741 lines
93 KiB
XML
741 lines
93 KiB
XML
<document id="9C4F00C52DD6E44E5E64740FC704FD9E" ID-CLB-Dataset="92619" ID-DOI="10.5852/ejt.2022.815.1749" ID-GBIF-Dataset="9a798cae-f10e-44c2-a65c-cdadbff28cb4" ID-ISSN="2118-9773" ID-Zenodo-Dep="6484153" ID-ZooBank="6AD0082E-7349-48DE-AFCA-1EE0BFBB3887" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1650913068508" checkinUser="felipe" docAuthor="Vranken, Nathan, Steenberge, Maarten Van, Heylen, Annelies, Decru, Eva & Snoeks, Jos" docDate="2022" docId="038F87A66110DA44FDA855B72794FF90" docLanguage="en" docName="ejt-815_1-94.pdf" docOrigin="European Journal of Taxonomy 815" docSource="http://dx.doi.org/10.5852/ejt.2022.815.1749" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Haplochromis rex Vranken & Steenberge & Heylen & Decru & Snoeks 2022, sp. nov." docType="treatment" docUuid="EF8B7189-BCA4-4A72-8387-498689D76CEB" docUuidSource="ZooBank" docVersion="10" lastPageNumber="29" masterDocId="FFB6FFDE6107DA58FFD257322629FC0A" masterDocTitle="From a pair to a dozen: the piscivorous species of Haplochromis (Cichlidae) from the Lake Edward system" masterLastPageNumber="94" masterPageNumber="1" pageNumber="24" updateTime="1699329164302" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="B57BF2B4139E0208642FCFBD706A247C">From a pair to a dozen: the piscivorous species of Haplochromis (Cichlidae) from the Lake Edward system</mods:title>
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<mods:namePart id="BB5EE1F1F1DDE7EAD70F12533795AAEE">Vranken, Nathan</mods:namePart>
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<mods:affiliation id="C01F7198D3FBADDE69FF821E6F6A7B1A">Leuven, Laboratory of Biodiversity and Evolutionary Genomics, Department of Biology, Charles Deberiotstraat 32, 3000 Leuven, Belgium. & Royal Museum for Central Africa, Biology department, Section Vertebrates, Leuvensesteenweg 13, 3080 Tervuren, Belgium.</mods:affiliation>
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<mods:nameIdentifier id="7423704C84239C40A565F0E66FB02A64" type="email">nathan.vranken@kuleuven.be</mods:nameIdentifier>
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<mods:namePart id="E2F77D550C09DC7CDF41F8B4BE790243">Steenberge, Maarten Van</mods:namePart>
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<mods:affiliation id="7977B6CD9C4713F893B19A2F4E7AB2B4">Royal Belgian Institute of Natural Sciences, Operational Directorate Taxonomy and Phylogeny, Vautierstraat 29, 1000 Brussels, Belgium. & Masaryk University, Department of Botany and Zoology, Kotlářská 2, 611 37 Brno, Czech Republic.</mods:affiliation>
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<mods:nameIdentifier id="8D9FA9B4AA87CCA6615319CA1DE0DF5E" type="email">mvansteenberge@naturalsciences.be</mods:nameIdentifier>
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<mods:namePart id="CAEEE9582E0A5A6E9F4722E242783040">Heylen, Annelies</mods:namePart>
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<mods:affiliation id="C41E29C3C1466A416BC0081DDEF5BAE7">KU Leuven, Laboratory of Biodiversity and Evolutionary Genomics, Department of Biology, Charles Deberiotstraat 32, 3000 Leuven, Belgium.</mods:affiliation>
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<mods:nameIdentifier id="7B0507FB9170FB3C0B454323B2468B9E" type="email">annelies.heylen@hotmail.com</mods:nameIdentifier>
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<mods:namePart id="7D6BD726DC0BBEA17D56510CC52B430C">Decru, Eva</mods:namePart>
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<mods:affiliation id="E5CB6C27CE36556B530855035D3964C9">KU Leuven, Laboratory of Biodiversity and Evolutionary Genomics, Department of Biology, Charles Deberiotstraat 32, 3000 Leuven, Belgium.</mods:affiliation>
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<mods:nameIdentifier id="9F9F8D0EA80E2CDF6D6C1B704BFCE3BA" type="email">eva.decru.icht@gmail.com</mods:nameIdentifier>
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<mods:namePart id="0608BDB5E9EA87E60F8027AC8273CDB4">Snoeks, Jos</mods:namePart>
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<mods:affiliation id="440B815B678078458659E61BFCD0B78A">Royal Museum for Central Africa, Biology department, Section Vertebrates, Leuvensesteenweg 13, 3080 Tervuren, Belgium.</mods:affiliation>
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<mods:nameIdentifier id="D282AC448C9B3FA765DB04EC7C3BE987" type="email">jos.snoeks@africamuseum.be</mods:nameIdentifier>
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<mods:title id="D266369DF535ACB7931DF592501EA90F">European Journal of Taxonomy</mods:title>
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<mods:date id="7B2441AF95640E0417257B5246F4F37B">2022</mods:date>
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<treatment id="038F87A66110DA44FDA855B72794FF90" ID-DOI="http://doi.org/10.5281/zenodo.6484175" ID-GBIF-Taxon="197197509" ID-Zenodo-Dep="6484175" LSID="urn:lsid:zoobank.org:act:EF8B7189-BCA4-4A72-8387-498689D76CEB" httpUri="http://treatment.plazi.org/id/038F87A66110DA44FDA855B72794FF90" lastPageId="28" lastPageNumber="29" pageId="23" pageNumber="24">
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<paragraph id="8B9936B06110DA4FFDA855B72593FEAA" blockId="23.[367,1220,645,742]" box="[634,954,645,672]" pageId="23" pageNumber="24">
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<heading id="D0D181DC6110DA4FFDA855B72593FEAA" box="[634,954,645,672]" centered="true" fontSize="11" level="2" pageId="23" pageNumber="24" reason="2">
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<taxonomicName id="4C264D336110DA4FFDA855B7257EFEAA" authority="Vranken & Steenberge & Heylen & Decru & Snoeks, 2022" authorityName="Vranken & Steenberge & Heylen & Decru & Snoeks" authorityYear="2022" box="[634,855,645,672]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="23" pageNumber="24" phylum="Chordata" rank="species" species="rex" status="sp. nov.">
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<emphasis id="B952EAA26110DA4FFDA855B7257EFEAA" bold="true" box="[634,855,645,672]" italics="true" pageId="23" pageNumber="24">Haplochromis rex</emphasis>
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<taxonomicNameLabel id="A26157D96110DA4FFC8D55B42593FEAA" box="[863,954,646,672]" pageId="23" pageNumber="24" rank="species">sp. nov.</taxonomicNameLabel>
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<subSubSection id="C33C653B6110DA4FFEBD559B22EDFEC9" box="[367,1220,681,707]" pageId="23" pageNumber="24" type="multiple">
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<uuid id="FF800C656110DA4FFD5E559B22EDFEC9" box="[652,1220,681,707]" pageId="23" pageNumber="24">EF8B7189-BCA4-4A72-8387-498689D76CEB</uuid>
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</paragraph>
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<subSubSection id="C33C653B6110DA4FFD5A55FE2582FEEC" box="[648,939,716,742]" pageId="23" pageNumber="24" type="description">
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<paragraph id="8B9936B06110DA4FFD5A55FE2582FEEC" blockId="23.[367,1220,645,742]" box="[648,939,716,742]" pageId="23" pageNumber="24">
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<figureCitation id="131D2A356110DA4FFD5A55FE24D8FEEC" box="[648,753,716,742]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="5.[190,234,1822,1848]" captionStartId-1="6.[189,231,1892,1918]" captionTargetBox-0="[193,1398,734,1726]" captionTargetBox-1="[208,1335,272,1830]" captionTargetId-0="figure-217@5.[189,1399,730,1726]" captionTargetId-1="figure-18@6.[208,1335,272,1830]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="Fig. 1. The Lake Edward system, consisting of Lakes Edward and George that are connected through the Kazinga Channel. The catch localities of all examined specimens are indicated: Haplochromis sp. ‘latifrons’ (), H. mentatus Regan, 1925 (), H. sp. ‘rex’ (), H. sp. ‘simba’ (▼), H. sp. ‘glaucus’ (), H. sp. ‘aquila’ (), H. sp. ‘kimondo’ (●), H. sp. ‘falcatus’ (), H. sp. ‘curvidens’ (), H. sp. ‘pardus’ (), H. sp. ‘quasimodo’ (▲), and H. squamipinnis Regan, 1921 (█). The type localities of H. mentatus and H. squamipinnis are not shown as the exact locations are unknown." captionText-1="Fig. 2. a. Differences in total number of outer teeth and premaxillary dentition between the macro- and microdontic groups of piscivores from the Lake Edward system.b. Schematic presentation of the dominant male colour patterns in life, in the various groups studied. For H. sp. ‘latifrons’ the female and for H. sp. ‘curvidens’ the non-dominant colour patterns are shown, as those of dominant males are unknown." figureDoi-0="http://doi.org/10.5281/zenodo.6484155" figureDoi-1="http://doi.org/10.5281/zenodo.6484157" httpUri-0="https://zenodo.org/record/6484155/files/figure.png" httpUri-1="https://zenodo.org/record/6484157/files/figure.png" pageId="23" pageNumber="24">Figs 1–2</figureCitation>
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,
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<figureCitation id="131D2A356110DA4FFD2F55FE2561FEEC" box="[765,840,716,742]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="24.[189,232,1963,1989]" captionStartId-1="26.[189,232,1892,1918]" captionStartId-2="27.[189,232,1963,1989]" captionTargetBox-0="[189,1398,1294,1855]" captionTargetBox-1="[207,1383,278,1848]" captionTargetBox-2="[190,1395,1029,1890]" captionTargetId-0="figure-316@24.[189,1399,1294,1855]" captionTargetId-1="figure-19@26.[207,1383,262,1848]" captionTargetId-2="figure-244@27.[190,1397,1029,1892]" captionTargetPageId-0="24" captionTargetPageId-1="26" captionTargetPageId-2="27" captionText-0="Fig. 11. Haplochromis rex sp. nov., holotype, ♂, 148.5 mm SL (RMCA 2018.008.P.0345). Drawn by N. Vranken." captionText-1="Fig. 12. Haplochromis rex sp. nov. a. Photograph of preserved holotype (RMCA 2018.008.P.0345; 148.5 mm SL). b. X-ray image of holotype c–d. Photographs of freshly caught specimens. c. Holotype, a dominant male. d. A female (RMCA 2017.006.P.0355; 135.7 mm SL) to illustrate the live colour patterns. The contrast was slightly enhanced." captionText-2="Fig. 13. Haplochromis rexsp. nov. (IRSNB 13480; 154.2 mm SL). a. Dorsal view of the lower pharyngeal jaw. b. Lateral view of the lower pharyngeal jaw." figureDoi-0="http://doi.org/10.5281/zenodo.6484187" figureDoi-1="http://doi.org/10.5281/zenodo.6484191" figureDoi-2="http://doi.org/10.5281/zenodo.6484195" httpUri-0="https://zenodo.org/record/6484187/files/figure.png" httpUri-1="https://zenodo.org/record/6484191/files/figure.png" httpUri-2="https://zenodo.org/record/6484195/files/figure.png" pageId="23" pageNumber="24">11–13</figureCitation>
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;
|
||
<tableCitation id="C6A4030B6110DA4FFC8655FE2582FEEC" box="[852,939,716,742]" captionStart="Table 1" captionStartId="9.[182,252,548,576]" captionTargetPageId="9" captionText="Table 1. (continued on the next three pages). Measurements and counts of the piscivorous species of Haplochromis from the Lake Edward system." httpUri="http://table.plazi.org/id/DF596638610EDA51FF6455162158FE4A" pageId="23" pageNumber="24" tableUuid="DF596638610EDA51FF6455162158FE4A">Table 1</tableCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C33C653B6110DA40FF6F543F251FFD86" lastPageId="24" lastPageNumber="25" pageId="23" pageNumber="24" type="diagnosis">
|
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<paragraph id="8B9936B06110DA4FFF6F543F27E0FF2D" blockId="23.[189,457,781,807]" box="[189,457,781,807]" pageId="23" pageNumber="24">
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<heading id="D0D181DC6110DA4FFF6F543F27E0FF2D" bold="true" box="[189,457,781,807]" fontSize="11" level="3" pageId="23" pageNumber="24" reason="3">
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<emphasis id="B952EAA26110DA4FFF6F543F27E0FF2D" bold="true" box="[189,457,781,807]" pageId="23" pageNumber="24">Differential diagnosis</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B9936B06110DA4FFF6F540E272DFFCA" blockId="23.[189,1400,827,960]" pageId="23" pageNumber="24">Species with a piscivorous morphology; head narrow [HW 36.8–41.6 (mean 39.2) % HL]; cheek deep [ChD 27.6–33.5 (31.1) % HL]; eye small [ED 22.2–28.3 (24.6) % HL]; outer oral teeth few and large [UOT 24–36 (median 29)]; dominant males cream-coloured with an orange operculum and a light blue snout.</paragraph>
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<paragraph id="8B9936B06110DA4FFF6F54D524FCF8BB" blockId="23.[188,1400,999,1201]" pageId="23" pageNumber="24">
|
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Amongst piscivorous species from the Lake Edward system,
|
||
<taxonomicName id="4C264D336110DA4FFC4F54DA25CFF80B" authority="Vranken & Steenberge & Heylen & Decru & Snoeks, 2022" authorityName="Vranken & Steenberge & Heylen & Decru & Snoeks" authorityYear="2022" box="[925,998,999,1025]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="23" pageNumber="24" phylum="Chordata" rank="species" species="rex" status="sp. nov.">
|
||
<emphasis id="B952EAA26110DA4FFC4F54DA2599F80B" box="[925,944,1000,1025]" italics="true" pageId="23" pageNumber="24">H</emphasis>
|
||
.
