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<document ID-DOI="10.11646/zootaxa.5249.2.6" ID-ISSN="1175-5326" ID-Zenodo-Dep="7687382" ID-ZooBank="051A50BF-8ABF-43EB-AE41-EE9CFA06AFAD" approvalRequired="1" approvalRequired_for_textStreams="1" checkinTime="1677655999792" checkinUser="plazi" docAuthor="Yang, Qiao-Zhi, Geshev, Georgi, Pathomwattananurak, Wuttipon &amp; Zhao, Ming-Zhi" docDate="2023" docId="0A057876FFAB2B2135939870FE03FE3D" docLanguage="en" docName="zootaxa.5249.2.6.pdf" docOrigin="Zootaxa 5249 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5249.2.6" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Eophileurus vietnamensis Yang, Pathomwattananurak &amp; Zhao 2023, new species" docType="treatment" docVersion="2" lastPageNumber="298" masterDocId="F63C000EFFAE2B2935049D01FFCDFFC8" masterDocTitle="Two new species of the genus Eophileurus Arrow, 1908 (Coleoptera: Scarabaeidae Dynastinae) from the Philippines and Vietnam, with first description of E quadratifovealis Yang &amp; Pathomwattananurak, 2022 female" masterLastPageNumber="300" masterPageNumber="290" pageNumber="295" updateTime="1677656497655" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Two new species of the genus Eophileurus Arrow, 1908 (Coleoptera: Scarabaeidae Dynastinae) from the Philippines and Vietnam, with first description of E quadratifovealis Yang &amp; Pathomwattananurak, 2022 female</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Yang, Qiao-Zhi</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Geshev, Georgi</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Pathomwattananurak, Wuttipon</mods:namePart>
<mods:affiliation>1051 / 38 Wiang, Mueang Dist., Chiang Rai Province, Thailand 57000.</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Zhao, Ming-Zhi</mods:namePart>
<mods:affiliation>College of Plant Protection, South China Agricultural University, Guangzhou, 510642, P. R. China.</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Zootaxa</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-03-01</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>5249</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>2</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>290</mods:start>
<mods:end>300</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.11646/zootaxa.5249.2.6</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.11646/zootaxa.5249.2.6</mods:identifier>
<mods:identifier type="ISSN">1175-5326</mods:identifier>
<mods:identifier type="Zenodo-Dep">7687382</mods:identifier>
<mods:identifier type="ZooBank">051A50BF-8ABF-43EB-AE41-EE9CFA06AFAD</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:0A057876FFAB2B2135939870FE03FE3D" httpUri="http://treatment.plazi.org/id/0A057876FFAB2B2135939870FE03FE3D" lastPageId="8" lastPageNumber="298" pageId="5" pageNumber="295">
<subSubSection box="[151,1089,1393,1420]" pageId="5" pageNumber="295" type="nomenclature">
<paragraph blockId="5.[151,1089,1393,1456]" box="[151,1089,1393,1420]" pageId="5" pageNumber="295">
<heading bold="true" box="[151,1089,1393,1420]" fontSize="11" level="1" pageId="5" pageNumber="295" reason="1">
<emphasis bold="true" box="[151,1089,1393,1420]" pageId="5" pageNumber="295">
<taxonomicName authority="Yang, Pathomwattananurak &amp; Zhao" authorityName="Yang, Pathomwattananurak &amp; Zhao" authorityYear="2023" box="[151,932,1393,1420]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="295" phylum="Arthropoda" rank="species" species="vietnamensis" status="new species">
<emphasis bold="true" box="[151,464,1393,1420]" italics="true" pageId="5" pageNumber="295">Eophileurus vietnamensis</emphasis>
Yang, Pathomwattananurak &amp; Zhao
</taxonomicName>
,
<taxonomicNameLabel box="[945,1089,1393,1420]" pageId="5" pageNumber="295" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[151,409,1429,1456]" pageId="5" pageNumber="295" type="description">
