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<document id="BE1E58D26D811E741236E403BD690A55" ID-CLB-Dataset="264199" ID-DOI="10.11646/zootaxa.5332.1.1" ID-GBIF-Dataset="66e0ec51-d494-43d4-965e-a2cd1462ef54" ID-ISSN="1175-5326" ID-Zenodo-Dep="8261502" ID-ZooBank="424F7439-4095-46A5-93E3-C4130E3B6D9A" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="juliana" checkinTime="1692350092877" checkinUser="plazi" docAuthor="POORANI, J." docDate="2023" docId="03C741621444462BBDDF36EBFD8AFC67" docLanguage="en" docName="zootaxa.5332.1.1.pdf" docOrigin="Zootaxa 5332 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5332.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Adalia simmondsi Kapur &amp; Sudha Rao" docType="treatment" docVersion="11" lastPageNumber="29" masterDocId="FFFE391A14584637BD48367CFFCDFF9B" masterDocTitle="An illustrated guide to lady beetles (Coleoptera: Coccinellidae) of the Indian Subcontinent. Part 1. Tribe Coccinellini" masterLastPageNumber="307" masterPageNumber="1" pageNumber="29" updateTime="1699593964221" updateUser="ExternalLinkService">
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<mods:title id="FBF0E5B5E4E52E91E022B6DECEF4859F">An illustrated guide to lady beetles (Coleoptera: Coccinellidae) of the Indian Subcontinent. Part 1. Tribe Coccinellini</mods:title>
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<treatment id="03C741621444462BBDDF36EBFD8AFC67" ID-DOI="http://doi.org/10.5281/zenodo.8261441" ID-GBIF-Taxon="212450162" ID-Zenodo-Dep="8261441" LSID="urn:lsid:plazi:treatment:03C741621444462BBDDF36EBFD8AFC67" httpUri="http://treatment.plazi.org/id/03C741621444462BBDDF36EBFD8AFC67" lastPageNumber="29" pageId="28" pageNumber="29">
<subSubSection id="C374A3FF1444462BBDDF36EBFDB1FF2A" box="[151,636,151,178]" pageId="28" pageNumber="29" type="nomenclature">
<paragraph id="8BD1F0741444462BBDDF36EBFDB1FF2A" blockId="28.[151,636,151,213]" box="[151,636,151,178]" pageId="28" pageNumber="29">
<heading id="D09947181444462BBDDF36EBFDB1FF2A" bold="true" box="[151,636,151,178]" fontSize="11" level="2" pageId="28" pageNumber="29" reason="4">
<taxonomicName id="4C6E8BF71444462BBDDF36EBFDB1FF2A" authority="Kapur &amp; Sudha Rao" authorityName="Kapur &amp; Sudha Rao" box="[151,636,151,178]" class="Insecta" family="Coccinellidae" genus="Adalia" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="simmondsi">
<emphasis id="B91A2C661444462BBDDF36EBFDB1FF2A" bold="true" box="[151,636,151,178]" pageId="28" pageNumber="29">
<emphasis id="B91A2C661444462BBDDF36EBFEBDFF29" bold="true" box="[151,368,151,178]" italics="true" pageId="28" pageNumber="29">Adalia simmondsi</emphasis>
Kapur &amp; Sudha Rao
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C374A3FF1444462BBDDF36C7FF37FF4E" box="[151,250,187,213]" pageId="28" pageNumber="29" type="description">
<paragraph id="8BD1F0741444462BBDDF36C7FF37FF4E" blockId="28.[151,636,151,213]" box="[151,250,187,213]" pageId="28" pageNumber="29">
(
<figureCitation id="1355ECF11444462BBDD736C7FF3CFF4E" box="[159,241,187,213]" captionStart="FIGURE 14" captionStartId="27.[151,250,1840,1864]" captionTargetBox="[166,1418,674,1808]" captionTargetId="figure-219@27.[151,1436,669,1816]" captionTargetPageId="27" captionText="FIGURE 14. Adalia simmondsi Kapur &amp; Sudha Rao: a. adult, dorsal view; b. spermatheca; c–e. male genitalia: c. tegmen, lateral view; d. tegmen, ventral view; e. penis." figureDoi="http://doi.org/10.5281/zenodo.8261548" httpUri="https://zenodo.org/record/8261548/files/figure.png" pageId="28" pageNumber="29">Fig. 14</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C374A3FF1444462BBDDF377CFE02FEEF" pageId="28" pageNumber="29" type="reference_group">
<paragraph id="8BD1F0741444462BBDDF377CFE36FEAD" blockId="28.[151,1437,255,372]" pageId="28" pageNumber="29">
<treatmentCitationGroup id="AB7ED75A1444462BBDDF377CFE36FEAD" pageId="28" pageNumber="29">
<treatmentCitation id="0ACFD6651444462BBDDF377CFD63FE8C" author="Kapur, A. P. &amp; Sudha Rao, V." box="[151,686,255,280]" page="51" pageId="28" pageNumber="29" year="1962">
