, C, frontal view of parameres; B, D, lateral view of aedeagus." figureDoi="http://doi.org/10.5281/zenodo.7687388" httpUri="https://zenodo.org/record/7687388/files/figure.png" pageId="1" pageNumber="291">Fig. 3A</figureCitation>
), slender and symmetric; lateral margin strongly convex at middle, and strongly concave on basal onefourth, with a pair of large, acute, slightly backward processes subapically; inner margin with dense and small teeth from posterior one-third to near apex, more than one layer close to apex; apex rounded; in lateral view (
<figureCitation captionStart="FIGURE 3" captionStartId="5.[151,250,1251,1275]" captionTargetBox="[179,1409,187,1219]" captionTargetId="figure-20@5.[151,1435,181,1227]" captionTargetPageId="5" captionText="FIGURE 3. Male genitalia of Eophileurus species. A–B, E. varipunctatus Yang, Pathomwattananurak &amp; Zhao, new species, holotype; C–D, E. iwasei Muramoto, 1995; A, C, frontal view of parameres; B, D, lateral view of aedeagus." figureDoi="http://doi.org/10.5281/zenodo.7687388" httpUri="https://zenodo.org/record/7687388/files/figure.png" pageId="1" pageNumber="291">Fig. 3B</figureCitation>
) apex curved downward, with a slightly backwards process near apex, with a backwardly curved ridge close to center, upper margin sinuate, rather strongly curved downward near apical one-third.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7687384" ID-Zenodo-Dep="7687384" httpUri="https://zenodo.org/record/7687384/files/figure.png" pageId="2" pageNumber="292" startId="2.[151,250,1702,1726]" targetBox="[179,1410,187,1675]" targetPageId="2">
<paragraph blockId="2.[151,1436,1702,1763]" pageId="2" pageNumber="292">
<emphasis bold="true" box="[151,278,1702,1726]" pageId="2" pageNumber="292">FIGURE 1.</emphasis>
Habitus of 
<taxonomicName authority="Yang, Pathomwattananurak &amp; Zhao" authorityName="Yang, Pathomwattananurak &amp; Zhao" authorityYear="2023" box="[406,1063,1702,1727]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="292" phylum="Arthropoda" rank="species" species="varipunctatus" status="new species">
<emphasis box="[406,678,1702,1726]" italics="true" pageId="2" pageNumber="292">Eophileurus varipunctatus</emphasis>
Yang, Pathomwattananurak &amp; Zhao
</taxonomicName>
, 
<emphasis bold="true" box="[1076,1204,1702,1726]" pageId="2" pageNumber="292">
<taxonomicNameLabel box="[1076,1204,1702,1726]" pageId="2" pageNumber="292" rank="species">new species</taxonomicNameLabel>
</emphasis>
. A–C, holotype; D–F, female; A, D, dorsal view; B, E, ventral view; C, F, oblique view.
</paragraph>
</caption>
<paragraph blockId="2.[151,1436,1808,2015]" box="[199,1281,1808,1835]" pageId="2" pageNumber="292">
<emphasis bold="true" box="[199,319,1809,1835]" pageId="2" pageNumber="292">
<typeStatus box="[199,319,1809,1835]" pageId="2" pageNumber="292">Paratypes</typeStatus>
</emphasis>
(male). Characters mainly stable, except protuberances at the margin of fovea absent.
