156 lines
18 KiB
XML
156 lines
18 KiB
XML
<document id="FA6BC6D549D5B7ED6DB1B09D3BDA4DD5" ID-DOI="10.11646/zootaxa.3956.2.3" ID-GBIF-Dataset="5dd27599-4955-4417-a887-5497706a61d6" ID-ISSN="1175-5326" ID-Zenodo-Dep="246048" ID-ZooBank="7926BAC8-2480-4AD3-A7ED-30554A29C49A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461054632739" checkinUser="plazi" docAuthor="Löcker, Birgit" docDate="2015" docId="03C7E50DFFDEFFEEFF44ED33FC6CFCDA" docLanguage="en" docName="zt03956p223.pdf" docOrigin="Zootaxa 3956 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Aka gwana Löcker, 2015, sp. nov." docType="treatment" docUuid="06E87AA3-58B5-4325-BCA5-A77F6C8B0F99" docUuidSource="ZooBank" docVersion="9" lastPageNumber="211" masterDocId="FFFE9D75FFD5FFE2FFD3E804FFB2FFE4" masterDocTitle="Revision of the Australian species of Aka White, 1879 (Fulgoromorpha: Cixiidae) with the description of a new genus" masterLastPageNumber="223" masterPageNumber="199" pageNumber="210" updateTime="1698580774322" updateUser="plazi">
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<mods:title id="B23C4DA9014D67BC52A18C1192586F0C">Revision of the Australian species of Aka White, 1879 (Fulgoromorpha: Cixiidae) with the description of a new genus</mods:title>
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<mods:name id="B04E352B7FCA2CFA8413B99BE710A06B" type="personal">
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<mods:namePart id="B1B05236E515EA1C1580EFFAF783A195">Löcker, Birgit</mods:namePart>
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<mods:title id="0DF6D897A9A604E8144C00A88AB0D42C">Zootaxa</mods:title>
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<mods:date id="4C183C05E8B6ED08D624F2CEB599978B">2015</mods:date>
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<mods:number id="A4B0FEAC93CC248BAC18F95A31BD5BCB">3956</mods:number>
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<mods:detail id="A0EC08A5E6B1A9382B2C3D67C13F5BED" type="issue">
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<mods:number id="DAEB4F35834E75CBCEFCFC99A281B774">2</mods:number>
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<mods:classification id="74814D5F75A382E4D160F5A2DC081BF1">journal article</mods:classification>
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<mods:identifier id="12D2E249C1B7C5FD65B66FDFD035FDB2" type="DOI">10.11646/zootaxa.3956.2.3</mods:identifier>
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<mods:identifier id="C4CBFFD478FEF0249760757C0F37FAFC" type="Zenodo-Dep">246048</mods:identifier>
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<mods:identifier id="EC3FC7171BAC03E4482EC332B3F05248" type="ZooBank">7926BAC8-2480-4AD3-A7ED-30554A29C49A</mods:identifier>
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<treatment id="03C7E50DFFDEFFEEFF44ED33FC6CFCDA" ID-DOI="http://doi.org/10.5281/zenodo.6105766" ID-GBIF-Taxon="119612742" ID-Zenodo-Dep="6105766" LSID="urn:lsid:zoobank.org:act:06E87AA3-58B5-4325-BCA5-A77F6C8B0F99" httpUri="http://treatment.plazi.org/id/03C7E50DFFDEFFEEFF44ED33FC6CFCDA" lastPageId="12" lastPageNumber="211" pageId="11" pageNumber="210">
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<subSubSection id="C3740790FFDEFFE9FF44ED33FAF0FA19" pageId="11" pageNumber="210" type="nomenclature">
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<paragraph id="8BD1541BFFDEFFE9FF44ED33FE36FAB5" blockId="11.[151,1325,1335,1428]" box="[151,388,1335,1361]" pageId="11" pageNumber="210">
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<heading id="D099E377FFDEFFE9FF44ED33FE36FAB5" bold="true" box="[151,388,1335,1361]" fontSize="11" level="1" pageId="11" pageNumber="210" reason="1">
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<emphasis id="B91A8809FFDEFFE9FF44ED33FE36FAB5" bold="true" box="[151,388,1335,1361]" pageId="11" pageNumber="210">
