treatments-xml/data/03/8F/87/038F87CF8F08487DDFED10DDA4E63CFB.xml

<document id="2EA7F3CB80E742F0051D8D0D0D7F15E7" ID-DOI="10.5281/zenodo.5041813" ID-ISSN="1942-1354" ID-Zenodo-Dep="5041813" ID-ZooBank="DF505A18-63A1-44BB-BF5D-13887FAE0DAD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1624979060254" checkinUser="carolina" docAuthor="Larochelle &amp; Larivière, Marie-Claude" docDate="2021" docId="038F87CF8F08487DDFED10DDA4E63CFB" docLanguage="en" docName="InsectaMundi.0864.1-96.pdf" docOrigin="Insecta Mundi 2021 (864)" docStyle="DocumentStyle:F9976E9B7F8A094D890004515A7A0592.3:InsectaMundi.2020-.journal_article" docStyleId="F9976E9B7F8A094D890004515A7A0592" docStyleName="InsectaMundi.2020-.journal_article" docStyleVersion="3" docTitle="Ctenognathus Fairmaire 1843" docType="key" docVersion="3" lastPageNumber="32" masterDocId="FFB6FFB78F17485CDF2D100BA17B386D" masterDocTitle="Synopsis of the tribe Platynini in New Zealand (Coleoptera: Carabidae)" masterLastPageNumber="96" masterPageNumber="1" pageNumber="30" updateTime="1698978488161" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
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<mods:title id="AE558D688CFCEB25F3C2793223A4B194">Synopsis of the tribe Platynini in New Zealand (Coleoptera: Carabidae)</mods:title>
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<mods:namePart id="016D08F0E6738C1BA6CD677B9F2CE962">Larochelle</mods:namePart>
<mods:affiliation id="F74314E3E28EF7A4B73543257C511200">Collection, Manaaki Whenua-Landcare Prosphodrus waimana Larochelle</mods:affiliation>
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<mods:namePart id="905177341670AC601FA389966DFB2D06">Larivière, Marie-Claude</mods:namePart>
<mods:affiliation id="A0C7CBCF26960A88F43962EC7210E9E3">- Landcare Research, Private Bag 92170, Auckland Larochelle and Larivière, new species</mods:affiliation>
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<mods:date id="178F8F3DE7EDD46D3CBE1E28F062E41A">2021</mods:date>
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Key to species of 
<taxonomicName id="4C264D5A8F084843DEB710DEA332389F" ID-CoL="8KWGZ" authorityName="Fairmaire" authorityYear="1843" box="[410,585,213,242]" class="Insecta" family="Carabidae" genus="Ctenognathus" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="30" phylum="Arthropoda" rank="genus">
<emphasis id="B952EACB8F084843DEB710DEA332389F" bold="true" box="[410,585,213,242]" italics="true" pageId="31" pageNumber="30">Ctenognathus</emphasis>
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1. Pronotum with a single setiferous puncture on each side (anteriorly)............................. 
<emphasis id="B952EACB8F084843DAA21109A4E63971" bold="true" box="[1423,1437,258,284]" pageId="31" pageNumber="30">2</emphasis>
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— Pronotum with two setiferous punctures on each side (anteriorly and posteriorly)................. 
<emphasis id="B952EACB8F084843DAA2112FA4E63953" bold="true" box="[1423,1437,292,318]" pageId="31" pageNumber="30">9</emphasis>
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<paragraph id="8B9936D98F084843DFED1145A4E63905" blockId="31.[192,1437,334,395]" box="[192,1437,334,361]" pageId="31" pageNumber="30">
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2(1). Meso- and metatarsomeres 1–4 (
<figureCitation id="131D2A5C8F084843DDA01145A3A73905" box="[653,732,334,360]" captionStart="Figures 27–36" captionStartId="69.[192,279,1765,1791]" captionTargetBox="[309,1386,262,1710]" captionTargetId="figure-85@69.[264,1406,242,1738]" captionTargetPageId="69" captionText="Figures 27–36. Meso- or metatarsomere, dorsal view: 27) not carinate; 28) tricarinate. Metatarsomere 4: 29) moderately bilobed and asymmetrical apically; 30) moderately bilobed and symmetrical; 31) strongly bilobed and asymmetrical. Abdominal sterna V–VI: 32) with 4 ambulatory setae; 33) with 2 ambulatory setae; 34) V–VII with numerous short setae; 35) VII, male with 2 ambulatory setae, female with 4 setae; 36) VII, male with 4 ambulatory setae, female with 8 setae." figureDoi="http://doi.org/10.5281/zenodo.5041827" httpUri="https://zenodo.org/record/5041827/files/figure.png" pageId="31" pageNumber="30">Fig. 27</figureCitation>
) not carinate dorsally................................... 
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— Meso- and metatarsomeres 1–4 (
<figureCitation id="131D2A5C8F084843DDA0117BA3A739E7" box="[653,732,368,394]" captionStart="Figures 27–36" captionStartId="69.[192,279,1765,1791]" captionTargetBox="[309,1386,262,1710]" captionTargetId="figure-85@69.[264,1406,242,1738]" captionTargetPageId="69" captionText="Figures 27–36. Meso- or metatarsomere, dorsal view: 27) not carinate; 28) tricarinate. Metatarsomere 4: 29) moderately bilobed and asymmetrical apically; 30) moderately bilobed and symmetrical; 31) strongly bilobed and asymmetrical. Abdominal sterna V–VI: 32) with 4 ambulatory setae; 33) with 2 ambulatory setae; 34) V–VII with numerous short setae; 35) VII, male with 2 ambulatory setae, female with 4 setae; 36) VII, male with 4 ambulatory setae, female with 8 setae." figureDoi="http://doi.org/10.5281/zenodo.5041827" httpUri="https://zenodo.org/record/5041827/files/figure.png" pageId="31" pageNumber="30">Fig. 28</figureCitation>
) tricarinate dorsally.................................... 
<emphasis id="B952EACB8F084843DAA2117BA4E639E7" bold="true" box="[1423,1437,368,394]" pageId="31" pageNumber="30">6</emphasis>
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3(2). Antennae entirely rufopiceous; abdomen black................................................ 
<emphasis id="B952EACB8F084843DAA21191A4E639D9" bold="true" box="[1423,1437,410,436]" pageId="31" pageNumber="30">4</emphasis>
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</paragraph>
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— Antennae bicolored; abdomen rufopiceous................................................... 
<emphasis id="B952EACB8F084843DAA211B6A4E639BA" bold="true" box="[1423,1437,445,471]" pageId="31" pageNumber="30">5</emphasis>
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4(3). Tempora not inflated; pronotum not sinuate posteriorly, sides moderately rounded, posterolateral angles obtusely rounded, base truncate, posterior bead present, complete; apices of elytra obtuse; 
<figureCitation id="131D2A5C8F084843DA611202A4E73A49" box="[1356,1436,521,548]" captionStart="Figures 53–56" captionStartId="74.[192,279,1866,1892]" captionTargetBox="[217,1433,320,1833]" captionTargetId="figure-45@74.[193,1440,280,1837]" captionTargetPageId="74" captionText="Figures 53–56. Dorsal habitus. 53) Tuiplatynus sophronitis, teneral. 54) T. libitus, teneral. 55) Ctenognathus novaezelandiae. 56) C. elevatus. Scale line = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5041841" httpUri="https://zenodo.org/record/5041841/files/figure.png" pageId="31" pageNumber="30">Fig. 55</figureCitation>
. [Body length 12.0– 
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; North Island (ND to TK–HB)]................................................................................. 
<taxonomicName id="4C264D5A8F084843DC951247A4E63A0A" ID-CoL="325SV" authority="(Fairmaire)" baseAuthorityName="Fairmaire" baseAuthorityYear="1843" box="[952,1437,588,615]" class="Insecta" family="Carabidae" genus="Ctenognathus" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="30" phylum="Arthropoda" rank="species" species="novaezelandiae">
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<emphasis id="B952EACB8F084843DC951247A46B3A0A" bold="true" box="[952,1296,588,615]" italics="true" pageId="31" pageNumber="30">Ctenognathus novaezelandiae</emphasis>
(Fairmaire)
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— Tempora moderately inflated (about two-thirds as long as eyes); pronotum moderately sinuate posteriorly, sides strongly rounded, posterolateral angles slightly acute, base convex, posterior bead absent; apices of elytra rounded; 
<figureCitation id="131D2A5C8F084843DD4512B8A3CD3AA3" box="[616,694,691,718]" captionStart="Figures 53–56" captionStartId="74.[192,279,1866,1892]" captionTargetBox="[217,1433,320,1833]" captionTargetId="figure-45@74.[193,1440,280,1837]" captionTargetPageId="74" captionText="Figures 53–56. Dorsal habitus. 53) Tuiplatynus sophronitis, teneral. 54) T. libitus, teneral. 55) Ctenognathus novaezelandiae. 56) C. elevatus. Scale line = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5041841" httpUri="https://zenodo.org/record/5041841/files/figure.png" pageId="31" pageNumber="30">Fig. 56</figureCitation>
. [Body length 12.0– 
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; southern North Island (WN)]............................................................... 