<taxonomicName id="4C6E8BF71444462BBDDF377CFD63FE8C" authority="Kapur &amp; Sudha Rao, 1962: 49 - 51" authorityName="Kapur &amp; Sudha Rao" authorityPageNumber="49 - 51" authorityYear="1962" box="[151,686,255,280]" class="Insecta" family="Coccinellidae" genus="Adalia" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="simmondsi">
<emphasis id="B91A2C661444462BBDDF377CFE80FE83" box="[151,333,256,280]" italics="true" pageId="28" pageNumber="29">Adalia simmondsi</emphasis>
<bibRefCitation id="EFFF8D851444462BBC1C377CFD63FE8C" author="Kapur, A. P. &amp; Sudha Rao, V." box="[340,686,255,280]" pageId="28" pageNumber="29" pagination="47 - 51" refId="ref115835" refString="Kapur, A. P. &amp; Sudha Rao, V. (1962) A new species of ladybird beetle (Col., Coccinellidae) feeding on Adelges spp. (Hemiptera: Adelgidae) in North western India. Entomophaga, 7, 47 - 51. https: // doi. org / 10.1007 / BF 02375991" type="journal article" year="1962">Kapur &amp; Sudha Rao, 1962: 49–51</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(
<typeStatus id="54D54ED61444462BBFF3377CFCD6FE83" box="[699,795,256,280]" pageId="28" pageNumber="29" type="holotype">Holotype</typeStatus>
male, ZSIK; Type locality: Kalatop Forest, Chamba,
<collectingRegion id="49AA3E961444462BB872377CFED1FEAD" country="India" name="Himachal Pradesh" pageId="28" pageNumber="29">Himachal Pradesh</collectingRegion>
);
<treatmentCitation id="0ACFD6651444462BBC653762FE3AFEAD" author="Poorani, J." box="[301,503,286,310]" page="320" pageId="28" pageNumber="29" year="2002">
<bibRefCitation id="EFFF8D851444462BBC653762FE3AFEAD" author="Poorani, J." box="[301,503,286,310]" pageId="28" pageNumber="29" pagination="307 - 383" refId="ref118931" refString="Poorani, J. (2002 a) An annotated checklist of the Coccinellidae (excluding Epilachninae) of the Indian subregion. Oriental Insects, 36, 307 - 383. https: // doi. org / 10.1080 / 00305316.2002.10417335" type="journal article" year="2002">Poorani 2002a: 320</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BD1F0741444462BBDDF3741FE02FEEF" blockId="28.[151,1437,255,372]" pageId="28" pageNumber="29">
<treatmentCitationGroup id="AB7ED75A1444462BBDDF3741FE02FEEF" pageId="28" pageNumber="29">
<treatmentCitation id="0ACFD6651444462BBDDF3741FDA1FECE" author="Iablokoff-Khnzorian, S. M." box="[151,620,317,341]" page="80" pageId="28" pageNumber="29" year="1986">
<taxonomicName id="4C6E8BF71444462BBDDF3741FDA1FECE" authority="Iablokoff-Khnzorian, 1986: 80" authorityName="Iablokoff-Khnzorian" authorityPageNumber="80" authorityYear="1986" box="[151,620,317,341]" class="Insecta" family="Coccinellidae" genus="Adalia" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="puetzi">
<emphasis id="B91A2C661444462BBDDF3741FEEFFECE" box="[151,290,317,341]" italics="true" pageId="28" pageNumber="29">Adalia puetzi</emphasis>
<materialsCitation id="3B06FA291444462BBC653741FDA1FECE" ID-GBIF-Occurrence="4399052346" box="[301,620,317,341]" collectorName="NW Murree" country="Pakistan" location="Institute de Zoologie" pageId="28" pageNumber="29" specimenCount="1" typeStatus="holotype">
<bibRefCitation id="EFFF8D851444462BBC653741FDA1FECE" author="Iablokoff-Khnzorian, S. M." box="[301,620,317,341]" pageId="28" pageNumber="29" pagination="80" refId="ref114111" refString="Iablokoff-Khnzorian, S. M. (1986) Adalia puetzi n. sp., nouvelle espece du Pakistan (Col. Coccinellidae). Nouvelle Revue d'Entomologie, New Series, 3 (1), 80." type="journal article" year="1986">Iablokoff-Khnzorian, 1986: 80</bibRefCitation>
</materialsCitation>
</taxonomicName>
</treatmentCitation>
(
<typeStatus id="54D54ED61444462BBF363741FD13FECE" box="[638,734,317,341]" pageId="28" pageNumber="29" type="holotype">Holotype</typeStatus>
male,
<location id="8EB1A6AF1444462BBE633741FBC7FECE" LSID="urn:lsid:plazi:treatment:03C741621444462BBDDF36EBFD8AFC67:8EB1A6AF1444462BBE633741FBC7FECE" box="[811,1034,317,341]" country="Pakistan" name="Institute de Zoologie" pageId="28" pageNumber="29">Institute de Zoologie</location>
, Yerevan; Type locality:
<collectorName id="269B95A21444462BB85F3741FA55FECF" box="[1303,1432,317,340]" pageId="28" pageNumber="29">NW Murree</collectorName>
,
<collectingCountry id="F379B0E41444462BBD8F3720FEEFFEEF" box="[199,290,348,372]" name="Pakistan" pageId="28" pageNumber="29">Pakistan</collectingCountry>
).