<paragraph blockId="5.[151,1089,1393,1456]" box="[151,409,1429,1456]" pageId="5" pageNumber="295">
(
<figureCitation box="[159,308,1430,1456]" captionStart="FIGURE 4" captionStartId="6.[151,250,1969,1993]" captionTargetBox="[160,1431,907,1928]" captionTargetId="figure-324@6.[151,1435,885,1945]" captionTargetPageId="6" captionText="FIGURE 4. Habitus of Eophileurus vietnamensis Yang, Pathomwattananurak &amp; Zhao, new species. AC, holotype; DE, male paratype; FH, female paratype; A, D, F, dorsal view; B, G, ventral view; C, E, H, oblique view." figureDoi="http://doi.org/10.5281/zenodo.7687390" httpUri="https://zenodo.org/record/7687390/files/figure.png" pageId="5" pageNumber="295">Figs. 4A4H</figureCitation>
,
<figureCitation box="[319,401,1429,1455]" captionStart="FIGURE 6" captionStartId="8.[151,250,1546,1570]" captionTargetBox="[188,1402,538,1503]" captionTargetId="figure-160@8.[177,1411,525,1523]" captionTargetPageId="8" captionText="FIGURE 6. Male genitalia of Eophileurus species. AB, E. vietnamensis Yang, Pathomwattananurak &amp; Zhao, new species, holotype; CD, E. chinensis (Faldermann, 1835); A, C, frontal view of parameres; B, D, lateral view of aedeagus." figureDoi="http://doi.org/10.5281/zenodo.7687394" httpUri="https://zenodo.org/record/7687394/files/figure.png" pageId="5" pageNumber="295">6A6B</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection lastPageId="7" lastPageNumber="297" pageId="5" pageNumber="295" type="materials_examined">
<paragraph blockId="5.[151,1437,1501,2032]" pageId="5" pageNumber="295">
<emphasis bold="true" box="[151,325,1501,1528]" pageId="5" pageNumber="295">Type material.</emphasis>
(4♁♁,
<specimenCount box="[412,468,1501,1528]" pageId="5" pageNumber="295" type="female">5♀♀</specimenCount>
).
<materialsCitation collectingDate="2022-03" collectionCode="CAU" country="Vietnam" elevation="1475" location="Pu Luong Nature Reserve" municipality="Ba Thuoc Dist." pageId="5" pageNumber="295" specimenCount="1" stateProvince="Thanh Hoa" typeStatus="holotype">
<emphasis bold="true" box="[491,610,1502,1528]" pageId="5" pageNumber="295">
<typeStatus box="[491,604,1502,1528]" pageId="5" pageNumber="295">Holotype</typeStatus>
:
</emphasis>
♁ (
<collectionCode box="[653,717,1502,1528]" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33074" lsid="urn:lsid:biocol.org:col:33074" name="China Agricultural University" pageId="5" pageNumber="295">CAU</collectionCode>
), “
<location LSID="urn:lsid:plazi:treatment:0A057876FFAB2B2135939870FE03FE3D:87739FBBFFAB2B2C37EF98DFFBDEFA30" box="[747,1043,1502,1528]" country="Vietnam" municipality="Ba Thuoc Dist." name="Pu Luong Nature Reserve" pageId="5" pageNumber="295" stateProvince="Thanh Hoa">Pu Luong Nature Reserve</location>
,
<collectingMunicipality box="[1054,1228,1502,1528]" pageId="5" pageNumber="295">Ba Thuoc Dist.</collectingMunicipality>
,
<collectingRegion box="[1238,1432,1502,1528]" country="Vietnam" name="Thanh Hoa" pageId="5" pageNumber="295">Thanh Hoa Prov.</collectingRegion>
,
<collectingCountry box="[151,249,1538,1564]" name="Vietnam" pageId="5" pageNumber="295">Vietnam</collectingCountry>
, alt.
<quantity box="[305,464,1537,1564]" metricMagnitude="3" metricUnit="m" metricValue="1.475" metricValueMax="1.75" metricValueMin="1.2" pageId="5" pageNumber="295" unit="m" value="1475.0" valueMax="1750.0" valueMin="1200.0">
<elevation box="[305,464,1537,1564]" metricMagnitude="3" metricUnit="m" metricValue="1.475" metricValueMax="1.75" metricValueMin="1.2" pageId="5" pageNumber="295" unit="m" value="1475.0" valueMax="1750.0" valueMin="1200.0">12001750 m</elevation>
</quantity>
,
<date box="[477,571,1538,1564]" pageId="5" pageNumber="295" value="2022-03">
<collectingDate box="[477,571,1538,1564]" pageId="5" pageNumber="295" value="2022-03">III.2022</collectingDate>
</date>
, local collector leg.”