</paragraph>
<paragraph blockId="2.[151,1436,1808,2015]" pageId="2" pageNumber="292">
<emphasis bold="true" box="[199,293,1845,1871]" pageId="2" pageNumber="292">Female.</emphasis>
Similar to male, except clypeus and frons much more densely rugopunctate throughout; clypeal horn much shorter, reduced to a tubercle; fovea absent, with an extremely shallow and small depression on anterior region of pronotum; pronotum more weakly rugopunctate anteriorly; anterior margin of pronotum entirely beaded; scutellum without inward curves at side near base; protarsus not thickened, inner protarsal claw not enlarged and elongated (
<figureCitation box="[275,420,1988,2015]" captionStart="FIGURE 1" captionStartId="2.[151,250,1702,1726]" captionTargetBox="[179,1410,187,1675]" captionTargetId="figure-18@2.[177,1411,179,1676]" captionTargetPageId="2" captionText="FIGURE 1. Habitus of Eophileurus varipunctatus Yang, Pathomwattananurak &amp; Zhao, new species. A–C, holotype; D–F, female; A, D, dorsal view; B, E, ventral view; C, F, oblique view." figureDoi="http://doi.org/10.5281/zenodo.7687384" httpUri="https://zenodo.org/record/7687384/files/figure.png" pageId="2" pageNumber="292">Figs. 1D–1F</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="293" type="description">
<paragraph blockId="3.[151,1437,150,1977]" pageId="3" pageNumber="293">
<emphasis bold="true" box="[199,382,151,177]" pageId="3" pageNumber="293">Measurements.</emphasis>
Length: male 
<quantity box="[541,703,150,177]" metricMagnitude="-2" metricUnit="m" metricValue="2.55" metricValueMax="2.58" metricValueMin="2.52" pageId="3" pageNumber="293" unit="mm" value="25.5" valueMax="25.8" valueMin="25.2">25.2–25.8 mm</quantity>
(
<typeStatus box="[716,816,151,177]" pageId="3" pageNumber="293">holotype</typeStatus>
<quantity box="[822,925,150,177]" metricMagnitude="-2" metricUnit="m" metricValue="2.57" pageId="3" pageNumber="293" unit="mm" value="25.7">25.7 mm</quantity>
), female 
<quantity box="[1025,1126,151,177]" metricMagnitude="-2" metricUnit="m" metricValue="2.67" pageId="3" pageNumber="293" unit="mm" value="26.7">26.7 mm</quantity>
; width: male 
<quantity box="[1275,1437,151,177]" metricMagnitude="-2" metricUnit="m" metricValue="1.305" metricValueMax="1.34" metricValueMin="1.27" pageId="3" pageNumber="293" unit="mm" value="13.05" valueMax="13.4" valueMin="12.7">12.7–13.4 mm</quantity>
(
<typeStatus box="[159,259,187,213]" pageId="3" pageNumber="293">holotype</typeStatus>
<quantity box="[266,370,187,213]" metricMagnitude="-2" metricUnit="m" metricValue="1.34" pageId="3" pageNumber="293" unit="mm" value="13.4">13.4 mm</quantity>
), female 
<quantity box="[472,575,187,213]" metricMagnitude="-2" metricUnit="m" metricValue="1.3699999999999999" pageId="3" pageNumber="293" unit="mm" value="13.7">13.7 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="293" type="diagnosis">
<paragraph blockId="3.[151,1437,150,1977]" pageId="3" pageNumber="293">
<emphasis bold="true" box="[199,322,222,248]" pageId="3" pageNumber="293">Diagnosis.</emphasis>
The new species is most closely related to 
<taxonomicName authority="Muramoto, 1995" authorityName="Muramoto" authorityYear="1995" box="[809,1220,222,249]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="293" phylum="Arthropoda" rank="species" species="iwasei">
<emphasis box="[809,1025,223,249]" italics="true" pageId="3" pageNumber="293">Eophileurus iwasei</emphasis>
<bibRefCitation author="Muramoto, R." box="[1032,1220,222,249]" pageId="3" pageNumber="293" pagination="223 - 226" refId="ref5515" refString="Muramoto, R. (1995) Two new species of the genus Eophileurus Arrow (Coleoptera, Scarabaeidae, Dynastinae) from Southeast Asia. Japanese Journal of Systematic Entomology, 1, 223 - 226." type="journal article" year="1995">Muramoto, 1995</bibRefCitation>
</taxonomicName>
(
<figureCitation captionStart="FIGURE 2" captionStartId="4.[151,250,1884,1908]" captionTargetBox="[158,1432,214,1854]" captionTargetId="figure-18@4.[151,1435,201,1861]" captionTargetPageId="4" captionText="FIGURE 2. Habitus of Eophileurus iwasei Muramoto, 1995. A–C, male (Mindanao Island); D–F, female (Leyte Island); A, D, dorsal view; B, E, ventral view; C, F, oblique view." figureDoi="http://doi.org/10.5281/zenodo.7687386" httpUri="https://zenodo.org/record/7687386/files/figure.png" pageId="3" pageNumber="293">Figs. 2A–2F, 2C– 2D</figureCitation>
