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<mods:title id="CC2257A525E9387489A9CF460624ED9B">Observations on the Biology of Afrotropical Hesperiidae (Lepidoptera). Part 5. Hesperiinae incertae sedis: Dicotyledon Feeders</mods:title>
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<mods:namePart id="2B0E1138D440AF4DA9E54EFFB867BF95">Cock, Matthew J. W.</mods:namePart>
<mods:affiliation id="92571A3FFB6E840EA91A90E84DC25DF0">C / o CABI Europe - UK, Bakeham Lane, Egham, TW 20 9 TY, UK (e-mail: m. cock @ cabi. org; mjwcock @ btinternet. com)</mods:affiliation>
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<subSubSection id="C3367BD7FFBAFFE49BFDFABDFE64BF06" box="[151,433,1324,1350]" pageId="4" pageNumber="5" type="nomenclature">
<paragraph id="8B93285CFFBAFFE49BFDFABDFE64BF06" blockId="4.[151,433,1324,1350]" box="[151,433,1324,1350]" pageId="4" pageNumber="5">
<heading id="D0DB9F30FFBAFFE49BFDFABDFE64BF06" bold="true" box="[151,433,1324,1350]" fontSize="11" level="2" pageId="4" pageNumber="5" reason="4">
<taxonomicName id="4C2C53DFFFBAFFE49BFDFABDFE64BF06" ID-CoL="929VK" authority="Holland, 1896" authorityName="Holland" authorityYear="1896" baseAuthorityName="Lindsey &amp; Miller" baseAuthorityYear="1965" box="[151,433,1324,1350]" class="Insecta" family="Hesperiidae" genus="Gorgyra" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B958F44EFFBAFFE49BFDFABDFE64BF06" bold="true" box="[151,433,1324,1350]" pageId="4" pageNumber="5">
<emphasis id="B958F44EFFBAFFE49BFDFABDFF29BF06" bold="true" box="[151,252,1324,1350]" italics="true" pageId="4" pageNumber="5">Gorgyra</emphasis>
<bibRefCitation id="EFBD55ADFFBAFFE49A69FABDFE64BF06" author="Holland" box="[259,433,1324,1350]" firstAuthor="Holland" pageId="4" pageNumber="5" pagination="2 - 107" refId="ref43303" refString="Holland, W. J. (1896) A preliminary revision and synonymic catalogue of the Hesperiidae of Africa and the adjacent islands, with descriptions of some apparently new species. Proceedings of the Zoological Society of London, 1896, 2 - 107, plates I - V. http: // dx. doi. org / 10.1111 / j. 1096 - 3642.1896. tb 03030. x" type="journal article" year="1896">
<collectingCountry id="F33B68CCFFBAFFE49A69FABDFEBFBF06" box="[259,362,1324,1350]" name="Netherlands" pageId="4" pageNumber="5">Holland</collectingCountry>
, 1896
</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C3367BD7FFBAFFE49BFDFAE3FB4BBFB7" pageId="4" pageNumber="5" type="discussion">
<paragraph id="8B93285CFFBAFFE49BFDFAE3FB4BBFB7" blockId="4.[151,1436,1394,2031]" pageId="4" pageNumber="5">
This is a genus of 20 or more species (
<bibRefCitation id="EFBD55ADFFBAFFE49930FAE3FD3EBFCB" author="Larsen" box="[602,747,1394,1419]" firstAuthor="Larsen" pageId="4" pageNumber="5" pagination="55 - 58" refId="ref44005" refString="Larsen, T. B. (2010) Gorgyra stewarti sp. nov. - a surprising new skipper butterfly from Cameroun and the democratic Republic of Congo (Lep.: Hesperiidae). Entomologist's Record and Journal of Variation, 122, 55 - 58." type="journal article" year="2010">Larsen 2010</bibRefCitation>
) restricted to Africa. These are secretive butterflies, seldom seen. They tend to be found in localised colonies, which may be more easily located as caterpillars. The known food plants of
<emphasis id="B958F44EFFBAFFE49A57FA2AFE44BF92" box="[317,401,1467,1490]" italics="true" pageId="4" pageNumber="5">Gorgya</emphasis>
spp. are all dicotyledons; most records are from species of
<taxonomicName id="4C2C53DFFFBAFFE49F52FA2BFB19BF93" box="[1080,1228,1466,1491]" class="Magnoliopsida" family="Connaraceae" kingdom="Plantae" order="Oxalidales" pageId="4" pageNumber="5" phylum="Tracheophyta" rank="family">Connaraceae</taxonomicName>
