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<document id="786784F1268A60793BFB3083382E6158" ID-CLB-Dataset="51205" ID-DOI="10.11646/zootaxa.4061.5.3" ID-GBIF-Dataset="8e4c370b-2a1d-4bc4-a714-16f9f3a1bd65" ID-ISSN="1175-5326" ID-Zenodo-Dep="264294" ID-ZooBank="6EEE4C42-D292-490A-9549-CB3428430920" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459680617424" checkinUser="plazi" docAuthor="Häckel, Martin" docDate="2016" docId="038F87EE01413017FF0BD276FB9CFCD0" docLanguage="en" docName="zootaxa.4061.5.3.pdf" docOrigin="Zootaxa 4061 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Craspedophorus stanleyi Alluaud 1930" docType="treatment" docVersion="10" lastPageNumber="521" masterDocId="FFB6FF9601513006FF9CD433FFDDFFEB" masterDocTitle="A contribution to the knowledge of the subfamily Panagaeinae Hope, 1838 from Africa. Part 1. Revision of the Craspedophorus reflexus group (Coleoptera: Carabidae)" masterLastPageNumber="526" masterPageNumber="504" pageNumber="520" updateTime="1698207076058" updateUser="ExternalLinkService">
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<mods:title id="B4E362DC1CA87DB92485DE96AA6A4C0C">A contribution to the knowledge of the subfamily Panagaeinae Hope, 1838 from Africa. Part 1. Revision of the Craspedophorus reflexus group (Coleoptera: Carabidae)</mods:title>
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<mods:namePart id="4009CD75AC93F269DC01DA3811C2B6E9">Häckel, Martin</mods:namePart>
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<treatment id="038F87EE01413017FF0BD276FB9CFCD0" ID-DOI="http://doi.org/10.5281/zenodo.6088145" ID-GBIF-Taxon="119333006" ID-Zenodo-Dep="6088145" LSID="urn:lsid:plazi:treatment:038F87EE01413017FF0BD276FB9CFCD0" httpUri="http://treatment.plazi.org/id/038F87EE01413017FF0BD276FB9CFCD0" lastPageId="17" lastPageNumber="521" pageId="16" pageNumber="520">
<subSubSection id="C33C657301413016FF0BD276FD7EF9B4" box="[151,675,1605,1631]" pageId="16" pageNumber="520" type="nomenclature">
<paragraph id="8B9936F801413016FF0BD276FD7EF9B4" blockId="16.[151,675,1605,1665]" box="[151,675,1605,1631]" pageId="16" pageNumber="520">
<heading id="D0D1819401413016FF0BD276FD7EF9B4" bold="true" box="[151,675,1605,1631]" fontSize="11" level="1" pageId="16" pageNumber="520" reason="1">
<emphasis id="B952EAEA01413016FF0BD276FD7EF9B4" bold="true" box="[151,675,1605,1631]" pageId="16" pageNumber="520">
12.
<taxonomicName id="4C264D7B01413016FF5FD276FD7EF9B4" ID-CoL="Z5ZF" authority="Alluaud, 1930" authorityName="Alluaud" authorityYear="1930" box="[195,675,1605,1631]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="520" phylum="Arthropoda" rank="species" species="stanleyi">
<emphasis id="B952EAEA01413016FF5FD276FE31F9B4" bold="true" box="[195,492,1605,1631]" italics="true" pageId="16" pageNumber="520">Craspedophorus stanleyi</emphasis>
<bibRefCitation id="EFB74B0901413016FE68D276FD7EF9B4" author="Alluaud" box="[500,675,1605,1631]" pageId="16" pageNumber="525" refString="Alluaud, C. (1930) Note sur les Panageides dAfrique tropicale. Afra, Cahiers d'Entomologie, 1, 1 - 9." type="journal article" year="1930">Alluaud, 1930</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33C657301413016FF0BD25BFDE7F96B" box="[151,570,1640,1665]" pageId="16" pageNumber="520" type="description">
<paragraph id="8B9936F801413016FF0BD25BFDE7F96B" blockId="16.[151,675,1605,1665]" box="[151,570,1640,1665]" pageId="16" pageNumber="520">(Plate 3, Figs 21, 22, Plate 4, Fig. 32)</paragraph>
</subSubSection>
<subSubSection id="C33C657301413016FF0BD283FD75F90F" pageId="16" pageNumber="520" type="reference_group">
<paragraph id="8B9936F801413016FF0BD283FD75F90F" blockId="16.[151,1437,1712,1764]" pageId="16" pageNumber="520">
<treatmentCitationGroup id="AB3611D601413016FF0BD283FD75F90F" pageId="16" pageNumber="520">