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<taxonomicName id="4C6E2F98FFDEFFE9FF44ED33FEADFAB5" ID-CoL="65V5V" box="[151,287,1335,1361]" class="Insecta" family="Cixiidae" genus="Aka" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="210" phylum="Arthropoda" rank="species" species="gwana" status="sp. nov.">
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<emphasis id="B91A8809FFDEFFE9FF44ED33FEADFAB5" bold="true" box="[151,287,1335,1361]" italics="true" pageId="11" pageNumber="210">Aka gwana</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2293572FFDEFFE9FEF5ED33FE36FAB5" box="[294,388,1335,1361]" pageId="11" pageNumber="210" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BD1541BFFDEFFE9FF44ED5EFECCFA97" blockId="11.[151,1325,1335,1428]" box="[151,382,1370,1395]" pageId="11" pageNumber="210">
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(
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<figureCitation id="1355489EFFDEFFE9FF4CED5EFF51FA97" box="[159,227,1370,1395]" captionStart="FIGURE 3" captionStartId="4.[151,250,1998,2020]" captionTargetBox="[151,1431,488,1976]" captionTargetId="figure@4.[151,1436,468,1977]" captionTargetPageId="4" captionText="FIGURE 3. Aka gwana: A habitus; B, C head. Aka hardyi: D, E head; F, G habitus. Scale bar 1 mm." httpUri="https://zenodo.org/record/246051/files/figure.png" pageId="11" pageNumber="210">Figs 3</figureCitation>
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A–C, 7A, 10)
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</paragraph>
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<paragraph id="8BD1541BFFDEFFE9FF44ED7AFA9FFA70" blockId="11.[151,1325,1335,1428]" box="[151,1325,1406,1428]" pageId="11" pageNumber="210">Zoobank Registration: http://zoobank.org/ urn:lsid:zoobank.org:act:06E87AA3-58B5-4325-BCA5-A77F6C8B0F99</paragraph>
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<paragraph id="8BD1541BFFDEFFE9FF44EDC4FAF0FA19" blockId="11.[151,1437,1472,2037]" pageId="11" pageNumber="210">
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<emphasis id="B91A8809FFDEFFE9FF44EDC4FF51FA3D" bold="true" box="[151,227,1472,1497]" pageId="11" pageNumber="210">
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<typeStatus id="54D5EAB9FFDEFFE9FF44EDC4FF6DFA3D" box="[151,223,1472,1497]" pageId="11" pageNumber="210">Types</typeStatus>
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.
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</emphasis>
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<typeStatus id="54D5EAB9FFDEFFE9FF3CEDC5FEE4FA3D" box="[239,342,1473,1497]" pageId="11" pageNumber="210" type="holotype">
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<emphasis id="B91A8809FFDEFFE9FF3CEDC5FEE4FA3D" box="[239,342,1473,1497]" italics="true" pageId="11" pageNumber="210">Holotype</emphasis>
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</typeStatus>
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, ♂,
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<emphasis id="B91A8809FFDEFFE9FE43EDC4FDCBFA3D" bold="true" box="[400,633,1472,1497]" pageId="11" pageNumber="210">
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<collectingCountry id="F379148BFFDEFFE9FE43EDC4FD85FA3D" box="[400,567,1472,1497]" name="Australia" pageId="11" pageNumber="210">AUSTRALIA</collectingCountry>
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, Tas:
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</emphasis>
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Mt Michael,
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<quantity id="4C96F9FEFFDEFFE9FCCCEDC5FCD0FA3D" box="[799,866,1473,1497]" metricMagnitude="2" metricUnit="m" metricValue="7.1" pageId="11" pageNumber="210" unit="m" value="710.0">710m</quantity>
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, u.v.l.,
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<date id="FFD072DBFFDEFFE9FC68EDC5FB9EFA3D" box="[955,1068,1473,1497]" pageId="11" pageNumber="210" value="1990-10-24">24.x.1990</date>
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(T. Semmens) (TAIC 145562).