) due to the parameres of similar shape, with a series of small teeth in the inner margins, and the similar external characters. We have only examined an underdeveloped male of the latter species. Although the original description of 
<taxonomicName authorityName="Muramoto" authorityYear="1995" box="[185,291,331,356]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="293" phylum="Arthropoda" rank="species" species="iwasei">
<emphasis box="[185,291,331,356]" italics="true" pageId="3" pageNumber="293">E. iwasei</emphasis>
</taxonomicName>
was based on a more developed male (body length 
<quantity box="[900,1006,331,357]" metricMagnitude="-2" metricUnit="m" metricValue="2.28" pageId="3" pageNumber="293" unit="mm" value="22.8">22.8 mm</quantity>
; 
<bibRefCitation author="Muramoto, R." box="[1021,1208,330,357]" pageId="3" pageNumber="293" pagination="223 - 226" refId="ref5515" refString="Muramoto, R. (1995) Two new species of the genus Eophileurus Arrow (Coleoptera, Scarabaeidae, Dynastinae) from Southeast Asia. Japanese Journal of Systematic Entomology, 1, 223 - 226." type="journal article" year="1995">Muramoto 1995</bibRefCitation>
), we remain unsure whether it is a well-developed individual. And the three examined male specimens of the new species are basically of the same size, which makes development unclear. However, the following characters allow an easy separation of the two species (those of 
<taxonomicName authorityName="Muramoto" authorityYear="1995" box="[435,538,439,464]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="293" phylum="Arthropoda" rank="species" species="iwasei">
<emphasis box="[435,538,439,464]" italics="true" pageId="3" pageNumber="293">E. iwasei</emphasis>
</taxonomicName>
are in parenthesis):
</paragraph>
<paragraph blockId="3.[151,1437,150,1977]" pageId="3" pageNumber="293">
Body distinctly wider (more slender in 
<taxonomicName authorityName="Muramoto" authorityYear="1995" box="[653,758,475,500]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="293" phylum="Arthropoda" rank="species" species="iwasei">
<emphasis box="[653,758,475,500]" italics="true" pageId="3" pageNumber="293">E. iwasei</emphasis>
</taxonomicName>
); scutellum with an inward curvature close to base at each side in male (absent in 
<taxonomicName authorityName="Muramoto" authorityYear="1995" box="[415,519,511,536]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="293" phylum="Arthropoda" rank="species" species="iwasei">
<emphasis box="[415,519,511,536]" italics="true" pageId="3" pageNumber="293">E. iwasei</emphasis>
</taxonomicName>
); elytral punctures in primary costae C, D, E and interstices 3, 4 distinctly larger and deeper than those in inner primary costae and interstices (all elytral punctures regularly and universally sized in 
<taxonomicName authorityName="Muramoto" authorityYear="1995" box="[182,287,583,608]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="293" phylum="Arthropoda" rank="species" species="iwasei">
<emphasis box="[182,287,583,608]" italics="true" pageId="3" pageNumber="293">E. iwasei</emphasis>
</taxonomicName>
); elytral punctures much larger than micropunctures (differences of puncture sizes less distinct in 
<taxonomicName authorityName="Muramoto" authorityYear="1995" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="295" phylum="Arthropoda" rank="species" species="iwasei">
<emphasis italics="true" pageId="3" pageNumber="293">
<emphasis bold="true" box="[1412,1436,583,609]" italics="true" pageId="3" pageNumber="293">E.</emphasis>
iwasei
</emphasis>
</taxonomicName>
); pygidium with sparser and slightly smaller punctures; mesosternum with denser punctures at each side, punctures slightly smaller close to center; apex of paramere semicircular, enlarged, with a sharp, short, outward and backward process at external side (apex of paramere flat, weakly enlarged, with a pair of sharp, rather short processes at external side in 
<taxonomicName authorityName="Muramoto" authorityYear="1995" box="[489,596,727,752]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="293" phylum="Arthropoda" rank="species" species="iwasei">
<emphasis box="[489,596,727,752]" italics="true" pageId="3" pageNumber="293">E. iwasei</emphasis>
</taxonomicName>
); female pronotum only with a very small and shallow fovea anteriorly (female pronotum with a rather big and deep sub-triangular fovea at anterior region and a shallow furrow behind it extending to base).