<taxonomicName id="4C264D7B01413016FF0BD283FD8EF92D" ID-CoL="Z5ZF" authority="Alluaud, 1930: 5" authorityName="Alluaud" authorityPageNumber="5" authorityYear="1930" box="[151,595,1712,1734]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="520" phylum="Arthropoda" rank="species" species="stanleyi">
<emphasis id="B952EAEA01413016FF0BD283FE45F92D" box="[151,408,1712,1734]" italics="true" pageId="16" pageNumber="520">Craspedophorus stanleyi</emphasis>
<treatmentCitation id="0A8710E901413016FE3DD283FD8EF92D" author="Alluaud" box="[417,595,1712,1734]" page="5" pageId="16" pageNumber="520" year="1930">
<bibRefCitation id="EFB74B0901413016FE3DD283FDE4F92D" author="Alluaud" box="[417,569,1712,1734]" pageId="16" pageNumber="525" refString="Alluaud, C. (1930) Note sur les Panageides dAfrique tropicale. Afra, Cahiers d'Entomologie, 1, 1 - 9." type="journal article" year="1930">Alluaud, 1930</bibRefCitation>
: 5
</treatmentCitation>
</taxonomicName>
(
<typeStatus id="549D885A01413016FDFFD283FD4DF92D" box="[611,656,1712,1734]" pageId="16" pageNumber="520">type</typeStatus>
loc. “Stanleyville” [=Democratic
<collectingCountry id="F331766801413016FC64D283FBE2F92D" box="[1016,1087,1712,1734]" name="Republic of the Congo" pageId="16" pageNumber="520">Congo</collectingCountry>
: Orientale Province: Kisangani]).
<treatmentCitation id="0A8710E901413016FF5BD2FCFE51F90F" author="Burgeon" box="[199,396,1742,1764]" page="181" pageId="16" pageNumber="520" year="1935">
<bibRefCitation id="EFB74B0901413016FF5BD2FCFE87F90F" author="Burgeon" box="[199,346,1742,1764]" pageId="16" pageNumber="526" refString="Burgeon, L. (1935) Catalogues Raisonnes de la Faune Entomologique Du Congo Belge. Zoologie, t. 3, fasc. 3. Coleopteres, Carabides (Prem. pt.). Annales Du Musee Du Congo Belge, 1935, 135 - 257." type="journal article" year="1935">Burgeon 1935</bibRefCitation>
: 181
</treatmentCitation>
,
<treatmentCitation id="0A8710E901413016FE0AD2FDFD79F90F" author="Hackel" box="[406,676,1742,1764]" page="83" pageId="16" pageNumber="520" year="2012">
<bibRefCitation id="EFB74B0901413016FE0AD2FDFDA2F90F" author="Hackel" box="[406,639,1742,1764]" pageId="16" pageNumber="526" refString="Hackel, M. &amp; Farkac, J. (2012) A check-list of the subfamily Panagaeinae Hope, 1838 of the World (Coleoptera: Carabidae). Studies and Reports, Taxonomical Series, 8 (1 - 2), 67 - 116." type="journal article" year="2012">Häckel et Farkač 2012</bibRefCitation>
: 83
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C33C657301413017FF0BD322FB8EFCFC" lastPageId="17" lastPageNumber="521" pageId="16" pageNumber="520" type="materials_examined">
<paragraph id="8B9936F801413016FF0BD322FE2CF899" blockId="16.[151,1437,1809,2014]" pageId="16" pageNumber="520">
<emphasis id="B952EAEA01413016FF0BD322FE99F8C1" bold="true" box="[151,324,1809,1834]" pageId="16" pageNumber="520">
<typeStatus id="549D885A01413016FF0BD322FF0FF8C1" box="[151,210,1809,1834]" pageId="16" pageNumber="520">Type</typeStatus>
material.
</emphasis>
<typeStatus id="549D885A01413016FED7D322FE6EF8C1" box="[331,435,1809,1834]" pageId="16" pageNumber="520" type="holotype">Holotype</typeStatus>
(♀): “Stanleyville [handwritten white label //
<taxonomicName id="4C264D7B01413016FC2BD322FBCDF8C1" box="[951,1040,1809,1834]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="525" phylum="Arthropoda" rank="species" species="stanleyi">
<emphasis id="B952EAEA01413016FC2BD322FBCDF8C1" box="[951,1040,1809,1834]" italics="true" pageId="16" pageNumber="520">Stanleyi</emphasis>
</taxonomicName>
/
<typeStatus id="549D885A01413016FBB9D321FBBDF8C1" box="[1061,1120,1810,1834]" pageId="16" pageNumber="520">Type</typeStatus>
. [handwritten in black ink] / Alluaud det. 1930 [printed in brown ink, combined with handwritten corrections in black ink]” (Plate 3, Fig. 22,
<collectionCode id="ED37AE3D01413016FF0BD369FF32F89A" LSID="urn:lsid:biocol.org:col:34988" box="[151,239,1882,1905]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="16" pageNumber="520">MNHN</collectionCode>
, Alluaud's Collection).
