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<document id="7396A2E7ABB321C99B48D07715241ABF" ID-DOI="10.11646/phytotaxa.629.1.3" ID-ISSN="1179-3163" ID-Zenodo-Dep="10254755" IM.bibliography_approvedBy="jonas" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="jonas" IM.treatments_approvedBy="jonas" checkinTime="1701682507275" checkinUser="plazi" docAuthor="Smith, Gideon F., Figueiredo, Estrela, Verloove, Filip, Klopper, Ronell R. &amp; Silva, Vasco" docDate="2023" docId="B5213461455FFFDC99A8FF30FBC80928" docLanguage="en" docName="phytotaxa.629.1.3.pdf" docOrigin="Phytotaxa 629 (1)" docSource="https://phytotaxa.mapress.com/pt/article/download/phytotaxa.629.1.3/51327" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.11:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="11" docTitle="Aloe maculata subsp. maculata" docType="treatment" docVersion="3" lastPageNumber="41" masterDocId="49184C19455BFFDA9920FFAEFFF30C3C" masterDocTitle="An annotated catalogue of Aloe and Aloiampelos (Asphodelaceae subfam. Alooideae) naturalised and escaped in continental Portugal" masterLastPageNumber="52" masterPageNumber="35" pageNumber="39" updateTime="1701715323976" updateUser="jonas" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="DED50E74E41440DCF551E015EF389B18">An annotated catalogue of Aloe and Aloiampelos (Asphodelaceae subfam. Alooideae) naturalised and escaped in continental Portugal</mods:title>
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<mods:namePart id="DDC6DAE1F5CA33CB57862D502E16022C">Smith, Gideon F.</mods:namePart>
<mods:nameIdentifier id="9DC90284B475B7F7646CE95D65C0C3EE" type="ORCID">0000-0002-5417-9208</mods:nameIdentifier>
<mods:affiliation id="E0BEA2225D1FC83B7929718343A2B33E">Ria Olivier Herbarium, Department of Botany, Nelson Mandela University, P. O. Box 77000, Gqeberha, 6031 South Africa. &amp; smithgideon 1 @ gmail. com; https: // orcid. org / 0000 - 0002 - 5417 - 9208</mods:affiliation>
<mods:nameIdentifier id="B43B8317E0EF5F48F58D6EA3FEEAF2FB" type="email">smithgideon1@gmail.com</mods:nameIdentifier>
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<mods:namePart id="13646527D0A07A2D54463EBA9A679999">Figueiredo, Estrela</mods:namePart>
<mods:nameIdentifier id="F68042F560D7E5EB5EDB40F7A577A891" type="ORCID">0000-0002-8511-8213</mods:nameIdentifier>
<mods:affiliation id="EE1756EE848334E4BFDC7AA1C783C052">Ria Olivier Herbarium, Department of Botany, Nelson Mandela University, P. O. Box 77000, Gqeberha, 6031 South Africa. &amp; epnfigueiredo @ gmail. com; https: // orcid. org / 0000 - 0002 - 8511 - 8213</mods:affiliation>
<mods:nameIdentifier id="ED5B206B918BCF724F7A3F51FAD5BE7E" type="email">epnfigueiredo@gmail.com</mods:nameIdentifier>
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<mods:namePart id="4BC942482AEF39EA174C7EB0BF83C9CB">Verloove, Filip</mods:namePart>
<mods:nameIdentifier id="6014AB39AA5D11EBA88541F77A638919" type="ORCID">0000-0003-4144-2422</mods:nameIdentifier>
<mods:affiliation id="EF99F3A3908CB14B19F665783BF95BDB">Meise Botanic Garden, Nieuwelaan 38, B- 1860 Meise, Belgium. &amp; filip. verloove @ plantentuinmeise. be; https: // orcid. org / 0000 - 0003 - 4144 - 2422</mods:affiliation>
<mods:nameIdentifier id="B6F8A016E020ADF438FB1626C32909FC" type="email">filip.verloove@plantentuinmeise.be</mods:nameIdentifier>
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<mods:namePart id="87732CE4A2D578AD48908E643E4DEC7D">Klopper, Ronell R.</mods:namePart>
<mods:affiliation id="5A7B13369A825EF14620E88B791C4593">Foundational Biodiversity Science Division, South African National Biodiversity Institute, Private Bag X 101, Pretoria, 0001 South Africa. &amp; H. G. W. J. Schweickerdt Herbarium, Department of Plant and Soil Sciences, University of Pretoria, Pretoria, 0002 South Africa.</mods:affiliation>
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<mods:namePart id="7BEFDCC8778D80E0B828DA8B481215C4">Silva, Vasco</mods:namePart>
<mods:affiliation id="6777155EF203F53BC2BEAEEF6353F231">Centre for Applied Ecology Prof. Baeta Neves, School of Agriculture, University of Lisbon, Tapada da Ajuda, 1349 - 017 Lisbon, Portugal. &amp; Egas Moniz Center for Interdisciplinary Research (CiiEM), Egas Moniz School of Health and Science, Campus Universitário, Quinta da Granja, Monte da Caparica, 2829 - 511 Caparica, Portugal.</mods:affiliation>
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<mods:title id="99C2F98EF30A5301393D94804820080C">Phytotaxa</mods:title>
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<mods:date id="E3AF76690F332B6BDE756E8CD66F3D51">2023</mods:date>
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<mods:number id="696416B2610BBF2F02F19C07CC6204CD">2023-12-04</mods:number>
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<mods:number id="616B328C6C2D9D55484DFFE59DF6953C">629</mods:number>
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<mods:url id="A5752F5FB1D55CEC2E65CD1B50D43FF8">https://phytotaxa.mapress.com/pt/article/download/phytotaxa.629.1.3/51327</mods:url>
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<paragraph id="3D378577455FFFDE99A8FF30FC870C84" blockId="4.[136,999,158,256]" box="[136,884,158,185]" pageId="4" pageNumber="39">
<heading id="667F321B455FFFDE99A8FF30FC870C84" box="[136,884,158,185]" fontSize="11" level="2" pageId="4" pageNumber="39" reason="4">
2.
<taxonomicName id="FA88FEF4455FFFDE9984FF30FCE10C85" ID-CoL="C3TJ" box="[164,786,158,185]" class="Liliopsida" family="Asphodelaceae" genus="Aloe" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="39" phylum="Tracheophyta" rank="subSpecies" species="maculata" subSpecies="maculata">
<emphasis id="0FFC5965455FFFDE9984FF30FEB80C85" bold="true" box="[164,331,158,185]" italics="true" pageId="4" pageNumber="39">Aloe maculata</emphasis>
<bibRefCitation id="5919F886455FFFDE9871FF30FDE00C85" author="Allioni" box="[337,531,158,185]" firstAuthor="Allioni" page="65" pageId="4" pageNumber="39" pagination="53 - 96" refId="ref8573" refString="Allioni, C. (1773 [?]) Auctarium ad synopsim methodicam stirpium horti reg. taurinensis. Melanges de Philosophie et de Mathematique de la Societe Royale de Turin pour les annees 1770 - 1773 5: 53 - 96. [Preprinted on 30 September 1773]. Available from: https: // www. biodiversitylibrary. org / item / 32648 # page / 85 / mode / 1 up (accessed 1 December 2023)." type="journal article" year="1773">Allioni (1773: 65</bibRefCitation>
[13]) subsp.
<emphasis id="0FFC5965455FFFDE9B87FF30FCE10C85" bold="true" box="[679,786,158,185]" italics="true" pageId="4" pageNumber="39">maculata</emphasis>
</taxonomicName>
(
<figureCitation id="A5B399F2455FFFDE9A02FF30FC940C84" box="[802,871,158,184]" captionStart="FIGURE 2" captionStartId="4.[136,229,1798,1820]" captionTargetBox="[171,1417,844,1755]" captionTargetId="figure-263@4.[151,1436,821,1773]" captionTargetPageId="4" captionText="FIGURE 2. Aloe maculata in Portugal. A. Naturalised at Almograve along the Baixo Alentejo coast. Photograph taken on 29 December 2017. B. Naturalised at Porto Covo along the Alentejo Litoral coast.Photograph taken on 30 December 2017. C. Naturalised at Malveira da Serra, Almoinhas Velhas, in Cascais, near metropolitan Lisbon. Photograph taken on 28 September 2014. D. Close-up of an inflorescence that carries salmon-orange flowers. The racemes are distinctly capitate and flowers have prominent basal swellings. All photographs by Gideon F. Smith." figureDoi="http://doi.org/10.5281/zenodo.10254759" httpUri="https://zenodo.org/record/10254759/files/figure.png" pageId="4" pageNumber="39">Fig. 2</figureCitation>
).
</heading>
</paragraph>
</subSubSection>
<subSubSection id="7592D6FC455FFFDE99A8FF68FC560CE7" box="[136,933,198,220]" pageId="4" pageNumber="39" type="reference_group">
<paragraph id="3D378577455FFFDE99A8FF68FC560CE7" blockId="4.[136,999,158,256]" box="[136,933,198,220]" pageId="4" pageNumber="39">
<emphasis id="0FFC5965455FFFDE99A8FF68FE870CE7" bold="true" box="[136,372,198,219]" pageId="4" pageNumber="39">Locally used synonyms</emphasis>
:—
<taxonomicName id="FA88FEF4455FFFDE98B3FF68FC520CE7" authority="(Aiton 1789: 467) Haworth (1804: 17)" authorityName="Haworth" authorityPageNumber="17" authorityYear="1804" baseAuthorityName="Aiton" baseAuthorityPageNumber="467" baseAuthorityYear="1789" box="[403,929,198,220]" class="Liliopsida" family="Asphodelaceae" genus="Aloe" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="39" phylum="Tracheophyta" rank="species" species="saponaria">
<emphasis id="0FFC5965455FFFDE98B3FF68FDD50CE7" box="[403,550,198,219]" italics="true" pageId="4" pageNumber="39">Aloe saponaria</emphasis>
(Aiton 1789: 467)
<bibRefCitation id="5919F886455FFFDE9BFFFF68FC520CE7" author="Haworth" box="[735,929,198,220]" firstAuthor="Haworth" page="17" pageId="4" pageNumber="39" pagination="1 - 28" refId="ref10946" refString="Haworth, A. H. (1804) A new arrangement of the genus Aloe, with a chronological sketch of the progressive knowledge of that genus, and of other succulent genera. Transactions of the Linnean Society of London 7 (1): 1 - 28. https: // doi. org / 10.1111 / j. 1096 - 3642.1804. tb 00276. x" type="journal article" year="1804">Haworth (1804: 17)</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="7592D6FC455FFFDE99A8FF44FC140CC3" box="[136,999,233,256]" pageId="4" pageNumber="39" type="vernacular_names">
<paragraph id="3D378577455FFFDE99A8FF44FC140CC3" blockId="4.[136,999,158,256]" box="[136,999,233,256]" pageId="4" pageNumber="39">
<emphasis id="0FFC5965455FFFDE99A8FF44FECD0CC3" bold="true" box="[136,318,233,256]" pageId="4" pageNumber="39">Common name(s)</emphasis>
:—
<emphasis id="0FFC5965455FFFDE987DFF44FE390CC3" box="[349,458,234,255]" italics="true" pageId="4" pageNumber="39">Portuguese</emphasis>
: áloe-manchado.
<emphasis id="0FFC5965455FFFDE9B55FF44FD4D0CC3" box="[629,702,234,255]" italics="true" pageId="4" pageNumber="39">English</emphasis>
: soap aloe (
<bibRefCitation id="5919F886455FFFDE9A12FF44FC2F0CC3" author="Grace" box="[818,988,234,255]" etAl="et al." firstAuthor="Grace" pageId="4" pageNumber="39" refId="ref10501" refString="Grace, O. M., Klopper, R. R., Figueiredo, E. &amp; Smith, G. F. (2011) The aloe names book. Strelitzia 29. South African National Biodiversity Institute, Pretoria, and Royal Botanic Gardens, Kew, 232 pp. Available from: http: // hdl. handle. net / 20.500.12143 / 270 (accessed 1 December 2023)." type="book" year="2011">
Grace
<emphasis id="0FFC5965455FFFDE9A53FF44FC6E0CC3" box="[883,925,234,255]" italics="true" pageId="4" pageNumber="39">et al</emphasis>
. 2011
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="7592D6FC455FFFDE99A8FE80FCEB0EB0" pageId="4" pageNumber="39" type="description">
<paragraph id="3D378577455FFFDE99A8FE80FCEB0EB0" blockId="4.[136,1452,302,796]" pageId="4" pageNumber="39">
<emphasis id="0FFC5965455FFFDE99A8FE80FEE10D74" bold="true" box="[136,274,302,328]" pageId="4" pageNumber="39">Description</emphasis>
:—Plants acaulescent or with
<emphasis id="0FFC5965455FFFDE9B47FE81FD690D74" box="[615,666,303,328]" italics="true" pageId="4" pageNumber="39">stem</emphasis>
up to
<quantity id="FA702892455FFFDE9BC5FE80FCD40D75" box="[741,807,302,329]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" pageId="4" pageNumber="39" unit="m" value="0.5">0.5 m</quantity>
tall;
<emphasis id="0FFC5965455FFFDE9A43FE81FC4A0D74" box="[867,953,303,328]" italics="true" pageId="4" pageNumber="39">rosettes</emphasis>
solitary or suckering to
<taxonomicName id="FA88FEF4455FFFDE9DF1FE80FAA30D74" box="[1233,1360,302,328]" form="dense" pageId="4" pageNumber="39" rank="form">form dense</taxonomicName>
groups.
<emphasis id="0FFC5965455FFFDE99A8FEFDFF250D50" box="[136,214,339,364]" italics="true" pageId="4" pageNumber="39">Leaves</emphasis>
densely rosulate, erectly spreading to slightly recurved, up to 2530 ×
<quantity id="FA702892455FFFDE9D14FEFCFB6D0D51" box="[1076,1182,338,365]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" metricValueMax="1.2" metricValueMin="0.8" pageId="4" pageNumber="39" unit="cm" value="10.0" valueMax="12.0" valueMin="8.0">812 cm</quantity>
, ovate-lanceolate, with dried twisted apex, adaxial surface pale to darker green, with numerous, dull, H-shaped or irregularly shaped, white maculations in irregular broken, wavy, transverse bands, abaxial surface paler green, obscurely lineate and usually without maculations;
<emphasis id="0FFC5965455FFFDE98A0FE11FE230DE4" box="[384,464,447,472]" italics="true" pageId="4" pageNumber="39">margin</emphasis>
horny, brown;
<emphasis id="0FFC5965455FFFDE9BA2FE10FCD40DE4" box="[642,807,446,472]" italics="true" pageId="4" pageNumber="39">marginal teeth</emphasis>
±
<quantity id="FA702892455FFFDE9A67FE10FC6A0DE5" box="[839,921,446,473]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="39" unit="mm" value="10.0">10 mm</quantity>
apart,
<quantity id="FA702892455FFFDE9ACAFE10FBB90DE5" box="[1002,1098,446,473]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="4" pageNumber="39" unit="mm" value="4.0" valueMax="5.0" valueMin="3.0">35 mm</quantity>
long, pungent, reddish brown;
<emphasis id="0FFC5965455FFFDE99A8FE4CFF2C0DC0" box="[136,223,482,508]" italics="true" pageId="4" pageNumber="39">exudate</emphasis>
clear.
<emphasis id="0FFC5965455FFFDE980FFE4CFE360DC0" box="[303,453,482,508]" italics="true" pageId="4" pageNumber="39">Inflorescence</emphasis>
0.41.0 m high, erect, branched.
<emphasis id="0FFC5965455FFFDE9A6DFE4DFC420DC0" box="[845,945,483,508]" italics="true" pageId="4" pageNumber="39">Racemes</emphasis>
<quantity id="FA702892455FFFDE9A9AFE4CFBDF0DC1" box="[954,1068,482,509]" metricMagnitude="-1" metricUnit="m" metricValue="1.1" metricValueMax="1.2" metricValueMin="1.0" pageId="4" pageNumber="39" unit="cm" value="11.0" valueMax="12.0" valueMin="10.0">1012 cm</quantity>
long, capitate-corymbose, dense.
<emphasis id="0FFC5965455FFFDE99A8FDA8FEED0E1C" box="[136,286,518,544]" italics="true" pageId="4" pageNumber="39">Floral bracts</emphasis>
± 1223 ×
<quantity id="FA702892455FFFDE9882FDA8FDF00E1D" box="[418,515,518,545]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="4" pageNumber="39" unit="mm" value="4.0" valueMax="5.0" valueMin="3.0">35 mm</quantity>
.
<emphasis id="0FFC5965455FFFDE9B2EFDA8FD9E0E1C" box="[526,621,518,544]" italics="true" pageId="4" pageNumber="39">Pedicels</emphasis>
<quantity id="FA702892455FFFDE9B55FDA8FD1C0E1D" box="[629,751,518,545]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" metricValueMax="4.5" metricValueMin="3.5" pageId="4" pageNumber="39" unit="mm" value="40.0" valueMax="45.0" valueMin="35.0">3545 mm</quantity>
long.
<emphasis id="0FFC5965455FFFDE9A17FDA8FC600E1C" box="[823,915,518,544]" italics="true" pageId="4" pageNumber="39">Flowers</emphasis>
:
<emphasis id="0FFC5965455FFFDE9A83FDA8FBF10E1C" box="[931,1026,518,544]" italics="true" pageId="4" pageNumber="39">perianth</emphasis>
<quantity id="FA702892455FFFDE9D2AFDA8FB770E1D" box="[1034,1156,518,545]" metricMagnitude="-2" metricUnit="m" metricValue="4.25" metricValueMax="5.0" metricValueMin="3.5" pageId="4" pageNumber="39" unit="mm" value="42.5" valueMax="50.0" valueMin="35.0">3550 mm</quantity>
long, up to
<quantity id="FA702892455FFFDE9C2EFDA8FAAD0E1D" box="[1294,1374,518,545]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="39" unit="mm" value="10.0">10 mm</quantity>
across ovary, abruptly constricted above ovary to form sub-globose basal swelling, enlarging towards wide open mouth, slightly decurved, usually salmon pink to orange;
<emphasis id="0FFC5965455FFFDE9B97FDE1FC930E54" box="[695,864,591,616]" italics="true" pageId="4" pageNumber="39">outer segments</emphasis>
free for
<quantity id="FA702892455FFFDE9AE1FDE0FBCF0E55" box="[961,1084,590,617]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="4" pageNumber="39" unit="mm" value="12.5" valueMax="15.0" valueMin="10.0">1015 mm</quantity>
;
<emphasis id="0FFC5965455FFFDE9D68FDE1FB510E54" box="[1096,1186,591,616]" italics="true" pageId="4" pageNumber="39">stamens</emphasis>
exserted
<quantity id="FA702892455FFFDE9C2DFDE0FA9E0E55" box="[1293,1389,590,617]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="4" pageNumber="39" unit="mm" value="2.0" valueMax="3.0" valueMin="1.0">13 mm</quantity>
;
<emphasis id="0FFC5965455FFFDE9C58FDE0FA580E54" box="[1400,1451,590,616]" italics="true" pageId="4" pageNumber="39">style</emphasis>
exserted to
<quantity id="FA702892455FFFDE9829FDDCFEBE0EB1" box="[265,333,626,653]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="4" pageNumber="39" unit="mm" value="5.0">5 mm</quantity>
.
<emphasis id="0FFC5965455FFFDE9878FDDCFE470EB0" box="[344,436,626,652]" italics="true" pageId="4" pageNumber="39">Capsule</emphasis>
(2.0)2.53.0 cm long, oblong.
</paragraph>
</subSubSection>
<subSubSection id="7592D6FC455FFFDE999DFD38FAC00F20" pageId="4" pageNumber="39" type="biology_ecology">
<paragraph id="3D378577455FFFDE999DFD38FDE70E8C" blockId="4.[136,1452,302,796]" box="[189,532,662,688]" pageId="4" pageNumber="39">
<emphasis id="0FFC5965455FFFDE999DFD38FE800E8C" bold="true" box="[189,371,662,688]" pageId="4" pageNumber="39">Flowering time</emphasis>
:—AprilJuly.
