treatments-xml/data/D2/5A/D0/D25AD024FFA3983601C7A83F70F6F96C.xml

<document id="4FFE2DE645BC410A78F75E728DE2F318" ID-CLB-Dataset="55626" ID-DOI="10.11646/zootaxa.5133.4.2" ID-GBIF-Dataset="0cd92d63-eeac-48d6-9c9d-270f866de7d3" ID-ISSN="1175-5326" ID-Zenodo-Dep="6530956" ID-ZooBank="BD6A434A-21D7-49FC-A09F-FEF4BC740B14" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1652085552068" checkinUser="plazi" docAuthor="Joharchi, Omid, Döker, Ismail &amp; Khaustov, Vladimir A." docDate="2022" docId="D25AD024FFA3983601C7A83F70F6F96C" docLanguage="en" docName="zootaxa.5133.4.2.pdf" docOrigin="Zootaxa 5133 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cosmolaelaps lasiophilus Joharchi 2022, sp. nov." docType="treatment" docVersion="7" lastPageNumber="494" masterDocId="2E63A85CFFA0983E0150AC5D7325FFD1" masterDocTitle="New species and new records of Cosmolaelaps Berlese (Acari: Laelapidae) from Russia, with a review of the Russian species of the genus" masterLastPageNumber="508" masterPageNumber="486" pageNumber="489" updateTime="1699329983419" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="CFF54445B5B95D5F33BB59EB57392D57">New species and new records of Cosmolaelaps Berlese (Acari: Laelapidae) from Russia, with a review of the Russian species of the genus</mods:title>
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<mods:namePart id="9029249C56F5EFD36802B6AA96913877">Joharchi, Omid</mods:namePart>
<mods:affiliation id="F93D9EF8691A598BBBC580D60CAD74E9">Institute of Environmental and Agricultural Biology (X-BIO), Tyumen State University, Semakova Str. 10, 625003 Tyumen, Russia.</mods:affiliation>
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<mods:namePart id="E71A064C8BE9C20CE99DE1C93C5BC37D">Döker, Ismail</mods:namePart>
<mods:nameIdentifier id="A6D4ADE5120ACC36621BB22034E946D0" type="ORCID">0000-0002-2741-4946</mods:nameIdentifier>
<mods:affiliation id="A48F3551AE81BBD549E8029ECC6D1273">Institute of Environmental and Agricultural Biology (X-BIO), Tyumen State University, Semakova Str. 10, 625003 Tyumen, Russia. &amp; Cukurova University, Agricultural Faculty, Department of Plant Protection, Acarology Laboratory, 01330, Adana, Turkey. https: // orcid. org / 0000 - 0002 - 2741 - 4946 (Joharchi) https: // orcid. org / 0000 - 0002 - 1412 - 1554 (Döker) https: // orcid. org / 0000 - 0003 - 2831 - 7213 (Khaustov)</mods:affiliation>
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<mods:namePart id="7A83C65CCB94420587B35145E6A4D9C0">Khaustov, Vladimir A.</mods:namePart>
<mods:affiliation id="BE1319F2086B9866B58CD121EF985CD8">Institute of Environmental and Agricultural Biology (X-BIO), Tyumen State University, Semakova Str. 10, 625003 Tyumen, Russia.</mods:affiliation>
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<treatment id="D25AD024FFA3983601C7A83F70F6F96C" ID-DOI="http://doi.org/10.5281/zenodo.6533186" ID-GBIF-Taxon="195277958" ID-Zenodo-Dep="6533186" LSID="urn:lsid:plazi:treatment:D25AD024FFA3983601C7A83F70F6F96C" httpUri="http://treatment.plazi.org/id/D25AD024FFA3983601C7A83F70F6F96C" lastPageId="8" lastPageNumber="494" pageId="3" pageNumber="489">
<subSubSection id="12E932B9FFA3983D01C7A83F7180FBAC" box="[151,677,1122,1149]" pageId="3" pageNumber="489" type="nomenclature">
<paragraph id="5A4C6132FFA3983D01C7A83F7180FBAC" blockId="3.[151,677,1122,1185]" box="[151,677,1122,1149]" pageId="3" pageNumber="489">
<heading id="0104D65EFFA3983D01C7A83F7180FBAC" bold="true" box="[151,677,1122,1149]" fontSize="11" level="1" pageId="3" pageNumber="489" reason="1">
<emphasis id="6887BD20FFA3983D01C7A83F7180FBAC" bold="true" box="[151,677,1122,1149]" pageId="3" pageNumber="489">
<taxonomicName id="9DF31AB1FFA3983D01C7A83F7165FBAD" authority="Joharchi" authorityName="Joharchi" authorityYear="2022" box="[151,576,1122,1149]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="489" phylum="Arthropoda" rank="species" species="lasiophilus" status="sp. nov.">
<emphasis id="6887BD20FFA3983D01C7A83F72EFFBAC" bold="true" box="[151,458,1122,1149]" italics="true" pageId="3" pageNumber="489">Cosmolaelaps lasiophilus</emphasis>
Joharchi
</taxonomicName>
<taxonomicNameLabel id="73B4005BFFA3983D0317A83E7180FBAC" box="[583,677,1123,1149]" pageId="3" pageNumber="489" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="12E932B9FFA3983D01C7A8DA7218FB70" box="[151,317,1159,1185]" pageId="3" pageNumber="489" type="description">
<paragraph id="5A4C6132FFA3983D01C7A8DA7218FB70" blockId="3.[151,677,1122,1185]" box="[151,317,1159,1185]" pageId="3" pageNumber="489">
(
<figureCitation id="C2C87DB7FFA3983D01CFA8DA7211FB70" box="[159,308,1159,1185]" captionStart-0="FIGURES 8–12" captionStart-1="FIGURES 13–20" captionStart-2="FIGURE 21–24" captionStartId-0="4.[151,264,1943,1967]" captionStartId-1="5.[151,264,1908,1932]" captionStartId-2="6.[151,250,1926,1950]" captionTargetBox-0="[181,1397,196,1901]" captionTargetBox-1="[198,1388,181,1884]" captionTargetBox-2="[161,1425,197,1887]" captionTargetId-0="figure-18@4.[167,1419,181,1919]" captionTargetId-1="figure-19@5.[198,1389,181,1884]" captionTargetId-2="figure-18@6.[152,1436,181,1902]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionText-0="FIGURES 8–12. Cosmolaelaps lasiophilus Joharchi sp. nov., female. 8, dorsal idiosoma, enlarged section not to scale; 9, ventral idiosoma; 10, subcapitulum; 11, epistome; 12, chelicera." captionText-1="FIGURES 13–20. DIC micrographs of Cosmolaelaps lasiophilus Joharchi sp. nov., female. 13, idiosoma in dorsal view; 14, idiosoma in ventral view; 15, general view of some podonotal setae; 16, general view of some opisthonotal setae; 17, general view of part of opisthogastric area; 18, subcapitulum; 19, distal portion of palp, with focus on tarsal claw; 20, insemination structures." captionText-2="FIGURE 21–24. Cosmolaelaps lasiophilus Joharchi sp. nov., female. 21, leg I (trochanter-tibia); 22, leg II; 23, leg III; 24, leg IV." figureDoi-0="http://doi.org/10.5281/zenodo.6530964" figureDoi-1="http://doi.org/10.5281/zenodo.6530968" figureDoi-2="http://doi.org/10.5281/zenodo.6530972" httpUri-0="https://zenodo.org/record/6530964/files/figure.png" httpUri-1="https://zenodo.org/record/6530968/files/figure.png" httpUri-2="https://zenodo.org/record/6530972/files/figure.png" pageId="3" pageNumber="489">Figures 8–24</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="12E932B9FFA3983D01C7A892700BFAE0" pageId="3" pageNumber="489" type="materials_examined">
<paragraph id="5A4C6132FFA3983D01C7A892700BFAE0" blockId="3.[151,1437,1230,2013]" pageId="3" pageNumber="489">
<emphasis id="6887BD20FFA3983D01C7A8927263FB39" bold="true" box="[151,326,1230,1257]" pageId="3" pageNumber="489">Type material.</emphasis>
<materialsCitation id="EA9B6B6FFFA3983D001DA89276B8FADC" ID-GBIF-Occurrence="3763889304" collectedFrom="in the nest of Lasius niger (L.) (Hymenoptera: Formicidae)" collectingDate="2021-10-18" collectionCode="TUMZ" collectorName="Alexander A. Khaustov" country="Russia" latitude="57.066666" location="Uspenka" longLatPrecision="1053" longitude="65.066666" municipality="Uspenka" pageId="3" pageNumber="489" specimenCount="1" specimenCount-female="1" stateProvince="Tyumen Province" typeStatus="holotype">
<typeStatus id="8548DF90FFA3983D001DA892729DFB38" box="[333,440,1231,1257]" pageId="3" pageNumber="489" type="holotype">Holotype</typeStatus>
, 
<specimenCount id="4CF5AABBFFA3983D0094A8927136FB38" box="[452,531,1231,1257]" pageId="3" pageNumber="489" type="female">female</specimenCount>
, vicinities of 
<collectingMunicipality id="BA28FB48FFA3983D03FCA8927034FB38" box="[684,785,1231,1257]" pageId="3" pageNumber="489">Uspenka</collectingMunicipality>
, 
<collectingRegion id="9837AFD0FFA3983D024DA89270C3FB38" box="[797,998,1231,1257]" country="Russia" name="Tyumen'" pageId="3" pageNumber="489">Tyumen Province</collectingRegion>
, 
<collectingCountry id="22E421A2FFA3983D02A2A892771AFB38" box="[1010,1087,1231,1257]" name="Russia" pageId="3" pageNumber="489">Russia</collectingCountry>
, 
<geoCoordinate id="3FC707F5FFA3983D051BA893778EFB38" box="[1099,1195,1230,1257]" degrees="57" direction="north" minutes="04" orientation="latitude" pageId="3" pageNumber="489" precision="925" value="57.066666">57°04'N</geoCoordinate>
, 
<geoCoordinate id="3FC707F5FFA3983D05E6A8937637FB38" box="[1206,1298,1230,1257]" degrees="65" direction="east" minutes="04" orientation="longitude" pageId="3" pageNumber="489" precision="925" value="65.066666">65°04'E</geoCoordinate>
, 
<date id="2E4D47F2FFA3983D044EA89273F7FADC" pageId="3" pageNumber="489" value="2021-10-18">
<collectingDate id="3E09BE1AFFA3983D044EA89273F7FADC" pageId="3" pageNumber="489" value="2021-10-18">18 October 2021</collectingDate>
</date>
, 
<collectorName id="F70604E4FFA3983D01B1A8AE72D6FADC" box="[225,499,1267,1293]" pageId="3" pageNumber="489">Alexander A. Khaustov</collectorName>
coll., 
<collectedFrom id="E9EF22CFFFA3983D0314A8AE7620FADF" box="[580,1285,1267,1294]" pageId="3" pageNumber="489">
in the nest of 
<taxonomicName id="9DF31AB1FFA3983D03A2A8AE70A6FADD" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[754,899,1267,1292]" class="Insecta" family="Formicidae" genus="Lasius" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="489" phylum="Arthropoda" rank="species" species="niger">
<emphasis id="6887BD20FFA3983D03A2A8AE70A6FADD" box="[754,899,1267,1292]" italics="true" pageId="3" pageNumber="489">Lasius niger</emphasis>
</taxonomicName>
(L.) (
<taxonomicName id="9DF31AB1FFA3983D029EA8AE7748FADC" box="[974,1133,1267,1293]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="489" phylum="Arthropoda" rank="order">Hymenoptera</taxonomicName>
: 
<taxonomicName id="9DF31AB1FFA3983D052CA8AE77DBFADF" box="[1148,1278,1267,1294]" class="Insecta" family="Formicidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="489" phylum="Arthropoda" rank="family">Formicidae</taxonomicName>
)
</collectedFrom>
(in 
<collectionCode id="3CE2F9F7FFA3983D046DA8AE76B7FADC" box="[1341,1426,1267,1293]" pageId="3" pageNumber="489">TUMZ</collectionCode>
).
</materialsCitation>
<materialsCitation id="EA9B6B6FFFA3983D01C7A94A700BFAE0" ID-GBIF-Occurrence="3763889301" box="[151,814,1303,1329]" pageId="3" pageNumber="489" typeStatus="paratype">
<typeStatus id="8548DF90FFA3983D01C7A94A722DFAE0" box="[151,264,1303,1329]" pageId="3" pageNumber="489" type="paratype">Paratypes</typeStatus>
: 
<specimenCount id="4CF5AABBFFA3983D0043A94A72BAFAE0" box="[275,415,1303,1329]" pageId="3" pageNumber="489" type="female">five females</specimenCount>
, same data as 
<typeStatus id="8548DF90FFA3983D036FA94A7184FAE0" box="[575,673,1303,1329]" pageId="3" pageNumber="489" type="holotype">holotype</typeStatus>
(in 
<collectionCode id="3CE2F9F7FFA3983D039DA94A7007FAE0" box="[717,802,1303,1329]" pageId="3" pageNumber="489">TUMZ</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="12E932B9FFA3983D0197A967773FF9A4" pageId="3" pageNumber="489" type="diagnosis">
<paragraph id="5A4C6132FFA3983D0197A967773FF9A4" blockId="3.[151,1437,1230,2013]" pageId="3" pageNumber="489">
<emphasis id="6887BD20FFA3983D0197A96772D2FA85" bold="true" box="[199,503,1338,1364]" pageId="3" pageNumber="489">Diagnosis (adult female).</emphasis>
Dorsal shield with weak reticulate ornamentation, bearing 39 pairs of relatively long (sub-equal), knife-shaped setae, mostly reaching base of next posterior setae in series, 
<emphasis id="6887BD20FFA3983D05C0A902778BFAA8" box="[1168,1198,1375,1401]" italics="true" pageId="3" pageNumber="489">Z5</emphasis>
shorter than 
<emphasis id="6887BD20FFA3983D046FA902767CFAA8" box="[1343,1369,1375,1401]" italics="true" pageId="3" pageNumber="489">J5</emphasis>
(ratio of 
<emphasis id="6887BD20FFA3983D01E7A9DE73F7FA4C" box="[183,210,1411,1437]" italics="true" pageId="3" pageNumber="489">J5</emphasis>
/ 
<emphasis id="6887BD20FFA3983D018AA9DE73D2FA4C" box="[218,247,1411,1437]" italics="true" pageId="3" pageNumber="489">Z5</emphasis>
length ≃1.5); presternal area lightly sclerotised, anterior margin of shield hardly conspicuous, posterior margin distinctly concave, ratio of shield length/width (at broadest level) ≃ 0.8; genital shield ratio of length/ width (at broadest level) ≃ 2.4, shield ornamented posteromedially by 7–8 cells flanked by a median Λ-shaped ornamentation, anal shield ratio of length/width (at broadest level) ≃ 1.2, circum-anal setae smooth and subequal, post-anal seta thickened; distance between genital and anal shields is shorter than length of anal shield. Fixed digit of chelicera with six teeth of various sizes. Legs without spatulate setae but most setae conspicuously thickened and some inserted on small tubercles. Legs I and IV shorter than length of idiosoma.