<taxonomicName id="4C264D5A8F084843DB1412DEA4E53A82" ID-CoL="8LCP4" authority="(White)" baseAuthorityName="White" baseAuthorityYear="1846" box="[1081,1438,725,752]" class="Insecta" family="Carabidae" genus="Ctenognathus" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="30" phylum="Arthropoda" rank="species" species="elevatus">
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<emphasis id="B952EACB8F084843DB1412DEA4403A9D" bold="true" box="[1081,1339,725,752]" italics="true" pageId="31" pageNumber="30">Ctenognathus elevatus</emphasis>
(White)
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5(3). Antennal segments 1–4 rufopiceous, 5–11 reddish; mandibles very long, slightly curved anteriorly; pronotum with lateral depressions narrow, slightly widened from apex to base, posterolateral angles obtuse; metatarsomeres 5 glabrous ventrally; 
<figureCitation id="131D2A5C8F084843DC70134FA2CB3B32" box="[861,944,836,863]" captionStart="Figures 57–60" captionStartId="75.[194,281,1866,1892]" captionTargetBox="[198,1431,256,1834]" captionTargetId="figure-41@75.[192,1439,232,1837]" captionTargetPageId="75" captionText="Figures 57–60. Dorsal habitus. 57) Ctenognathus garnerae new species. 58) C. tepaki new species. 59) C. urewera new species. 60) C. cardiophorus. Scale line = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5041845" httpUri="https://zenodo.org/record/5041845/files/figure.png" pageId="31" pageNumber="30">Fig. 57</figureCitation>
. [Body length 
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; Three Kings Islands (TH)].................. 
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<taxonomicName id="4C264D5A8F084843DDF7136DA4703BED" ID-CoL="8MN57" authority="Larochelle and Lariviere" authorityName="Larochelle and Lariviere E" authorityYear="2021" box="[730,1291,869,897]" class="Insecta" family="Carabidae" genus="Ctenognathus" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="30" phylum="Arthropoda" rank="species" species="garnerae" status="sp. nov.">
<emphasis id="B952EACB8F084843DDF7136DA29D3BED" bold="true" box="[730,998,870,897]" italics="true" pageId="31" pageNumber="30">Ctenognathus garnerae</emphasis>
Larochelle and Larivière
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, 
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— Antennal segments 1 rufopiceous, 2–11 reddish; mandibles moderately long and curved anteriorly; pronotum with lateral depressions wide throughout, posterolateral angles rectangular; metatarsomeres 5 setose ventrally (with long setae); 
<figureCitation id="131D2A5C8F084843DDDD13C7A23A3B8A" box="[752,833,972,999]" captionStart="Figures 57–60" captionStartId="75.[194,281,1866,1892]" captionTargetBox="[198,1431,256,1834]" captionTargetId="figure-41@75.[192,1439,232,1837]" captionTargetPageId="75" captionText="Figures 57–60. Dorsal habitus. 57) Ctenognathus garnerae new species. 58) C. tepaki new species. 59) C. urewera new species. 60) C. cardiophorus. Scale line = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5041845" httpUri="https://zenodo.org/record/5041845/files/figure.png" pageId="31" pageNumber="30">Fig. 58</figureCitation>
. [Body length 
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; northern North Island (ND, top of Aupouri Peninsula)].... 
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<taxonomicName id="4C264D5A8F084843DDD413E5A4703C64" ID-CoL="8MBX5" authority="Larochelle and Lariviere" authorityName="Larochelle and Lariviere E" authorityYear="2021" box="[761,1291,1006,1033]" class="Insecta" family="Carabidae" genus="Ctenognathus" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="30" phylum="Arthropoda" rank="species" species="tepaki" status="sp. nov.">
<emphasis id="B952EACB8F084843DDD413E5A29D3C64" bold="true" box="[761,998,1006,1033]" italics="true" pageId="31" pageNumber="30">Ctenognathus tepaki</emphasis>
Larochelle and Larivière
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, 
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6(2). Head, pronotum, and elytra coal black; femora rufopiceous; elytra without visible scutellar setiferous pore, microsculpture very transverse (with microlines); terminal segment of palpi obtuse apically (
<figureCitation id="131D2A5C8F084843DE771456A0D33C1A" box="[346,424,1117,1143]" captionStart="Figures 1–13" captionStartId="67.[192,279,1833,1859]" captionTargetBox="[247,1417,213,1793]" captionTargetId="figure-11@67.[207,1425,213,1806]" captionTargetPageId="67" captionText="Figures 1–13. Microsculpture: 1) granulate; 2) isodiametric; 3) moderately transverse; 4) very transverse. Neck constriction, dorsolateral view: 5) deep; 6) shallow; 7) absent. Eye and buccal fissure, lateral view: 8) touching; 9) separated. Terminal segment of palpi: 10) obtuse apically; 11) truncate. Paraglossae: 12) pubescent; 13) glabrous." figureDoi="http://doi.org/10.5281/zenodo.5041815" httpUri="https://zenodo.org/record/5041815/files/figure.png" pageId="31" pageNumber="30">Fig. 10</figureCitation>
)............................................................................ 
<emphasis id="B952EACB8F084843DAA21456A4E63C1A" bold="true" box="[1423,1437,1117,1143]" pageId="31" pageNumber="30">7</emphasis>
</keyLead>
</paragraph>
<paragraph id="8B9936D98F084843DFED148BA4E63CD6" blockId="31.[192,1441,1049,1212]" pageId="31" pageNumber="30">
<keyLead id="30D795EC8F084843DFED148BA4E63CD6" pageId="31" pageNumber="30">
— Head and pronotum pure black, elytra piceous black; femora reddish; elytra with visible scutellar setiferous pore, microsculpture isodiametric; terminal segment of palpi truncate apically (
<figureCitation id="131D2A5C8F084843DBDF14AAA43B3CD6" box="[1266,1344,1185,1211]" captionStart="Figures 1–13" captionStartId="67.[192,279,1833,1859]" captionTargetBox="[247,1417,213,1793]" captionTargetId="figure-11@67.[207,1425,213,1806]" captionTargetPageId="67" captionText="Figures 1–13. Microsculpture: 1) granulate; 2) isodiametric; 3) moderately transverse; 4) very transverse. Neck constriction, dorsolateral view: 5) deep; 6) shallow; 7) absent. Eye and buccal fissure, lateral view: 8) touching; 9) separated. Terminal segment of palpi: 10) obtuse apically; 11) truncate. Paraglossae: 12) pubescent; 13) glabrous." figureDoi="http://doi.org/10.5281/zenodo.5041815" httpUri="https://zenodo.org/record/5041815/files/figure.png" pageId="31" pageNumber="30">Fig. 11</figureCitation>
).... 
<emphasis id="B952EACB8F084843DAA214AAA4E63CD6" bold="true" box="[1423,1437,1185,1211]" pageId="31" pageNumber="30">8</emphasis>
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</paragraph>
</keyStep>
<keyStep id="30D22E7C8F084843DFED14C0A4E53DB9" pageId="31" pageNumber="30">
<paragraph id="8B9936D98F084843DFED14C0A4E63D21" blockId="31.[192,1440,1227,1493]" pageId="31" pageNumber="30">
<keyLead id="30D795EC8F084843DFED14C0A4E63D21" pageId="31" pageNumber="30">
7(6). Pronotum widest before middle, slightly wrinkled on disc, sides strongly sinuate posteriorly, posterolateral angles rectangular; interval 3 of elytra with a minute subapical setiferous puncture; 
<materialsCitation id="3B4E3C848F084843DA6014E5A4E63D21" location="eastern central North Island" pageId="31" pageNumber="30" specimenCount="1" stateProvince="Gabu">
<figureCitation id="131D2A5C8F084843DA6014E5A4E63D65" box="[1357,1437,1261,1288]" captionStart="Figures 57–60" captionStartId="75.[194,281,1866,1892]" captionTargetBox="[198,1431,256,1834]" captionTargetId="figure-41@75.[192,1439,232,1837]" captionTargetPageId="75" captionText="Figures 57–60. Dorsal habitus. 57) Ctenognathus garnerae new species. 58) C. tepaki new species. 59) C. urewera new species. 60) C. cardiophorus. Scale line = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5041845" httpUri="https://zenodo.org/record/5041845/files/figure.png" pageId="31" pageNumber="30">Fig. 59</figureCitation>
. [Body length 
<quantity id="4CDE9B3C8F084843DEC7151BA3F73D47" box="[490,652,1296,1322]" metricMagnitude="-2" metricUnit="m" metricValue="1.325" metricValueMax="1.38" metricValueMin="1.27" pageId="31" pageNumber="30" unit="mm" value="13.25" valueMax="13.8" valueMin="12.7">12.7–13.8 mm</quantity>
; 
<location id="8EF960028F084843DDBA151BA2AB3D47" LSID="urn:lsid:plazi:treatment:038F87CF8F08487DDFED10DDA4E63CFB:8EF960028F084843DDBA151BA2AB3D47" box="[663,976,1296,1322]" name="eastern central North Island" pageId="31" pageNumber="30" stateProvince="Gabu">eastern central North Island</location>
(BP, 
<collectingRegion id="49E2F83B8F084843DB24151BA5543D47" box="[1033,1071,1296,1322]" country="Guinea-Bissau" name="Gabu" pageId="31" pageNumber="30">GB</collectingRegion>
, HB)]..................................................... 
<emphasis id="B952EACB8F084843DDCF153AA4E63D21" bold="true" box="[738,1437,1329,1356]" pageId="31" pageNumber="30">
<taxonomicName id="4C264D5A8F084843DDCF153AA4703D21" ID-CoL="8MLCX" authority="Larochelle and Lariviere" authorityName="Larochelle and Lariviere E" authorityYear="2021" box="[738,1291,1329,1356]" class="Insecta" family="Carabidae" genus="Ctenognathus" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="30" phylum="Arthropoda" rank="species" species="urewera" status="sp. nov.">
<emphasis id="B952EACB8F084843DDCF153AA29E3D21" bold="true" box="[738,997,1329,1356]" italics="true" pageId="31" pageNumber="30">Ctenognathus urewera</emphasis>
Larochelle and Larivière
</taxonomicName>
, 
<taxonomicNameLabel id="A26157B08F084843DA3B1539A4E63D21" box="[1302,1437,1330,1356]" pageId="31" pageNumber="30" rank="species">new species</taxonomicNameLabel>
</emphasis>
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</keyLead>
</paragraph>
<paragraph id="8B9936D98F084843DFED155EA4E53DB9" blockId="31.[192,1440,1227,1493]" pageId="31" pageNumber="30">
<keyLead id="30D795EC8F084843DFED155EA4E53DB9" pageId="31" pageNumber="30">
— Pronotum widest about middle, deeply wrinkled on disc, sides not sinuate posteriorly, posterolateral angles acute and projected laterally; interval 3 of elytra without setiferous punctures; 
<figureCitation id="131D2A5C8F084843DA2E157DA4293DFD" box="[1283,1362,1398,1424]" captionStart="Figures 57–60" captionStartId="75.[194,281,1866,1892]" captionTargetBox="[198,1431,256,1834]" captionTargetId="figure-41@75.[192,1439,232,1837]" captionTargetPageId="75" captionText="Figures 57–60. Dorsal habitus. 57) Ctenognathus garnerae new species. 58) C. tepaki new species. 59) C. urewera new species. 60) C. cardiophorus. Scale line = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5041845" httpUri="https://zenodo.org/record/5041845/files/figure.png" pageId="31" pageNumber="30">Fig. 60</figureCitation>
. [Body length 
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; northern North Island (ND to WO)].................................................................................... 
