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<document id="E5B2E3B9C9DED7D084ABB0922A80B9EC" ID-CLB-Dataset="92619" ID-DOI="10.5852/ejt.2022.815.1749" ID-GBIF-Dataset="9a798cae-f10e-44c2-a65c-cdadbff28cb4" ID-ISSN="2118-9773" ID-Zenodo-Dep="6484153" ID-ZooBank="6AD0082E-7349-48DE-AFCA-1EE0BFBB3887" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1650913068508" checkinUser="felipe" docAuthor="Vranken, Nathan, Steenberge, Maarten Van, Heylen, Annelies, Decru, Eva &amp; Snoeks, Jos" docDate="2022" docId="038F87A6611BDA79FDB954EB24ADFD74" docLanguage="en" docName="ejt-815_1-94.pdf" docOrigin="European Journal of Taxonomy 815" docSource="http://dx.doi.org/10.5852/ejt.2022.815.1749" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Haplochromis simba Vranken &amp; Steenberge &amp; Heylen &amp; Decru &amp; Snoeks 2022, sp. nov." docType="treatment" docUuid="266E8987-E220-4EC4-AD72-0A46BE085EC8" docUuidSource="ZooBank" docVersion="10" lastPageNumber="34" masterDocId="FFB6FFDE6107DA58FFD257322629FC0A" masterDocTitle="From a pair to a dozen: the piscivorous species of Haplochromis (Cichlidae) from the Lake Edward system" masterLastPageNumber="94" masterPageNumber="1" pageNumber="29" updateTime="1699329164302" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="2CE2585BCB8CF0D62BC6208727776C00">From a pair to a dozen: the piscivorous species of Haplochromis (Cichlidae) from the Lake Edward system</mods:title>
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<mods:namePart id="C4FA99AB6080919B533FAD657FD42581">Vranken, Nathan</mods:namePart>
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<mods:affiliation id="8F168CFDFFE7504CF53E3F7D1472BC7F">Leuven, Laboratory of Biodiversity and Evolutionary Genomics, Department of Biology, Charles Deberiotstraat 32, 3000 Leuven, Belgium. &amp; Royal Museum for Central Africa, Biology department, Section Vertebrates, Leuvensesteenweg 13, 3080 Tervuren, Belgium.</mods:affiliation>
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<mods:namePart id="8CCAEC3140E3AE5E6325FD8B521CABC2">Steenberge, Maarten Van</mods:namePart>
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<mods:affiliation id="02A4414859CB41E3DBA8DAE6C679567D">Royal Belgian Institute of Natural Sciences, Operational Directorate Taxonomy and Phylogeny, Vautierstraat 29, 1000 Brussels, Belgium. &amp; Masaryk University, Department of Botany and Zoology, Kotlářská 2, 611 37 Brno, Czech Republic.</mods:affiliation>
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<mods:namePart id="D8DFD19F53E73525F1779F765C9C9316">Heylen, Annelies</mods:namePart>
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<mods:affiliation id="0AB60CCB0A47E890F1B8D697AFA72582">KU Leuven, Laboratory of Biodiversity and Evolutionary Genomics, Department of Biology, Charles Deberiotstraat 32, 3000 Leuven, Belgium.</mods:affiliation>
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<mods:namePart id="8DF4AB98BE59A510DCF83BE96E156F5A">Decru, Eva</mods:namePart>
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<mods:affiliation id="B9CFB5ABB94AE73C38B7E8C9EAB54991">KU Leuven, Laboratory of Biodiversity and Evolutionary Genomics, Department of Biology, Charles Deberiotstraat 32, 3000 Leuven, Belgium.</mods:affiliation>
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<mods:affiliation id="D6B127558C9D10D6D5AF16A1853B9EA0">Royal Museum for Central Africa, Biology department, Section Vertebrates, Leuvensesteenweg 13, 3080 Tervuren, Belgium.</mods:affiliation>
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<heading id="D0D181DC611BDA44FDB954EB25E0FFFE" box="[619,969,985,1012]" centered="true" fontSize="11" level="2" pageId="28" pageNumber="29" reason="2">
<taxonomicName id="4C264D33611BDA44FDB954EB254EFFFE" authority="Vranken &amp; Steenberge &amp; Heylen &amp; Decru &amp; Snoeks, 2022" authorityName="Vranken &amp; Steenberge &amp; Heylen &amp; Decru &amp; Snoeks" authorityYear="2022" box="[619,871,985,1012]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="simba" status="sp. nov.">
<emphasis id="B952EAA2611BDA44FDB954EB254EFFFE" bold="true" box="[619,871,985,1012]" italics="true" pageId="28" pageNumber="29">Haplochromis simba</emphasis>
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<taxonomicNameLabel id="A26157D9611BDA44FCBC54E825E0FFFE" box="[878,969,986,1012]" pageId="28" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
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<subSubSection id="C33C653B611BDA44FD5A53112585F837" box="[648,940,1059,1085]" pageId="28" pageNumber="29" type="description">
<paragraph id="8B9936B0611BDA44FD5A53112585F837" blockId="28.[367,1220,985,1085]" box="[648,940,1059,1085]" pageId="28" pageNumber="29">
<figureCitation id="131D2A35611BDA44FD5A531124D9F837" box="[648,752,1059,1085]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="5.[190,234,1822,1848]" captionStartId-1="6.[189,231,1892,1918]" captionTargetBox-0="[193,1398,734,1726]" captionTargetBox-1="[208,1335,272,1830]" captionTargetId-0="figure-217@5.[189,1399,730,1726]" captionTargetId-1="figure-18@6.[208,1335,272,1830]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="Fig. 1. The Lake Edward system, consisting of Lakes Edward and George that are connected through the Kazinga Channel. The catch localities of all examined specimens are indicated: Haplochromis sp. latifrons (), H. mentatus Regan, 1925 (), H. sp. rex (), H. sp. simba (▼), H. sp. glaucus (), H. sp. aquila (), H. sp. kimondo (●), H. sp. falcatus (), H. sp. curvidens (), H. sp. pardus (), H. sp. quasimodo (▲), and H. squamipinnis Regan, 1921 (█). The type localities of H. mentatus and H. squamipinnis are not shown as the exact locations are unknown." captionText-1="Fig. 2. a. Differences in total number of outer teeth and premaxillary dentition between the macro- and microdontic groups of piscivores from the Lake Edward system.b. Schematic presentation of the dominant male colour patterns in life, in the various groups studied. For H. sp. latifrons the female and for H. sp. curvidens the non-dominant colour patterns are shown, as those of dominant males are unknown." figureDoi-0="http://doi.org/10.5281/zenodo.6484155" figureDoi-1="http://doi.org/10.5281/zenodo.6484157" httpUri-0="https://zenodo.org/record/6484155/files/figure.png" httpUri-1="https://zenodo.org/record/6484157/files/figure.png" pageId="28" pageNumber="29">Figs 12</figureCitation>
,
<figureCitation id="131D2A35611BDA44FD2E53112563F837" box="[764,842,1059,1085]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="30.[189,232,1963,1989]" captionStartId-1="31.[189,232,1892,1918]" captionStartId-2="32.[189,232,1963,1989]" captionTargetBox-0="[189,1398,1326,1868]" captionTargetBox-1="[189,1398,271,1801]" captionTargetBox-2="[189,1397,960,1907]" captionTargetId-0="figure-310@30.[189,1399,1326,1868]" captionTargetId-1="figure-12@31.[186,1402,265,1803]" captionTargetId-2="figure-181@32.[189,1398,960,1909]" captionTargetPageId-0="30" captionTargetPageId-1="31" captionTargetPageId-2="32" captionText-0="Fig. 14. Haplochromis simba sp. nov., holotype, ♂, 105.8 mm SL (RMCA 2016.035.P.0225). Drawn by N. Vranken." captionText-1="Fig. 15. Haplochromis simba sp. nov. a. Photograph of preserved holotype (RMCA 2016.035.P.0225; 105.8 mm SL). b. X-ray image of holotype. cd. Photographs of freshly caught specimens. c. Dominant male (RMCA 2016.035.P.0224; 97.9 mm SL). d. Female (RMCA 2018.008.P.0348; 109.0 mm SL) to illustrate the live colour patterns. The contrast was slightly enhanced." captionText-2="Fig. 16. Haplochromis simba sp. nov. (RMCA 2016.035.P.0223; 105.5 mm SL). a. Dorsal view of the lower pharyngeal jaw. b. Lateral view of the lower pharyngeal jaw." figureDoi-0="http://doi.org/10.5281/zenodo.6484197" figureDoi-1="http://doi.org/10.5281/zenodo.6484199" figureDoi-2="http://doi.org/10.5281/zenodo.6484201" httpUri-0="https://zenodo.org/record/6484197/files/figure.png" httpUri-1="https://zenodo.org/record/6484199/files/figure.png" httpUri-2="https://zenodo.org/record/6484201/files/figure.png" pageId="28" pageNumber="29">1416</figureCitation>
;
<tableCitation id="C6A4030B611BDA44FC8753112585F837" box="[853,940,1059,1085]" captionStart="Table 1" captionStartId="9.[182,252,548,576]" captionTargetPageId="9" captionText="Table 1. (continued on the next three pages). Measurements and counts of the piscivorous species of Haplochromis from the Lake Edward system." httpUri="http://table.plazi.org/id/DF596638610EDA51FF6455162158FE4A" pageId="28" pageNumber="29" tableUuid="DF596638610EDA51FF6455162158FE4A">Table 1</tableCitation>
</paragraph>
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<subSubSection id="C33C653B611BDA45FF6F53B22374FE1E" lastPageId="29" lastPageNumber="30" pageId="28" pageNumber="29" type="diagnosis">
<paragraph id="8B9936B0611BDA44FF6F53B227C8F897" blockId="28.[189,481,1152,1181]" box="[189,481,1152,1181]" pageId="28" pageNumber="29">
<emphasis id="B952EAA2611BDA44FF6F53B227C8F897" bold="true" box="[189,481,1152,1181]" pageId="28" pageNumber="29">Differential diagnosis</emphasis>
</paragraph>
<paragraph id="8B9936B0611BDA44FF6F53852439F910" blockId="28.[189,1399,1207,1307]" pageId="28" pageNumber="29">Species with a piscivorous morphology; premaxillary pedicel and mental prominences strongly developed; outer oral teeth few and large [UOT 2231 (median 27)]; dominant males yellow with an orange anterior part of flank.</paragraph>
<paragraph id="8B9936B0611BDA44FF6F527126DEFAAC" blockId="28.[188,1399,1347,1702]" pageId="28" pageNumber="29">
Amongst piscivorous species from the Lake Edward system,
<taxonomicName id="4C264D33611BDA44FC5B527625DDF957" authority="Vranken &amp; Steenberge &amp; Heylen &amp; Decru &amp; Snoeks, 2022" authorityName="Vranken &amp; Steenberge &amp; Heylen &amp; Decru &amp; Snoeks" authorityYear="2022" box="[905,1012,1347,1373]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="simba" status="sp. nov.">
<emphasis id="B952EAA2611BDA44FC5B527625B7F957" box="[905,926,1348,1373]" italics="true" pageId="28" pageNumber="29">H</emphasis>
.