|
||
<emphasis id="B952EAA26110DA4FFC1354DA25CFF80B" box="[961,998,1000,1025]" italics="true" pageId="23" pageNumber="24">rex</emphasis>
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||
</taxonomicName>
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||
<taxonomicNameLabel id="A26157D96110DA4FFC2054D52267F80B" box="[1010,1102,999,1025]" pageId="23" pageNumber="24" rank="species">sp. nov.</taxonomicNameLabel>
|
||
differs from
|
||
<taxonomicName id="4C264D336110DA4FFB3D54DA235FF80B" authority="Vranken & Steenberge & Heylen & Decru & Snoeks, 2022" authorityName="Vranken & Steenberge & Heylen & Decru & Snoeks" authorityYear="2022" box="[1263,1398,999,1025]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="23" pageNumber="24" phylum="Chordata" rank="species" species="latifrons" status="sp. nov.">
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||
<emphasis id="B952EAA26110DA4FFB3D54DA232BF80B" box="[1263,1282,1000,1025]" italics="true" pageId="23" pageNumber="24">H</emphasis>
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||
.
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||
<emphasis id="B952EAA26110DA4FFAC154D5235FF80B" box="[1299,1398,999,1025]" italics="true" pageId="23" pageNumber="24">latifrons</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A26157D96110DA4FFF6D53392735F82F" box="[191,284,1035,1061]" pageId="23" pageNumber="24" rank="species">sp. nov.</taxonomicNameLabel>
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and
|
||
<taxonomicName id="4C264D336110DA4FFE8A533927C3F82E" authorityName="Regan" authorityYear="1925" box="[344,490,1035,1061]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="23" pageNumber="24" phylum="Chordata" rank="species" species="mentatus">
|
||
<emphasis id="B952EAA26110DA4FFE8A53392744F82E" box="[344,365,1035,1060]" italics="true" pageId="23" pageNumber="24">H</emphasis>
|
||
.
|
||
<emphasis id="B952EAA26110DA4FFEAD533927C3F82E" box="[383,490,1035,1060]" italics="true" pageId="23" pageNumber="24">mentatus</emphasis>
|
||
</taxonomicName>
|
||
by the combination of a shorter caudal peduncle [CPL 13.5–16.2 (14.8) vs 15.7–18.0 (16.6–17.0) % SL] and a narrower interorbital area [IOW 44.9–52.7 (48.9) vs 51.3–63.3 (55.5–60.0) % HW]; further from
|
||
<taxonomicName id="4C264D336110DA4FFD8A536324C9F860" authority="Vranken & Steenberge & Heylen & Decru & Snoeks, 2022" authorityName="Vranken & Steenberge & Heylen & Decru & Snoeks" authorityYear="2022" box="[600,736,1104,1131]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="23" pageNumber="24" phylum="Chordata" rank="species" species="latifrons" status="sp. nov.">
|
||
<emphasis id="B952EAA26110DA4FFD8A53632445F860" box="[600,620,1105,1130]" italics="true" pageId="23" pageNumber="24">H</emphasis>
|
||
.
|
||
<emphasis id="B952EAA26110DA4FFDAE536224C9F860" box="[636,736,1104,1130]" italics="true" pageId="23" pageNumber="24">latifrons</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26157D96110DA4FFD395363256EF861" box="[747,839,1105,1131]" pageId="23" pageNumber="24" rank="species">sp. nov.</taxonomicNameLabel>
|
||
by absence vs presence of a well-defined midlateral band; further from
|
||
<taxonomicName id="4C264D336110DA4FFE3953472453F884" authorityName="Regan" authorityYear="1925" box="[491,634,1140,1166]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="23" pageNumber="24" phylum="Chordata" rank="species" species="mentatus">
|
||
<emphasis id="B952EAA26110DA4FFE3953472429F884" box="[491,512,1141,1166]" italics="true" pageId="23" pageNumber="24">H</emphasis>
|
||
.
|
||
<emphasis id="B952EAA26110DA4FFDDD53472453F884" box="[527,634,1141,1166]" italics="true" pageId="23" pageNumber="24">mentatus</emphasis>
|
||
</taxonomicName>
|
||
by dominant males cream-coloured with an orange operculum vs yellow-green with a red anterior part of flank.
|
||
</paragraph>
|
||
<paragraph id="8B9936B06110DA4FFF6F53EB24DBF98A" blockId="23.[189,1401,1240,1408]" pageId="23" pageNumber="24">
|
||
It differs from
|
||
<taxonomicName id="4C264D336110DA4FFEA753EB27CDF8F8" authority="Vranken & Steenberge & Heylen & Decru & Snoeks, 2022" authorityName="Vranken & Steenberge & Heylen & Decru & Snoeks" authorityYear="2022" box="[373,484,1240,1267]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="23" pageNumber="24" phylum="Chordata" rank="species" species="simba" status="sp. nov.">
|
||
<emphasis id="B952EAA26110DA4FFEA753EB27A3F8F8" box="[373,394,1241,1266]" italics="true" pageId="23" pageNumber="24">H</emphasis>
|
||
.
|
||
<emphasis id="B952EAA26110DA4FFE4F53EA27CDF8F8" box="[413,484,1240,1266]" italics="true" pageId="23" pageNumber="24">simba</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26157D96110DA4FFE3D53EB2467F8F9" box="[495,590,1241,1267]" pageId="23" pageNumber="24" rank="species">sp. nov.</taxonomicNameLabel>
|
||
by the combination of a larger number of longitudinal line scales (LongL 34–38 vs 32–33), a larger number of scales between first anal-fin spine and upper lateral line (ULL-A 12–16 vs 9–11), absent or weakly developed vs strongly developed mental prominence, and dominant males cream-coloured with an orange operculum and a light blue snout vs yellow with an orange anterior part of flank and a yellow snout.
|
||
</paragraph>
|
||
<paragraph id="8B9936B06110DA4FFF6F529524BEFA21" blockId="23.[189,1400,1446,1579]" pageId="23" pageNumber="24">
|
||
It differs from
|
||
<taxonomicName id="4C264D336110DA4FFEBC529527C7F9CB" authority="Vranken & Steenberge & Heylen & Decru & Snoeks, 2022" authorityName="Vranken & Steenberge & Heylen & Decru & Snoeks" authorityYear="2022" box="[366,494,1447,1473]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="23" pageNumber="24" phylum="Chordata" rank="species" species="glaucus" status="sp. nov.">
|
||
<emphasis id="B952EAA26110DA4FFEBC529527ABF9CA" box="[366,386,1447,1472]" italics="true" pageId="23" pageNumber="24">H</emphasis>
|
||
.
|
||
<emphasis id="B952EAA26110DA4FFE40529527C7F9CB" box="[402,494,1447,1473]" italics="true" pageId="23" pageNumber="24">glaucus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26157D96110DA4FFE2B5295247DF9CB" box="[505,596,1447,1473]" pageId="23" pageNumber="24" rank="species">sp. nov.</taxonomicNameLabel>
|
||
by the combination of a narrower interorbital area [IOW 44.9–52.7 (48.9) vs 50.9–57.1 (53.8) % HW], a steeper snout (40–50° vs 30–40°), rounded vs acute oral jaws in dorsal view, and dominant males cream-coloured with an orange operculum and a light blue snout vs uniformly light blue with a dusky snout.
|
||
</paragraph>
|
||
<paragraph id="8B9936B06110DA4FFF6F5160225DFAB8" blockId="23.[189,1399,1617,1715]" pageId="23" pageNumber="24">
|
||
It differs from
|
||
<taxonomicName id="4C264D336110DA4FFEA0516027CCFA66" authority="Vranken & Steenberge & Heylen & Decru & Snoeks, 2022" authorityName="Vranken & Steenberge & Heylen & Decru & Snoeks" authorityYear="2022" box="[370,485,1618,1644]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="23" pageNumber="24" phylum="Chordata" rank="species" species="aquila" status="sp. nov.">
|
||
<emphasis id="B952EAA26110DA4FFEA0516027AEFA61" box="[370,391,1618,1643]" italics="true" pageId="23" pageNumber="24">H</emphasis>
|
||
.
|
||
<emphasis id="B952EAA26110DA4FFE4B516027CCFA66" box="[409,485,1618,1644]" italics="true" pageId="23" pageNumber="24">aquila</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26157D96110DA4FFE3D51602465FA66" box="[495,588,1618,1644]" pageId="23" pageNumber="24" rank="species">sp. nov.</taxonomicNameLabel>
|
||
by the combination of a deeper lacrimal [LaD 18.9–22.5 (20.8) vs 17.0–19.1 (18.3) % HL], smaller eye [ED 22.2–28.3 (24.6) vs 30.0–31.5 (30.6) % HL], and dominant males cream-coloured with an orange operculum vs light grey with a black head.
|
||
</paragraph>
|
||
<paragraph id="8B9936B06110DA4FFF6F51E8237FFB30" blockId="23.[188,1399,1753,1850]" pageId="23" pageNumber="24">
|
||
It differs from
|
||
<taxonomicName id="4C264D336110DA4FFEA151E82428FAF9" authority="Vranken & Steenberge & Heylen & Decru & Snoeks, 2022" authorityName="Vranken & Steenberge & Heylen & Decru & Snoeks" authorityYear="2022" box="[371,513,1753,1780]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="23" pageNumber="24" phylum="Chordata" rank="species" species="kimondo" status="sp. nov.">
|
||
<emphasis id="B952EAA26110DA4FFEA151E827A1FAF9" box="[371,392,1754,1779]" italics="true" pageId="23" pageNumber="24">H</emphasis>
|
||
.
|
||
<emphasis id="B952EAA26110DA4FFE4951EB2428FAF9" box="[411,513,1753,1779]" italics="true" pageId="23" pageNumber="24">kimondo</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26157D96110DA4FFDD951E82445FAFE" box="[523,620,1754,1780]" pageId="23" pageNumber="24" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="4C264D336110DA4FFDA951E82529FAF9" authority="Vranken & Steenberge & Heylen & Decru & Snoeks, 2022" authorityName="Vranken & Steenberge & Heylen & Decru & Snoeks" authorityYear="2022" box="[635,768,1753,1780]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="23" pageNumber="24" phylum="Chordata" rank="species" species="falcatus" status="sp. nov.">
|
||
<emphasis id="B952EAA26110DA4FFDA951E824B9FAF9" box="[635,656,1754,1779]" italics="true" pageId="23" pageNumber="24">H</emphasis>
|
||
.
|
||
<emphasis id="B952EAA26110DA4FFD7051EB2529FAF9" box="[674,768,1753,1779]" italics="true" pageId="23" pageNumber="24">falcatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26157D96110DA4FFCD951E82542FAFE" box="[779,875,1754,1780]" pageId="23" pageNumber="24" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="4C264D336110DA4FFCA851E8223DFAF9" authority="Vranken & Steenberge & Heylen & Decru & Snoeks, 2022" authorityName="Vranken & Steenberge & Heylen & Decru & Snoeks" authorityYear="2022" box="[890,1044,1753,1780]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="23" pageNumber="24" phylum="Chordata" rank="species" species="curvidens" status="sp. nov.">
|
||
<emphasis id="B952EAA26110DA4FFCA851E825A6FAF9" box="[890,911,1754,1779]" italics="true" pageId="23" pageNumber="24">H</emphasis>
|
||
.
|
||
<emphasis id="B952EAA26110DA4FFC7351EB223DFAF9" box="[929,1044,1753,1779]" italics="true" pageId="23" pageNumber="24">curvidens</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26157D96110DA4FFBCD51E822A9FAFE" box="[1055,1152,1754,1780]" pageId="23" pageNumber="24" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="4C264D336110DA4FFB5C51E8232EFAF9" authority="Vranken & Steenberge & Heylen & Decru & Snoeks, 2022" authorityName="Vranken & Steenberge & Heylen & Decru & Snoeks" authorityYear="2022" box="[1166,1287,1753,1780]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="23" pageNumber="24" phylum="Chordata" rank="species" species="pardus" status="sp. nov.">
|
||
<emphasis id="B952EAA26110DA4FFB5C51E8228AFAF9" box="[1166,1187,1754,1779]" italics="true" pageId="23" pageNumber="24">H</emphasis>
|
||
.