<taxonomicName id="4C264D5A8F084843DCF815B1A4E53DB9" ID-CoL="325S9" authority="(Chaudoir)" baseAuthorityName="Chaudoir" baseAuthorityYear="1878" box="[981,1438,1466,1493]" class="Insecta" family="Carabidae" genus="Ctenognathus" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="30" phylum="Arthropoda" rank="species" species="cardiophorus">
<emphasis id="B952EACB8F084843DCF815B1A4E53DB9" bold="true" box="[981,1438,1466,1493]" pageId="31" pageNumber="30">
<emphasis id="B952EACB8F084843DCF815B1A4683DB8" bold="true" box="[981,1299,1466,1493]" italics="true" pageId="31" pageNumber="30">Ctenognathus cardiophorus</emphasis>
(Chaudoir)
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</paragraph>
</keyStep>
<keyStep id="30D22E7C8F084843DFED15EFA4E63EA6" pageId="31" pageNumber="30">
<paragraph id="8B9936D98F084843DFED15EFA4E53E2F" blockId="31.[192,1442,1508,1740]" pageId="31" pageNumber="30">
<keyLead id="30D795EC8F084843DFED15EFA4E53E2F" pageId="31" pageNumber="30">
8(6). Pronotum subquadrate, moderately convex, sides moderately sinuate posteriorly, posterolateral angles rectangular, moderately long; interval 3 of elytra without setiferous punctures; 
<figureCitation id="131D2A5C8F084843DB94160CA47D3E4C" box="[1209,1286,1543,1569]" captionStart="Figures 61–64" captionStartId="76.[192,279,1866,1892]" captionTargetBox="[210,1433,218,1837]" captionTargetId="figure-45@76.[195,1440,206,1837]" captionTargetPageId="76" captionText="Figures 61–64. Dorsal habitus. 61) Ctenognathus adamsi. 62) C. pictonensis. 63) C. tawanui new species. 64) C. perumalae new species. Scale line = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5041851" httpUri="https://zenodo.org/record/5041851/files/figure.png" pageId="31" pageNumber="30">Fig. 61</figureCitation>
. [Body length 
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; North Island (AK–CL to WN)]................. 
<taxonomicName id="4C264D5A8F084843DB121623A4E53E2F" ID-CoL="325S5" authority="(Broun)" baseAuthorityName="Broun" baseAuthorityYear="1886" box="[1087,1438,1576,1603]" class="Insecta" family="Carabidae" genus="Ctenognathus" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="30" phylum="Arthropoda" rank="species" species="adamsi">
<emphasis id="B952EACB8F084843DB121623A4E53E2F" bold="true" box="[1087,1438,1576,1603]" pageId="31" pageNumber="30">
<emphasis id="B952EACB8F084843DB121623A4413E2E" bold="true" box="[1087,1338,1576,1603]" italics="true" pageId="31" pageNumber="30">Ctenognathus adamsi</emphasis>
(Broun)
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</keyLead>
</paragraph>
<paragraph id="8B9936D98F084843DFED1640A4E63EA6" blockId="31.[192,1442,1508,1740]" pageId="31" pageNumber="30">
<keyLead id="30D795EC8F084843DFED1640A4E63EA6" pageId="31" pageNumber="30">
— Pronotum elongate, slightly convex, sides strongly sinuate posteriorly, posterolateral angles acute (projected laterally), very long; interval 3 of elytra with a large (foveate) subapical setiferous puncture; 
<figureCitation id="131D2A5C8F084843DE7C1684A0DB3EC4" box="[337,416,1679,1705]" captionStart="Figures 61–64" captionStartId="76.[192,279,1866,1892]" captionTargetBox="[210,1433,218,1837]" captionTargetId="figure-45@76.[195,1440,206,1837]" captionTargetPageId="76" captionText="Figures 61–64. Dorsal habitus. 61) Ctenognathus adamsi. 62) C. pictonensis. 63) C. tawanui new species. 64) C. perumalae new species. Scale line = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5041851" httpUri="https://zenodo.org/record/5041851/files/figure.png" pageId="31" pageNumber="30">Fig. 62</figureCitation>
. [Body length 11.0– 
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; southern North Island (WI to WN–WA) and northern South Island (SD, NN, MB)]...................................... 
<taxonomicName id="4C264D5A8F084843DB0016BAA4E63EA6" ID-CoL="325T3" authority="Sharp" authorityName="Sharp" authorityYear="1886" box="[1069,1437,1713,1740]" class="Insecta" family="Carabidae" genus="Ctenognathus" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="30" phylum="Arthropoda" rank="species" species="pictonensis">
<emphasis id="B952EACB8F084843DB0016BAA4E63EA6" bold="true" box="[1069,1437,1713,1740]" pageId="31" pageNumber="30">
<emphasis id="B952EACB8F084843DB0016BAA4293EA6" bold="true" box="[1069,1362,1713,1740]" italics="true" pageId="31" pageNumber="30">Ctenognathus pictonensis</emphasis>
Sharp
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</paragraph>
</keyStep>
<keyStep id="30D22E7C8F084843DFED16D7A4E63F7A" pageId="31" pageNumber="30">
<paragraph id="8B9936D98F084843DFED16D7A4E63E98" blockId="31.[192,1437,1755,1816]" box="[192,1437,1755,1782]" pageId="31" pageNumber="30">
<keyLead id="30D795EC8F084843DFED16D7A4E63E98" box="[192,1437,1755,1782]" pageId="31" pageNumber="30">
9(1). Femora dark, pure black, piceous black or dark rufous....................................... 
<emphasis id="B952EACB8F084843DAAC16D0A4E63E98" bold="true" box="[1409,1437,1755,1781]" pageId="31" pageNumber="30">10</emphasis>
</keyLead>
</paragraph>
<paragraph id="8B9936D98F084843DFED16F5A4E63F7A" blockId="31.[192,1437,1755,1816]" box="[192,1437,1789,1816]" pageId="31" pageNumber="30">
<keyLead id="30D795EC8F084843DFED16F5A4E63F7A" box="[192,1437,1789,1816]" pageId="31" pageNumber="30">
— Femora paler, yellow or rufotestaceous..................................................... 
<emphasis id="B952EACB8F084843DAAC16F6A4E63F7A" bold="true" box="[1409,1437,1789,1815]" pageId="31" pageNumber="30">15</emphasis>
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</keyStep>
<keyStep id="30D22E7C8F08487CDFED1723A4E6389D" lastPageId="32" lastPageNumber="31" pageId="31" pageNumber="30">
<paragraph id="8B9936D98F084843DFED1723A4E63FEB" blockId="31.[192,1440,1832,1927]" pageId="31" pageNumber="30">
<keyLead id="30D795EC8F084843DFED1723A4E63FEB" pageId="31" pageNumber="30">
10(9). Scutellar setiferous pore of elytra invisible; legs short; 
<figureCitation id="131D2A5C8F084843DCAF1723A2A93F2F" box="[898,978,1832,1858]" captionStart="Figures 61–64" captionStartId="76.[192,279,1866,1892]" captionTargetBox="[210,1433,218,1837]" captionTargetId="figure-45@76.[195,1440,206,1837]" captionTargetPageId="76" captionText="Figures 61–64. Dorsal habitus. 61) Ctenognathus adamsi. 62) C. pictonensis. 63) C. tawanui new species. 64) C. perumalae new species. Scale line = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5041851" httpUri="https://zenodo.org/record/5041851/files/figure.png" pageId="31" pageNumber="30">Fig. 63</figureCitation>
. [Rather small, body length 7.0– 
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; terminal segment of palpi truncate apically; southern South Island (SL)]..................................................... 
<emphasis id="B952EACB8F084843DDCF1767A4E63FEB" bold="true" box="[738,1437,1899,1927]" pageId="31" pageNumber="30">
<taxonomicName id="4C264D5A8F084843DDCF1767A4703FEB" ID-CoL="8MC3H" authority="Larochelle and Lariviere" authorityName="Larochelle and Lariviere E" authorityYear="2021" box="[738,1291,1899,1927]" class="Insecta" family="Carabidae" genus="Ctenognathus" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="30" phylum="Arthropoda" rank="species" species="tawanui" status="sp. nov.">
<emphasis id="B952EACB8F084843DDCF1767A29D3FEB" bold="true" box="[738,998,1900,1927]" italics="true" pageId="31" pageNumber="30">Ctenognathus tawanui</emphasis>
Larochelle and Larivière
</taxonomicName>
, 
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</emphasis>
</keyLead>
</paragraph>
<paragraph id="8B9936D98F37487CDFED10DDA4E6389D" blockId="32.[192,1437,214,240]" box="[192,1437,214,240]" pageId="32" pageNumber="31">
<keyLead id="30D795EC8F37487CDFED10DDA4E6389D" box="[192,1437,214,240]" pageId="32" pageNumber="31">
— Scutellar setiferous pore of elytra visible; legs moderately long or very long..................... 