</materialsCitation>
<materialsCitation collectingDate="2022-04" collectingDateMax="2022-06" collectingDateMin="2022-04" collectionCode="CAU, CQZY" collectorName="Van Dang" country="Vietnam" county="Ba Thuoc Dist." elevation="1475" location="Pu Luong Nature Reserve" municipality="Pu Luong Nature Reserve" pageId="5" pageNumber="295" specimenCount="5" specimenCount-female="5" stateProvince="Thanh Hoa" typeStatus="paratype">
<emphasis bold="true" box="[815,944,1538,1564]" pageId="5" pageNumber="295">
<typeStatus box="[815,939,1538,1564]" pageId="5" pageNumber="295">Paratypes</typeStatus>
:
</emphasis>
<specimenCount box="[953,987,1538,1564]" pageId="5" pageNumber="295" type="female">1♀</specimenCount>
(
<collectionCode box="[1004,1066,1538,1564]" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33074" lsid="urn:lsid:biocol.org:col:33074" name="China Agricultural University" pageId="5" pageNumber="295">CAU</collectionCode>
) “
<location LSID="urn:lsid:plazi:treatment:0A057876FFAB2B2135939870FE03FE3D:87739FBBFFAB2B2C31419B03FABDF9D4" box="[1093,1392,1538,1564]" country="Vietnam" county="Ba Thuoc Dist." municipality="Pu Luong Nature Reserve" name="Pu Luong Nature Reserve" pageId="5" pageNumber="295" stateProvince="Thanh Hoa">Pu Luong Nature Reserve</location>
,
<location LSID="urn:lsid:plazi:treatment:0A057876FFAB2B2135939870FE03FE3D:87739FBBFFAB2B2C30799B03FED2F988" country="Vietnam" county="Ba Thuoc Dist." municipality="Pu Luong Nature Reserve" name="Ba Thuoc Dist." pageId="5" pageNumber="295" stateProvince="Thanh Hoa">Ba Thuoc Dist.</location>
,
<collectingRegion box="[298,492,1574,1600]" country="Vietnam" name="Thanh Hoa" pageId="5" pageNumber="295">Thanh Hoa Prov.</collectingRegion>
,
<collectingCountry box="[502,600,1574,1600]" name="Vietnam" pageId="5" pageNumber="295">Vietnam</collectingCountry>
, alt.
<quantity box="[654,811,1573,1600]" metricMagnitude="3" metricUnit="m" metricValue="1.475" metricValueMax="1.75" metricValueMin="1.2" pageId="5" pageNumber="295" unit="m" value="1475.0" valueMax="1750.0" valueMin="1200.0">
<elevation box="[654,811,1573,1600]" metricMagnitude="3" metricUnit="m" metricValue="1.475" metricValueMax="1.75" metricValueMin="1.2" pageId="5" pageNumber="295" unit="m" value="1475.0" valueMax="1750.0" valueMin="1200.0">12001750 m</elevation>
</quantity>
,
<date box="[823,918,1574,1600]" pageId="5" pageNumber="295" value="2022-06">
<collectingDate box="[823,918,1574,1600]" pageId="5" pageNumber="295" value="2022-06">VI.2022</collectingDate>
</date>
, local collector leg.”; 1♁ (
<collectionCode box="[1216,1298,1574,1600]" pageId="5" pageNumber="295">CQZY</collectionCode>
), same label as the
<typeStatus box="[225,327,1610,1636]" pageId="5" pageNumber="295">holotype</typeStatus>
; 2♁♁,
<specimenCount box="[410,465,1610,1636]" pageId="5" pageNumber="295" type="female">3♀♀</specimenCount>
(
<collectionCode box="[482,562,1610,1636]" pageId="5" pageNumber="295">CQZY</collectionCode>
) “
<collectingMunicipality box="[587,885,1610,1636]" pageId="5" pageNumber="295">Pu Luong Nature Reserve</collectingMunicipality>
,
<collectingCounty box="[898,1073,1610,1636]" pageId="5" pageNumber="295">Ba Thuoc Dist.</collectingCounty>
,
<collectingRegion box="[1084,1280,1610,1636]" country="Vietnam" name="Thanh Hoa" pageId="5" pageNumber="295">Thanh Hoa Prov.</collectingRegion>
,
<collectingCountry box="[1291,1389,1610,1636]" name="Vietnam" pageId="5" pageNumber="295">Vietnam</collectingCountry>
, alt.