|
||
<emphasis id="B952EAA26110DA4FFB6451EB232EFAF9" box="[1206,1287,1753,1779]" italics="true" pageId="23" pageNumber="24">pardus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26157D96110DA4FFAC051E8235BFAFE" box="[1298,1394,1754,1780]" pageId="23" pageNumber="24" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="4C264D336110DA4FFF6F51CF274BFB1D" authority="Vranken & Steenberge & Heylen & Decru & Snoeks, 2022" authorityName="Vranken & Steenberge & Heylen & Decru & Snoeks" authorityYear="2022" box="[189,354,1789,1815]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="23" pageNumber="24" phylum="Chordata" rank="species" species="quasimodo" status="sp. nov.">
|
||
<emphasis id="B952EAA26110DA4FFF6F51CF26FBFB1C" box="[189,210,1789,1814]" italics="true" pageId="23" pageNumber="24">H</emphasis>
|
||
.
|
||
<emphasis id="B952EAA26110DA4FFF3251CF274BFB1D" box="[224,354,1789,1815]" italics="true" pageId="23" pageNumber="24">quasimodo</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26157D96110DA4FFEB551CF27EBFB1D" box="[359,450,1789,1815]" pageId="23" pageNumber="24" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, and
|
||
<taxonomicName id="4C264D336110DA4FFE2F51CF2496FB1C" authorityName="Regan" authorityYear="1921" box="[509,703,1789,1815]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="23" pageNumber="24" phylum="Chordata" rank="species" species="squamipinnis">
|
||
<emphasis id="B952EAA26110DA4FFE2F51CF243BFB1C" box="[509,530,1789,1814]" italics="true" pageId="23" pageNumber="24">H</emphasis>
|
||
.
|
||
<emphasis id="B952EAA26110DA4FFDF351CF2496FB1C" box="[545,703,1789,1814]" italics="true" pageId="23" pageNumber="24">squamipinnis</emphasis>
|
||
</taxonomicName>
|
||
by the combination of stout vs slim oral jaws, large vs small outer oral teeth, and a smaller number of outer upper jaw teeth [UOT 24–36 (29) vs 39–79 (45–58)].
|
||
</paragraph>
|
||
<paragraph id="8B9936B06110DA4FFF6F50502494FBEF" blockId="23.[188,1398,1889,2021]" pageId="23" pageNumber="24">
|
||
It further differs from
|
||
<taxonomicName id="4C264D336110DA4FFE1D50502474FB71" authority="Vranken & Steenberge & Heylen & Decru & Snoeks, 2022" authorityName="Vranken & Steenberge & Heylen & Decru & Snoeks" authorityYear="2022" box="[463,605,1889,1916]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="23" pageNumber="24" phylum="Chordata" rank="species" species="kimondo" status="sp. nov.">
|
||
<emphasis id="B952EAA26110DA4FFE1D505027CDFB71" box="[463,484,1890,1915]" italics="true" pageId="23" pageNumber="24">H</emphasis>
|
||
.
|
||
<emphasis id="B952EAA26110DA4FFE2550532474FB71" box="[503,605,1889,1915]" italics="true" pageId="23" pageNumber="24">kimondo</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26157D96110DA4FFDBA505024E0FB76" box="[616,713,1890,1916]" pageId="23" pageNumber="24" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="4C264D336110DA4FFD0A5050255AFB71" authority="Vranken & Steenberge & Heylen & Decru & Snoeks, 2022" authorityName="Vranken & Steenberge & Heylen & Decru & Snoeks" authorityYear="2022" box="[728,883,1889,1916]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="23" pageNumber="24" phylum="Chordata" rank="species" species="curvidens" status="sp. nov.">
|
||
<emphasis id="B952EAA26110DA4FFD0A505024C4FB71" box="[728,749,1890,1915]" italics="true" pageId="23" pageNumber="24">H</emphasis>
|
||
.
|
||
<emphasis id="B952EAA26110DA4FFCD25053255AFB71" box="[768,883,1889,1915]" italics="true" pageId="23" pageNumber="24">curvidens</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26157D96110DA4FFCAC505025F6FB76" box="[894,991,1890,1916]" pageId="23" pageNumber="24" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, and
|
||
<taxonomicName id="4C264D336110DA4FFBF6505022E4FB71" authority="Vranken & Steenberge & Heylen & Decru & Snoeks, 2022" authorityName="Vranken & Steenberge & Heylen & Decru & Snoeks" authorityYear="2022" box="[1060,1229,1889,1916]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="23" pageNumber="24" phylum="Chordata" rank="species" species="quasimodo" status="sp. nov.">
|
||
<emphasis id="B952EAA26110DA4FFBF650502210FB71" box="[1060,1081,1890,1915]" italics="true" pageId="23" pageNumber="24">H</emphasis>
|
||
.
|
||
<emphasis id="B952EAA26110DA4FFB99505322E4FB71" box="[1099,1229,1889,1915]" italics="true" pageId="23" pageNumber="24">quasimodo</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26157D96110DA4FFB0A5050231EFB76" box="[1240,1335,1890,1916]" pageId="23" pageNumber="24" rank="species">sp. nov.</taxonomicNameLabel>
|
||
by a narrower head [HW 36.8–41.6 (39.2) vs 42.0–48.1 (43.4–45.3) % HL]; further from
|
||
<taxonomicName id="4C264D336110DA4FFB5F50B7233EFB94" authority="Vranken & Steenberge & Heylen & Decru & Snoeks, 2022" authorityName="Vranken & Steenberge & Heylen & Decru & Snoeks" authorityYear="2022" box="[1165,1303,1924,1951]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="23" pageNumber="24" phylum="Chordata" rank="species" species="kimondo" status="sp. nov.">
|
||
<emphasis id="B952EAA26110DA4FFB5F50B7228BFB94" box="[1165,1186,1925,1950]" italics="true" pageId="23" pageNumber="24">H</emphasis>
|
||
.
|
||
<emphasis id="B952EAA26110DA4FFB6350B6233EFB94" box="[1201,1303,1924,1950]" italics="true" pageId="23" pageNumber="24">kimondo</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26157D96110DA4FFACE50B7235FFB95" box="[1308,1398,1925,1951]" pageId="23" pageNumber="24" rank="species">sp. nov.</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="4C264D336110DA4FFF21509B27B7FBC8" authority="Vranken & Steenberge & Heylen & Decru & Snoeks, 2022" authorityName="Vranken & Steenberge & Heylen & Decru & Snoeks" authorityYear="2022" box="[243,414,1960,1986]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="23" pageNumber="24" phylum="Chordata" rank="species" species="quasimodo" status="sp. nov.">
|
||
<emphasis id="B952EAA26110DA4FFF21509B2721FBC8" box="[243,264,1961,1986]" italics="true" pageId="23" pageNumber="24">H</emphasis>
|
||
.
|
||
<emphasis id="B952EAA26110DA4FFECE509A27B7FBC8" box="[284,414,1960,1986]" italics="true" pageId="23" pageNumber="24">quasimodo</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26157D96110DA4FFE79509A2423FBC8" box="[427,522,1960,1986]" pageId="23" pageNumber="24" rank="species">sp. nov.</taxonomicNameLabel>
|
||
by dominant males cream-coloured with an orange operculum vs grey dorsally and yellow or blue-black ventrally.
|
||
</paragraph>
|
||
<paragraph id="8B9936B0611FDA40FF6F563B251FFD86" blockId="24.[188,1399,265,397]" pageId="24" pageNumber="25">
|
||
It further differs from
|
||
<taxonomicName id="4C264D33611FDA40FE105638246DFD29" authority="Vranken & Steenberge & Heylen & Decru & Snoeks, 2022" authorityName="Vranken & Steenberge & Heylen & Decru & Snoeks" authorityYear="2022" box="[450,580,265,291]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="falcatus" status="sp. nov.">
|
||
<emphasis id="B952EAA2611FDA40FE10563827FEFD29" box="[450,471,266,291]" italics="true" pageId="24" pageNumber="25">H</emphasis>
|
||
.
|
||
<emphasis id="B952EAA2611FDA40FE34563B246DFD29" box="[486,580,265,291]" italics="true" pageId="24" pageNumber="25">falcatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26157D9611FDA40FD9E563B2483FD29" box="[588,682,265,291]" pageId="24" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="4C264D33611FDA40FD6756382502FD29" authority="Vranken & Steenberge & Heylen & Decru & Snoeks, 2022" authorityName="Vranken & Steenberge & Heylen & Decru & Snoeks" authorityYear="2022" box="[693,811,265,291]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="pardus" status="sp. nov.">
|
||
<emphasis id="B952EAA2611FDA40FD67563824E3FD29" box="[693,714,266,291]" italics="true" pageId="24" pageNumber="25">H</emphasis>
|
||
.
|
||
<emphasis id="B952EAA2611FDA40FD08563B2502FD29" box="[730,811,265,291]" italics="true" pageId="24" pageNumber="25">pardus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26157D9611FDA40FCE1563B25B9FD29" box="[819,912,265,291]" pageId="24" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, and
|
||
<taxonomicName id="4C264D33611FDA40FC1C563822B9FD29" authorityName="Regan" authorityYear="1921" box="[974,1168,265,291]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="squamipinnis">
|
||
<emphasis id="B952EAA2611FDA40FC1C563825CAFD29" box="[974,995,266,291]" italics="true" pageId="24" pageNumber="25">H</emphasis>
|
||
.
|
||
<emphasis id="B952EAA2611FDA40FC20563822B9FD29" box="[1010,1168,266,291]" italics="true" pageId="24" pageNumber="25">squamipinnis</emphasis>
|
||
</taxonomicName>
|
||
by a larger number of longitudinal line scales (LongL 34–38 vs 29–33, rarely 34) and dominant males cream-coloured with an orange operculum vs uniformly olive-green with an orange-red anterior part of flank, speckled to uniformly black, or uniformly slate blue, respectively.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33C653B611FDA40FF6F56862762FE6D" pageId="24" pageNumber="25" type="etymology">
|
||
<paragraph id="8B9936B0611FDA40FF6F5686276DFDC4" blockId="24.[189,324,436,462]" box="[189,324,436,462]" pageId="24" pageNumber="25">
|
||
<heading id="D0D181DC611FDA40FF6F5686276DFDC4" bold="true" box="[189,324,436,462]" fontSize="11" level="3" pageId="24" pageNumber="25" reason="3">
|
||
<emphasis id="B952EAA2611FDA40FF6F5686276DFDC4" bold="true" box="[189,324,436,462]" pageId="24" pageNumber="25">Etymology</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8B9936B0611FDA40FF6F56D02762FE6D" blockId="24.[188,1399,482,615]" pageId="24" pageNumber="25">
|
||
Specific name from the Latin ‘
|
||
<taxonomicName id="4C264D33611FDA40FDFF56D1247BFDF6" authorityName="Vranken & Steenberge & Heylen & Decru & Snoeks" authorityYear="2022" box="[557,594,483,508]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="rex">
|
||
<emphasis id="B952EAA2611FDA40FDFF56D1247BFDF6" box="[557,594,483,508]" italics="true" pageId="24" pageNumber="25">rex</emphasis>
|
||
</taxonomicName>
|
||
’ for ‘king’ (one that holds a preeminent position); referring to very small eyes, deep cheeks, and strong jaws set with large and acute teeth indicating this piscivore has most specialised morphology among all piscivores from the Lake Edward system to hunt on large prey (
|
||
<bibRefCitation id="EFB74B41611FDA40FAE5551B2712FE6C" author="Barel C. D. N. & van Oijen M. J. P. & Witte F. & Witte-Maas E. L. M." pageId="24" pageNumber="25" pagination="333 - 380" refId="ref48472" refString="Barel C. D. N., van Oijen M. J. P., Witte F. & Witte-Maas E. L. M. 1977. An introduction to the taxonomy and morphology of the haplochromine Cichlidae from Lake Victoria. Netherlands Journal of Zoology 27: 333 - 380." type="journal article" year="1977">
|
||
Barel
|
||
<emphasis id="B952EAA2611FDA40FF6F557F26D8FE6C" box="[189,241,588,614]" italics="true" pageId="24" pageNumber="25">et al</emphasis>
|
||
. 1977
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33C653B611FDA41FF6F55BF236BFE9A" lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25" type="materials_examined">
|
||
<paragraph id="8B9936B0611FDA40FF6F55BF2783FEAD" blockId="24.[189,426,653,679]" box="[189,426,653,679]" pageId="24" pageNumber="25">
|
||
<heading id="D0D181DC611FDA40FF6F55BF2783FEAD" bold="true" box="[189,426,653,679]" fontSize="11" level="3" pageId="24" pageNumber="25" reason="3">
|
||
<emphasis id="B952EAA2611FDA40FF6F55BF2783FEAD" bold="true" box="[189,426,653,679]" pageId="24" pageNumber="25">Material examined</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8B9936B0611FDA40FF38558E25EDFF17" blockId="24.[189,1399,700,797]" pageId="24" pageNumber="25">
|
||
<materialsCitation id="3B4E3CED611FDA40FF38558E25EDFF17" ID-GBIF-Occurrence="3861203312" collectingDate="2018-01-24" collectionCode="RMCA" collectorName="HIPE 3 exped." country="Uganda" latitude="-0.40444446" location="Rwenshama" longLatPrecision="1" longitude="29.773556" pageId="24" pageNumber="25" specimenCode="RMCA 2018.008.P.0345" specimenCount="1" specimenCount-male="1" stateProvince="Lake Edward" typeStatus="holotype">
|
||
<emphasis id="B952EAA2611FDA40FF38558E2775FEDC" bold="true" box="[234,348,700,726]" pageId="24" pageNumber="25">
|
||
<typeStatus id="549D8812611FDA40FF38558E2775FEDC" box="[234,348,700,726]" pageId="24" pageNumber="25" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
<collectingCountry id="F3317620611FDA40FF6F55ED2713FEF3" box="[189,314,735,761]" name="Uganda" pageId="24" pageNumber="25">UGANDA</collectingCountry>
|
||
•
|
||
<specimenCount id="9D20FD39611FDA40FE8155D22742FEF0" box="[339,363,736,762]" pageId="24" pageNumber="25" type="male">♂</specimenCount>
|
||
,
|
||
<quantity id="4CDE9B55611FDA40FEAA55ED27C6FEF0" box="[376,495,735,762]" metricMagnitude="-1" metricUnit="m" metricValue="1.4849999999999999" pageId="24" pageNumber="25" unit="mm" value="148.5">148.5 mm</quantity>
|
||
SL;
|
||
<collectingRegion id="49E2F852611FDA40FDFA55ED24E0FEF3" box="[552,713,735,761]" pageId="24" pageNumber="25">Lake Edward</collectingRegion>
|
||
;
|
||
<geoCoordinate id="EE125077611FDA40FD0655EC2542FEF3" box="[724,875,734,762]" degrees="0" direction="south" minutes="24" orientation="latitude" pageId="24" pageNumber="25" precision="1" seconds="16.0" value="-0.40444446">0°24′16.0″ S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE125077611FDA40FCAA55EC2208FEF0" box="[888,1057,734,762]" degrees="29" direction="east" minutes="46" orientation="longitude" pageId="24" pageNumber="25" precision="1" seconds="24.8" value="29.773556">29°46′24.8″ E</geoCoordinate>
|
||
;
|
||
<date id="FF981070611FDA40FBFF55ED22EFFEF3" box="[1069,1222,735,762]" pageId="24" pageNumber="25" value="2018-01-24">
|
||
<collectingDate id="EFDCE998611FDA40FBFF55ED22EFFEF3" box="[1069,1222,735,762]" pageId="24" pageNumber="25" value="2018-01-24">24 Jan. 2018</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="26D35366611FDA40FB0055ED235EFEF3" box="[1234,1399,735,761]" pageId="24" pageNumber="25">HIPE3 exped.</collectorName>
|
||
leg.; bought at
|
||
<location id="8EF9606B611FDA40FEBF543027D5FF16" LSID="urn:lsid:plazi:treatment:038F87A66110DA44FDA855B72794FF90:8EF9606B611FDA40FEBF543027D5FF16" box="[365,508,770,796]" country="Uganda" latitude="-0.40444446" longLatPrecision="1" longitude="29.773556" name="Rwenshama" pageId="24" pageNumber="25" stateProvince="Lake Edward">Rwenshama</location>
|
||
landing site;
|
||
<specimenCode id="DB809ECB611FDA40FD49543125E9FF17" box="[667,960,770,797]" pageId="24" pageNumber="25">RMCA 2018.008.P.0345</specimenCode>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8B9936B0611FDA40FF385476257DF85F" blockId="24.[189,1399,836,1109]" pageId="24" pageNumber="25">
|
||
<materialsCitation id="3B4E3CED611FDA40FF38547624AFFFC2" ID-GBIF-Occurrence="3861203388" collectingDate="1953-03-26" collectionCode="IRSNB" collectorName="KEA exped." country="Democratic Republic of the Congo" elevation="500" latitude="-0.68072224" location="Vitshumbi" longLatPrecision="1" longitude="29.38961" pageId="24" pageNumber="25" specimenCode="IRSNB 13474" specimenCount="1" specimenCount-male="1" stateProvince="Lake Edward" typeStatus="paratype">
|
||
<emphasis id="B952EAA2611FDA40FF385476274EFF54" bold="true" box="[234,359,836,862]" pageId="24" pageNumber="25">
|
||
<typeStatus id="549D8812611FDA40FF385476274EFF54" box="[234,359,836,862]" pageId="24" pageNumber="25" type="paratype">Paratypes</typeStatus>
|
||
</emphasis>
|
||
<collectingCountry id="F3317620611FDA40FF6F545524C5FF8B" box="[189,748,871,898]" name="Democratic Republic of the Congo" pageId="24" pageNumber="25">DEMOCRATIC REPUBLIC OF THE CONGO</collectingCountry>
|
||
•
|
||
<specimenCount id="9D20FD39611FDA40FCD75455251AFF88" box="[773,819,871,898]" pageId="24" pageNumber="25" type="male">1 ♂</specimenCount>
|
||
,
|
||
<quantity id="4CDE9B55611FDA40FC925455259EFF88" box="[832,951,871,898]" metricMagnitude="-1" metricUnit="m" metricValue="1.315" pageId="24" pageNumber="25" unit="mm" value="131.5">131.5 mm</quantity>
|
||
SL; “Lac Edouard: Vitshumbi (contre le pier)” [
|
||
<collectingRegion id="49E2F852611FDA40FEFC54B827FDFFAE" box="[302,468,906,932]" pageId="24" pageNumber="25">Lake Edward</collectingRegion>
|
||
:
|
||
<location id="8EF9606B611FDA40FE0D54B82470FFAE" LSID="urn:lsid:plazi:treatment:038F87A66110DA44FDA855B72794FF90:8EF9606B611FDA40FE0D54B82470FFAE" box="[479,601,906,932]" country="Democratic Republic of the Congo" latitude="-0.68072224" longLatPrecision="1" longitude="29.38961" municipality="Vitshumbi" name="Vitshumbi" pageId="24" pageNumber="25" stateProvince="Lake Edward">Vitshumbi</location>
|
||
(against the pier)];
|
||
<geoCoordinate id="EE125077611FDA40FC9654BB25F5FFAE" box="[836,988,905,933]" degrees="0" direction="south" minutes="40" orientation="latitude" pageId="24" pageNumber="25" precision="1" seconds="50.6" value="-0.68072224">0°40′50.6″ S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE125077611FDA40FC3854BB22B8FFAF" box="[1002,1169,905,933]" degrees="29" direction="east" minutes="23" orientation="longitude" pageId="24" pageNumber="25" precision="1" seconds="22.6" value="29.38961">29°23′22.6″ E</geoCoordinate>
|
||
(inferred);
|
||
<date id="FF981070611FDA40FAC854B826D2FFC2" pageId="24" pageNumber="25" value="1953-03-26">
|
||
<collectingDate id="EFDCE998611FDA40FAC854B826D2FFC2" pageId="24" pageNumber="25" value="1953-03-26">26 Mar. 1953</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="26D35366611FDA40FEDA549C27B3FFCD" box="[264,410,941,968]" pageId="24" pageNumber="25">KEA exped.</collectorName>
|
||
leg.;
|
||
<specimenCode id="DB809ECB611FDA40FE0C549C24AFFFC2" box="[478,646,942,968]" collectionCode="IRSNB" country="Belgium" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="24" pageNumber="25">IRSNB 13474</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3B4E3CED611FDA40FD70549C25FAF804" ID-GBIF-Occurrence="3861203409" collectingDate="1953-06-01" collectionCode="IRSNB" collectorName="KEA exped." country="Democratic Republic of the Congo" elevation="500" latitude="-0.68072224" location="Kiavinionge" longLatPrecision="1" longitude="29.38961" pageId="24" pageNumber="25" specimenCode="IRSNB 13480" specimenCount="1" specimenCount-male="1" stateProvince="Lake Edward" typeStatus="paratype">
|
||
<specimenCount id="9D20FD39611FDA40FD70549C24F8FFC2" box="[674,721,942,968]" pageId="24" pageNumber="25" type="male">1 ♂</specimenCount>
|
||
,
|
||
<quantity id="4CDE9B55611FDA40FD0D549F257EFFC2" box="[735,855,941,968]" metricMagnitude="-1" metricUnit="m" metricValue="1.5419999999999998" pageId="24" pageNumber="25" unit="mm" value="154.2">154.2 mm</quantity>
|
||
SL; “Lac Edouard:
|
||
<quantity id="4CDE9B55611FDA40FB9A549C228BFFCD" box="[1096,1186,941,968]" metricMagnitude="3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="24" pageNumber="25" unit="km" value="2.5" valueMax="3.0" valueMin="2.0">2–3 km</quantity>
|
||
±
|
||
<quantity id="4CDE9B55611FDA40FB16549F2339FFC2" box="[1220,1296,941,968]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="24" pageNumber="25" unit="m" value="500.0">500 m</quantity>
|
||
au large à l’Ouest de Kiavinionge” [
|
||
<collectingRegion id="49E2F852611FDA40FDD354E3248FFFE1" box="[513,678,977,1003]" pageId="24" pageNumber="25">Lake Edward</collectingRegion>
|
||
:
|
||
<locationDeviation id="9A2B2F92611FDA40FD6054E325F6FFE1" box="[690,991,977,1004]" pageId="24" pageNumber="25">
|
||
<quantity id="4CDE9B55611FDA40FD6054E32522FFE1" box="[690,779,977,1003]" metricMagnitude="3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="24" pageNumber="25" unit="km" value="2.5" valueMax="3.0" valueMin="2.0">2–3 km</quantity>
|
||
±
|
||
<quantity id="4CDE9B55611FDA40FCF954E3255FFFE1" box="[811,886,977,1004]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="24" pageNumber="25" unit="m" value="500.0">500 m</quantity>
|
||
offshore
|
||
</locationDeviation>
|
||
west of
|
||
<location id="8EF9606B611FDA40FB9754E222F4FFE1" LSID="urn:lsid:plazi:treatment:038F87A66110DA44FDA855B72794FF90:8EF9606B611FDA40FB9754E222F4FFE1" box="[1093,1245,976,1003]" country="Democratic Republic of the Congo" latitude="-0.68072224" longLatPrecision="1" longitude="29.38961" municipality="Vitshumbi" name="Kiavinionge" pageId="24" pageNumber="25" stateProvince="Lake Edward">Kiavinionge</location>
|
||
];
|
||
<geoCoordinate id="EE125077611FDA40FB2254FD2358FFE1" box="[1264,1393,975,1003]" degrees="0" direction="south" minutes="11" orientation="latitude" pageId="24" pageNumber="25" precision="15" seconds="39" value="-0.19416668">0°11′39″ S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE125077611FDA40FF6F54C02762F804" box="[189,331,1010,1038]" degrees="29" direction="east" minutes="32" orientation="longitude" pageId="24" pageNumber="25" precision="15" seconds="31" value="29.541945">29°32′31″ E</geoCoordinate>
|
||
(inferred);
|
||
<date id="FF981070611FDA40FE1D54C6247EF805" box="[463,599,1012,1039]" pageId="24" pageNumber="25" value="1953-06-01">
|
||
<collectingDate id="EFDCE998611FDA40FE1D54C6247EF805" box="[463,599,1012,1039]" pageId="24" pageNumber="25" value="1953-06-01">1 Jun. 1953</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="26D35366611FDA40FDB054C624D9F804" box="[610,752,1012,1038]" pageId="24" pageNumber="25">KEA exped.</collectorName>
|
||
leg.;
|
||
<specimenCode id="DB809ECB611FDA40FCFC54C625FAF804" box="[814,979,1012,1038]" collectionCode="IRSNB" country="Belgium" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="24" pageNumber="25">IRSNB 13480</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3B4E3CED611FDA40FC3B54C62579F85F" ID-GBIF-Occurrence="3861203383" collectingDate="1953-06-01" collectionCode="IRSNB" collectorName="KEA exped." country="Democratic Republic of the Congo" elevation="500" latitude="-0.68072224" location="Kiavinionge" longLatPrecision="1" longitude="29.38961" pageId="24" pageNumber="25" specimenCode="IRSNB 13485" specimenCount="1" specimenCount-female="1" stateProvince="Lake Edward" typeStatus="paratype">
|
||
<specimenCount id="9D20FD39611FDA40FC3B54C6223DF804" box="[1001,1044,1012,1038]" pageId="24" pageNumber="25" type="female">1 ♀</specimenCount>
|
||
,
|
||
<quantity id="4CDE9B55611FDA40FBF354C622BEF804" box="[1057,1175,1012,1039]" metricMagnitude="-1" metricUnit="m" metricValue="1.548" pageId="24" pageNumber="25" unit="mm" value="154.8">154.8 mm</quantity>
|
||
SL; “Lac Edouard:
|
||
<quantity id="4CDE9B55611FDA40FF6F5325273DF83B" box="[189,276,1047,1073]" metricMagnitude="3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="24" pageNumber="25" unit="km" value="2.5" valueMax="3.0" valueMin="2.0">2–3 km</quantity>
|
||
à l’Ouest de Kiavinionge” [
|
||
<collectingRegion id="49E2F852611FDA40FD86532524DEF83B" box="[596,759,1047,1073]" pageId="24" pageNumber="25">Lake Edward</collectingRegion>
|
||
:
|
||
<locationDeviation id="9A2B2F92611FDA40FCD2532525BAF838" box="[768,915,1047,1074]" pageId="24" pageNumber="25">
|
||
<quantity id="4CDE9B55611FDA40FCD25325257EF83B" box="[768,855,1047,1073]" metricMagnitude="3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="24" pageNumber="25" unit="km" value="2.5" valueMax="3.0" valueMin="2.0">2–3 km</quantity>
|
||
west
|
||
</locationDeviation>
|
||
of
|
||
<location id="8EF9606B611FDA40FC6453252267F838" LSID="urn:lsid:plazi:treatment:038F87A66110DA44FDA855B72794FF90:8EF9606B611FDA40FC6453252267F838" box="[950,1102,1047,1074]" country="Democratic Republic of the Congo" latitude="-0.68072224" longLatPrecision="1" longitude="29.38961" municipality="Vitshumbi" name="Kiavinionge" pageId="24" pageNumber="25" stateProvince="Lake Edward">Kiavinionge</location>
|
||
];
|
||
<geoCoordinate id="EE125077611FDA40FB8D532422F4F83B" box="[1119,1245,1046,1074]" degrees="0" direction="south" minutes="11" orientation="latitude" pageId="24" pageNumber="25" precision="15" seconds="39" value="-0.19416668">0°11′39″ S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE125077611FDA40FB3B5324235FF838" box="[1257,1398,1046,1074]" degrees="29" direction="east" minutes="32" orientation="longitude" pageId="24" pageNumber="25" precision="15" seconds="31" value="29.541945">29°32′31″ E</geoCoordinate>
|
||
(inferred);
|
||
<date id="FF981070611FDA40FEEF530827E1F85F" box="[317,456,1082,1109]" pageId="24" pageNumber="25" value="1953-06-01">
|
||
<collectingDate id="EFDCE998611FDA40FEEF530827E1F85F" box="[317,456,1082,1109]" pageId="24" pageNumber="25" value="1953-06-01">1 Jun. 1953</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="26D35366611FDA40FE065309244DF85E" box="[468,612,1082,1109]" pageId="24" pageNumber="25">KEA exped.</collectorName>
|
||
leg.;
|
||
<specimenCode id="DB809ECB611FDA40FD7453082579F85F" box="[678,848,1082,1109]" collectionCode="IRSNB" country="Belgium" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="24" pageNumber="25">IRSNB 13485</specimenCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9936B0611FDA41FF6F534E236BFE9A" blockId="24.[189,1399,1146,1210]" lastBlockId="25.[189,1399,263,657]" lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25">
|
||
<materialsCitation id="3B4E3CED611FDA40FF6F534E2272F8B0" ID-GBIF-Occurrence="3861203372" collectingDate="2016-10-23" collectionCode="RMCA" collectorName="HIPE 1 exped." country="Uganda" elevation="-20" lastPageId="24" lastPageNumber="26" latitude="-0.2" location="Lake Edward" longLatPrecision="1" longitude="29.793999" pageId="24" pageNumber="25" specimenCode="RMCA 2016.035.P.0220" specimenCount="1" specimenCount-male="1" stateProvince="Lake Edward">
|
||
<collectingCountry id="F3317620611FDA40FF6F534E2713F89C" box="[189,314,1148,1174]" name="Uganda" pageId="24" pageNumber="25">UGANDA</collectingCountry>
|
||
–
|
||
<emphasis id="B952EAA2611FDA40FE8F534E2425F89C" bold="true" box="[349,524,1148,1174]" pageId="24" pageNumber="25">
|
||
<collectingRegion id="49E2F852611FDA40FE8F534E2425F89C" box="[349,524,1148,1174]" pageId="24" pageNumber="25">Lake Edward</collectingRegion>
|
||
</emphasis>
|
||
•
|
||
<specimenCount id="9D20FD39611FDA40FDF5534E247EF89C" box="[551,599,1148,1174]" pageId="24" pageNumber="25" type="male">1 ♂</specimenCount>
|
||
,
|
||
<quantity id="4CDE9B55611FDA40FDB7534E24F4F89C" box="[613,733,1148,1174]" metricMagnitude="-1" metricUnit="m" metricValue="1.144" pageId="24" pageNumber="25" unit="mm" value="114.4">114.4 mm</quantity>
|
||
SL;
|
||
<geoCoordinate id="EE125077611FDA40FCCB5348259BF89C" box="[793,946,1146,1174]" degrees="0" direction="south" minutes="12" orientation="latitude" pageId="24" pageNumber="25" precision="1" seconds="00.0" value="-0.2">0°12′00.0″ S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE125077611FDA40FC1353482242F89C" box="[961,1131,1146,1174]" degrees="29" direction="east" minutes="47" orientation="longitude" pageId="24" pageNumber="25" precision="1" seconds="38.4" value="29.793999">29°47′38.4″ E</geoCoordinate>
|
||
;
|
||
<date id="FF981070611FDA40FBA8534E2330F89C" box="[1146,1305,1148,1174]" pageId="24" pageNumber="25" value="2016-10-23">
|
||
<collectingDate id="EFDCE998611FDA40FBA8534E2330F89C" box="[1146,1305,1148,1174]" pageId="24" pageNumber="25" value="2016-10-23">23 Oct. 2016</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="26D35366611FDA40FAF5534E2722F8B3" pageId="24" pageNumber="25">HIPE1 exped.</collectorName>
|
||
leg.; deep catch, open water ±
|
||
<quantity id="4CDE9B55611FDA40FD4E53AD24F4F8B0" box="[668,733,1183,1210]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="24" pageNumber="25" unit="m" value="20.0">
|
||
<elevation id="000BD183611FDA40FD4E53AD24F4F8B0" box="[668,733,1183,1210]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="24" pageNumber="25" unit="m" value="20.0">20 m</elevation>
|
||
</quantity>
|
||
deep;
|
||
<specimenCode id="DB809ECB611FDA40FCE753AD2272F8B0" box="[821,1115,1183,1210]" pageId="24" pageNumber="25">RMCA 2016.035.P.0220</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3B4E3CED611FDA41FBAC53AD27ABFD42" ID-GBIF-Occurrence="3861203377" collectingDate="2016-10-24" collectionCode="RMCA" collectorName="HIPE 1 exped." country="Uganda" elevation="20" lastPageId="25" lastPageNumber="26" latitude="-0.2" location="Katwe" longLatPrecision="1" longitude="29.793999" pageId="24" pageNumber="25" specimenCode="RMCA 2016.035.P.0221" specimenCount="1" specimenCount-male="1" stateProvince="Lake Edward">
|
||
<specimenCount id="9D20FD39611FDA40FBAC53AD229BF8B0" box="[1150,1202,1183,1210]" pageId="24" pageNumber="25" type="male">1 ♂</specimenCount>
|
||
,
|
||
<quantity id="4CDE9B55611FDA40FB1153AD2369F8B0" box="[1219,1344,1183,1210]" metricMagnitude="-1" metricUnit="m" metricValue="1.545" pageId="24" pageNumber="25" unit="mm" value="154.5">154.5 mm</quantity>
|
||
SL;
|
||
<geoCoordinate id="EE125077611EDA41FF6F5635277FFD29" box="[189,342,263,291]" degrees="0" direction="south" minutes="08" orientation="latitude" pageId="25" pageNumber="26" precision="1" seconds="51.9" value="-0.14775">0°08′51.9″ S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE125077611EDA41FEB756352426FD29" box="[357,527,263,291]" degrees="29" direction="east" minutes="52" orientation="longitude" pageId="25" pageNumber="26" precision="1" seconds="59.6" value="29.883223">29°52′59.6″ E</geoCoordinate>
|
||
;
|
||
<date id="FF981070611EDA41FDCC563B2494FD29" box="[542,701,265,291]" pageId="25" pageNumber="26" value="2016-10-24">
|
||
<collectingDate id="EFDCE998611EDA41FDCC563B2494FD29" box="[542,701,265,291]" pageId="25" pageNumber="26" value="2016-10-24">24 Oct. 2016</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="26D35366611EDA41FD19563B255BFD29" box="[715,882,265,291]" pageId="25" pageNumber="26">HIPE1 exped.</collectorName>
|
||
leg.; bought at
|
||
<location id="8EF9606B611EDA41FBE3563B2257FD29" LSID="urn:lsid:plazi:treatment:038F87A66110DA44FDA855B72794FF90:8EF9606B611EDA41FBE3563B2257FD29" box="[1073,1150,265,291]" country="Uganda" latitude="-0.2" longLatPrecision="1" longitude="29.793999" name="Katwe" pageId="25" pageNumber="26" stateProvince="Lake Edward">Katwe</location>
|
||
landing site;
|
||
<specimenCode id="DB809ECB611EDA41FAF0563B27ABFD42" pageId="25" pageNumber="26">RMCA 2016.035.P.0221</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3B4E3CED611EDA41FE76561C2573FD67" ID-GBIF-Occurrence="3861203385" collectingDate="2017-03-26" collectionCode="RMCA" collectorName="HIPE 2 exped." country="Uganda" elevation="20" lastPageId="25" lastPageNumber="26" latitude="-0.2" location="Rwenshama" longLatPrecision="1" longitude="29.793999" pageId="25" pageNumber="25" specimenCode="RMCA 2017.006.P.0352" specimenCount="1" specimenCount-male="1" stateProvince="Lake Edward">
|
||
<specimenCount id="9D20FD39611EDA41FE76561C27FEFD42" box="[420,471,302,328]" pageId="25" pageNumber="26" type="male">1 ♂</specimenCount>
|
||
,
|
||
<quantity id="4CDE9B55611EDA41FE3B561C244DFD42" box="[489,612,302,328]" metricMagnitude="-1" metricUnit="m" metricValue="1.147" pageId="25" pageNumber="26" unit="mm" value="114.7">114.7 mm</quantity>
|
||
SL;
|
||
<location id="8EF9606B611EDA41FD74561F2511FD4D" LSID="urn:lsid:plazi:treatment:038F87A66110DA44FDA855B72794FF90:8EF9606B611EDA41FD74561F2511FD4D" box="[678,824,301,327]" country="Uganda" latitude="-0.2" longLatPrecision="1" longitude="29.793999" name="Rwenshama" pageId="25" pageNumber="26" stateProvince="Lake Edward">Rwenshama</location>
|
||
, rocky shore;
|
||
<geoCoordinate id="EE125077611EDA41FC3E561E22A1FD42" box="[1004,1160,300,328]" degrees="0" direction="south" minutes="24" orientation="latitude" pageId="25" pageNumber="26" precision="1" seconds="05.7" value="-0.40158334">0°24′05.7″ S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE125077611EDA41FB48561E2361FD42" box="[1178,1352,300,328]" degrees="29" direction="east" minutes="46" orientation="longitude" pageId="25" pageNumber="26" precision="1" seconds="35.1" value="29.776417">29°46′35.1″ E</geoCoordinate>
|
||
;
|
||
<date id="FF981070611EDA41FA8B561F2712FD66" pageId="25" pageNumber="26" value="2017-03-26">
|
||
<collectingDate id="EFDCE998611EDA41FA8B561F2712FD66" pageId="25" pageNumber="26" value="2017-03-26">26 Mar. 2017</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="26D35366611EDA41FE98566027D8FD66" box="[330,497,338,364]" pageId="25" pageNumber="26">HIPE2 exped.</collectorName>
|
||
leg.;
|
||
<specimenCode id="DB809ECB611EDA41FDE556602573FD67" box="[567,858,338,365]" pageId="25" pageNumber="26">RMCA 2017.006.P.0352</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3B4E3CED611EDA41FCAA566026D1FDBC" ID-GBIF-Occurrence="3861203311" collectingDate="2017-03-31" collectionCode="RMCA" collectorName="HIPE 2 exped." country="Uganda" elevation="20" lastPageId="25" lastPageNumber="26" latitude="-0.2" location="Kayanja" longLatPrecision="1" longitude="29.793999" pageId="25" pageNumber="25" specimenCode="RMCA 2017.006.P.0353, RMCA 2017.006.P.0354, RMCA 2017.006.P.0355" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Lake Edward">
|
||
<specimenCount id="9D20FD39611EDA41FCAA56602581FD67" box="[888,936,338,365]" pageId="25" pageNumber="26" type="male">1 ♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D20FD39611EDA41FC65566025D4FD67" box="[951,1021,338,365]" pageId="25" pageNumber="26" type="female">2 ♀♀</specimenCount>
|
||
, 101.0–140.0 mm SL;
|
||
<location id="8EF9606B611EDA41FAC75660235EFD66" LSID="urn:lsid:plazi:treatment:038F87A66110DA44FDA855B72794FF90:8EF9606B611EDA41FAC75660235EFD66" box="[1301,1399,338,364]" country="Uganda" latitude="-0.2" longLatPrecision="1" longitude="29.793999" name="Kayanja" pageId="25" pageNumber="26" stateProvince="Lake Edward">Kayanja</location>
|
||
offshore;
|
||
<geoCoordinate id="EE125077611EDA41FEE2564727E0FD9B" box="[304,457,373,401]" degrees="0" direction="south" minutes="05" orientation="latitude" pageId="25" pageNumber="26" precision="1" seconds="34.8" value="-0.093">0°05′34.8″ S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE125077611EDA41FE0B564724ADFD9B" box="[473,644,373,401]" degrees="29" direction="east" minutes="45" orientation="longitude" pageId="25" pageNumber="26" precision="1" seconds="28.8" value="29.758">29°45′28.8″ E</geoCoordinate>
|
||
;
|
||
<date id="FF981070611EDA41FD405645251EFD9B" box="[658,823,375,401]" pageId="25" pageNumber="26" value="2017-03-31">
|
||
<collectingDate id="EFDCE998611EDA41FD405645251EFD9B" box="[658,823,375,401]" pageId="25" pageNumber="26" value="2017-03-31">31 Mar. 2017</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="26D35366611EDA41FC94564525C4FD9A" box="[838,1005,374,401]" pageId="25" pageNumber="26">HIPE2 exped.</collectorName>
|
||
leg.;
|
||
<specimenCode id="DB809ECB611EDA41FBE1564526D1FDBC" pageId="25" pageNumber="26">RMCA 2017.006.P.0353 to 0355</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3B4E3CED611EDA41FEC056A92435FDD0" ID-GBIF-Occurrence="3861203323" collectingDate="2018-01-20" collectionCode="RMCA" collectorName="HIPE 3 exped." country="Uganda" elevation="20" lastPageId="25" lastPageNumber="26" latitude="-0.2" location="Kayanja" longLatPrecision="1" longitude="29.793999" pageId="25" pageNumber="25" specimenCode="RMCA 2018.008.P.0342" specimenCount="1" specimenCount-female="1" stateProvince="Lake Edward">
|
||
<specimenCount id="9D20FD39611EDA41FEC056A92716FDBC" box="[274,319,411,438]" pageId="25" pageNumber="26" type="female">1 ♀</specimenCount>
|
||
,
|
||
<quantity id="4CDE9B55611EDA41FE9D56A92791FDBC" box="[335,440,411,438]" metricMagnitude="-2" metricUnit="m" metricValue="8.05" pageId="25" pageNumber="26" unit="mm" value="80.5">80.5 mm</quantity>
|
||
SL;
|
||
<location id="8EF9606B611EDA41FE2056A9247DFDBF" LSID="urn:lsid:plazi:treatment:038F87A66110DA44FDA855B72794FF90:8EF9606B611EDA41FE2056A9247DFDBF" box="[498,596,411,437]" country="Uganda" latitude="-0.2" longLatPrecision="1" longitude="29.793999" name="Kayanja" pageId="25" pageNumber="26" stateProvince="Lake Edward">Kayanja</location>
|
||
offshore;
|
||
<geoCoordinate id="EE125077611EDA41FD1C56AB254FFDBF" box="[718,870,409,437]" degrees="0" direction="south" minutes="05" orientation="latitude" pageId="25" pageNumber="26" precision="1" seconds="31.2" value="-0.092">0°05′31.2″ S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE125077611EDA41FCA156AB2234FDBC" box="[883,1053,409,438]" degrees="29" direction="east" minutes="45" orientation="longitude" pageId="25" pageNumber="26" precision="1" seconds="30.3" value="29.758417">29°45′30.3″ E</geoCoordinate>
|
||
;
|
||
<date id="FF981070611EDA41FBF856A922EDFDBF" box="[1066,1220,411,438]" pageId="25" pageNumber="26" value="2018-01-20">
|
||
<collectingDate id="EFDCE998611EDA41FBF856A922EDFDBF" box="[1066,1220,411,438]" pageId="25" pageNumber="26" value="2018-01-20">20 Jan. 2018</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="26D35366611EDA41FB0356A9235EFDBF" box="[1233,1399,411,437]" pageId="25" pageNumber="26">HIPE3 exped.</collectorName>
|
||
leg.;
|
||
<specimenCode id="DB809ECB611EDA41FF2856F22435FDD0" box="[250,540,448,474]" pageId="25" pageNumber="26">RMCA 2018.008.P.0342</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3B4E3CED611EDA41FDEB56F2230FFDF4" ID-GBIF-Occurrence="3861203338" collectingDate="2018-01-24" collectionCode="RMCA" collectorName="HIPE 3 exped." country="Uganda" elevation="20" lastPageId="25" lastPageNumber="26" latitude="-0.2" location="Rwenshama" longLatPrecision="1" longitude="29.793999" pageId="25" pageNumber="25" specimenCode="RMCA 2018.008.P.0343 to 0344" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Lake Edward">
|
||
<specimenCount id="9D20FD39611EDA41FDEB56F22440FDD0" box="[569,617,448,474]" pageId="25" pageNumber="26" type="male">1 ♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D20FD39611EDA41FDAA56F2248FFDD0" box="[632,678,448,474]" pageId="25" pageNumber="26" type="female">1 ♀</specimenCount>
|
||
, 134.6,
|
||
<quantity id="4CDE9B55611EDA41FCD856F225AAFDD0" box="[778,899,448,474]" metricMagnitude="-1" metricUnit="m" metricValue="1.4769999999999999" pageId="25" pageNumber="26" unit="mm" value="147.7">147.7 mm</quantity>
|
||
SL;
|
||
<geoCoordinate id="EE125077611EDA41FC12568C2270FDD0" box="[960,1113,446,474]" degrees="0" direction="south" minutes="24" orientation="latitude" pageId="25" pageNumber="26" precision="1" seconds="16.0" value="-0.40444446">0°24′16.0″ S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE125077611EDA41FBBA568C233AFDD0" box="[1128,1299,446,474]" degrees="29" direction="east" minutes="46" orientation="longitude" pageId="25" pageNumber="26" precision="1" seconds="24.8" value="29.773556">29°46′24.8″ E</geoCoordinate>
|
||
;
|
||
<date id="FF981070611EDA41FAF356F226D2FDF4" pageId="25" pageNumber="26" value="2018-01-24">
|
||
<collectingDate id="EFDCE998611EDA41FAF356F226D2FDF4" pageId="25" pageNumber="26" value="2018-01-24">24 Jan. 2018</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="26D35366611EDA41FEDB56D62786FDF4" box="[265,431,484,510]" pageId="25" pageNumber="26">HIPE3 exped.</collectorName>
|
||
leg.; bought at
|
||
<location id="8EF9606B611EDA41FDBF56D624D5FDF4" LSID="urn:lsid:plazi:treatment:038F87A66110DA44FDA855B72794FF90:8EF9606B611EDA41FDBF56D624D5FDF4" box="[621,764,484,510]" country="Uganda" latitude="-0.2" longLatPrecision="1" longitude="29.793999" name="Rwenshama" pageId="25" pageNumber="26" stateProvince="Lake Edward">Rwenshama</location>
|
||
landing site;
|
||
<specimenCode id="DB809ECB611EDA41FC7256D6230FFDF4" box="[928,1318,484,511]" pageId="25" pageNumber="26">RMCA 2018.008.P.0343 to 0344</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3B4E3CED611EDA41FA9356D627C9FE4D" ID-GBIF-Occurrence="3861203358" collectingDate="2018-01-24" collectionCode="RMCA" collectorName="HIPE 3 exped." country="Uganda" elevation="20" lastPageId="25" lastPageNumber="26" latitude="-0.2" location="Rwenshama" longLatPrecision="1" longitude="29.793999" pageId="25" pageNumber="25" specimenCode="RMCA 2018.008.P.0346" specimenCount="1" specimenCount-female="1" stateProvince="Lake Edward">
|
||
<specimenCount id="9D20FD39611EDA41FA9356D62346FDF5" box="[1345,1391,484,511]" pageId="25" pageNumber="26" type="female">1 ♀</specimenCount>
|
||
,
|
||
<quantity id="4CDE9B55611EDA41FF6F553B2700FE29" box="[189,297,521,547]" metricMagnitude="-2" metricUnit="m" metricValue="8.42" pageId="25" pageNumber="26" unit="mm" value="84.2">84.2 mm</quantity>
|
||
SL;
|
||
<location id="8EF9606B611EDA41FEBA553B27D3FE29" LSID="urn:lsid:plazi:treatment:038F87A66110DA44FDA855B72794FF90:8EF9606B611EDA41FEBA553B27D3FE29" box="[360,506,521,547]" country="Uganda" latitude="-0.2" longLatPrecision="1" longitude="29.793999" name="Rwenshama" pageId="25" pageNumber="26" stateProvince="Lake Edward">Rwenshama</location>
|
||
, offshore;
|
||
<geoCoordinate id="EE125077611EDA41FDAF5535253EFE29" box="[637,791,519,547]" degrees="0" direction="south" minutes="24" orientation="latitude" pageId="25" pageNumber="26" precision="1" seconds="14.4" value="-0.404">0°24′14.4″ S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE125077611EDA41FCF5553525FAFE29" box="[807,979,519,547]" degrees="29" direction="east" minutes="45" orientation="longitude" pageId="25" pageNumber="26" precision="1" seconds="57.0" value="29.765833">29°45′57.0″ E</geoCoordinate>
|
||
;
|
||
<date id="FF981070611EDA41FC31553B22ABFE29" box="[995,1154,521,547]" pageId="25" pageNumber="26" value="2018-01-24">
|
||
<collectingDate id="EFDCE998611EDA41FC31553B22ABFE29" box="[995,1154,521,547]" pageId="25" pageNumber="26" value="2018-01-24">24 Jan. 2018</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="26D35366611EDA41FB43553B2310FE29" box="[1169,1337,521,547]" pageId="25" pageNumber="26">HIPE3 exped.</collectorName>
|
||
leg.;
|
||
<specimenCode id="DB809ECB611EDA41FF6F551F27C9FE4D" box="[189,480,557,583]" pageId="25" pageNumber="26">RMCA 2018.008.P.0346</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3B4E3CED611EDA41FDD2551F222AFE66" ID-GBIF-Occurrence="3861203366" collectingDate="2018-02-04" collectionCode="RMCA" collectorName="HIPE 3 exped." country="Uganda" elevation="20" lastPageId="25" lastPageNumber="26" latitude="-0.2" location="Kagoro" longLatPrecision="1" longitude="29.793999" pageId="25" pageNumber="25" specimenCode="RMCA 2018.008.P.0347" specimenCount="1" specimenCount-male="1" stateProvince="Lake Edward">
|
||
<specimenCount id="9D20FD39611EDA41FDD2551F2418FE42" box="[512,561,557,584]" pageId="25" pageNumber="26" type="male">1 ♂</specimenCount>
|
||
,
|
||
<quantity id="4CDE9B55611EDA41FD92551F2492FE42" box="[576,699,557,584]" metricMagnitude="-1" metricUnit="m" metricValue="1.283" pageId="25" pageNumber="26" unit="mm" value="128.3">128.3 mm</quantity>
|
||
SL;
|
||
<location id="8EF9606B611EDA41FD2B551C2578FE42" LSID="urn:lsid:plazi:treatment:038F87A66110DA44FDA855B72794FF90:8EF9606B611EDA41FD2B551C2578FE42" box="[761,849,558,584]" country="Uganda" latitude="-0.2" longLatPrecision="1" longitude="29.793999" name="Kagoro" pageId="25" pageNumber="26" stateProvince="Lake Edward">Kagoro</location>
|
||
fishing ground;
|
||
<geoCoordinate id="EE125077611EDA41FBCE551E229FFE4D" box="[1052,1206,556,584]" degrees="0" direction="south" minutes="12" orientation="latitude" pageId="25" pageNumber="26" precision="1" seconds="50.1" value="-0.21391667">0°12′50.1″ S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE125077611EDA41FB14551E2358FE42" box="[1222,1393,556,584]" degrees="29" direction="east" minutes="49" orientation="longitude" pageId="25" pageNumber="26" precision="1" seconds="19.7" value="29.82214">29°49′19.7″ E</geoCoordinate>
|
||
;
|
||
<date id="FF981070611EDA41FF6F55602779FE66" box="[189,336,594,620]" pageId="25" pageNumber="26" value="2018-02-04">
|
||
<collectingDate id="EFDCE998611EDA41FF6F55602779FE66" box="[189,336,594,620]" pageId="25" pageNumber="26" value="2018-02-04">4 Feb. 2018</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="26D35366611EDA41FE8C5560242CFE66" box="[350,517,594,620]" pageId="25" pageNumber="26">HIPE3 exped.</collectorName>
|
||
leg.; open water;
|
||
<specimenCode id="DB809ECB611EDA41FD325560222AFE66" box="[736,1027,594,620]" pageId="25" pageNumber="26">RMCA 2018.008.P.0347</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3B4E3CED611EDA41FBF255602317FE9A" ID-GBIF-Occurrence="3861203357" collectingDate="2019-03-21" collectionCode="RMCA" collectorName="HIPE 4 exped." country="Uganda" elevation="20" lastPageId="25" lastPageNumber="26" latitude="-0.2" location="Kayanja" longLatPrecision="1" longitude="29.793999" pageId="25" pageNumber="25" specimenCode="RMCA 2019.002.P.0014" specimenCount="1" specimenCount-female="1" stateProvince="Lake Edward">
|
||
<specimenCount id="9D20FD39611EDA41FBF255602267FE66" box="[1056,1102,594,620]" pageId="25" pageNumber="26" type="female">1 ♀</specimenCount>
|
||
,
|
||
<quantity id="4CDE9B55611EDA41FB8C556022F1FE66" box="[1118,1240,594,620]" metricMagnitude="-1" metricUnit="m" metricValue="1.3419999999999999" pageId="25" pageNumber="26" unit="mm" value="134.2">134.2 mm</quantity>
|
||
SL;
|
||
<location id="8EF9606B611EDA41FAC75560235EFE66" LSID="urn:lsid:plazi:treatment:038F87A66110DA44FDA855B72794FF90:8EF9606B611EDA41FAC75560235EFE66" box="[1301,1399,594,620]" country="Uganda" latitude="-0.2" longLatPrecision="1" longitude="29.793999" name="Kayanja" pageId="25" pageNumber="26" stateProvince="Lake Edward">Kayanja</location>
|
||
offshore;
|
||
<geoCoordinate id="EE125077611EDA41FEFC554727ECFE9A" box="[302,453,629,657]" degrees="0" direction="south" minutes="05" orientation="latitude" pageId="25" pageNumber="26" precision="1" seconds="34.8" value="-0.093">0°05′34.8″ S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE125077611EDA41FE0055472453FE9B" box="[466,634,629,657]" degrees="29" direction="east" minutes="45" orientation="longitude" pageId="25" pageNumber="26" precision="1" seconds="28.8" value="29.758">29°45′28.8″ E</geoCoordinate>
|
||
;
|
||
<date id="FF981070611EDA41FD555544250EFE9A" box="[647,807,630,657]" pageId="25" pageNumber="26" value="2019-03-21">
|
||
<collectingDate id="EFDCE998611EDA41FD555544250EFE9A" box="[647,807,630,657]" pageId="25" pageNumber="26" value="2019-03-21">21 Mar. 2019</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="26D35366611EDA41FCE6554425F0FE9A" box="[820,985,630,656]" pageId="25" pageNumber="26">HIPE4 exped.</collectorName>
|
||
leg.;
|
||
<specimenCode id="DB809ECB611EDA41FBC855442317FE9A" box="[1050,1342,630,656]" pageId="25" pageNumber="26">RMCA 2019.002.P.0014</specimenCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF596638611FDA40FF6F50992762FBE2" ID-DOI="http://doi.org/10.5281/zenodo.6484187" ID-Zenodo-Dep="6484187" httpUri="https://zenodo.org/record/6484187/files/figure.png" pageId="24" pageNumber="25" startId="24.[189,232,1963,1989]" targetBox="[189,1398,1294,1855]" targetPageId="24">
|
||
<paragraph id="8B9936B0611FDA40FF6F50992762FBE2" blockId="24.[189,1400,1963,2025]" pageId="24" pageNumber="25">
|
||
<emphasis id="B952EAA2611FDA40FF6F50992732FBCF" bold="true" box="[189,283,1963,1989]" pageId="24" pageNumber="25">Fig. 11.</emphasis>
|
||
<taxonomicName id="4C264D33611FDA40FEF6509927D3FBCF" authority="Vranken & Steenberge & Heylen & Decru & Snoeks, 2022" authorityName="Vranken & Steenberge & Heylen & Decru & Snoeks" authorityYear="2022" box="[292,506,1963,1989]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="rex" status="sp. nov.">
|
||
<emphasis id="B952EAA2611FDA40FEF6509927D3FBCF" box="[292,506,1963,1989]" italics="true" pageId="24" pageNumber="25">Haplochromis rex</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26157D9611FDA40FDD7509E244DFBCC" box="[517,612,1964,1990]" pageId="24" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype, ♂, 148.5 mm SL (RMCA 2018.008.P.0345). Drawn by N. Vranken.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C33C653B611EDA44FF6F558B2593FF0E" lastPageId="28" lastPageNumber="29" pageId="25" pageNumber="26" type="description">
|
||
<paragraph id="8B9936B0611EDA41FF6F558B2767FED9" blockId="25.[189,334,697,723]" box="[189,334,697,723]" pageId="25" pageNumber="26">
|
||
<heading id="D0D181DC611EDA41FF6F558B2767FED9" bold="true" box="[189,334,697,723]" fontSize="11" level="3" pageId="25" pageNumber="26" reason="3">
|
||
<emphasis id="B952EAA2611EDA41FF6F558B2767FED9" bold="true" box="[189,334,697,723]" pageId="25" pageNumber="26">Description</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8B9936B0611EDA41FF6F55DB237CF87B" blockId="25.[187,1401,745,1137]" pageId="25" pageNumber="26">
|
||
Based on
|
||
<specimenCount id="9D20FD39611EDA41FEE355DB27F9FF09" box="[305,464,745,771]" pageId="25" pageNumber="26" type="generic">16 specimens</specimenCount>
|
||
(
|
||
<quantity id="4CDE9B55611EDA41FE0D55DB24B2FF09" box="[479,667,745,772]" metricMagnitude="-1" metricUnit="m" metricValue="1.1765" metricValueMax="1.548" metricValueMin="0.805" pageId="25" pageNumber="26" unit="mm" value="117.65" valueMax="154.8" valueMin="80.5">80.5–154.8 mm</quantity>
|
||
SL); body shallow (
|
||
<tableCitation id="C6A4030B611EDA41FC5855DB25C8FF09" box="[906,993,745,771]" captionStart="Table 1" captionStartId="9.[182,252,548,576]" captionTargetPageId="9" captionText="Table 1. (continued on the next three pages). Measurements and counts of the piscivorous species of Haplochromis from the Lake Edward system." httpUri="http://table.plazi.org/id/DF596638610EDA51FF6455162158FE4A" pageId="25" pageNumber="26" tableUuid="DF596638610EDA51FF6455162158FE4A">Table 1</tableCitation>
|
||
) and oval to rectangular (
|
||
<figureCitation id="131D2A35611EDA41FAC055DB2341FF09" box="[1298,1384,745,771]" captionStart="Fig" captionStartId="24.[189,232,1963,1989]" captionTargetBox="[189,1398,1294,1855]" captionTargetId="figure-316@24.[189,1399,1294,1855]" captionTargetPageId="24" captionText="Fig. 11. Haplochromis rex sp. nov., holotype, ♂, 148.5 mm SL (RMCA 2018.008.P.0345). Drawn by N. Vranken." figureDoi="http://doi.org/10.5281/zenodo.6484187" httpUri="https://zenodo.org/record/6484187/files/figure.png" pageId="25" pageNumber="26">Fig. 11</figureCitation>
|
||
). Head long, very narrow, and with a (weakly) convex dorsal outline; cheek deep; lacrimal very deep; eye very small and very high on head; interorbital area very narrow. Snout long, rounded in dorsal view, blunt, and slopes at 40–50°; premaxillary pedicel long and strongly prominent. Jaws isognathous to weakly prognathous, long, very narrow, stout, and rounded in dorsal view; jaws expand slightly laterally, hereby often broader than snout. Gape large and slopes gently at 15–25°; maxilla extends to vertical through anterior margin of orbit. Lower jaw relatively deep and with a straight ventral outline in lateral view, mental prominence absent or weakly developed, and lower jaw side with an inclination of 30–35° to horizontal in anterior view. Upper jaw strongly expanded anteriorly. Lips and oral mucosa large. Neurocranium shallow, ethmo-vomerine block decurved, preorbital region very shallow (18–20% NL), orbital region shallow (26–29% NL), and supraoccipital crest shallow and wedge-shaped (
|
||
<figureCitation id="131D2A35611EDA41FB3353642361F87B" box="[1249,1352,1110,1137]" captionStart="Fig" captionStartId="26.[189,232,1892,1918]" captionTargetBox="[207,1383,278,1848]" captionTargetId="figure-19@26.[207,1383,262,1848]" captionTargetPageId="26" captionText="Fig. 12. Haplochromis rex sp. nov. a. Photograph of preserved holotype (RMCA 2018.008.P.0345; 148.5 mm SL). b. X-ray image of holotype c–d. Photographs of freshly caught specimens. c. Holotype, a dominant male. d. A female (RMCA 2017.006.P.0355; 135.7 mm SL) to illustrate the live colour patterns. The contrast was slightly enhanced." figureDoi="http://doi.org/10.5281/zenodo.6484191" httpUri="https://zenodo.org/record/6484191/files/figure.png" pageId="25" pageNumber="26">Fig. 12b</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9936B0611EDA41FF6F53AB275DF9DD" blockId="25.[188,1400,1177,1495]" pageId="25" pageNumber="26">
|
||
Outer oral teeth few, unicuspid, and large. Necks very stout, conical, and straight; crowns weakly recurved, rarely straight, and acutely pointed. Dental arcades rounded. Outer teeth widely and regularly set with neck-distances of 1–2 neck-widths. No enlarged teeth posterior in upper jaw. Inner teeth straight to recurved, unicuspid, and relatively small in small specimens (<
|
||
<quantity id="4CDE9B55611EDA41FC3D5234227DF92B" box="[1007,1108,1286,1313]" metricMagnitude="-1" metricUnit="m" metricValue="1.25" pageId="25" pageNumber="26" unit="mm" value="125.0">125 mm</quantity>
|
||
SL), and large in large specimens (>
|
||
<quantity id="4CDE9B55611EDA41FEB2521927E8F94C" box="[352,449,1323,1350]" metricMagnitude="-1" metricUnit="m" metricValue="1.25" pageId="25" pageNumber="26" unit="mm" value="125.0">125 mm</quantity>
|
||
SL). Tooth bands very slender crescent-shaped with 1–3 rows of inner teeth, and narrow posteriorly until only outer row remains past ⅔ length of tooth band in upper jaw, past ½ length of tooth band in lower jaw. Inner teeth widely and regularly set on 1–2 outer neck-widths from outer row; implantation erect in first row and recumbent in subsequent rows; size decreases slightly buccally and posteriorly.
|
||
</paragraph>
|
||
<paragraph id="8B9936B0611EDA41FF6F51332787FADB" blockId="25.[187,1400,1536,1745]" pageId="25" pageNumber="26">
|
||
Lower pharyngeal bone average in length, narrow, slim, and shallow over whole length (
|
||
<figureCitation id="131D2A35611EDA41FADE51322341FA10" box="[1292,1384,1536,1562]" captionStart="Fig" captionStartId="27.[189,232,1963,1989]" captionTargetBox="[190,1395,1029,1890]" captionTargetId="figure-244@27.[190,1397,1029,1892]" captionTargetPageId="27" captionText="Fig. 13. Haplochromis rexsp. nov. (IRSNB 13480; 154.2 mm SL). a. Dorsal view of the lower pharyngeal jaw. b. Lateral view of the lower pharyngeal jaw." figureDoi="http://doi.org/10.5281/zenodo.6484195" httpUri="https://zenodo.org/record/6484195/files/figure.png" pageId="25" pageNumber="26">Fig. 13</figureCitation>
|
||
). Pharyngeal teeth relatively large and slender; major cusps acutely pointed; cusp gaps concave; minor cusps and cusp protuberances mostly absent. Teeth in two median longitudinal rows equal in size and form to lateral teeth,
|
||
<quantity id="4CDE9B55611EDA41FE64515C27D8FA82" box="[438,497,1646,1672]" metricMagnitude="-1" metricUnit="m" metricValue="2.794" pageId="25" pageNumber="26" unit="in" value="11.0">11 in</quantity>
|
||
each row. posterior transverse row with 15–16 teeth, implanted recumbently with a lateral inclination; major cusps weakly recurved, bluntly pointed, and laterally compressed; minor cusps mostly absent.