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<typeStatus id="54D5EAB9FFDEFFE9FF44EDE2FEBBFA19" box="[151,265,1510,1533]" pageId="11" pageNumber="210" type="paratype">
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<emphasis id="B91A8809FFDEFFE9FF44EDE2FEBBFA19" box="[151,265,1510,1533]" italics="true" pageId="11" pageNumber="210">Paratypes</emphasis>
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</typeStatus>
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,
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<specimenCount id="9D689F92FFDEFFE9FEC5EDE1FEF2FA19" box="[278,320,1509,1533]" pageId="11" pageNumber="210" type="generic">1 ♂</specimenCount>
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, same data as
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<typeStatus id="54D5EAB9FFDEFFE9FE30EDE0FDFAFA19" box="[483,584,1508,1533]" pageId="11" pageNumber="210" type="holotype">holotype</typeStatus>
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; 1 ♀, Mt Barrow summit,
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<quantity id="4C96F9FEFFDEFFE9FCA2EDE1FC73FA19" box="[881,961,1509,1533]" metricMagnitude="3" metricUnit="m" metricValue="1.28" pageId="11" pageNumber="210" unit="m" value="1280.0">1280m</quantity>
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, UVL,
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<date id="FFD072DBFFDEFFE9FBC7EDE0FBCFFA19" box="[1044,1149,1508,1533]" pageId="11" pageNumber="210" value="1990-01-25">25.i.1990</date>
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(L. Hill) (TAIC).
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</paragraph>
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</subSubSection>
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<subSubSection id="C3740790FFDEFFE9FF14EE0CFDD7F98D" pageId="11" pageNumber="210" type="etymology">
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<paragraph id="8BD1541BFFDEFFE9FF14EE0CFDD7F98D" blockId="11.[151,1437,1472,2037]" pageId="11" pageNumber="210">
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<emphasis id="B91A8809FFDEFFE9FF14EE0CFEFCF9C5" bold="true" box="[199,334,1544,1569]" pageId="11" pageNumber="210">Etymology.</emphasis>
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The term „
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<taxonomicName id="4C6E2F98FFDEFFE9FE05EE0DFD91F9C5" box="[470,547,1545,1569]" class="Insecta" family="Cixiidae" genus="Aka" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="210" phylum="Arthropoda" rank="species" species="gwana">gwana</taxonomicName>
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“ means „goanna“ in Torres Strait Creole, an indigenous language spoken in Queensland (
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<bibRefCitation id="EFFF29EAFFDEFFE9FEF8EE28FD3AF9A1" author="Thieberger" box="[299,648,1580,1605]" pageId="11" pageNumber="210" refString="Thieberger, N. & McGregor, W. (1994) Macquarie Aboriginal words. The Macquarie Library Pty Ltd, Sydney, 724 pp." type="book" year="1994">Thieberger & McGregor 1994</bibRefCitation>
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). Named after the long, forked spine on the apex of flagellum which resembles the forked tongue of a goanna.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3740790FFDEFFEEFF14EE71FBADFD12" lastPageId="12" lastPageNumber="211" pageId="11" pageNumber="210" type="diagnosis">
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<paragraph id="8BD1541BFFDEFFE9FF14EE71FE11F955" blockId="11.[151,1437,1472,2037]" pageId="11" pageNumber="210">
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<emphasis id="B91A8809FFDEFFE9FF14EE71FEF3F96A" bold="true" box="[199,321,1653,1678]" pageId="11" pageNumber="210">Diagnosis.</emphasis>
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This species can be distinguished from all other Australian
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<taxonomicName id="4C6E2F98FFDEFFE9FC3FEE71FBAAF969" box="[1004,1048,1653,1677]" class="Insecta" family="Cixiidae" genus="Aka" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="210" phylum="Arthropoda" rank="genus">
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<emphasis id="B91A8809FFDEFFE9FC3FEE71FBAAF969" box="[1004,1048,1653,1677]" italics="true" pageId="11" pageNumber="210">Aka</emphasis>
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</taxonomicName>
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by the presence of more than two spines on the flagellum.
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</paragraph>
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<paragraph id="8BD1541BFFDEFFE9FF14EEB9FAAEF86D" blockId="11.[151,1437,1472,2037]" pageId="11" pageNumber="210">
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<emphasis id="B91A8809FFDEFFE9FF14EEB9FE92F932" bold="true" box="[199,288,1725,1750]" pageId="11" pageNumber="210">Colour.</emphasis>
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Vertex mid brown, carinae paler, with a whitish spot where subapical carinae meet lateral borders of vertex or vertex entirely light brown. Face light to mid brown, darker near vertex; carinae concolorous; frons often with a whitish spot at each lateral end of frontoclypeal suture. Pro- and mesonotum brown with slightly paler carinae; pronotum either paler than mesonotum or both the same colour but lateral parts darker. Forewings light brown to whitish, mottled with darker spots; veins concolorous with cells; tubercles darker, in distinct contrast to cells; pterostigma whitish. Abdominal sternites light brown. Legs light brown with a few darker marks.