<emphasis id="B952EACB8F37487CDAAC10DDA4E6389D" bold="true" box="[1409,1437,214,240]" pageId="32" pageNumber="31">11</emphasis>
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</paragraph>
</keyStep>
<keyStep id="30D22E7C8F37487CDFED110BA4E639CF" pageId="32" pageNumber="31">
<paragraph id="8B9936D98F37487CDFED110BA4E63930" blockId="32.[192,1440,256,418]" pageId="32" pageNumber="31">
<keyLead id="30D795EC8F37487CDFED110BA4E63930" pageId="32" pageNumber="31">
11(10). Pronotum strongly cordate; elytra subovate, strongly convex; 
<figureCitation id="131D2A5C8F37487CDCEF110BA5743977" box="[962,1039,256,282]" captionStart="Figures 61–64" captionStartId="76.[192,279,1866,1892]" captionTargetBox="[210,1433,218,1837]" captionTargetId="figure-45@76.[195,1440,206,1837]" captionTargetPageId="76" captionText="Figures 61–64. Dorsal habitus. 61) Ctenognathus adamsi. 62) C. pictonensis. 63) C. tawanui new species. 64) C. perumalae new species. Scale line = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5041851" httpUri="https://zenodo.org/record/5041851/files/figure.png" pageId="32" pageNumber="31">Fig. 64</figureCitation>
. [Body length 
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; southwestern South Island (southern WD)]................................................................................ 
<emphasis id="B952EACB8F37487CDDE81148A4E63930" bold="true" box="[709,1437,323,350]" pageId="32" pageNumber="31">
<taxonomicName id="4C264D5A8F37487CDDE81148A4703933" ID-CoL="8M6G4" authority="Larochelle and Lariviere" authorityName="Larochelle and Lariviere E" authorityYear="2021" box="[709,1291,323,350]" class="Insecta" family="Carabidae" genus="Ctenognathus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="31" phylum="Arthropoda" rank="species" species="perumalae" status="sp. nov.">
<emphasis id="B952EACB8F37487CDDE81148A29D3933" bold="true" box="[709,998,323,350]" italics="true" pageId="32" pageNumber="31">Ctenognathus perumalae</emphasis>
Larochelle and Larivière
</taxonomicName>
, 
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</emphasis>
</keyLead>
</paragraph>
<paragraph id="8B9936D98F37487CDFED116DA4E639CF" blockId="32.[192,1440,256,418]" pageId="32" pageNumber="31">
<keyLead id="30D795EC8F37487CDFED116DA4E639CF" pageId="32" pageNumber="31">
— Pronotum moderately cordate, subquadrate, or subrectangular; elytra oblong, slightly convex or subdepressed............................................................................ 
<emphasis id="B952EACB8F37487CDAAC1183A4E639CF" bold="true" box="[1409,1437,392,418]" pageId="32" pageNumber="31">12</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D22E7C8F37487CDFED11B9A4E63983" pageId="32" pageNumber="31">
<paragraph id="8B9936D98F37487CDFED11B9A4E639A1" blockId="32.[192,1437,434,494]" box="[192,1437,434,460]" pageId="32" pageNumber="31">
<keyLead id="30D795EC8F37487CDFED11B9A4E639A1" box="[192,1437,434,460]" pageId="32" pageNumber="31">
12(11). Antennal segments 1–3(4) black, (3)4–11 dark rufous........................................ 
<emphasis id="B952EACB8F37487CDAAC11B9A4E639A1" bold="true" box="[1409,1437,434,460]" pageId="32" pageNumber="31">13</emphasis>
</keyLead>
</paragraph>
<paragraph id="8B9936D98F37487CDFED11DFA4E63983" blockId="32.[192,1437,434,494]" box="[192,1437,468,494]" pageId="32" pageNumber="31">
<keyLead id="30D795EC8F37487CDFED11DFA4E63983" box="[192,1437,468,494]" pageId="32" pageNumber="31">
— Antennae entirely dark rufous............................................................ 
<emphasis id="B952EACB8F37487CDAAC11DFA4E63983" bold="true" box="[1409,1437,468,494]" pageId="32" pageNumber="31">14</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D22E7C8F37487CDFED11F5A4E53AAC" pageId="32" pageNumber="31">
<paragraph id="8B9936D98F37487CDFED11F5A4E63A31" blockId="32.[192,1440,510,706]" pageId="32" pageNumber="31">
<keyLead id="30D795EC8F37487CDFED11F5A4E63A31" pageId="32" pageNumber="31">
13(12). Femora and tibiae black; pronotum moderately cordate, base emarginate; elytra subdepressed, microsculpture isodiametric, striae finely punctate; 
<figureCitation id="131D2A5C8F37487CDC11122BA2F23A56" box="[828,905,544,571]" captionStart="Figures 65–68" captionStartId="77.[192,279,1866,1892]" captionTargetBox="[214,1431,237,1836]" captionTargetId="figure-12@77.[192,1439,208,1839]" captionTargetPageId="77" captionText="Figures 65–68. Dorsal habitus. 65) Ctenognathus takahe new species. 66) C. arnaudensis. 67) C. marieclaudiae new species. 68) C. edwardsii. Scale line = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5041855" httpUri="https://zenodo.org/record/5041855/files/figure.png" pageId="32" pageNumber="31">Fig. 65</figureCitation>
. [Body length 
<quantity id="4CDE9B3C8F37487CDB0A122BA5C03A57" box="[1063,1211,544,571]" metricMagnitude="-3" metricUnit="m" metricValue="9.95" metricValueMax="11.4" metricValueMin="8.5" pageId="32" pageNumber="31" unit="mm" value="9.95" valueMax="11.4" valueMin="8.5">8.5–11.4 mm</quantity>
; southwestern South Island (southern FD)]............ 
<emphasis id="B952EACB8F37487CDDDE124AA4E63A31" bold="true" box="[755,1437,577,604]" pageId="32" pageNumber="31">
<taxonomicName id="4C264D5A8F37487CDDDE124AA4703A31" ID-CoL="8ML7L" authority="Larochelle and Lariviere" authorityName="Larochelle and Lariviere E" authorityYear="2021" box="[755,1291,577,604]" class="Insecta" family="Carabidae" genus="Ctenognathus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="31" phylum="Arthropoda" rank="species" species="takahe" status="sp. nov.">
<emphasis id="B952EACB8F37487CDDDE124AA29E3A31" bold="true" box="[755,997,577,604]" italics="true" pageId="32" pageNumber="31">Ctenognathus takahe</emphasis>
Larochelle and Larivière
</taxonomicName>
, 
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</paragraph>
<paragraph id="8B9936D98F37487CDFED126FA4E53AAC" blockId="32.[192,1440,510,706]" pageId="32" pageNumber="31">
<keyLead id="30D795EC8F37487CDFED126FA4E53AAC" pageId="32" pageNumber="31">
— Femora and tibiae dark rufous; pronotum subrectangular, base subtruncate; elytra slightly convex, microsculpture moderately transverse, striae impunctate; 
<figureCitation id="131D2A5C8F37487CDCC3128DA5393ACD" box="[1006,1090,646,672]" captionStart="Figures 65–68" captionStartId="77.[192,279,1866,1892]" captionTargetBox="[214,1431,237,1836]" captionTargetId="figure-12@77.[192,1439,208,1839]" captionTargetPageId="77" captionText="Figures 65–68. Dorsal habitus. 65) Ctenognathus takahe new species. 66) C. arnaudensis. 67) C. marieclaudiae new species. 68) C. edwardsii. Scale line = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5041855" httpUri="https://zenodo.org/record/5041855/files/figure.png" pageId="32" pageNumber="31">Fig. 66</figureCitation>
. [Body length 
<quantity id="4CDE9B3C8F37487CDBD9128DA4E73ACD" box="[1268,1436,646,672]" metricMagnitude="-2" metricUnit="m" metricValue="1.035" metricValueMax="1.06" metricValueMin="1.01" pageId="32" pageNumber="31" unit="mm" value="10.35" valueMax="10.6" valueMin="10.1">10.1–10.6 mm</quantity>
; northern South Island (NN, BR, MB, KA)]............... 
<taxonomicName id="4C264D5A8F37487CDB2A12ACA4E53AAC" ID-CoL="325S6" authority="(Broun)" baseAuthorityName="Broun" baseAuthorityYear="1921" box="[1031,1438,679,706]" class="Insecta" family="Carabidae" genus="Ctenognathus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="31" phylum="Arthropoda" rank="species" species="arnaudensis">
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<emphasis id="B952EACB8F37487CDB2A12ACA4423AAF" bold="true" box="[1031,1337,679,706]" italics="true" pageId="32" pageNumber="31">Ctenognathus arnaudensis</emphasis>
(Broun)
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<keyStep id="30D22E7C8F37487CDFED12D9A4E53BB4" pageId="32" pageNumber="31">
<paragraph id="8B9936D98F37487CDFED12D9A4E63B3F" blockId="32.[192,1440,722,986]" pageId="32" pageNumber="31">
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14(12). Pronotum unwrinkled, moderately cordate, laterobasal foveae moderately wide (not meeting toward pronotal midline); elytra very shiny, slightly convex, interval 3 with three or four large (foveate) setiferous punctures; 
<figureCitation id="131D2A5C8F37487CDD13131DA3F63B5D" box="[574,653,790,816]" captionStart="Figures 65–68" captionStartId="77.[192,279,1866,1892]" captionTargetBox="[214,1431,237,1836]" captionTargetId="figure-12@77.[192,1439,208,1839]" captionTargetPageId="77" captionText="Figures 65–68. Dorsal habitus. 65) Ctenognathus takahe new species. 66) C. arnaudensis. 67) C. marieclaudiae new species. 68) C. edwardsii. Scale line = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5041855" httpUri="https://zenodo.org/record/5041855/files/figure.png" pageId="32" pageNumber="31">Fig. 67</figureCitation>
. [Body length 
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; western South Island (BR, WD, OL)]........................................ 