<quantity box="[151,308,1645,1672]" metricMagnitude="3" metricUnit="m" metricValue="1.475" metricValueMax="1.75" metricValueMin="1.2" pageId="5" pageNumber="295" unit="m" value="1475.0" valueMax="1750.0" valueMin="1200.0">
<elevation box="[151,308,1645,1672]" metricMagnitude="3" metricUnit="m" metricValue="1.475" metricValueMax="1.75" metricValueMin="1.2" pageId="5" pageNumber="295" unit="m" value="1475.0" valueMax="1750.0" valueMin="1200.0">12001750 m</elevation>
</quantity>
,
<date box="[319,415,1646,1672]" pageId="5" pageNumber="295" value="2022-06">
<collectingDate box="[319,415,1646,1672]" pageId="5" pageNumber="295" value="2022-06">VI.2022</collectingDate>
</date>
, local collector leg.”;
<specimenCount box="[661,696,1646,1672]" pageId="5" pageNumber="295" type="female">1♀</specimenCount>
(
<collectionCode box="[711,793,1646,1672]" pageId="5" pageNumber="295">CQZY</collectionCode>
), “
<collectingRegion box="[822,993,1646,1672]" country="Vietnam" name="Nghe An" pageId="5" pageNumber="295">Nghe An Prov.</collectingRegion>
,
<collectingCountry box="[1003,1101,1646,1672]" name="Vietnam" pageId="5" pageNumber="295">Vietnam</collectingCountry>
,
<date box="[1112,1203,1646,1672]" pageId="5" pageNumber="295" value="2022-04">
<collectingDate box="[1112,1203,1646,1672]" pageId="5" pageNumber="295" value="2022-04">IV.2022</collectingDate>
</date>
,
<collectorName box="[1214,1326,1646,1672]" pageId="5" pageNumber="295">Van Dang</collectorName>
leg.”
</materialsCitation>
</paragraph>
<paragraph blockId="5.[151,1437,1501,2032]" lastBlockId="6.[151,1437,151,861]" lastPageId="6" lastPageNumber="296" pageId="5" pageNumber="295">
<emphasis bold="true" box="[199,308,1682,1708]" pageId="5" pageNumber="295">
<typeStatus box="[199,308,1682,1708]" pageId="5" pageNumber="295">Holotype</typeStatus>
</emphasis>
(male).
<emphasis bold="true" box="[409,505,1681,1707]" pageId="5" pageNumber="295">General</emphasis>
(
<figureCitation box="[524,673,1682,1708]" captionStart="FIGURE 4" captionStartId="6.[151,250,1969,1993]" captionTargetBox="[160,1431,907,1928]" captionTargetId="figure-324@6.[151,1435,885,1945]" captionTargetPageId="6" captionText="FIGURE 4. Habitus of Eophileurus vietnamensis Yang, Pathomwattananurak &amp; Zhao, new species. AC, holotype; DE, male paratype; FH, female paratype; A, D, F, dorsal view; B, G, ventral view; C, E, H, oblique view." figureDoi="http://doi.org/10.5281/zenodo.7687390" httpUri="https://zenodo.org/record/7687390/files/figure.png" pageId="5" pageNumber="295">Figs. 4A4C</figureCitation>
): Body black, metasternum with slight reddish color, body shape oblong, flattened, surface shiny; ventral surface with sparse, short, reddish-yellow setae.
<emphasis bold="true" box="[1163,1236,1718,1744]" pageId="5" pageNumber="295">Head:</emphasis>
Clypeus sparsely punctate, with micropunctures on anterior region and larger punctures near sides; center with a long, slightly backward horn and a weak ridge from horn to apex; clypeal apex acute. Frons moderately depressed, disk with sparse micropunctures. Mandible sinuate on outer margin.
<emphasis bold="true" box="[831,962,1826,1852]" pageId="5" pageNumber="295">Pronotum:</emphasis>
Surface with dense punctures, punctures becoming denser and larger near all margins. Disc with a large, deep, round fovea; punctures in fovea dense and large, round or almost U-shaped; with 2 weak protuberances on each side. All margins beaded except before fovea, more strongly beaded at sides.Anterior angle protruding, apex rounded, posterior angle rounded.
<emphasis bold="true" box="[1217,1346,1933,1959]" pageId="5" pageNumber="295">Scutellum:</emphasis>
Surface with dense punctures on disc and sparse micropunctures near base.
<emphasis bold="true" box="[928,1013,1970,1996]" pageId="5" pageNumber="295">Elytra:</emphasis>
Surface with sparse micropunctures, and dense and large, round punctures. Interstice 1 with punctures in 3 irregular rows combined to 2 irregular rows at middle then to 1 irregular row near posterior one-fourth, interstice 2 with punctures in 1 irregular row. Primary costa A distinct, primary costa B indistinct. Subapical umbone prominent. Lateral margin of elytron slightly curved outward, weakly and horizontally dilated from base to about posterior one-third.
<emphasis bold="true" box="[1046,1170,222,248]" pageId="6" pageNumber="296">Pygidium:</emphasis>
Surface convex, almost impunctate on disc, strongly rugopunctate near base and densely punctate on remaining area.
<emphasis bold="true" box="[1179,1350,259,285]" pageId="6" pageNumber="296">Metasternum:</emphasis>
Region near sides with dense and large punctures; median portion with sparse and small punctures; setae reddish-yellow, short, on anterior region excluding the middle, setae longer near base.
<emphasis bold="true" box="[943,1198,330,356]" pageId="6" pageNumber="296">Abdominal ventrites:</emphasis>
Surfaces with sparse micropunctures throughout and larger punctures on each side; ventrite 5 and 6 each with a long, shallow horizontal depression at center.
<emphasis bold="true" box="[384,450,403,429]" pageId="6" pageNumber="296">Legs:</emphasis>
Protibia tridentate, protarsi moderately thickened, inner protibial claw strongly enlarged and moderately elongated. Inner metatibial spur long, slightly curved outward at middle; outer metatibial spur shorter, slightly curved outwards more strongly so than inner metatibial spur.
<emphasis bold="true" box="[1033,1162,475,501]" pageId="6" pageNumber="296">Parameres</emphasis>
(
<figureCitation box="[1178,1325,475,501]" captionStart="FIGURE 6" captionStartId="8.[151,250,1546,1570]" captionTargetBox="[188,1402,538,1503]" captionTargetId="figure-160@8.[177,1411,525,1523]" captionTargetPageId="8" captionText="FIGURE 6. Male genitalia of Eophileurus species. AB, E. vietnamensis Yang, Pathomwattananurak &amp; Zhao, new species, holotype; CD, E. chinensis (Faldermann, 1835); A, C, frontal view of parameres; B, D, lateral view of aedeagus." figureDoi="http://doi.org/10.5281/zenodo.7687394" httpUri="https://zenodo.org/record/7687394/files/figure.png" pageId="6" pageNumber="296">Figs. 6A6B</figureCitation>
): Slender and symmetric; in frontal view (
<figureCitation box="[521,613,511,537]" captionStart="FIGURE 6" captionStartId="8.[151,250,1546,1570]" captionTargetBox="[188,1402,538,1503]" captionTargetId="figure-160@8.[177,1411,525,1523]" captionTargetPageId="8" captionText="FIGURE 6. Male genitalia of Eophileurus species. AB, E. vietnamensis Yang, Pathomwattananurak &amp; Zhao, new species, holotype; CD, E. chinensis (Faldermann, 1835); A, C, frontal view of parameres; B, D, lateral view of aedeagus." figureDoi="http://doi.org/10.5281/zenodo.7687394" httpUri="https://zenodo.org/record/7687394/files/figure.png" pageId="6" pageNumber="296">Fig. 6A</figureCitation>
), outer side prominent on basal one-third and at middle, then parallel at sides; inner margins parallel, with a small, acute and frontward process, then curved backward and outward, and then inward near apex; apex rounded, pointed inward; in lateral view (
<figureCitation box="[960,1050,583,609]" captionStart="FIGURE 6" captionStartId="8.[151,250,1546,1570]" captionTargetBox="[188,1402,538,1503]" captionTargetId="figure-160@8.[177,1411,525,1523]" captionTargetPageId="8" captionText="FIGURE 6. Male genitalia of Eophileurus species. AB, E. vietnamensis Yang, Pathomwattananurak &amp; Zhao, new species, holotype; CD, E. chinensis (Faldermann, 1835); A, C, frontal view of parameres; B, D, lateral view of aedeagus." figureDoi="http://doi.org/10.5281/zenodo.7687394" httpUri="https://zenodo.org/record/7687394/files/figure.png" pageId="6" pageNumber="296">Fig. 6B</figureCitation>
), upper margin slightly depressed near base; inner processes invisible, outer prominent part downward, almost at right angle at its base; apex slightly bent backward.