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</paragraph>
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<paragraph id="8BD1541BFFDEFFE9FF14EF91FCE9F849" blockId="11.[151,1437,1472,2037]" box="[199,859,1941,1966]" pageId="11" pageNumber="210">
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<emphasis id="B91A8809FFDEFFE9FF14EF91FED2F84A" bold="true" box="[199,352,1941,1966]" pageId="11" pageNumber="210">Morphology.</emphasis>
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Body length: ♂
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<quantity id="4C96F9FEFFDEFFE9FDCFEF91FD17F849" box="[540,677,1941,1965]" metricMagnitude="-3" metricUnit="m" metricValue="4.25" metricValueMax="4.3" metricValueMin="4.2" pageId="11" pageNumber="210" unit="mm" value="4.25" valueMax="4.3" valueMin="4.2">4.2–4.3 mm</quantity>
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; ♀
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<quantity id="4C96F9FEFFDEFFE9FD1EEF91FCE5F849" box="[717,855,1941,1966]" metricMagnitude="-3" metricUnit="m" metricValue="5.35" metricValueMax="5.6" metricValueMin="5.1" pageId="11" pageNumber="210" unit="mm" value="5.35" valueMax="5.6" valueMin="5.1">5.1–5.6 mm</quantity>
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.
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</paragraph>
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<paragraph id="8BD1541BFFDEFFEEFF14EFBCFC20FF1C" blockId="11.[151,1437,1472,2037]" lastBlockId="12.[151,1437,151,830]" lastPageId="12" lastPageNumber="211" pageId="11" pageNumber="210">
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Head: Vertex 1.3–1.4 x wider than long; median carina of vertex covering about 3/4 of basal compartment of vertex. Frons 1.2–1.3 x longer than wide; maximum width no more than
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<date id="FFD072DBFFDEFFE9FC37EFD9FC4DF811" box="[996,1023,2013,2037]" pageId="11" pageNumber="210">2x</date>
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apical width; position of maximum width distinctly dorsad of centre of frontoclypeal suture; median carina forked in basal (near frontoclypeal suture) third to half of frons. Median ocellus present (indistinct). Anteclypeus with very indistinct median carina. Subapical segment of rostrum 1.3–1.7 x longer than apical segment.
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</paragraph>
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<paragraph id="8BD1541BFFD9FFEEFF14E900FC8BFDA5" blockId="12.[151,1437,151,830]" pageId="12" pageNumber="211">
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Thorax: Mesonotum with indistinct sublateral carinae. Forewing 2.9–3.3 x longer than wide; costal margin with 8–11 distinct tubercles; tubercles aligned alongside veins, but reaching to some extent into cell area; tubercles large, dark, in distinct contrast to cells; many tubercles bearing very long setae; Sc+R+M near basal cell fused, forming a minute common stem Sc+R+M; fork of ScRA+
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<collectingCountry id="F379148BFFD9FFEEFCE1E974FCE7FE6C" box="[818,853,368,392]" name="Poland" pageId="12" pageNumber="211">RP</collectingCountry>
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distinctly basad, slightly basad or at same level as fork CuA1+CuA2; distance tegula to ScR+M fork distinctly (about ½ to ¾) shorter than distance between this fork and ScRA+
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<collectingCountry id="F379148BFFD9FFEEFECBE9BCFE89FE34" box="[280,315,440,464]" name="Poland" pageId="12" pageNumber="211">RP</collectingCountry>
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fork; position of r-m moderately basad of fork MA+MP; fork MA+MP situated around apical 1/3 of forewing; icu slightly distad of apex of clavus;
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<collectingCountry id="F379148BFFD9FFEEFD74E9D9FD7BFE10" box="[679,713,477,500]" name="Poland" pageId="12" pageNumber="211">RP</collectingCountry>
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apically bifid; MA bifid; MP unforked; nodus of y-vein more or less in centre of clavus; y-vein with A1 moderately elevated; 7 apical cells.
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<collectingCountry id="F379148BFFD9FFEEFC2EEA06FB87FDFF" box="[1021,1077,514,539]" name="India" pageId="12" pageNumber="211">Hind</collectingCountry>
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leg: 1st tarsomere with 7 apical teeth; 2nd tarsomere with 7–8 apical teeth and 4–5 fine setae.