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<taxonomicName id="4C264D5A8F37487CDC10133CA4703B3F" ID-CoL="8MHLF" authority="Larochelle" authorityName="Larochelle &amp; Larivière" authorityYear="2021" box="[829,1291,823,850]" class="Insecta" family="Carabidae" genus="Ctenognathus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="31" phylum="Arthropoda" rank="species" species="marieclaudiae" status="sp. nov.">
<emphasis id="B952EACB8F37487CDC10133CA5FC3B3F" bold="true" box="[829,1159,823,850]" italics="true" pageId="32" pageNumber="31">Ctenognathus marieclaudiae</emphasis>
Larochelle
</taxonomicName>
, 
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</emphasis>
</keyLead>
</paragraph>
<paragraph id="8B9936D98F37487CDFED1351A4E53BB4" blockId="32.[192,1440,722,986]" pageId="32" pageNumber="31">
<keyLead id="30D795EC8F37487CDFED1351A4E53BB4" pageId="32" pageNumber="31">
— Pronotum wrinkled on disc and across base, subquadrate, laterobasal foveae very wide (meeting toward pronotal midline); elytra dull, subdepressed, interval 3 with three shallow, minute setiferous punctures; 
<figureCitation id="131D2A5C8F37487CDEBF1395A0A43BD5" box="[402,479,926,952]" captionStart="Figures 65–68" captionStartId="77.[192,279,1866,1892]" captionTargetBox="[214,1431,237,1836]" captionTargetId="figure-12@77.[192,1439,208,1839]" captionTargetPageId="77" captionText="Figures 65–68. Dorsal habitus. 65) Ctenognathus takahe new species. 66) C. arnaudensis. 67) C. marieclaudiae new species. 68) C. edwardsii. Scale line = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5041855" httpUri="https://zenodo.org/record/5041855/files/figure.png" pageId="32" pageNumber="31">Fig. 68</figureCitation>
. [Smaller, body length 8.5–11.0 mm; southern South Island (MC to FD–SL) and Stewart Island]................................................... 
<taxonomicName id="4C264D5A8F37487CDB1D13B4A4E53BB4" ID-CoL="6BMK2" authority="(Bates)" baseAuthorityName="Bates" baseAuthorityYear="1874" box="[1072,1438,959,986]" class="Insecta" family="Carabidae" genus="Ctenognathus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="31" phylum="Arthropoda" rank="species" species="edwardsii">
<emphasis id="B952EACB8F37487CDB1D13B4A4E53BB4" bold="true" box="[1072,1438,959,986]" pageId="32" pageNumber="31">
<emphasis id="B952EACB8F37487CDB1D13B4A43E3BB7" bold="true" box="[1072,1349,959,986]" italics="true" pageId="32" pageNumber="31">Ctenognathus edwardsii</emphasis>
(Bates)
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</paragraph>
</keyStep>
<keyStep id="30D22E7C8F37487CDFED13E1A4E63C07" pageId="32" pageNumber="31">
<paragraph id="8B9936D98F37487CDFED13E1A4E63C25" blockId="32.[192,1440,1002,1130]" pageId="32" pageNumber="31">
<keyLead id="30D795EC8F37487CDFED13E1A4E63C25" pageId="32" pageNumber="31">
15(9). Meso- and metatarsomeres 1–4 (
<figureCitation id="131D2A5C8F37487CDDB313E1A38A3C69" box="[670,753,1002,1028]" captionStart="Figures 27–36" captionStartId="69.[192,279,1765,1791]" captionTargetBox="[309,1386,262,1710]" captionTargetId="figure-85@69.[264,1406,242,1738]" captionTargetPageId="69" captionText="Figures 27–36. Meso- or metatarsomere, dorsal view: 27) not carinate; 28) tricarinate. Metatarsomere 4: 29) moderately bilobed and asymmetrical apically; 30) moderately bilobed and symmetrical; 31) strongly bilobed and asymmetrical. Abdominal sterna V–VI: 32) with 4 ambulatory setae; 33) with 2 ambulatory setae; 34) V–VII with numerous short setae; 35) VII, male with 2 ambulatory setae, female with 4 setae; 36) VII, male with 4 ambulatory setae, female with 8 setae." figureDoi="http://doi.org/10.5281/zenodo.5041827" httpUri="https://zenodo.org/record/5041827/files/figure.png" pageId="32" pageNumber="31">Fig. 27</figureCitation>
) not carinate dorsally; 
<figureCitation id="131D2A5C8F37487CDB2F13E1A52F3C69" box="[1026,1108,1002,1028]" captionStart="Figures 69–72" captionStartId="78.[192,279,1866,1892]" captionTargetBox="[212,1431,236,1836]" captionTargetId="figure-17@78.[192,1439,208,1839]" captionTargetPageId="78" captionText="Figures 69–72. Dorsal habitus. 69) Ctenognathus kaikoura new species. 70) C. sandageri. 71) C. helmsi. 72) C. xanthomelus. Scale line = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5041857" httpUri="https://zenodo.org/record/5041857/files/figure.png" pageId="32" pageNumber="31">Fig. 69</figureCitation>
. [Body length 
<quantity id="4CDE9B3C8F37487CDA2E13E1A4E73C69" box="[1283,1436,1002,1029]" metricMagnitude="-3" metricUnit="m" metricValue="9.8" metricValueMax="10.5" metricValueMin="9.1" pageId="32" pageNumber="31" unit="mm" value="9.8" valueMax="10.5" valueMin="9.1">9.1–10.5 mm</quantity>
; northeastern South Island (MB, KA)]................................................................................... 
<emphasis id="B952EACB8F37487CDDF51426A4E63C25" bold="true" box="[728,1437,1069,1096]" pageId="32" pageNumber="31">
<taxonomicName id="4C264D5A8F37487CDDF51426A4703C25" ID-CoL="8MFPR" authority="Larochelle and Lariviere" authorityName="Larochelle and Lariviere E" authorityYear="2021" box="[728,1291,1069,1096]" class="Insecta" family="Carabidae" genus="Ctenognathus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="31" phylum="Arthropoda" rank="species" species="kaikoura" status="sp. nov.">
<emphasis id="B952EACB8F37487CDDF51426A29D3C25" bold="true" box="[728,998,1069,1096]" italics="true" pageId="32" pageNumber="31">Ctenognathus kaikoura</emphasis>
Larochelle and Larivière
</taxonomicName>
, 
<taxonomicNameLabel id="A26157B08F37487CDA3B1425A4E63C25" box="[1302,1437,1070,1096]" pageId="32" pageNumber="31" rank="species">new species</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8B9936D98F37487CDFED145BA4E63C07" blockId="32.[192,1440,1002,1130]" box="[192,1437,1104,1130]" pageId="32" pageNumber="31">
<keyLead id="30D795EC8F37487CDFED145BA4E63C07" box="[192,1437,1104,1130]" pageId="32" pageNumber="31">
— Meso- and metatarsomeres 1–4 (
<figureCitation id="131D2A5C8F37487CDDA0145BA3A73C07" box="[653,732,1104,1130]" captionStart="Figures 27–36" captionStartId="69.[192,279,1765,1791]" captionTargetBox="[309,1386,262,1710]" captionTargetId="figure-85@69.[264,1406,242,1738]" captionTargetPageId="69" captionText="Figures 27–36. Meso- or metatarsomere, dorsal view: 27) not carinate; 28) tricarinate. Metatarsomere 4: 29) moderately bilobed and asymmetrical apically; 30) moderately bilobed and symmetrical; 31) strongly bilobed and asymmetrical. Abdominal sterna V–VI: 32) with 4 ambulatory setae; 33) with 2 ambulatory setae; 34) V–VII with numerous short setae; 35) VII, male with 2 ambulatory setae, female with 4 setae; 36) VII, male with 4 ambulatory setae, female with 8 setae." figureDoi="http://doi.org/10.5281/zenodo.5041827" httpUri="https://zenodo.org/record/5041827/files/figure.png" pageId="32" pageNumber="31">Fig. 28</figureCitation>
) tricarinate dorsally................................... 
<emphasis id="B952EACB8F37487CDAAC145BA4E63C07" bold="true" box="[1409,1437,1104,1130]" pageId="32" pageNumber="31">16</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D22E7C8F37487CDFED1471A4E63C97" pageId="32" pageNumber="31">
<paragraph id="8B9936D98F37487CDFED1471A4E63CDB" blockId="32.[192,1440,1146,1274]" pageId="32" pageNumber="31">
<keyLead id="30D795EC8F37487CDFED1471A4E63CDB" pageId="32" pageNumber="31">
16(15). Sternum VII (last visible sternum of abdomen; 
<figureCitation id="131D2A5C8F37487CDC6D1471A2EA3CF8" box="[832,913,1146,1173]" captionStart="Figures 27–36" captionStartId="69.[192,279,1765,1791]" captionTargetBox="[309,1386,262,1710]" captionTargetId="figure-85@69.[264,1406,242,1738]" captionTargetPageId="69" captionText="Figures 27–36. Meso- or metatarsomere, dorsal view: 27) not carinate; 28) tricarinate. Metatarsomere 4: 29) moderately bilobed and asymmetrical apically; 30) moderately bilobed and symmetrical; 31) strongly bilobed and asymmetrical. Abdominal sterna V–VI: 32) with 4 ambulatory setae; 33) with 2 ambulatory setae; 34) V–VII with numerous short setae; 35) VII, male with 2 ambulatory setae, female with 4 setae; 36) VII, male with 4 ambulatory setae, female with 8 setae." figureDoi="http://doi.org/10.5281/zenodo.5041827" httpUri="https://zenodo.org/record/5041827/files/figure.png" pageId="32" pageNumber="31">Fig. 35</figureCitation>
): male with two long apical ambulatory setae, female with four long apical ambulatory setae............................................ 