</paragraph>
</subSubSection>
<subSubSection id="12E932B9FFA398360197AA2372ADFE90" lastPageId="8" lastPageNumber="494" pageId="3" pageNumber="489" type="description">
<paragraph id="5A4C6132FFA3983D0197AA2372D9F94B" blockId="3.[151,1437,1230,2013]" box="[199,508,1662,1690]" pageId="3" pageNumber="489">
<emphasis id="6887BD20FFA3983D0197AA237292F948" bold="true" box="[199,439,1662,1689]" pageId="3" pageNumber="489">Description. Female</emphasis>
(n=6)
</paragraph>
<paragraph id="5A4C6132FFA3983D0197AAFE7167F80C" blockId="3.[151,1437,1230,2013]" pageId="3" pageNumber="489">
<emphasis id="6887BD20FFA3983D0197AAFE72A6F96C" box="[199,387,1699,1725]" italics="true" pageId="3" pageNumber="489">Dorsal idiosoma</emphasis>
(
<figureCitation id="C2C87DB7FFA3983D00C1AAFE72FCF96C" box="[401,473,1699,1725]" captionStart="FIGURES 8–12" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[181,1397,196,1901]" captionTargetId="figure-18@4.[167,1419,181,1919]" captionTargetPageId="4" captionText="FIGURES 8–12. Cosmolaelaps lasiophilus Joharchi sp. nov., female. 8, dorsal idiosoma, enlarged section not to scale; 9, ventral idiosoma; 10, subcapitulum; 11, epistome; 12, chelicera." figureDoi="http://doi.org/10.5281/zenodo.6530964" httpUri="https://zenodo.org/record/6530964/files/figure.png" pageId="3" pageNumber="489">Figs 8</figureCitation>
, 
<figureCitation id="C2C87DB7FFA3983D00B6AAFE717DF96F" box="[486,600,1698,1726]" captionStart="FIGURES 13–20" captionStartId="5.[151,264,1908,1932]" captionTargetBox="[198,1388,181,1884]" captionTargetId="figure-19@5.[198,1389,181,1884]" captionTargetPageId="5" captionText="FIGURES 13–20. DIC micrographs of Cosmolaelaps lasiophilus Joharchi sp. nov., female. 13, idiosoma in dorsal view; 14, idiosoma in ventral view; 15, general view of some podonotal setae; 16, general view of some opisthonotal setae; 17, general view of part of opisthogastric area; 18, subcapitulum; 19, distal portion of palp, with focus on tarsal claw; 20, insemination structures." figureDoi="http://doi.org/10.5281/zenodo.6530968" httpUri="https://zenodo.org/record/6530968/files/figure.png" pageId="3" pageNumber="489">13, 15, 16</figureCitation>
). Dorsal shield oval shaped, small, 389–406 long, 256–265 wide, covering entire dorsal idiosoma, shield with weak reticulation, more distinct in opisthonotal and lateral regions. Shield with 39 pairs of almost uniform length setae (28–40), including two pairs of 
<emphasis id="6887BD20FFA3983D029AAAB670C3F8D5" box="[970,998,1771,1796]" italics="true" pageId="3" pageNumber="489">Zx</emphasis>
setae between 
<emphasis id="6887BD20FFA3983D05C5AAB67784F8D5" box="[1173,1185,1771,1796]" italics="true" pageId="3" pageNumber="489">J</emphasis>
and 
<emphasis id="6887BD20FFA3983D058BAAB677CEF8D5" box="[1243,1259,1771,1796]" italics="true" pageId="3" pageNumber="489">Z</emphasis>
setae and three unpaired supernumerary seta 
<emphasis id="6887BD20FFA3983D00B7AB527125F8F9" box="[487,512,1807,1832]" italics="true" pageId="3" pageNumber="489">Jx</emphasis>
between 
<emphasis id="6887BD20FFA3983D033EAB5271ADF8F8" box="[622,648,1807,1833]" italics="true" pageId="3" pageNumber="489">J2</emphasis>
and 
<emphasis id="6887BD20FFA3983D0392AB5271F8F8F8" box="[706,733,1807,1833]" italics="true" pageId="3" pageNumber="489">J4</emphasis>
, except 
<emphasis id="6887BD20FFA3983D026EAB527071F8F8" box="[830,852,1807,1833]" italics="true" pageId="3" pageNumber="489">j1</emphasis>
, 
<emphasis id="6887BD20FFA3983D0234AB5270A4F8F8" box="[868,897,1807,1833]" italics="true" pageId="3" pageNumber="489">Z5</emphasis>
(14–18) simple and thickened, 
<emphasis id="6887BD20FFA3983D05BCAB527627F8F8" box="[1260,1282,1807,1833]" italics="true" pageId="3" pageNumber="489">j1</emphasis>
apically bent (hook shaped), 
<emphasis id="6887BD20FFA3983D0014AB6E7279F89C" box="[324,348,1843,1869]" italics="true" pageId="3" pageNumber="489">z1</emphasis>
(15–18) and 
<emphasis id="6887BD20FFA3983D00A1AB6E7129F89C" box="[497,524,1843,1869]" italics="true" pageId="3" pageNumber="489">J5</emphasis>
(21–24) shorter (
<figureCitation id="C2C87DB7FFA3983D039DAB6E706BF89F" box="[717,846,1842,1870]" captionStart="FIGURES 13–20" captionStartId="5.[151,264,1908,1932]" captionTargetBox="[198,1388,181,1884]" captionTargetId="figure-19@5.[198,1389,181,1884]" captionTargetPageId="5" captionText="FIGURES 13–20. DIC micrographs of Cosmolaelaps lasiophilus Joharchi sp. nov., female. 13, idiosoma in dorsal view; 14, idiosoma in ventral view; 15, general view of some podonotal setae; 16, general view of some opisthonotal setae; 17, general view of part of opisthogastric area; 18, subcapitulum; 19, distal portion of palp, with focus on tarsal claw; 20, insemination structures." figureDoi="http://doi.org/10.5281/zenodo.6530968" httpUri="https://zenodo.org/record/6530968/files/figure.png" pageId="3" pageNumber="489">Figs 15, 16</figureCitation>
). All setae slightly swollen near the base and pointed at tip (knife-shaped), reaching base of following setae, setae of central area shorter (not reaching base of following setae) (
<figureCitation id="C2C87DB7FFA3983D01B6AB267209F844" box="[230,300,1915,1941]" captionStart="FIGURES 8–12" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[181,1397,196,1901]" captionTargetId="figure-18@4.[167,1419,181,1919]" captionTargetPageId="4" captionText="FIGURES 8–12. Cosmolaelaps lasiophilus Joharchi sp. nov., female. 8, dorsal idiosoma, enlarged section not to scale; 9, ventral idiosoma; 10, subcapitulum; 11, epistome; 12, chelicera." figureDoi="http://doi.org/10.5281/zenodo.6530964" httpUri="https://zenodo.org/record/6530964/files/figure.png" pageId="3" pageNumber="489">Figs 8</figureCitation>
, 
<figureCitation id="C2C87DB7FFA3983D0067AB267282F847" box="[311,423,1914,1942]" captionStart="FIGURES 13–20" captionStartId="5.[151,264,1908,1932]" captionTargetBox="[198,1388,181,1884]" captionTargetId="figure-19@5.[198,1389,181,1884]" captionTargetPageId="5" captionText="FIGURES 13–20. DIC micrographs of Cosmolaelaps lasiophilus Joharchi sp. nov., female. 13, idiosoma in dorsal view; 14, idiosoma in ventral view; 15, general view of some podonotal setae; 16, general view of some opisthonotal setae; 17, general view of part of opisthogastric area; 18, subcapitulum; 19, distal portion of palp, with focus on tarsal claw; 20, insemination structures." figureDoi="http://doi.org/10.5281/zenodo.6530968" httpUri="https://zenodo.org/record/6530968/files/figure.png" pageId="3" pageNumber="489">13, 15, 16</figureCitation>
). Dorsal shield with 22 pairs of pore-like structures, including seven pairs of gland openings (
<emphasis id="6887BD20FFA3983D01F1ABC273EFF868" box="[161,202,1951,1977]" italics="true" pageId="3" pageNumber="489">gd1</emphasis>
, 
<emphasis id="6887BD20FFA3983D0189ABC27226F868" box="[217,259,1951,1977]" italics="true" pageId="3" pageNumber="489">gd2</emphasis>
, 
<emphasis id="6887BD20FFA3983D0042ABC27219F868" box="[274,316,1951,1977]" italics="true" pageId="3" pageNumber="489">gd4</emphasis>
, 
<emphasis id="6887BD20FFA3983D001BABC27250F868" box="[331,373,1951,1977]" italics="true" pageId="3" pageNumber="489">gd5</emphasis>
, 
<emphasis id="6887BD20FFA3983D00D3ABC37288F869" box="[387,429,1950,1976]" italics="true" pageId="3" pageNumber="489">gd6</emphasis>
, 
<emphasis id="6887BD20FFA3983D00ECABC272C3F868" box="[444,486,1951,1977]" italics="true" pageId="3" pageNumber="489">gd8</emphasis>
, 
<emphasis id="6887BD20FFA3983D00A5ABC2713BF868" box="[501,542,1951,1977]" italics="true" pageId="3" pageNumber="489">gd9</emphasis>
) and 15 pairs of poroids, 
<emphasis id="6887BD20FFA3983D0213ABC37042F869" box="[835,871,1950,1976]" italics="true" pageId="3" pageNumber="489">id6</emphasis>
indistinct. Shape, position and relative length and shape of setae shown in 
<figureCitation id="C2C87DB7FFA3983D00F8AB9E7137F80C" box="[424,530,1987,2013]" captionStart="FIGURES 8–12" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[181,1397,196,1901]" captionTargetId="figure-18@4.[167,1419,181,1919]" captionTargetPageId="4" captionText="FIGURES 8–12. Cosmolaelaps lasiophilus Joharchi sp. nov., female. 8, dorsal idiosoma, enlarged section not to scale; 9, ventral idiosoma; 10, subcapitulum; 11, epistome; 12, chelicera." figureDoi="http://doi.org/10.5281/zenodo.6530964" httpUri="https://zenodo.org/record/6530964/files/figure.png" pageId="3" pageNumber="489">Figures 8</figureCitation>
, 
<figureCitation id="C2C87DB7FFA3983D034FAB9E711BF80C" box="[543,574,1987,2013]" captionStart="FIGURES 13–20" captionStartId="5.[151,264,1908,1932]" captionTargetBox="[198,1388,181,1884]" captionTargetId="figure-19@5.[198,1389,181,1884]" captionTargetPageId="5" captionText="FIGURES 13–20. DIC micrographs of Cosmolaelaps lasiophilus Joharchi sp. nov., female. 13, idiosoma in dorsal view; 14, idiosoma in ventral view; 15, general view of some podonotal setae; 16, general view of some opisthonotal setae; 17, general view of part of opisthogastric area; 18, subcapitulum; 19, distal portion of palp, with focus on tarsal claw; 20, insemination structures." figureDoi="http://doi.org/10.5281/zenodo.6530968" httpUri="https://zenodo.org/record/6530968/files/figure.png" pageId="3" pageNumber="489">13</figureCitation>
.
</paragraph>
<caption id="0E8C31BAFFA4983A01C7ABCA7004F805" ID-DOI="http://doi.org/10.5281/zenodo.6530964" ID-Zenodo-Dep="6530964" httpUri="https://zenodo.org/record/6530964/files/figure.png" pageId="4" pageNumber="490" startId="4.[151,264,1943,1967]" targetBox="[181,1397,196,1901]" targetPageId="4">
<paragraph id="5A4C6132FFA4983A01C7ABCA7004F805" blockId="4.[151,1437,1943,2004]" pageId="4" pageNumber="490">
<emphasis id="6887BD20FFA4983A01C7ABCA726FF87E" bold="true" box="[151,330,1943,1967]" pageId="4" pageNumber="490">FIGURES 8–12.</emphasis>
<taxonomicName id="9DF31AB1FFA4983A0004ABC5719EF861" authority="Joharchi" authorityName="Joharchi" authorityYear="2022" box="[340,699,1944,1968]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="490" phylum="Arthropoda" rank="species" species="lasiophilus" status="sp. nov.">
<emphasis id="6887BD20FFA4983A0004ABC5717EF861" box="[340,603,1944,1968]" italics="true" pageId="4" pageNumber="490">Cosmolaelaps lasiophilus</emphasis>
Joharchi
</taxonomicName>
<taxonomicNameLabel id="73B4005BFFA4983A0394ABC5703CF87E" box="[708,793,1944,1968]" pageId="4" pageNumber="490" rank="species">
<emphasis id="6887BD20FFA4983A0394ABC57036F861" bold="true" box="[708,787,1944,1968]" pageId="4" pageNumber="490">sp. nov</emphasis>
.
</taxonomicNameLabel>
, female. 8, dorsal idiosoma, enlarged section not to scale; 9, ventral idiosoma; 10, subcapitulum; 11, epistome; 12, chelicera.
</paragraph>
</caption>
<caption id="0E8C31BAFFA5983B01C7AB297224F829" ID-DOI="http://doi.org/10.5281/zenodo.6530968" ID-Zenodo-Dep="6530968" httpUri="https://zenodo.org/record/6530968/files/figure.png" pageId="5" pageNumber="491" startId="5.[151,264,1908,1932]" targetBox="[198,1388,181,1884]" targetPageId="5">
<paragraph id="5A4C6132FFA5983B01C7AB297224F829" blockId="5.[151,1437,1908,2040]" pageId="5" pageNumber="491">
<emphasis id="6887BD20FFA5983B01C7AB297270F85D" bold="true" box="[151,341,1908,1932]" pageId="5" pageNumber="491">FIGURES 13–20.</emphasis>
DIC micrographs of 
<taxonomicName id="9DF31AB1FFA5983B0362AB2970B3F85D" authority="Joharchi" authorityName="Joharchi" authorityYear="2022" box="[562,918,1908,1932]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="5" pageNumber="491" phylum="Arthropoda" rank="species" species="lasiophilus" status="sp. nov.">
<emphasis id="6887BD20FFA5983B0362AB29701DF85D" box="[562,824,1908,1932]" italics="true" pageId="5" pageNumber="491">Cosmolaelaps lasiophilus</emphasis>
Joharchi
</taxonomicName>
<taxonomicNameLabel id="73B4005BFFA5983B02CDAB2870D5F85D" box="[925,1008,1909,1933]" pageId="5" pageNumber="491" rank="species">
<emphasis id="6887BD20FFA5983B02CDAB2870CFF85C" bold="true" box="[925,1002,1909,1933]" pageId="5" pageNumber="491">sp. nov</emphasis>
.
</taxonomicNameLabel>
, female. 13, idiosoma in dorsal view; 14, idiosoma in ventral view; 15, general view of some podonotal setae; 16, general view of some opisthonotal setae; 17, general view of part of opisthogastric area; 18, subcapitulum; 19, distal portion of palp, with focus on tarsal claw; 20, insemination structures.
</paragraph>
</caption>
<caption id="0E8C31BAFFA6983801C7ABDB7393F813" ID-DOI="http://doi.org/10.5281/zenodo.6530972" ID-Zenodo-Dep="6530972" httpUri="https://zenodo.org/record/6530972/files/figure.png" pageId="6" pageNumber="492" startId="6.[151,250,1926,1950]" targetBox="[161,1425,197,1887]" targetPageId="6">
<paragraph id="5A4C6132FFA6983801C7ABDB7393F813" blockId="6.[151,1436,1926,1986]" pageId="6" pageNumber="492">
<emphasis id="6887BD20FFA6983801C7ABDB7263F84F" bold="true" box="[151,326,1926,1950]" pageId="6" pageNumber="492">FIGURE 21–24.</emphasis>
<taxonomicName id="9DF31AB1FFA69838001CABDB718BF84F" authority="Joharchi" authorityName="Joharchi" authorityYear="2022" box="[332,686,1926,1950]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="492" phylum="Arthropoda" rank="species" species="lasiophilus" status="sp. nov.">
<emphasis id="6887BD20FFA69838001CABDB7174F84F" box="[332,593,1926,1950]" italics="true" pageId="6" pageNumber="492">Cosmolaelaps lasiophilus</emphasis>
Joharchi
</taxonomicName>
<taxonomicNameLabel id="73B4005BFFA6983803E5ABDB7022F84F" box="[693,775,1926,1950]" pageId="6" pageNumber="492" rank="species">
<emphasis id="6887BD20FFA6983803E5ABDB7025F84F" bold="true" box="[693,768,1926,1950]" pageId="6" pageNumber="492">sp. nov</emphasis>
.
</taxonomicNameLabel>
, female. 21, leg I (trochanter-tibia); 22, leg II; 23, leg III; 24, leg IV.
</paragraph>
</caption>
<paragraph id="5A4C6132FFA798390197ACCA7050FBAC" blockId="7.[151,1437,150,2014]" pageId="7" pageNumber="493">
<emphasis id="6887BD20FFA798390197ACCA72AEFF60" box="[199,395,151,177]" italics="true" pageId="7" pageNumber="493">Ventral idiosoma</emphasis>
(
<figureCitation id="C2C87DB7FFA7983900CFACCA72CBFF60" box="[415,494,151,177]" captionStart="FIGURES 8–12" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[181,1397,196,1901]" captionTargetId="figure-18@4.[167,1419,181,1919]" captionTargetPageId="4" captionText="FIGURES 8–12. Cosmolaelaps lasiophilus Joharchi sp. nov., female. 8, dorsal idiosoma, enlarged section not to scale; 9, ventral idiosoma; 10, subcapitulum; 11, epistome; 12, chelicera." figureDoi="http://doi.org/10.5281/zenodo.6530964" httpUri="https://zenodo.org/record/6530964/files/figure.png" pageId="7" pageNumber="493">Figs 9</figureCitation>
, 
<figureCitation id="C2C87DB7FFA7983900AFACCA7168FF60" box="[511,589,151,177]" captionStart="FIGURES 13–20" captionStartId="5.[151,264,1908,1932]" captionTargetBox="[198,1388,181,1884]" captionTargetId="figure-19@5.[198,1389,181,1884]" captionTargetPageId="5" captionText="FIGURES 13–20. DIC micrographs of Cosmolaelaps lasiophilus Joharchi sp. nov., female. 13, idiosoma in dorsal view; 14, idiosoma in ventral view; 15, general view of some podonotal setae; 16, general view of some opisthonotal setae; 17, general view of part of opisthogastric area; 18, subcapitulum; 19, distal portion of palp, with focus on tarsal claw; 20, insemination structures." figureDoi="http://doi.org/10.5281/zenodo.6530968" httpUri="https://zenodo.org/record/6530968/files/figure.png" pageId="7" pageNumber="493">14, 17</figureCitation>
). Tritosternum with paired pilose laciniae (64–68), fused basally (3–5), columnar base 23–27 × 12–15 wide; presternal area lightly sclerotised, with a few transverse curved lines, merged to sternal shield. Sternal shield (length 92–96) narrowest between coxae II (74–80), widest (114–118), anterior margin of shield hardly conspicuous, posterior margin distinctly concave, anterolateral corners narrowly extending between coxae I–II, distally bearing gland pores 
<emphasis id="6887BD20FFA798390331AD7A71ACFE90" box="[609,649,295,321]" italics="true" pageId="7" pageNumber="493">gvb</emphasis>
; endopodal elements between coxae II and III fused with shield and endopodal elements III/IV elongate, narrow and curved. Sternal shield with three pairs of smooth setae (
<emphasis id="6887BD20FFA798390478AD16766DFEB4" box="[1320,1352,331,357]" italics="true" pageId="7" pageNumber="493">st1</emphasis>
31–33, 
<emphasis id="6887BD20FFA7983901C7AD327392FE58" box="[151,183,367,393]" italics="true" pageId="7" pageNumber="493">st2</emphasis>
28–30, 
<emphasis id="6887BD20FFA798390046AD327212FE58" box="[278,311,367,393]" italics="true" pageId="7" pageNumber="493">st3</emphasis>
24–27) and two pairs of slit-shaped pore-like structures (
<emphasis id="6887BD20FFA798390299AD3270C9FE58" box="[969,1004,367,393]" italics="true" pageId="7" pageNumber="493">iv1</emphasis>
adjacent to setae 
<emphasis id="6887BD20FFA7983905E9AD3277FFFE58" box="[1209,1242,367,393]" italics="true" pageId="7" pageNumber="493">st1</emphasis>
; 
<emphasis id="6887BD20FFA7983905BBAD327628FE58" box="[1259,1293,367,393]" italics="true" pageId="7" pageNumber="493">iv2</emphasis>
between 
<emphasis id="6887BD20FFA79839042CAD3276B9FE58" box="[1404,1436,367,393]" italics="true" pageId="7" pageNumber="493">st2</emphasis>
and 
<emphasis id="6887BD20FFA798390197ADCE73CDFE7C" box="[199,232,403,429]" italics="true" pageId="7" pageNumber="493">st3</emphasis>
), setae not reaching base of following setae, shield with lineate-reticulate ornamentation on lateral surface, longitudinally median part smooth (or faintly reticulated) (
<figureCitation id="C2C87DB7FFA79839024DADEA7040FE00" box="[797,869,439,465]" captionStart="FIGURES 8–12" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[181,1397,196,1901]" captionTargetId="figure-18@4.[167,1419,181,1919]" captionTargetPageId="4" captionText="FIGURES 8–12. Cosmolaelaps lasiophilus Joharchi sp. nov., female. 8, dorsal idiosoma, enlarged section not to scale; 9, ventral idiosoma; 10, subcapitulum; 11, epistome; 12, chelicera." figureDoi="http://doi.org/10.5281/zenodo.6530964" httpUri="https://zenodo.org/record/6530964/files/figure.png" pageId="7" pageNumber="493">Figs 9</figureCitation>
, 
<figureCitation id="C2C87DB7FFA79839023FADEA70ABFE00" box="[879,910,439,465]" captionStart="FIGURES 13–20" captionStartId="5.[151,264,1908,1932]" captionTargetBox="[198,1388,181,1884]" captionTargetId="figure-19@5.[198,1389,181,1884]" captionTargetPageId="5" captionText="FIGURES 13–20. DIC micrographs of Cosmolaelaps lasiophilus Joharchi sp. nov., female. 13, idiosoma in dorsal view; 14, idiosoma in ventral view; 15, general view of some podonotal setae; 16, general view of some opisthonotal setae; 17, general view of part of opisthogastric area; 18, subcapitulum; 19, distal portion of palp, with focus on tarsal claw; 20, insemination structures." figureDoi="http://doi.org/10.5281/zenodo.6530968" httpUri="https://zenodo.org/record/6530968/files/figure.png" pageId="7" pageNumber="493">14</figureCitation>
). Metasternal setae 
<emphasis id="6887BD20FFA798390539ADEA77ACFE00" box="[1129,1161,439,465]" italics="true" pageId="7" pageNumber="493">st4</emphasis>
(24–27) and metasternal poroids located on soft integument; metasternal platelets absent. Genital shield flask-shaped, slightly expanded laterally past level of setae 
<emphasis id="6887BD20FFA798390097ADA272C2FDC8" box="[455,487,511,537]" italics="true" pageId="7" pageNumber="493">st5</emphasis>
, length 165–180, maximum width 71–75, posterior margin rounded, anterior hyaline margin of shield irregularly convex, overlapping posterior area of sternal shield (extending beyond base of setae 
<emphasis id="6887BD20FFA7983901C7AE1A739DFDB0" box="[151,184,583,609]" italics="true" pageId="7" pageNumber="493">st3</emphasis>
), shield ornamented posteromedially by 7–8 cells flanked by a median Λ-shaped ornamentation, bearing a pair of simple setae 
<emphasis id="6887BD20FFA798390001AE367254FD54" box="[337,369,619,645]" italics="true" pageId="7" pageNumber="493">st5</emphasis>
(22–24) (
<figureCitation id="C2C87DB7FFA7983900B7AE367116FD54" box="[487,563,619,645]" captionStart="FIGURES 8–12" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[181,1397,196,1901]" captionTargetId="figure-18@4.[167,1419,181,1919]" captionTargetPageId="4" captionText="FIGURES 8–12. Cosmolaelaps lasiophilus Joharchi sp. nov., female. 8, dorsal idiosoma, enlarged section not to scale; 9, ventral idiosoma; 10, subcapitulum; 11, epistome; 12, chelicera." figureDoi="http://doi.org/10.5281/zenodo.6530964" httpUri="https://zenodo.org/record/6530964/files/figure.png" pageId="7" pageNumber="493">Figs 9</figureCitation>
, 
<figureCitation id="C2C87DB7FFA798390313AE367147FD54" box="[579,610,619,645]" captionStart="FIGURES 13–20" captionStartId="5.[151,264,1908,1932]" captionTargetBox="[198,1388,181,1884]" captionTargetId="figure-19@5.[198,1389,181,1884]" captionTargetPageId="5" captionText="FIGURES 13–20. DIC micrographs of Cosmolaelaps lasiophilus Joharchi sp. nov., female. 13, idiosoma in dorsal view; 14, idiosoma in ventral view; 15, general view of some podonotal setae; 16, general view of some opisthonotal setae; 17, general view of part of opisthogastric area; 18, subcapitulum; 19, distal portion of palp, with focus on tarsal claw; 20, insemination structures." figureDoi="http://doi.org/10.5281/zenodo.6530968" httpUri="https://zenodo.org/record/6530968/files/figure.png" pageId="7" pageNumber="493">14</figureCitation>
). Shield flanked by two pairs of minute, elongate paragenital platelets; paragenital poroids 
<emphasis id="6887BD20FFA79839002CAED272BBFD78" box="[380,414,655,681]" italics="true" pageId="7" pageNumber="493">iv5</emphasis>
located on soft cuticle lateral to shield near seta 
<emphasis id="6887BD20FFA79839028FAED27725FD78" box="[991,1024,655,681]" italics="true" pageId="7" pageNumber="493">st5</emphasis>
. Anal shield subtriangular, rounded anteriorly, length 65–70, width 52–55, anterior half lineate-reticulate, para-anal setae and post-anal seta subequal in length (15–18), post-anal seta thicker, cribrum consisting of a terminal tuft with 3–4 irregular rows of spicules, anteriormost rows of spicules extending slightly beyond post-anal seta (
<figureCitation id="C2C87DB7FFA798390294AEA6772BFCC4" box="[964,1038,763,789]" captionStart="FIGURES 8–12" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[181,1397,196,1901]" captionTargetId="figure-18@4.[167,1419,181,1919]" captionTargetPageId="4" captionText="FIGURES 8–12. Cosmolaelaps lasiophilus Joharchi sp. nov., female. 8, dorsal idiosoma, enlarged section not to scale; 9, ventral idiosoma; 10, subcapitulum; 11, epistome; 12, chelicera." figureDoi="http://doi.org/10.5281/zenodo.6530964" httpUri="https://zenodo.org/record/6530964/files/figure.png" pageId="7" pageNumber="493">Figs 9</figureCitation>
, 
<figureCitation id="C2C87DB7FFA79839054BAEA67740FCC4" box="[1051,1125,763,789]" captionStart="FIGURES 13–20" captionStartId="5.[151,264,1908,1932]" captionTargetBox="[198,1388,181,1884]" captionTargetId="figure-19@5.[198,1389,181,1884]" captionTargetPageId="5" captionText="FIGURES 13–20. DIC micrographs of Cosmolaelaps lasiophilus Joharchi sp. nov., female. 13, idiosoma in dorsal view; 14, idiosoma in ventral view; 15, general view of some podonotal setae; 16, general view of some opisthonotal setae; 17, general view of part of opisthogastric area; 18, subcapitulum; 19, distal portion of palp, with focus on tarsal claw; 20, insemination structures." figureDoi="http://doi.org/10.5281/zenodo.6530968" httpUri="https://zenodo.org/record/6530968/files/figure.png" pageId="7" pageNumber="493">14, 17</figureCitation>
); anal gland pores (
<emphasis id="6887BD20FFA798390416AEA6764AFCC4" box="[1350,1391,763,789]" italics="true" pageId="7" pageNumber="493">gv3</emphasis>
) on anterolateral margin of anal shield. Soft opisthogastric cuticle surrounding genital and anal shields with one pair of suboval metapodal plates (17–19 long × 4–6 wide), six pairs of poroids (five 
<emphasis id="6887BD20FFA798390556AF1E770DFC8D" box="[1030,1064,835,860]" italics="true" pageId="7" pageNumber="493">ivo</emphasis>
; 
<emphasis id="6887BD20FFA798390568AF1E777FFC8D" box="[1080,1114,835,860]" italics="true" pageId="7" pageNumber="493">ivp</emphasis>
) and 16 pairs of moderately thick setae (
<emphasis id="6887BD20FFA798390075AF3A72A4FC50" box="[293,385,871,897]" italics="true" pageId="7" pageNumber="493">Jv1–Jv5</emphasis>
, 
<emphasis id="6887BD20FFA7983900C2AF3A72D5FC51" box="[402,496,871,897]" italics="true" pageId="7" pageNumber="493">Zv1– Zv</emphasis>
5, 
<emphasis id="6887BD20FFA79839035FAF3A7179FC50" box="[527,604,871,897]" italics="true" pageId="7" pageNumber="493">R1–R4</emphasis>
, 
<emphasis id="6887BD20FFA79839033DAF3A71C4FC50" box="[621,737,871,897]" italics="true" pageId="7" pageNumber="493">UR2–UR3</emphasis>
); almost all setae uniform in length (11–16), and arising on small sclerotised platelet, except 
<emphasis id="6887BD20FFA798390340AFD67176FC74" box="[528,595,907,933]" italics="true" pageId="7" pageNumber="493">Jv1–3</emphasis>
and 
<emphasis id="6887BD20FFA7983903DDAFD67192FC74" box="[653,695,907,933]" italics="true" pageId="7" pageNumber="493">Zv1</emphasis>
longer (20–24) and setiform (
<figureCitation id="C2C87DB7FFA798390540AFD6777FFC74" box="[1040,1114,907,933]" captionStart="FIGURES 8–12" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[181,1397,196,1901]" captionTargetId="figure-18@4.[167,1419,181,1919]" captionTargetPageId="4" captionText="FIGURES 8–12. Cosmolaelaps lasiophilus Joharchi sp. nov., female. 8, dorsal idiosoma, enlarged section not to scale; 9, ventral idiosoma; 10, subcapitulum; 11, epistome; 12, chelicera." figureDoi="http://doi.org/10.5281/zenodo.6530964" httpUri="https://zenodo.org/record/6530964/files/figure.png" pageId="7" pageNumber="493">Figs 9</figureCitation>
, 
<figureCitation id="C2C87DB7FFA798390537AFD67797FC74" box="[1127,1202,907,933]" captionStart="FIGURES 13–20" captionStartId="5.[151,264,1908,1932]" captionTargetBox="[198,1388,181,1884]" captionTargetId="figure-19@5.[198,1389,181,1884]" captionTargetPageId="5" captionText="FIGURES 13–20. DIC micrographs of Cosmolaelaps lasiophilus Joharchi sp. nov., female. 13, idiosoma in dorsal view; 14, idiosoma in ventral view; 15, general view of some podonotal setae; 16, general view of some opisthonotal setae; 17, general view of part of opisthogastric area; 18, subcapitulum; 19, distal portion of palp, with focus on tarsal claw; 20, insemination structures." figureDoi="http://doi.org/10.5281/zenodo.6530968" httpUri="https://zenodo.org/record/6530968/files/figure.png" pageId="7" pageNumber="493">14, 17</figureCitation>
). Peritrematal shield well developed laterad peritremes, bearing one gland pore and one poroid at level near coxae II–III, post-stigmatic extension of shield relatively wide, not reaching posterior level of coxae IV, with two pairs of poroids and one pair of gland pores; anterior region of peritrematal shield fused to anterior margin of dorsal shield behind setae 
<emphasis id="6887BD20FFA7983901C7A8467395FBE4" box="[151,176,1051,1077]" italics="true" pageId="7" pageNumber="493">z1</emphasis>
. Peritremes long, extending to mid-level of coxa I (near 
<emphasis id="6887BD20FFA79839026DA8467072FBE4" box="[829,855,1051,1077]" italics="true" pageId="7" pageNumber="493">s1</emphasis>
). Two exopodal platelets between coxae II–III and III–IV present, anterior platelet small and subtriangular, those between coxae III–IV slightly extending posteriorly; parapodal platelets strip-like, bearing gland pore 
<emphasis id="6887BD20FFA7983903EDA83E71C0FBAC" box="[701,741,1123,1149]" italics="true" pageId="7" pageNumber="493">gv2</emphasis>
(
<figureCitation id="C2C87DB7FFA7983903A4A83E7018FBAC" box="[756,829,1123,1149]" captionStart="FIGURES 8–12" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[181,1397,196,1901]" captionTargetId="figure-18@4.[167,1419,181,1919]" captionTargetPageId="4" captionText="FIGURES 8–12. Cosmolaelaps lasiophilus Joharchi sp. nov., female. 8, dorsal idiosoma, enlarged section not to scale; 9, ventral idiosoma; 10, subcapitulum; 11, epistome; 12, chelicera." figureDoi="http://doi.org/10.5281/zenodo.6530964" httpUri="https://zenodo.org/record/6530964/files/figure.png" pageId="7" pageNumber="493">Figs 9</figureCitation>
, 
<figureCitation id="C2C87DB7FFA798390219A83E704DFBAC" box="[841,872,1123,1149]" captionStart="FIGURES 13–20" captionStartId="5.[151,264,1908,1932]" captionTargetBox="[198,1388,181,1884]" captionTargetId="figure-19@5.[198,1389,181,1884]" captionTargetPageId="5" captionText="FIGURES 13–20. DIC micrographs of Cosmolaelaps lasiophilus Joharchi sp. nov., female. 13, idiosoma in dorsal view; 14, idiosoma in ventral view; 15, general view of some podonotal setae; 16, general view of some opisthonotal setae; 17, general view of part of opisthogastric area; 18, subcapitulum; 19, distal portion of palp, with focus on tarsal claw; 20, insemination structures." figureDoi="http://doi.org/10.5281/zenodo.6530968" httpUri="https://zenodo.org/record/6530968/files/figure.png" pageId="7" pageNumber="493">14</figureCitation>
).
</paragraph>
<paragraph id="5A4C6132FFA798390197A8DA779AF9FC" blockId="7.[151,1437,150,2014]" pageId="7" pageNumber="493">
<taxonomicName id="9DF31AB1FFA798390197A8DA7273FB70" authorityName="Krefft" authorityYear="1871" box="[199,342,1159,1185]" class="Chromadorea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="7" pageNumber="493" phylum="Nematoda" rank="genus">
<emphasis id="6887BD20FFA798390197A8DA7273FB70" box="[199,342,1159,1185]" italics="true" pageId="7" pageNumber="493">Gnathosoma</emphasis>
</taxonomicName>
(
<figureCitation id="C2C87DB7FFA798390037A8DA72CFFB70" box="[359,490,1159,1185]" captionStart="FIGURES 8–12" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[181,1397,196,1901]" captionTargetId="figure-18@4.[167,1419,181,1919]" captionTargetPageId="4" captionText="FIGURES 8–12. Cosmolaelaps lasiophilus Joharchi sp. nov., female. 8, dorsal idiosoma, enlarged section not to scale; 9, ventral idiosoma; 10, subcapitulum; 11, epistome; 12, chelicera." figureDoi="http://doi.org/10.5281/zenodo.6530964" httpUri="https://zenodo.org/record/6530964/files/figure.png" pageId="7" pageNumber="493">Figs 10–12</figureCitation>
, 
<figureCitation id="C2C87DB7FFA7983900A7A8DA7167FB70" box="[503,578,1159,1185]" captionStart="FIGURES 13–20" captionStartId="5.[151,264,1908,1932]" captionTargetBox="[198,1388,181,1884]" captionTargetId="figure-19@5.[198,1389,181,1884]" captionTargetPageId="5" captionText="FIGURES 13–20. DIC micrographs of Cosmolaelaps lasiophilus Joharchi sp. nov., female. 13, idiosoma in dorsal view; 14, idiosoma in ventral view; 15, general view of some podonotal setae; 16, general view of some opisthonotal setae; 17, general view of part of opisthogastric area; 18, subcapitulum; 19, distal portion of palp, with focus on tarsal claw; 20, insemination structures." figureDoi="http://doi.org/10.5281/zenodo.6530968" httpUri="https://zenodo.org/record/6530968/files/figure.png" pageId="7" pageNumber="493">18, 19</figureCitation>
). Anterior margin of epistome convex, irregularly denticulate, with smooth median spike (
<figureCitation id="C2C87DB7FFA798390015A8F672BEFB14" box="[325,411,1195,1221]" captionStart="FIGURES 8–12" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[181,1397,196,1901]" captionTargetId="figure-18@4.[167,1419,181,1919]" captionTargetPageId="4" captionText="FIGURES 8–12. Cosmolaelaps lasiophilus Joharchi sp. nov., female. 8, dorsal idiosoma, enlarged section not to scale; 9, ventral idiosoma; 10, subcapitulum; 11, epistome; 12, chelicera." figureDoi="http://doi.org/10.5281/zenodo.6530964" httpUri="https://zenodo.org/record/6530964/files/figure.png" pageId="7" pageNumber="493">Fig. 11</figureCitation>
). Hypostomal groove with six transverse rows of denticles, each row with 8–19 small denticles, with smooth posterior transverse line, groove wider anteriorly, progressively narrowing from anterior to posterior, anteriormost row slightly convex (
<figureCitation id="C2C87DB7FFA7983903DAA8AE71C5FADC" box="[650,736,1267,1293]" captionStart="FIGURES 8–12" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[181,1397,196,1901]" captionTargetId="figure-18@4.[167,1419,181,1919]" captionTargetPageId="4" captionText="FIGURES 8–12. Cosmolaelaps lasiophilus Joharchi sp. nov., female. 8, dorsal idiosoma, enlarged section not to scale; 9, ventral idiosoma; 10, subcapitulum; 11, epistome; 12, chelicera." figureDoi="http://doi.org/10.5281/zenodo.6530964" httpUri="https://zenodo.org/record/6530964/files/figure.png" pageId="7" pageNumber="493">Figs 10</figureCitation>
, 
<figureCitation id="C2C87DB7FFA7983903BCA8AE702EFADC" box="[748,779,1267,1293]" captionStart="FIGURES 13–20" captionStartId="5.[151,264,1908,1932]" captionTargetBox="[198,1388,181,1884]" captionTargetId="figure-19@5.[198,1389,181,1884]" captionTargetPageId="5" captionText="FIGURES 13–20. DIC micrographs of Cosmolaelaps lasiophilus Joharchi sp. nov., female. 13, idiosoma in dorsal view; 14, idiosoma in ventral view; 15, general view of some podonotal setae; 16, general view of some opisthonotal setae; 17, general view of part of opisthogastric area; 18, subcapitulum; 19, distal portion of palp, with focus on tarsal claw; 20, insemination structures." figureDoi="http://doi.org/10.5281/zenodo.6530968" httpUri="https://zenodo.org/record/6530968/files/figure.png" pageId="7" pageNumber="493">18</figureCitation>
). Corniculi horn-like, internal malae longer than corniculi, comprising a pair of pilose median projections, and 7–9 lateral, longer, tentacle-like, smooth projections (
<figureCitation id="C2C87DB7FFA79839046AA94A76ABFAE0" box="[1338,1422,1303,1329]" captionStart="FIGURES 8–12" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[181,1397,196,1901]" captionTargetId="figure-18@4.[167,1419,181,1919]" captionTargetPageId="4" captionText="FIGURES 8–12. Cosmolaelaps lasiophilus Joharchi sp. nov., female. 8, dorsal idiosoma, enlarged section not to scale; 9, ventral idiosoma; 10, subcapitulum; 11, epistome; 12, chelicera." figureDoi="http://doi.org/10.5281/zenodo.6530964" httpUri="https://zenodo.org/record/6530964/files/figure.png" pageId="7" pageNumber="493">Fig. 10</figureCitation>
); labrum acuminate, considerably longer than internal malae, fringed; hypostomal and capitular setae smooth, 
<emphasis id="6887BD20FFA7983904D0A96676B9FA84" box="[1408,1436,1339,1365]" italics="true" pageId="7" pageNumber="493">h3</emphasis>
(34–40)&gt; 
<emphasis id="6887BD20FFA79839005EA902720FFAA8" box="[270,298,1375,1401]" italics="true" pageId="7" pageNumber="493">h1</emphasis>
(28–31)&gt; 
<emphasis id="6887BD20FFA7983900F8A90272E7FAA9" box="[424,450,1375,1400]" italics="true" pageId="7" pageNumber="493">pc</emphasis>
(21–25)&gt; 
<emphasis id="6887BD20FFA798390310A9027179FAA8" box="[576,604,1375,1401]" italics="true" pageId="7" pageNumber="493">h2</emphasis>
(11–14) (
<figureCitation id="C2C87DB7FFA798390399A9027005FAA8" box="[713,800,1375,1401]" captionStart="FIGURES 8–12" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[181,1397,196,1901]" captionTargetId="figure-18@4.[167,1419,181,1919]" captionTargetPageId="4" captionText="FIGURES 8–12. Cosmolaelaps lasiophilus Joharchi sp. nov., female. 8, dorsal idiosoma, enlarged section not to scale; 9, ventral idiosoma; 10, subcapitulum; 11, epistome; 12, chelicera." figureDoi="http://doi.org/10.5281/zenodo.6530964" httpUri="https://zenodo.org/record/6530964/files/figure.png" pageId="7" pageNumber="493">Figs 10</figureCitation>
, 
<figureCitation id="C2C87DB7FFA79839027CA902706EFAA8" box="[812,843,1375,1401]" captionStart="FIGURES 13–20" captionStartId="5.[151,264,1908,1932]" captionTargetBox="[198,1388,181,1884]" captionTargetId="figure-19@5.[198,1389,181,1884]" captionTargetPageId="5" captionText="FIGURES 13–20. DIC micrographs of Cosmolaelaps lasiophilus Joharchi sp. nov., female. 13, idiosoma in dorsal view; 14, idiosoma in ventral view; 15, general view of some podonotal setae; 16, general view of some opisthonotal setae; 17, general view of part of opisthogastric area; 18, subcapitulum; 19, distal portion of palp, with focus on tarsal claw; 20, insemination structures." figureDoi="http://doi.org/10.5281/zenodo.6530968" httpUri="https://zenodo.org/record/6530968/files/figure.png" pageId="7" pageNumber="493">18</figureCitation>
). Chaetotaxy of palps: trochanter 2, femur 5, genu 6, tibia 14, tarsus 15, all setae smooth except palpfemur with seta 
<emphasis id="6887BD20FFA798390208A9DE7051FA4C" box="[856,884,1411,1437]" italics="true" pageId="7" pageNumber="493">d3</emphasis>
thickened and inserted on small tubercles, seta 
<emphasis id="6887BD20FFA7983904D6A9DE76B9FA4C" box="[1414,1436,1411,1437]" italics="true" pageId="7" pageNumber="493">al</emphasis>
thickened and apically subspatulate; palpgenu with 
<emphasis id="6887BD20FFA79839038EA9FA7027FA10" box="[734,770,1447,1473]" italics="true" pageId="7" pageNumber="493">al1</emphasis>
stout, 
<emphasis id="6887BD20FFA79839021EA9FA7057FA10" box="[846,882,1447,1473]" italics="true" pageId="7" pageNumber="493">al2</emphasis>
paddle-like; palp tarsal claw two-tined (
<figureCitation id="C2C87DB7FFA79839046BA9FA76AAFA10" box="[1339,1423,1447,1473]" captionStart="FIGURES 13–20" captionStartId="5.[151,264,1908,1932]" captionTargetBox="[198,1388,181,1884]" captionTargetId="figure-19@5.[198,1389,181,1884]" captionTargetPageId="5" captionText="FIGURES 13–20. DIC micrographs of Cosmolaelaps lasiophilus Joharchi sp. nov., female. 13, idiosoma in dorsal view; 14, idiosoma in ventral view; 15, general view of some podonotal setae; 16, general view of some opisthonotal setae; 17, general view of part of opisthogastric area; 18, subcapitulum; 19, distal portion of palp, with focus on tarsal claw; 20, insemination structures." figureDoi="http://doi.org/10.5281/zenodo.6530968" httpUri="https://zenodo.org/record/6530968/files/figure.png" pageId="7" pageNumber="493">Fig. 19</figureCitation>
). Supralabral process indistinct. Fixed digit of chelicera with an offset and subapical tooth (gabelzhan), followed by five teeth, a setaceous pilus dentilis, dorsal cheliceral seta prostrate, arthrodial membrane with a rounded flap and normal filaments; cheliceral lyrifissures distinct, movable digit of chelicera bidentate (
<figureCitation id="C2C87DB7FFA79839050EAA4E7797F9FC" box="[1118,1202,1555,1581]" captionStart="FIGURES 8–12" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[181,1397,196,1901]" captionTargetId="figure-18@4.[167,1419,181,1919]" captionTargetPageId="4" captionText="FIGURES 8–12. Cosmolaelaps lasiophilus Joharchi sp. nov., female. 8, dorsal idiosoma, enlarged section not to scale; 9, ventral idiosoma; 10, subcapitulum; 11, epistome; 12, chelicera." figureDoi="http://doi.org/10.5281/zenodo.6530964" httpUri="https://zenodo.org/record/6530964/files/figure.png" pageId="7" pageNumber="493">Fig. 12</figureCitation>
).
</paragraph>
<paragraph id="5A4C6132FFA798390197AA6A7613F948" blockId="7.[151,1437,150,2014]" pageId="7" pageNumber="493">
<emphasis id="6887BD20FFA798390197AA6A72F5F981" box="[199,464,1591,1616]" italics="true" pageId="7" pageNumber="493">Insemination structures</emphasis>
(
<figureCitation id="C2C87DB7FFA7983900B1AA6A7113F980" box="[481,566,1591,1617]" captionStart="FIGURES 13–20" captionStartId="5.[151,264,1908,1932]" captionTargetBox="[198,1388,181,1884]" captionTargetId="figure-19@5.[198,1389,181,1884]" captionTargetPageId="5" captionText="FIGURES 13–20. DIC micrographs of Cosmolaelaps lasiophilus Joharchi sp. nov., female. 13, idiosoma in dorsal view; 14, idiosoma in ventral view; 15, general view of some podonotal setae; 16, general view of some opisthonotal setae; 17, general view of part of opisthogastric area; 18, subcapitulum; 19, distal portion of palp, with focus on tarsal claw; 20, insemination structures." figureDoi="http://doi.org/10.5281/zenodo.6530968" httpUri="https://zenodo.org/record/6530968/files/figure.png" pageId="7" pageNumber="493">Fig. 20</figureCitation>
). Laelapid-type sperm access system, tubulus long, wider at the solenostome level of coxa III and entering sacculus via a pair of circular openings. Sacculus irregular, the proximal ends of the tubulus slightly swollen at the junction with ramus, ends of ramus with considerably horn-like projection.
</paragraph>
<paragraph id="5A4C6132FFA798360197AAFE7677FECC" blockId="7.[151,1437,150,2014]" lastBlockId="8.[151,1437,151,1725]" lastPageId="8" lastPageNumber="494" pageId="7" pageNumber="493">
<emphasis id="6887BD20FFA798390197AAFE73D9F96D" box="[199,252,1699,1724]" italics="true" pageId="7" pageNumber="493">Legs</emphasis>
(
<figureCitation id="C2C87DB7FFA79839005CAAFE72AAF96C" box="[268,399,1699,1725]" captionStart="FIGURE 21–24" captionStartId="6.[151,250,1926,1950]" captionTargetBox="[161,1425,197,1887]" captionTargetId="figure-18@6.[152,1436,181,1902]" captionTargetPageId="6" captionText="FIGURE 21–24. Cosmolaelaps lasiophilus Joharchi sp. nov., female. 21, leg I (trochanter-tibia); 22, leg II; 23, leg III; 24, leg IV." figureDoi="http://doi.org/10.5281/zenodo.6530972" httpUri="https://zenodo.org/record/6530972/files/figure.png" pageId="7" pageNumber="493">Figs 21–24</figureCitation>
). Legs II and III short (260–278, 240–255), I and IV longer (365–385, 347–364) (excluding pre-tarsus). Chaetotaxy normal for free-living 
<taxonomicName id="9DF31AB1FFA7983903F0AA9A703EF930" box="[672,795,1735,1761]" class="Arachnida" family="Laelapidae" kingdom="Animalia" order="Mesostigmata" pageId="7" pageNumber="493" phylum="Arthropoda" rank="family">Laelapidae</taxonomicName>
(
<emphasis id="6887BD20FFA79839027BAA9A704CF931" box="[811,873,1735,1760]" italics="true" pageId="7" pageNumber="493">sensu</emphasis>
<bibRefCitation id="3E621CC3FFA798390221AA9A7729F933" author="Evans, G. O." box="[881,1036,1735,1762]" pageId="7" pageNumber="493" pagination="275 - 303" refId="ref14188" refString="Evans, G. O. (1963 a) Observations on the chaetotaxy of the legs in the free-living Gamasina (Acari: Mesostigmata). Bulletin of the British Museum (Natural History) Zoology, 10 (5), 275 - 303. https: // doi. org / 10.5962 / bhl. part. 20528" type="journal article" year="1963">Evans, 1963a</bibRefCitation>
): Leg I (
<figureCitation id="C2C87DB7FFA79839053CAA9A77E5F930" box="[1132,1216,1735,1761]" captionStart="FIGURE 21–24" captionStartId="6.[151,250,1926,1950]" captionTargetBox="[161,1425,197,1887]" captionTargetId="figure-18@6.[152,1436,181,1902]" captionTargetPageId="6" captionText="FIGURE 21–24. Cosmolaelaps lasiophilus Joharchi sp. nov., female. 21, leg I (trochanter-tibia); 22, leg II; 23, leg III; 24, leg IV." figureDoi="http://doi.org/10.5281/zenodo.6530972" httpUri="https://zenodo.org/record/6530972/files/figure.png" pageId="7" pageNumber="493">Fig. 21</figureCitation>
): coxa 0-0/1, 0/1-0, trochanter 1-1/1, 0/2-1 (
<emphasis id="6887BD20FFA7983900FCAAB672E7F8D4" box="[428,450,1771,1797]" italics="true" pageId="7" pageNumber="493">al</emphasis>
and 
<emphasis id="6887BD20FFA7983900ADAAB6713CF8D4" box="[509,537,1771,1797]" italics="true" pageId="7" pageNumber="493">ad</emphasis>
thickened), femur 2-3/1, 2/3-2 (
<emphasis id="6887BD20FFA7983902DFAAB67091F8D4" box="[911,948,1771,1797]" italics="true" pageId="7" pageNumber="493">al1</emphasis>
, 
<emphasis id="6887BD20FFA798390294AAB670CBF8D4" box="[964,1006,1771,1797]" italics="true" pageId="7" pageNumber="493">ad3</emphasis>
and 
<emphasis id="6887BD20FFA798390579AAB67776F8D4" box="[1065,1107,1771,1797]" italics="true" pageId="7" pageNumber="493">pd2</emphasis>
thickened), genu 2-3/2, 3/1- 2 (dorsals and laterals thickened and 
<emphasis id="6887BD20FFA798390319AB5271B5F8F8" box="[585,656,1807,1833]" italics="true" pageId="7" pageNumber="493">ad1–3</emphasis>
inserted on small tubercles), tibia 2-3/2, 3/1-2 (dorsals and laterals thickened). Leg II (
<figureCitation id="C2C87DB7FFA798390026AB6E72E9F89C" box="[374,460,1843,1869]" captionStart="FIGURE 21–24" captionStartId="6.[151,250,1926,1950]" captionTargetBox="[161,1425,197,1887]" captionTargetId="figure-18@6.[152,1436,181,1902]" captionTargetPageId="6" captionText="FIGURE 21–24. Cosmolaelaps lasiophilus Joharchi sp. nov., female. 21, leg I (trochanter-tibia); 22, leg II; 23, leg III; 24, leg IV." figureDoi="http://doi.org/10.5281/zenodo.6530972" httpUri="https://zenodo.org/record/6530972/files/figure.png" pageId="7" pageNumber="493">Fig. 22</figureCitation>
): coxa 0-0/1, 0/1-0, trochanter 1-0/1, 0/2-1 (
<emphasis id="6887BD20FFA798390299AB6E70FAF89C" box="[969,991,1843,1869]" italics="true" pageId="7" pageNumber="493">al</emphasis>
slightly thickened), femur 2-3/1, 2/2-1 (
<emphasis id="6887BD20FFA7983901F1AB0A73EFF8A0" box="[161,202,1879,1905]" italics="true" pageId="7" pageNumber="493">ad1</emphasis>
, 
<emphasis id="6887BD20FFA798390188AB0A7227F8A0" box="[216,258,1879,1905]" italics="true" pageId="7" pageNumber="493">pd1</emphasis>
and 
<emphasis id="6887BD20FFA798390068AB0A7244F8A0" box="[312,353,1879,1905]" italics="true" pageId="7" pageNumber="493">pd2</emphasis>
thickened and inserted on small tubercles, 
<emphasis id="6887BD20FFA798390212AB0A7049F8A0" box="[834,876,1879,1905]" italics="true" pageId="7" pageNumber="493">pd2</emphasis>
longest), genu 2-3/1, 2/1-2 (dorsals thickened and inserted on small tubercles), tibia 2-2/1, 2/1-2. Leg III (
<figureCitation id="C2C87DB7FFA7983903A2AB267060F844" box="[754,837,1915,1941]" captionStart="FIGURE 21–24" captionStartId="6.[151,250,1926,1950]" captionTargetBox="[161,1425,197,1887]" captionTargetId="figure-18@6.[152,1436,181,1902]" captionTargetPageId="6" captionText="FIGURE 21–24. Cosmolaelaps lasiophilus Joharchi sp. nov., female. 21, leg I (trochanter-tibia); 22, leg II; 23, leg III; 24, leg IV." figureDoi="http://doi.org/10.5281/zenodo.6530972" httpUri="https://zenodo.org/record/6530972/files/figure.png" pageId="7" pageNumber="493">Fig. 23</figureCitation>
): coxa 0-0/1, 0/1-0, trochanter 1-1/1, 0/2-0 (
<emphasis id="6887BD20FFA798390477AB267666F844" box="[1319,1347,1915,1941]" italics="true" pageId="7" pageNumber="493">ad</emphasis>
slightly thickened), femur 1-2/1, 1/0-1 (
<emphasis id="6887BD20FFA7983900A5ABC2713AF868" box="[501,543,1951,1977]" italics="true" pageId="7" pageNumber="493">ad1</emphasis>
, 
<emphasis id="6887BD20FFA79839037CABC2716DF868" box="[556,584,1951,1977]" italics="true" pageId="7" pageNumber="493">pd</emphasis>
and 
<emphasis id="6887BD20FFA79839032CABC271B7F868" box="[636,658,1951,1977]" italics="true" pageId="7" pageNumber="493">pl</emphasis>
thickened, 
<emphasis id="6887BD20FFA798390241ABC2701EF868" box="[785,827,1951,1977]" italics="true" pageId="7" pageNumber="493">ad1</emphasis>
inserted on small tubercle), genu 2-2/1, 2/1-1 (all setae on segment slightly thickened), tibia: 2-1/1, 2/1-1 (all setae on segment slightly thickened). Leg IV (
<figureCitation id="C2C87DB7FFA7983905AEAB9E7676F80C" box="[1278,1363,1987,2013]" captionStart="FIGURE 21–24" captionStartId="6.[151,250,1926,1950]" captionTargetBox="[161,1425,197,1887]" captionTargetId="figure-18@6.[152,1436,181,1902]" captionTargetPageId="6" captionText="FIGURE 21–24. Cosmolaelaps lasiophilus Joharchi sp. nov., female. 21, leg I (trochanter-tibia); 22, leg II; 23, leg III; 24, leg IV." figureDoi="http://doi.org/10.5281/zenodo.6530972" httpUri="https://zenodo.org/record/6530972/files/figure.png" pageId="7" pageNumber="493">Fig. 24</figureCitation>
): coxa 0-0/1, 0/0-0, trochanter 1-1/1, 0/2-0 (
<emphasis id="6887BD20FFA898360361ACCA7168FF60" box="[561,589,151,177]" italics="true" pageId="8" pageNumber="494">ad</emphasis>
thickened), femur 1-2/1, 1/0-1 (
<emphasis id="6887BD20FFA8983602E0ACCA70FFFF60" box="[944,986,151,177]" italics="true" pageId="8" pageNumber="494">ad1</emphasis>
and 
<emphasis id="6887BD20FFA89836055EACCA771CFF60" box="[1038,1081,151,177]" italics="true" pageId="8" pageNumber="494">ad2</emphasis>
thickened and inserted on small tubercles), genu 2-2/1, 3/0-1 (dorsals thickened), tibia 2-1/1, 3/1-2 (dorsals and laterals thickened, 
<emphasis id="6887BD20FFA8983605A4ACE67632FF04" box="[1268,1303,187,213]" italics="true" pageId="8" pageNumber="494">al2</emphasis>
inserted on small tubercle). Tarsi II–IV with 18 setae (3- 3/2, 3/2-3 + 
<emphasis id="6887BD20FFA898360278AC82706CFF29" box="[808,841,223,248]" italics="true" pageId="8" pageNumber="494">mv</emphasis>
, 
<emphasis id="6887BD20FFA898360207AC82705CFF28" box="[855,889,223,249]" italics="true" pageId="8" pageNumber="494">md</emphasis>
); almost all setae on tarsus II–IV thickened, see 
<figureCitation id="C2C87DB7FFA8983601C7AD5E7219FECC" box="[151,316,259,285]" captionStart="FIGURE 21–24" captionStartId="6.[151,250,1926,1950]" captionTargetBox="[161,1425,197,1887]" captionTargetId="figure-18@6.[152,1436,181,1902]" captionTargetPageId="6" captionText="FIGURE 21–24. Cosmolaelaps lasiophilus Joharchi sp. nov., female. 21, leg I (trochanter-tibia); 22, leg II; 23, leg III; 24, leg IV." figureDoi="http://doi.org/10.5281/zenodo.6530972" httpUri="https://zenodo.org/record/6530972/files/figure.png" pageId="8" pageNumber="494">Figures 22–24</figureCitation>
. All pretarsi with well-developed paired claws, rounded pulvilli and normal ambulacral stalk.
</paragraph>
<paragraph id="5A4C6132FFA898360197AD7A72ADFE90" blockId="8.[151,1437,151,1725]" box="[199,392,295,321]" pageId="8" pageNumber="494">
<emphasis id="6887BD20FFA898360197AD7A722EFE90" bold="true" box="[199,267,295,321]" pageId="8" pageNumber="494">Male.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="12E932B9FFA898360197AD1670BEFE58" pageId="8" pageNumber="494" type="etymology">
<paragraph id="5A4C6132FFA898360197AD1670BEFE58" blockId="8.[151,1437,151,1725]" pageId="8" pageNumber="494">
<emphasis id="6887BD20FFA898360197AD16726DFEB4" bold="true" box="[199,328,331,357]" pageId="8" pageNumber="494">Etymology</emphasis>
. The specific name of the new species is derived from its occurrence in nests of ants of the genus 
<taxonomicName id="9DF31AB1FFA8983601C7AD3273C5FE59" authorityName="Fabricius" authorityYear="1804" box="[151,224,367,392]" class="Insecta" family="Formicidae" genus="Lasius" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="494" phylum="Arthropoda" rank="genus">
<emphasis id="6887BD20FFA8983601C7AD3273C5FE59" box="[151,224,367,392]" italics="true" pageId="8" pageNumber="494">Lasius</emphasis>
</taxonomicName>
and Greek 
<emphasis id="6887BD20FFA898360032AD3372B9FE58" box="[354,412,366,393]" italics="true" pageId="8" pageNumber="494">ΦΙΛία</emphasis>
(philia), meaning &quot;friendship&quot; or &quot;fondness&quot;.
</paragraph>
</subSubSection>
<subSubSection id="12E932B9FFA898360197ADCF70F6F96C" pageId="8" pageNumber="494" type="diagnosis">
<paragraph id="5A4C6132FFA898360197ADCF762AFDB0" blockId="8.[151,1437,151,1725]" pageId="8" pageNumber="494">
<emphasis id="6887BD20FFA898360197ADCF72F5FE7D" bold="true" box="[199,464,402,428]" pageId="8" pageNumber="494">Differential diagnosis.</emphasis>
<taxonomicName id="9DF31AB1FFA89836008AADCE71D8FE7C" authorityName="Joharchi" authorityYear="2022" box="[474,765,403,429]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="494" phylum="Arthropoda" rank="species" species="lasiophilus">
<emphasis id="6887BD20FFA89836008AADCE71D8FE7C" box="[474,765,403,429]" italics="true" pageId="8" pageNumber="494">Cosmolaelaps lasiophilus</emphasis>
</taxonomicName>
is closely related to 
<taxonomicName id="9DF31AB1FFA8983602A1ADCE72E4FE03" authority="(Michael, 1891) sensu Evans &amp; Till (1966)" authorityName="sensu Evans &amp; Till" authorityYear="1966" baseAuthorityName="Michael" baseAuthorityYear="1891" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="494" phylum="Arthropoda" rank="species" species="vacua">
<emphasis id="6887BD20FFA8983602A1ADCE77F9FE7D" box="[1009,1244,403,429]" italics="true" pageId="8" pageNumber="494">Cosmolaelaps vacua</emphasis>
(
<bibRefCitation id="3E621CC3FFA8983605BDADCE76B1FE7C" author="Michael, A. D." box="[1261,1428,403,429]" pageId="8" pageNumber="494" pagination="638 - 654" refId="ref15960" refString="Michael, A. D. (1891) On the association of gamasids with ants. Proceedings of the Zoological Society of London, 4, 638 - 654." type="journal article" year="1891">Michael, 1891</bibRefCitation>
) 
<emphasis id="6887BD20FFA8983601C7ADEA73F0FE01" box="[151,213,439,464]" italics="true" pageId="8" pageNumber="494">sensu</emphasis>
<bibRefCitation id="3E621CC3FFA89836018DADEA72E4FE03" author="Evans, G. O. &amp; Till, W. M." box="[221,449,439,466]" pageId="8" pageNumber="494" pagination="107 - 370" refId="ref14300" refString="Evans, G. O. &amp; Till, W. M. (1966) Studies on the British Dermanyssidae (Acari: Mesostigmata). Part II. Classification. Bulletin of the British Museum (Natural History) Zoology, 14, 107 - 370." type="journal article" year="1966">Evans &amp; Till (1966)</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="9DF31AB1FFA89836009DADEA71D9FE00" authority="Zeman, 1982" authorityName="Zeman" authorityYear="1982" box="[461,764,439,465]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="494" phylum="Arthropoda" rank="species" species="thysanifer">
<emphasis id="6887BD20FFA89836009DADEA72C5FE00" box="[461,480,439,465]" italics="true" pageId="8" pageNumber="494">C</emphasis>
. 
<emphasis id="6887BD20FFA8983600BEADEA717BFE00" box="[494,606,439,465]" italics="true" pageId="8" pageNumber="494">thysanifer</emphasis>
<bibRefCitation id="3E621CC3FFA898360335ADEA71D9FE00" author="Zeman, P." box="[613,764,439,465]" pageId="8" pageNumber="494" pagination="231 - 237" refId="ref16896" refString="Zeman, P. (1982) Two new species of Hypoaspidinae (Acari: Mesostigmata: Dermanysside) associated with ants. Vestnik Ceskoslovenske Spolecnosti Zoologicke, 46, 231 - 237." type="journal article" year="1982">Zeman, 1982</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="9DF31AB1FFA898360258ADEA7770FE00" authority="Ryke, 1963" authorityName="Ryke" authorityYear="1963" box="[776,1109,439,465]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="494" phylum="Arthropoda" rank="species" species="transvaalensis">
<emphasis id="6887BD20FFA898360258ADEA703EFE00" box="[776,795,439,465]" italics="true" pageId="8" pageNumber="494">C</emphasis>
. 
<emphasis id="6887BD20FFA898360279ADEA70EEFE00" box="[809,971,439,465]" italics="true" pageId="8" pageNumber="494">transvaalensis</emphasis>
<bibRefCitation id="3E621CC3FFA898360282ADEA7770FE00" author="Ryke, P. A. J." box="[978,1109,439,465]" pageId="8" pageNumber="494" pagination="1 - 15" refId="ref16407" refString="Ryke, P. A. J. (1963) Some free-living Hypoaspidinae (Acari: Mesostigmata) from South Africa. Revista de Biologia, 5, 1 - 15." type="journal article" year="1963">Ryke, 1963</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="9DF31AB1FFA898360530ADEA76B2FE00" authority="(Berlese, 1905)" baseAuthorityName="Berlese" baseAuthorityYear="1905" box="[1120,1431,438,465]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="494" phylum="Arthropoda" rank="species" species="robustus">
<emphasis id="6887BD20FFA898360530ADEA7756FE00" box="[1120,1139,439,465]" italics="true" pageId="8" pageNumber="494">C</emphasis>
. 
<emphasis id="6887BD20FFA8983605D1ADEA77C5FE00" box="[1153,1248,439,465]" italics="true" pageId="8" pageNumber="494">robustus</emphasis>
(
<bibRefCitation id="3E621CC3FFA8983605A0ADEA76AAFE00" author="Berlese, A." box="[1264,1423,438,465]" pageId="8" pageNumber="494" pagination="154 - 176" refId="ref13741" refString="Berlese, A. (1905) Acari nuovi. Manipulus IV. Redia, 2, 154 - 176." type="journal article" year="1905">Berlese, 1905</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicName id="9DF31AB1FFA8983601C7AD8672FEFE24" authority="Joharchi 2019" authorityName="Joharchi" authorityYear="2019" box="[151,475,475,501]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="494" phylum="Arthropoda" rank="species" species="siberiensis">
<emphasis id="6887BD20FFA8983601C7AD86738FFE24" box="[151,170,475,501]" italics="true" pageId="8" pageNumber="494">C</emphasis>
. 
<emphasis id="6887BD20FFA8983601EAAD867217FE24" box="[186,306,475,501]" italics="true" pageId="8" pageNumber="494">siberiensis</emphasis>
<bibRefCitation id="3E621CC3FFA89836006BAD8672FEFE24" author="Joharchi, O &amp; Trach, V. A." box="[315,475,475,501]" pageId="8" pageNumber="494" pagination="486 - 494" refId="ref14677" refString="Joharchi, O &amp; Trach, V. A. (2019) A new species of Cosmolaelaps Berlese (Acari: Laelapidae) from Ukraine. Zootaxa, 4647 (1), 486 - 494. https: // doi. org / 10.11646 / zootaxa. 4647.1.30" type="journal article" year="2019">Joharchi 2019</bibRefCitation>
</taxonomicName>
(in 
<bibRefCitation id="3E621CC3FFA89836035DAD8671D9FE24" author="Joharchi, O &amp; Trach, V. A." box="[525,764,475,501]" pageId="8" pageNumber="494" pagination="486 - 494" refId="ref14677" refString="Joharchi, O &amp; Trach, V. A. (2019) A new species of Cosmolaelaps Berlese (Acari: Laelapidae) from Ukraine. Zootaxa, 4647 (1), 486 - 494. https: // doi. org / 10.11646 / zootaxa. 4647.1.30" type="journal article" year="2019">
Joharchi 
<emphasis id="6887BD20FFA898360325AD86718CFE24" box="[629,681,475,501]" italics="true" pageId="8" pageNumber="494">et al</emphasis>
., 2019
</bibRefCitation>
), 
<taxonomicName id="9DF31AB1FFA898360242AD86777BFE27" authority="(Shcherbak, 1971 b)" baseAuthorityName="Shcherbak" baseAuthorityYear="1971" box="[786,1118,475,502]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="494" phylum="Arthropoda" rank="species" species="mixta">
<emphasis id="6887BD20FFA898360242AD867000FE24" box="[786,805,475,501]" italics="true" pageId="8" pageNumber="494">C</emphasis>
. 
<emphasis id="6887BD20FFA898360265AD867056FE25" box="[821,883,475,500]" italics="true" pageId="8" pageNumber="494">mixta</emphasis>
(
<bibRefCitation id="3E621CC3FFA8983602D4AD867770FE27" author="Shcherbak, G. I." box="[900,1109,475,502]" pageId="8" pageNumber="494" pagination="20 - 30" refId="ref16506" refString="Shcherbak, G. I. (1971 b) Species of Gamasoidea mites from the genus Hypoaspis Can. (Parasitiformes, Gamasoidea) new for the USSR fauna. Collection of papers from the Zoological Museum of the Institute of Zoology of SSR Ukraine Academy of Sciences, 34, 20 - 30. [in Ukrainian]" type="journal article" year="1971">Shcherbak, 1971b</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicName id="9DF31AB1FFA89836053CAD867651FE24" authority="(Karg, 1988)" baseAuthorityName="Karg" baseAuthorityYear="1988" box="[1132,1396,475,501]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="494" phylum="Arthropoda" rank="species" species="pumili">
<emphasis id="6887BD20FFA89836053CAD86775AFE24" box="[1132,1151,475,501]" italics="true" pageId="8" pageNumber="494">C</emphasis>
. 
<emphasis id="6887BD20FFA8983605DFAD8677F2FE24" box="[1167,1239,475,501]" italics="true" pageId="8" pageNumber="494">pumili</emphasis>
(
<bibRefCitation id="3E621CC3FFA8983605B8AD867649FE24" author="Karg, W." box="[1256,1388,475,501]" pageId="8" pageNumber="494" pagination="509 - 525" refId="ref15324" refString="Karg, W. (1988) Die Arten der Raubmilbengattung Cosmolaelaps Berlese, 1903 (Gattung Hypoaspis Canestrini, 1884, Acarina, Parasitiformes). Zoologische Jahrbucher Abteilung fur Systematik, Okologie und Geographie der Tiere, 115, 509 - 525." type="journal article" year="1988">Karg, 1988</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicName id="9DF31AB1FFA8983604D3AD86725EFDC8" authority="(Karg, 1987)" baseAuthorityName="Karg" baseAuthorityYear="1987" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="494" phylum="Arthropoda" rank="species" species="lingua">
<emphasis id="6887BD20FFA8983604D3AD8676B3FE24" box="[1411,1430,475,501]" italics="true" pageId="8" pageNumber="494">C</emphasis>
. 
<emphasis id="6887BD20FFA8983601C7ADA273FAFDC8" box="[151,223,511,537]" italics="true" pageId="8" pageNumber="494">lingua</emphasis>
(
<bibRefCitation id="3E621CC3FFA8983601BFADA27256FDC8" author="Karg, W." box="[239,371,511,537]" pageId="8" pageNumber="494" pagination="289 - 302" refId="ref15285" refString="Karg, W. (1987) Neue Raubmilbenarten der Gattung Hypoaspis Can. 1884 (Acari: Parasitiformes). Zoologische Jahrbucher Abteilung fur Systematik, Okologie und Geographie der Tiere, 114, 289 - 302." type="journal article" year="1987">Karg, 1987</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicName id="9DF31AB1FFA8983600D8ADA271B4FDCB" authority="(Ma, 1996)" baseAuthorityName="Ma" baseAuthorityYear="1996" box="[392,657,511,538]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="494" phylum="Arthropoda" rank="species" species="sungaris">
<emphasis id="6887BD20FFA8983600D8ADA272BEFDC8" box="[392,411,511,537]" italics="true" pageId="8" pageNumber="494">C</emphasis>
. 
<emphasis id="6887BD20FFA8983600F9ADA2712CFDC9" box="[425,521,511,536]" italics="true" pageId="8" pageNumber="494">sungaris</emphasis>
(
<bibRefCitation id="3E621CC3FFA89836034AADA271ACFDCB" author="Ma, L. - M." box="[538,649,511,538]" pageId="8" pageNumber="494" pagination="48 - 54" refId="ref15688" refString="Ma, L. - M. (1996) Three new species of the genus Hypoaspis from Jilin Province, China (Acari: Laelapidae). Acta Zootaxonomica Sinica, 21, 48 - 54. [in Chinese]" type="journal article" year="1996">Ma, 1996</bibRefCitation>
)
</taxonomicName>
and 
<taxonomicName id="9DF31AB1FFA89836039AADA2777CFDC8" authority="(Bei et al., 2003)" baseAuthorityName="Bei" baseAuthorityYear="2003" box="[714,1113,511,537]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="494" phylum="Arthropoda" rank="species" species="shenyangensis">
<emphasis id="6887BD20FFA89836039AADA271F8FDC8" box="[714,733,511,537]" italics="true" pageId="8" pageNumber="494">C</emphasis>
. 
<emphasis id="6887BD20FFA8983603BCADA270ABFDC8" box="[748,910,511,537]" italics="true" pageId="8" pageNumber="494">shenyangensis</emphasis>
(
<bibRefCitation id="3E621CC3FFA8983602CEADA27774FDC8" author="Bei, N. - X. &amp; Shi, C. - M. &amp; Yin, S. - G." box="[926,1105,511,537]" pageId="8" pageNumber="494" pagination="648 - 650" refId="ref13627" refString="Bei, N. - X., Shi, C. - M. &amp; Yin, S. - G. (2003) A new species of the genus Hypoaspis from China (Acari, Laelapidae). Acta Zootaxonomica Sinica, 28, 648 - 650. [in Chinese]" type="journal article" year="2003">
Bei 
<emphasis id="6887BD20FFA89836029FADA27724FDC8" box="[975,1025,511,537]" italics="true" pageId="8" pageNumber="494">et al</emphasis>
., 2003
</bibRefCitation>
)
</taxonomicName>
in the shape of dorsal setae, which are slightly swollen near the base and pointed at tip (knife-shaped) (see 
<figureCitation id="C2C87DB7FFA898360552AE7E7744FDEC" box="[1026,1121,547,573]" captionStart="FIGURES 8–12" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[181,1397,196,1901]" captionTargetId="figure-18@4.[167,1419,181,1919]" captionTargetPageId="4" captionText="FIGURES 8–12. Cosmolaelaps lasiophilus Joharchi sp. nov., female. 8, dorsal idiosoma, enlarged section not to scale; 9, ventral idiosoma; 10, subcapitulum; 11, epistome; 12, chelicera." figureDoi="http://doi.org/10.5281/zenodo.6530964" httpUri="https://zenodo.org/record/6530964/files/figure.png" pageId="8" pageNumber="494">Figure 8</figureCitation>
), with all dorsal setae of the same shape and sub-equal in length (at most reaching base of following setae), except setae 
<emphasis id="6887BD20FFA8983605CEAE1A7791FDB0" box="[1182,1204,583,609]" italics="true" pageId="8" pageNumber="494">j1</emphasis>
and 
<emphasis id="6887BD20FFA8983605BAAE1A762DFDB0" box="[1258,1288,583,609]" italics="true" pageId="8" pageNumber="494">Z5</emphasis>
.
</paragraph>
<paragraph id="5A4C6132FFA898360197AE3670F6F96C" blockId="8.[151,1437,151,1725]" pageId="8" pageNumber="494">
<taxonomicName id="9DF31AB1FFA898360197AE3672CAFD54" authorityName="Joharchi" authorityYear="2022" box="[199,495,619,645]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="494" phylum="Arthropoda" rank="species" species="lasiophilus">
<emphasis id="6887BD20FFA898360197AE3672CAFD54" box="[199,495,619,645]" italics="true" pageId="8" pageNumber="494">Cosmolaelaps lasiophilus</emphasis>
</taxonomicName>
differs from 
<taxonomicName id="9DF31AB1FFA8983603C7AE367024FD55" authorityName="sensu Evans &amp; Till" authorityYear="1966" baseAuthorityName="Michael" baseAuthorityYear="1891" box="[663,769,619,645]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="494" phylum="Arthropoda" rank="species" species="vacua">
<emphasis id="6887BD20FFA8983603C7AE36718FFD54" box="[663,682,619,645]" italics="true" pageId="8" pageNumber="494">C</emphasis>
. 
<emphasis id="6887BD20FFA8983603EEAE367024FD55" box="[702,769,619,644]" italics="true" pageId="8" pageNumber="494">vacua</emphasis>
</taxonomicName>
by lacking spatulate setae on legs, most setae on legs conspicuously thickened and some inserted on small tubercles, while in 
<taxonomicName id="9DF31AB1FFA898360282AED27712FD79" authorityName="sensu Evans &amp; Till" authorityYear="1966" baseAuthorityName="Michael" baseAuthorityYear="1891" box="[978,1079,655,681]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="494" phylum="Arthropoda" rank="species" species="vacua">
<emphasis id="6887BD20FFA898360282AED270C0FD78" box="[978,997,655,681]" italics="true" pageId="8" pageNumber="494">C</emphasis>
. 
<emphasis id="6887BD20FFA8983602A4AED27712FD79" box="[1012,1079,655,680]" italics="true" pageId="8" pageNumber="494">vacua</emphasis>
</taxonomicName>
two spatulate setae are present on femora II and IV. 
<bibRefCitation id="3E621CC3FFA8983600D9AEEE714BFD1F" author="Evans, G. O. &amp; Till, W. M." box="[393,622,691,718]" pageId="8" pageNumber="494" pagination="107 - 370" refId="ref14300" refString="Evans, G. O. &amp; Till, W. M. (1966) Studies on the British Dermanyssidae (Acari: Mesostigmata). Part II. Classification. Bulletin of the British Museum (Natural History) Zoology, 14, 107 - 370." type="journal article" year="1966">Evans &amp; Till (1966)</bibRefCitation>
did not provide any information about the shape of the leg setae for 
<taxonomicName id="9DF31AB1FFA8983604D3AEEE73FFFD21" authorityName="sensu Evans &amp; Till" authorityYear="1966" baseAuthorityName="Michael" baseAuthorityYear="1891" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="494" phylum="Arthropoda" rank="species" species="vacua">
<emphasis id="6887BD20FFA8983604D3AEEE76B3FD1C" box="[1411,1430,691,717]" italics="true" pageId="8" pageNumber="494">C</emphasis>
. 
<emphasis id="6887BD20FFA8983601C7AE8A73FFFD21" box="[151,218,727,752]" italics="true" pageId="8" pageNumber="494">vacua</emphasis>
</taxonomicName>
, but according to the original description (
<bibRefCitation id="3E621CC3FFA8983603E4AE8A707DFD20" author="Michael, A. D." box="[692,856,727,753]" pageId="8" pageNumber="494" pagination="638 - 654" refId="ref15960" refString="Michael, A. D. (1891) On the association of gamasids with ants. Proceedings of the Zoological Society of London, 4, 638 - 654." type="journal article" year="1891">Michael, 1891</bibRefCitation>
) of 
<taxonomicName id="9DF31AB1FFA8983602D5AE8A7748FD21" authorityName="sensu Evans &amp; Till" authorityYear="1966" baseAuthorityName="Michael" baseAuthorityYear="1891" box="[901,1133,727,753]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="494" phylum="Arthropoda" rank="species" species="vacua">
<emphasis id="6887BD20FFA8983602D5AE8A7748FD21" box="[901,1133,727,753]" italics="true" pageId="8" pageNumber="494">Cosmolaelaps vacua</emphasis>
</taxonomicName>
, femora II and IV bear two spatulate setae (
<bibRefCitation id="3E621CC3FFA89836001DAEA672F5FCC4" author="Karg, W." box="[333,464,763,789]" pageId="8" pageNumber="494" pagination="213 - 232" refId="ref15239" refString="Karg, W. (1981) Die Untergattung Cosmolaelaps Berlese, 1903 der Raubmilbengattung Hypoaspis Canestrini, 1884. Mitteilungen aus dem Zoologischen Museum in Berlin, 57, 213 - 232. https: // doi. org / 10.1002 / mmnz. 19810570203" type="journal article" year="1981">Karg, 1981</bibRefCitation>
, 
<figureCitation id="C2C87DB7FFA89836008EAEA67114FCC4" box="[478,561,763,789]" captionStart="FIGURES 8–12" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[181,1397,196,1901]" captionTargetId="figure-18@4.[167,1419,181,1919]" captionTargetPageId="4" captionText="FIGURES 8–12. Cosmolaelaps lasiophilus Joharchi sp. nov., female. 8, dorsal idiosoma, enlarged section not to scale; 9, ventral idiosoma; 10, subcapitulum; 11, epistome; 12, chelicera." figureDoi="http://doi.org/10.5281/zenodo.6530964" httpUri="https://zenodo.org/record/6530964/files/figure.png" pageId="8" pageNumber="494">Fig. 8a</figureCitation>
, also illustrated this character). It differs from 
<taxonomicName id="9DF31AB1FFA89836051BAEA677FBFCC4" authorityName="Zeman" authorityYear="1982" box="[1099,1246,763,789]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="494" phylum="Arthropoda" rank="species" species="thysanifer">
<emphasis id="6887BD20FFA89836051BAEA6777BFCC4" box="[1099,1118,763,789]" italics="true" pageId="8" pageNumber="494">C</emphasis>
. 
<emphasis id="6887BD20FFA89836053EAEA677FBFCC4" box="[1134,1246,763,789]" italics="true" pageId="8" pageNumber="494">thysanifer</emphasis>
</taxonomicName>
by setae 
<emphasis id="6887BD20FFA89836041DAEA6764EFCC4" box="[1357,1387,763,789]" italics="true" pageId="8" pageNumber="494">Z5</emphasis>
and some opisthogastric setae thickened, versus setae 
<emphasis id="6887BD20FFA898360399AF4271C3FCE8" box="[713,742,799,825]" italics="true" pageId="8" pageNumber="494">Z5</emphasis>
brush-like and all opisthogastric setae simple and setiform in 
<taxonomicName id="9DF31AB1FFA8983601C7AF1E720FFC8C" authorityName="Zeman" authorityYear="1982" box="[151,298,835,861]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="494" phylum="Arthropoda" rank="species" species="thysanifer">
<emphasis id="6887BD20FFA8983601C7AF1E738FFC8C" box="[151,170,835,861]" italics="true" pageId="8" pageNumber="494">C</emphasis>
. 
<emphasis id="6887BD20FFA8983601EAAF1E720FFC8C" box="[186,298,835,861]" italics="true" pageId="8" pageNumber="494">thysanifer</emphasis>
</taxonomicName>
. It differs from 
<taxonomicName id="9DF31AB1FFA8983600B4AF1E7142FC8D" baseAuthorityName="Ma" baseAuthorityYear="1996" box="[484,615,835,861]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="494" phylum="Arthropoda" rank="species" species="sungaris">
<emphasis id="6887BD20FFA8983600B4AF1E72D2FC8C" box="[484,503,835,861]" italics="true" pageId="8" pageNumber="494">C</emphasis>
. 
<emphasis id="6887BD20FFA898360357AF1E7142FC8D" box="[519,615,835,860]" italics="true" pageId="8" pageNumber="494">sungaris</emphasis>
</taxonomicName>
and 
<taxonomicName id="9DF31AB1FFA8983603F2AF1E7043FC8C" box="[674,870,835,861]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="494" phylum="Arthropoda" rank="species" species="shenyangensis">
<emphasis id="6887BD20FFA8983603F2AF1E7190FC8C" box="[674,693,835,861]" italics="true" pageId="8" pageNumber="494">C</emphasis>
. 
<emphasis id="6887BD20FFA898360394AF1E7043FC8C" box="[708,870,835,861]" italics="true" pageId="8" pageNumber="494">shenyangensis</emphasis>
</taxonomicName>
by its much smaller size of body (389–406 long, 256–265wide), versus body size much larger in both of 
<taxonomicName id="9DF31AB1FFA89836025AAF3A70AEFC51" baseAuthorityName="Ma" baseAuthorityYear="1996" box="[778,907,871,897]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="494" phylum="Arthropoda" rank="species" species="sungaris">
<emphasis id="6887BD20FFA89836025AAF3A7038FC50" box="[778,797,871,897]" italics="true" pageId="8" pageNumber="494">C</emphasis>
. 
<emphasis id="6887BD20FFA89836027BAF3A70AEFC51" box="[811,907,871,896]" italics="true" pageId="8" pageNumber="494">sungaris</emphasis>
</taxonomicName>
and 
<taxonomicName id="9DF31AB1FFA898360293AF3A77A3FC50" box="[963,1158,871,897]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="494" phylum="Arthropoda" rank="species" species="shenyangensis">
<emphasis id="6887BD20FFA898360293AF3A70F3FC50" box="[963,982,871,897]" italics="true" pageId="8" pageNumber="494">C</emphasis>
. 
<emphasis id="6887BD20FFA8983602B4AF3A77A3FC50" box="[996,1158,871,897]" italics="true" pageId="8" pageNumber="494">shenyangensis</emphasis>
</taxonomicName>
(566 long × 383 wide in 
<taxonomicName id="9DF31AB1FFA8983601C7AFD67232FC75" baseAuthorityName="Ma" baseAuthorityYear="1996" box="[151,279,907,933]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="494" phylum="Arthropoda" rank="species" species="sungaris">
<emphasis id="6887BD20FFA8983601C7AFD6738FFC74" box="[151,170,907,933]" italics="true" pageId="8" pageNumber="494">C</emphasis>
. 
<emphasis id="6887BD20FFA8983601E7AFD67232FC75" box="[183,279,907,932]" italics="true" pageId="8" pageNumber="494">sungaris</emphasis>
</taxonomicName>
, and 567 long × 413 wide in 
<taxonomicName id="9DF31AB1FFA89836030AAFD671D9FC74" box="[602,764,907,933]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="494" phylum="Arthropoda" rank="species" species="shenyangensis">
<emphasis id="6887BD20FFA89836030AAFD671D9FC74" box="[602,764,907,933]" italics="true" pageId="8" pageNumber="494">shenyangensis</emphasis>
</taxonomicName>
) and also differs from 
<taxonomicName id="9DF31AB1FFA8983602A8AFD6775DFC75" baseAuthorityName="Ma" baseAuthorityYear="1996" box="[1016,1144,907,933]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="494" phylum="Arthropoda" rank="species" species="sungaris">
<emphasis id="6887BD20FFA8983602A8AFD6772EFC74" box="[1016,1035,907,933]" italics="true" pageId="8" pageNumber="494">C</emphasis>
. 
<emphasis id="6887BD20FFA898360548AFD6775DFC75" box="[1048,1144,907,932]" italics="true" pageId="8" pageNumber="494">sungaris</emphasis>
</taxonomicName>
by its longer dorsal shield setae (reaching base of following setae), versus dorsal shield setae much shorter in 
<taxonomicName id="9DF31AB1FFA898360507AFF277FFFC19" baseAuthorityName="Ma" baseAuthorityYear="1996" box="[1111,1242,943,969]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="494" phylum="Arthropoda" rank="species" species="sungaris">
<emphasis id="6887BD20FFA898360507AFF2774FFC18" box="[1111,1130,943,969]" italics="true" pageId="8" pageNumber="494">C</emphasis>
. 
<emphasis id="6887BD20FFA89836052AAFF277FFFC19" box="[1146,1242,943,968]" italics="true" pageId="8" pageNumber="494">sungaris</emphasis>
</taxonomicName>
(barely reaching more than half of distance of following setae in series). 
<taxonomicName id="9DF31AB1FFA898360251AF8E7707FC3C" authorityName="Joharchi" authorityYear="2022" box="[769,1058,979,1005]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="494" phylum="Arthropoda" rank="species" species="lasiophilus">
<emphasis id="6887BD20FFA898360251AF8E7707FC3C" box="[769,1058,979,1005]" italics="true" pageId="8" pageNumber="494">Cosmolaelaps lasiophilus</emphasis>
</taxonomicName>
can be readily distinguished from 
<taxonomicName id="9DF31AB1FFA8983601C7AFAA7279FBC0" authorityName="Ryke" authorityYear="1963" box="[151,348,1015,1041]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="494" phylum="Arthropoda" rank="species" species="transvaalensis">
<emphasis id="6887BD20FFA8983601C7AFAA738FFBC0" box="[151,170,1015,1041]" italics="true" pageId="8" pageNumber="494">C</emphasis>
. 