</paragraph>
<paragraph blockId="6.[151,1437,151,861]" pageId="6" pageNumber="296">
<emphasis bold="true" box="[199,319,691,717]" pageId="6" pageNumber="296">
<typeStatus box="[199,319,691,717]" pageId="6" pageNumber="296">Paratypes</typeStatus>
</emphasis>
(male). Similarly or slightly less developed than the
<typeStatus box="[993,1095,691,717]" pageId="6" pageNumber="296">holotype</typeStatus>
; characters stable, the most underdeveloped male with differences as follow: clypeal horn shorter, clypeus more densely punctate, frons with rugopunctures near base; fovea smaller, almost longitudinally oval in shape, distinctly wider anteriorly, widest on anterior one-third; anterior margin of pronotum almost entirely beaded, with a small gap at middle; scutellum more sparsely punctate, with only a few punctures near base (
<figureCitation box="[777,913,835,861]" captionStart="FIGURE 4" captionStartId="6.[151,250,1969,1993]" captionTargetBox="[160,1431,907,1928]" captionTargetId="figure-324@6.[151,1435,885,1945]" captionTargetPageId="6" captionText="FIGURE 4. Habitus of Eophileurus vietnamensis Yang, Pathomwattananurak &amp; Zhao, new species. AC, holotype; DE, male paratype; FH, female paratype; A, D, F, dorsal view; B, G, ventral view; C, E, H, oblique view." figureDoi="http://doi.org/10.5281/zenodo.7687390" httpUri="https://zenodo.org/record/7687390/files/figure.png" pageId="6" pageNumber="296">Fig. 4D4E</figureCitation>
).
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7687390" ID-Zenodo-Dep="7687390" httpUri="https://zenodo.org/record/7687390/files/figure.png" pageId="6" pageNumber="296" startId="6.[151,250,1969,1993]" targetBox="[160,1431,907,1928]" targetPageId="6">
<paragraph blockId="6.[151,1436,1969,2029]" pageId="6" pageNumber="296">
<emphasis bold="true" box="[151,274,1969,1993]" pageId="6" pageNumber="296">FIGURE 4.</emphasis>
Habitus of
<taxonomicName authority="Yang, Pathomwattananurak &amp; Zhao" authorityName="Yang, Pathomwattananurak &amp; Zhao" authorityYear="2023" box="[392,1023,1969,1993]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="296" phylum="Arthropoda" rank="species" species="vietnamensis" status="new species">
<emphasis box="[392,653,1969,1993]" italics="true" pageId="6" pageNumber="296">Eophileurus vietnamensis</emphasis>
Yang, Pathomwattananurak &amp; Zhao
</taxonomicName>
,
<emphasis bold="true" box="[1032,1157,1969,1993]" pageId="6" pageNumber="296">
<taxonomicNameLabel box="[1032,1157,1969,1993]" pageId="6" pageNumber="296" rank="species">new species</taxonomicNameLabel>
</emphasis>
. AC, holotype; DE, male paratype; FH, female paratype; A, D, F, dorsal view; B, G, ventral view; C, E, H, oblique view.
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7687392" ID-Zenodo-Dep="7687392" httpUri="https://zenodo.org/record/7687392/files/figure.png" pageId="7" pageNumber="297" startId="7.[151,250,1545,1569]" targetBox="[165,1423,190,1512]" targetPageId="7">
<paragraph blockId="7.[151,1436,1545,1605]" pageId="7" pageNumber="297">
<emphasis bold="true" box="[151,275,1545,1569]" pageId="7" pageNumber="297">FIGURE 5.</emphasis>
Habitus of
<taxonomicName authority="(Faldermann, 1835)" baseAuthorityName="Faldermann" baseAuthorityYear="1835" box="[393,828,1545,1569]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="297" phylum="Arthropoda" rank="species" species="chinensis">
<emphasis box="[393,617,1545,1569]" italics="true" pageId="7" pageNumber="297">Eophileurus chinensis</emphasis>
(
<bibRefCitation author="Faldermann, F." box="[630,820,1545,1569]" pageId="7" pageNumber="297" pagination="337 - 464" refId="ref5230" refString="Faldermann, F. (1835) Coleopterorum ab illustrissimo Bungio in China boreali, Mongolia et montibus altaicis collectorum nec non ab ill. Turchaninoffio et Stschukinoe e provincia Irkutsk missorum illustrations. Memoires de l'Academie imperiale des Sciences de Saint Petersbourg, 6 (2), 337 - 464." type="journal article" year="1835">Faldermann, 1835</bibRefCitation>
)
</taxonomicName>
. AD, male; EF, female; A, CE, dorsal view; B, F, ventral view.