</paragraph>
<paragraph blockId="3.[151,1437,150,1977]" pageId="3" pageNumber="293">
The parameres of the new species also resembles 
<taxonomicName authority="Yamaya &amp; Muramoto, 2008" authorityName="Yamaya &amp; Muramoto" authorityYear="2008" box="[778,1295,835,861]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="293" phylum="Arthropoda" rank="species" species="niii">
<emphasis box="[778,962,835,861]" italics="true" pageId="3" pageNumber="293">Eophileurus niii</emphasis>
Yamaya &amp; Muramoto, 2008
</taxonomicName>
(
<bibRefCitation author="Yamaya, S. &amp; Muramoto, R." pageId="3" pageNumber="293" pagination="95 - 97" refId="ref5733" refString="Yamaya, S. &amp; Muramoto, R. (2008 b) A new species of the genus Eophileurus Arrow, 1908 from Java, Indonesia. Kogane, 9, 95 - 97." type="journal article" year="2008">Yamaya &amp; Muramoto, 2008b</bibRefCitation>
) since both are slender and have similar apices with small, backward process at each side. 
<taxonomicName authorityName="Yamaya &amp; Muramoto" authorityYear="2008" box="[1368,1436,871,896]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="293" phylum="Arthropoda" rank="species" species="niii">
<emphasis box="[1368,1436,871,896]" italics="true" pageId="3" pageNumber="293">E. niii</emphasis>
</taxonomicName>
is, however, not considered closely related to the new species due to the base of its parameres slightly overlapping and the different size of the teeth on the inner margin as well as the different external characters.
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="293" type="etymology">
<paragraph blockId="3.[151,1437,150,1977]" pageId="3" pageNumber="293">
<emphasis bold="true" box="[199,334,979,1005]" pageId="3" pageNumber="293">Etymology.</emphasis>
The specific epithet is a combination of the Latin prefix “ 
<emphasis box="[1021,1066,979,1004]" italics="true" pageId="3" pageNumber="293">vari</emphasis>
” and the adjective “ 
<emphasis box="[1308,1417,979,1004]" italics="true" pageId="3" pageNumber="293">punctatus</emphasis>
”, referring to the peculiar arrangement of elytral punctures, which distinctly varies in sizes on different striae.
</paragraph>
</subSubSection>
<subSubSection box="[199,927,1050,1077]" pageId="3" pageNumber="293" type="distribution">
<paragraph blockId="3.[151,1437,150,1977]" box="[199,927,1050,1077]" pageId="3" pageNumber="293">
<emphasis bold="true" box="[199,353,1050,1076]" pageId="3" pageNumber="293">Distribution.</emphasis>
The 
<collectingCountry box="[410,536,1051,1077]" name="Philippines" pageId="3" pageNumber="293">Philippines</collectingCountry>
(Mindoro Island, 
<collectingRegion box="[740,835,1051,1077]" country="Philippines" name="Palawan" pageId="3" pageNumber="293">Palawan</collectingRegion>
Island).
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="293" type="discussion">
<paragraph blockId="3.[151,1437,150,1977]" pageId="3" pageNumber="293">
<emphasis bold="true" box="[199,315,1087,1113]" pageId="3" pageNumber="293">Remarks.</emphasis>
All the specimens of the new species were purchased from the local collector N. Mohagan from Mindoro, the 
<collectingCountry box="[306,436,1123,1149]" name="Philippines" pageId="3" pageNumber="293">Philippines</collectingCountry>
. Although geographically close, many groups widespread through the Philippine Modile Belt (PMB, including Mindoro Island) are absent from 
<collectingRegion box="[768,863,1159,1185]" country="Philippines" name="Palawan" pageId="3" pageNumber="293">Palawan</collectingRegion>
Island (
<bibRefCitation author="Heads, M." box="[952,1092,1159,1185]" pageId="3" pageNumber="293" pagination="356 - 401" refId="ref5283" refString="Heads, M. (2013) Biogeography of the Philippines. In: Heads, M. (Ed.), Biogeography of Australasia: A Molecular Analysis. Cambridge University Press, Cambridge, pp. 356 - 401. https: // doi. org / 10.1017 / CBO 9781139644464.011" type="book chapter" year="2013">Heads 2013</bibRefCitation>
). It is rare to see shared taxa of pleurostict scarabs between the two regions, e.g., 
<taxonomicName authority="(Moser, 1915)" baseAuthorityName="Moser" baseAuthorityYear="1915" box="[709,1194,1194,1221]" class="Insecta" family="Melolonthidae" genus="Carlschoenherria" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="293" phylum="Arthropoda" rank="species" species="palawana">
<emphasis box="[709,1025,1195,1221]" italics="true" pageId="3" pageNumber="293">Carlschoenherria palawana</emphasis>