, but there are also records from
<taxonomicName id="4C2C53DFFFBAFFE49A44FA4FFE30BFB7" box="[302,485,1502,1527]" class="Magnoliopsida" family="Dichapetalaceae" kingdom="Plantae" order="Malpighiales" pageId="4" pageNumber="5" phylum="Tracheophyta" rank="family">Dichapetalaceae</taxonomicName>
and unconfirmed records from
<taxonomicName id="4C2C53DFFFBAFFE49822FA4FFC38BFB7" box="[840,1005,1502,1527]" class="Magnoliopsida" family="Putranjivaceae" kingdom="Plantae" order="Malpighiales" pageId="4" pageNumber="5" phylum="Tracheophyta" rank="family">Putranjivaceae</taxonomicName>
and
<taxonomicName id="4C2C53DFFFBAFFE49F49FA4FFB4FBFB7" box="[1059,1178,1502,1527]" class="Magnoliopsida" family="Meliaceae" kingdom="Plantae" order="Sapindales" pageId="4" pageNumber="5" phylum="Tracheophyta" rank="family">Meliaceae</taxonomicName>
.
</paragraph>
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<subSubSection id="C3367BD7FFBAFFE59BADF992FACAB858" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5" type="description">
<paragraph id="8B93285CFFBAFFE49BADF992FC56BD57" blockId="4.[151,1436,1394,2031]" pageId="4" pageNumber="5">
We present information on four species that we have reared. In addition to these, at least four other species have been reared. Sevastopulo (unpublished) reared
<taxonomicName id="4C2C53DFFFBAFFE49999F9B6FC40BC7E" authority="Evans" authorityName="Evans" authorityYear="1937" box="[755,917,1574,1599]" class="Insecta" family="Hesperiidae" genus="Gorgyra" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="bina">
<emphasis id="B958F44EFFBAFFE49999F9B6FC92BC7F" box="[755,839,1574,1599]" italics="true" pageId="4" pageNumber="5">G. bina</emphasis>
Evans
</taxonomicName>
from Zika Forest, Entebbe,
<collectingCountry id="F33B68CCFFBAFFE49F8AF9B7FAE2BC7F" box="[1248,1335,1574,1599]" name="Uganda" pageId="4" pageNumber="5">Uganda</collectingCountry>
from an unidentified food plant which in his photographs can be seen to have quite large (c.
<quantity id="4CD485B9FFBAFFE49F2DF9DBFB5BBC23" box="[1095,1166,1610,1635]" metricMagnitude="-2" metricUnit="m" metricValue="6.0" pageId="4" pageNumber="5" unit="mm" value="60.0">60mm</quantity>
long) shiny leaves. The folded leaf shelter resembles that of the mature caterpillar of
<taxonomicName id="4C2C53DFFFBAFFE49857F9FEFC25BCC7" authority="Riley" authorityName="Riley" authorityYear="1929" box="[829,1008,1646,1671]" class="Insecta" family="Hesperiidae" genus="Gorgyra" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="bibulus">
<emphasis id="B958F44EFFBAFFE49857F9FEFC7BBCC7" box="[829,942,1646,1671]" italics="true" pageId="4" pageNumber="5">G. bibulus</emphasis>
Riley
</taxonomicName>
(
<figureCitation id="131734D9FFBAFFE49895F9FFFBB5BCC7" box="[1023,1120,1646,1671]" captionStart="FIGURE 6" captionStartId="9.[151,249,817,839]" captionTargetBox="[159,1428,207,796]" captionTargetId="figure-246@9.[151,1436,195,796]" captionTargetPageId="9" captionText="FIGURE 6. Leaf shelters of Gorgyra bibulus on Rourea thomsonii, collected 4 Jul 1998, Gatamayu Forest, Kenya; the lines in the background are 8mm apart. 1, stage 1 shelter on left, 98/201E; stage 2 shelter on right, 98/201F; 2, stage 3 shelter, 98/201C; 3, pupal shelter, 98/201A." figureDoi="http://doi.org/10.5281/zenodo.5267847" httpUri="https://zenodo.org/record/5267847/files/figure.png" pageId="4" pageNumber="5">Figure 6</figureCitation>
). The form of the caterpillar is similar to other
<taxonomicName id="4C2C53DFFFBAFFE49A09F902FE2DBCEA" box="[355,504,1683,1706]" class="Insecta" family="Hesperiidae" genus="Gorgyra" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B958F44EFFBAFFE49A09F902FE17BCEA" box="[355,450,1683,1706]" italics="true" pageId="4" pageNumber="5">Gorgyra</emphasis>
spp.
</taxonomicName>
treated here: head black; body grey-green, inter-segmental folds more yellow; legs brown; anal flap large, rounded, tinged with olive. The pupa is formed in a slight cocoon in a folded leaf, lined with white waxy powder: reddish brown, abdomen tinged olive; covered with white waxy powder, which adheres to the setae of head and thorax; short, deeply bifurcate frontal projection.
</paragraph>
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<bibRefCitation id="EFBD55ADFFBAFFE49BADF8B3FE4CBD7B" author="Sevastopulo" box="[199,409,1826,1851]" firstAuthor="Sevastopulo" pageId="4" pageNumber="5" pagination="131 - 136" refId="ref45528" refString="Sevastopulo, D. G. (1974) Butterflies of the Shimba Hills. Entomologist's Record and Journal of Variation, 86, 131 - 136." type="journal article" year="1974">Sevastopulo (1974</bibRefCitation>
,
<bibRefCitation id="EFBD55ADFFBAFFE49ACFF8B3FE0ABD7B" author="Sevastopulo, D. G." bookContentInfo="34, 84 - 92, 124 - 132, 175 - 184; 35, 94 - 100, 177 - 195; 36, 45 - 50; 37, 147 - 172, 202 - 216." box="[421,479,1826,1851]" editor="Sevastopulo, D. G." journalOrPublisher="Bulletin of the Amateur Entomologists' Society" pageId="4" pageNumber="5" pagination="131 - 132" part="34" refId="ref45556" refString="Sevastopulo, D. G. (1975) Hesperiidae. Bulletin of the Amateur Entomologists' Society, 34, 131 - 132. In: Sevastopulo, D. G. (1975 - 1978) A list of the food-plants of East African Macrolepidoptera. Bulletin of the Amateur Entomologists' Society, 34, 84 - 92, 124 - 132, 175 - 184; 35, 94 - 100, 177 - 195; 36, 45 - 50; 37, 147 - 172, 202 - 216." title="Hesperiidae" type="book chapter" volumeTitle="A list of the food-plants of East African Macrolepidoptera. Bulletin of the Amateur Entomologists' Society" year="1975">1975</bibRefCitation>
, unpublished) reared
<taxonomicName id="4C2C53DFFFBAFFE499A4F8B2FCBDBD7A" authority="Evans" authorityName="Evans" authorityYear="1937" box="[718,872,1826,1851]" class="Insecta" family="Hesperiidae" genus="Gorgyra" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="diva">
<emphasis id="B958F44EFFBAFFE499A4F8B2FCC8BD7B" box="[718,797,1826,1851]" italics="true" pageId="4" pageNumber="5">G. diva</emphasis>
Evans
</taxonomicName>
from
<taxonomicName id="4C2C53DFFFBAFFE498C6F8B2FBBBBD7B" box="[940,1134,1826,1851]" class="Magnoliopsida" family="Connaraceae" genus="Rourea" kingdom="Plantae" order="Oxalidales" pageId="4" pageNumber="5" phylum="Tracheophyta" rank="species" species="orientalis">
<emphasis id="B958F44EFFBAFFE498C6F8B2FBBBBD7B" box="[940,1134,1826,1851]" italics="true" pageId="4" pageNumber="5">Rourea orientalis</emphasis>
</taxonomicName>
(=
<taxonomicName id="4C2C53DFFFBAFFE49FFEF8B2FA46BD7B" box="[1172,1427,1826,1851]" class="Magnoliopsida" family="Connaraceae" genus="Byrsocarpus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Oxalidales" pageId="4" pageNumber="5" phylum="Tracheophyta" rank="species" species="orientalis">
<emphasis id="B958F44EFFBAFFE49FFEF8B2FA46BD7B" box="[1172,1427,1826,1851]" italics="true" pageId="4" pageNumber="5">Byrsocarpus orientalis</emphasis>
</taxonomicName>
) (
<taxonomicName id="4C2C53DFFFBAFFE49BF4F8D7FEE1BD1F" box="[158,308,1862,1887]" class="Magnoliopsida" family="Connaraceae" kingdom="Plantae" order="Oxalidales" pageId="4" pageNumber="5" phylum="Tracheophyta" rank="family">Connaraceae</taxonomicName>
), in the Shimba Hills. Armed with this information, MJWC examined herbarium specimens in the National Herbarium,
<collectingCountry id="F33B68CCFFBAFFE49AE9F8FAFE1ABDC3" box="[387,463,1899,1923]" name="Kenya" pageId="4" pageNumber="5">Kenya</collectingCountry>
, and so tracked down the food plant (
<figureCitation id="131734D9FFBAFFE49803F8FBFC12BDC3" box="[873,967,1898,1923]" captionStart="FIGURE 1" captionStartId="5.[151,249,1446,1468]" captionTargetBox="[151,1436,581,1425]" captionTargetId="figure-268@5.[151,1436,581,1425]" captionTargetPageId="5" captionText="FIGURE 1. Rourea orientalis, a food plant of Gorgyra diva and G. johnstoni, Shimba Hills, 13 Apr 1990." figureDoi="http://doi.org/10.5281/zenodo.5267835" httpUri="https://zenodo.org/record/5267835/files/figure.png" pageId="4" pageNumber="5">Figure 1</figureCitation>
), but the caterpillars that he reared proved to be
<taxonomicName id="4C2C53DFFFBAFFE49BBEF81EFE6DBDE7" authority="(Butler)" baseAuthorityName="Butler" box="[212,440,1934,1959]" class="Insecta" family="Hesperiidae" genus="Gorgyra" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="johnstoni">
<emphasis id="B958F44EFFBAFFE49BBEF81EFE8CBDE7" box="[212,345,1934,1959]" italics="true" pageId="4" pageNumber="5">G. johnstoni</emphasis>
(Butler)
</taxonomicName>
(below). Sevastopulos caterpillar had the head reddish mahogany; body yellowish green, minutely speckled white, intersegmental areas yellowish; anal flap rounded, brownish; all legs green. Pupa in a silk lined cell in a folded leaf lined with white waxy powder; pale greenish amber, the head and thorax tinged chestnut; head with two projecting tubercles between the eyes; thickly coated with white waxy powder which adheres to the setae of the thorax and head. Sevastopulos food plant record has been repeated by subsequent authors:
<bibRefCitation id="EFBD55ADFFBBFFE59E56FF2DFF36BAB8" author="Kielland" firstAuthor="Kielland" pageId="5" pageNumber="6" refId="ref43661" refString="Kielland, J. (1990) Butterflies of Tanzania. Hill House, Melbourne, Australia, 363 pp." type="book" year="1990">Kielland (1990)</bibRefCitation>
,
<bibRefCitation id="EFBD55ADFFBBFFE59B9AFF71FE47BAB8" author="Larsen" box="[240,402,223,248]" firstAuthor="Larsen" pageId="5" pageNumber="6" refId="ref43862" refString="Larsen, T. B. (1991) The Butterflies of Kenya and their Natural History. Oxford University Press, Oxford, UK, 490 pp." type="book" year="1991">Larsen (1991)</bibRefCitation>
,
<bibRefCitation id="EFBD55ADFFBBFFE59AF4FF4EFD52BAB8" author="Ackery" box="[414,647,223,248]" etAl="et al." firstAuthor="Ackery" pageId="5" pageNumber="6" refId="ref41160" refString="Ackery, P. R., Smith, C. R. &amp; Vane-Wright, R. I. (1995) Carcasson's African Butterflies. An annotated catalogue of the Papilionoidea and Hesperioidea of the Afrotropical Region. CSIRO Publications, Melbourne, Australia, 803 pp." type="book" year="1995">
Ackery
<emphasis id="B958F44EFFBBFFE59A93FF70FDFEBAB8" box="[505,555,223,248]" italics="true" pageId="5" pageNumber="6">et al</emphasis>
. (1995)
</bibRefCitation>
, and
<bibRefCitation id="EFBD55ADFFBBFFE599AFFF4EFC75BAB8" author="Heath" box="[709,928,223,248]" etAl="et al." firstAuthor="Heath" pageId="5" pageNumber="6" refId="ref42672" refString="Heath, A., Newport, M. A. &amp; Hancock, D. (2002) The Butterflies of Zambia. African Butterfly Research Institute and The Lepidopterists' Society of Africa, Nairobi, Kenya, 137 pp. + CD." type="book" year="2002">
Heath
<emphasis id="B958F44EFFBBFFE5987AFF70FC96BAB8" box="[784,835,223,248]" italics="true" pageId="5" pageNumber="6">et al</emphasis>
. (2002)
</bibRefCitation>
.
<bibRefCitation id="EFBD55ADFFBBFFE598C6FF71FB9FBAB8" author="Larsen" box="[940,1098,223,248]" firstAuthor="Larsen" pageId="5" pageNumber="6" refId="ref43862" refString="Larsen, T. B. (1991) The Butterflies of Kenya and their Natural History. Oxford University Press, Oxford, UK, 490 pp." type="book" year="1991">Larsen (1991)</bibRefCitation>
states that the caterpillars are frequently parasitized (based on MJWCs observations); at that time, MJWC thought he had reared
<taxonomicName id="4C2C53DFFFBBFFE59E6BFE95FA84BB5D" authorityName="Evans" authorityYear="1937" box="[1281,1361,260,285]" class="Insecta" family="Hesperiidae" genus="Gorgyra" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="diva">
<emphasis id="B958F44EFFBBFFE59E6BFE95FA84BB5D" box="[1281,1361,260,285]" italics="true" pageId="5" pageNumber="6">G. diva</emphasis>
</taxonomicName>
, but it was
<taxonomicName id="4C2C53DFFFBBFFE59BA3FEB9FE9BBB00" box="[201,334,295,320]" class="Insecta" family="Hesperiidae" genus="Gorgyra" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="johnstoni">
<emphasis id="B958F44EFFBBFFE59BA3FEB9FE9BBB00" box="[201,334,295,320]" italics="true" pageId="5" pageNumber="6">G. johnstoni</emphasis>
</taxonomicName>
, so these observations should be associated with the latter species (below).
</paragraph>
<paragraph id="8B93285CFFBBFFE59BADFEDDFACAB858" blockId="5.[151,1436,151,536]" pageId="5" pageNumber="6">
<bibRefCitation id="EFBD55ADFFBBFFE59BADFEDDFEAABB25" author="Vuattoux" box="[199,383,332,357]" firstAuthor="Vuattoux" pageId="5" pageNumber="6" pagination="562 - 568" refId="ref46285" refString="Vuattoux, R. (1999) Les lepidopteres hesperiides de la Station de Lamto (Cote d'Ivoire). Lambillionea, 94, 562 - 568." type="journal article" year="1999">Vuattoux (1999)</bibRefCitation>
reared
<taxonomicName id="4C2C53DFFFBBFFE59AB9FEDDFD56BB23" authority="Druce" authorityName="Druce" authorityYear="1909" box="[467,643,332,357]" class="Insecta" family="Hesperiidae" genus="Gorgyra" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="afikpo">
<emphasis id="B958F44EFFBBFFE59AB9FEDDFDEDBB25" box="[467,568,332,357]" italics="true" pageId="5" pageNumber="6">G. afikpo</emphasis>
Druce
</taxonomicName>
from three different food plants in
<collectingCountry id="F33B68CCFFBBFFE59F64FEDDFB7DBB25" box="[1038,1192,332,357]" name="Ivory Coast" pageId="5" pageNumber="6">Côte dIvoire</collectingCountry>
:
<taxonomicName id="4C2C53DFFFBBFFE59FD9FEDDFEE8BBC8" class="Magnoliopsida" family="Connaraceae" genus="Connarus" kingdom="Plantae" order="Oxalidales" pageId="5" pageNumber="6" phylum="Tracheophyta" rank="species" species="thonningii">
<emphasis id="B958F44EFFBBFFE59FD9FEDDFA49BB25" box="[1203,1436,332,357]" italics="true" pageId="5" pageNumber="6">Connarus thonningii</emphasis>
(Connaraceae)
</taxonomicName>
,
<taxonomicName id="4C2C53DFFFBBFFE59A27FEFEFD68BBC8" box="[333,701,367,392]" class="Magnoliopsida" family="Dichapetalaceae" genus="Dichapetalum" kingdom="Plantae" order="Malpighiales" pageId="5" pageNumber="6" phylum="Tracheophyta" rank="species" species="madagascariense">
<emphasis id="B958F44EFFBBFFE59A27FEFEFD68BBC8" box="[333,701,367,392]" italics="true" pageId="5" pageNumber="6">Dichapetalum madagascariense</emphasis>
</taxonomicName>
(=
<taxonomicName id="4C2C53DFFFBBFFE59988FEE0FCAFBBC7" box="[738,890,368,392]" class="Magnoliopsida" family="Dichapetalaceae" genus="Dichapetalum" kingdom="Plantae" order="Malpighiales" pageId="5" pageNumber="6" phylum="Tracheophyta" rank="species" species="guineense">
<emphasis id="B958F44EFFBBFFE59988FEE0FCAFBBC7" box="[738,890,368,392]" italics="true" pageId="5" pageNumber="6">D. guineense</emphasis>
</taxonomicName>
) (
<taxonomicName id="4C2C53DFFFBBFFE598FDFEFEFB80BBC8" box="[919,1109,367,392]" class="Magnoliopsida" family="Dichapetalaceae" kingdom="Plantae" order="Malpighiales" pageId="5" pageNumber="6" phylum="Tracheophyta" rank="family">Dichapetalaceae</taxonomicName>
), and an unidentified plant.
<bibRefCitation id="EFBD55ADFFBBFFE59BFDFE05FE81BBED" author="Vuattoux" box="[151,340,404,429]" firstAuthor="Vuattoux" pageId="5" pageNumber="6" pagination="562 - 568" refId="ref46285" refString="Vuattoux, R. (1999) Les lepidopteres hesperiides de la Station de Lamto (Cote d'Ivoire). Lambillionea, 94, 562 - 568." type="journal article" year="1999">Vuattoux (1999)</bibRefCitation>
also reared
<taxonomicName id="4C2C53DFFFBBFFE59A81FE05FD45BBEB" authority="Evans" authorityName="Evans" authorityYear="1937" box="[491,656,404,428]" class="Insecta" family="Hesperiidae" genus="Gorgyra" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="sara">
<emphasis id="B958F44EFFBBFFE59A81FE05FD95BBEC" box="[491,576,404,428]" italics="true" pageId="5" pageNumber="6">G. sara</emphasis>
Evans
</taxonomicName>
from
<taxonomicName id="4C2C53DFFFBBFFE599B5FE05FBB7BBED" box="[735,1122,404,429]" class="Magnoliopsida" family="Connaraceae" genus="Cnestis" kingdom="Plantae" order="Oxalidales" pageId="5" pageNumber="6" phylum="Tracheophyta" rank="species" species="ferruginea">
<emphasis id="B958F44EFFBBFFE599B5FE05FC66BBED" box="[735,947,404,429]" italics="true" pageId="5" pageNumber="6">Cnestis ferruginea</emphasis>
(Connaraceae)
</taxonomicName>
and an unidentified plant. MJWC found a
<taxonomicName id="4C2C53DFFFBBFFE59A26FE29FE06BB8F" box="[332,467,440,463]" class="Insecta" family="Hesperiidae" genus="Gorgyra" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B958F44EFFBBFFE59A26FE29FE7EBB8F" box="[332,427,440,463]" italics="true" pageId="5" pageNumber="6">Gorgyra</emphasis>
sp.