<emphasis id="B952EAA2611BDA44FC7F527125DDF957" box="[941,1012,1347,1373]" italics="true" pageId="28" pageNumber="29">simba</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26157D9611BDA44FC295271227CF957" box="[1019,1109,1347,1373]" pageId="28" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
differs from
<taxonomicName id="4C264D33611BDA44FB3D5276235EF957" authority="Vranken &amp; Steenberge &amp; Heylen &amp; Decru &amp; Snoeks, 2022" authorityName="Vranken &amp; Steenberge &amp; Heylen &amp; Decru &amp; Snoeks" authorityYear="2022" box="[1263,1399,1347,1373]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="latifrons" status="sp. nov.">
<emphasis id="B952EAA2611BDA44FB3D5276232DF957" box="[1263,1284,1348,1373]" italics="true" pageId="28" pageNumber="29">H</emphasis>
.
<emphasis id="B952EAA2611BDA44FAC15271235EF957" box="[1299,1399,1347,1373]" italics="true" pageId="28" pageNumber="29">latifrons</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26157D9611BDA44FF6F525A2733F988" box="[189,282,1384,1410]" pageId="28" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="4C264D33611BDA44FEF4525A279CF98B" authorityName="Regan" authorityYear="1925" box="[294,437,1384,1410]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="mentatus">
<emphasis id="B952EAA2611BDA44FEF4525A2712F98B" box="[294,315,1384,1409]" italics="true" pageId="28" pageNumber="29">H</emphasis>
.
<emphasis id="B952EAA2611BDA44FE98525A279CF98B" box="[330,437,1384,1409]" italics="true" pageId="28" pageNumber="29">mentatus</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C264D33611BDA44FE24525A245EF98B" authority="Vranken &amp; Steenberge &amp; Heylen &amp; Decru &amp; Snoeks, 2022" authorityName="Vranken &amp; Steenberge &amp; Heylen &amp; Decru &amp; Snoeks" authorityYear="2022" box="[502,631,1383,1410]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="glaucus" status="sp. nov.">
<emphasis id="B952EAA2611BDA44FE24525A2422F98B" box="[502,523,1384,1409]" italics="true" pageId="28" pageNumber="29">H</emphasis>
.
<emphasis id="B952EAA2611BDA44FDC95255245EF98B" box="[539,631,1383,1409]" italics="true" pageId="28" pageNumber="29">glaucus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26157D9611BDA44FDAC525A24F0F988" box="[638,729,1384,1410]" pageId="28" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
by the combination of a narrower interorbital area [IOW 45.550.4 (mean 48.1) vs 50.963.3 (53.860.0) % HW] and a strongly developed vs absent or weakly developed mental prominence. It further differs from
<taxonomicName id="4C264D33611BDA44FCFA52832599F9C0" authority="Vranken &amp; Steenberge &amp; Heylen &amp; Decru &amp; Snoeks, 2022" authorityName="Vranken &amp; Steenberge &amp; Heylen &amp; Decru &amp; Snoeks" authorityYear="2022" box="[808,944,1456,1483]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="latifrons" status="sp. nov.">
<emphasis id="B952EAA2611BDA44FCFA52832514F9C0" box="[808,829,1457,1482]" italics="true" pageId="28" pageNumber="29">H</emphasis>
.
<emphasis id="B952EAA2611BDA44FC9E52822599F9C0" box="[844,944,1456,1482]" italics="true" pageId="28" pageNumber="29">latifrons</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26157D9611BDA44FC6752832226F9C1" box="[949,1039,1457,1483]" pageId="28" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
by a longer anal fin base [AFB 17.319.0 (18.0) vs 14.717.3 (15.7) % SL] and absence vs presence of a well-defined mid-lateral band; further from
<taxonomicName id="4C264D33611BDA44FE8052C827C8FA19" authorityName="Regan" authorityYear="1925" box="[338,481,1530,1556]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="mentatus">
<emphasis id="B952EAA2611BDA44FE8052C8274EFA19" box="[338,359,1530,1555]" italics="true" pageId="28" pageNumber="29">H</emphasis>
.
<emphasis id="B952EAA2611BDA44FEA452C827C8FA19" box="[374,481,1530,1555]" italics="true" pageId="28" pageNumber="29">mentatus</emphasis>
</taxonomicName>
by a smaller number of outer upper jaw teeth [UOT 2231 (27) vs 2846 (36)] and dominant males yellow with an orange anterior part of flank vs green-yellow with a red anterior part of flank; further from
<taxonomicName id="4C264D33611BDA44FE1251712468FA57" authority="Vranken &amp; Steenberge &amp; Heylen &amp; Decru &amp; Snoeks, 2022" authorityName="Vranken &amp; Steenberge &amp; Heylen &amp; Decru &amp; Snoeks" authorityYear="2022" box="[448,577,1603,1629]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="glaucus" status="sp. nov.">
<emphasis id="B952EAA2611BDA44FE12517127FCFA56" box="[448,469,1603,1628]" italics="true" pageId="28" pageNumber="29">H</emphasis>
.
<emphasis id="B952EAA2611BDA44FE3751712468FA57" box="[485,577,1603,1629]" italics="true" pageId="28" pageNumber="29">glaucus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26157D9611BDA44FD985171248CFA57" box="[586,677,1603,1629]" pageId="28" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
by a narrower lower pharyngeal element [LPW 83.887.9 vs 93.395.1% LPL] and dominant males yellow with an orange anterior part of flank vs uniformly lightblue.
</paragraph>
<paragraph id="8B9936B0611BDA44FF6F51FD2422FBE2" blockId="28.[188,1399,1742,2025]" pageId="28" pageNumber="29">
It differs from
<taxonomicName id="4C264D33611BDA44FEBF51FD279FFAE2" authority="Vranken &amp; Steenberge &amp; Heylen &amp; Decru &amp; Snoeks, 2022" authorityName="Vranken &amp; Steenberge &amp; Heylen &amp; Decru &amp; Snoeks" authorityYear="2022" box="[365,438,1743,1769]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="rex" status="sp. nov.">
<emphasis id="B952EAA2611BDA44FEBF51FD27ABFAE2" box="[365,386,1743,1768]" italics="true" pageId="28" pageNumber="29">H</emphasis>
.
<emphasis id="B952EAA2611BDA44FE4351FD279FFAE2" box="[401,438,1743,1768]" italics="true" pageId="28" pageNumber="29">rex</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26157D9611BDA44FE6D51FD2433FAE3" box="[447,538,1743,1769]" pageId="28" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="4C264D33611BDA44FD8651FD24EDFAE2" authority="Vranken &amp; Steenberge &amp; Heylen &amp; Decru &amp; Snoeks, 2022" authorityName="Vranken &amp; Steenberge &amp; Heylen &amp; Decru &amp; Snoeks" authorityYear="2022" box="[596,708,1742,1769]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="aquila" status="sp. nov.">
<emphasis id="B952EAA2611BDA44FD8651FD2440FAE2" box="[596,617,1743,1768]" italics="true" pageId="28" pageNumber="29">H</emphasis>
.
<emphasis id="B952EAA2611BDA44FDAA51FC24EDFAE2" box="[632,708,1742,1768]" italics="true" pageId="28" pageNumber="29">aquila</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26157D9611BDA44FD1F51FD2501FAE3" box="[717,808,1743,1769]" pageId="28" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
by the combination of a smaller number of scales between first anal-fin spine and upper lateral line (ULL-A 911 vs 1216, rarely 11) and a strongly developed vs absent or weakly developed mental prominence. It further differs from
<taxonomicName id="4C264D33611BDA44FB11502A2326FB3B" authority="Vranken &amp; Steenberge &amp; Heylen &amp; Decru &amp; Snoeks, 2022" authorityName="Vranken &amp; Steenberge &amp; Heylen &amp; Decru &amp; Snoeks" authorityYear="2022" box="[1219,1295,1816,1842]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="rex" status="sp. nov.">
<emphasis id="B952EAA2611BDA44FB11502A22F1FB3B" box="[1219,1240,1816,1841]" italics="true" pageId="28" pageNumber="29">H</emphasis>
.
<emphasis id="B952EAA2611BDA44FB38502A2326FB3B" box="[1258,1295,1816,1841]" italics="true" pageId="28" pageNumber="29">rex</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26157D9611BDA44FACB502A235EFB38" box="[1305,1399,1816,1842]" pageId="28" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
by a larger number of longitudinal line scales (LongL 3233 vs 3438) and dominant males uniformly yellow with an orange anterior part of flank vs cream-coloured with an orange operculum and a light blue snout; further from
<taxonomicName id="4C264D33611BDA44FE0B50B42460FB95" authority="Vranken &amp; Steenberge &amp; Heylen &amp; Decru &amp; Snoeks, 2022" authorityName="Vranken &amp; Steenberge &amp; Heylen &amp; Decru &amp; Snoeks" authorityYear="2022" box="[473,585,1925,1952]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="aquila" status="sp. nov.">
<emphasis id="B952EAA2611BDA44FE0B50B427C7FB95" box="[473,494,1926,1951]" italics="true" pageId="28" pageNumber="29">H</emphasis>
.
<emphasis id="B952EAA2611BDA44FE2F50B72460FB95" box="[509,585,1925,1951]" italics="true" pageId="28" pageNumber="29">aquila</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26157D9611BDA44FD9D50B42480FBAA" box="[591,681,1926,1952]" pageId="28" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
by a smaller eye [ED 26.729.5 (28.3) vs 30.031.5 (30.6) % HL] and dominant males yellow with an orange anterior part of flank and a yellow head vs uniformly light grey with a black head.
</paragraph>
<paragraph id="8B9936B0611ADA45FF6F563B2278FD60" blockId="29.[189,1399,265,362]" pageId="29" pageNumber="30">
It differs from
<taxonomicName id="4C264D33611ADA45FEA156382428FD29" authority="Vranken &amp; Steenberge &amp; Heylen &amp; Decru &amp; Snoeks, 2022" authorityName="Vranken &amp; Steenberge &amp; Heylen &amp; Decru &amp; Snoeks" authorityYear="2022" box="[371,513,265,291]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="kimondo" status="sp. nov.">
<emphasis id="B952EAA2611ADA45FEA1563827A1FD29" box="[371,392,266,291]" italics="true" pageId="29" pageNumber="30">H</emphasis>
.
<emphasis id="B952EAA2611ADA45FE49563B2428FD29" box="[411,513,265,291]" italics="true" pageId="29" pageNumber="30">kimondo</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26157D9611ADA45FDD9563B2445FD29" box="[523,620,265,291]" pageId="29" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="4C264D33611ADA45FDA956382529FD29" authority="Vranken &amp; Steenberge &amp; Heylen &amp; Decru &amp; Snoeks, 2022" authorityName="Vranken &amp; Steenberge &amp; Heylen &amp; Decru &amp; Snoeks" authorityYear="2022" box="[635,768,265,291]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="falcatus" status="sp. nov.">
<emphasis id="B952EAA2611ADA45FDA9563824B9FD29" box="[635,656,266,291]" italics="true" pageId="29" pageNumber="30">H</emphasis>
.
<emphasis id="B952EAA2611ADA45FD70563B2529FD29" box="[674,768,265,291]" italics="true" pageId="29" pageNumber="30">falcatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26157D9611ADA45FCD9563B2542FD29" box="[779,875,265,291]" pageId="29" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="4C264D33611ADA45FCA85638223DFD29" authority="Vranken &amp; Steenberge &amp; Heylen &amp; Decru &amp; Snoeks, 2022" authorityName="Vranken &amp; Steenberge &amp; Heylen &amp; Decru &amp; Snoeks" authorityYear="2022" box="[890,1044,265,291]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="curvidens" status="sp. nov.">
<emphasis id="B952EAA2611ADA45FCA8563825A6FD29" box="[890,911,266,291]" italics="true" pageId="29" pageNumber="30">H</emphasis>
.
<emphasis id="B952EAA2611ADA45FC73563B223DFD29" box="[929,1044,265,291]" italics="true" pageId="29" pageNumber="30">curvidens</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26157D9611ADA45FBCD563B22A9FD29" box="[1055,1152,265,291]" pageId="29" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="4C264D33611ADA45FB5C5638232EFD29" authority="Vranken &amp; Steenberge &amp; Heylen &amp; Decru &amp; Snoeks, 2022" authorityName="Vranken &amp; Steenberge &amp; Heylen &amp; Decru &amp; Snoeks" authorityYear="2022" box="[1166,1287,265,291]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="pardus" status="sp. nov.">
<emphasis id="B952EAA2611ADA45FB5C5638228AFD29" box="[1166,1187,266,291]" italics="true" pageId="29" pageNumber="30">H</emphasis>
.
<emphasis id="B952EAA2611ADA45FB64563B232EFD29" box="[1206,1287,265,291]" italics="true" pageId="29" pageNumber="30">pardus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26157D9611ADA45FAC0563B235BFD29" box="[1298,1394,265,291]" pageId="29" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="4C264D33611ADA45FF6F561F274AFD4C" authority="Vranken &amp; Steenberge &amp; Heylen &amp; Decru &amp; Snoeks, 2022" authorityName="Vranken &amp; Steenberge &amp; Heylen &amp; Decru &amp; Snoeks" authorityYear="2022" box="[189,355,300,327]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="quasimodo" status="sp. nov.">
<emphasis id="B952EAA2611ADA45FF6F561F26FBFD4C" box="[189,210,301,326]" italics="true" pageId="29" pageNumber="30">H</emphasis>
.
<emphasis id="B952EAA2611ADA45FF33561E274AFD4C" box="[225,355,300,326]" italics="true" pageId="29" pageNumber="30">quasimodo</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26157D9611ADA45FEB8561F27EEFD4D" box="[362,455,301,327]" pageId="29" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
, and
<taxonomicName id="4C264D33611ADA45FDD6561F24EFFD4C" authorityName="Regan" authorityYear="1921" box="[516,710,301,327]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="squamipinnis">
<emphasis id="B952EAA2611ADA45FDD6561F2430FD4C" box="[516,537,301,326]" italics="true" pageId="29" pageNumber="30">H</emphasis>
.
<emphasis id="B952EAA2611ADA45FDFA561F24EFFD4C" box="[552,710,301,326]" italics="true" pageId="29" pageNumber="30">squamipinnis</emphasis>
</taxonomicName>
by the combination of large vs small outer oral teeth and a smaller number of outer upper jaw teeth [UOT 2231 (27) vs 3979 (4558)].
</paragraph>
<paragraph id="8B9936B0611ADA45FF6F56A32374FE1E" blockId="29.[189,1400,401,533]" pageId="29" pageNumber="30">
It further differs from
<taxonomicName id="4C264D33611ADA45FE1856A02464FDA1" authority="Vranken &amp; Steenberge &amp; Heylen &amp; Decru &amp; Snoeks, 2022" authorityName="Vranken &amp; Steenberge &amp; Heylen &amp; Decru &amp; Snoeks" authorityYear="2022" box="[458,589,401,427]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="falcatus" status="sp. nov.">
<emphasis id="B952EAA2611ADA45FE1856A027F4FDA1" box="[458,477,402,427]" italics="true" pageId="29" pageNumber="30">H</emphasis>
.
<emphasis id="B952EAA2611ADA45FE3D56A32464FDA1" box="[495,589,401,427]" italics="true" pageId="29" pageNumber="30">falcatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26157D9611ADA45FD8A56A3249EFDA1" box="[600,695,401,427]" pageId="29" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="4C264D33611ADA45FD1756A02575FDA1" authority="Vranken &amp; Steenberge &amp; Heylen &amp; Decru &amp; Snoeks, 2022" authorityName="Vranken &amp; Steenberge &amp; Heylen &amp; Decru &amp; Snoeks" authorityYear="2022" box="[709,860,401,427]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="curvidens" status="sp. nov.">
<emphasis id="B952EAA2611ADA45FD1756A024F1FDA1" box="[709,728,402,427]" italics="true" pageId="29" pageNumber="30">H</emphasis>
.
<emphasis id="B952EAA2611ADA45FD3B56A32575FDA1" box="[745,860,401,427]" italics="true" pageId="29" pageNumber="30">curvidens</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26157D9611ADA45FCB556A325EFFDA1" box="[871,966,401,427]" pageId="29" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
, and
<taxonomicName id="4C264D33611ADA45FBD556A02254FDA1" authority="Vranken &amp; Steenberge &amp; Heylen &amp; Decru &amp; Snoeks, 2022" authorityName="Vranken &amp; Steenberge &amp; Heylen &amp; Decru &amp; Snoeks" authorityYear="2022" box="[1031,1149,401,427]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="pardus" status="sp. nov.">
<emphasis id="B952EAA2611ADA45FBD556A02233FDA1" box="[1031,1050,402,427]" italics="true" pageId="29" pageNumber="30">H</emphasis>
.
<emphasis id="B952EAA2611ADA45FBFE56A32254FDA1" box="[1068,1149,401,427]" italics="true" pageId="29" pageNumber="30">pardus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26157D9611ADA45FB5A56A322CCFDA1" box="[1160,1253,401,427]" pageId="29" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
by a deeper lacrimal [LaD 18.720.5 (19.5) vs 16.018.8 (16.718.0) % HL]; further from
<taxonomicName id="4C264D33611ADA45FB8B568722CAFDC4" authority="Vranken &amp; Steenberge &amp; Heylen &amp; Decru &amp; Snoeks, 2022" authorityName="Vranken &amp; Steenberge &amp; Heylen &amp; Decru &amp; Snoeks" authorityYear="2022" box="[1113,1251,436,463]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="kimondo" status="sp. nov.">
<emphasis id="B952EAA2611ADA45FB8B56872247FDC4" box="[1113,1134,437,462]" italics="true" pageId="29" pageNumber="30">H</emphasis>
.
<emphasis id="B952EAA2611ADA45FBAF568622CAFDC4" box="[1149,1251,436,462]" italics="true" pageId="29" pageNumber="30">kimondo</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26157D9611ADA45FB395687236CFDC5" box="[1259,1349,437,463]" pageId="29" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="4C264D33611ADA45FF6F56EA274FFDFB" authority="Vranken &amp; Steenberge &amp; Heylen &amp; Decru &amp; Snoeks, 2022" authorityName="Vranken &amp; Steenberge &amp; Heylen &amp; Decru &amp; Snoeks" authorityYear="2022" box="[189,358,471,498]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="quasimodo" status="sp. nov.">
<emphasis id="B952EAA2611ADA45FF6F56EA26FBFDFB" box="[189,210,472,497]" italics="true" pageId="29" pageNumber="30">H</emphasis>
.
<emphasis id="B952EAA2611ADA45FF3656E5274FFDFB" box="[228,358,471,497]" italics="true" pageId="29" pageNumber="30">quasimodo</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26157D9611ADA45FEA356EA27E7FDF8" box="[369,462,472,498]" pageId="29" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
by a narrower head [HW 39.541.5 (40.8) vs 42.048.1 (45.145.3) % HL]; further from
<taxonomicName id="4C264D33611ADA45FE8656C9243EFE1E" authorityName="Regan" authorityYear="1921" box="[340,535,507,533]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="squamipinnis">
<emphasis id="B952EAA2611ADA45FE8656C92740FE1E" box="[340,361,507,532]" italics="true" pageId="29" pageNumber="30">H</emphasis>
.
<emphasis id="B952EAA2611ADA45FEAB56C9243EFE1E" box="[377,535,507,532]" italics="true" pageId="29" pageNumber="30">squamipinnis</emphasis>
</taxonomicName>
by absence vs presence of minute scales on proximal part of dorsal fin.
</paragraph>
</subSubSection>
<subSubSection id="C33C653B611ADA45FF6F550E247CFEAD" pageId="29" pageNumber="30" type="etymology">
<paragraph id="8B9936B0611ADA45FF6F550E276DFE5C" blockId="29.[189,324,572,598]" box="[189,324,572,598]" pageId="29" pageNumber="30">
<heading id="D0D181DC611ADA45FF6F550E276DFE5C" bold="true" box="[189,324,572,598]" fontSize="11" level="3" pageId="29" pageNumber="30" reason="3">
<emphasis id="B952EAA2611ADA45FF6F550E276DFE5C" bold="true" box="[189,324,572,598]" pageId="29" pageNumber="30">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8B9936B0611ADA45FF6F5558247CFEAD" blockId="29.[189,1399,618,680]" pageId="29" pageNumber="30">Specific name from Swahili simba for lion; referring to yellow body, orange cheeks that resemble manes, and predatory morphology.</paragraph>
</subSubSection>
<subSubSection id="C33C653B611ADA45FF6F55FD25EDF8F1" pageId="29" pageNumber="30" type="materials_examined">
<paragraph id="8B9936B0611ADA45FF6F55FD2783FEE3" blockId="29.[189,426,719,745]" box="[189,426,719,745]" pageId="29" pageNumber="30">
<heading id="D0D181DC611ADA45FF6F55FD2783FEE3" bold="true" box="[189,426,719,745]" fontSize="11" level="3" pageId="29" pageNumber="30" reason="3">
<emphasis id="B952EAA2611ADA45FF6F55FD2783FEE3" bold="true" box="[189,426,719,745]" pageId="29" pageNumber="30">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8B9936B0611ADA45FF3855CC25EDFF55" blockId="29.[189,1399,766,863]" pageId="29" pageNumber="30">
<materialsCitation id="3B4E3CED611ADA45FF3855CC25EDFF55" ID-GBIF-Occurrence="3861203350" collectingDate="2016-11-09" collectionCode="RMCA" collectorName="HIPE 1 exped." country="Uganda" latitude="-0.40444446" location="Rwenshama" longLatPrecision="1" longitude="29.773556" pageId="29" pageNumber="30" specimenCode="RMCA 2016.035.P.0225" specimenCount="1" specimenCount-male="1" stateProvince="Lake Edward" typeStatus="holotype">
<emphasis id="B952EAA2611ADA45FF3855CC2775FF12" bold="true" box="[234,348,766,792]" pageId="29" pageNumber="30">
<typeStatus id="549D8812611ADA45FF3855CC2775FF12" box="[234,348,766,792]" pageId="29" pageNumber="30" type="holotype">Holotype</typeStatus>
</emphasis>
<collectingCountry id="F3317620611ADA45FF6F54132713FF31" box="[189,314,801,827]" name="Uganda" pageId="29" pageNumber="30">UGANDA</collectingCountry>
<specimenCount id="9D20FD39611ADA45FE8654132745FF31" box="[340,364,801,827]" pageId="29" pageNumber="30" type="male"></specimenCount>
,
<quantity id="4CDE9B55611ADA45FEAB541327D8FF31" box="[377,497,801,828]" metricMagnitude="-1" metricUnit="m" metricValue="1.058" pageId="29" pageNumber="30" unit="mm" value="105.8">105.8 mm</quantity>
SL;
<collectingRegion id="49E2F852611ADA45FDF8541324E2FF31" box="[554,715,801,827]" pageId="29" pageNumber="30">Lake Edward</collectingRegion>
;
<geoCoordinate id="EE125077611ADA45FD05542D2547FF31" box="[727,878,799,827]" degrees="0" direction="south" minutes="24" orientation="latitude" pageId="29" pageNumber="30" precision="1" seconds="16.0" value="-0.40444446">0°2416.0″ S</geoCoordinate>
,
<geoCoordinate id="EE125077611ADA45FCAE542D220CFF31" box="[892,1061,799,827]" degrees="29" direction="east" minutes="46" orientation="longitude" pageId="29" pageNumber="30" precision="1" seconds="24.8" value="29.773556">29°4624.8″ E</geoCoordinate>
;
<date id="FF981070611ADA45FBE3541322ECFF31" box="[1073,1221,801,827]" pageId="29" pageNumber="30" value="2016-11-09">
<collectingDate id="EFDCE998611ADA45FBE3541322ECFF31" box="[1073,1221,801,827]" pageId="29" pageNumber="30" value="2016-11-09">9 Nov. 2016</collectingDate>
</date>
;
<collectorName id="26D35366611ADA45FB005413235EFF31" box="[1234,1399,801,827]" pageId="29" pageNumber="30">HIPE1 exped.</collectorName>
leg.; bought at Rwenshama landing site;
<specimenCode id="DB809ECB611ADA45FD49547625E9FF55" box="[667,960,836,863]" pageId="29" pageNumber="30">RMCA 2016.035.P.0225</specimenCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8B9936B0611ADA45FF3854B727D2F803" blockId="29.[189,1399,901,1033]" pageId="29" pageNumber="30">
<materialsCitation id="3B4E3CED611ADA45FF3854B727DEF803" ID-GBIF-Occurrence="3861203340" collectingDate="1953-07-02" collectionCode="IRSNB" collectorName="KEA exped." country="Democratic Republic of the Congo" latitude="-0.68072224" location="Vitshumbi" longLatPrecision="1" longitude="29.38961" pageId="29" pageNumber="30" specimenCode="IRSNB 13488" specimenCount="1" specimenCount-male="1" stateProvince="Lake Edward" typeStatus="paratype">
<emphasis id="B952EAA2611ADA45FF3854B7274EFF95" bold="true" box="[234,359,901,927]" pageId="29" pageNumber="30">
<typeStatus id="549D8812611ADA45FF3854B7274EFF95" box="[234,359,901,927]" pageId="29" pageNumber="30" type="paratype">Paratypes</typeStatus>
</emphasis>
<collectingCountry id="F3317620611ADA45FF6F549B24D7FFC9" box="[189,766,937,963]" name="Democratic Republic of the Congo" pageId="29" pageNumber="30">DEMOCRATIC REPUBLIC OF THE CONGO</collectingCountry>
<specimenCount id="9D20FD39611ADA45FCCD549B2578FFC9" box="[799,849,937,963]" pageId="29" pageNumber="30" type="male">1 ♂</specimenCount>
,
<quantity id="4CDE9B55611ADA45FCB0549B25E6FFC9" box="[866,975,937,963]" metricMagnitude="-2" metricUnit="m" metricValue="9.84" pageId="29" pageNumber="30" unit="mm" value="98.4">98.4 mm</quantity>
SL; “Lac Edouard: Vitshumbi (au Nord)” [
<collectingRegion id="49E2F852611ADA45FECD54FE27EFFFEC" box="[287,454,972,998]" pageId="29" pageNumber="30">Lake Edward</collectingRegion>
: north of
<location id="8EF9606B611ADA45FDE854F92492FFEC" LSID="urn:lsid:plazi:treatment:038F87A6611BDA79FDB954EB24ADFD74:8EF9606B611ADA45FDE854F92492FFEC" box="[570,699,971,998]" country="Democratic Republic of the Congo" latitude="-0.68072224" longLatPrecision="1" longitude="29.38961" name="Vitshumbi" pageId="29" pageNumber="30" stateProvince="Lake Edward">Vitshumbi</location>
];
<geoCoordinate id="EE125077611ADA45FD1C54F8254EFFEC" box="[718,871,970,998]" degrees="0" direction="south" minutes="40" orientation="latitude" pageId="29" pageNumber="30" precision="1" seconds="50.6" value="-0.68072224">0°4050.6″ S</geoCoordinate>
,
<geoCoordinate id="EE125077611ADA45FCA754F82235FFEC" box="[885,1052,970,998]" degrees="29" direction="east" minutes="23" orientation="longitude" pageId="29" pageNumber="30" precision="1" seconds="22.6" value="29.38961">29°2322.6″ E</geoCoordinate>
(inferred);
<date id="FF981070611ADA45FB7454FE2307FFED" box="[1190,1326,972,999]" pageId="29" pageNumber="30" value="1953-07-02">
<collectingDate id="EFDCE998611ADA45FB7454FE2307FFED" box="[1190,1326,972,999]" pageId="29" pageNumber="30" value="1953-07-02">2 Jul. 1953</collectingDate>
</date>
;
<collectorName id="26D35366611ADA45FAEE54FE2722F803" pageId="29" pageNumber="30">KEA exped.</collectorName>
leg.;
<specimenCode id="DB809ECB611ADA45FE9E54DD27DEF803" box="[332,503,1007,1033]" collectionCode="IRSNB" country="Belgium" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="29" pageNumber="30">IRSNB 13488</specimenCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8B9936B0611ADA45FF6F530325EDF8F1" blockId="29.[189,1399,1071,1275]" pageId="29" pageNumber="30">
<materialsCitation id="3B4E3CED611ADA45FF6F5303230FF864" ID-GBIF-Occurrence="3861203324" collectingDate="2016-11-09" collectionCode="RMCA" collectorName="HIPE 1 exped." country="Uganda" latitude="-0.40444446" location="Rwenshama" longLatPrecision="1" longitude="29.773556" pageId="29" pageNumber="30" specimenCode="RMCA 2016.035.P.0222, RMCA 2016.035.P.0223, RMCA 2016.035.P.0224" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Lake Edward">
<collectingCountry id="F3317620611ADA45FF6F53032713F841" box="[189,314,1073,1099]" name="Uganda" pageId="29" pageNumber="30">UGANDA</collectingCountry>
<emphasis id="B952EAA2611ADA45FE8C53032426F841" bold="true" box="[350,527,1073,1099]" pageId="29" pageNumber="30">
<collectingRegion id="49E2F852611ADA45FE8C53032426F841" box="[350,527,1073,1099]" pageId="29" pageNumber="30">Lake Edward</collectingRegion>
</emphasis>
<specimenCount id="9D20FD39611ADA45FDFC5303245CF841" box="[558,629,1073,1099]" pageId="29" pageNumber="30" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="9D20FD39611ADA45FD565303249AF841" box="[644,691,1073,1099]" pageId="29" pageNumber="30" type="female">1 ♀</specimenCount>
,
<quantity id="4CDE9B55611ADA45FD17530225A8F841" box="[709,897,1072,1099]" metricMagnitude="-1" metricUnit="m" metricValue="1.017" metricValueMax="1.055" metricValueMin="0.979" pageId="29" pageNumber="30" unit="mm" value="101.7" valueMax="105.5" valueMin="97.9">97.9105.5 mm</quantity>
SL;
<geoCoordinate id="EE125077611ADA45FC12531D2273F841" box="[960,1114,1071,1099]" degrees="0" direction="south" minutes="24" orientation="latitude" pageId="29" pageNumber="30" precision="1" seconds="16.0" value="-0.40444446">0°2416.0″ S</geoCoordinate>
,
<geoCoordinate id="EE125077611ADA45FBB8531D233FF841" box="[1130,1302,1071,1099]" degrees="29" direction="east" minutes="46" orientation="longitude" pageId="29" pageNumber="30" precision="1" seconds="24.8" value="29.773556">29°4624.8″ E</geoCoordinate>
;
<date id="FF981070611ADA45FAF7530326D2F867" pageId="29" pageNumber="30" value="2016-11-09">
<collectingDate id="EFDCE998611ADA45FAF7530326D2F867" pageId="29" pageNumber="30" value="2016-11-09">9 Nov. 2016</collectingDate>
</date>
;
<collectorName id="26D35366611ADA45FEDB53662786F864" box="[265,431,1108,1134]" pageId="29" pageNumber="30">HIPE1 exped.</collectorName>
leg.; bought at
<location id="8EF9606B611ADA45FDBF536624D5F864" LSID="urn:lsid:plazi:treatment:038F87A6611BDA79FDB954EB24ADFD74:8EF9606B611ADA45FDBF536624D5F864" box="[621,764,1108,1134]" country="Uganda" latitude="-0.40444446" longLatPrecision="1" longitude="29.773556" name="Rwenshama" pageId="29" pageNumber="30" stateProvince="Lake Edward">Rwenshama</location>
landing site;
<specimenCode id="DB809ECB611ADA45FC725366230FF864" box="[928,1318,1107,1134]" pageId="29" pageNumber="30">RMCA 2016.035.P.0222 to 0224</specimenCode>
</materialsCitation>
<materialsCitation id="3B4E3CED611ADA45FA93536627C9F8BF" ID-GBIF-Occurrence="3861203373" collectingDate="2017-03-25" collectionCode="RMCA" collectorName="HIPE 2 exped." country="Uganda" latitude="-0.40158334" location="Rwenshama" longLatPrecision="1" longitude="29.776417" pageId="29" pageNumber="30" specimenCode="RMCA 2017.006.P.0356" specimenCount="1" specimenCount-female="1" stateProvince="Lake Edward">
<specimenCount id="9D20FD39611ADA45FA9353662346F864" box="[1345,1391,1108,1134]" pageId="29" pageNumber="30" type="female">1 ♀</specimenCount>
,
<quantity id="4CDE9B55611ADA45FF6F5345270CF89B" box="[189,293,1143,1169]" metricMagnitude="-2" metricUnit="m" metricValue="8.709999999999999" pageId="29" pageNumber="30" unit="mm" value="87.1">87.1 mm</quantity>
SL;
<location id="8EF9606B611ADA45FE8C534527D9F89B" LSID="urn:lsid:plazi:treatment:038F87A6611BDA79FDB954EB24ADFD74:8EF9606B611ADA45FE8C534527D9F89B" box="[350,496,1143,1169]" country="Uganda" latitude="-0.40158334" longLatPrecision="1" longitude="29.776417" name="Rwenshama" pageId="29" pageNumber="30" stateProvince="Lake Edward">Rwenshama</location>
, rocky shore;
<geoCoordinate id="EE125077611ADA45FD4653472502F89B" box="[660,811,1141,1169]" degrees="0" direction="south" minutes="24" orientation="latitude" pageId="29" pageNumber="30" precision="1" seconds="05.7" value="-0.40158334">0°2405.7″ S</geoCoordinate>
,
<geoCoordinate id="EE125077611ADA45FCE5534725C9F89B" box="[823,992,1141,1169]" degrees="29" direction="east" minutes="46" orientation="longitude" pageId="29" pageNumber="30" precision="1" seconds="35.1" value="29.776417">29°4635.1″ E</geoCoordinate>
;
<date id="FF981070611ADA45FC3E534522A5F89B" box="[1004,1164,1143,1170]" pageId="29" pageNumber="30" value="2017-03-25">
<collectingDate id="EFDCE998611ADA45FC3E534522A5F89B" box="[1004,1164,1143,1170]" pageId="29" pageNumber="30" value="2017-03-25">25 Mar. 2017</collectingDate>
</date>
;
<collectorName id="26D35366611ADA45FB4A53452314F89B" box="[1176,1341,1143,1169]" pageId="29" pageNumber="30">HIPE2 exped.</collectorName>
leg.;
<specimenCode id="DB809ECB611ADA45FF6F53A827C9F8BF" box="[189,480,1178,1205]" pageId="29" pageNumber="30">RMCA 2017.006.P.0356</specimenCode>
</materialsCitation>
<materialsCitation id="3B4E3CED611ADA45FE2C53A827C9F8D2" ID-GBIF-Occurrence="3861203375" collectingDate="2017-03-25" collectionCode="RMCA" collectorName="HIPE 2 exped." country="Uganda" latitude="-0.40158334" location="Rwenshama" longLatPrecision="1" longitude="29.776417" pageId="29" pageNumber="30" specimenCode="RMCA 2017.006.P.0357, RMCA 2017.006.P.0358, RMCA 2017.006.P.0359" specimenCount="3" specimenCount-male="3" stateProvince="Lake Edward">
<specimenCount id="9D20FD39611ADA45FE2C53A8246CF8BF" box="[510,581,1178,1205]" pageId="29" pageNumber="30" type="male">3 ♂♂</specimenCount>
, 90.0
<quantity id="4CDE9B55611ADA45FD4A53A82539F8BF" box="[664,784,1178,1205]" metricMagnitude="-1" metricUnit="m" metricValue="1.003" pageId="29" pageNumber="30" unit="mm" value="100.3">100.3 mm</quantity>
SL; same collection data as for preceding;
<specimenCode id="DB809ECB611ADA45FAF053A827C9F8D2" pageId="29" pageNumber="30">RMCA 2017.006.P.0357 to 0359</specimenCode>
</materialsCitation>
<materialsCitation id="3B4E3CED611ADA45FE24538F25E9F8F1" ID-GBIF-Occurrence="3861203369" collectingDate="2018-01-24" collectionCode="RMCA" collectorName="HIPE 3 exped." country="Uganda" latitude="-0.40444446" location="Rwenshama" longLatPrecision="1" longitude="29.773556" pageId="29" pageNumber="30" specimenCode="RMCA 2018.008.P.0348" specimenCount="1" specimenCount-female="1" stateProvince="Lake Edward">
<specimenCount id="9D20FD39611ADA45FE24538F2408F8D2" box="[502,545,1213,1240]" pageId="29" pageNumber="30" type="female">1 ♀</specimenCount>
, 109.0 mm SL;
<geoCoordinate id="EE125077611ADA45FD0E538E255BF8DD" box="[732,882,1212,1240]" degrees="0" direction="south" minutes="24" orientation="latitude" pageId="29" pageNumber="30" precision="1" seconds="16.0" value="-0.40444446">0°2416.0″ S</geoCoordinate>
,
<geoCoordinate id="EE125077611ADA45FCAC538E220FF8D2" box="[894,1062,1212,1240]" degrees="29" direction="east" minutes="46" orientation="longitude" pageId="29" pageNumber="30" precision="1" seconds="24.8" value="29.773556">29°4624.8″ E</geoCoordinate>
;
<date id="FF981070611ADA45FBE3538F22E1F8DD" box="[1073,1224,1213,1240]" pageId="29" pageNumber="30" value="2018-01-24">
<collectingDate id="EFDCE998611ADA45FBE3538F22E1F8DD" box="[1073,1224,1213,1240]" pageId="29" pageNumber="30" value="2018-01-24">24 Jan. 2018</collectingDate>
</date>
;
<collectorName id="26D35366611ADA45FB01538F235EF8DD" box="[1235,1399,1213,1239]" pageId="29" pageNumber="30">HIPE3 exped.</collectorName>
leg.; bought at
<location id="8EF9606B611ADA45FEBF53D227D5F8F0" LSID="urn:lsid:plazi:treatment:038F87A6611BDA79FDB954EB24ADFD74:8EF9606B611ADA45FEBF53D227D5F8F0" box="[365,508,1248,1274]" country="Uganda" latitude="-0.40444446" longLatPrecision="1" longitude="29.773556" name="Rwenshama" pageId="29" pageNumber="30" stateProvince="Lake Edward">Rwenshama</location>
landing site;
<specimenCode id="DB809ECB611ADA45FD4953D325E9F8F1" box="[667,960,1249,1275]" pageId="29" pageNumber="30">RMCA 2018.008.P.0348</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33C653B611ADA78FF6F52102438FF18" lastPageId="32" lastPageNumber="33" pageId="29" pageNumber="30" type="description">
<paragraph id="8B9936B0611ADA45FF6F52102767F936" blockId="29.[189,334,1314,1340]" box="[189,334,1314,1340]" pageId="29" pageNumber="30">
<heading id="D0D181DC611ADA45FF6F52102767F936" bold="true" box="[189,334,1314,1340]" fontSize="11" level="3" pageId="29" pageNumber="30" reason="3">
<emphasis id="B952EAA2611ADA45FF6F52102767F936" bold="true" box="[189,334,1314,1340]" pageId="29" pageNumber="30">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8B9936B0611ADA45FF6F52622494FAE4" blockId="29.[188,1399,1360,1774]" pageId="29" pageNumber="30">
Based on
<specimenCount id="9D20FD39611ADA45FEE4526327F1F960" box="[310,472,1360,1387]" pageId="29" pageNumber="30" type="generic">10 specimens</specimenCount>
(87.1109.0 mm SL); body shallow (
<tableCitation id="C6A4030B611ADA45FC70526225D5F961" box="[930,1020,1360,1387]" captionStart="Table 1" captionStartId="9.[182,252,548,576]" captionTargetPageId="9" captionText="Table 1. (continued on the next three pages). Measurements and counts of the piscivorous species of Haplochromis from the Lake Edward system." httpUri="http://table.plazi.org/id/DF596638610EDA51FF6455162158FE4A" pageId="29" pageNumber="30" tableUuid="DF596638610EDA51FF6455162158FE4A">Table 1</tableCitation>
) and oval to slightly pyriform; caudal peduncle shallow (
<figureCitation id="131D2A35611ADA45FE2A5241247DF984" box="[504,596,1395,1422]" captionStart="Fig" captionStartId="30.[189,232,1963,1989]" captionTargetBox="[189,1398,1326,1868]" captionTargetId="figure-310@30.[189,1399,1326,1868]" captionTargetPageId="30" captionText="Fig. 14. Haplochromis simba sp. nov., holotype, ♂, 105.8 mm SL (RMCA 2016.035.P.0225). Drawn by N. Vranken." figureDoi="http://doi.org/10.5281/zenodo.6484197" httpUri="https://zenodo.org/record/6484197/files/figure.png" pageId="29" pageNumber="30">Fig. 14</figureCitation>
). Head long, narrow, and with a straight to weakly convex dorsal outline and a weak concavity above eye; eye small and high on head; interorbital area narrow; cheek and lacrimal deep. Snout long, rounded in dorsal outline, acute in lateral view, and slopes gently at 3045°; premaxillary pedicel average in length and strongly prominent. Jaws isognathous to prognathous, long, slim, very narrow, and rounded in dorsal view; gape large and slopes gently at 2025°; maxilla extends to between verticals through anterior point of orbit and just past this point. Lower jaw relatively shallow and with a straight ventral outline in lateral view, mental prominence strongly developed, and lower jaw side steep with an inclination of 3540° to horizontal in anterior view. Upper jaw expanded anteriorly and ventrally. Lips and oral mucosa thin. Neurocranium shallow, ethmo-vomerine block decurved, preorbital region very shallow (1820% NL), orbital region shallow (2628% NL), and supraoccipital crest shallow and wedge-shaped (
<figureCitation id="131D2A35611ADA45FD9B51E12486FAE4" box="[585,687,1747,1774]" captionStart="Fig" captionStartId="31.[189,232,1892,1918]" captionTargetBox="[189,1398,271,1801]" captionTargetId="figure-12@31.[186,1402,265,1803]" captionTargetPageId="31" captionText="Fig. 15. Haplochromis simba sp. nov. a. Photograph of preserved holotype (RMCA 2016.035.P.0225; 105.8 mm SL). b. X-ray image of holotype. cd. Photographs of freshly caught specimens. c. Dominant male (RMCA 2016.035.P.0224; 97.9 mm SL). d. Female (RMCA 2018.008.P.0348; 109.0 mm SL) to illustrate the live colour patterns. The contrast was slightly enhanced." figureDoi="http://doi.org/10.5281/zenodo.6484199" httpUri="https://zenodo.org/record/6484199/files/figure.png" pageId="29" pageNumber="30">Fig. 15b</figureCitation>
).
</paragraph>
<paragraph id="8B9936B0611ADA46FF6F5027276BFD4C" blockId="29.[189,1401,1813,2016]" lastBlockId="30.[189,1398,265,326]" lastPageId="30" lastPageNumber="31" pageId="29" pageNumber="30">Outer oral teeth few, unicuspid, and very large. Necks stout, conical, and straight; crowns straight to weakly recurved, and acutely pointed. Dental arcades rounded. Outer teeth widely and irregularly set with neck-distances of 14 neck-widths. In upper jaw, 12 posteriormost teeth enlarged. Inner teeth small, recurved, unicuspid, and acutely pointed. Tooth bands very slender crescent-shaped with 12 rows of inner teeth, and narrow posteriorly until only outer row remains past ⅔ length of tooth band. Inner teeth widely and irregularly set on 12 outer tooth neck-widths from outer row; implantation erect in first row and recumbent in subsequent rows; in upper jaw, size decreases slightly buccally and posteriorly.</paragraph>
<paragraph id="8B9936B06119DA46FF6F565C2727FE32" blockId="30.[187,1400,365,568]" pageId="30" pageNumber="31">
Lower pharyngeal bone long, very narrow, slim, and shallow over whole length (
<figureCitation id="131D2A356119DA46FB51565C22F5FD82" box="[1155,1244,365,392]" captionStart="Fig" captionStartId="32.[189,232,1963,1989]" captionTargetBox="[189,1397,960,1907]" captionTargetId="figure-181@32.[189,1398,960,1909]" captionTargetPageId="32" captionText="Fig. 16. Haplochromis simba sp. nov. (RMCA 2016.035.P.0223; 105.5 mm SL). a. Dorsal view of the lower pharyngeal jaw. b. Lateral view of the lower pharyngeal jaw." figureDoi="http://doi.org/10.5281/zenodo.6484201" httpUri="https://zenodo.org/record/6484201/files/figure.png" pageId="30" pageNumber="31">Fig. 16</figureCitation>
). Pharyngeal teeth relatively large and slender; major cusps acutely pointed; cusp gaps concave; minor cusps and cusp protuberances very small. Teeth in two median longitudinal rows equal in size and form to lateral teeth,
<quantity id="4CDE9B556119DA46FED456E5276AFDFB" box="[262,323,471,497]" metricMagnitude="-1" metricUnit="m" metricValue="3.048" pageId="30" pageNumber="31" unit="in" value="12.0">12 in</quantity>
each row. Posterior transverse row with 1618 teeth, implanted recumbently with a lateral inclination; major cusps weakly recurved, bluntly pointed, and laterally compressed; minor cusps mostly absent.
</paragraph>
<paragraph id="8B9936B06119DA46FF6F556D235EFE73" blockId="30.[189,1399,607,634]" box="[189,1399,607,634]" pageId="30" pageNumber="31">Chest scales small; transition to larger flank scales gradual. Minute scales on proximal half of caudal fin.</paragraph>
<paragraph id="8B9936B06119DA46FF6F5592271EFF2E" blockId="30.[189,1399,672,804]" pageId="30" pageNumber="31">Caudal fin emarginate; dorsal and anal fins reach to between verticals through two scales anterior to and one scale posterior to caudal-fin base. Pectoral fin reaches to between anal opening and second analfin spine; pelvic fin reaches to between first and second anal-fin spines; first branched pelvic-fin ray elongated.</paragraph>
<paragraph id="8B9936B06119DA46FF6F5479223EFF83" blockId="30.[189,1399,843,905]" pageId="30" pageNumber="31">Ceratobranchial gill rakers in outer row of first gill arch short, stout, and simple; posteriormost rakers anvil-shaped to weakly bifid. Epibranchial gill rakers slender and simple.</paragraph>
<paragraph id="8B9936B06119DA46FF6F5482278AFFC0" blockId="30.[189,419,944,970]" box="[189,419,944,970]" pageId="30" pageNumber="31">
<heading id="D0D181DC6119DA46FF6F5482278AFFC0" bold="true" box="[189,419,944,970]" fontSize="11" level="3" pageId="30" pageNumber="31" reason="3">
<emphasis id="B952EAA26119DA46FF6F5482278AFFC0" bold="true" box="[189,419,944,970]" pageId="30" pageNumber="31">Colouration in life</emphasis>
</heading>
</paragraph>
<paragraph id="8B9936B06119DA78FF6F54ED2429FD4C" blockId="30.[189,1401,991,1193]" lastBlockId="32.[189,1398,265,326]" lastPageId="32" lastPageNumber="33" pageId="30" pageNumber="31">
Dominant males: body yellow; operculum, cheek, and anterior part of flank orange; belly and chest white (
<figureCitation id="131D2A356119DA46FEC253302751F817" box="[272,376,1026,1053]" captionStart="Fig" captionStartId="31.[189,232,1892,1918]" captionTargetBox="[189,1398,271,1801]" captionTargetId="figure-12@31.[186,1402,265,1803]" captionTargetPageId="31" captionText="Fig. 15. Haplochromis simba sp. nov. a. Photograph of preserved holotype (RMCA 2016.035.P.0225; 105.8 mm SL). b. X-ray image of holotype. cd. Photographs of freshly caught specimens. c. Dominant male (RMCA 2016.035.P.0224; 97.9 mm SL). d. Female (RMCA 2018.008.P.0348; 109.0 mm SL) to illustrate the live colour patterns. The contrast was slightly enhanced." figureDoi="http://doi.org/10.5281/zenodo.6484199" httpUri="https://zenodo.org/record/6484199/files/figure.png" pageId="30" pageNumber="31">Fig. 15c</figureCitation>
). Snout dusky; lower jaw and lips pink; eye with (dark) grey outer ring and silver to golden inner ring. Nostril and lacrimal stripes faint; mental blotch present. pectoral fin yellow; pelvic fin black; dorsal fin hyaline to dusky and with black lappets and a dusky posterior part; anal fin faint crimson and with 23 very large orange egg-spots with hyaline rings; caudal fin dusky and with a faint crimson ventral part and a maculated dorsal part. Non-dominant males: anterior part of flank, operculum, and cheek with pink sheen; anal fin with crimson anterior part and hyaline posterior part; caudal fin uniformly dusky.
</paragraph>
<caption id="DF5966386119DA46FF6F50992762FBE2" ID-DOI="http://doi.org/10.5281/zenodo.6484197" ID-Zenodo-Dep="6484197" httpUri="https://zenodo.org/record/6484197/files/figure.png" pageId="30" pageNumber="31" startId="30.[189,232,1963,1989]" targetBox="[189,1398,1326,1868]" targetPageId="30">
<paragraph id="8B9936B06119DA46FF6F50992762FBE2" blockId="30.[189,1398,1963,2025]" pageId="30" pageNumber="31">
<emphasis id="B952EAA26119DA46FF6F50992733FBCF" bold="true" box="[189,282,1963,1989]" pageId="30" pageNumber="31">Fig. 14.</emphasis>
<taxonomicName id="4C264D336119DA46FEF25099243FFBCF" authority="Vranken &amp; Steenberge &amp; Heylen &amp; Decru &amp; Snoeks, 2022" authorityName="Vranken &amp; Steenberge &amp; Heylen &amp; Decru &amp; Snoeks" authorityYear="2022" box="[288,534,1963,1989]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="30" pageNumber="31" phylum="Chordata" rank="species" species="simba" status="sp. nov.">
<emphasis id="B952EAA26119DA46FEF25099243FFBCF" box="[288,534,1963,1989]" italics="true" pageId="30" pageNumber="31">Haplochromis simba</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26157D96119DA46FDCE509E2450FBCC" box="[540,633,1964,1990]" pageId="30" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, ♂, 105.8 mm SL (RMCA 2016.035.P.0225). Drawn by N. Vranken.
</paragraph>
</caption>
<caption id="DF5966386118DA47FF6F505625C4FBE2" ID-DOI="http://doi.org/10.5281/zenodo.6484199" ID-Zenodo-Dep="6484199" httpUri="https://zenodo.org/record/6484199/files/figure.png" pageId="31" pageNumber="32" startId="31.[189,232,1892,1918]" targetBox="[189,1398,271,1801]" targetPageId="31">
<paragraph id="8B9936B06118DA47FF6F505625C4FBE2" blockId="31.[188,1399,1892,2025]" pageId="31" pageNumber="32">
<emphasis id="B952EAA26118DA47FF6F50562735FB75" bold="true" box="[189,284,1892,1919]" pageId="31" pageNumber="32">Fig. 15.</emphasis>
<taxonomicName id="4C264D336118DA47FEF650572435FB75" authority="Vranken &amp; Steenberge &amp; Heylen &amp; Decru &amp; Snoeks, 2022" authorityName="Vranken &amp; Steenberge &amp; Heylen &amp; Decru &amp; Snoeks" authorityYear="2022" box="[292,540,1893,1919]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="31" pageNumber="32" phylum="Chordata" rank="species" species="simba" status="sp. nov.">
<emphasis id="B952EAA26118DA47FEF650572435FB75" box="[292,540,1893,1919]" italics="true" pageId="31" pageNumber="32">Haplochromis simba</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26157D96118DA47FDF4505724ABFB75" box="[550,642,1893,1919]" pageId="31" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B952EAA26118DA47FD5B505724B1FB75" bold="true" box="[649,664,1893,1919]" pageId="31" pageNumber="32">a</emphasis>
. Photograph of preserved holotype (RMCA 2016.035.P.0225; 105.8 mm SL).
<emphasis id="B952EAA26118DA47FEA150BA27AAFBA8" bold="true" box="[371,387,1928,1954]" pageId="31" pageNumber="32">b</emphasis>
. X-ray image of holotype.
<emphasis id="B952EAA26118DA47FD6C50BA24C2FBA8" bold="true" box="[702,747,1928,1954]" pageId="31" pageNumber="32">cd</emphasis>
. Photographs of freshly caught specimens.
<emphasis id="B952EAA26118DA47FB3750BA22DBFBA8" bold="true" box="[1253,1266,1928,1954]" pageId="31" pageNumber="32">c</emphasis>
. Dominant male (RMCA 2016.035.P.0224; 97.9 mm SL).
<emphasis id="B952EAA26118DA47FD3F509924D5FBCF" bold="true" box="[749,764,1963,1989]" pageId="31" pageNumber="32">d</emphasis>
. Female (RMCA 2018.008.P.0348; 109.0 mm SL) to illustrate the live colour patterns. The contrast was slightly enhanced.
</paragraph>
</caption>
<paragraph id="8B9936B06127DA78FF6F565C26DBFE1F" blockId="32.[189,1399,366,533]" pageId="32" pageNumber="33">
Females and juveniles: body, operculum, and cheek yellow; belly and chest white; lacrimal and lower jaw pink; snout dusky; eye with (dark) grey outer ring and silver to golden inner ring (
<figureCitation id="131D2A356127DA78FB7E56A3233BFDA6" box="[1196,1298,401,428]" captionStart="Fig" captionStartId="31.[189,232,1892,1918]" captionTargetBox="[189,1398,271,1801]" captionTargetId="figure-12@31.[186,1402,265,1803]" captionTargetPageId="31" captionText="Fig. 15. Haplochromis simba sp. nov. a. Photograph of preserved holotype (RMCA 2016.035.P.0225; 105.8 mm SL). b. X-ray image of holotype. cd. Photographs of freshly caught specimens. c. Dominant male (RMCA 2016.035.P.0224; 97.9 mm SL). d. Female (RMCA 2018.008.P.0348; 109.0 mm SL) to illustrate the live colour patterns. The contrast was slightly enhanced." figureDoi="http://doi.org/10.5281/zenodo.6484199" httpUri="https://zenodo.org/record/6484199/files/figure.png" pageId="32" pageNumber="33">Fig. 15d</figureCitation>
). Nostril and interorbital stripes faint; mental blotch present. Pectoral, anal, and caudal fins yellow; anal fin with a white base; caudal fin with a dusky base; dorsal fin hyaline and with black lappets and a dusky distal part.
</paragraph>
<paragraph id="8B9936B06127DA78FF6F550E27F9FE5C" blockId="32.[189,464,572,598]" box="[189,464,572,598]" pageId="32" pageNumber="33">
<heading id="D0D181DC6127DA78FF6F550E27F9FE5C" bold="true" box="[189,464,572,598]" fontSize="11" level="3" pageId="32" pageNumber="33" reason="3">
<emphasis id="B952EAA26127DA78FF6F550E27F9FE5C" bold="true" box="[189,464,572,598]" pageId="32" pageNumber="33">Preserved colouration</emphasis>
</heading>
</paragraph>
<paragraph id="8B9936B06127DA78FF6F55582438FF18" blockId="32.[189,1399,618,786]" pageId="32" pageNumber="33">
Body whitish; dorsum and cheek brown; belly, chest, and operculum white; snout dusky (
<figureCitation id="131D2A356127DA78FAD355582340FE8F" box="[1281,1385,618,645]" captionStart="Fig" captionStartId="31.[189,232,1892,1918]" captionTargetBox="[189,1398,271,1801]" captionTargetId="figure-12@31.[186,1402,265,1803]" captionTargetPageId="31" captionText="Fig. 15. Haplochromis simba sp. nov. a. Photograph of preserved holotype (RMCA 2016.035.P.0225; 105.8 mm SL). b. X-ray image of holotype. cd. Photographs of freshly caught specimens. c. Dominant male (RMCA 2016.035.P.0224; 97.9 mm SL). d. Female (RMCA 2018.008.P.0348; 109.0 mm SL) to illustrate the live colour patterns. The contrast was slightly enhanced." figureDoi="http://doi.org/10.5281/zenodo.6484199" httpUri="https://zenodo.org/record/6484199/files/figure.png" pageId="32" pageNumber="33">Fig. 15a</figureCitation>
). Nostril stripe well-defined; lacrimal stripe and mental blotch present; interorbital stripe and nape band faint. Pectoral fin dusky; pelvic fin yellowish in females, black in males; dorsal fin dusky and with black lappets and sooty posterior part; anal fin yellowish and with 12 large egg-spots; caudal fin dusky and with a maculated dorsal part.
</paragraph>
</subSubSection>
<caption id="DF5966386127DA78FF6F509925F8FBE2" ID-DOI="http://doi.org/10.5281/zenodo.6484201" ID-Zenodo-Dep="6484201" httpUri="https://zenodo.org/record/6484201/files/figure.png" pageId="32" pageNumber="33" startId="32.[189,232,1963,1989]" targetBox="[189,1397,960,1907]" targetPageId="32">
<paragraph id="8B9936B06127DA78FF6F509925F8FBE2" blockId="32.[189,1399,1963,2025]" pageId="32" pageNumber="33">
<emphasis id="B952EAA26127DA78FF6F50992732FBCF" bold="true" box="[189,283,1963,1989]" pageId="32" pageNumber="33">Fig. 16.</emphasis>
<taxonomicName id="4C264D336127DA78FEF150992432FBCF" authority="Vranken &amp; Steenberge &amp; Heylen &amp; Decru &amp; Snoeks, 2022" authorityName="Vranken &amp; Steenberge &amp; Heylen &amp; Decru &amp; Snoeks" authorityYear="2022" box="[291,539,1963,1989]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="simba" status="sp. nov.">
<emphasis id="B952EAA26127DA78FEF150992432FBCF" box="[291,539,1963,1989]" italics="true" pageId="32" pageNumber="33">Haplochromis simba</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26157D96127DA78FDF6509E2456FBCC" box="[548,639,1964,1990]" pageId="32" pageNumber="33" rank="species">sp. nov.</taxonomicNameLabel>
(RMCA 2016.035.P.0223; 105.5 mm SL).
<emphasis id="B952EAA26127DA78FBAF509922A2FBCF" bold="true" box="[1149,1163,1963,1989]" pageId="32" pageNumber="33">a</emphasis>
. Dorsal view of the lower pharyngeal jaw.
<emphasis id="B952EAA26127DA78FE1A50FC27F1FBE2" bold="true" box="[456,472,1998,2024]" pageId="32" pageNumber="33">b</emphasis>
. Lateral view of the lower pharyngeal jaw.
</paragraph>
</caption>
<subSubSection id="C33C653B6126DA79FF6F563A24ADFD74" pageId="33" pageNumber="34" type="distribution">
<paragraph id="8B9936B06126DA79FF6F563A27D8FD29" blockId="33.[189,497,264,291]" box="[189,497,264,291]" pageId="33" pageNumber="34">
<heading id="D0D181DC6126DA79FF6F563A27D8FD29" bold="true" box="[189,497,264,291]" fontSize="11" level="3" pageId="33" pageNumber="34" reason="3">
<emphasis id="B952EAA26126DA79FF6F563A27D8FD29" bold="true" box="[189,497,264,291]" pageId="33" pageNumber="34">Distribution and ecology</emphasis>
</heading>
</paragraph>
<paragraph id="8B9936B06126DA79FF6F560824ADFD74" blockId="33.[189,1398,314,382]" pageId="33" pageNumber="34">Only known from Lake Edward, found in inshore areas over hard substrates. Based on its morphology, most probably a piscivorous species.</paragraph>
</subSubSection>
</treatment>
</document>