<emphasis id="B91A2C661444462BBC7A3720FE02FEEF" bold="true" box="[306,463,348,372]" pageId="28" pageNumber="29">New synonym.</emphasis>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C374A3FF1444462BBDDF37DDFB3CFD0F" pageId="28" pageNumber="29" type="diagnosis">
<paragraph id="8BD1F0741444462BBDDF37DDFB3CFD0F" blockId="28.[151,1437,417,1020]" pageId="28" pageNumber="29">
<emphasis id="B91A2C661444462BBDDF37DDFEDFFE20" bold="true" box="[151,274,417,443]" pageId="28" pageNumber="29">Diagnosis.</emphasis>
Length:
<quantity id="4C965D911444462BBC3437DEFDE8FE27" box="[380,549,418,444]" metricMagnitude="-3" metricUnit="m" metricValue="4.05" metricValueMax="4.3" metricValueMin="3.8" pageId="28" pageNumber="29" unit="mm" value="4.05" valueMax="4.3" valueMin="3.8">3.80–4.30 mm</quantity>
; width:
<quantity id="4C965D911444462BBFCC37DEFCE0FE27" box="[644,813,418,444]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" metricValueMax="2.8" metricValueMin="2.6" pageId="28" pageNumber="29" unit="mm" value="2.7" valueMax="2.8" valueMin="2.6">2.60–2.80 mm</quantity>
. Form (
<figureCitation id="1355ECF11444462BBEC137DEFC24FE27" box="[905,1001,418,444]" captionStart="FIGURE 14" captionStartId="27.[151,250,1840,1864]" captionTargetBox="[166,1418,674,1808]" captionTargetId="figure-219@27.[151,1436,669,1816]" captionTargetPageId="27" captionText="FIGURE 14. Adalia simmondsi Kapur &amp; Sudha Rao: a. adult, dorsal view; b. spermatheca; c–e. male genitalia: c. tegmen, lateral view; d. tegmen, ventral view; e. penis." figureDoi="http://doi.org/10.5281/zenodo.8261548" httpUri="https://zenodo.org/record/8261548/files/figure.png" pageId="28" pageNumber="29">Fig. 14a</figureCitation>
) elongate oval, weakly convex. Head yellowish testaceous, with a black marking covering most of middle and lateral sides. Pronotum pale yellowish to ochreous, with at least 5 and a maximum of 11 black maculae of varying size and shape, sometimes fused to form larger markings. Scutellar shield light yellowish brown or with darker borders. Elytra uniformly pale ochreous to yellowish brown, elytral punctation somewhat coarse, clearly dual and dense. Ventral side more or less dark pitchy brown, except outer half of prothoracic hypomeron, elytral epipleura and posterior abdominal ventrites pale ochreous to yellowish brown. Male genitalia (
<figureCitation id="1355ECF11444462BBFD23406FCDEFD0F" box="[666,787,634,660]" captionStart="FIGURE 14" captionStartId="27.[151,250,1840,1864]" captionTargetBox="[166,1418,674,1808]" captionTargetId="figure-219@27.[151,1436,669,1816]" captionTargetPageId="27" captionText="FIGURE 14. Adalia simmondsi Kapur &amp; Sudha Rao: a. adult, dorsal view; b. spermatheca; c–e. male genitalia: c. tegmen, lateral view; d. tegmen, ventral view; e. penis." figureDoi="http://doi.org/10.5281/zenodo.8261548" httpUri="https://zenodo.org/record/8261548/files/figure.png" pageId="28" pageNumber="29">Fig. 14c–e</figureCitation>
) and spermatheca (
<figureCitation id="1355ECF11444462BBEA43406FB86FD0F" box="[1004,1099,634,660]" captionStart="FIGURE 14" captionStartId="27.[151,250,1840,1864]" captionTargetBox="[166,1418,674,1808]" captionTargetId="figure-219@27.[151,1436,669,1816]" captionTargetPageId="27" captionText="FIGURE 14. Adalia simmondsi Kapur &amp; Sudha Rao: a. adult, dorsal view; b. spermatheca; c–e. male genitalia: c. tegmen, lateral view; d. tegmen, ventral view; e. penis." figureDoi="http://doi.org/10.5281/zenodo.8261548" httpUri="https://zenodo.org/record/8261548/files/figure.png" pageId="28" pageNumber="29">Fig. 14b</figureCitation>
) as illustrated.