</paragraph>
</caption>
<paragraph blockId="7.[151,1437,1651,2038]" pageId="7" pageNumber="297">
<emphasis bold="true" box="[199,319,1652,1678]" pageId="7" pageNumber="297">
<typeStatus box="[199,319,1652,1678]" pageId="7" pageNumber="297">Paratypes</typeStatus>
</emphasis>
(female). Similar to male, but clypeus and frons densely rugopunctate throughout; clypeal horn shorter; fovea absent, with a shallow and densely punctate furrow on anterior region of pronotum, punctures in furrow distinctly larger and deeper than the others, round or nearly U-shape; anterior margin of pronotum entirely beaded; scutellum sparsely punctate near base; pygidium flattened from middle to base, with a clear protuberance between flattened area and apex, apex with a weak, flat and wide protuberance, punctures on pygidium denser on disc; ventrite 5 and 6 without depression, ventrite 6 with sparser punctures, rugopunctate near base; protarsi not thickened, inner claw not enlarged and elongated (
<figureCitation box="[716,850,1867,1894]" captionStart="FIGURE 4" captionStartId="6.[151,250,1969,1993]" captionTargetBox="[160,1431,907,1928]" captionTargetId="figure-324@6.[151,1435,885,1945]" captionTargetPageId="6" captionText="FIGURE 4. Habitus of Eophileurus vietnamensis Yang, Pathomwattananurak &amp; Zhao, new species. AC, holotype; DE, male paratype; FH, female paratype; A, D, F, dorsal view; B, G, ventral view; C, E, H, oblique view." figureDoi="http://doi.org/10.5281/zenodo.7687390" httpUri="https://zenodo.org/record/7687390/files/figure.png" pageId="7" pageNumber="297">Fig. 4F4H</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="297" type="description">
<paragraph blockId="7.[151,1437,1651,2038]" pageId="7" pageNumber="297">
<emphasis bold="true" box="[199,382,1904,1930]" pageId="7" pageNumber="297">Measurements.</emphasis>
Length: male 22.0
<quantity box="[614,714,1904,1930]" metricMagnitude="-2" metricUnit="m" metricValue="2.41" pageId="7" pageNumber="297" unit="mm" value="24.1">24.1 mm</quantity>
(
<typeStatus box="[729,829,1903,1929]" pageId="7" pageNumber="297">holotype</typeStatus>
<quantity box="[838,944,1904,1930]" metricMagnitude="-2" metricUnit="m" metricValue="2.41" pageId="7" pageNumber="297" unit="mm" value="24.1">24.1 mm</quantity>
), female
<quantity box="[1049,1217,1903,1930]" metricMagnitude="-2" metricUnit="m" metricValue="2.36" metricValueMax="2.46" metricValueMin="2.26" pageId="7" pageNumber="297" unit="mm" value="23.6" valueMax="24.6" valueMin="22.6">22.624.6 mm</quantity>
; width: male
<quantity metricMagnitude="-2" metricUnit="m" metricValue="1.11" metricValueMax="1.19" metricValueMin="1.03" pageId="7" pageNumber="297" unit="mm" value="11.1" valueMax="11.9" valueMin="10.3">10.3 11.9 mm</quantity>
(
<typeStatus box="[264,364,1939,1965]" pageId="7" pageNumber="297">holotype</typeStatus>
<quantity box="[371,474,1940,1966]" metricMagnitude="-2" metricUnit="m" metricValue="1.19" pageId="7" pageNumber="297" unit="mm" value="11.9">11.9 mm</quantity>
), female
<quantity box="[576,741,1940,1966]" metricMagnitude="-2" metricUnit="m" metricValue="1.1649999999999998" metricValueMax="1.21" metricValueMin="1.1199999999999999" pageId="7" pageNumber="297" unit="mm" value="11.649999999999999" valueMax="12.1" valueMin="11.2">11.212.1 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection lastPageId="8" lastPageNumber="298" pageId="7" pageNumber="297" type="diagnosis">
<paragraph blockId="7.[151,1437,1651,2038]" lastBlockId="8.[151,1436,151,501]" lastPageId="8" lastPageNumber="298" pageId="7" pageNumber="297">
<emphasis bold="true" box="[199,322,1975,2001]" pageId="7" pageNumber="297">Diagnosis.</emphasis>
The new species is closely allied to
<taxonomicName authority="(Faldermann, 1835)" baseAuthorityName="Faldermann" baseAuthorityYear="1835" box="[742,1223,1975,2002]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="297" phylum="Arthropoda" rank="species" species="chinensis">
<emphasis box="[742,991,1975,2001]" italics="true" pageId="7" pageNumber="297">Eophileurus chinensis</emphasis>
(
<bibRefCitation author="Faldermann, F." box="[1007,1215,1975,2002]" pageId="7" pageNumber="297" pagination="337 - 464" refId="ref5230" refString="Faldermann, F. (1835) Coleopterorum ab illustrissimo Bungio in China boreali, Mongolia et montibus altaicis collectorum nec non ab ill. Turchaninoffio et Stschukinoe e provincia Irkutsk missorum illustrations. Memoires de l'Academie imperiale des Sciences de Saint Petersbourg, 6 (2), 337 - 464." type="journal article" year="1835">Faldermann, 1835</bibRefCitation>