</paragraph>
<paragraph id="8B9936F801413016FF5BD34EFEFAF851" blockId="16.[151,1437,1809,2014]" pageId="16" pageNumber="520">
<emphasis id="B952EAEA01413016FF5BD34EFDA6F87D" bold="true" box="[199,635,1917,1942]" pageId="16" pageNumber="520">
Other material examined.
<collectingCountry id="F331766801413016FD9AD34EFDABF87D" box="[518,630,1917,1942]" name="Tanzania" pageId="16" pageNumber="520">Tanzania</collectingCountry>
.
</emphasis>
<specimenCount id="9D20FD7101413016FD1ED34DFD6DF87E" box="[642,688,1918,1942]" pageId="16" pageNumber="520" type="generic">1 ♂</specimenCount>
: “Africa or., Kidugala [Iringa Region], 63., n. sp.?“ (Plate 3, Fig. 21,
<collectionCode id="ED37AE3D01413016FF5DD391FEC1F851" LSID="urn:lsid:biocol.org:col:34988" box="[193,284,1954,1978]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="16" pageNumber="520">MNHN</collectionCode>
).
</paragraph>
<paragraph id="8B9936F801413016FF5BD3F6FA40F835" blockId="16.[151,1437,1809,2014]" box="[199,1437,1989,2014]" pageId="16" pageNumber="520">
<emphasis id="B952EAEA01413016FF5BD3F6FEDBF835" bold="true" box="[199,262,1989,2014]" pageId="16" pageNumber="520">Note.</emphasis>
The original description of this species was based on a broken specimen, labelled “Stanleyville“.
</paragraph>
<paragraph id="8B9936F801403017FF0BD4A4FCE7FD88" blockId="17.[151,1437,151,827]" pageId="17" pageNumber="521">
Description (in part, see
<bibRefCitation id="EFB74B0901403017FE5DD4A4FDB9FF5B" author="Alluaud" box="[449,612,151,176]" pageId="17" pageNumber="525" refString="Alluaud, C. (1930) Note sur les Panageides dAfrique tropicale. Afra, Cahiers d'Entomologie, 1, 1 - 9." type="journal article" year="1930">Alluaud 1930</bibRefCitation>
: 5). “Length
<quantity id="4CDE9B1D01403017FC94D4A4FCA9FF44" box="[776,884,151,176]" metricMagnitude="-2" metricUnit="m" metricValue="2.25" pageId="17" pageNumber="521" unit="mm" value="22.5">22.5 mm</quantity>
, width
<quantity id="4CDE9B1D01403017FC4ED4ABFBF4FF44" box="[978,1065,152,176]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="17" pageNumber="521" unit="mm" value="10.0">10 mm</quantity>
. Very special form, because of disproportion between maximum width of pronotum (
<quantity id="4CDE9B1D01403017FC9FD48FFC94FF3F" box="[771,841,188,213]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="17" pageNumber="521" unit="mm" value="6.0">6 mm</quantity>
) and elytra (
<quantity id="4CDE9B1D01403017FC42D48FFBE8FF3F" box="[990,1077,188,212]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="17" pageNumber="521" unit="mm" value="10.0">10 mm</quantity>
), widely ovoid, and because of configuration of elytral maculae, with spots interleaved with black intervals. Palps and antennae absent in the only specimen I have. Head small, with diminutive mandibles. Pronotum with lateral margins wide, elevated, disc coarsely punctured; hind angles rounded, with a small indentation at base level. Elytral striae very fairly punctured, intervals throughout densely punctured except maculae. Humeral macula comprised of three elongated spots on intervals 4, 6 and 8, preapical macula also comprised of three spots, each of different length, located on same intervals as in humeral macula, in addition with a small yellow spot on 7th interval; spot on 6th interval twice longer than spots on intervals 4 and 8. Ventrites without crenulation on posterior margins. Metepisterna twice longer than wide. Sternites more coarsely and less densely punctured near margins, more finely punctured at middle. Tarsi with ventraldiffuse brushes of setae. It fairly resembles
<taxonomicName id="4C264D7B01403017FD4CD630FBCCFDF0" authority="Fabricius, 1781" authorityName="Fabricius" authorityYear="1781" box="[720,1041,514,539]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="521" phylum="Arthropoda" rank="species" species="reflexus">
<emphasis id="B952EAEA01403017FD4CD630FC94FDF0" box="[720,841,514,539]" italics="true" pageId="17" pageNumber="521">C. reflexus</emphasis>
(
<bibRefCitation id="EFB74B0901403017FCC4D631FBD4FDF0" author="Fabricius" box="[856,1033,514,539]" pageId="17" pageNumber="526" refString="Fabricius, J. C. (1781) Species Insectorum inhibentes eorum differenties specificas, synonyma, auctorum, loca natalia, metamorphosim adiectis observationibus, descriptionibus. Tomus I. C. E. Bohn, Hamburg et Kilonii, vii + 522 pp." type="book" year="1781">Fabricius, 1781</bibRefCitation>
)
</taxonomicName>
by similarly shaped pronotum, but because of disproportion between diminutive head with pronotum and bloated back,
<taxonomicName id="4C264D7B01403017FBCDD614FB14FDD4" box="[1105,1225,550,575]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="521" phylum="Arthropoda" rank="species" species="stanleyi">
<emphasis id="B952EAEA01403017FBCDD614FB14FDD4" box="[1105,1225,550,575]" italics="true" pageId="17" pageNumber="521">C. stanleyi</emphasis>
</taxonomicName>
does not resemble any other species of the genus known to me” [from French].
</paragraph>
<paragraph id="8B9936F801403017FF5BD65DFB8EFCFC" blockId="17.[151,1437,151,827]" pageId="17" pageNumber="521">
This
<typeStatus id="549D885A01403017FE9CD643FEEDFD63" box="[256,304,624,648]" pageId="17" pageNumber="521">type</typeStatus>
could not be sexed with certainty. The shape of the preserved labial palps seems to suggest a female (Plate 3, Fig. 22). Alluauds collection (
<collectionCode id="ED37AE3D01403017FDECD6A0FD14FD40" LSID="urn:lsid:biocol.org:col:34988" box="[624,713,659,683]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="17" pageNumber="521">MNHN</collectionCode>
) contains a male specimen from Kidugala (Iringa,
<collectingCountry id="F331766801403017FABBD6A1FA4DFD40" box="[1319,1424,658,683]" name="Tanzania" pageId="17" pageNumber="521">Tanzania</collectingCountry>
), which I dissected and which in my opinion has the pronotal shape and elytral coloration approaching the (so far unknown) male of
<taxonomicName id="4C264D7B01403017FEF6D6E8FE3FFD18" box="[362,482,730,755]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="521" phylum="Arthropoda" rank="species" species="stanleyi">
<emphasis id="B952EAEA01403017FEF6D6E8FE3FFD18" box="[362,482,730,755]" italics="true" pageId="17" pageNumber="521">C. stanleyi</emphasis>
</taxonomicName>
(Plate 3, Fig. 21). Until a female becomes available from the same locality, or a male is found to match the
<typeStatus id="549D885A01403017FE16D733FE67FCF3" box="[394,442,768,792]" pageId="17" pageNumber="521">type</typeStatus>
female, I provisionally regard this specimen as
<taxonomicName id="4C264D7B01403017FC48D6CCFB96FCFC" box="[980,1099,766,791]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="521" phylum="Arthropoda" rank="species" species="stanleyi">
<emphasis id="B952EAEA01403017FC48D6CCFB96FCFC" box="[980,1099,766,791]" italics="true" pageId="17" pageNumber="521">C. stanleyi</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C33C657301403017FF5BD711FB9CFCD0" box="[199,1089,802,827]" pageId="17" pageNumber="521" type="distribution">
<paragraph id="8B9936F801403017FF5BD711FB9CFCD0" blockId="17.[151,1437,151,827]" box="[199,1089,802,827]" pageId="17" pageNumber="521">
<emphasis id="B952EAEA01403017FF5BD711FEBDFCD0" bold="true" box="[199,352,802,827]" pageId="17" pageNumber="521">Distribution.</emphasis>
Democratic
<collectingCountry id="F331766801403017FE6DD710FD9DFCD0" box="[497,576,803,827]" name="Republic of the Congo" pageId="17" pageNumber="521">Congo</collectingCountry>
: Orientale Province;
<collectingCountry id="F331766801403017FCB0D711FC48FCD0" box="[812,917,802,827]" name="Tanzania" pageId="17" pageNumber="521">Tanzania</collectingCountry>
: Iringa Region.
</paragraph>
</subSubSection>
</treatment>
</document>