</paragraph>
</subSubSection>
<subSubSection id="C374A3FF1444462BBD8F34E1FE8CFD47" pageId="28" pageNumber="29" type="distribution">
<paragraph id="8BD1F0741444462BBD8F34E1FE8CFD47" blockId="28.[151,1437,417,1020]" pageId="28" pageNumber="29">
<emphasis id="B91A2C661444462BBD8F34E1FE94FD2C" bold="true" box="[199,345,669,695]" pageId="28" pageNumber="29">Distribution</emphasis>
:
<collectingCountry id="F379B0E41444462BBC2134E2FE6AFD23" box="[361,423,670,696]" name="India" pageId="28" pageNumber="29">India</collectingCountry>
: Northwestern Himalayas (
<collectingRegion id="49AA3E961444462BBF9F34E2FC66FD23" box="[727,939,670,696]" country="India" name="Himachal Pradesh" pageId="28" pageNumber="29">Himachal Pradesh</collectingRegion>
). It appears to be endemic to this region and extremely rare.
</paragraph>
</subSubSection>
<subSubSection id="C374A3FF1444462BBD8F3499FA95FCBF" pageId="28" pageNumber="29" type="biology_ecology">
<paragraph id="8BD1F0741444462BBD8F3499FBC2FC9B" blockId="28.[151,1437,417,1020]" box="[199,1039,741,768]" pageId="28" pageNumber="29">
<emphasis id="B91A2C661444462BBD8F3499FE2EFD64" bold="true" box="[199,483,741,767]" pageId="28" pageNumber="29">Prey/associated habitat.</emphasis>
<taxonomicName id="4C6E8BF71444462BBCA1349AFDACFC9B" box="[489,609,742,768]" class="Insecta" family="Adelgidae" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="family">Adelgidae</taxonomicName>
:
<taxonomicName id="4C6E8BF71444462BBF23349AFD34FC9B" box="[619,761,742,768]" class="Insecta" family="Adelgidae" genus="Adelges" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B91A2C661444462BBF23349AFD09FC9B" box="[619,708,742,768]" italics="true" pageId="28" pageNumber="29">Adelges</emphasis>
spp.
</taxonomicName>
on silver fir (label data).
</paragraph>
<paragraph id="8BD1F0741444462BBD8F3575FA95FCBF" blockId="28.[151,1437,417,1020]" box="[199,1368,777,804]" pageId="28" pageNumber="29">
<emphasis id="B91A2C661444462BBD8F3575FE0CFCBF" bold="true" box="[199,449,777,804]" pageId="28" pageNumber="29">Seasonal occurrence.</emphasis>
Collected during July, November, and January, in northwestern
<collectingCountry id="F379B0E41444462BB9D93576FB06FCBF" box="[1169,1227,778,804]" name="India" pageId="28" pageNumber="29">India</collectingCountry>
(label data).
</paragraph>
</subSubSection>
<subSubSection id="C374A3FF1444462BBD8F3552FD8AFC67" pageId="28" pageNumber="29" type="discussion">
<paragraph id="8BD1F0741444462BBD8F3552FD8AFC67" blockId="28.[151,1437,417,1020]" pageId="28" pageNumber="29">
<emphasis id="B91A2C661444462BBD8F3552FEC7FCD3" bold="true" box="[199,266,814,840]" pageId="28" pageNumber="29">Notes</emphasis>
.