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</paragraph>
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<paragraph id="8BD1541BFFD9FFEEFF14EA49FBADFD12" blockId="12.[151,1437,151,830]" pageId="12" pageNumber="211">
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Male genitalia: Anal tube (
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<figureCitation id="1355489EFFD9FFEEFE29EA49FDE3FD82" box="[506,593,589,614]" captionStart="FIGURE 10" captionStartId="12.[151,250,1485,1507]" captionTargetBox="[331,1266,889,1441]" captionTargetId="figure@12.[321,1266,859,1464]" captionTargetPageId="12" captionText="FIGURE 10. Aka gwana: A aedeagus ventral; B aedeagus right lateral; C – D anal tube; E pygofer; F – G genital styles." httpUri="https://zenodo.org/record/246058/files/figure.png" pageId="12" pageNumber="211">Figs 10</figureCitation>
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C,D) apically slightly indented, forming two rounded lobes. Pygofer as in
|
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<figureCitation id="1355489EFFD9FFEEFF44EA75FF5EFD6E" box="[151,236,625,650]" captionStart="FIGURE 10" captionStartId="12.[151,250,1485,1507]" captionTargetBox="[331,1266,889,1441]" captionTargetId="figure@12.[321,1266,859,1464]" captionTargetPageId="12" captionText="FIGURE 10. Aka gwana: A aedeagus ventral; B aedeagus right lateral; C – D anal tube; E pygofer; F – G genital styles." httpUri="https://zenodo.org/record/246058/files/figure.png" pageId="12" pageNumber="211">Fig. 10</figureCitation>
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E. Genital styles apically rounded, laterally produced resembling the beak of a bird as in
|
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<figureCitation id="1355489EFFD9FFEEFACBEA75FADDFD6E" box="[1304,1391,625,650]" captionStart="FIGURE 10" captionStartId="12.[151,250,1485,1507]" captionTargetBox="[331,1266,889,1441]" captionTargetId="figure@12.[321,1266,859,1464]" captionTargetPageId="12" captionText="FIGURE 10. Aka gwana: A aedeagus ventral; B aedeagus right lateral; C – D anal tube; E pygofer; F – G genital styles." httpUri="https://zenodo.org/record/246058/files/figure.png" pageId="12" pageNumber="211">Figs 10</figureCitation>
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F,G. Aedeagus (
|
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<figureCitation id="1355489EFFD9FFEEFEC7EA91FED9FD4A" box="[276,363,661,686]" captionStart="FIGURE 10" captionStartId="12.[151,250,1485,1507]" captionTargetBox="[331,1266,889,1441]" captionTargetId="figure@12.[321,1266,859,1464]" captionTargetPageId="12" captionText="FIGURE 10. Aka gwana: A aedeagus ventral; B aedeagus right lateral; C – D anal tube; E pygofer; F – G genital styles." httpUri="https://zenodo.org/record/246058/files/figure.png" pageId="12" pageNumber="211">Figs 10</figureCitation>
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A,B): Phallotheca left laterally with spines (a,b) and right laterally with a curved spine (c). Base of phallotheca ventrally with 2 small sclerotised teeth. Flagellum near base with 3 sclerotised spines (d,e,f), apically widening into a large disc which gives rise to a long, bifurcate spine (g).
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</paragraph>
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</subSubSection>
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<subSubSection id="C3740790FFD9FFEEFF14EB05FC6CFCDA" pageId="12" pageNumber="211" type="discussion">
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<paragraph id="8BD1541BFFD9FFEEFF14EB05FC6CFCDA" blockId="12.[151,1437,151,830]" pageId="12" pageNumber="211">
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<emphasis id="B91A8809FFD9FFEEFF14EB05FE89FCFE" bold="true" box="[199,315,769,794]" pageId="12" pageNumber="211">Remarks.</emphasis>
|
||
In one of the specimens the CuA1+CuA2 fork is in an unusual position near the apex of the forewing, therefore the ScRA+
|
||
<collectingCountry id="F379148BFFD9FFEEFE22EB22FDA6FCDA" box="[497,532,806,830]" name="Poland" pageId="12" pageNumber="211">RP</collectingCountry>
|
||
fork is distinctly basad in this specimen.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |