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<document id="7D02BFB72CD67F5028CC031D9067B6D3" ID-DOI="10.5281/zenodo.5041813" ID-ISSN="1942-1354" ID-Zenodo-Dep="5041813" ID-ZooBank="DF505A18-63A1-44BB-BF5D-13887FAE0DAD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1624979060254" checkinUser="carolina" docAuthor="Larochelle &amp; Larivière, Marie-Claude" docDate="2021" docId="038F87CF8F304874DFED16E8A02E3FEA" docLanguage="en" docName="InsectaMundi.0864.1-96.pdf" docOrigin="Insecta Mundi 2021 (864)" docStyle="DocumentStyle:F9976E9B7F8A094D890004515A7A0592.3:InsectaMundi.2020-.journal_article" docStyleId="F9976E9B7F8A094D890004515A7A0592" docStyleName="InsectaMundi.2020-.journal_article" docStyleVersion="3" docTitle="Ctenognathus cardiophorus" docType="treatment" docVersion="3" lastPageNumber="39" masterDocId="FFB6FFB78F17485CDF2D100BA17B386D" masterDocTitle="Synopsis of the tribe Platynini in New Zealand (Coleoptera: Carabidae)" masterLastPageNumber="96" masterPageNumber="1" pageNumber="38" updateTime="1698978488161" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
<mods:mods id="1AF12F0CC157A8CE76C59A59FEFDBEEF" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="DC0E50173F229619A527E87D6D62D5D3">
<mods:title id="BF79C1B9B7830E5E2AEE580772B4C117">Synopsis of the tribe Platynini in New Zealand (Coleoptera: Carabidae)</mods:title>
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<mods:name id="EC4DBDC10F01DB986A6876461596BDEF" type="personal">
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<mods:roleTerm id="059487B34DFA6D28A087829910C24A3D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7D608239EE0248443C0849B399E97BCA">Larochelle</mods:namePart>
<mods:affiliation id="34874F09B5B1C8365E6723FA53F3A6F2">Collection, Manaaki Whenua-Landcare Prosphodrus waimana Larochelle</mods:affiliation>
</mods:name>
<mods:name id="785437C22CC73325011767650E99F0CD" type="personal">
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</mods:role>
<mods:namePart id="13A2A6070D2258088392DAF2BD6F4E42">Larivière, Marie-Claude</mods:namePart>
<mods:affiliation id="453EDAC0E77F9296CED71D03F61A0208">- Landcare Research, Private Bag 92170, Auckland Larochelle and Larivière, new species</mods:affiliation>
</mods:name>
<mods:typeOfResource id="15E492888B19962F0F1AE913A3A1948B">text</mods:typeOfResource>
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<mods:title id="3B0F0669263EA509F84BC84AE3065986">Insecta Mundi</mods:title>
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<mods:part id="964BB2EB15C89F03E4EA1E884B2438D7">
<mods:date id="4AB2B99D23BCDAE4DFEA5B09ECC11C1C">2021</mods:date>
<mods:detail id="7E48F30C1891844D06E52A6A59C2814E" type="pubDate">
<mods:number id="4252A8854817EFCA5865DBE387D4DB8D">2021-04-30</mods:number>
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<mods:detail id="AD596A745ABE63146B97B2BB13B5BE21" type="volume">
<mods:number id="8C487A57F35847AF67AEA42EAB55DCE0">2021</mods:number>
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<mods:detail id="C8A48E8CE76BAD02EFAF2A205C98C626" type="issue">
<mods:number id="99377C7AA97CCF2DE276BD4AC3A3CED0">864</mods:number>
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<mods:start id="ECDDCBF88BDA317A11D07ABD2D1C224F">1</mods:start>
<mods:end id="DD1282BBDE72431019BF45CA5384E362">96</mods:end>
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<mods:classification id="441AF20830D7B01D515515B41E7ADD1C">journal article</mods:classification>
<mods:identifier id="B52F902CC46745F85CB35E604F83E658" type="DOI">10.5281/zenodo.5041813</mods:identifier>
<mods:identifier id="892A04C816F29565AAAC6AE90FB56F56" type="ISSN">1942-1354</mods:identifier>
<mods:identifier id="DD2E1AA9F163D4C98167E2B21770AD64" type="Zenodo-Dep">5041813</mods:identifier>
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<treatment id="038F87CF8F304874DFED16E8A02E3FEA" LSID="urn:lsid:plazi:treatment:038F87CF8F304874DFED16E8A02E3FEA" httpUri="http://treatment.plazi.org/id/038F87CF8F304874DFED16E8A02E3FEA" lastPageId="40" lastPageNumber="39" pageId="39" pageNumber="38">
<subSubSection id="C33C65528F30487BDFED16E8A3853F6D" box="[192,766,1763,1792]" pageId="39" pageNumber="38" type="nomenclature">
<paragraph id="8B9936D98F30487BDFED16E8A3853F6D" blockId="39.[192,766,1763,1826]" box="[192,766,1763,1792]" pageId="39" pageNumber="38">
<heading id="D0D181B58F30487BDFED16E8A3853F6D" box="[192,766,1763,1792]" fontSize="12" level="2" pageId="39" pageNumber="38" reason="2">
<taxonomicName id="4C264D5A8F30487BDFED16E8A3853F6D" ID-CoL="325S9" authority="(Chaudoir, 1878)" baseAuthorityName="Chaudoir" baseAuthorityYear="1878" box="[192,766,1763,1792]" class="Insecta" family="Carabidae" genus="Ctenognathus" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="38" phylum="Arthropoda" rank="species" species="cardiophorus">
<emphasis id="B952EACB8F30487BDFED16E8A3853F6D" bold="true" box="[192,766,1763,1792]" pageId="39" pageNumber="38">
<emphasis id="B952EACB8F30487BDFED16E8A3673F6D" bold="true" box="[192,540,1763,1792]" italics="true" pageId="39" pageNumber="38">Ctenognathus cardiophorus</emphasis>
(
<bibRefCitation id="EFB74B288F30487BDD0616E8A38F3F6D" author="Chaudoir" box="[555,756,1763,1792]" pageId="39" pageNumber="38" pagination="337 - 382" refId="ref54909" refString="Chaudoir M de. 1878 - 1879. Revision des genres Onychopterygia, Dicranoncus et Colpodes. Annales de la Societe Entomologique de France (5) 8: 275 - 336 (1878), 337 - 382 (1879)." type="journal article" year="1878">Chaudoir, 1878</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33C65528F30487BDFED170CA00E3F4C" box="[192,373,1799,1826]" pageId="39" pageNumber="38" type="description">
<paragraph id="8B9936D98F30487BDFED170CA00E3F4C" blockId="39.[192,766,1763,1826]" box="[192,373,1799,1826]" pageId="39" pageNumber="38">
<figureCitation id="131D2A5C8F30487BDFED170CA0763F4C" box="[192,269,1799,1825]" captionStart="Figures 57–60" captionStartId="75.[194,281,1866,1892]" captionTargetBox="[198,1431,256,1834]" captionTargetId="figure-41@75.[192,1439,232,1837]" captionTargetPageId="75" captionText="Figures 57–60. Dorsal habitus. 57) Ctenognathus garnerae new species. 58) C. tepaki new species. 59) C. urewera new species. 60) C. cardiophorus. Scale line = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5041845" httpUri="https://zenodo.org/record/5041845/files/figure.png" pageId="39" pageNumber="38">Fig. 60</figureCitation>
,
<figureCitation id="131D2A5C8F30487BDE351703A0393F4F" box="[280,322,1800,1826]" captionStart="Figures 101–107" captionStartId="84.[192,279,1866,1892]" captionTargetBox="[252,1384,254,1815]" captionTargetId="figure-54@84.[191,1440,208,1841]" captionTargetPageId="84" captionText="Figures 101–107. Aedeagus, lateral. 101) Ctenognathus tepaki new species. 102) C. urewera new species. 103) C. cardiophorus. 104) C. adamsi. 105) C. pictonensis. 106) C. tawanui new species. 107) C. perumalae new species." figureDoi="http://doi.org/10.5281/zenodo.5041881" httpUri="https://zenodo.org/record/5041881/files/figure.png" pageId="39" pageNumber="38">103</figureCitation>
,
<figureCitation id="131D2A5C8F30487BDE60170CA00E3F4C" box="[333,373,1799,1825]" captionStart="Figures 124–129" captionStartId="88.[192,279,1866,1892]" captionTargetBox="[206,1426,210,1836]" captionTargetId="figure-49@88.[206,1426,210,1836]" captionTargetPageId="88" captionText="Figures 124–129. Species distribution maps. 124) Ctenognathus adamsi. 125) C. arnaudensis. 126) C. cardiophorus. 127) C. colensonis. 128) C. davidsoni new species. 129) C. earlyi new species." figureDoi="http://doi.org/10.5281/zenodo.5041893" httpUri="https://zenodo.org/record/5041893/files/figure.png" pageId="39" pageNumber="38">126</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C33C65528F304874DFCD173BA3C13924" lastPageId="40" lastPageNumber="39" pageId="39" pageNumber="38" type="reference_group">