)
</taxonomicName>
(
<figureCitation box="[1240,1384,1975,2002]" captionStart="FIGURE 5" captionStartId="7.[151,250,1545,1569]" captionTargetBox="[165,1423,190,1512]" captionTargetId="figure-20@7.[164,1423,181,1521]" captionTargetPageId="7" captionText="FIGURE 5. Habitus of Eophileurus chinensis (Faldermann, 1835). AD, male; EF, female; A, CE, dorsal view; B, F, ventral view." figureDoi="http://doi.org/10.5281/zenodo.7687392" httpUri="https://zenodo.org/record/7687392/files/figure.png" pageId="7" pageNumber="297">Figs. 5A5F</figureCitation>
) but can be distinguished by the following features: clypeus of similarly developed male usually slightly more densely punctate with slightly larger punctures; clypeus and frons of female usually with larger but shallower rugopunctures; fovea of well-developed males usually larger, more rounded; interstices 1 and 2 with denser punctures, row in interstice 2 irregular (usually regular in
<taxonomicName baseAuthorityName="Faldermann" baseAuthorityYear="1835" box="[599,735,223,249]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="298" phylum="Arthropoda" rank="species" species="chinensis">
<emphasis box="[599,735,223,249]" italics="true" pageId="8" pageNumber="298">E. chinensis</emphasis>
</taxonomicName>
, rarely slightly irregular); elytral margin more weakly dilated; protuberance at apex of female pygidium indistinct, much smaller and wider (rather distinct and strong in
<taxonomicName baseAuthorityName="Faldermann" baseAuthorityYear="1835" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="298" phylum="Arthropoda" rank="species" species="chinensis">
<emphasis italics="true" pageId="8" pageNumber="298">
<emphasis bold="true" box="[1412,1436,259,285]" italics="true" pageId="8" pageNumber="298">E.</emphasis>
chinensis
</emphasis>
</taxonomicName>
); outer margin of parameres parallel in apical half (diverging apicad toward the subapical curvature in
<taxonomicName baseAuthorityName="Faldermann" baseAuthorityYear="1835" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="298" phylum="Arthropoda" rank="species" species="chinensis">
<emphasis italics="true" pageId="8" pageNumber="298">
<emphasis bold="true" box="[1412,1436,295,321]" italics="true" pageId="8" pageNumber="298">E.</emphasis>
chinensis
</emphasis>
</taxonomicName>
) with much wider gap between the inner process and the apex of paramere, inner process much smaller and sharper, not visible in lateral view (inner process large and somewhat rounded at apex, clearly visible in lateral view in
<taxonomicName baseAuthorityName="Faldermann" baseAuthorityYear="1835" box="[241,376,403,429]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="298" phylum="Arthropoda" rank="species" species="chinensis">
<emphasis box="[241,376,403,429]" italics="true" pageId="8" pageNumber="298">E. chinensis</emphasis>
</taxonomicName>
,
<figureCitation box="[391,539,403,429]" captionStart="FIGURE 6" captionStartId="8.[151,250,1546,1570]" captionTargetBox="[188,1402,538,1503]" captionTargetId="figure-160@8.[177,1411,525,1523]" captionTargetPageId="8" captionText="FIGURE 6. Male genitalia of Eophileurus species. AB, E. vietnamensis Yang, Pathomwattananurak &amp; Zhao, new species, holotype; CD, E. chinensis (Faldermann, 1835); A, C, frontal view of parameres; B, D, lateral view of aedeagus." figureDoi="http://doi.org/10.5281/zenodo.7687394" httpUri="https://zenodo.org/record/7687394/files/figure.png" pageId="8" pageNumber="298">Figs. 6C6D</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection box="[199,882,439,465]" pageId="8" pageNumber="298" type="etymology">
<paragraph blockId="8.[151,1436,151,501]" box="[199,882,439,465]" pageId="8" pageNumber="298">
<emphasis bold="true" box="[199,334,439,465]" pageId="8" pageNumber="298">Etymology.</emphasis>
This species is named after its country of origin.
</paragraph>
</subSubSection>
<subSubSection box="[199,462,474,501]" pageId="8" pageNumber="298" type="distribution">
<paragraph blockId="8.[151,1436,151,501]" box="[199,462,474,501]" pageId="8" pageNumber="298">
<emphasis bold="true" box="[199,353,474,500]" pageId="8" pageNumber="298">Distribution.</emphasis>
<collectingCountry box="[359,458,475,501]" name="Vietnam" pageId="8" pageNumber="298">Vietnam</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>