(Moser, 1915)
</taxonomicName>
, 
<taxonomicName authority="(Keith, 2006)" baseAuthorityName="Keith" baseAuthorityYear="2006" class="Insecta" family="Scarabaeidae" genus="Philgertia" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="293" phylum="Arthropoda" rank="species" species="lii">
<emphasis box="[1206,1350,1195,1221]" italics="true" pageId="3" pageNumber="293">Philgertia lii</emphasis>
(
<bibRefCitation author="Keith, D." pageId="3" pageNumber="293" pagination="90 - 94" refId="ref5412" refString="Keith, D. (2006) Reflexions sur le gener Engertia et description d'une nouvelle espece des Philippines (Col. Scarabaeoidea Melolonthidae). Lambillionea, 106 (1), 90 - 94." type="journal article" year="2006">Keith, 2006</bibRefCitation>
)
</taxonomicName>
and 
<taxonomicName authority="(Guerin-Meneville, 1843)" baseAuthorityName="Guerin-Meneville" baseAuthorityYear="1843" box="[277,974,1231,1257]" class="Insecta" family="Rutelidae" genus="Parastasia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="293" phylum="Arthropoda" rank="subSpecies" species="bimaculata" subSpecies="bimaculata">
<emphasis box="[277,671,1231,1257]" italics="true" pageId="3" pageNumber="293">Parastasia bimaculata bimaculata</emphasis>
(Guérin-Méneville, 1843)
</taxonomicName>
(
<bibRefCitation author="Itoh, T." box="[992,1108,1231,1257]" pageId="3" pageNumber="293" pagination="3 - 20" refId="ref5337" refString="Itoh, T. (1993) A taxonomical study on the genus Schoenherria Burmeister (Melolonthinae: Melolonthidae) II. Lamellicornia, 9, 3 - 20." type="journal article" year="1993">Itoh 1993</bibRefCitation>
; 
<bibRefCitation author="Keith, D." box="[1123,1256,1231,1257]" pageId="3" pageNumber="293" pagination="90 - 94" refId="ref5412" refString="Keith, D. (2006) Reflexions sur le gener Engertia et description d'une nouvelle espece des Philippines (Col. Scarabaeoidea Melolonthidae). Lambillionea, 106 (1), 90 - 94." type="journal article" year="2006">Keith 2006</bibRefCitation>
; 
<bibRefCitation author="Kuijten, P." box="[1270,1424,1231,1257]" pageId="3" pageNumber="293" pagination="3 - 176" refId="ref5449" refString="Kuijten, P. (1992) A revision of the genus Parastasia in the Indo-Australian region (Coleoptera, Scarabaeidae, Rutelinae). Zoologische Verhandelingen, 275, 3 - 176." type="journal article" year="1992">Kuijten 1992</bibRefCitation>
). Since the distribution of 
<taxonomicName authorityName="Muramoto" authorityYear="1995" box="[442,662,1267,1293]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="293" phylum="Arthropoda" rank="species" species="iwasei">
<emphasis box="[442,662,1267,1293]" italics="true" pageId="3" pageNumber="293">Eophileurus iwasei</emphasis>
</taxonomicName>
covers a large majority of PMB (
<bibRefCitation author="Yamaya, S. &amp; Muramoto, R." box="[1061,1398,1267,1293]" pageId="3" pageNumber="293" pagination="1 - 34" refId="ref5690" refString="Yamaya, S. &amp; Muramoto, R. (2008 a) On the genus Eophileurus Arrow, 1908 (Scarabaeidae, Dynastinae), with descriptions of five new species. Kogane, Supplementum, 2, 1 - 34." type="journal article" year="2008">Yamaya &amp; Muramoto 2008a</bibRefCitation>
), it is surprising to find a similar species in both PMB and 
<collectingRegion box="[786,881,1303,1329]" country="Philippines" name="Palawan" pageId="3" pageNumber="293">Palawan</collectingRegion>
regions. The female from 
<collectingRegion box="[1187,1282,1303,1329]" country="Philippines" name="Palawan" pageId="3" pageNumber="293">Palawan</collectingRegion>
is tentatively assigned to the new species due to its external affinity to the male specimens, especially from the characteristically enlarged elytral punctures in primary costae C, D, E and interstices 3, 4. Given that females between closely related 
<taxonomicName box="[151,374,1411,1437]" pageId="3" pageNumber="293">
<emphasis box="[151,288,1411,1437]" italics="true" pageId="3" pageNumber="293">Eophileurus</emphasis>
species
</taxonomicName>
are often indistinguishable from each other externally and neither associated male from 
<collectingRegion box="[1341,1436,1411,1437]" country="Philippines" name="Palawan" pageId="3" pageNumber="293">Palawan</collectingRegion>
nor female from Mindoro are available, we therefore do not treat the female as a 
<typeStatus box="[1061,1161,1447,1473]" pageId="3" pageNumber="293">paratype</typeStatus>
. We also cannot exclude the possibility that these specimens were partly mislabeled by the dealer.