</taxonomicName>
caterpillar (94/105) on
<taxonomicName id="4C2C53DFFFBBFFE599B4FE29FCA0BB90" box="[734,885,439,464]" class="Magnoliopsida" family="Connaraceae" genus="Cnestis" kingdom="Plantae" order="Oxalidales" pageId="5" pageNumber="6" phylum="Tracheophyta" rank="species" species="ferruginea">
<emphasis id="B958F44EFFBBFFE599B4FE29FCA0BB90" box="[734,885,439,464]" italics="true" pageId="5" pageNumber="6">C. ferruginea</emphasis>
</taxonomicName>
(
<figureCitation id="131734D9FFBBFFE598EEFE26FC2EBB90" box="[900,1019,439,464]" captionStart="FIGURE 2" captionStartId="5.[151,249,1978,2000]" captionTargetBox="[151,1436,1518,1956]" captionTargetId="figure-272@5.[151,1436,1518,1956]" captionTargetPageId="5" captionText="FIGURE 2. Cnestis ferruginea, a food plant of Gorgyra sara, Ibadan, Nigeria, 17 Mar 1994. 1, fruiting plant; 2, stages 1 and 2 leaf shelters of a Gorgyra sp., perhaps G.sara." figureDoi="http://doi.org/10.5281/zenodo.5267837" httpUri="https://zenodo.org/record/5267837/files/figure.png" pageId="5" pageNumber="6">Figure 2.1</figureCitation>
) at Ibadan,
<collectingCountry id="F33B68CCFFBBFFE59FEBFE26FB0DBB90" box="[1153,1240,439,464]" name="Nigeria" pageId="5" pageNumber="6">Nigeria</collectingCountry>
, which may have been
<taxonomicName id="4C2C53DFFFBBFFE59BB8FE4DFEF6BBB4" authorityName="Evans" authorityYear="1937" box="[210,291,476,500]" class="Insecta" family="Hesperiidae" genus="Gorgyra" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="sara">
<emphasis id="B958F44EFFBBFFE59BB8FE4DFEF6BBB4" box="[210,291,476,500]" italics="true" pageId="5" pageNumber="6">G. sara</emphasis>
</taxonomicName>
, but it died as a crippled pupa when the final instar head capsule did not split and detach properly. The leaf shelters (
<figureCitation id="131734D9FFBBFFE59A46FE6EFE77B858" box="[300,418,511,536]" captionStart="FIGURE 2" captionStartId="5.[151,249,1978,2000]" captionTargetBox="[151,1436,1518,1956]" captionTargetId="figure-272@5.[151,1436,1518,1956]" captionTargetPageId="5" captionText="FIGURE 2. Cnestis ferruginea, a food plant of Gorgyra sara, Ibadan, Nigeria, 17 Mar 1994. 1, fruiting plant; 2, stages 1 and 2 leaf shelters of a Gorgyra sp., perhaps G.sara." figureDoi="http://doi.org/10.5281/zenodo.5267837" httpUri="https://zenodo.org/record/5267837/files/figure.png" pageId="5" pageNumber="6">Figure 2.2</figureCitation>
) and caterpillar (
<figureCitation id="131734D9FFBBFFE5990AFE6EFD15B858" box="[608,704,511,536]" captionStart="FIGURE 3" captionStartId="6.[151,249,1145,1167]" captionTargetBox="[151,1436,187,1124]" captionTargetId="figure-322@6.[151,1436,187,1124]" captionTargetPageId="6" captionText="FIGURE 3. Final instar caterpillar of a Gorgyra sp., perhaps G.sara, collected on Cnestis ferruginea; Ibadan, Nigeria, 17 Mar 1994; died; ref. 94/105. 1, dorsolateral view; 2, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5267841" httpUri="https://zenodo.org/record/5267841/files/figure.png" pageId="5" pageNumber="6">Figure 3</figureCitation>
) were similar to those of the
<taxonomicName id="4C2C53DFFFBBFFE59F63FD91FBAEB858" authorityName="Riley" authorityYear="1929" box="[1033,1147,511,536]" class="Insecta" family="Hesperiidae" genus="Gorgyra" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="bibulus">
<emphasis id="B958F44EFFBBFFE59F63FD91FBAEB858" box="[1033,1147,511,536]" italics="true" pageId="5" pageNumber="6">G. bibulus</emphasis>
</taxonomicName>
treated below.
</paragraph>
</subSubSection>
</treatment>
</document>