<taxonomicName id="4C6E8BF71444462BBC513552FD53FCD3" authority="Iablokoff-Khnzorian" authorityName="Iablokoff-Khnzorian" authorityYear="1986" box="[281,670,814,840]" class="Insecta" family="Coccinellidae" genus="Adalia" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="puetzi">
<emphasis id="B91A2C661444462BBC513552FE62FCD3" box="[281,431,814,840]" italics="true" pageId="28" pageNumber="29">Adalia puetzi</emphasis>
Iablokoff-Khnzorian
</taxonomicName>
(
<emphasis id="B91A2C661444462BBFF83552FC99FCD3" bold="true" box="[688,852,814,840]" pageId="28" pageNumber="29">new synonym</emphasis>
) is conspecific with this species and the specimens identified by Dr Roger Booth as
<taxonomicName id="4C6E8BF71444462BBF4C352EFDABFCF7" authorityName="Iablokoff-Khnzorian" authorityYear="1986" box="[516,614,850,876]" class="Insecta" family="Coccinellidae" genus="Adalia" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="puetzi">
<emphasis id="B91A2C661444462BBF4C352EFDABFCF7" box="[516,614,850,876]" italics="true" pageId="28" pageNumber="29">A. puetzi</emphasis>
</taxonomicName>
in the collections of BMNH (examined) match
<taxonomicName id="4C6E8BF71444462BB933352EFADFFCF7" box="[1147,1298,850,876]" class="Insecta" family="Coccinellidae" genus="Adalia" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="simmondsi">
<emphasis id="B91A2C661444462BB933352EFADFFCF7" box="[1147,1298,850,876]" italics="true" pageId="28" pageNumber="29">A. simmondsi</emphasis>
</taxonomicName>
and in both, elytral punctation is somewhat coarse, clearly dual and dense and the interspaces are not smooth and shiny as in
<taxonomicName id="4C6E8BF71444462BB8CC350AFEC2FC2F" authorityName=": Mulsant" authorityYear="1846" class="Insecta" family="Coccinellidae" genus="Adalia" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="bipunctata">
<emphasis id="B91A2C661444462BB8CC350AFEC2FC2F" italics="true" pageId="28" pageNumber="29">A. bipunctata</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6E8BF71444462BBC0035E6FE24FC2F" box="[328,489,922,948]" class="Insecta" family="Coccinellidae" genus="Adalia" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="tetraspilota">
<emphasis id="B91A2C661444462BBC0035E6FE24FC2F" box="[328,489,922,948]" italics="true" pageId="28" pageNumber="29">A. tetraspilota</emphasis>
</taxonomicName>
. In general,
<taxonomicName id="4C6E8BF71444462BBF3235E6FCDFFC2F" box="[634,786,922,948]" class="Insecta" family="Coccinellidae" genus="Adalia" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="simmondsi">
<emphasis id="B91A2C661444462BBF3235E6FCDFFC2F" box="[634,786,922,948]" italics="true" pageId="28" pageNumber="29">A. simmondsi</emphasis>
</taxonomicName>
appears to be smaller in size and museum specimens are dull yellow.
<bibRefCitation id="EFFF8D851444462BBC6835C2FE23FC42" author="Kovar, I." box="[288,494,958,985]" pageId="28" pageNumber="29" pagination="1807" refId="ref116365" refString="Kovar, I. (2007) Family Coccinellidae Latreille, 1807. In: L ˆ bl, I. &amp; Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 4. Apollo Books, Stenstrup, pp. 568 - 631." type="book chapter" year="2007">Kovář (2007: 605)</bibRefCitation>
wrongly listed it as a synonym of
<taxonomicName id="4C6E8BF71444462BBE2135C2FBACFC43" authority="(Hope)" baseAuthorityName="Hope" box="[873,1121,958,984]" class="Insecta" family="Coccinellidae" genus="Adalia" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="tetraspilota">
<emphasis id="B91A2C661444462BBE2135C2FBC4FC43" box="[873,1033,958,984]" italics="true" pageId="28" pageNumber="29">A. tetraspilota</emphasis>
(Hope)
</taxonomicName>
.
<bibRefCitation id="EFFF8D851444462BB92335C2FA51FC43" author="Kapur, A. P. &amp; Sudha Rao, V." box="[1131,1436,958,984]" pageId="28" pageNumber="29" pagination="47 - 51" refId="ref115835" refString="Kapur, A. P. &amp; Sudha Rao, V. (1962) A new species of ladybird beetle (Col., Coccinellidae) feeding on Adelges spp. (Hemiptera: Adelgidae) in North western India. Entomophaga, 7, 47 - 51. https: // doi. org / 10.1007 / BF 02375991" type="journal article" year="1962">Kapur &amp; Sudha Rao (1962)</bibRefCitation>
described it in detail with illustrations.
</paragraph>
</subSubSection>
</treatment>
</document>
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