154 lines
18 KiB
XML
154 lines
18 KiB
XML
<document id="7C35DCE2C640E062832EB72DFD7E22A7" ID-CLB-Dataset="3791" ID-DOI="10.11646/zootaxa.5060.1.1" ID-GBIF-Dataset="40f89bbe-7f7b-4abd-aa02-a45e410da232" ID-ISSN="1175-5326" ID-Zenodo-Dep="5606671" ID-ZooBank="14126B35-AEEA-40F5-A892-FE6A8D64E5D6" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1635406161957" checkinUser="plazi" docAuthor="Mikhaljova, Elena V." docDate="2021" docId="038F87D5FF9EFF8FFF6D2148FD609AA0" docLanguage="en" docName="zootaxa.5060.1.1.pdf" docOrigin="Zootaxa 5060 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Litovkia Mikhaljova 2021, gen. nov." docType="treatment" docVersion="7" lastPageNumber="19" masterDocId="FFB6FFADFF8FFF9DFFFA2217FF8C9A76" masterDocTitle="The millipede family Diplomaragnidae Attems, 1907 in the Asian part of Russia with the descriptions of two new genera and seven new species (Diplopoda, Chordeumatida)" masterLastPageNumber="44" masterPageNumber="1" pageNumber="18" updateTime="1699419728057" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="1EA409C8FAC3EB279201B4409D2F8AD3">The millipede family Diplomaragnidae Attems, 1907 in the Asian part of Russia with the descriptions of two new genera and seven new species (Diplopoda, Chordeumatida)</mods:title>
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<mods:namePart id="8145F73242C3AC4CFCBEA51C2F3EE05C">Mikhaljova, Elena V.</mods:namePart>
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<mods:title id="28D8EE339F9F47865A0048C079E81D30">Zootaxa</mods:title>
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<mods:part id="8D98C3C61B05215E096313F4F9781F76">
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<mods:date id="3F30DA5494A0D1529ECA26B16AD6BC73">2021</mods:date>
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<mods:number id="B73E1967CBFA1A20ECB03CA8F60AFB85">2021-10-28</mods:number>
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<mods:number id="0E6F7C405EECB02B747A487C17CAA153">5060</mods:number>
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<mods:identifier id="1D77A61102528219C9E6E4E2F357DCF8" type="DOI">10.11646/zootaxa.5060.1.1</mods:identifier>
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<treatment id="038F87D5FF9EFF8FFF6D2148FD609AA0" ID-DOI="http://doi.org/10.5281/zenodo.5608349" ID-GBIF-Taxon="189548872" ID-Zenodo-Dep="5608349" LSID="urn:lsid:plazi:treatment:038F87D5FF9EFF8FFF6D2148FD609AA0" httpUri="http://treatment.plazi.org/id/038F87D5FF9EFF8FFF6D2148FD609AA0" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18">
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<subSubSection id="C33C6548FF9EFF8CFF6D2148FEFC990C" box="[151,368,863,890]" pageId="17" pageNumber="18" type="nomenclature">
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<paragraph id="8B9936C3FF9EFF8CFF6D2148FEFC990C" blockId="17.[151,368,863,890]" box="[151,368,863,890]" pageId="17" pageNumber="18">
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<heading id="D0D181AFFF9EFF8CFF6D2148FEFC990C" bold="true" box="[151,368,863,890]" fontSize="11" level="1" pageId="17" pageNumber="18" reason="1">
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<emphasis id="B952EAD1FF9EFF8CFF6D2148FEFC990C" bold="true" box="[151,368,863,890]" pageId="17" pageNumber="18">
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<taxonomicName id="4C264D40FF9EFF8CFF6D2148FF76990C" authority="Mikhaljova, 2021" authorityName="Mikhaljova" authorityYear="2021" box="[151,250,863,890]" class="Diplopoda" family="Diplomaragnidae" genus="Litovkia" kingdom="Animalia" order="Chordeumatida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B952EAD1FF9EFF8CFF6D2148FF76990C" bold="true" box="[151,250,863,890]" italics="true" pageId="17" pageNumber="18">Litovkia</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A26157AAFF9EFF8CFEF82177FEFC990C" box="[258,368,864,890]" pageId="17" pageNumber="18" rank="genus">gen. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C33C6548FF9EFF8CFF6D21B2FC2599AD" pageId="17" pageNumber="18" type="materials_examined">
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<paragraph id="8B9936C3FF9EFF8CFF6D21B2FDC699CB" blockId="17.[151,937,932,987]" box="[151,586,932,957]" pageId="17" pageNumber="18">
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<typeStatus id="549D8861FF9EFF8CFF6D21B2FF4599CB" box="[151,201,933,957]" pageId="17" pageNumber="18">Type</typeStatus>
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species:
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<taxonomicName id="4C264D40FF9EFF8CFEDC21B3FE7F99CA" authority="Mikhaljova, 2021" authorityName="Mikhaljova" authorityYear="2021" box="[294,499,932,956]" class="Diplopoda" family="Diplomaragnidae" genus="Litovkia" kingdom="Animalia" order="Chordeumatida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="anisimovka" status="sp. nov.">
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<emphasis id="B952EAD1FF9EFF8CFEDC21B3FE7F99CA" box="[294,499,932,956]" italics="true" pageId="17" pageNumber="18">Litovkia anisimovka</emphasis>
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</taxonomicName>
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<emphasis id="B952EAD1FF9EFF8CFE0321B2FDC699CB" bold="true" box="[505,586,933,957]" pageId="17" pageNumber="18">
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<taxonomicNameLabel id="A26157AAFF9EFF8CFE0321B2FDC699CB" box="[505,586,933,957]" pageId="17" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</paragraph>
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<paragraph id="8B9936C3FF9EFF8CFF6D21D4FC2599AD" blockId="17.[151,937,932,987]" box="[151,937,963,987]" pageId="17" pageNumber="18">
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Species included:
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<taxonomicName id="4C264D40FF9EFF8CFEAA21D4FD9099AD" authority="Mikhaljova, 2021" authorityName="Mikhaljova" authorityYear="2021" box="[336,540,963,987]" class="Diplopoda" family="Diplomaragnidae" genus="Litovkia" kingdom="Animalia" order="Chordeumatida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="anisimovka" status="sp. nov.">
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<emphasis id="B952EAD1FF9EFF8CFEAA21D4FD9099AD" box="[336,540,963,987]" italics="true" pageId="17" pageNumber="18">Litovkia anisimovka</emphasis>
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</taxonomicName>
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<emphasis id="B952EAD1FF9EFF8CFDD921D4FDFF99AD" bold="true" box="[547,627,963,987]" pageId="17" pageNumber="18">
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<taxonomicNameLabel id="A26157AAFF9EFF8CFDD921D4FDFF99AD" box="[547,627,963,987]" pageId="17" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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,
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<taxonomicName id="4C264D40FF9EFF8CFD7A21D4FC2599AD" authority="(Shear, 1990)" authorityName="Mikhaljova" authorityYear="2021" baseAuthorityName="Shear" baseAuthorityYear="1990" box="[640,937,963,987]" class="Diplopoda" family="Diplomaragnidae" genus="Litovkia" kingdom="Animalia" order="Chordeumatida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="lysaya">
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<emphasis id="B952EAD1FF9EFF8CFD7A21D4FC9799AD" box="[640,795,963,987]" italics="true" pageId="17" pageNumber="18">Litovkia lysaya</emphasis>
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(
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<bibRefCitation id="EFB74B32FF9EFF8CFCD221D4FC2E99AD" author="Shear, W. A." box="[808,930,963,987]" pageId="17" pageNumber="18" pagination="1 - 40" refId="ref33264" refString="Shear, W. A. (1990) On the Central and East Asian milliped family Diplomaragnidae (Diplopoda, Chordeumatida, Diplomaragnoidea). American Museum Novitates, 2977, 1 - 40." type="journal article" year="1990">Shear, 1990</bibRefCitation>
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)
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</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="C33C6548FF9EFF8CFF6D261EFE3C9CDD" pageId="17" pageNumber="18" type="diagnosis">
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<paragraph id="8B9936C3FF9EFF8CFF6D261EFB459EF9" blockId="17.[151,1437,1033,2032]" pageId="17" pageNumber="18">
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<emphasis id="B952EAD1FF9EFF8CFF6D261EFE9E9E55" bold="true" box="[151,274,1033,1059]" pageId="17" pageNumber="18">Diagnosis.</emphasis>
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The genus differs from other genera of the family mainly by the anterior angiocoxal processes of the posterior gonopods, which pass through lateral foramina in the colpocoxites, by the clavate femora of leg 4 with a depressed glabrous area, framed by an eyelashlike group of setae and by the distal bifurcation of the anterior angiocoxal processes. The structure of the male leg 4 is unique for the family
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<taxonomicName id="4C264D40FF9EFF8CFC042662FB489EF9" baseAuthorityName="Mikhaljova" baseAuthorityYear="2017" box="[1022,1220,1141,1167]" class="Diplopoda" family="Diplomaragnidae" kingdom="Animalia" order="Chordeumatida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="family">Diplomaragnidae</taxonomicName>
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.
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</paragraph>
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<paragraph id="8B9936C3FF9EFF8CFF3D268EFE509FA2" blockId="17.[151,1437,1033,2032]" pageId="17" pageNumber="18">
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The new genus seems more closely related to the East Asian genus
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<taxonomicName id="4C264D40FF9EFF8CFC4C268EFBD79EC5" authorityName="Attems" authorityYear="1907" box="[950,1115,1177,1203]" class="Diplopoda" family="Diplomaragnidae" genus="Diplomaragna" kingdom="Animalia" order="Chordeumatida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
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<emphasis id="B952EAD1FF9EFF8CFC4C268EFBD79EC5" box="[950,1115,1177,1203]" italics="true" pageId="17" pageNumber="18">Diplomaragna</emphasis>
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</taxonomicName>
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and the South Korean genus
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<taxonomicName id="4C264D40FF9EFF8CFF6D26A9FDB49EA1" authority="Mikhaljova and Lim, 2008" authorityName="Mikhaljova and Lim" authorityYear="2008" box="[151,568,1213,1239]" class="Diplopoda" family="Diplomaragnidae" genus="Koreagna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chordeumatida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
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<emphasis id="B952EAD1FF9EFF8CFF6D26A9FE8A9EA1" box="[151,262,1214,1239]" italics="true" pageId="17" pageNumber="18">Koreagna</emphasis>
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<bibRefCitation id="EFB74B32FF9EFF8CFEF626AAFDB49EA1" author="Mikhaljova, E. V. & Lim, K. - Y." box="[268,568,1213,1239]" pageId="17" pageNumber="18" pagination="51 - 61" refId="ref31153" refString="Mikhaljova, E. V. & Lim, K. - Y. (2008) The millipede family Diplomaragnidae Attems 1907 in the Korean Peninsula, with the descriptions of a new genus and new species (Diplopoda, Chordeumatida). Zootaxa, 1925, 51 - 61." type="journal article" year="2008">Mikhaljova and Lim, 2008</bibRefCitation>
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</taxonomicName>
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. The new genus resembles
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<taxonomicName id="4C264D40FF9EFF8CFC9826AAFB8B9EA1" authorityName="Attems" authorityYear="1907" box="[866,1031,1213,1239]" class="Diplopoda" family="Diplomaragnidae" genus="Diplomaragna" kingdom="Animalia" order="Chordeumatida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
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<emphasis id="B952EAD1FF9EFF8CFC9826AAFB8B9EA1" box="[866,1031,1213,1239]" italics="true" pageId="17" pageNumber="18">Diplomaragna</emphasis>
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</taxonomicName>
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in having both angiocoxal processes (anterior and posterior) of which the anterior one is long, passing through colpocoxites. However, the new genus differs from it by the anterior angiocoxal processes, which pass through lateral foramina in colpocoxites and by the bifurcation of the distal part of these processes (in contrast to the non-bifurcated anterior angiocoxal processes, passing between the colpocoxites along the midline in
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<taxonomicName id="4C264D40FF9EFF8CFCF1275AFC3C9F11" authorityName="Attems" authorityYear="1907" box="[779,944,1357,1383]" class="Diplopoda" family="Diplomaragnidae" genus="Diplomaragna" kingdom="Animalia" order="Chordeumatida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
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<emphasis id="B952EAD1FF9EFF8CFCF1275AFC3C9F11" box="[779,944,1357,1383]" italics="true" pageId="17" pageNumber="18">Diplomaragna</emphasis>
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</taxonomicName>
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), as well as by the clavate femora of leg 4 with a depressed glabrous area, framed by an eyelashlike group of setae and by the presence of small mesal sheath processes (in contrast to the absence of a depressed glabrous area on the leg 4 femora and the absence of any sheath processes in
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<taxonomicName id="4C264D40FF9EFF8CFEDC27AEFE479FA5" authorityName="Attems" authorityYear="1907" box="[294,459,1465,1491]" class="Diplopoda" family="Diplomaragnidae" genus="Diplomaragna" kingdom="Animalia" order="Chordeumatida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
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<emphasis id="B952EAD1FF9EFF8CFEDC27AEFE479FA5" box="[294,459,1465,1491]" italics="true" pageId="17" pageNumber="18">Diplomaragna</emphasis>
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</taxonomicName>
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).
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</paragraph>
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<paragraph id="8B9936C3FF9EFF8CFF3D27CAFE3C9CDD" blockId="17.[151,1437,1033,2032]" pageId="17" pageNumber="18">
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The new genus resembles
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<taxonomicName id="4C264D40FF9EFF8CFE0427C9FDE19F81" authorityName="Mikhaljova and Lim" authorityYear="2008" box="[510,621,1502,1527]" class="Diplopoda" family="Diplomaragnidae" genus="Koreagna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chordeumatida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
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<emphasis id="B952EAD1FF9EFF8CFE0427C9FDE19F81" box="[510,621,1502,1527]" italics="true" pageId="17" pageNumber="18">Koreagna</emphasis>
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</taxonomicName>
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mainly in the bifurcation of the distal part of the anterior angiocoxal process but differs by the presence of posterior angiocoxal processes and by the absence of lateral sheath processes of the colpocoxite, as well as by the absence of a depressed glabrous area on the leg 4 femora. It should also be noted that the anterior angiocoxite is completely divided into two branches (outer outgrowth and mesal long process piercing the colpocoxite or sheathed on its anterior face) in the species of the Japanese genus
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<taxonomicName id="4C264D40FF9EFF8CFB5E2479FE209CDD" authority="Miyosi, 1958" authorityName="Miyosi" authorityYear="1958" class="Diplopoda" family="Diplomaragnidae" genus="Pterygostegia" kingdom="Animalia" order="Chordeumatida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
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<emphasis id="B952EAD1FF9EFF8CFB5E2479FACC9CF1" box="[1188,1344,1646,1671]" italics="true" pageId="17" pageNumber="18">Pterygostegia</emphasis>
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Miyosi, 1958 (
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<bibRefCitation id="EFB74B32FF9EFF8CFF242486FE299CDD" author="Mikhaljova, E. V." box="[222,421,1681,1707]" pageId="17" pageNumber="18" pagination="153 - 181" refId="ref29738" refString="Mikhaljova, E. V. (2000 [for 1999]) Review of the millipede family Diplomaragnidae (Diplopoda: Chordeumatida). Arthropoda Selecta, 8 (3), 153 - 181." type="journal article" year="2000">Mikhaljova 2000</bibRefCitation>
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)
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C33C6548FF9EFF8CFF3D24A2FD719D99" pageId="17" pageNumber="18" type="description">
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<paragraph id="8B9936C3FF9EFF8CFF3D24A2FCC49D61" blockId="17.[151,1437,1033,2032]" pageId="17" pageNumber="18">
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<emphasis id="B952EAD1FF9EFF8CFF3D24A2FED59CB9" bold="true" box="[199,345,1717,1743]" pageId="17" pageNumber="18">Description.</emphasis>
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Body with 32 rings. Eyes present, in a triangular fields. Medium-sized paraterga well-developed, horizontal. Leg pairs 3–7 more or less enlarged. Legs 4 with femora clavate, depressed glabrous area, framed by eyelashlike group of setae. Legs 10 and 11 with coxal glands.
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</paragraph>
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<paragraph id="8B9936C3FF9EFF8CFF3D2536FD719D99" blockId="17.[151,1437,1033,2032]" pageId="17" pageNumber="18">Telopodites of anterior gonopods flagelliform, 1-segmented, each positioned on posterior face of posterior gonopod colpocoxites inside a narrow and deep sheath groove. Posterior gonopod colpocoxites fused medially basally or almost to midway. Each colpocoxite entire. Colpocoxite sheath groove with very small mesal processes. Posterior gonopod angiocoxite with a subglobule in posterior view. Anterior angiocoxal processes distally bifid, pass though lateral foramina in colpocoxites. Posterior angiocoxal processes present. Posterior gonopod telopodite 2-segmented, setose, distal segment of medium length.</paragraph>
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</subSubSection>
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<subSubSection id="C33C6548FF9DFF8FFF3D2280FB6A9AC7" box="[199,1254,151,177]" pageId="18" pageNumber="19" type="etymology">
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<paragraph id="8B9936C3FF9DFF8FFF3D2280FB6A9AC7" blockId="18.[199,1254,151,214]" box="[199,1254,151,177]" pageId="18" pageNumber="19">
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<emphasis id="B952EAD1FF9DFF8FFF3D2280FEC29AC7" bold="true" box="[199,334,151,177]" pageId="18" pageNumber="19">Etymology.</emphasis>
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The generic name refers to the
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<typeStatus id="549D8861FF9DFF8FFD4B2280FD6D9AC7" box="[689,737,151,177]" pageId="18" pageNumber="19">type</typeStatus>
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locality, Mountain Litovka. Gender feminine.
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</paragraph>
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</subSubSection>
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<subSubSection id="C33C6548FF9DFF8FFF3D22ADFD609AA0" box="[199,748,186,214]" pageId="18" pageNumber="19" type="distribution">
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||
<paragraph id="8B9936C3FF9DFF8FFF3D22ADFD609AA0" blockId="18.[199,1254,151,214]" box="[199,748,186,214]" pageId="18" pageNumber="19">
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<emphasis id="B952EAD1FF9DFF8FFF3D22ADFEED9AA2" bold="true" box="[199,353,186,212]" pageId="18" pageNumber="19">Distribution.</emphasis>
|
||
<collectingCountry id="F3317653FF9DFF8FFE9D22ACFE3A9AA3" box="[359,438,187,213]" name="Russia" pageId="18" pageNumber="19">Russia</collectingCountry>
|
||
: Far East (
|
||
<collectingRegion id="49E2F821FF9DFF8FFDD622ACFD539AA0" box="[556,735,187,214]" country="Russia" name="Primorskiy" pageId="18" pageNumber="19">Primorsky Krai</collectingRegion>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |