treatments-xml/data/03/85/87/038587A1FF80FF97FF60FF54F98DF8BB.xml
2024-06-21 12:22:17 +02:00

530 lines
74 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="5A5313251CB8C2B547124F0EDAD76147" ID-CLB-Dataset="45884" ID-DOI="10.11646/zootaxa.3793.5.4" ID-GBIF-Dataset="5f8cac29-bb09-4a52-b618-503b5fc8c954" ID-ISSN="1175-5326" ID-Zenodo-Dep="252346" ID-ZooBank="F9A1E47F-653E-48AD-8FDF-D7D0D59A0A3A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460414883683" checkinUser="plazi" docAuthor="Hämäläinen, Matti" docDate="2014" docId="038587A1FF80FF97FF60FF54F98DF8BB" docLanguage="en" docName="zt03793p572.pdf" docOrigin="Zootaxa 3793 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Atrocalopteryx auco Hämäläinen, 2014, spec. nov." docType="treatment" docVersion="9" lastPageNumber="566" masterDocId="FFBCFFD9FF81FF92FFF7FFC3FC01FFF6" masterDocTitle="Atrocalopteryx auco spec. nov. from Vietnam, with taxonomic notes on its congeners (Odonata: Calopterygidae)" masterLastPageNumber="572" masterPageNumber="561" pageNumber="562" updateTime="1698328446632" updateUser="ExternalLinkService">
<mods:mods id="C55CF26FC86EFFD49A4C1DDC6C08942C" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="88848AEB5744C89D4083F1DE4E47681D">
<mods:title id="49FF556B98976E81170EE990488B5AA5">Atrocalopteryx auco spec. nov. from Vietnam, with taxonomic notes on its congeners (Odonata: Calopterygidae)</mods:title>
</mods:titleInfo>
<mods:name id="42E2BB924C8B16D288888274E19BF6F1" type="personal">
<mods:role id="D1E7624B5F20E99C6F8817895AE7AE1E">
<mods:roleTerm id="07622A2A4548402BD96AEFC23EE3288F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="757246DDA7679AAE13C5DAA0CE3F37E8">Hämäläinen, Matti</mods:namePart>
</mods:name>
<mods:typeOfResource id="32842AD405F69D777A3A9721D9500E15">text</mods:typeOfResource>
<mods:relatedItem id="E586D478302F4066B791D5733226AC37" type="host">
<mods:titleInfo id="8465A269B7E84D77D26C80212B27526D">
<mods:title id="26AE34B3090945746AFC1DAC853017BC">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="EF6908F46E4DC455047C87B8C787B79B">
<mods:date id="9A936FD6182C88CA126AE9533B2F10E7">2014</mods:date>
<mods:detail id="ABBE32672E2754F2C6B1F1F3AA5FC834" type="volume">
<mods:number id="94C7BE1A0DCBF8434B436F25942AFDD0">3793</mods:number>
</mods:detail>
<mods:detail id="6CAEF6991E31F51F26E90B2BBBA10DF0" type="issue">
<mods:number id="F7E7DCA109CE5EA80A26C2E3D900DFC1">5</mods:number>
</mods:detail>
<mods:extent id="D4CD1189C3C0BDB82686C1C1BAA15E6F" unit="page">
<mods:start id="9805DF41791206227FBDB6B752CC555F">561</mods:start>
<mods:end id="85EFD4DE718FE7A15B60615806AE0F11">572</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="5F793801EF13100911210A8C11F0C482">journal article</mods:classification>
<mods:identifier id="69FE83B14D8B46AE02D9D89BF9D920FE" type="CLB-Dataset">45884</mods:identifier>
<mods:identifier id="72F6ED6D0107A31770CF7277D9DADCA9" type="DOI">10.11646/zootaxa.3793.5.4</mods:identifier>
<mods:identifier id="119E719C9286836433FB327DCAF8B64C" type="GBIF-Dataset">5f8cac29-bb09-4a52-b618-503b5fc8c954</mods:identifier>
<mods:identifier id="D1F8389C6774510425393D1AC67EE313" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="45A33E08F2EA94A71DB12C072BD4614F" type="Zenodo-Dep">252346</mods:identifier>
<mods:identifier id="308B0409EB39548109E1A313807F797C" type="ZooBank">F9A1E47F-653E-48AD-8FDF-D7D0D59A0A3A</mods:identifier>
</mods:mods>
<treatment id="038587A1FF80FF97FF60FF54F98DF8BB" ID-DOI="http://doi.org/10.5281/zenodo.6133204" ID-GBIF-Taxon="119424651" ID-Zenodo-Dep="6133204" LSID="urn:lsid:plazi:treatment:038587A1FF80FF97FF60FF54F98DF8BB" httpUri="http://treatment.plazi.org/id/038587A1FF80FF97FF60FF54F98DF8BB" lastPageId="5" lastPageNumber="566" pageId="1" pageNumber="562">
<subSubSection id="C336653CFF80FF93FF60FF54FD13FF25" pageId="1" pageNumber="562" type="nomenclature">
<paragraph id="8B9336B7FF80FF93FF60FF54FE0BFF44" blockId="1.[151,522,151,211]" box="[151,522,151,178]" pageId="1" pageNumber="562">
<heading id="D0DB81DBFF80FF93FF60FF54FE0BFF44" bold="true" box="[151,522,151,178]" fontSize="11" level="1" pageId="1" pageNumber="562" reason="1">
<emphasis id="B958EAA5FF80FF93FF60FF54FE0BFF44" bold="true" box="[151,522,151,178]" pageId="1" pageNumber="562">
<taxonomicName id="4C2C4D34FF80FF93FF60FF54FD8DFF47" ID-CoL="9X24F" box="[151,396,151,177]" class="Insecta" family="Calopterygidae" genus="Atrocalopteryx" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="562" phylum="Arthropoda" rank="species" species="auco" status="sp. nov.">
<emphasis id="B958EAA5FF80FF93FF60FF54FD8DFF47" bold="true" box="[151,396,151,177]" italics="true" pageId="1" pageNumber="562">Atrocalopteryx auco</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26B57DEFF80FF93FE64FF5BFE0BFF44" box="[403,522,152,178]" pageId="1" pageNumber="562" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B9336B7FF80FF93FF60FF79FD13FF25" blockId="1.[151,522,151,211]" box="[151,274,186,211]" pageId="1" pageNumber="562">
(
<figureCitation id="13172A32FF80FF93FF68FF79FD08FF25" box="[159,265,186,211]" captionStart-0="FIGURES 1 3" captionStart-1="FIGURES 4 5" captionStart-2="FIGURES 6 9" captionStartId-0="2.[151,264,1792,1815]" captionStartId-1="3.[151,264,1835,1858]" captionStartId-2="4.[151,264,1964,1987]" captionTargetBox-0="[151,1436,193,1771]" captionTargetBox-1="[151,1436,210,1794]" captionTargetBox-2="[261,1286,241,1963]" captionTargetId-0="figure@2.[151,1436,193,1771]" captionTargetId-1="figure@3.[151,1436,193,1814]" captionTargetId-2="figure@4.[261,1286,193,1980]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="FIGURES 1 3. Atrocalopteryx auco spec. nov. photographed in the field. 1) Male holotype photographed by Matti Hämäläinen at the type locality before it was collected; 2 3) Male (2) and female (3) photographed by Tom Kompier at Huu Lien (Tan Lai) on 17 November 2013. Specimens not collected." captionText-1="FIGURES 4 5. Atrocalopteryx auco spec. nov. 4) habitus of holotype male; 5) habitus paratype female." captionText-2="FIGURES 6 9. Atrocalopteryx auco spec. nov., paratype male photographed alive by Sébastien Delonglée on 9 November 2013. 6) head; 7) head, thorax and bases of wing; 8) details of venation at wing base; 9) tip of abdomen, lateroventral view." httpUri-0="https://zenodo.org/record/252347/files/figure.png" httpUri-1="https://zenodo.org/record/252348/files/figure.png" httpUri-2="https://zenodo.org/record/252349/files/figure.png" pageId="1" pageNumber="562">Figs. 19</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C336653CFF80FF93FF60FEC1F949FE71" pageId="1" pageNumber="562" type="materials_examined">
<paragraph id="8B9336B7FF80FF93FF60FEC1F949FE71" blockId="1.[151,1437,258,1903]" pageId="1" pageNumber="562">
<emphasis id="B958EAA5FF80FF93FF60FEC1FD5EFEED" bold="true" box="[151,351,258,283]" pageId="1" pageNumber="562">Material studied</emphasis>
:
<typeStatus id="54978815FF80FF93FE8EFEC1FDE7FEED" box="[377,486,258,283]" pageId="1" pageNumber="562" type="holotype">
<emphasis id="B958EAA5FF80FF93FE8EFEC1FDE7FEED" bold="true" box="[377,486,258,283]" pageId="1" pageNumber="562">Holotype</emphasis>
</typeStatus>
♂:
<collectingCountry id="F33B7627FF80FF93FDE3FEC1FE77FEED" box="[532,630,258,283]" name="Vietnam" pageId="1" pageNumber="562">Vietnam</collectingCountry>
, Lang Son province, Huu Lien, Lan Chau community, a sidestream of Boc stream,
<geoCoordinate id="EE185070FF80FF93FED2FEE4FDCAFEC8" box="[293,459,295,319]" direction="north" orientation="latitude" pageId="1" pageNumber="562" precision="15" value="21.726112">21° 43 34 N</geoCoordinate>
,
<geoCoordinate id="EE185070FF80FF93FE20FEE5FE88FEC8" box="[471,649,294,319]" direction="east" orientation="longitude" pageId="1" pageNumber="562" precision="15" value="106.3875">106° 23 15 E</geoCoordinate>
; alt. ca
<quantity id="4CD49B52FF80FF93FD17FEE5FF28FEC8" box="[736,809,294,319]" metricMagnitude="2" metricUnit="m" metricValue="2.6" pageId="1" pageNumber="562" unit="m" value="260.0">260 m</quantity>
,
<date id="FF921077FF80FF93FCC1FEE4FFC0FEC9" box="[822,961,295,319]" pageId="1" pageNumber="562" value="2008-06-09">9 June 2008</date>
, M. Hämäläinen leg. Deposited at
<collectionCode id="ED3DAE72FF80FF93FABFFEE4F99DFEC8" box="[1352,1436,295,318]" pageId="1" pageNumber="562">RNMH</collectionCode>
(Leiden).
<typeStatus id="54978815FF80FF93FEF1FE89FD7FFE95" box="[262,382,330,355]" pageId="1" pageNumber="562" type="paratype">
<emphasis id="B958EAA5FF80FF93FEF1FE89FD7FFE95" bold="true" box="[262,382,330,355]" pageId="1" pageNumber="562">Paratypes</emphasis>
</typeStatus>
:
<specimenCount id="9D2AFD3EFF80FF93FE62FE88FDC1FE94" box="[405,448,331,355]" pageId="1" pageNumber="562" type="generic">2 ♂</specimenCount>
, 1 ♀, same data as for the
<typeStatus id="54978815FF80FF93FD05FE88FF59FE95" box="[754,856,331,355]" pageId="1" pageNumber="562" type="holotype">holotype</typeStatus>
;
<specimenCount id="9D2AFD3EFF80FF93FC9BFE88FF99FE94" box="[876,920,331,355]" pageId="1" pageNumber="562" type="generic">1 ♂</specimenCount>
,
<collectingCountry id="F33B7627FF80FF93FC52FE89F806FE95" box="[933,1031,330,355]" name="Vietnam" pageId="1" pageNumber="562">Vietnam</collectingCountry>
, Lang Son province, Huu Lien, Tan Lai community,
<geoCoordinate id="EE185070FF80FF93FEB8FEACFDF5FE70" box="[335,500,367,391]" direction="north" orientation="latitude" pageId="1" pageNumber="562" precision="15" value="21.677776">21° 40 40 N</geoCoordinate>
,
<geoCoordinate id="EE185070FF80FF93FE08FEADFEAEFE70" box="[511,687,366,391]" direction="east" orientation="longitude" pageId="1" pageNumber="562" precision="15" value="106.378334">106° 22 42 E</geoCoordinate>
; alt. ca
<quantity id="4CD49B52FF80FF93FCF2FEACFF4FFE70" box="[773,846,367,391]" metricMagnitude="2" metricUnit="m" metricValue="2.2" pageId="1" pageNumber="562" unit="m" value="220.0">220 m</quantity>
,
<date id="FF921077FF80FF93FCADFEACF826FE71" box="[858,1063,366,391]" pageId="1" pageNumber="562" value="2013-11-09">9 November 2013</date>
, Sebastien Delonglee leg.
</paragraph>
</subSubSection>
<subSubSection id="C336653CFF80FF93FF30FE51F991FDE1" pageId="1" pageNumber="562" type="etymology">
<paragraph id="8B9336B7FF80FF93FF30FE51F991FDE1" blockId="1.[151,1437,258,1903]" pageId="1" pageNumber="562">
<emphasis id="B958EAA5FF80FF93FF30FE51FD47FE5D" bold="true" box="[199,326,402,427]" pageId="1" pageNumber="562">Etymology</emphasis>
. The specific epithet
<taxonomicName id="4C2C4D34FF80FF93FDCDFE50FE71FE5C" box="[570,624,403,426]" class="Insecta" family="Calopterygidae" genus="Atrocalopteryx" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="562" phylum="Arthropoda" rank="species" species="auco">
<emphasis id="B958EAA5FF80FF93FDCDFE50FE71FE5C" box="[570,624,403,426]" italics="true" pageId="1" pageNumber="562">auco</emphasis>
</taxonomicName>
is named after Âu Cơ, a character in the Vietnamese mythology. Âu Cơ was a young, beautiful mountain fairy who fell in love with Lac Long Quân (the Dragon Lord of Lac). They married and she gave birth to an egg sac from which hatched a hundred children known collectively as the Bach Viet, the ancestors of the Vietnamese people. Âu Cơ is widely honoured as the mother of Vietnamese civilization.
</paragraph>
</subSubSection>
<subSubSection id="C336653CFF80FF97FF30FDE1FFD9FE03" lastPageId="5" lastPageNumber="566" pageId="1" pageNumber="562" type="description">
<paragraph id="8B9336B7FF80FF93FF30FDE1F8E6FD51" blockId="1.[151,1437,258,1903]" pageId="1" pageNumber="562">
<emphasis id="B958EAA5FF80FF93FF30FDE1FE1FFDCD" bold="true" box="[199,542,546,571]" pageId="1" pageNumber="562">
Description of
<typeStatus id="54978815FF80FF93FE81FDE1FDDCFDCD" box="[374,477,546,571]" pageId="1" pageNumber="562" type="holotype">holotype</typeStatus>
male
</emphasis>
. Head: Eyes in living specimen dark brown above, pale greenish below (
<figureCitation id="13172A32FF80FF93FAA5FDE1F994FDCD" box="[1362,1429,546,571]" captionStart="FIGURES 1 3" captionStartId="2.[151,264,1792,1815]" captionTargetBox="[151,1436,193,1771]" captionTargetId="figure@2.[151,1436,193,1771]" captionTargetPageId="2" captionText="FIGURES 1 3. Atrocalopteryx auco spec. nov. photographed in the field. 1) Male holotype photographed by Matti Hämäläinen at the type locality before it was collected; 2 3) Male (2) and female (3) photographed by Tom Kompier at Huu Lien (Tan Lai) on 17 November 2013. Specimens not collected." httpUri="https://zenodo.org/record/252347/files/figure.png" pageId="1" pageNumber="562">Fig. 1</figureCitation>
; cf.
<figureCitation id="13172A32FF80FF93FF4AFD85FD0FFDA9" box="[189,270,582,607]" captionStart="FIGURES 1 3" captionStartId="2.[151,264,1792,1815]" captionTargetBox="[151,1436,193,1771]" captionTargetId="figure@2.[151,1436,193,1771]" captionTargetPageId="2" captionText="FIGURES 1 3. Atrocalopteryx auco spec. nov. photographed in the field. 1) Male holotype photographed by Matti Hämäläinen at the type locality before it was collected; 2 3) Male (2) and female (3) photographed by Tom Kompier at Huu Lien (Tan Lai) on 17 November 2013. Specimens not collected." httpUri="https://zenodo.org/record/252347/files/figure.png" pageId="1" pageNumber="562">Figs. 2</figureCitation>
,
<figureCitation id="13172A32FF80FF93FEEAFD85FD4BFDA9" box="[285,330,582,607]" captionStart="FIGURES 6 9" captionStartId="4.[151,264,1964,1987]" captionTargetBox="[261,1286,241,1963]" captionTargetId="figure@4.[261,1286,193,1980]" captionTargetPageId="4" captionText="FIGURES 6 9. Atrocalopteryx auco spec. nov., paratype male photographed alive by Sébastien Delonglée on 9 November 2013. 6) head; 7) head, thorax and bases of wing; 8) details of venation at wing base; 9) tip of abdomen, lateroventral view." httpUri="https://zenodo.org/record/252349/files/figure.png" pageId="1" pageNumber="562">67</figureCitation>
). Labium largely yellowish brown, tips of lateral lobes black. Labrum black with yellow lateral spots; base of mandibles with pale spots (cf.
<figureCitation id="13172A32FF80FF93FD7DFDA9FECCFD75" box="[650,717,618,643]" captionStart="FIGURES 6 9" captionStartId="4.[151,264,1964,1987]" captionTargetBox="[261,1286,241,1963]" captionTargetId="figure@4.[261,1286,193,1980]" captionTargetPageId="4" captionText="FIGURES 6 9. Atrocalopteryx auco spec. nov., paratype male photographed alive by Sébastien Delonglée on 9 November 2013. 6) head; 7) head, thorax and bases of wing; 8) details of venation at wing base; 9) tip of abdomen, lateroventral view." httpUri="https://zenodo.org/record/252349/files/figure.png" pageId="1" pageNumber="562">Fig. 6</figureCitation>
). Clypeus, frons and vertex dark metallic green. Antennae black with the anterior surface of pedicel yellow. A distinct postocular tubercle is present on the occiput.
</paragraph>
<paragraph id="8B9336B7FF80FF93FF30FD71FD6DFCAD" blockId="1.[151,1437,258,1903]" pageId="1" pageNumber="562">
Thorax: Prothorax dark metallic green. Synthorax dark, lustrous metallic green with yellowish markings restricted to a distinct narrow yellow stripe on the lower half of metepimeron bordering the second lateral suture and to a yellow triangle on lower part of metinfraepisternum (cf.
<figureCitation id="13172A32FF80FF93FC8AFD39FFCCFCE5" box="[893,973,762,787]" captionStart="FIGURES 4 5" captionStartId="3.[151,264,1835,1858]" captionTargetBox="[151,1436,210,1794]" captionTargetId="figure@3.[151,1436,193,1814]" captionTargetPageId="3" captionText="FIGURES 4 5. Atrocalopteryx auco spec. nov. 4) habitus of holotype male; 5) habitus paratype female." httpUri="https://zenodo.org/record/252348/files/figure.png" pageId="1" pageNumber="562">Figs. 4</figureCitation>
,
<figureCitation id="13172A32FF80FF93FC2DFD38FFEBFCE5" box="[986,1002,763,787]" captionStart="FIGURES 6 9" captionStartId="4.[151,264,1964,1987]" captionTargetBox="[261,1286,241,1963]" captionTargetId="figure@4.[261,1286,193,1980]" captionTargetPageId="4" captionText="FIGURES 6 9. Atrocalopteryx auco spec. nov., paratype male photographed alive by Sébastien Delonglée on 9 November 2013. 6) head; 7) head, thorax and bases of wing; 8) details of venation at wing base; 9) tip of abdomen, lateroventral view." httpUri="https://zenodo.org/record/252349/files/figure.png" pageId="1" pageNumber="562">7</figureCitation>
). Ventral side of synthorax dark. Legs black, middle and hind coxae with small yellowish markings.
<collectingCountry id="F33B7627FF80FF93FCAEFCDDFF90FCC1" box="[857,913,798,823]" name="India" pageId="1" pageNumber="562">Hind</collectingCountry>
tibiae distinctly curved, middle tibiae slightly curved (
<figureCitation id="13172A32FF80FF93FF05FC81FD41FCAD" box="[242,320,834,859]" captionStart="FIGURES 1 3" captionStartId="2.[151,264,1792,1815]" captionTargetBox="[151,1436,193,1771]" captionTargetId="figure@2.[151,1436,193,1771]" captionTargetPageId="2" captionText="FIGURES 1 3. Atrocalopteryx auco spec. nov. photographed in the field. 1) Male holotype photographed by Matti Hämäläinen at the type locality before it was collected; 2 3) Male (2) and female (3) photographed by Tom Kompier at Huu Lien (Tan Lai) on 17 November 2013. Specimens not collected." httpUri="https://zenodo.org/record/252347/files/figure.png" pageId="1" pageNumber="562">Figs. 1</figureCitation>
,
<figureCitation id="13172A32FF80FF93FEBAFC80FD5FFCAD" box="[333,350,835,859]" captionStart="FIGURES 4 5" captionStartId="3.[151,264,1835,1858]" captionTargetBox="[151,1436,210,1794]" captionTargetId="figure@3.[151,1436,193,1814]" captionTargetPageId="3" captionText="FIGURES 4 5. Atrocalopteryx auco spec. nov. 4) habitus of holotype male; 5) habitus paratype female." httpUri="https://zenodo.org/record/252348/files/figure.png" pageId="1" pageNumber="562">4</figureCitation>
).
</paragraph>
<paragraph id="8B9336B7FF80FF93FF30FCA5FF67FB69" blockId="1.[151,1437,258,1903]" pageId="1" pageNumber="562">
Wings: Wings moderately broad; in hind wing the length/breadth ratio is 3.5. Both wings dark opaque brown throughout (
<figureCitation id="13172A32FF80FF93FED4FC49FD73FC55" box="[291,370,906,931]" captionStart="FIGURES 1 3" captionStartId="2.[151,264,1792,1815]" captionTargetBox="[151,1436,193,1771]" captionTargetId="figure@2.[151,1436,193,1771]" captionTargetPageId="2" captionText="FIGURES 1 3. Atrocalopteryx auco spec. nov. photographed in the field. 1) Male holotype photographed by Matti Hämäläinen at the type locality before it was collected; 2 3) Male (2) and female (3) photographed by Tom Kompier at Huu Lien (Tan Lai) on 17 November 2013. Specimens not collected." httpUri="https://zenodo.org/record/252347/files/figure.png" pageId="1" pageNumber="562">Figs. 1</figureCitation>
,
<figureCitation id="13172A32FF80FF93FE75FC48FD92FC55" box="[386,403,907,931]" captionStart="FIGURES 4 5" captionStartId="3.[151,264,1835,1858]" captionTargetBox="[151,1436,210,1794]" captionTargetId="figure@3.[151,1436,193,1814]" captionTargetPageId="3" captionText="FIGURES 4 5. Atrocalopteryx auco spec. nov. 4) habitus of holotype male; 5) habitus paratype female." httpUri="https://zenodo.org/record/252348/files/figure.png" pageId="1" pageNumber="562">4</figureCitation>
). Under surface with a distinct reddish brown sheen viewed from certain angles; on upper side the sheen is less distinct. On upper side all main veins and crossveins are dark. On underside of hind wing main veins paler and most crossveins yellowish (cf.
<figureCitation id="13172A32FF80FF93FCF2FC11FF56FC1D" box="[773,855,978,1003]" captionStart="FIGURES 1 3" captionStartId="2.[151,264,1792,1815]" captionTargetBox="[151,1436,193,1771]" captionTargetId="figure@2.[151,1436,193,1771]" captionTargetPageId="2" captionText="FIGURES 1 3. Atrocalopteryx auco spec. nov. photographed in the field. 1) Male holotype photographed by Matti Hämäläinen at the type locality before it was collected; 2 3) Male (2) and female (3) photographed by Tom Kompier at Huu Lien (Tan Lai) on 17 November 2013. Specimens not collected." httpUri="https://zenodo.org/record/252347/files/figure.png" pageId="1" pageNumber="562">Figs. 2</figureCitation>
,
<figureCitation id="13172A32FF80FF93FC9FFC10FF79FC1D" box="[872,888,979,1003]" captionStart="FIGURES 6 9" captionStartId="4.[151,264,1964,1987]" captionTargetBox="[261,1286,241,1963]" captionTargetId="figure@4.[261,1286,193,1980]" captionTargetPageId="4" captionText="FIGURES 6 9. Atrocalopteryx auco spec. nov., paratype male photographed alive by Sébastien Delonglée on 9 November 2013. 6) head; 7) head, thorax and bases of wing; 8) details of venation at wing base; 9) tip of abdomen, lateroventral view." httpUri="https://zenodo.org/record/252349/files/figure.png" pageId="1" pageNumber="562">7</figureCitation>
), more distinctly so in basal half of wing. On underside of fore wing veins only paler than wing membrane at base; yellow crossveins occur only at base, except in costal and subcostal fields yellow crossveins present slightly apicad to nodus. Median space open, without crossveins. Cubital field with 1718 crossveins in fore wing, 22
<quantity id="4CD49B52FF80FF93FC5AFBFCFFEDFBA1" box="[941,1004,1086,1111]" metricMagnitude="-1" metricUnit="m" metricValue="5.588" pageId="1" pageNumber="562" unit="in" value="22.0">22 in</quantity>
hind wing. Quadrangle with 1315 crossveins in fore wing, 17
<quantity id="4CD49B52FF80FF93FE2BFBA1FE17FB8D" box="[476,534,1122,1147]" metricMagnitude="-1" metricUnit="m" metricValue="3.81" pageId="1" pageNumber="562" unit="in" value="15.0">15 in</quantity>
hind wing. Antenodals (the first series)
<quantity id="4CD49B52FF80FF93FC10FBA1F84DFB8D" box="[999,1100,1122,1147]" metricMagnitude="-1" metricUnit="m" metricValue="9.271" metricValueMax="9.398" metricValueMin="9.144" pageId="1" pageNumber="562" unit="in" value="36.5" valueMax="37.0" valueMin="36.0">3637 in</quantity>
fore wing and
<quantity id="4CD49B52FF80FF93FB0AFBA0F963FB8D" box="[1277,1378,1122,1147]" metricMagnitude="-1" metricUnit="m" metricValue="8.128" metricValueMax="8.382" metricValueMin="7.874" pageId="1" pageNumber="562" unit="in" value="32.0" valueMax="33.0" valueMin="31.0">3133 in</quantity>
hind wing. Vein IR2 runs parallel with RA after its origin (cf.
<figureCitation id="13172A32FF80FF93FCE3FB45FF56FB69" box="[788,855,1158,1183]" captionStart="FIGURES 6 9" captionStartId="4.[151,264,1964,1987]" captionTargetBox="[261,1286,241,1963]" captionTargetId="figure@4.[261,1286,193,1980]" captionTargetPageId="4" captionText="FIGURES 6 9. Atrocalopteryx auco spec. nov., paratype male photographed alive by Sébastien Delonglée on 9 November 2013. 6) head; 7) head, thorax and bases of wing; 8) details of venation at wing base; 9) tip of abdomen, lateroventral view." httpUri="https://zenodo.org/record/252349/files/figure.png" pageId="1" pageNumber="562">Fig. 8</figureCitation>
).
</paragraph>
<paragraph id="8B9336B7FF80FF93FF30FB69FFE8FAD8" blockId="1.[151,1437,258,1903]" pageId="1" pageNumber="562">
Abdomen shining metallic green on dorsum and on sides, except ventrolateral edge of S10 and fine ventral and caudal margins on tergites of S89 yellow. Underside dark with apical end of S37 obscurely paler, underside of S810 conspicuously yellow (cf.
<figureCitation id="13172A32FF80FF93FDEDFB31FE60FAFD" box="[538,609,1266,1291]" captionStart="FIGURES 6 9" captionStartId="4.[151,264,1964,1987]" captionTargetBox="[261,1286,241,1963]" captionTargetId="figure@4.[261,1286,193,1980]" captionTargetPageId="4" captionText="FIGURES 6 9. Atrocalopteryx auco spec. nov., paratype male photographed alive by Sébastien Delonglée on 9 November 2013. 6) head; 7) head, thorax and bases of wing; 8) details of venation at wing base; 9) tip of abdomen, lateroventral view." httpUri="https://zenodo.org/record/252349/files/figure.png" pageId="1" pageNumber="562">Fig. 9</figureCitation>
). Anal appendages black, with basal part of inferiors distinctly yellow; appendages of typical structure for genus. Penis similar to that of
<emphasis id="B958EAA5FF80FF93FC8AFAD4FFE8FAD8" box="[893,1001,1303,1326]" italics="true" pageId="1" pageNumber="562">
<taxonomicName id="4C2C4D34FF80FF93FC8AFAD4FFE5FAD8" box="[893,996,1303,1326]" class="Insecta" family="Calopterygidae" genus="Calopteryx" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="562" phylum="Arthropoda" rank="species" species="atrata">A. atrata</taxonomicName>
.
</emphasis>
</paragraph>
<paragraph id="8B9336B7FF80FF93FF30FAF9FFD9FAA5" blockId="1.[151,1437,258,1903]" box="[199,984,1338,1363]" pageId="1" pageNumber="562">
<emphasis id="B958EAA5FF80FF93FF30FAF9FD77FAA5" bold="true" box="[199,374,1338,1363]" pageId="1" pageNumber="562">Measurements</emphasis>
(mm):
<collectingCountry id="F33B7627FF80FF93FE3DFAF9FE03FAA5" box="[458,514,1338,1363]" name="India" pageId="1" pageNumber="562">Hind</collectingCountry>
wing 40; abdomen (incl. appendages) 54.
</paragraph>
<paragraph id="8B9336B7FF80FF93FF30FA9DF832F985" blockId="1.[151,1437,258,1903]" pageId="1" pageNumber="562">
<emphasis id="B958EAA5FF80FF93FF30FA9DFE1DFA81" bold="true" box="[199,540,1374,1399]" pageId="1" pageNumber="562">
Variation in male
<typeStatus id="54978815FF80FF93FE51FA9DFE1DFA81" box="[422,540,1374,1399]" pageId="1" pageNumber="562" type="paratype">paratypes</typeStatus>
</emphasis>
. One of the
<typeStatus id="54978815FF80FF93FD44FA9CFF1FFA81" box="[691,798,1375,1399]" pageId="1" pageNumber="562" type="paratype">paratypes</typeStatus>
collected in 2008 is teneral, the other not fully mature (semi-mature). The
<typeStatus id="54978815FF80FF93FE8AFA40FDDCFA6D" box="[381,477,1411,1435]" pageId="1" pageNumber="562" type="paratype">paratype</typeStatus>
collected in 2013 is a mature specimen;
<figureCitation id="13172A32FF80FF93FC32FA41F830FA6D" box="[965,1073,1410,1435]" captionStart="FIGURES 6 9" captionStartId="4.[151,264,1964,1987]" captionTargetBox="[261,1286,241,1963]" captionTargetId="figure@4.[261,1286,193,1980]" captionTargetPageId="4" captionText="FIGURES 6 9. Atrocalopteryx auco spec. nov., paratype male photographed alive by Sébastien Delonglée on 9 November 2013. 6) head; 7) head, thorax and bases of wing; 8) details of venation at wing base; 9) tip of abdomen, lateroventral view." httpUri="https://zenodo.org/record/252349/files/figure.png" pageId="1" pageNumber="562">Figs. 69</figureCitation>
illustrate this specimen, which resembles the
<typeStatus id="54978815FF80FF93FECFFA65FD9BFA49" box="[312,410,1446,1471]" pageId="1" pageNumber="562" type="holotype">holotype</typeStatus>
in the colour of body and wings. In the teneral specimen the metallic colour of the body is already weakly developed, legs are brownish and wings pale brownish with distinct violet shine. In the semimature
<typeStatus id="54978815FF80FF93FF18FA2CFD4EF9F1" box="[239,335,1519,1543]" pageId="1" pageNumber="562" type="paratype">paratype</typeStatus>
the wings have greenish or violet iridescent sheen depending on angle of view, except on the wingtip; on upper surface the tip is blackish brown, but on under surface it has a distinct coppery sheen. The body colour of this
<typeStatus id="54978815FF80FF93FEB7F9F4FDA1F9B9" box="[320,416,1591,1615]" pageId="1" pageNumber="562" type="paratype">paratype</typeStatus>
resembles that of the
<typeStatus id="54978815FF80FF93FD5AF9F5FF12F9B9" box="[685,787,1590,1615]" pageId="1" pageNumber="562" type="holotype">holotype</typeStatus>
. The venation in the
<typeStatus id="54978815FF80FF93FBE4F9F4F87EF9B9" box="[1043,1151,1591,1615]" pageId="1" pageNumber="562" type="paratype">paratypes</typeStatus>
is similar to that of the
<typeStatus id="54978815FF80FF93FF60F999FCFDF985" box="[151,252,1626,1651]" pageId="1" pageNumber="562" type="holotype">holotype</typeStatus>
, with insignificant variation in the numbers of crossveins and antenodals.
</paragraph>
<paragraph id="8B9336B7FF80FF93FF30F9BDF82DF961" blockId="1.[151,1437,258,1903]" box="[199,1068,1662,1687]" pageId="1" pageNumber="562">
<emphasis id="B958EAA5FF80FF93FF30F9BDFD77F961" bold="true" box="[199,374,1662,1687]" pageId="1" pageNumber="562">Measurements</emphasis>
(mm):
<collectingCountry id="F33B7627FF80FF93FE3DF9BDFE03F961" box="[458,514,1662,1687]" name="India" pageId="1" pageNumber="562">Hind</collectingCountry>
wing 4143; abdomen (incl. appendages) 5356.
</paragraph>
<paragraph id="8B9336B7FF80FF93FF30F961FF88F8F4" blockId="1.[151,1437,258,1903]" pageId="1" pageNumber="562">
<emphasis id="B958EAA5FF80FF93FF30F961FDCCF94D" bold="true" box="[199,461,1698,1723]" pageId="1" pageNumber="562">Description of female</emphasis>
(
<figureCitation id="13172A32FF80FF93FE17F961FE32F94D" box="[480,563,1698,1723]" captionStart="FIGURES 1 3" captionStartId="2.[151,264,1792,1815]" captionTargetBox="[151,1436,193,1771]" captionTargetId="figure@2.[151,1436,193,1771]" captionTargetPageId="2" captionText="FIGURES 1 3. Atrocalopteryx auco spec. nov. photographed in the field. 1) Male holotype photographed by Matti Hämäläinen at the type locality before it was collected; 2 3) Male (2) and female (3) photographed by Tom Kompier at Huu Lien (Tan Lai) on 17 November 2013. Specimens not collected." httpUri="https://zenodo.org/record/252347/files/figure.png" pageId="1" pageNumber="562">Figs. 3</figureCitation>
,
<figureCitation id="13172A32FF80FF93FDB5F961FE53F94D" box="[578,594,1698,1723]" captionStart="FIGURES 4 5" captionStartId="3.[151,264,1835,1858]" captionTargetBox="[151,1436,210,1794]" captionTargetId="figure@3.[151,1436,193,1814]" captionTargetPageId="3" captionText="FIGURES 4 5. Atrocalopteryx auco spec. nov. 4) habitus of holotype male; 5) habitus paratype female." httpUri="https://zenodo.org/record/252348/files/figure.png" pageId="1" pageNumber="562">5</figureCitation>
). Head: Eyes in living specimens above paler brown than in male, eyes below pale yellowish (
<figureCitation id="13172A32FF80FF93FE6DF905FDE1F929" box="[410,480,1734,1759]" captionStart="FIGURES 1 3" captionStartId="2.[151,264,1792,1815]" captionTargetBox="[151,1436,193,1771]" captionTargetId="figure@2.[151,1436,193,1771]" captionTargetPageId="2" captionText="FIGURES 1 3. Atrocalopteryx auco spec. nov. photographed in the field. 1) Male holotype photographed by Matti Hämäläinen at the type locality before it was collected; 2 3) Male (2) and female (3) photographed by Tom Kompier at Huu Lien (Tan Lai) on 17 November 2013. Specimens not collected." httpUri="https://zenodo.org/record/252347/files/figure.png" pageId="1" pageNumber="562">Fig. 3</figureCitation>
). Colour pattern of head similar to male, but yellow on labrum more extensive, not divided into two separate dots. A small postocular tubercle present.
</paragraph>
<paragraph id="8B9336B7FF80FF93FF30F8CDFE6EF899" blockId="1.[151,1437,258,1903]" pageId="1" pageNumber="562">
Thorax: Metallic green, yellow on sides more extensive than in male. Also lower border of metepimeron and anterioventral edge of mesepimeron obscurely yellowish, and metinfraepisternum pale (
<figureCitation id="13172A32FF80FF93FB37F8F1F90FF8BD" box="[1216,1294,1842,1867]" captionStart="FIGURES 4 5" captionStartId="3.[151,264,1835,1858]" captionTargetBox="[151,1436,210,1794]" captionTargetId="figure@3.[151,1436,193,1814]" captionTargetPageId="3" captionText="FIGURES 4 5. Atrocalopteryx auco spec. nov. 4) habitus of holotype male; 5) habitus paratype female." httpUri="https://zenodo.org/record/252348/files/figure.png" pageId="1" pageNumber="562">Fig. 5</figureCitation>
). Venter of synthorax pale brownish. Legs as in male.
</paragraph>
<caption id="DF53663FFF83FF90FF60F8C3FF1CF8A2" httpUri="https://zenodo.org/record/252347/files/figure.png" pageId="2" pageNumber="563" targetBox="[151,1436,193,1771]" targetPageId="2">
<paragraph id="8B9336B7FF83FF90FF60F8C3FF1CF8A2" blockId="2.[151,1436,1792,1876]" pageId="2" pageNumber="563">
<emphasis id="B958EAA5FF83FF90FF60F8C3FD43F8E0" bold="true" box="[151,322,1792,1815]" pageId="2" pageNumber="563">FIGURES 13.</emphasis>
<taxonomicName id="4C2C4D34FF83FF90FEB8F8C2FE24F8E1" box="[335,549,1793,1815]" class="Insecta" family="Calopterygidae" genus="Atrocalopteryx" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="563" phylum="Arthropoda" rank="species" species="auco" status="sp. nov.">
<emphasis id="B958EAA5FF83FF90FEB8F8C2FE24F8E1" box="[335,549,1793,1815]" italics="true" pageId="2" pageNumber="563">Atrocalopteryx auco</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26B57DEFF83FF90FDC4F8C2FEA0F8E0" box="[563,673,1793,1814]" pageId="2" pageNumber="563" rank="species">
<emphasis id="B958EAA5FF83FF90FDC4F8C2FEA0F8E0" bold="true" box="[563,673,1793,1814]" pageId="2" pageNumber="563">spec. nov.</emphasis>
</taxonomicNameLabel>
photographed in the field.
<emphasis id="B958EAA5FF83FF90FC2DF8C3FFEEF8E1" bold="true" box="[986,1007,1792,1815]" pageId="2" pageNumber="563">1)</emphasis>
Male holotype photographed by Matti Hämäläinen at the type locality before it was collected;
<emphasis id="B958EAA5FF83FF90FD20F8DCFF04F8C0" bold="true" box="[727,773,1823,1846]" pageId="2" pageNumber="563">23)</emphasis>
Male (2) and female (3) photographed by Tom Kompier at Huu Lien (Tan Lai) on 17 November 2013. Specimens not collected.
</paragraph>
</caption>
<caption id="DF53663FFF82FF91FF60F8E8F8BAF8B4" box="[151,1211,1835,1858]" httpUri="https://zenodo.org/record/252348/files/figure.png" pageId="3" pageNumber="564" targetBox="[151,1436,210,1794]" targetPageId="3">
<paragraph id="8B9336B7FF82FF91FF60F8E8F8BAF8B4" blockId="3.[151,1211,1835,1858]" box="[151,1211,1835,1858]" pageId="3" pageNumber="564">
<emphasis id="B958EAA5FF82FF91FF60F8E8FD3BF8B7" bold="true" box="[151,314,1835,1858]" pageId="3" pageNumber="564">FIGURES 45.</emphasis>
<taxonomicName id="4C2C4D34FF82FF91FEB7F8EFFE0FF8B4" box="[320,526,1836,1858]" class="Insecta" family="Calopterygidae" genus="Atrocalopteryx" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="564" phylum="Arthropoda" rank="species" species="auco" status="sp. nov.">
<emphasis id="B958EAA5FF82FF91FEB7F8EFFE0FF8B4" box="[320,526,1836,1858]" italics="true" pageId="3" pageNumber="564">Atrocalopteryx auco</emphasis>
</taxonomicName>
<emphasis id="B958EAA5FF82FF91FDE2F8EFFE99F8B4" bold="true" box="[533,664,1835,1858]" pageId="3" pageNumber="564">
<taxonomicNameLabel id="A26B57DEFF82FF91FDE2F8EFFE7AF8B7" box="[533,635,1836,1857]" pageId="3" pageNumber="564" rank="species">spec. nov.</taxonomicNameLabel>
4)
</emphasis>
habitus of holotype male;
<emphasis id="B958EAA5FF82FF91FC51F8E8FFBDF8B4" bold="true" box="[934,956,1835,1858]" pageId="3" pageNumber="564">5)</emphasis>
habitus paratype female.
</paragraph>
</caption>
<caption id="DF53663FFF85FF96FF60F86FF979F817" httpUri="https://zenodo.org/record/252349/files/figure.png" pageId="4" pageNumber="565" targetBox="[261,1286,241,1963]" targetPageId="4">
<paragraph id="8B9336B7FF85FF96FF60F86FF979F817" blockId="4.[151,1436,1964,2017]" pageId="4" pageNumber="565">
<emphasis id="B958EAA5FF85FF96FF60F86FFD3DF837" bold="true" box="[151,316,1964,1987]" pageId="4" pageNumber="565">FIGURES 69.</emphasis>
<taxonomicName id="4C2C4D34FF85FF96FEB2F86FFE17F834" box="[325,534,1964,1986]" class="Insecta" family="Calopterygidae" genus="Atrocalopteryx" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="565" phylum="Arthropoda" rank="species" species="auco" status="sp. nov.">
<emphasis id="B958EAA5FF85FF96FEB2F86FFE17F834" box="[325,534,1964,1986]" italics="true" pageId="4" pageNumber="565">Atrocalopteryx auco</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26B57DEFF85FF96FDE9F86EFE86F834" box="[542,647,1965,1986]" pageId="4" pageNumber="565" rank="species">
<emphasis id="B958EAA5FF85FF96FDE9F86EFE86F834" bold="true" box="[542,647,1965,1986]" pageId="4" pageNumber="565">spec. nov.</emphasis>
</taxonomicNameLabel>
, paratype male photographed alive by Sébastien Delonglée on 9 November 2013.
<emphasis id="B958EAA5FF85FF96FF21F809FCEBF817" bold="true" box="[214,234,1994,2017]" pageId="4" pageNumber="565">6)</emphasis>
head;
<emphasis id="B958EAA5FF85FF96FED8F809FD42F817" bold="true" box="[303,323,1994,2017]" pageId="4" pageNumber="565">7)</emphasis>
head, thorax and bases of wing;
<emphasis id="B958EAA5FF85FF96FD66F809FEA7F817" bold="true" box="[657,678,1994,2017]" pageId="4" pageNumber="565">8)</emphasis>
details of venation at wing base;
<emphasis id="B958EAA5FF85FF96FC0DF809F80EF817" bold="true" box="[1018,1039,1994,2017]" pageId="4" pageNumber="565">9)</emphasis>
tip of abdomen, lateroventral view.
</paragraph>
</caption>
<paragraph id="8B9336B7FF84FF97FF30FF54FD5AFE7E" blockId="5.[151,1437,151,1869]" pageId="5" pageNumber="566">
Wings (
<figureCitation id="13172A32FF84FF97FEEAFF54FD6DFF46" box="[285,364,151,176]" captionStart="FIGURES 1 3" captionStartId="2.[151,264,1792,1815]" captionTargetBox="[151,1436,193,1771]" captionTargetId="figure@2.[151,1436,193,1771]" captionTargetPageId="2" captionText="FIGURES 1 3. Atrocalopteryx auco spec. nov. photographed in the field. 1) Male holotype photographed by Matti Hämäläinen at the type locality before it was collected; 2 3) Male (2) and female (3) photographed by Tom Kompier at Huu Lien (Tan Lai) on 17 November 2013. Specimens not collected." httpUri="https://zenodo.org/record/252347/files/figure.png" pageId="5" pageNumber="566">Figs. 3</figureCitation>
,
<figureCitation id="13172A32FF84FF97FE8FFF54FD89FF46" box="[376,392,151,176]" captionStart="FIGURES 4 5" captionStartId="3.[151,264,1835,1858]" captionTargetBox="[151,1436,210,1794]" captionTargetId="figure@3.[151,1436,193,1814]" captionTargetPageId="3" captionText="FIGURES 4 5. Atrocalopteryx auco spec. nov. 4) habitus of holotype male; 5) habitus paratype female." httpUri="https://zenodo.org/record/252348/files/figure.png" pageId="5" pageNumber="566">5</figureCitation>
): Dark brownish throughout, but subhyaline at basal 1/4, below costal and subcostal fields. In some angles wings show slight iridescence. Underside of hind wing with pale veins and distinctly yellowish reticulation as in male. White pseudopterostigma in both wings, ventral border slightly curved in middle, extending length of 78 underlying cells. Length/breadth ratio in hind wing 3.45. Median space open, without crossveins. Cubital space with 2122 crossveins in fore wing,
<quantity id="4CD49B52FF84FF97FD21FEE4FF3BFEB6" box="[726,826,295,320]" metricMagnitude="-1" metricUnit="m" metricValue="6.223" metricValueMax="6.35" metricValueMin="6.096" pageId="5" pageNumber="566" unit="in" value="24.5" valueMax="25.0" valueMin="24.0">2425 in</quantity>
hind wing. Quadrangle with 1618 crossveins in fore wing, 19
<quantity id="4CD49B52FF84FF97FEFFFE8FFD41FE93" box="[264,320,332,357]" metricMagnitude="-1" metricUnit="m" metricValue="4.826" pageId="5" pageNumber="566" unit="in" value="19.0">19 in</quantity>
hind wing. Antenodals (first series) 38
<quantity id="4CD49B52FF84FF97FCF2FE8FFF3FFE93" box="[773,830,332,357]" metricMagnitude="-1" metricUnit="m" metricValue="9.144" pageId="5" pageNumber="566" unit="in" value="36.0">36 in</quantity>
fore wing and
<quantity id="4CD49B52FF84FF97FC1EFE8FF84DFE93" box="[1001,1100,332,357]" metricMagnitude="-1" metricUnit="m" metricValue="8.763" metricValueMax="8.89" metricValueMin="8.636" pageId="5" pageNumber="566" unit="in" value="34.5" valueMax="35.0" valueMin="34.0">3435 in</quantity>
hind wing. Position of IR2 as in
<typeStatus id="54978815FF84FF97FF43FEACFD17FE7E" box="[180,278,367,392]" pageId="5" pageNumber="566" type="holotype">holotype</typeStatus>
male.
</paragraph>
<paragraph id="8B9336B7FF84FF97FF30FE57FEDCFE26" blockId="5.[151,1437,151,1869]" pageId="5" pageNumber="566">Abdomen black with slight metallic shine; S89 with brownish dorsal stripe, lower 2/3rd of lateral side and venter of S89 brownish, S10 brownish throughout.</paragraph>
<paragraph id="8B9336B7FF84FF97FF30FE1FFFD9FE03" blockId="5.[151,1437,151,1869]" box="[199,984,476,501]" pageId="5" pageNumber="566">
<emphasis id="B958EAA5FF84FF97FF30FE1FFD77FE03" bold="true" box="[199,374,476,501]" pageId="5" pageNumber="566">Measurements</emphasis>
(mm):
<collectingCountry id="F33B7627FF84FF97FE3DFE1FFE03FE03" box="[458,514,476,501]" name="India" pageId="5" pageNumber="566">Hind</collectingCountry>
wing 45; abdomen (incl. appendages) 52.
</paragraph>
</subSubSection>
<subSubSection id="C336653CFF84FF97FF30FE3CF89EF96E" pageId="5" pageNumber="566" type="distribution">
<paragraph id="8B9336B7FF84FF97FF30FE3CF89CFDEE" blockId="5.[151,1437,151,1869]" box="[199,1181,511,536]" pageId="5" pageNumber="566">
<emphasis id="B958EAA5FF84FF97FF30FE3CFD61FDEE" bold="true" box="[199,352,511,536]" pageId="5" pageNumber="566">Distribution.</emphasis>
So far known only at Huu Lien, Lang Son province, in northern
<collectingCountry id="F33B7627FF84FF97FBC0FE3CF898FDEE" box="[1079,1177,511,536]" name="Vietnam" pageId="5" pageNumber="566">Vietnam</collectingCountry>
.
</paragraph>
<paragraph id="8B9336B7FF84FF97FF30FDE7FFA5FD06" blockId="5.[151,1437,151,1869]" pageId="5" pageNumber="566">
<emphasis id="B958EAA5FF84FF97FF30FDE7FE4EFDCB" bold="true" box="[199,591,548,573]" pageId="5" pageNumber="566">Notes on the habitat and biology.</emphasis>
<taxonomicName id="4C2C4D34FF84FF97FDA1FDE7FEAAFDCA" box="[598,683,548,572]" class="Insecta" family="Calopterygidae" genus="Atrocalopteryx" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="566" phylum="Arthropoda" rank="species" species="auco">
<emphasis id="B958EAA5FF84FF97FDA1FDE7FEAAFDCA" box="[598,683,548,572]" italics="true" pageId="5" pageNumber="566">A. auco</emphasis>
</taxonomicName>
has been found along streams (
<quantity id="4CD49B52FF84FF97FBF9FDE7F857FDCA" box="[1038,1110,548,572]" metricMagnitude="0" metricUnit="m" metricValue="5.0" metricValueMax="7.0" metricValueMin="3.0" pageId="5" pageNumber="566" unit="m" value="5.0" valueMax="7.0" valueMin="3.0">37 m</quantity>
wide) with moderate flow in rather open surroundings in a valley (alt. ca
<quantity id="4CD49B52FF84FF97FD75FD84FF03FD96" box="[642,770,583,608]" metricMagnitude="2" metricUnit="m" metricValue="2.3" metricValueMax="2.6" metricValueMin="2.0" pageId="5" pageNumber="566" unit="m" value="230.0" valueMax="260.0" valueMin="200.0">200260 m</quantity>
) surrounded by limestone karst peaks. At the site where the
<typeStatus id="54978815FF84FF97FF60FDAFFCF8FD73" box="[151,249,620,645]" pageId="5" pageNumber="566" type="holotype">holotype</typeStatus>
was collected a stream emerges from underground beneath a karst peak. The immediate surroundings are cultivated land, but at least in
<date id="FF921077FF84FF97FE0BFD53FE7BFD5E" box="[508,634,656,680]" pageId="5" pageNumber="566" value="2008" valueMax="2009">20082009</date>
there was a small bamboo thicket left, where these damselflies took shelter. The flight period of
<taxonomicName id="4C2C4D34FF84FF97FE21FD77FE2FFD3A" box="[470,558,692,716]" class="Insecta" family="Calopterygidae" genus="Atrocalopteryx" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="566" phylum="Arthropoda" rank="species" species="auco">
<emphasis id="B958EAA5FF84FF97FE21FD77FE2FFD3A" box="[470,558,692,716]" italics="true" pageId="5" pageNumber="566">A. auco</emphasis>
</taxonomicName>
extends from early June to early December at least. The damselflies are alert and difficult to approach, unlike
<taxonomicName id="4C2C4D34FF84FF97FDF2FD1BFE95FD06" box="[517,660,728,752]" class="Insecta" family="Calopterygidae" genus="Calopteryx" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="566" phylum="Arthropoda" rank="species" species="atrocyana">
<emphasis id="B958EAA5FF84FF97FDF2FD1BFE95FD06" box="[517,660,728,752]" italics="true" pageId="5" pageNumber="566">A. atrocyana</emphasis>
</taxonomicName>
, which are more docile.
</paragraph>
<paragraph id="8B9336B7FF84FF97FF30FD3EFEEFFC52" blockId="5.[151,1437,151,1869]" pageId="5" pageNumber="566">
Huu Lien has been designated as a nature reserve covering ca 120 sq km. Much of the area of the reserve is forested, but since the two valleys inside the reserve have a population of ca 3000 people, the forests in the reserve are suffering from continuing exploitation. In some sites
<taxonomicName id="4C2C4D34FF84FF97FCEDFC87FF6EFCAA" box="[794,879,836,860]" class="Insecta" family="Calopterygidae" genus="Atrocalopteryx" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="566" phylum="Arthropoda" rank="species" species="auco">
<emphasis id="B958EAA5FF84FF97FCEDFC87FF6EFCAA" box="[794,879,836,860]" italics="true" pageId="5" pageNumber="566">A. auco</emphasis>
</taxonomicName>
can be seen together with
<taxonomicName id="4C2C4D34FF84FF97FB69FC87F92EFCAA" box="[1182,1327,836,860]" class="Insecta" family="Calopterygidae" genus="Calopteryx" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="566" phylum="Arthropoda" rank="species" species="atrocyana">
<emphasis id="B958EAA5FF84FF97FB69FC87F92EFCAA" box="[1182,1327,836,860]" italics="true" pageId="5" pageNumber="566">A. atrocyana</emphasis>
</taxonomicName>
, which is locally abundant in Huu Lien. However, the latter species prefers shadier streams in good forest environments. It can often be seen in shady places away from streams.
</paragraph>
<paragraph id="8B9336B7FF84FF97FF30FC6CF96FFA8E" blockId="5.[151,1437,151,1869]" pageId="5" pageNumber="566">
<emphasis id="B958EAA5FF84FF97FF30FC6CFE4BFC3E" bold="true" box="[199,586,943,968]" pageId="5" pageNumber="566">Distinguishing characters. Male</emphasis>
—Only two of the earlier known
<taxonomicName id="4C2C4D34FF84FF97FC22FC73F87DFC3E" box="[981,1148,944,968]" class="Insecta" family="Calopterygidae" genus="Atrocalopteryx" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="566" phylum="Arthropoda" rank="genus">
<emphasis id="B958EAA5FF84FF97FC22FC73F87DFC3E" box="[981,1148,944,968]" italics="true" pageId="5" pageNumber="566">Atrocalopteryx</emphasis>
</taxonomicName>
species,
<taxonomicName id="4C2C4D34FF84FF97FB1EFC73F951FC3E" box="[1257,1360,944,968]" class="Insecta" family="Calopterygidae" genus="Calopteryx" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="566" phylum="Arthropoda" rank="species" species="atrata">
<emphasis id="B958EAA5FF84FF97FB1EFC73F951FC3E" box="[1257,1360,944,968]" italics="true" pageId="5" pageNumber="566">A. atrata</emphasis>
</taxonomicName>
(
<figureCitation id="13172A32FF84FF97FA93FC6CFCDCFC1A" captionStart="FIGURES 10 15" captionStartId="6.[151,264,1712,1735]" captionTargetBox="[151,1436,193,1690]" captionTargetId="figure@6.[151,1436,193,1691]" captionTargetPageId="6" captionText="FIGURES 10 15. Males of Atrocalopteryx species photographed in the field. 10) A. atrata, China, Guangdong, by Hao-miao Zhang; 11) A. atrata, Japan, Chiba by Roy Futahashi; 12 13) A. atrocyana, Vietnam, Lang Son, Huu Lien, by Sébastien Delonglée; 14) A. coomani, Vietnam, Hoa Binh, Thuong Tien, by Sébastien Delonglée; 15) A. melli, China, Guangdong, by Shan-lian Mo." httpUri="https://zenodo.org/record/252350/files/figure.png" pageId="5" pageNumber="566">Figs. 1011</figureCitation>
) and
<taxonomicName id="4C2C4D34FF84FF97FEE9FC17FDB1FC1A" box="[286,432,980,1004]" class="Insecta" family="Calopterygidae" genus="Calopteryx" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="566" phylum="Arthropoda" rank="species" species="atrocyana">
<emphasis id="B958EAA5FF84FF97FEE9FC17FDB1FC1A" box="[286,432,980,1004]" italics="true" pageId="5" pageNumber="566">A. atrocyana</emphasis>
</taxonomicName>
(
<figureCitation id="13172A32FF84FF97FE36FC17FE4BFC1A" box="[449,586,980,1005]" captionStart="FIGURES 10 15" captionStartId="6.[151,264,1712,1735]" captionTargetBox="[151,1436,193,1690]" captionTargetId="figure@6.[151,1436,193,1691]" captionTargetPageId="6" captionText="FIGURES 10 15. Males of Atrocalopteryx species photographed in the field. 10) A. atrata, China, Guangdong, by Hao-miao Zhang; 11) A. atrata, Japan, Chiba by Roy Futahashi; 12 13) A. atrocyana, Vietnam, Lang Son, Huu Lien, by Sébastien Delonglée; 14) A. coomani, Vietnam, Hoa Binh, Thuong Tien, by Sébastien Delonglée; 15) A. melli, China, Guangdong, by Shan-lian Mo." httpUri="https://zenodo.org/record/252350/files/figure.png" pageId="5" pageNumber="566">Figs. 1213</figureCitation>
), have completely opaque wings in the male sex. In all other species there are hyaline portions in the wings (
<figureCitation id="13172A32FF84FF97FDEDFC34FEA2FBE6" box="[538,675,1015,1040]" captionStart-0="FIGURES 10 15" captionStart-1="FIGURES 16 21" captionStartId-0="6.[151,264,1712,1735]" captionStartId-1="7.[151,264,1791,1814]" captionTargetBox-0="[151,1436,193,1690]" captionTargetBox-1="[151,1436,193,1769]" captionTargetId-0="figure@6.[151,1436,193,1691]" captionTargetId-1="figure@7.[151,1436,193,1770]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionText-0="FIGURES 10 15. Males of Atrocalopteryx species photographed in the field. 10) A. atrata, China, Guangdong, by Hao-miao Zhang; 11) A. atrata, Japan, Chiba by Roy Futahashi; 12 13) A. atrocyana, Vietnam, Lang Son, Huu Lien, by Sébastien Delonglée; 14) A. coomani, Vietnam, Hoa Binh, Thuong Tien, by Sébastien Delonglée; 15) A. melli, China, Guangdong, by Shan-lian Mo." captionText-1="FIGURES 16 21. Atrocalopteryx species photographed in the field. 16) A. laosica male, Vietnam, Lao Chai, Sapa, by Cuong Do; 17) A. oberthueri male, China Yunnan, Dali, by Hao-miao Zhang; 18) A. atrata female, China, Guizhou, by Hao-miao Zhang; 19) A. atrocyana female, Vietnam, Lang Son, Huu Lien, by Matti Hämäläinen; 20) A. coomani, Vietnam, Hoa Binh, Thuong Tien, by Sébastien Delonglée; 21) A. melli, China, Guangdong, Nankunshan, by Hong-dao Wu." httpUri-0="https://zenodo.org/record/252350/files/figure.png" httpUri-1="https://zenodo.org/record/252351/files/figure.png" pageId="5" pageNumber="566">Figs. 1417</figureCitation>
). The mature male of
<taxonomicName id="4C2C4D34FF84FF97FC69FC3BFFF5FBE6" box="[926,1012,1016,1040]" class="Insecta" family="Calopterygidae" genus="Atrocalopteryx" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="566" phylum="Arthropoda" rank="species" species="auco" status="sp. nov.">
<emphasis id="B958EAA5FF84FF97FC69FC3BFFF5FBE6" box="[926,1012,1016,1040]" italics="true" pageId="5" pageNumber="566">A. auco</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26B57DEFF84FF97FC0BFC34F871FBE6" box="[1020,1136,1015,1040]" pageId="5" pageNumber="566" rank="species">
<emphasis id="B958EAA5FF84FF97FC0BFC34F871FBE6" bold="true" box="[1020,1136,1015,1040]" pageId="5" pageNumber="566">spec. nov.</emphasis>
</taxonomicNameLabel>
differs from the other two opaque-winged species by having yellowish crossveins on the under surface of the wings (
<figureCitation id="13172A32FF84FF97FB41FBDFF908FBC2" box="[1206,1289,1052,1077]" captionStart="FIGURES 1 3" captionStartId="2.[151,264,1792,1815]" captionTargetBox="[151,1436,193,1771]" captionTargetId="figure@2.[151,1436,193,1771]" captionTargetPageId="2" captionText="FIGURES 1 3. Atrocalopteryx auco spec. nov. photographed in the field. 1) Male holotype photographed by Matti Hämäläinen at the type locality before it was collected; 2 3) Male (2) and female (3) photographed by Tom Kompier at Huu Lien (Tan Lai) on 17 November 2013. Specimens not collected." httpUri="https://zenodo.org/record/252347/files/figure.png" pageId="5" pageNumber="566">Figs. 2</figureCitation>
,
<figureCitation id="13172A32FF84FF97FAEFFBDFF929FBC2" box="[1304,1320,1052,1076]" captionStart="FIGURES 6 9" captionStartId="4.[151,264,1964,1987]" captionTargetBox="[261,1286,241,1963]" captionTargetId="figure@4.[261,1286,193,1980]" captionTargetPageId="4" captionText="FIGURES 6 9. Atrocalopteryx auco spec. nov., paratype male photographed alive by Sébastien Delonglée on 9 November 2013. 6) head; 7) head, thorax and bases of wing; 8) details of venation at wing base; 9) tip of abdomen, lateroventral view." httpUri="https://zenodo.org/record/252349/files/figure.png" pageId="5" pageNumber="566">7</figureCitation>
); the pale crossveins are present throughout the hind wing, but in the fore wing present only in the basal area, extending apicad to the nodus at the upper wing border. On the upper surface of both wings the crossveins are dark. Also in
<taxonomicName id="4C2C4D34FF84FF97FF60FB4AFCDAFB56" box="[151,219,1161,1184]" class="Insecta" family="Calopterygidae" genus="Calopteryx" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="566" phylum="Arthropoda" rank="species" species="atrata">
<emphasis id="B958EAA5FF84FF97FF60FB4AFCDAFB56" box="[151,219,1161,1184]" italics="true" pageId="5" pageNumber="566">atrata</emphasis>
</taxonomicName>
and
<taxonomicName id="4C2C4D34FF84FF97FEE2FB4AFD87FB56" box="[277,390,1161,1184]" class="Insecta" family="Calopterygidae" genus="Calopteryx" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="566" phylum="Arthropoda" rank="species" species="atrocyana">
<emphasis id="B958EAA5FF84FF97FEE2FB4AFD87FB56" box="[277,390,1161,1184]" italics="true" pageId="5" pageNumber="566">atrocyana</emphasis>
</taxonomicName>
the crossveins are dark coloured on the upper surface of the wings, but in these species the under surface of the wings does not have yellowish crossveins. In
<taxonomicName id="4C2C4D34FF84FF97FC64FB6EFFD6FB32" box="[915,983,1197,1220]" class="Insecta" family="Calopterygidae" genus="Calopteryx" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="566" phylum="Arthropoda" rank="species" species="atrata">
<emphasis id="B958EAA5FF84FF97FC64FB6EFFD6FB32" box="[915,983,1197,1220]" italics="true" pageId="5" pageNumber="566">atrata</emphasis>
</taxonomicName>
and
<taxonomicName id="4C2C4D34FF84FF97FBE5FB6EF882FB32" box="[1042,1155,1197,1220]" class="Insecta" family="Calopterygidae" genus="Calopteryx" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="566" phylum="Arthropoda" rank="species" species="atrocyana">
<emphasis id="B958EAA5FF84FF97FBE5FB6EF882FB32" box="[1042,1155,1197,1220]" italics="true" pageId="5" pageNumber="566">atrocyana</emphasis>
</taxonomicName>
there are obscurely pale longitudinal veins on the basal part of the under surface of wings. Moreover, at some angles the crossveins of the costal field look pale, but not yellowish. In
<taxonomicName id="4C2C4D34FF84FF97FD76FB36FEF3FAFA" box="[641,754,1269,1292]" class="Insecta" family="Calopterygidae" genus="Calopteryx" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="566" phylum="Arthropoda" rank="species" species="atrocyana">
<emphasis id="B958EAA5FF84FF97FD76FB36FEF3FAFA" box="[641,754,1269,1292]" italics="true" pageId="5" pageNumber="566">atrocyana</emphasis>
</taxonomicName>
the wings are proportionally much broader than in
<taxonomicName id="4C2C4D34FF84FF97FAC2FB36F96DFAFA" box="[1333,1388,1269,1292]" class="Insecta" family="Calopterygidae" genus="Atrocalopteryx" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="566" phylum="Arthropoda" rank="species" species="auco">
<emphasis id="B958EAA5FF84FF97FAC2FB36F96DFAFA" box="[1333,1388,1269,1292]" italics="true" pageId="5" pageNumber="566">auco</emphasis>
</taxonomicName>
and
<taxonomicName id="4C2C4D34FF84FF97FF60FADAFCDAFAC6" box="[151,219,1305,1328]" class="Insecta" family="Calopterygidae" genus="Calopteryx" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="566" phylum="Arthropoda" rank="species" species="atrata">
<emphasis id="B958EAA5FF84FF97FF60FADAFCDAFAC6" box="[151,219,1305,1328]" italics="true" pageId="5" pageNumber="566">atrata</emphasis>
</taxonomicName>
. In mature males of
<taxonomicName id="4C2C4D34FF84FF97FE32FADAFE37FAC6" box="[453,566,1305,1328]" class="Insecta" family="Calopterygidae" genus="Calopteryx" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="566" phylum="Arthropoda" rank="species" species="atrocyana">
<emphasis id="B958EAA5FF84FF97FE32FADAFE37FAC6" box="[453,566,1305,1328]" italics="true" pageId="5" pageNumber="566">atrocyana</emphasis>
</taxonomicName>
and
<taxonomicName id="4C2C4D34FF84FF97FD9BFADAFEB1FAC6" box="[620,688,1305,1328]" class="Insecta" family="Calopterygidae" genus="Calopteryx" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="566" phylum="Arthropoda" rank="species" species="atrata">
<emphasis id="B958EAA5FF84FF97FD9BFADAFEB1FAC6" box="[620,688,1305,1328]" italics="true" pageId="5" pageNumber="566">atrata</emphasis>
</taxonomicName>
both surfaces of the blackish wings reflect a bluish metallic sheen in certain angles, but in
<taxonomicName id="4C2C4D34FF84FF97FE56FAFEFE13FAA2" box="[417,530,1341,1364]" class="Insecta" family="Calopterygidae" genus="Calopteryx" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="566" phylum="Arthropoda" rank="species" species="atrocyana">
<emphasis id="B958EAA5FF84FF97FE56FAFEFE13FAA2" box="[417,530,1341,1364]" italics="true" pageId="5" pageNumber="566">atrocyana</emphasis>
</taxonomicName>
the bluish sheen is more conspicuous. In mature males of
<taxonomicName id="4C2C4D34FF84FF97FB68FAFEF8D4FAA2" box="[1183,1237,1341,1364]" class="Insecta" family="Calopterygidae" genus="Atrocalopteryx" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="566" phylum="Arthropoda" rank="species" species="auco">
<emphasis id="B958EAA5FF84FF97FB68FAFEF8D4FAA2" box="[1183,1237,1341,1364]" italics="true" pageId="5" pageNumber="566">auco</emphasis>
</taxonomicName>
the upper surface of the wings is dark brown with a slight sheen, while the under surface reflects a reddish-brown sheen (
<figureCitation id="13172A32FF84FF97FAEBFA9CF95CFA8E" box="[1308,1373,1375,1400]" captionStart="FIGURES 1 3" captionStartId="2.[151,264,1792,1815]" captionTargetBox="[151,1436,193,1771]" captionTargetId="figure@2.[151,1436,193,1771]" captionTargetPageId="2" captionText="FIGURES 1 3. Atrocalopteryx auco spec. nov. photographed in the field. 1) Male holotype photographed by Matti Hämäläinen at the type locality before it was collected; 2 3) Male (2) and female (3) photographed by Tom Kompier at Huu Lien (Tan Lai) on 17 November 2013. Specimens not collected." httpUri="https://zenodo.org/record/252347/files/figure.png" pageId="5" pageNumber="566">Fig. 1</figureCitation>
).
</paragraph>
<paragraph id="8B9336B7FF84FF97FF30FA47FE46F9DA" blockId="5.[151,1437,151,1869]" pageId="5" pageNumber="566">
<taxonomicName id="4C2C4D34FF84FF97FF30FA47FD00FA6D" box="[199,257,1412,1435]" class="Insecta" family="Calopterygidae" genus="Atrocalopteryx" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="571" phylum="Arthropoda" rank="species" species="auco">
<emphasis id="B958EAA5FF84FF97FF30FA47FD00FA6D" box="[199,257,1412,1435]" italics="true" pageId="5" pageNumber="566">Auco</emphasis>
</taxonomicName>
male differs from the other two species by having the underside of abdominal segments 810 strikingly yellowish (
<figureCitation id="13172A32FF84FF97FEECFA64FD69FA36" box="[283,360,1447,1472]" captionStart="FIGURES 6 9" captionStartId="4.[151,264,1964,1987]" captionTargetBox="[261,1286,241,1963]" captionTargetId="figure@4.[261,1286,193,1980]" captionTargetPageId="4" captionText="FIGURES 6 9. Atrocalopteryx auco spec. nov., paratype male photographed alive by Sébastien Delonglée on 9 November 2013. 6) head; 7) head, thorax and bases of wing; 8) details of venation at wing base; 9) tip of abdomen, lateroventral view." httpUri="https://zenodo.org/record/252349/files/figure.png" pageId="5" pageNumber="566">Fig. 9</figureCitation>
), the yellow extending along the lateroventral border of terga and covering part of the intersegmental rings of S8 and S9, lower half of sides of S10, and base of inferior appendages. In
<taxonomicName id="4C2C4D34FF84FF97FAD3FA0EF969FA12" box="[1316,1384,1485,1508]" class="Insecta" family="Calopterygidae" genus="Calopteryx" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="566" phylum="Arthropoda" rank="species" species="atrata">
<emphasis id="B958EAA5FF84FF97FAD3FA0EF969FA12" box="[1316,1384,1485,1508]" italics="true" pageId="5" pageNumber="566">atrata</emphasis>
</taxonomicName>
and
<taxonomicName id="4C2C4D34FF84FF97FF60FA32FD09F9FE" box="[151,264,1521,1544]" class="Insecta" family="Calopterygidae" genus="Calopteryx" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="566" phylum="Arthropoda" rank="species" species="atrocyana">
<emphasis id="B958EAA5FF84FF97FF60FA32FD09F9FE" box="[151,264,1521,1544]" italics="true" pageId="5" pageNumber="566">atrocyana</emphasis>
</taxonomicName>
these are all black. Unlike the other two species, in
<taxonomicName id="4C2C4D34FF84FF97FCAFFA32FF8EF9FE" box="[856,911,1521,1544]" class="Insecta" family="Calopterygidae" genus="Atrocalopteryx" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="566" phylum="Arthropoda" rank="species" species="auco">
<emphasis id="B958EAA5FF84FF97FCAFFA32FF8EF9FE" box="[856,911,1521,1544]" italics="true" pageId="5" pageNumber="566">auco</emphasis>
</taxonomicName>
there is a short yellow stripe along the second lateral suture of the synthorax (
<figureCitation id="13172A32FF84FF97FE02F9D7FE38F9DA" box="[501,569,1556,1581]" captionStart="FIGURES 6 9" captionStartId="4.[151,264,1964,1987]" captionTargetBox="[261,1286,241,1963]" captionTargetId="figure@4.[261,1286,193,1980]" captionTargetPageId="4" captionText="FIGURES 6 9. Atrocalopteryx auco spec. nov., paratype male photographed alive by Sébastien Delonglée on 9 November 2013. 6) head; 7) head, thorax and bases of wing; 8) details of venation at wing base; 9) tip of abdomen, lateroventral view." httpUri="https://zenodo.org/record/252349/files/figure.png" pageId="5" pageNumber="566">Fig. 7</figureCitation>
).
</paragraph>
<paragraph id="8B9336B7FF84FF97FF30F9FBF89EF96E" blockId="5.[151,1437,151,1869]" pageId="5" pageNumber="566">
On the head of
<taxonomicName id="4C2C4D34FF84FF97FE80F9FAFDACF9A6" box="[375,429,1593,1616]" class="Insecta" family="Calopterygidae" genus="Atrocalopteryx" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="566" phylum="Arthropoda" rank="species" species="auco">
<emphasis id="B958EAA5FF84FF97FE80F9FAFDACF9A6" box="[375,429,1593,1616]" italics="true" pageId="5" pageNumber="566">auco</emphasis>
</taxonomicName>
(
<figureCitation id="13172A32FF84FF97FE4AF9F4FE03F9A6" box="[445,514,1591,1616]" captionStart="FIGURES 6 9" captionStartId="4.[151,264,1964,1987]" captionTargetBox="[261,1286,241,1963]" captionTargetId="figure@4.[261,1286,193,1980]" captionTargetPageId="4" captionText="FIGURES 6 9. Atrocalopteryx auco spec. nov., paratype male photographed alive by Sébastien Delonglée on 9 November 2013. 6) head; 7) head, thorax and bases of wing; 8) details of venation at wing base; 9) tip of abdomen, lateroventral view." httpUri="https://zenodo.org/record/252349/files/figure.png" pageId="5" pageNumber="566">Fig. 6</figureCitation>
) there are two yellow lateral spots on the labrum and a yellow spot at the base of the mandibles. In
<taxonomicName id="4C2C4D34FF84FF97FE9FF99EFDADF982" box="[360,428,1629,1652]" class="Insecta" family="Calopterygidae" genus="Calopteryx" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="566" phylum="Arthropoda" rank="species" species="atrata">
<emphasis id="B958EAA5FF84FF97FE9FF99EFDADF982" box="[360,428,1629,1652]" italics="true" pageId="5" pageNumber="566">atrata</emphasis>
</taxonomicName>
and
<taxonomicName id="4C2C4D34FF84FF97FE1EF99EFE58F982" box="[489,601,1629,1652]" class="Insecta" family="Calopterygidae" genus="Calopteryx" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="566" phylum="Arthropoda" rank="species" species="atrocyana">
<emphasis id="B958EAA5FF84FF97FE1EF99EFE58F982" box="[489,601,1629,1652]" italics="true" pageId="5" pageNumber="566">atrocyana</emphasis>
</taxonomicName>
these yellow markings are absent. In
<taxonomicName id="4C2C4D34FF84FF97FBE2F99EF84AF982" box="[1045,1099,1629,1652]" class="Insecta" family="Calopterygidae" genus="Atrocalopteryx" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="566" phylum="Arthropoda" rank="species" species="auco">
<emphasis id="B958EAA5FF84FF97FBE2F99EF84AF982" box="[1045,1099,1629,1652]" italics="true" pageId="5" pageNumber="566">auco</emphasis>
</taxonomicName>
there is a distinct postocular tubercle on the occiput, while in
<taxonomicName id="4C2C4D34FF84FF97FDF1F942FE4BF96E" box="[518,586,1665,1688]" class="Insecta" family="Calopterygidae" genus="Calopteryx" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="566" phylum="Arthropoda" rank="species" species="atrata">
<emphasis id="B958EAA5FF84FF97FDF1F942FE4BF96E" box="[518,586,1665,1688]" italics="true" pageId="5" pageNumber="566">atrata</emphasis>
</taxonomicName>
the tubercle is absent and in
<taxonomicName id="4C2C4D34FF84FF97FC66F942F800F96E" box="[913,1025,1665,1688]" class="Insecta" family="Calopterygidae" genus="Calopteryx" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="566" phylum="Arthropoda" rank="species" species="atrocyana">
<emphasis id="B958EAA5FF84FF97FC66F942F800F96E" box="[913,1025,1665,1688]" italics="true" pageId="5" pageNumber="566">atrocyana</emphasis>
</taxonomicName>
it is vestigial.
</paragraph>
</subSubSection>
<subSubSection id="C336653CFF84FF97FF30F967F98DF8BB" pageId="5" pageNumber="566" type="description">
<paragraph id="8B9336B7FF84FF97FF30F967F98DF8BB" blockId="5.[151,1437,151,1869]" pageId="5" pageNumber="566">
<emphasis id="B958EAA5FF84FF97FF30F967FD1FF94B" bold="true" box="[199,286,1700,1725]" pageId="5" pageNumber="566">Female</emphasis>
—The female of
<taxonomicName id="4C2C4D34FF84FF97FE13F966FE1BF94A" box="[484,538,1701,1724]" class="Insecta" family="Calopterygidae" genus="Atrocalopteryx" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="566" phylum="Arthropoda" rank="species" species="auco">
<emphasis id="B958EAA5FF84FF97FE13F966FE1BF94A" box="[484,538,1701,1724]" italics="true" pageId="5" pageNumber="566">auco</emphasis>
</taxonomicName>
(
<figureCitation id="13172A32FF84FF97FDDBF967FE71F94A" box="[556,624,1700,1725]" captionStart="FIGURES 1 3" captionStartId="2.[151,264,1792,1815]" captionTargetBox="[151,1436,193,1771]" captionTargetId="figure@2.[151,1436,193,1771]" captionTargetPageId="2" captionText="FIGURES 1 3. Atrocalopteryx auco spec. nov. photographed in the field. 1) Male holotype photographed by Matti Hämäläinen at the type locality before it was collected; 2 3) Male (2) and female (3) photographed by Tom Kompier at Huu Lien (Tan Lai) on 17 November 2013. Specimens not collected." httpUri="https://zenodo.org/record/252347/files/figure.png" pageId="5" pageNumber="566">Fig. 3</figureCitation>
) can be easily separated from
<taxonomicName id="4C2C4D34FF84FF97FC26F966F814F94A" box="[977,1045,1701,1724]" class="Insecta" family="Calopterygidae" genus="Calopteryx" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="566" phylum="Arthropoda" rank="species" species="atrata">
<emphasis id="B958EAA5FF84FF97FC26F966F814F94A" box="[977,1045,1701,1724]" italics="true" pageId="5" pageNumber="566">atrata</emphasis>
</taxonomicName>
and
<taxonomicName id="4C2C4D34FF84FF97FBA7F966F8C1F94A" box="[1104,1216,1701,1724]" class="Insecta" family="Calopterygidae" genus="Calopteryx" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="566" phylum="Arthropoda" rank="species" species="atrocyana">
<emphasis id="B958EAA5FF84FF97FBA7F966F8C1F94A" box="[1104,1216,1701,1724]" italics="true" pageId="5" pageNumber="566">atrocyana</emphasis>
</taxonomicName>
by the presence of whitish pseudopterostigma in both wings; this is absent in
<taxonomicName id="4C2C4D34FF84FF97FCC3F90AFF79F916" box="[820,888,1737,1760]" class="Insecta" family="Calopterygidae" genus="Calopteryx" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="566" phylum="Arthropoda" rank="species" species="atrata">
<emphasis id="B958EAA5FF84FF97FCC3F90AFF79F916" box="[820,888,1737,1760]" italics="true" pageId="5" pageNumber="566">atrata</emphasis>
</taxonomicName>
(
<figureCitation id="13172A32FF84FF97FC7EF904FFDDF916" box="[905,988,1735,1760]" captionStart="FIGURES 16 21" captionStartId="7.[151,264,1791,1814]" captionTargetBox="[151,1436,193,1769]" captionTargetId="figure@7.[151,1436,193,1770]" captionTargetPageId="7" captionText="FIGURES 16 21. Atrocalopteryx species photographed in the field. 16) A. laosica male, Vietnam, Lao Chai, Sapa, by Cuong Do; 17) A. oberthueri male, China Yunnan, Dali, by Hao-miao Zhang; 18) A. atrata female, China, Guizhou, by Hao-miao Zhang; 19) A. atrocyana female, Vietnam, Lang Son, Huu Lien, by Matti Hämäläinen; 20) A. coomani, Vietnam, Hoa Binh, Thuong Tien, by Sébastien Delonglée; 21) A. melli, China, Guangdong, Nankunshan, by Hong-dao Wu." httpUri="https://zenodo.org/record/252351/files/figure.png" pageId="5" pageNumber="566">Fig. 18</figureCitation>
),
<taxonomicName id="4C2C4D34FF84FF97FC04F90AF865F916" box="[1011,1124,1737,1760]" class="Insecta" family="Calopterygidae" genus="Calopteryx" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="566" phylum="Arthropoda" rank="species" species="atrocyana">
<emphasis id="B958EAA5FF84FF97FC04F90AF865F916" box="[1011,1124,1737,1760]" italics="true" pageId="5" pageNumber="566">atrocyana</emphasis>
</taxonomicName>
(
<figureCitation id="13172A32FF84FF97FB82F904F8C9F916" box="[1141,1224,1735,1760]" captionStart="FIGURES 16 21" captionStartId="7.[151,264,1791,1814]" captionTargetBox="[151,1436,193,1769]" captionTargetId="figure@7.[151,1436,193,1770]" captionTargetPageId="7" captionText="FIGURES 16 21. Atrocalopteryx species photographed in the field. 16) A. laosica male, Vietnam, Lao Chai, Sapa, by Cuong Do; 17) A. oberthueri male, China Yunnan, Dali, by Hao-miao Zhang; 18) A. atrata female, China, Guizhou, by Hao-miao Zhang; 19) A. atrocyana female, Vietnam, Lang Son, Huu Lien, by Matti Hämäläinen; 20) A. coomani, Vietnam, Hoa Binh, Thuong Tien, by Sébastien Delonglée; 21) A. melli, China, Guangdong, Nankunshan, by Hong-dao Wu." httpUri="https://zenodo.org/record/252351/files/figure.png" pageId="5" pageNumber="566">Fig. 19</figureCitation>
), and
<taxonomicName id="4C2C4D34FF84FF97FAE7F90BF960F916" box="[1296,1377,1736,1760]" class="Insecta" family="Calopterygidae" genus="Calopteryx" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="566" phylum="Arthropoda" rank="species" species="laosica">
<emphasis id="B958EAA5FF84FF97FAE7F90BF960F916" box="[1296,1377,1736,1760]" italics="true" pageId="5" pageNumber="566">laosica</emphasis>
</taxonomicName>
. The females of both
<taxonomicName id="4C2C4D34FF84FF97FEBCF92FFDC8F8F2" box="[331,457,1772,1796]" class="Insecta" family="Calopterygidae" genus="Atrocalopteryx" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="566" phylum="Arthropoda" rank="species" species="coomani">
<emphasis id="B958EAA5FF84FF97FEBCF92FFDC8F8F2" box="[331,457,1772,1796]" italics="true" pageId="5" pageNumber="566">A. coomani</emphasis>
</taxonomicName>
(
<figureCitation id="13172A32FF84FF97FE2EF92FFE2BF8F2" box="[473,554,1772,1797]" captionStart="FIGURES 16 21" captionStartId="7.[151,264,1791,1814]" captionTargetBox="[151,1436,193,1769]" captionTargetId="figure@7.[151,1436,193,1770]" captionTargetPageId="7" captionText="FIGURES 16 21. Atrocalopteryx species photographed in the field. 16) A. laosica male, Vietnam, Lao Chai, Sapa, by Cuong Do; 17) A. oberthueri male, China Yunnan, Dali, by Hao-miao Zhang; 18) A. atrata female, China, Guizhou, by Hao-miao Zhang; 19) A. atrocyana female, Vietnam, Lang Son, Huu Lien, by Matti Hämäläinen; 20) A. coomani, Vietnam, Hoa Binh, Thuong Tien, by Sébastien Delonglée; 21) A. melli, China, Guangdong, Nankunshan, by Hong-dao Wu." httpUri="https://zenodo.org/record/252351/files/figure.png" pageId="5" pageNumber="566">Fig. 20</figureCitation>
) and
<taxonomicName id="4C2C4D34FF84FF97FD9FF92FFEBFF8F2" box="[616,702,1772,1796]" class="Insecta" family="Calopterygidae" genus="Calopteryx" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="566" phylum="Arthropoda" rank="species" species="melli">
<emphasis id="B958EAA5FF84FF97FD9FF92FFEBFF8F2" box="[616,702,1772,1796]" italics="true" pageId="5" pageNumber="566">A. melli</emphasis>
</taxonomicName>
(
<figureCitation id="13172A32FF84FF97FD3AF92FFF1FF8F2" box="[717,798,1772,1797]" captionStart="FIGURES 16 21" captionStartId="7.[151,264,1791,1814]" captionTargetBox="[151,1436,193,1769]" captionTargetId="figure@7.[151,1436,193,1770]" captionTargetPageId="7" captionText="FIGURES 16 21. Atrocalopteryx species photographed in the field. 16) A. laosica male, Vietnam, Lao Chai, Sapa, by Cuong Do; 17) A. oberthueri male, China Yunnan, Dali, by Hao-miao Zhang; 18) A. atrata female, China, Guizhou, by Hao-miao Zhang; 19) A. atrocyana female, Vietnam, Lang Son, Huu Lien, by Matti Hämäläinen; 20) A. coomani, Vietnam, Hoa Binh, Thuong Tien, by Sébastien Delonglée; 21) A. melli, China, Guangdong, Nankunshan, by Hong-dao Wu." httpUri="https://zenodo.org/record/252351/files/figure.png" pageId="5" pageNumber="566">Fig. 21</figureCitation>
) have quite similarly shaped white pseudopterostigma as
<taxonomicName id="4C2C4D34FF84FF97FF60F8D2FCCCF8DE" box="[151,205,1809,1832]" class="Insecta" family="Calopterygidae" genus="Atrocalopteryx" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="566" phylum="Arthropoda" rank="species" species="auco">
<emphasis id="B958EAA5FF84FF97FF60F8D2FCCCF8DE" box="[151,205,1809,1832]" italics="true" pageId="5" pageNumber="566">auco</emphasis>
</taxonomicName>
, but the three species can be separated by characters presented in the key (p. 571). Moreover,
<taxonomicName id="4C2C4D34FF84FF97FB00F8D2F92FF8DE" box="[1271,1326,1809,1832]" class="Insecta" family="Calopterygidae" genus="Atrocalopteryx" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="566" phylum="Arthropoda" rank="species" species="auco">
<emphasis id="B958EAA5FF84FF97FB00F8D2F92FF8DE" box="[1271,1326,1809,1832]" italics="true" pageId="5" pageNumber="566">auco</emphasis>
</taxonomicName>
female is smaller than the other two species, and in
<taxonomicName id="4C2C4D34FF84FF97FD9AF8F6FEA5F8BA" box="[621,676,1845,1868]" class="Insecta" family="Calopterygidae" genus="Atrocalopteryx" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="566" phylum="Arthropoda" rank="species" species="auco">
<emphasis id="B958EAA5FF84FF97FD9AF8F6FEA5F8BA" box="[621,676,1845,1868]" italics="true" pageId="5" pageNumber="566">auco</emphasis>
</taxonomicName>
the yellow on metepisternum and metepimeron is more restricted.
</paragraph>
</subSubSection>
</treatment>
</document>