|
||
</paragraph>
|
||
<paragraph id="8B9936B0611EDA41FF6F51CB2711FB8B" blockId="25.[189,1398,1785,1921]" pageId="25" pageNumber="26">All scales relatively small, many scale rows between anal fin and upper lateral line (12–16), infraorbital scales on cheek (4–7), and scales around caudal peduncle (18–20); many scales in longitudinal line (34–38); chest scales small; transition to larger flank scales gradual. Minute scales on proximal half of caudal fin.</paragraph>
|
||
<paragraph id="8B9936B0611EDA43FF6F50982452FD4C" blockId="25.[189,1399,1962,2025]" lastBlockId="27.[189,1399,265,327]" lastPageId="27" lastPageNumber="28" pageId="25" pageNumber="26">Caudal fin emarginate; dorsal and anal fins reach to between verticals through two scales anterior to and one scale posterior to caudal-fin base. Pectoral fin reaches to between anal opening and three scales anterior to this point; pelvic fin reaches to anal opening in females, to first anal-fin spine in males; first branched pelvic-fin ray not elongated.</paragraph>
|
||
<caption id="DF596638611DDA42FF6F505624E5FBE2" ID-DOI="http://doi.org/10.5281/zenodo.6484191" ID-Zenodo-Dep="6484191" httpUri="https://zenodo.org/record/6484191/files/figure.png" pageId="26" pageNumber="27" startId="26.[189,232,1892,1918]" targetBox="[207,1383,278,1848]" targetPageId="26">
|
||
<paragraph id="8B9936B0611DDA42FF6F505624E5FBE2" blockId="26.[189,1399,1892,2025]" pageId="26" pageNumber="27">
|
||
<emphasis id="B952EAA2611DDA42FF6F50562737FB75" bold="true" box="[189,286,1892,1919]" pageId="26" pageNumber="27">Fig. 12.</emphasis>
|
||
<taxonomicName id="4C264D33611DDA42FEF85057242AFB75" authority="Vranken & Steenberge & Heylen & Decru & Snoeks, 2022" authorityName="Vranken & Steenberge & Heylen & Decru & Snoeks" authorityYear="2022" box="[298,515,1893,1919]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="rex" status="sp. nov.">
|
||
<emphasis id="B952EAA2611DDA42FEF85057242AFB75" box="[298,515,1893,1919]" italics="true" pageId="26" pageNumber="27">Haplochromis rex</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26157D9611DDA42FDDC50572444FB75" box="[526,621,1893,1919]" pageId="26" pageNumber="27" rank="species">sp. nov.</taxonomicNameLabel>
|
||
<emphasis id="B952EAA2611DDA42FDAA505724AEFB75" bold="true" box="[632,647,1893,1919]" pageId="26" pageNumber="27">a</emphasis>
|
||
. Photograph of preserved holotype (RMCA 2018.008.P.0345; 148.5 mm SL).
|
||
<emphasis id="B952EAA2611DDA42FEA750BA27ACFBA8" bold="true" box="[373,389,1928,1954]" pageId="26" pageNumber="27">b</emphasis>
|
||
. X-ray image of holotype
|
||
<emphasis id="B952EAA2611DDA42FD6E50BA24C1FBA8" bold="true" box="[700,744,1928,1954]" pageId="26" pageNumber="27">c–d</emphasis>
|
||
. Photographs of freshly caught specimens.
|
||
<emphasis id="B952EAA2611DDA42FB3450BA22DAFBA8" bold="true" box="[1254,1267,1928,1954]" pageId="26" pageNumber="27">c</emphasis>
|
||
. Holotype, a dominant male.
|
||
<emphasis id="B952EAA2611DDA42FE4850992783FBCF" bold="true" box="[410,426,1963,1989]" pageId="26" pageNumber="27">d</emphasis>
|
||
. A female (RMCA 2017.006.P.0355; 135.7 mm SL) to illustrate the live colour patterns. The contrast was slightly enhanced.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B9936B0611CDA43FF6F565C22B8FDA1" blockId="27.[189,1399,366,427]" pageId="27" pageNumber="28">Ceratobranchial gill rakers in outer row of first gill arch short, stout, and simple; posteriormost rakers anvil-shaped to weakly trifid. Epibranchial gill rakers relatively slender and simple.</paragraph>
|
||
<paragraph id="8B9936B0611CDA43FF6F56E0278AFDE6" blockId="27.[189,419,466,492]" box="[189,419,466,492]" pageId="27" pageNumber="28">
|
||
<heading id="D0D181DC611CDA43FF6F56E0278AFDE6" bold="true" box="[189,419,466,492]" fontSize="11" level="3" pageId="27" pageNumber="28" reason="3">
|
||
<emphasis id="B952EAA2611CDA43FF6F56E0278AFDE6" bold="true" box="[189,419,466,492]" pageId="27" pageNumber="28">Colouration in life</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8B9936B0611CDA43FF6F55332496FF18" blockId="27.[188,1401,513,786]" pageId="27" pageNumber="28">
|
||
Dominant males: body cream-coloured with faint orange sheen; dorsum greyish; belly and chest black; caudal peduncle with blue sheen (
|
||
<figureCitation id="131D2A35611CDA43FD915516248FFE34" box="[579,678,548,574]" captionStart="Fig" captionStartId="26.[189,232,1892,1918]" captionTargetBox="[207,1383,278,1848]" captionTargetId="figure-19@26.[207,1383,262,1848]" captionTargetPageId="26" captionText="Fig. 12. Haplochromis rex sp. nov. a. Photograph of preserved holotype (RMCA 2018.008.P.0345; 148.5 mm SL). b. X-ray image of holotype c–d. Photographs of freshly caught specimens. c. Holotype, a dominant male. d. A female (RMCA 2017.006.P.0355; 135.7 mm SL) to illustrate the live colour patterns. The contrast was slightly enhanced." figureDoi="http://doi.org/10.5281/zenodo.6484191" httpUri="https://zenodo.org/record/6484191/files/figure.png" pageId="27" pageNumber="28">Fig. 12c</figureCitation>
|
||
). Operculum, dorsal part of head, and interorbital area orange; snout, lower jaw, and lips light blue; cheek white and speckled black; eye with (dark) grey outer ring and silver inner ring. Nostril, interorbital, supraorbital, and lacrimal stripes faint; mental blotch present. Pectoral and dorsal fin hyaline; dorsal fin with a dusky base, a dusky and maculated posterior part, and orange lappets, except for black anteriormost lappets. Pelvic fin black; anal and caudal fins orange-red and with dusky bases; anal fin with dusky posterior part and 1–3 small orange egg-spots with dusky rings; caudal fin dorsally maculated orange.
|
||
</paragraph>
|
||
<paragraph id="8B9936B0611CDA44FF6F540B27B3FD4D" blockId="27.[188,1400,825,957]" lastBlockId="28.[189,1398,265,327]" lastPageId="28" lastPageNumber="29" pageId="27" pageNumber="28">
|
||
Females and juveniles: body cream-coloured with yellow sheen; dorsum and dorsal part of head greyish; belly, chest, operculum, cheek, and lower jaw white; snout dusky and faint light blue; eye with (dark) grey outer ring and silver inner ring (
|
||
<figureCitation id="131D2A35611CDA43FDAC544D24CEFF90" box="[638,743,895,922]" captionStart="Fig" captionStartId="26.[189,232,1892,1918]" captionTargetBox="[207,1383,278,1848]" captionTargetId="figure-19@26.[207,1383,262,1848]" captionTargetPageId="26" captionText="Fig. 12. Haplochromis rex sp. nov. a. Photograph of preserved holotype (RMCA 2018.008.P.0345; 148.5 mm SL). b. X-ray image of holotype c–d. Photographs of freshly caught specimens. c. Holotype, a dominant male. d. A female (RMCA 2017.006.P.0355; 135.7 mm SL) to illustrate the live colour patterns. The contrast was slightly enhanced." figureDoi="http://doi.org/10.5281/zenodo.6484191" httpUri="https://zenodo.org/record/6484191/files/figure.png" pageId="27" pageNumber="28">Fig. 12d</figureCitation>
|
||
). Nostril, interorbital, supraorbital, and lacrimal stripes slender and well-defined; mental blotch present. Pectoral, pelvic, anal, and caudal fins yellowish; anal fin with 2 spots resembling egg-spots; caudal fin with dusky base and dorsal part; dorsal fin dusky and with black lappets.
|
||
</paragraph>
|
||
<caption id="DF596638611CDA43FF6F509924D4FBE2" ID-DOI="http://doi.org/10.5281/zenodo.6484195" ID-Zenodo-Dep="6484195" httpUri="https://zenodo.org/record/6484195/files/figure.png" pageId="27" pageNumber="28" startId="27.[189,232,1963,1989]" targetBox="[190,1395,1029,1890]" targetPageId="27">
|
||
<paragraph id="8B9936B0611CDA43FF6F509924D4FBE2" blockId="27.[186,1398,1963,2025]" pageId="27" pageNumber="28">
|
||
<emphasis id="B952EAA2611CDA43FF6F50992731FBCF" bold="true" box="[189,280,1963,1989]" pageId="27" pageNumber="28">Fig. 13.</emphasis>
|
||
<taxonomicName id="4C264D33611CDA43FECE509927C6FBCF" authority="Vranken & Steenberge & Heylen & Decru & Snoeks, 2022" authorityName="Vranken & Steenberge & Heylen & Decru & Snoeks" authorityYear="2022" box="[284,495,1963,1989]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="27" pageNumber="28" phylum="Chordata" rank="species" species="rex" status="sp. nov.">
|
||
<emphasis id="B952EAA2611CDA43FECE509927C6FBCF" box="[284,495,1963,1989]" italics="true" pageId="27" pageNumber="28">Haplochromis rex</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26157D9611CDA43FE23509E2460FBCC" box="[497,585,1964,1990]" pageId="27" pageNumber="28" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(IRSNB 13480; 154.2 mm SL).
|
||
<emphasis id="B952EAA2611CDA43FC6B509925E1FBCF" bold="true" box="[953,968,1963,1989]" pageId="27" pageNumber="28">a</emphasis>
|
||
. Dorsal view of the lower pharyngeal jaw.
|
||
<emphasis id="B952EAA2611CDA43FF2650FC272DFBE2" bold="true" box="[244,260,1998,2024]" pageId="27" pageNumber="28">b</emphasis>
|
||
. Lateral view of the lower pharyngeal jaw.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B9936B0611BDA44FF6F564227F9FD80" blockId="28.[189,464,368,394]" box="[189,464,368,394]" pageId="28" pageNumber="29">
|
||
<heading id="D0D181DC611BDA44FF6F564227F9FD80" bold="true" box="[189,464,368,394]" fontSize="11" level="3" pageId="28" pageNumber="29" reason="3">
|
||
<emphasis id="B952EAA2611BDA44FF6F564227F9FD80" bold="true" box="[189,464,368,394]" pageId="28" pageNumber="29">Preserved colouration</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8B9936B0611BDA44FF6F56922593FF0E" blockId="28.[187,1400,416,772]" pageId="28" pageNumber="29">
|
||
Body, belly, operculum, and cheek yellowish; dorsum brown; chest yellowish in females, black in males (
|
||
<figureCitation id="131D2A35611BDA44FF1756F72705FDD5" box="[197,300,453,479]" captionStart="Fig" captionStartId="26.[189,232,1892,1918]" captionTargetBox="[207,1383,278,1848]" captionTargetId="figure-19@26.[207,1383,262,1848]" captionTargetPageId="26" captionText="Fig. 12. Haplochromis rex sp. nov. a. Photograph of preserved holotype (RMCA 2018.008.P.0345; 148.5 mm SL). b. X-ray image of holotype c–d. Photographs of freshly caught specimens. c. Holotype, a dominant male. d. A female (RMCA 2017.006.P.0355; 135.7 mm SL) to illustrate the live colour patterns. The contrast was slightly enhanced." figureDoi="http://doi.org/10.5281/zenodo.6484191" httpUri="https://zenodo.org/record/6484191/files/figure.png" pageId="28" pageNumber="29">Fig. 12a</figureCitation>
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). Flank rarely with 7–8 very faint narrow vertical stripes. Snout dusky; in males, lower jaw black. In females, all head stripes faint and lacrimal stripe reduced to blotch below eye; in males, head stripes as described here. Nostril and interorbital stripes narrow, well-defined, and horizontally inclined in lateral view, while positioned anterior of nostrils and orbits, respectively, in dorsal view; lacrimal stripe distinct, broad (may cover whole lacrimal), and vaguely delineated; vertical preopercular stripe welldefined; supraorbital stripe present; nape band and mental blotch faint. Pectoral and anal fins yellowish; anal fin with dusky base and posterior margin and 2 small egg-spots in males; pelvic fin yellowish with a black first ray in females, black in males. Dorsal and caudal fins dusky and with yellowish posterior parts; dorsal fin with black lappets and maculated posterior parts.
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</paragraph>
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</subSubSection>
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<subSubSection id="C33C653B611BDA44FF6F541E2794FF90" pageId="28" pageNumber="29" type="distribution">
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<paragraph id="8B9936B0611BDA44FF6F541E27D8FF4C" blockId="28.[189,497,812,838]" box="[189,497,812,838]" pageId="28" pageNumber="29">
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<heading id="D0D181DC611BDA44FF6F541E27D8FF4C" bold="true" box="[189,497,812,838]" fontSize="11" level="3" pageId="28" pageNumber="29" reason="3">
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<emphasis id="B952EAA2611BDA44FF6F541E27D8FF4C" bold="true" box="[189,497,812,838]" pageId="28" pageNumber="29">Distribution and ecology</emphasis>
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</heading>
|
||
</paragraph>
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||
<paragraph id="8B9936B0611BDA44FF6F546E2794FF90" blockId="28.[189,1398,860,923]" pageId="28" pageNumber="29">Only known from Lake Edward, found over sandy substrates. Based on its morphology, most probably a piscivorous species.</paragraph>
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</subSubSection>
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</treatment>
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</document> |