</paragraph>
<paragraph blockId="3.[151,1437,150,1977]" pageId="3" pageNumber="293">
A similar case of such distribution pattern occurs in Java Island, i.e., 
<taxonomicName authority="Prell, 1913" authorityName="Prell" authorityYear="1913" box="[969,1217,1519,1545]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="293" phylum="Arthropoda" rank="species" species="javanus">
<emphasis box="[969,1087,1519,1544]" italics="true" pageId="3" pageNumber="293">E. javanus</emphasis>
<bibRefCitation author="Prell, H." box="[1094,1217,1519,1545]" pageId="3" pageNumber="293" pagination="103 - 124" refId="ref5630" refString="Prell, H. (1913) Beitrage zur Kenntnis der Dynastinen. VII. Ueber das Genus Eophilurus Arrow. Memoires de la Societe Entomologique de Belgique, 22, 103 - 124." type="journal article" year="1913">Prell, 1913</bibRefCitation>
</taxonomicName>
and 
<taxonomicName authority="Yamaya &amp; Muramoto, 2008" authorityName="Yamaya &amp; Muramoto" authorityYear="2008" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="293" phylum="Arthropoda" rank="species" species="niii">
<emphasis box="[1272,1340,1519,1544]" italics="true" pageId="3" pageNumber="293">E. niii</emphasis>
Yamaya &amp; Muramoto, 2008
</taxonomicName>
(
<bibRefCitation author="Yamaya, S. &amp; Muramoto, R." box="[384,715,1555,1581]" pageId="3" pageNumber="293" pagination="95 - 97" refId="ref5733" refString="Yamaya, S. &amp; Muramoto, R. (2008 b) A new species of the genus Eophileurus Arrow, 1908 from Java, Indonesia. Kogane, 9, 95 - 97." type="journal article" year="2008">Yamaya &amp; Muramoto 2008b</bibRefCitation>
). However, they may not form a monophyletic clade since there are several species morphologically more similar to 
<taxonomicName authorityName="Prell" authorityYear="1913" box="[740,858,1591,1616]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="293" phylum="Arthropoda" rank="species" species="javanus">
<emphasis box="[740,858,1591,1616]" italics="true" pageId="3" pageNumber="293">E. javanus</emphasis>
</taxonomicName>
in nearby island groups. Another hypothesis is that 
<taxonomicName authorityName="Yang, Pathomwattananurak &amp; Zhao" authorityYear="2023" box="[151,340,1627,1652]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="293" phylum="Arthropoda" rank="species" species="varipunctatus">
<emphasis box="[151,340,1627,1652]" italics="true" pageId="3" pageNumber="293">E. varipunctatus</emphasis>
</taxonomicName>
and 
<taxonomicName authorityName="Muramoto" authorityYear="1995" box="[401,508,1627,1652]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="293" phylum="Arthropoda" rank="species" species="iwasei">
<emphasis box="[401,508,1627,1652]" italics="true" pageId="3" pageNumber="293">E. iwasei</emphasis>
</taxonomicName>
are not vicariant siblings, their ancestors dispersed into the 
<collectingCountry box="[1213,1339,1627,1653]" name="Philippines" pageId="3" pageNumber="293">Philippines</collectingCountry>
through different paths, e.g., by crossing 
<collectingRegion box="[517,612,1663,1689]" country="Philippines" name="Palawan" pageId="3" pageNumber="293">Palawan</collectingRegion>
or 
<collectingRegion box="[649,704,1663,1689]" country="Philippines" name="Sulu" pageId="3" pageNumber="293">Sulu</collectingRegion>
.