<emphasis id="6887BD20FFA8983601EAAFAA7279FBC0" box="[186,348,1015,1041]" italics="true" pageId="8" pageNumber="494">transvaalensis</emphasis>
</taxonomicName>
by some opisthogastric setae thickened and most setae on legs conspicuously thickened (some inserted on small tubercles), versus setae on both opisthogastric cuticle and legs simple in 
<taxonomicName id="9DF31AB1FFA8983605F2A8467642FBE4" authorityName="Ryke" authorityYear="1963" box="[1186,1383,1051,1077]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="494" phylum="Arthropoda" rank="species" species="transvaalensis">
<emphasis id="6887BD20FFA8983605F2A8467790FBE4" box="[1186,1205,1051,1077]" italics="true" pageId="8" pageNumber="494">C</emphasis>
. 
<emphasis id="6887BD20FFA898360595A8467642FBE4" box="[1221,1383,1051,1077]" italics="true" pageId="8" pageNumber="494">transvaalensis</emphasis>
</taxonomicName>
(see 
<bibRefCitation id="3E621CC3FFA8983601C7A862723EFB8B" author="Ryke, P. A. J." box="[151,283,1087,1114]" pageId="8" pageNumber="494" pagination="1 - 15" refId="ref16407" refString="Ryke, P. A. J. (1963) Some free-living Hypoaspidinae (Acari: Mesostigmata) from South Africa. Revista de Biologia, 5, 1 - 15." type="journal article" year="1963">Ryke, 1963</bibRefCitation>
). It differs from 
<taxonomicName id="9DF31AB1FFA898360082A862711FFB88" baseAuthorityName="Karg" baseAuthorityYear="1988" box="[466,570,1087,1113]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="494" phylum="Arthropoda" rank="species" species="pumili">
<emphasis id="6887BD20FFA898360082A86272C0FB88" box="[466,485,1087,1113]" italics="true" pageId="8" pageNumber="494">C</emphasis>
. 
<emphasis id="6887BD20FFA8983600A2A862711FFB88" box="[498,570,1087,1113]" italics="true" pageId="8" pageNumber="494">pumili</emphasis>
</taxonomicName>
because setae 
<emphasis id="6887BD20FFA89836038EA86271D9FB88" box="[734,764,1087,1113]" italics="true" pageId="8" pageNumber="494">Z5</emphasis>
are thickened, shorter than 
<emphasis id="6887BD20FFA898360560A862776EFB88" box="[1072,1099,1087,1113]" italics="true" pageId="8" pageNumber="494">J5</emphasis>
and legs I, IV are shorter than length of idiosoma, while in 
<taxonomicName id="9DF31AB1FFA898360082A83E711CFBAC" baseAuthorityName="Karg" baseAuthorityYear="1988" box="[466,569,1123,1149]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="494" phylum="Arthropoda" rank="species" species="pumili">
<emphasis id="6887BD20FFA898360082A83E72C0FBAC" box="[466,485,1123,1149]" italics="true" pageId="8" pageNumber="494">C</emphasis>
. 
<emphasis id="6887BD20FFA8983600A1A83E711CFBAC" box="[497,569,1123,1149]" italics="true" pageId="8" pageNumber="494">pumili</emphasis>
</taxonomicName>
setae 
<emphasis id="6887BD20FFA89836032BA83E71BCFBAC" box="[635,665,1123,1149]" italics="true" pageId="8" pageNumber="494">Z5</emphasis>
setiform, longer than 
<emphasis id="6887BD20FFA8983602DBA83E7080FBAC" box="[907,933,1123,1149]" italics="true" pageId="8" pageNumber="494">J5</emphasis>
and legs I, IV longer than length of idiosoma. 
<taxonomicName id="9DF31AB1FFA8983601C7A8DA7211FB70" authorityName="Joharchi" authorityYear="2022" box="[151,308,1159,1185]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="494" phylum="Arthropoda" rank="species" species="lasiophilus">
<emphasis id="6887BD20FFA8983601C7A8DA738FFB70" box="[151,170,1159,1185]" italics="true" pageId="8" pageNumber="494">C</emphasis>
. 
<emphasis id="6887BD20FFA8983601E9A8DA7211FB70" box="[185,308,1159,1185]" italics="true" pageId="8" pageNumber="494">lasiophilus</emphasis>
</taxonomicName>
can be easily distinguished from 
<taxonomicName id="9DF31AB1FFA8983603E2A8DA7017FB70" baseAuthorityName="Berlese" baseAuthorityYear="1905" box="[690,818,1159,1185]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="494" phylum="Arthropoda" rank="species" species="robustus">
<emphasis id="6887BD20FFA8983603E2A8DA71E0FB70" box="[690,709,1159,1185]" italics="true" pageId="8" pageNumber="494">C</emphasis>
. 
<emphasis id="6887BD20FFA898360383A8DA7017FB70" box="[723,818,1159,1185]" italics="true" pageId="8" pageNumber="494">robustus</emphasis>
</taxonomicName>
by the smaller size of the genital shield (there are two pairs of setae between genital and anal shields) and presternal area lightly sclerotised (with a few transverse curved lines and merged to the sternal shield), while genital shield expanded posteriorly in 
<taxonomicName id="9DF31AB1FFA898360516A89277E3FB38" baseAuthorityName="Berlese" baseAuthorityYear="1905" box="[1094,1222,1231,1257]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="494" phylum="Arthropoda" rank="species" species="robustus">
<emphasis id="6887BD20FFA898360516A892777CFB38" box="[1094,1113,1231,1257]" italics="true" pageId="8" pageNumber="494">C</emphasis>
. 
<emphasis id="6887BD20FFA898360537A89277E3FB38" box="[1127,1222,1231,1257]" italics="true" pageId="8" pageNumber="494">robustus</emphasis>
</taxonomicName>
, setae 
<emphasis id="6887BD20FFA898360444A892761BFB38" box="[1300,1342,1231,1257]" italics="true" pageId="8" pageNumber="494">Zv1</emphasis>
and 
<emphasis id="6887BD20FFA898360425A89276B9FB38" box="[1397,1436,1231,1257]" italics="true" pageId="8" pageNumber="494">Jv1</emphasis>
inserted on lateral margins of shield (in addition of 
<emphasis id="6887BD20FFA89836038AA8AE71DEFADC" box="[730,763,1267,1293]" italics="true" pageId="8" pageNumber="494">st5</emphasis>
) and two triangular presternal plates present in 
<taxonomicName id="9DF31AB1FFA898360445A8AE76B0FADC" baseAuthorityName="Berlese" baseAuthorityYear="1905" box="[1301,1429,1267,1293]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="494" phylum="Arthropoda" rank="species" species="robustus">
<emphasis id="6887BD20FFA898360445A8AE760DFADC" box="[1301,1320,1267,1293]" italics="true" pageId="8" pageNumber="494">C</emphasis>
. 
<emphasis id="6887BD20FFA898360466A8AE76B0FADC" box="[1334,1429,1267,1293]" italics="true" pageId="8" pageNumber="494">robustus</emphasis>
</taxonomicName>
. 
<taxonomicName id="9DF31AB1FFA8983601C7A94A7293FAE0" authorityName="Joharchi" authorityYear="2022" box="[151,438,1303,1329]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="494" phylum="Arthropoda" rank="species" species="lasiophilus">
<emphasis id="6887BD20FFA8983601C7A94A7293FAE0" box="[151,438,1303,1329]" italics="true" pageId="8" pageNumber="494">Cosmolaelaps lasiophilus</emphasis>
</taxonomicName>
can be distinguished from 
<taxonomicName id="9DF31AB1FFA89836038FA94A7062FAE0" baseAuthorityName="Karg" baseAuthorityYear="1987" box="[735,839,1303,1329]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="494" phylum="Arthropoda" rank="species" species="lingua">
<emphasis id="6887BD20FFA89836038FA94A71D7FAE0" box="[735,754,1303,1329]" italics="true" pageId="8" pageNumber="494">C</emphasis>
. 
<emphasis id="6887BD20FFA8983603AFA94A7062FAE0" box="[767,839,1303,1329]" italics="true" pageId="8" pageNumber="494">lingua</emphasis>
</taxonomicName>
by its small body size (389–406 long, 256–265 wide), thick and short opisthogastric setae and presternal area lightly sclerotised (anterior margin of sternal shield hardly conspicuous), versus body larger (480–500 long, 300–330 wide), opisthogastric setae simple (setiform) and long, sclerotised platelets absent and presternal plates joined to sternal shield in 
<taxonomicName id="9DF31AB1FFA8983602B6A9DE7775FA4C" baseAuthorityName="Karg" baseAuthorityYear="1987" box="[998,1104,1411,1437]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="494" phylum="Arthropoda" rank="species" species="lingua">
<emphasis id="6887BD20FFA8983602B6A9DE70DCFA4C" box="[998,1017,1411,1437]" italics="true" pageId="8" pageNumber="494">C</emphasis>
. 
<emphasis id="6887BD20FFA898360558A9DE7775FA4C" box="[1032,1104,1411,1437]" italics="true" pageId="8" pageNumber="494">lingua</emphasis>
</taxonomicName>
. 
<taxonomicName id="9DF31AB1FFA89836050FA9DE76A4FA4C" authorityName="Joharchi" authorityYear="2022" box="[1119,1409,1411,1437]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="494" phylum="Arthropoda" rank="species" species="lasiophilus">
<emphasis id="6887BD20FFA89836050FA9DE76A4FA4C" box="[1119,1409,1411,1437]" italics="true" pageId="8" pageNumber="494">Cosmolaelaps lasiophilus</emphasis>
</taxonomicName>
is most similar to 
<taxonomicName id="9DF31AB1FFA89836001BA9FA72C0FA10" authorityName="Joharchi" authorityYear="2019" box="[331,485,1447,1473]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="494" phylum="Arthropoda" rank="species" species="siberiensis">
<emphasis id="6887BD20FFA89836001BA9FA727BFA10" box="[331,350,1447,1473]" italics="true" pageId="8" pageNumber="494">C</emphasis>
. 
<emphasis id="6887BD20FFA89836003DA9FA72C0FA10" box="[365,485,1447,1473]" italics="true" pageId="8" pageNumber="494">siberiensis</emphasis>
</taxonomicName>
and 
<taxonomicName id="9DF31AB1FFA89836034EA9FA715BFA11" baseAuthorityName="Shcherbak" baseAuthorityYear="1971" box="[542,638,1447,1473]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="494" phylum="Arthropoda" rank="species" species="mixta">
<emphasis id="6887BD20FFA89836034EA9FA7114FA10" box="[542,561,1447,1473]" italics="true" pageId="8" pageNumber="494">C</emphasis>
. 
<emphasis id="6887BD20FFA898360310A9FA715BFA11" box="[576,638,1447,1472]" italics="true" pageId="8" pageNumber="494">mixta</emphasis>
</taxonomicName>
, but can be readily distinguished from these two species by its longer dorsal setae, some of them reach to the base of next setae, while in both 
<taxonomicName id="9DF31AB1FFA898360281A996774EFA34" authorityName="Joharchi" authorityYear="2019" box="[977,1131,1483,1509]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="494" phylum="Arthropoda" rank="species" species="siberiensis">
<emphasis id="6887BD20FFA898360281A99670C1FA34" box="[977,996,1483,1509]" italics="true" pageId="8" pageNumber="494">C</emphasis>
. 
<emphasis id="6887BD20FFA8983602A3A996774EFA34" box="[1011,1131,1483,1509]" italics="true" pageId="8" pageNumber="494">siberiensis</emphasis>
</taxonomicName>
<emphasis id="6887BD20FFA898360523A99677B8FA34" box="[1139,1181,1483,1509]" italics="true" pageId="8" pageNumber="494">and</emphasis>
<taxonomicName id="9DF31AB1FFA8983605F5A9967620FA35" baseAuthorityName="Shcherbak" baseAuthorityYear="1971" box="[1189,1285,1483,1509]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="494" phylum="Arthropoda" rank="species" species="mixta">
<emphasis id="6887BD20FFA8983605F5A996779DFA34" box="[1189,1208,1483,1509]" italics="true" pageId="8" pageNumber="494">C</emphasis>
. 
<emphasis id="6887BD20FFA898360597A9967620FA35" box="[1223,1285,1483,1508]" italics="true" pageId="8" pageNumber="494">mixta</emphasis>
</taxonomicName>
dorsal shield setae considerably shorter and none of them reach base of next. Overall 
<taxonomicName id="9DF31AB1FFA898360299A9B27740F9D8" authorityName="Joharchi" authorityYear="2022" box="[969,1125,1519,1545]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="494" phylum="Arthropoda" rank="species" species="lasiophilus">
<emphasis id="6887BD20FFA898360299A9B270F9F9D8" box="[969,988,1519,1545]" italics="true" pageId="8" pageNumber="494">C</emphasis>
. 
<emphasis id="6887BD20FFA8983602BAA9B27740F9D8" box="[1002,1125,1519,1545]" italics="true" pageId="8" pageNumber="494">lasiophilus</emphasis>
</taxonomicName>
can be easily distinguished from all other congeners by the following combination of characters: (1) dorsal shield bearing 39 pairs of relatively long setae (some reach base of next posterior setae in series), 
<emphasis id="6887BD20FFA898360200AA6A7043F980" box="[848,870,1591,1617]" italics="true" pageId="8" pageNumber="494">j1</emphasis>
, 
<emphasis id="6887BD20FFA898360225AA6A70ABF980" box="[885,910,1591,1617]" italics="true" pageId="8" pageNumber="494">z1</emphasis>
and 
<emphasis id="6887BD20FFA898360295AA6A70C6F980" box="[965,995,1591,1617]" italics="true" pageId="8" pageNumber="494">Z5</emphasis>
simple, 
<emphasis id="6887BD20FFA898360514AA6A7747F980" box="[1092,1122,1591,1617]" italics="true" pageId="8" pageNumber="494">Z5</emphasis>
shorter than 
<emphasis id="6887BD20FFA8983605A6AA6A7634F980" box="[1270,1297,1591,1617]" italics="true" pageId="8" pageNumber="494">J5</emphasis>
(ratio of J5/ Z5 length ≃ 1.5); (2) post-anal seta thickened; (3) almost all opisthogastric setae thickened and arising on minute sclerotised platelet; (4) fixed digit of chelicera with six various size of teeth; (5) legs without spatulate setae but most setae conspicuously thickened and some inserted on small tubercles.
</paragraph>
</subSubSection>
</treatment>
</document>