<paragraph id="8B9936D98F30487BDFCD173BA2613F0A" blockId="39.[224,1440,1840,1926]" pageId="39" pageNumber="38">
<treatmentCitationGroup id="AB3611F78F30487BDFCD173BA2613F0A" pageId="39" pageNumber="38">
<treatmentCitation id="0A8710C88F30487BDFCD173BA3EB3F25" author="Chaudoir" box="[224,656,1840,1864]" page="305" pageId="39" pageNumber="38" year="1878">
<taxonomicName id="4C264D5A8F30487BDFCD173BA3EB3F25" ID-CoL="5ZMBB" authority="Chaudoir, 1878: 305" authorityName="Chaudoir" authorityPageNumber="305" authorityYear="1878" box="[224,656,1840,1864]" class="Insecta" family="Carabidae" genus="Colpodes" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="38" phylum="Arthropoda" rank="species" species="cardiophorus">
<emphasis id="B952EACB8F30487BDFCD173BA0C43F25" box="[224,447,1840,1864]" italics="true" pageId="39" pageNumber="38">Colpodes cardiophorus</emphasis>
<materialsCitation id="3B4E3C848F30487BDEE9173BA3EB3F25" box="[452,656,1840,1864]" collectionCode="MNHN" pageId="39" pageNumber="38" specimenCount="1" typeStatus="holotype">
<bibRefCitation id="EFB74B288F30487BDEE9173BA3EB3F25" author="Chaudoir" box="[452,656,1840,1864]" pageId="39" pageNumber="38" pagination="337 - 382" refId="ref54909" refString="Chaudoir M de. 1878 - 1879. Revision des genres Onychopterygia, Dicranoncus et Colpodes. Annales de la Societe Entomologique de France (5) 8: 275 - 336 (1878), 337 - 382 (1879)." type="journal article" year="1878">Chaudoir, 1878: 305</bibRefCitation>
</materialsCitation>
</taxonomicName>
</treatmentCitation>
.
<typeStatus id="549D887B8F30487BDDBA173BA3833F25" box="[663,760,1840,1864]" pageId="39" pageNumber="38" type="holotype">Holotype</typeStatus>
: female (MNHN) labeled “ExMusaeo Chaudoir (typed) /
<taxonomicName id="4C264D5A8F30487BDA19173BA30A3F0A" ID-CoL="5ZMBB" authority="Chaud. Nouv. Zelande C. Laferte" class="Insecta" family="Carabidae" genus="Colpodes" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="38" phylum="Arthropoda" rank="species" species="cardiophorus">
<emphasis id="B952EACB8F30487BDA19173BA05B3F0A" italics="true" pageId="39" pageNumber="38">cardiophorus</emphasis>
Chaud. Nouv. Zélande C. Laferté
</taxonomicName>
(hand-written).”
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B9936D98F30487BDFCD1765A4E43FEB" blockId="39.[224,1440,1840,1926]" box="[224,1439,1902,1926]" pageId="39" pageNumber="38">
<treatmentCitationGroup id="AB3611F78F30487BDFCD1765A4E43FEB" box="[224,1439,1902,1926]" pageId="39" pageNumber="38">
<treatmentCitation id="0A8710C88F30487BDFCD1765A3243FEB" author="Broun T." box="[224,607,1902,1926]" page="20" pageId="39" pageNumber="38" year="1880">
<taxonomicName id="4C264D5A8F30487BDFCD1765A3243FEB" ID-CoL="8L2KH" authority="Broun, 1880: 20" authorityName="Broun" authorityPageNumber="20" authorityYear="1880" box="[224,607,1902,1926]" class="Insecta" family="Carabidae" genus="Anchomenus" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="38" phylum="Arthropoda" rank="species" species="parabilis">
<emphasis id="B952EACB8F30487BDFCD1765A0C13FEB" box="[224,442,1902,1926]" italics="true" pageId="39" pageNumber="38">Anchomenus parabilis</emphasis>
<materialsCitation id="3B4E3C848F30487BDE921765A3243FEB" box="[447,607,1902,1926]" collectionCode="BMNH" pageId="39" pageNumber="38" specimenCount="1" typeStatus="holotype">
<bibRefCitation id="EFB74B288F30487BDE921765A3243FEB" author="Broun T." box="[447,607,1902,1926]" pageId="39" pageNumber="38" pagination="1 - 651" refId="ref54484" refString="Broun T. 1880. Manual of the New Zealand Coleoptera. Part I. Government Printer; Wellington. XIX + 1 - 651 p. + VIII." type="book chapter" year="1880">Broun, 1880: 20</bibRefCitation>
</materialsCitation>
</taxonomicName>
</treatmentCitation>
.
<typeStatus id="549D887B8F30487BDD4A1765A3B33FEB" box="[615,712,1902,1926]" pageId="39" pageNumber="38" type="holotype">Holotype</typeStatus>
: “Whangarei Heads” [ND] (BMNH could not be located).
<emphasis id="B952EACB8F30487BDA261765A4E43FEB" bold="true" box="[1291,1439,1902,1926]" pageId="39" pageNumber="38">New synonym</emphasis>
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B9936D98F3F4874DFCD10DDA3063883" blockId="40.[224,698,214,329]" box="[224,637,214,238]" pageId="40" pageNumber="39">
<treatmentCitationGroup id="AB3611F78F3F4874DFCD10DDA3063883" box="[224,637,214,238]" pageId="40" pageNumber="39">
<treatmentCitation id="0A8710C88F3F4874DFCD10DDA3033883" author="Broun T." box="[224,632,214,238]" page="986" pageId="40" pageNumber="39" year="1893">
<taxonomicName id="4C264D5A8F3F4874DFCD10DDA3033883" ID-CoL="8LCPH" authority=": Broun 1893: 986" authorityName=": Broun" authorityPageNumber="986" authorityYear="1893" box="[224,632,214,238]" class="Insecta" family="Carabidae" genus="Ctenognathus" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="39" phylum="Arthropoda" rank="species" species="parabilis">
<emphasis id="B952EACB8F3F4874DFCD10DDA0BF3883" box="[224,452,214,238]" italics="true" pageId="40" pageNumber="39">Ctenognathus parabilis</emphasis>
:
<bibRefCitation id="EFB74B288F3F4874DEFD10DDA3033883" author="Broun T." box="[464,632,214,238]" pageId="40" pageNumber="39" pagination="975 - 1504" refId="ref54633" refString="Broun T. 1893. Manual of the New Zealand Coleoptera. Parts V - VII. Government Printer; Wellington. XVII + 975 - 1504 p." type="book chapter" year="1893">Broun 1893: 986</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B9936D98F3F4874DFCD10FFA3C93961" blockId="40.[224,698,214,329]" box="[224,690,244,268]" pageId="40" pageNumber="39">
<treatmentCitationGroup id="AB3611F78F3F4874DFCD10FFA3C93961" box="[224,690,244,268]" pageId="40" pageNumber="39">
<treatmentCitation id="0A8710C88F3F4874DFCD10FFA3D53961" author="Hutton FW" box="[224,686,244,268]" page="144" pageId="40" pageNumber="39" year="1904">
<taxonomicName id="4C264D5A8F3F4874DFCD10FFA3D53961" ID-CoL="325S9" authority=": Hutton 1904: 144" authorityName="Hutton" authorityPageNumber="144" authorityYear="1904" box="[224,686,244,268]" class="Insecta" family="Carabidae" genus="Ctenognathus" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="39" phylum="Arthropoda" rank="species" species="cardiophorus">
<emphasis id="B952EACB8F3F4874DFCD10FFA08B3961" box="[224,496,244,268]" italics="true" pageId="40" pageNumber="39">Ctenognathus cardiophorus</emphasis>
:
<bibRefCitation id="EFB74B288F3F4874DED110FEA3D53961" author="Hutton FW" box="[508,686,244,268]" pageId="40" pageNumber="39" refId="ref55393" refString="Hutton FW. 1904. Index Faunae Novae Zealandiae. Dulau; London. 372 p." type="book" year="1904">Hutton 1904: 144</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B9936D98F3F4874DFCD1118A3153946" blockId="40.[224,698,214,329]" box="[224,622,275,299]" pageId="40" pageNumber="39">
<treatmentCitationGroup id="AB3611F78F3F4874DFCD1118A3153946" box="[224,622,275,299]" pageId="40" pageNumber="39">
<treatmentCitation id="0A8710C88F3F4874DFCD1118A3123946" author="Csiki E." box="[224,617,275,299]" page="749" pageId="40" pageNumber="39" year="1931">
<taxonomicName id="4C264D5A8F3F4874DFCD1118A3123946" ID-CoL="5ZMBB" authority=": Csiki 1931: 749" authorityName="Csiki" authorityPageNumber="749" authorityYear="1931" box="[224,617,275,299]" class="Insecta" family="Carabidae" genus="Colpodes" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="39" phylum="Arthropoda" rank="species" species="cardiophorus">
<emphasis id="B952EACB8F3F4874DFCD1118A0BB3946" box="[224,448,275,299]" italics="true" pageId="40" pageNumber="39">Colpodes cardiophorus</emphasis>
:
<bibRefCitation id="EFB74B288F3F4874DEE11118A3123946" author="Csiki E." box="[460,617,275,299]" pageId="40" pageNumber="39" pagination="739 - 1022" refId="ref55080" refString="Csiki E. 1931. Carabidae: Harpalinae V. Pars 115. p. 739 - 1022. In: Junk W, Schenkling S (eds.). Coleopterorum Catalogus. Junk; Berlin. 1933 p." type="book chapter" year="1931">Csiki 1931: 749</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B9936D98F3F4874DFCD113AA3C13924" blockId="40.[224,698,214,329]" box="[224,698,305,329]" pageId="40" pageNumber="39">
<treatmentCitationGroup id="AB3611F78F3F4874DFCD113AA3C13924" box="[224,698,305,329]" pageId="40" pageNumber="39">