<emphasis id="B952EACB8F37487CDAAC1497A4E63CDB" bold="true" box="[1409,1437,1180,1206]" pageId="32" pageNumber="31">17</emphasis>
</keyLead>
</paragraph>
<paragraph id="8B9936D98F37487CDFED14B5A4E63C97" blockId="32.[192,1440,1146,1274]" pageId="32" pageNumber="31">
<keyLead id="30D795EC8F37487CDFED14B5A4E63C97" pageId="32" pageNumber="31">
— Sternum VII (
<figureCitation id="131D2A5C8F37487CDE9114B5A3753CB4" box="[444,526,1214,1241]" captionStart="Figures 27–36" captionStartId="69.[192,279,1765,1791]" captionTargetBox="[309,1386,262,1710]" captionTargetId="figure-85@69.[264,1406,242,1738]" captionTargetPageId="69" captionText="Figures 27–36. Meso- or metatarsomere, dorsal view: 27) not carinate; 28) tricarinate. Metatarsomere 4: 29) moderately bilobed and asymmetrical apically; 30) moderately bilobed and symmetrical; 31) strongly bilobed and asymmetrical. Abdominal sterna V–VI: 32) with 4 ambulatory setae; 33) with 2 ambulatory setae; 34) V–VII with numerous short setae; 35) VII, male with 2 ambulatory setae, female with 4 setae; 36) VII, male with 4 ambulatory setae, female with 8 setae." figureDoi="http://doi.org/10.5281/zenodo.5041827" httpUri="https://zenodo.org/record/5041827/files/figure.png" pageId="32" pageNumber="31">Fig. 36</figureCitation>
): male with four long apical ambulatory setae, female with eight to ten long apical ambulatory setae..................................................................... 
<emphasis id="B952EACB8F37487CDAAC14EBA4E63C97" bold="true" box="[1409,1437,1248,1274]" pageId="32" pageNumber="31">20</emphasis>
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</paragraph>
</keyStep>
<keyStep id="30D22E7C8F37487CDFED1501A4E53D05" pageId="32" pageNumber="31">
<paragraph id="8B9936D98F37487CDFED1501A4E63D2B" blockId="32.[192,1440,1290,1384]" pageId="32" pageNumber="31">
<keyLead id="30D795EC8F37487CDFED1501A4E63D2B" pageId="32" pageNumber="31">
17(16). Posterolateral angles of pronotum acute and projected laterally; terminal segment of palpi obtuse apically (
<figureCitation id="131D2A5C8F37487CDE751527A0DE3D2B" box="[344,421,1324,1350]" captionStart="Figures 1–13" captionStartId="67.[192,279,1833,1859]" captionTargetBox="[247,1417,213,1793]" captionTargetId="figure-11@67.[207,1425,213,1806]" captionTargetPageId="67" captionText="Figures 1–13. Microsculpture: 1) granulate; 2) isodiametric; 3) moderately transverse; 4) very transverse. Neck constriction, dorsolateral view: 5) deep; 6) shallow; 7) absent. Eye and buccal fissure, lateral view: 8) touching; 9) separated. Terminal segment of palpi: 10) obtuse apically; 11) truncate. Paraglossae: 12) pubescent; 13) glabrous." figureDoi="http://doi.org/10.5281/zenodo.5041815" httpUri="https://zenodo.org/record/5041815/files/figure.png" pageId="32" pageNumber="31">Fig. 10</figureCitation>
).............................................................................. 
<emphasis id="B952EACB8F37487CDAAC1527A4E63D2B" bold="true" box="[1409,1437,1324,1350]" pageId="32" pageNumber="31">18</emphasis>
</keyLead>
</paragraph>
<paragraph id="8B9936D98F37487CDFED1545A4E53D05" blockId="32.[192,1440,1290,1384]" box="[192,1438,1358,1384]" pageId="32" pageNumber="31">
<keyLead id="30D795EC8F37487CDFED1545A4E53D05" box="[192,1438,1358,1384]" pageId="32" pageNumber="31">
— Posterolateral angles of pronotum rectangular; terminal segment of palpi truncate apically (
<figureCitation id="131D2A5C8F37487CDBD21545A4303D05" box="[1279,1355,1358,1384]" captionStart="Figures 1–13" captionStartId="67.[192,279,1833,1859]" captionTargetBox="[247,1417,213,1793]" captionTargetId="figure-11@67.[207,1425,213,1806]" captionTargetPageId="67" captionText="Figures 1–13. Microsculpture: 1) granulate; 2) isodiametric; 3) moderately transverse; 4) very transverse. Neck constriction, dorsolateral view: 5) deep; 6) shallow; 7) absent. Eye and buccal fissure, lateral view: 8) touching; 9) separated. Terminal segment of palpi: 10) obtuse apically; 11) truncate. Paraglossae: 12) pubescent; 13) glabrous." figureDoi="http://doi.org/10.5281/zenodo.5041815" httpUri="https://zenodo.org/record/5041815/files/figure.png" pageId="32" pageNumber="31">Fig. 11</figureCitation>
)... 
<emphasis id="B952EACB8F37487CDAAF1545A4E53D05" bold="true" box="[1410,1438,1358,1384]" pageId="32" pageNumber="31">19</emphasis>
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</paragraph>
</keyStep>
<keyStep id="30D22E7C8F37487CDFED1573A4E53E33" pageId="32" pageNumber="31">
<paragraph id="8B9936D98F37487CDFED1573A4E53D95" blockId="32.[192,1442,1400,1631]" pageId="32" pageNumber="31">
<keyLead id="30D795EC8F37487CDFED1573A4E53D95" pageId="32" pageNumber="31">
18(17). Femora and tibiae pale yellow; pronotum with anterolateral angles poorly developed, posterior bead incomplete (obsolete medially), base emarginate; legs moderately long; elytra very convex; 
<figureCitation id="131D2A5C8F37487CDA601591A4E63DD9" box="[1357,1437,1434,1460]" captionStart="Figures 69–72" captionStartId="78.[192,279,1866,1892]" captionTargetBox="[212,1431,236,1836]" captionTargetId="figure-17@78.[192,1439,208,1839]" captionTargetPageId="78" captionText="Figures 69–72. Dorsal habitus. 69) Ctenognathus kaikoura new species. 70) C. sandageri. 71) C. helmsi. 72) C. xanthomelus. Scale line = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5041857" httpUri="https://zenodo.org/record/5041857/files/figure.png" pageId="32" pageNumber="31">Fig. 70</figureCitation>
. [Body length 
<quantity id="4CDE9B3C8F37487CDEC615B7A3F43DBA" box="[491,655,1468,1495]" metricMagnitude="-2" metricUnit="m" metricValue="1.065" metricValueMax="1.1099999999999999" metricValueMin="1.02" pageId="32" pageNumber="31" unit="mm" value="10.649999999999999" valueMax="11.1" valueMin="10.2">10.2–11.1 mm</quantity>
; southern North Island (WN, WA)].................................................................................. 
<taxonomicName id="4C264D5A8F37487CDB3215D6A4E53D95" ID-CoL="325T6" authority="(Broun)" baseAuthorityName="Broun" baseAuthorityYear="1882" box="[1055,1438,1501,1528]" class="Insecta" family="Carabidae" genus="Ctenognathus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="31" phylum="Arthropoda" rank="species" species="sandageri">
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<emphasis id="B952EACB8F37487CDB3215D6A4433D95" bold="true" box="[1055,1336,1501,1528]" italics="true" pageId="32" pageNumber="31">Ctenognathus sandageri</emphasis>
(Broun)
</emphasis>
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</paragraph>
<paragraph id="8B9936D98F37487CDFED160AA4E53E33" blockId="32.[192,1442,1400,1631]" pageId="32" pageNumber="31">
<keyLead id="30D795EC8F37487CDFED160AA4E53E33" pageId="32" pageNumber="31">
— Femora dark yellow, tibiae reddish; pronotum with anterolateral angles well developed, posterior bead complete, base subtruncate; legs very long; elytra moderately convex; 
<figureCitation id="131D2A5C8F37487CDB791629A5D93E51" box="[1108,1186,1570,1596]" captionStart="Figures 69–72" captionStartId="78.[192,279,1866,1892]" captionTargetBox="[212,1431,236,1836]" captionTargetId="figure-17@78.[192,1439,208,1839]" captionTargetPageId="78" captionText="Figures 69–72. Dorsal habitus. 69) Ctenognathus kaikoura new species. 70) C. sandageri. 71) C. helmsi. 72) C. xanthomelus. Scale line = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5041857" httpUri="https://zenodo.org/record/5041857/files/figure.png" pageId="32" pageNumber="31">Fig. 71</figureCitation>
. [Body length 
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; western South Island (NN–SD to WD)]................... 