</paragraph>
<paragraph blockId="3.[151,1437,150,1977]" pageId="3" pageNumber="293">
Apart from the two Philippine species mentioned above, there is also a doubtful record of 
<taxonomicName authority="(Faldermann, 1835)" baseAuthorityName="Faldermann" baseAuthorityYear="1835" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="293" phylum="Arthropoda" rank="species" species="chinensis">
<emphasis box="[1295,1436,1699,1725]" italics="true" pageId="3" pageNumber="293">E. chinensis</emphasis>
(
<bibRefCitation author="Faldermann, F." box="[159,367,1734,1761]" pageId="3" pageNumber="293" pagination="337 - 464" refId="ref5230" refString="Faldermann, F. (1835) Coleopterorum ab illustrissimo Bungio in China boreali, Mongolia et montibus altaicis collectorum nec non ab ill. Turchaninoffio et Stschukinoe e provincia Irkutsk missorum illustrations. Memoires de l'Academie imperiale des Sciences de Saint Petersbourg, 6 (2), 337 - 464." type="journal article" year="1835">Faldermann, 1835</bibRefCitation>
)
</taxonomicName>
from “Agusan River, Mindanao Is.” (
<bibRefCitation author="Schultze, W." box="[810,976,1735,1761]" pageId="3" pageNumber="293" pagination="154 - 180" refId="ref5665" refString="Schultze, W. (1916) A catalogue of Philippine Coleoptera. The Philippine Journal of Science, 11, 154 - 180." type="journal article" year="1916">Schultze 1916</bibRefCitation>
; 
<bibRefCitation author="Muramoto, R." box="[989,1176,1734,1761]" pageId="3" pageNumber="293" pagination="223 - 226" refId="ref5515" refString="Muramoto, R. (1995) Two new species of the genus Eophileurus Arrow (Coleoptera, Scarabaeidae, Dynastinae) from Southeast Asia. Japanese Journal of Systematic Entomology, 1, 223 - 226." type="journal article" year="1995">Muramoto 1995</bibRefCitation>
). This record might be based on a misidentified specimen of 
<taxonomicName authorityName="Muramoto" authorityYear="1995" box="[568,670,1771,1796]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="293" phylum="Arthropoda" rank="species" species="iwasei">
<emphasis box="[568,670,1771,1796]" italics="true" pageId="3" pageNumber="293">E. iwasei</emphasis>
</taxonomicName>
. However, the exact identity of the specimen might not be confirmed due to the destruction of Bureau of Science of Manilla during World War II, which housed the specimen cited by 
<bibRefCitation author="Schultze, W." box="[151,330,1843,1869]" pageId="3" pageNumber="293" pagination="154 - 180" refId="ref5665" refString="Schultze, W. (1916) A catalogue of Philippine Coleoptera. The Philippine Journal of Science, 11, 154 - 180." type="journal article" year="1916">Schultze (1916)</bibRefCitation>
(but some material originally deposited in Bureau of Science which belongs to Baker’s collection was moved to the 
<collectingCountry box="[359,511,1879,1905]" name="United States of America" pageId="3" pageNumber="293">United States</collectingCountry>
before its destruction). Although 
<taxonomicName baseAuthorityName="Faldermann" baseAuthorityYear="1835" box="[894,1030,1879,1905]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="293" phylum="Arthropoda" rank="species" species="chinensis">
<emphasis box="[894,1030,1879,1905]" italics="true" pageId="3" pageNumber="293">E. chinensis</emphasis>
</taxonomicName>
can be found on 
<collectingCountry box="[1229,1387,1879,1905]" name="Taiwan Island" pageId="3" pageNumber="293">Taiwan Island</collectingCountry>
and small islands of Okinawa, there are no records in the Taiwan-Luzon Volcanic Arc, a path for the interchange between 
<collectingCountry box="[151,308,1951,1977]" name="Taiwan Island" pageId="3" pageNumber="293">Taiwan Island</collectingCountry>
and the 
<collectingCountry box="[404,534,1951,1977]" name="Philippines" pageId="3" pageNumber="293">Philippines</collectingCountry>
. Thus, the distribution of 
<taxonomicName baseAuthorityName="Faldermann" baseAuthorityYear="1835" box="[821,957,1951,1977]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="293" phylum="Arthropoda" rank="species" species="chinensis">
<emphasis box="[821,957,1951,1977]" italics="true" pageId="3" pageNumber="293">E. chinensis</emphasis>
</taxonomicName>
in the 
<collectingCountry box="[1034,1160,1951,1977]" name="Philippines" pageId="3" pageNumber="293">Philippines</collectingCountry>
should be omitted.
</paragraph>
</subSubSection>
</treatment>
</document>