<treatmentCitation id="0A8710C88F3F4874DFCD113AA3CD3924" author="Hudson GV" box="[224,694,305,329]" page="174" pageId="40" pageNumber="39" year="1934">
<taxonomicName id="4C264D5A8F3F4874DFCD113AA3CD3924" ID-CoL="325S9" authority=": Hudson 1934: 174" authorityName=": Hudson" authorityPageNumber="174" authorityYear="1934" box="[224,694,305,329]" class="Insecta" family="Carabidae" genus="Ctenognathus" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="39" phylum="Arthropoda" rank="species" species="cardiophorus">
<emphasis id="B952EACB8F3F4874DFCD113AA08B3924" box="[224,496,305,329]" italics="true" pageId="40" pageNumber="39">Ctenognathus cardiophorus</emphasis>
:
<bibRefCitation id="EFB74B288F3F4874DED1113AA3CD3924" author="Hudson GV" box="[508,694,305,329]" pageId="40" pageNumber="39" refId="ref55357" refString="Hudson GV. 1934. New Zealand beetles and their larvae: an elementary introduction to the study of our native Coleoptera. Ferguson and Osborn; Wellington. 236 p. + 18 pl." type="book" year="1934">Hudson 1934: 174</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C33C65528F3F4874DFED116AA35C3CA5" pageId="40" pageNumber="39" type="description">
<paragraph id="8B9936D98F3F4874DFED116AA35C3CA5" blockId="40.[191,1441,353,1224]" pageId="40" pageNumber="39">
<emphasis id="B952EACB8F3F4874DFED116AA02E3916" bold="true" box="[192,341,353,379]" pageId="40" pageNumber="39">Description.</emphasis>
Body length
<quantity id="4CDE9B3C8F3F4874DEC1116AA3E93911" box="[492,658,353,380]" metricMagnitude="-2" metricUnit="m" metricValue="1.2550000000000001" metricValueMax="1.3800000000000001" metricValueMin="1.1300000000000001" pageId="40" pageNumber="39" unit="mm" value="12.55" valueMax="13.8" valueMin="11.3">11.3–13.8 mm</quantity>
. Head, pronotum, and elytra coal black; abdomen blackish; antennae and palpi reddish; femora rufopiceous; tibiae and tarsi reddish. Microsculpture very transverse (with microlines) on head, pronotum, and elytra. Iridescence absent. Moderately shiny. Forebody narrow in comparison to elytra.
<emphasis id="B952EACB8F3F4874DE1A11CCA006398C" bold="true" box="[311,381,455,481]" pageId="40" pageNumber="39">Head.</emphasis>
Narrow. Mandibles moderately long and curved anteriorly. Labrum strongly transverse, slightly emarginate anteriorly. Eyes moderately convex; two setiferous punctures on inner side of each eye. Tempora not inflated. Neck constriction shallow dorsolaterally. Mentum: medial tooth truncate apically; two setae. Submentum with four setae. Palpi with terminal segment obtuse apically.
<emphasis id="B952EACB8F3F4874DB441227A5BD3A2B" bold="true" box="[1129,1222,556,582]" pageId="40" pageNumber="39">Thorax.</emphasis>
Pronotum slightly convex, impunctate, deeply wrinkled on disc, narrow, strongly cordate, widest about middle; apex slightly emarginate; anterolateral angles well developed, angulate; anterior bead complete; sides strongly rounded, not sinuate posteriorly; lateral beads complete; lateral depressions wide, more so posteriorly; a single setiferous puncture on each side (anteriorly); posterolateral angles acute, projected laterally; laterobasal foveae unusually depressed, shallow, moderately wide, prolonged forward; posterior bead complete, less impressed near posterolateral angles; base emarginate.
<emphasis id="B952EACB8F3F4874DD5612FCA3C23B7C" bold="true" box="[635,697,759,785]" pageId="40" pageNumber="39">Legs.</emphasis>
Very long. Metafemora with two posteroventral setae. Meso- and metatarsomeres 1–4 well developed, tricarinate dorsally, deeply bisulcate laterally. Metatarsomeres 4 moderately bilobed and asymmetrical apically. Metatarsomeres 5 glabrous ventrally.
<emphasis id="B952EACB8F3F4874DB491331A5CE3B39" bold="true" box="[1124,1205,826,852]" pageId="40" pageNumber="39">Elytra.</emphasis>
Moderately convex, not sloping down toward apex, subovate, widest about middle. Shoulders moderately rounded. Sides strongly rounded. Scutellar setiferous pore invisible. Scutellar striole short, impunctate. Striae obsolete, impunctate. Intervals moderately convex; interval 3 without setiferous punctures. Umbilicate series with 18–21 setiferous punctures. Subapical sinuations moderately strong. Apices obtusely rounded.
<emphasis id="B952EACB8F3F4874DB6D13CAA5C63BB6" bold="true" box="[1088,1213,961,987]" pageId="40" pageNumber="39">Abdomen.</emphasis>
Sterna IV–VI: both sexes with two long apical ambulatory setae. Sternum VII (last visible sternum): male with two long apical ambulatory setae; female with eight long apical ambulatory setae.
<emphasis id="B952EACB8F3F4874DC9E140EA5563C72" bold="true" box="[947,1069,1029,1055]" pageId="40" pageNumber="39">Aedeagus.</emphasis>
Lateral view (
<figureCitation id="131D2A5C8F3F4874DBE3140EA4543C72" box="[1230,1327,1029,1055]" captionStart="Figures 101–107" captionStartId="84.[192,279,1866,1892]" captionTargetBox="[252,1384,254,1815]" captionTargetId="figure-54@84.[191,1440,208,1841]" captionTargetPageId="84" captionText="Figures 101–107. Aedeagus, lateral. 101) Ctenognathus tepaki new species. 102) C. urewera new species. 103) C. cardiophorus. 104) C. adamsi. 105) C. pictonensis. 106) C. tawanui new species. 107) C. perumalae new species." figureDoi="http://doi.org/10.5281/zenodo.5041881" httpUri="https://zenodo.org/record/5041881/files/figure.png" pageId="40" pageNumber="39">Fig. 103</figureCitation>
): strongly arcuate; base strongly convex, with basal lobe very wide; middle strongly convex dorsally, moderately concave ventrally, with dorsal membranous area biconvex, strongly widened in apical half and very long; apex moderately convex dorsally, rather straight ventrally, with extreme tip moderately wide and moderately long. Dorsal view: moderately wide, asymmetrical (ostium of membranous area deflected to the left); basal bulb moderately distant from membranous area.
</paragraph>
</subSubSection>
<subSubSection id="C33C65528F3F4874DFED14D4A2B83D46" pageId="40" pageNumber="39" type="materials_examined">
<paragraph id="8B9936D98F3F4874DFED14D4A5883C97" blockId="40.[192,1267,1247,1274]" box="[192,1267,1247,1274]" pageId="40" pageNumber="39">
<emphasis id="B952EACB8F3F4874DFED14D4A0D83C94" bold="true" box="[192,419,1247,1273]" pageId="40" pageNumber="39">Material examined.</emphasis>
<specimenCount id="9D20FD508F3F4874DE8414D4A3373C94" box="[425,588,1247,1274]" pageId="40" pageNumber="39" type="generic">519 specimens</specimenCount>
(
<collectionCode id="ED37AE1C8F3F4874DD7714EBA3CB3C97" box="[602,688,1248,1274]" country="New Zealand" httpUri="http://grbio.org/cool/76et-m1kt" name="Auckland Institute and Museum" pageId="40" pageNumber="39" type="Museum">AMNZ</collectionCode>
,
<collectionCode id="ED37AE1C8F3F4874DD9614EBA26A3C97" box="[699,785,1248,1274]" pageId="40" pageNumber="39">BMNH</collectionCode>
,
<collectionCode id="ED37AE1C8F3F4874DC3614EBA2083C97" box="[795,883,1248,1274]" country="USA" httpUri="http://grbio.org/cool/daw7-8h44" name="The Cleveland Museum of Natural History" pageId="40" pageNumber="39" type="Museum">CMNH</collectionCode>
,
<collectionCode id="ED37AE1C8F3F4874DC5314EBA2B23C97" box="[894,969,1248,1274]" country="New Zealand" httpUri="http://grbio.org/cool/n5zm-bhpt" name="Lincoln University Entomology Research Museum" pageId="40" pageNumber="39">LUNZ</collectionCode>
,
<collectionCode id="ED37AE1C8F3F4874DCFE14EBA5563C97" box="[979,1069,1248,1274]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="40" pageNumber="39">MNHN</collectionCode>
,
<collectionCode id="ED37AE1C8F3F4874DB1A14EBA5F73C97" box="[1079,1164,1248,1274]" country="New Zealand" httpUri="http://grbio.org/cool/15kb-kde6" name="Museum of New Zealand Te Papa Tongarewa - Entomology" pageId="40" pageNumber="39" type="Museum">MONZ</collectionCode>
,
<collectionCode id="ED37AE1C8F3F4874DBBA14EBA59C3C97" box="[1175,1255,1248,1274]" country="New Zealand" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34070" lsid="urn:lsid:biocol.org:col:34070" name="New Zealand Arthropod Collection" pageId="40" pageNumber="39" type="Other institutional type">NZAC</collectionCode>
).
</paragraph>
<paragraph id="8B9936D98F3F4874DFED151AA2B83D46" blockId="40.[192,963,1297,1324]" box="[192,963,1297,1324]" pageId="40" pageNumber="39">
<emphasis id="B952EACB8F3F4874DFED151AA0A73D46" bold="true" box="[192,476,1297,1323]" pageId="40" pageNumber="39">Geographic distribution</emphasis>
(
<figureCitation id="131D2A5C8F3F4874DEC6151AA3323D41" box="[491,585,1297,1324]" captionStart="Figures 124–129" captionStartId="88.[192,279,1866,1892]" captionTargetBox="[206,1426,210,1836]" captionTargetId="figure-49@88.[206,1426,210,1836]" captionTargetPageId="88" captionText="Figures 124–129. Species distribution maps. 124) Ctenognathus adamsi. 125) C. arnaudensis. 126) C. cardiophorus. 127) C. colensonis. 128) C. davidsoni new species. 129) C. earlyi new species." figureDoi="http://doi.org/10.5281/zenodo.5041893" httpUri="https://zenodo.org/record/5041893/files/figure.png" pageId="40" pageNumber="39">Fig. 126</figureCitation>
). North Island: AK, CL, ND, WO.
</paragraph>
</subSubSection>
<subSubSection id="C33C65528F3F4874DFED1548A5333E35" pageId="40" pageNumber="39" type="biology_ecology">
<paragraph id="8B9936D98F3F4874DFED1548A34C3DCC" blockId="40.[192,1441,1347,1441]" pageId="40" pageNumber="39">
<emphasis id="B952EACB8F3F4874DFED1548A0583D30" bold="true" box="[192,291,1347,1373]" pageId="40" pageNumber="39">Ecology.</emphasis>
Lowland. Epigean, fossorial. Wet forests (broadleaf, podocarp) and tree plantations (pine). Associated with streams and slopes. Shaded ground; wet, soaked soil. Nocturnal; hides during the day under logs, fallen branches, and stones. Gregarious.
</paragraph>
<paragraph id="8B9936D98F3F4874DFED15B3A2023D99" blockId="40.[192,1440,1464,1525]" pageId="40" pageNumber="39">
<emphasis id="B952EACB8F3F4874DFED15B3A05B3DBF" bold="true" box="[192,288,1464,1490]" pageId="40" pageNumber="39">Biology.</emphasis>
Seasonality: Throughout the year. Tenerals: November–January. Predacious (based on mouthpart morphology). Predators: Spiders. Occasionally infested with mites.
</paragraph>
<paragraph id="8B9936D98F3F4874DFED1607A2E83E4A" blockId="40.[192,915,1548,1575]" box="[192,915,1548,1575]" pageId="40" pageNumber="39">
<emphasis id="B952EACB8F3F4874DFED1607A0F83E4B" bold="true" box="[192,387,1548,1574]" pageId="40" pageNumber="39">Dispersal power.</emphasis>
Subapterous (incapable of flight). Swift runner.
</paragraph>
<paragraph id="8B9936D98F3F4874DFED1635A5333E35" blockId="40.[192,1096,1598,1624]" box="[192,1096,1598,1624]" pageId="40" pageNumber="39">
<emphasis id="B952EACB8F3F4874DFED1635A0BD3E35" bold="true" box="[192,454,1598,1624]" pageId="40" pageNumber="39">Collecting techniques.</emphasis>
Pitfall trapping; turning logs, fallen branches, and stones.
</paragraph>
</subSubSection>
<subSubSection id="C33C65528F3F4874DFED1664A25A3EA3" pageId="40" pageNumber="39" type="reference_group">
<paragraph id="8B9936D98F3F4874DFED1664A25A3EA3" blockId="40.[192,1440,1647,1742]" pageId="40" pageNumber="39">
<emphasis id="B952EACB8F3F4874DFED1664A03E3EE4" bold="true" box="[192,325,1647,1673]" pageId="40" pageNumber="39">References.</emphasis>
<bibRefCitation id="EFB74B288F3F4874DE66167BA3B53EE7" author="Larochelle A &amp; Lariviere M-C." box="[331,718,1648,1674]" pageId="40" pageNumber="39" pagination="1 - 281" refId="ref55539" refString="Larochelle A, Lariviere M-C. 2001. Carabidae (Insecta: Coleoptera): catalogue. Fauna of New Zealand 43: 1 - 281." type="journal article" year="2001">Larochelle and Larivière 2001: 134</bibRefCitation>
, 136 (as
<taxonomicName id="4C264D5A8F3F4874DC1D167BA5263EE7" ID-CoL="325S9" authorityName=": Hudson" authorityYear="1934" box="[816,1117,1648,1674]" class="Insecta" family="Carabidae" genus="Ctenognathus" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="39" phylum="Arthropoda" rank="species" species="cardiophorus">
<emphasis id="B952EACB8F3F4874DC1D167BA5263EE7" box="[816,1117,1648,1674]" italics="true" pageId="40" pageNumber="39">Ctenognathus cardiophorus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C264D5A8F3F4874DB47167BA59C3EE7" ID-CoL="8LCPH" authorityName=": Broun" authorityYear="1893" box="[1130,1255,1648,1674]" class="Insecta" family="Carabidae" genus="Ctenognathus" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="39" phylum="Arthropoda" rank="species" species="parabilis">
<emphasis id="B952EACB8F3F4874DB47167BA59C3EE7" box="[1130,1255,1648,1674]" italics="true" pageId="40" pageNumber="39">C. parabilis</emphasis>
</taxonomicName>
, catalogue; biology, dispersal power, ecology, geographic distribution, references), 2007: 111 (as
<taxonomicName id="4C264D5A8F3F4874DB771699A4703EC6" ID-CoL="325S9" authorityName=": Hudson" authorityYear="1934" box="[1114,1291,1681,1708]" class="Insecta" family="Carabidae" genus="Ctenognathus" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="39" phylum="Arthropoda" rank="species" species="cardiophorus">
<emphasis id="B952EACB8F3F4874DB771699A4703EC6" box="[1114,1291,1681,1708]" italics="true" pageId="40" pageNumber="39">C. cardiophorus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C264D5A8F3F4874DA371699A4E23EC6" ID-CoL="8LCPH" authorityName=": Broun" authorityYear="1893" box="[1306,1433,1681,1708]" class="Insecta" family="Carabidae" genus="Ctenognathus" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="39" phylum="Arthropoda" rank="species" species="parabilis">
<emphasis id="B952EACB8F3F4874DA371699A4E23EC6" box="[1306,1433,1681,1708]" italics="true" pageId="40" pageNumber="39">C. parabilis</emphasis>
</taxonomicName>
; list), 2016: 36–37 (as
<taxonomicName id="4C264D5A8F3F4874DE8616BFA3223EA0" ID-CoL="325S9" authorityName=": Hudson" authorityYear="1934" box="[427,601,1715,1742]" class="Insecta" family="Carabidae" genus="Ctenognathus" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="39" phylum="Arthropoda" rank="species" species="cardiophorus">
<emphasis id="B952EACB8F3F4874DE8616BFA3223EA0" box="[427,601,1715,1742]" italics="true" pageId="40" pageNumber="39">C. cardiophorus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C264D5A8F3F4874DD4B16BFA3983EA0" ID-CoL="8LCPH" authorityName=": Broun" authorityYear="1893" box="[614,739,1715,1742]" class="Insecta" family="Carabidae" genus="Ctenognathus" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="39" phylum="Arthropoda" rank="species" species="parabilis">
<emphasis id="B952EACB8F3F4874DD4B16BFA3983EA0" box="[614,739,1715,1742]" italics="true" pageId="40" pageNumber="39">C. parabilis</emphasis>
</taxonomicName>
; list).
</paragraph>
</subSubSection>
<subSubSection id="C33C65528F3F4874DFED16EEA02E3FEA" pageId="40" pageNumber="39" type="discussion">
<paragraph id="8B9936D98F3F4874DFED16EEA02E3FEA" blockId="40.[192,1442,1765,1927]" pageId="40" pageNumber="39">
<emphasis id="B952EACB8F3F4874DFED16EEA0543E92" bold="true" box="[192,303,1765,1791]" pageId="40" pageNumber="39">Remarks.</emphasis>
The
<typeStatus id="549D887B8F3F4874DE4516EEA0B13E92" box="[360,458,1765,1791]" pageId="40" pageNumber="39" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C264D5A8F3F4874DEDF16EEA39D3E92" authorityName="Broun" authorityYear="1880" box="[498,742,1765,1791]" class="Insecta" family="Carabidae" genus="Anchomenus" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="39" phylum="Arthropoda" rank="species" species="parabilis">
<emphasis id="B952EACB8F3F4874DEDF16EEA39D3E92" box="[498,742,1765,1791]" italics="true" pageId="40" pageNumber="39">Anchomenus parabilis</emphasis>
</taxonomicName>
could not be located in BMNH where Broun’s specimens can usually be found. However, there is no ambiguity regarding the identity of this taxon. Furthermore, a non-type male specimen from the Broun Collection (BMNH) was examined. This specimen is from Hunua-Wairoa [= Wairoa, Hunua Ranges] (AK) and its morphology corresponds to Broun’s original description of
<taxonomicName id="4C264D5A8F3F4874DA271741A0343FEB" authorityName=": Hudson" authorityYear="1934" class="Insecta" family="Carabidae" genus="Ctenognathus" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="39" phylum="Arthropoda" rank="species" species="cardiophorus">
<emphasis id="B952EACB8F3F4874DA271741A0343FEB" italics="true" pageId="40" pageNumber="39">Ctenognathus cardiophorus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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