<taxonomicName id="4C264D5A8F37487CDB631648A4E53E33" ID-CoL="325SH" authority="(Sharp)" baseAuthorityName="Sharp" baseAuthorityYear="1881" box="[1102,1438,1603,1630]" class="Insecta" family="Carabidae" genus="Ctenognathus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="31" phylum="Arthropoda" rank="species" species="helmsi">
<emphasis id="B952EACB8F37487CDB631648A4E53E33" bold="true" box="[1102,1438,1603,1630]" pageId="32" pageNumber="31">
<emphasis id="B952EACB8F37487CDB631648A4443E33" bold="true" box="[1102,1343,1603,1630]" italics="true" pageId="32" pageNumber="31">Ctenognathus helmsi</emphasis>
(Sharp)
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</paragraph>
</keyStep>
<keyStep id="30D22E7C8F37487CDFED1665A4E53E83" pageId="32" pageNumber="31">
<paragraph id="8B9936D98F37487CDFED1665A4E53EC7" blockId="32.[192,1440,1646,1775]" pageId="32" pageNumber="31">
<keyLead id="30D795EC8F37487CDFED1665A4E53EC7" pageId="32" pageNumber="31">
19(17). Elytra ovate; pronotum widest about middle, sides strongly rounded; 
<figureCitation id="131D2A5C8F37487CDB171665A5F03EE5" box="[1082,1163,1646,1672]" captionStart="Figures 69–72" captionStartId="78.[192,279,1866,1892]" captionTargetBox="[212,1431,236,1836]" captionTargetId="figure-17@78.[192,1439,208,1839]" captionTargetPageId="78" captionText="Figures 69–72. Dorsal habitus. 69) Ctenognathus kaikoura new species. 70) C. sandageri. 71) C. helmsi. 72) C. xanthomelus. Scale line = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5041857" httpUri="https://zenodo.org/record/5041857/files/figure.png" pageId="32" pageNumber="31">Fig. 72</figureCitation>
. [Body length 10.0– 
<quantity id="4CDE9B3C8F37487CDA5C1665A0FB3EC6" metricMagnitude="-2" metricUnit="m" metricValue="1.14" pageId="32" pageNumber="31" unit="mm" value="11.4">11.4 mm</quantity>
; North Island (WO–BP to WN)].................... 
<taxonomicName id="4C264D5A8F37487CDB2D1684A4E53EC7" ID-CoL="8MF6G" authority="(Broun)" baseAuthorityName="Broun" baseAuthorityYear="1908" box="[1024,1438,1679,1706]" class="Insecta" family="Carabidae" genus="Ctenognathus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="31" phylum="Arthropoda" rank="species" species="xanthomelus">
<emphasis id="B952EACB8F37487CDB2D1684A4E53EC7" bold="true" box="[1024,1438,1679,1706]" pageId="32" pageNumber="31">
<emphasis id="B952EACB8F37487CDB2D1684A4423EC7" bold="true" box="[1024,1337,1679,1706]" italics="true" pageId="32" pageNumber="31">Ctenognathus xanthomelus</emphasis>
(Broun)
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</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B9936D98F37487CDFED16B8A4E53E83" blockId="32.[192,1440,1646,1775]" pageId="32" pageNumber="31">
<keyLead id="30D795EC8F37487CDFED16B8A4E53E83" pageId="32" pageNumber="31">
— Elytra subovate; pronotum widest before middle, sides moderately rounded; 
<figureCitation id="131D2A5C8F37487CDB5E16B9A5BA3EA1" box="[1139,1217,1714,1740]" captionStart="Figures 73–76" captionStartId="79.[194,281,1866,1892]" captionTargetBox="[214,1431,210,1834]" captionTargetId="figure-37@79.[192,1439,206,1837]" captionTargetPageId="79" captionText="Figures 73–76. Dorsal habitus. 73) Ctenognathus intermedius. 74) C. otagoensis. 75) C. colensonis. 76) C. earlyi new species. Scale line = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5041861" httpUri="https://zenodo.org/record/5041861/files/figure.png" pageId="32" pageNumber="31">Fig. 73</figureCitation>
. [Body length 10.5– 12.0 mm; southern North Island (TO to WN)]............. 
<taxonomicName id="4C264D5A8F37487CDB2716D8A4E53E83" ID-CoL="325SK" authority="(Broun)" baseAuthorityName="Broun" baseAuthorityYear="1908" box="[1034,1438,1747,1774]" class="Insecta" family="Carabidae" genus="Ctenognathus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="31" phylum="Arthropoda" rank="species" species="intermedius">
<emphasis id="B952EACB8F37487CDB2716D8A4E53E83" bold="true" box="[1034,1438,1747,1774]" pageId="32" pageNumber="31">
<emphasis id="B952EACB8F37487CDB2716D8A4413E83" bold="true" box="[1034,1338,1747,1774]" italics="true" pageId="32" pageNumber="31">Ctenognathus intermedius</emphasis>
(Broun)
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</paragraph>
</keyStep>
<keyStep id="30D22E7C8F37487CDFED16F5A4E63F57" pageId="32" pageNumber="31">
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<keyLead id="30D795EC8F37487CDFED16F5A4E63F75" box="[192,1437,1790,1817]" pageId="32" pageNumber="31">
20(16). Femora and tibiae pale yellow............................................................ 
<emphasis id="B952EACB8F37487CDAAC16F5A4E63F75" bold="true" box="[1409,1437,1790,1816]" pageId="32" pageNumber="31">21</emphasis>
</keyLead>
</paragraph>
<paragraph id="8B9936D98F37487CDFED172AA4E63F57" blockId="32.[192,1437,1790,1851]" box="[192,1437,1824,1851]" pageId="32" pageNumber="31">
<keyLead id="30D795EC8F37487CDFED172AA4E63F57" box="[192,1437,1824,1851]" pageId="32" pageNumber="31">
— Femora dark yellow or reddish; tibiae reddish................................................ 
<emphasis id="B952EACB8F37487CDAAC172BA4E63F57" bold="true" box="[1409,1437,1824,1850]" pageId="32" pageNumber="31">23</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D22E7C8F37487DDFED1741A4E63938" lastPageId="33" lastPageNumber="32" pageId="32" pageNumber="31">
<paragraph id="8B9936D98F37487DDFED1741A4E5397C" blockId="32.[192,1440,1866,1927]" lastBlockId="33.[192,1440,213,341]" lastPageId="33" lastPageNumber="32" pageId="32" pageNumber="31">
<keyLead id="30D795EC8F37487DDFED1741A4E5397C" lastPageId="33" lastPageNumber="32" pageId="32" pageNumber="31">
21(20). Elytra shiny; posterolateral angles of pronotum slightly obtuse; mentum tooth subtruncate apically; submentum with ten long setae; 
<figureCitation id="131D2A5C8F37487CDDB91767A3983FEA" box="[660,739,1900,1927]" captionStart="Figures 73–76" captionStartId="79.[194,281,1866,1892]" captionTargetBox="[214,1431,210,1834]" captionTargetId="figure-37@79.[192,1439,206,1837]" captionTargetPageId="79" captionText="Figures 73–76. Dorsal habitus. 73) Ctenognathus intermedius. 74) C. otagoensis. 75) C. colensonis. 76) C. earlyi new species. Scale line = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5041861" httpUri="https://zenodo.org/record/5041861/files/figure.png" pageId="32" pageNumber="31">Fig. 75</figureCitation>
. [Body length 
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; pronotum slightly cordate; elytra slightly convex, striae finely punctate; South Island (NN–BR to NC–MK)]............................................................................. 
<taxonomicName id="4C264D5A8F36487DDB0D10FCA4E5397C" ID-CoL="325SB" authority="(White)" baseAuthorityName="White" baseAuthorityYear="1846" box="[1056,1438,247,274]" class="Insecta" family="Carabidae" genus="Ctenognathus" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="32" phylum="Arthropoda" rank="species" species="colensonis">
<emphasis id="B952EACB8F36487DDB0D10FCA4E5397C" bold="true" box="[1056,1438,247,274]" pageId="33" pageNumber="32">
<emphasis id="B952EACB8F36487DDB0D10FCA447397F" bold="true" box="[1056,1340,247,274]" italics="true" pageId="33" pageNumber="32">Ctenognathus colensonis</emphasis>
(White)
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</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B9936D98F36487DDFED1111A4E63938" blockId="33.[192,1440,213,341]" pageId="33" pageNumber="32">
<keyLead id="30D795EC8F36487DDFED1111A4E63938" pageId="33" pageNumber="32">
— Elytra dull; posterolateral angles of pronotum rectangular; mentum tooth emarginate apically; submentum with six long setae............................................................... 
<emphasis id="B952EACB8F36487DDAAC1130A4E63938" bold="true" box="[1409,1437,315,341]" pageId="33" pageNumber="32">22</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D22E7C8F36487DDFED116EA4E63A4A" pageId="33" pageNumber="32">
<paragraph id="8B9936D98F36487DDFED116EA20239AF" blockId="33.[192,1442,357,552]" pageId="33" pageNumber="32">
<keyLead id="30D795EC8F36487DDFED116EA20239AF" pageId="33" pageNumber="32">
22(21). Pronotum slightly cordate; elytra subdepressed, striae finely punctate; eyes strongly rounded; terminal segment of palpi obtuse apically; 
<figureCitation id="131D2A5C8F36487DDDEB118DA26E39CD" box="[710,789,390,416]" captionStart="Figures 73–76" captionStartId="79.[194,281,1866,1892]" captionTargetBox="[214,1431,210,1834]" captionTargetId="figure-37@79.[192,1439,206,1837]" captionTargetPageId="79" captionText="Figures 73–76. Dorsal habitus. 73) Ctenognathus intermedius. 74) C. otagoensis. 75) C. colensonis. 76) C. earlyi new species. Scale line = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5041861" httpUri="https://zenodo.org/record/5041861/files/figure.png" pageId="33" pageNumber="32">Fig. 74</figureCitation>
. [Body length 
<quantity id="4CDE9B3C8F36487DDC95118DA52039CC" box="[952,1115,390,417]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" metricValueMax="1.27" metricValueMin="1.03" pageId="33" pageNumber="32" unit="mm" value="11.5" valueMax="12.7" valueMin="10.3">10.3–12.7 mm</quantity>
; southern South Island (OL– DN to FD–SL)] 
<taxonomicName id="4C264D5A8F36487DDD2911A3A20239AF" ID-CoL="325SX" authority="(Bates)" baseAuthorityName="Bates" baseAuthorityYear="1878" box="[516,889,424,451]" class="Insecta" family="Carabidae" genus="Ctenognathus" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="32" phylum="Arthropoda" rank="species" species="otagoensis">
<emphasis id="B952EACB8F36487DDD2911A3A20239AF" bold="true" box="[516,889,424,451]" pageId="33" pageNumber="32">
<emphasis id="B952EACB8F36487DDD2911A3A25A39AF" bold="true" box="[516,801,424,451]" italics="true" pageId="33" pageNumber="32">Ctenognathus otagoensis</emphasis>
(Bates)
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B9936D98F36487DDFED11C1A4E63A4A" blockId="33.[192,1442,357,552]" pageId="33" pageNumber="32">
<keyLead id="30D795EC8F36487DDFED11C1A4E63A4A" pageId="33" pageNumber="32">
— Pronotum subquadrate; elytra moderately convex, striae impunctate; eyes moderately rounded; terminal segment of palpi truncate apically; 
<figureCitation id="131D2A5C8F36487DDDF911E0A2593A68" box="[724,802,491,517]" captionStart="Figures 73–76" captionStartId="79.[194,281,1866,1892]" captionTargetBox="[214,1431,210,1834]" captionTargetId="figure-37@79.[192,1439,206,1837]" captionTargetPageId="79" captionText="Figures 73–76. Dorsal habitus. 73) Ctenognathus intermedius. 74) C. otagoensis. 75) C. colensonis. 76) C. earlyi new species. Scale line = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5041861" httpUri="https://zenodo.org/record/5041861/files/figure.png" pageId="33" pageNumber="32">Fig. 76</figureCitation>
. [Body length 
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; Stewart Island]........................................... 
<emphasis id="B952EACB8F36487DDDD31206A4E63A4A" bold="true" box="[766,1437,524,552]" pageId="33" pageNumber="32">
<taxonomicName id="4C264D5A8F36487DDDD31206A4703A4A" ID-CoL="8M979" authority="Larochelle and Lariviere" authorityName="Larochelle and Lariviere E" authorityYear="2021" box="[766,1291,524,552]" class="Insecta" family="Carabidae" genus="Ctenognathus" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="32" phylum="Arthropoda" rank="species" species="earlyi" status="sp. nov.">
<emphasis id="B952EACB8F36487DDDD31206A29D3A45" bold="true" box="[766,998,525,552]" italics="true" pageId="33" pageNumber="32">Ctenognathus earlyi</emphasis>
Larochelle and Larivière
</taxonomicName>
, 
<taxonomicNameLabel id="A26157B08F36487DDA3B1206A4E63A4A" box="[1302,1437,525,551]" pageId="33" pageNumber="32" rank="species">new species</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D22E7C8F36487DDFED123CA4E63B32" pageId="33" pageNumber="32">
<paragraph id="8B9936D98F36487DDFED123CA4E63AB5" blockId="33.[192,1441,567,863]" pageId="33" pageNumber="32">
<keyLead id="30D795EC8F36487DDFED123CA4E63AB5" pageId="33" pageNumber="32">
23(20). Antennal segments 1–3 rufotestaceous, 4–11 dark piceous; femora rufotestaceous; sides of pronotum not sinuate posteriorly, lateral depressions wide throughout, laterobasal foveae very wide (almost meeting toward pronotal midline); elytra with sides slightly rounded, interval 3 with three very deep, large (foveate) setiferous punctures; 
<figureCitation id="131D2A5C8F36487DDC0D1297A2153ADA" box="[800,878,668,695]" captionStart="Figures 77–79" captionStartId="80.[192,279,1866,1892]" captionTargetBox="[215,1432,219,1836]" captionTargetId="figure-43@80.[193,1440,206,1837]" captionTargetPageId="80" captionText="Figures 77–79. Dorsal habitus. 77) Ctenognathus davidsoni new species. 78) C. oreobius. 79) C. hoarei new species. Scale line = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5041865" httpUri="https://zenodo.org/record/5041865/files/figure.png" pageId="33" pageNumber="32">Fig. 77</figureCitation>
. [Body length 
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; western South Island (BR–SL)]..................... 
<emphasis id="B952EACB8F36487DDDE212B6A4E63AB5" bold="true" box="[719,1437,701,728]" pageId="33" pageNumber="32">
<taxonomicName id="4C264D5A8F36487DDDE212B6A4703AB5" ID-CoL="8MDS3" authority="Larochelle and Lariviere" authorityName="Larochelle and Lariviere E" authorityYear="2021" box="[719,1291,701,728]" class="Insecta" family="Carabidae" genus="Ctenognathus" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="32" phylum="Arthropoda" rank="species" species="davidsoni" status="sp. nov.">
<emphasis id="B952EACB8F36487DDDE212B6A29D3AB5" bold="true" box="[719,998,701,728]" italics="true" pageId="33" pageNumber="32">Ctenognathus davidsoni</emphasis>
Larochelle and Larivière
</taxonomicName>
, 
<taxonomicNameLabel id="A26157B08F36487DDA3B12B5A4E63AB5" box="[1302,1437,702,728]" pageId="33" pageNumber="32" rank="species">new species</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8B9936D98F36487DDFED12EBA4E63B32" blockId="33.[192,1441,567,863]" pageId="33" pageNumber="32">
<keyLead id="30D795EC8F36487DDFED12EBA4E63B32" pageId="33" pageNumber="32">
— Antennae entirely dark rufous or rufotestaceous; femora dark yellow; sides of pronotum moderately sinuate posteriorly, lateral depressions narrow anteriorly and widened posteriorly, laterobasal foveae moderately wide (well separated from each other); elytra with sides moderately rounded, interval 3 with three shallow, minute setiferous punctures......................................... 
<emphasis id="B952EACB8F36487DDAAC134EA4E63B32" bold="true" box="[1409,1437,837,863]" pageId="33" pageNumber="32">24</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D22E7C8F36487DDFED1364A4E63CFB" pageId="33" pageNumber="32">
<paragraph id="8B9936D98F36487DDFED1364A4E53B80" blockId="33.[192,1441,878,1175]" pageId="33" pageNumber="32">
<keyLead id="30D795EC8F36487DDFED1364A4E53B80" pageId="33" pageNumber="32">
24(23). Antennae dark rufous; terminal segment of palpi obtuse apically; eyes strongly convex; pronotum obsoletely wrinkled on disc, moderately cordate, posterolateral angles rectangular; elytra oblong, sides moderately rounded, striae deep and impunctate; 
<figureCitation id="131D2A5C8F36487DDCAC13B9A2AB3BA1" box="[897,976,946,972]" captionStart="Figures 77–79" captionStartId="80.[192,279,1866,1892]" captionTargetBox="[215,1432,219,1836]" captionTargetId="figure-43@80.[193,1440,206,1837]" captionTargetPageId="80" captionText="Figures 77–79. Dorsal habitus. 77) Ctenognathus davidsoni new species. 78) C. oreobius. 79) C. hoarei new species. Scale line = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5041865" httpUri="https://zenodo.org/record/5041865/files/figure.png" pageId="33" pageNumber="32">Fig. 78</figureCitation>
. [Body length 10.5–13.00 mm; southeastern South Island (MK–SL)]................................. 
<taxonomicName id="4C264D5A8F36487DDB1E13D8A4E53B80" ID-CoL="325SW" authority="(Broun)" baseAuthorityName="Broun" baseAuthorityYear="1886" box="[1075,1438,979,1006]" class="Insecta" family="Carabidae" genus="Ctenognathus" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="32" phylum="Arthropoda" rank="species" species="oreobius">
<emphasis id="B952EACB8F36487DDB1E13D8A4E53B80" bold="true" box="[1075,1438,979,1006]" pageId="33" pageNumber="32">
<emphasis id="B952EACB8F36487DDB1E13D8A4423B83" bold="true" box="[1075,1337,979,1006]" italics="true" pageId="33" pageNumber="32">Ctenognathus oreobius</emphasis>
(Broun)
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B9936D98F36487DDFED13FDA4E63CFB" blockId="33.[192,1441,878,1175]" pageId="33" pageNumber="32">
<keyLead id="30D795EC8F36487DDFED13FDA4E63CFB" pageId="33" pageNumber="32">
— Antennae rufotestaceous; terminal segment of palpi truncate apically; eyes moderately convex; pronotum deeply wrinkled on disc, slightly cordate, posterolateral angles acute and slightly projected laterally; elytra subovate, sides strongly rounded, striae shallow and finely punctate; 
<figureCitation id="131D2A5C8F36487DDA2E1432A4293C3E" box="[1283,1362,1081,1107]" captionStart="Figures 77–79" captionStartId="80.[192,279,1866,1892]" captionTargetBox="[215,1432,219,1836]" captionTargetId="figure-43@80.[193,1440,206,1837]" captionTargetPageId="80" captionText="Figures 77–79. Dorsal habitus. 77) Ctenognathus davidsoni new species. 78) C. oreobius. 79) C. hoarei new species. Scale line = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5041865" httpUri="https://zenodo.org/record/5041865/files/figure.png" pageId="33" pageNumber="32">Fig. 79</figureCitation>
. [Body length 
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; Banks Peninsula (MC)]............................................................................. 
<emphasis id="B952EACB8F36487DDDDA1477A4E63CFB" bold="true" box="[759,1437,1147,1175]" pageId="33" pageNumber="32">
<taxonomicName id="4C264D5A8F36487DDDDA1477A4703CFB" ID-CoL="8M8B3" authority="Larochelle and Lariviere" authorityName="Larochelle and Lariviere E" authorityYear="2021" box="[759,1291,1147,1175]" class="Insecta" family="Carabidae" genus="Ctenognathus" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="32" phylum="Arthropoda" rank="species" species="hoarei" status="sp. nov.">
<emphasis id="B952EACB8F36487DDDDA1477A29D3CFA" bold="true" box="[759,998,1148,1175]" italics="true" pageId="33" pageNumber="32">Ctenognathus hoarei</emphasis>
Larochelle and Larivière
</taxonomicName>
, 
<taxonomicNameLabel id="A26157B08F36487DDA3B1477A4E63CFB" box="[1302,1437,1148,1174]" pageId="33" pageNumber="32" rank="species">new species</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>