</paragraph>
<paragraph id="3D378577455FFFDE999DFD14FD150EC4" blockId="4.[136,1452,302,796]" pageId="4" pageNumber="39">
<emphasis id="0FFC5965455FFFDE999DFD14FEEA0EE8" bold="true" box="[189,281,698,724]" pageId="4" pageNumber="39">Habitat</emphasis>
:—On old (stone) walls and in abandoned gardens (
<bibRefCitation id="5919F886455FFFDE9A5AFD14FB920EE8" author="Almeida" box="[890,1121,698,724]" firstAuthor="Almeida" page="209" pageId="4" pageNumber="39" refId="ref8656" refString="Almeida, J. D. de. (2012) Flora exotica subespontanea de Portugal continental (plantas vasculares). 5. a edicao. M. Sc. thesis. University of Coimbra, Coimbra, 281 pp. Available from: https: // docplayer. com. br / 48643620 - Flora-exotica-subespontanea-de-portugalcontinental-plantas-vasculares. html (accessed 1 December 2023)." type="book" year="2012">Almeida 2012: 209</bibRefCitation>
). A range of sites in natural vegetation, including on coastal cliffs and sand dunes.
</paragraph>
<paragraph id="3D378577455FFFDE999DFCACFAC00F20" blockId="4.[136,1452,302,796]" box="[189,1331,770,796]" pageId="4" pageNumber="39">
<emphasis id="0FFC5965455FFFDE999DFCACFCB30F20" bold="true" box="[189,832,770,796]" pageId="4" pageNumber="39">
First mention of the species as naturalised in
<collectingCountry id="459FC5E7455FFFDE9BF8FCACFCB30F20" box="[728,832,770,796]" name="Portugal" pageId="4" pageNumber="39">Portugal</collectingCountry>
</emphasis>
:—1998 (
<emphasis id="0FFC5965455FFFDE9A8CFCACFC250F20" box="[940,982,770,796]" italics="true" pageId="4" pageNumber="39">fide</emphasis>
<bibRefCitation id="5919F886455FFFDE9AFBFCACFAD40F20" author="Almeida &amp; Freitas" box="[987,1319,770,796]" firstAuthor="Almeida" page="119" pageId="4" pageNumber="39" pagination="117 - 130" refId="ref8722" refString="Almeida, J. D. de &amp; Freitas, H. (2006) Exotic naturalized flora of continental Portugal. A reassessment. Botanica Complutensis 30: 117 - 130." type="journal article" year="2006">Almeida &amp; Freitas 2006: 119</bibRefCitation>
).
</paragraph>
</subSubSection>
<caption id="69F7D5FF455FFFDE99A8F8A8FEDE0B90" ID-DOI="http://doi.org/10.5281/zenodo.10254759" ID-Zenodo-Dep="10254759" httpUri="https://zenodo.org/record/10254759/files/figure.png" pageId="4" pageNumber="39" startId="4.[136,229,1798,1820]" targetBox="[171,1417,844,1755]" targetPageId="4" targetType="figure">
<paragraph id="3D378577455FFFDE99A8F8A8FEDE0B90" blockId="4.[136,1452,1798,1964]" pageId="4" pageNumber="39">
<emphasis id="0FFC5965455FFFDE99A8F8A8FF0D0B27" bold="true" box="[136,254,1798,1820]" pageId="4" pageNumber="39">FIGURE 2.</emphasis>
<taxonomicName id="FA88FEF4455FFFDE9824F8A8FE7C0B27" authorityName="Allioni" authorityYear="1773" box="[260,399,1798,1819]" class="Liliopsida" family="Asphodelaceae" genus="Aloe" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="39" phylum="Tracheophyta" rank="species" species="maculata">
<emphasis id="0FFC5965455FFFDE9824F8A8FE7C0B27" box="[260,399,1798,1819]" italics="true" pageId="4" pageNumber="39">Aloe maculata</emphasis>
</taxonomicName>
in Portugal.
<emphasis id="0FFC5965455FFFDE9B2DF8A8FDD60B27" bold="true" box="[525,549,1798,1819]" pageId="4" pageNumber="39">A.</emphasis>
Naturalised at Almograve along the Baixo Alentejo coast. Photograph taken on 29 December 2017.
<emphasis id="0FFC5965455FFFDE99E3F884FF2A0B03" bold="true" box="[195,217,1834,1855]" pageId="4" pageNumber="39">B.</emphasis>
Naturalised at Porto Covo along the Alentejo Litoral coast. Photograph taken on 30 December 2017.
<emphasis id="0FFC5965455FFFDE9DB4F884FB580B7C" bold="true" box="[1172,1195,1834,1856]" pageId="4" pageNumber="39">C.</emphasis>
Naturalised at Malveira da Serra, Almoinhas Velhas, in Cascais, near metropolitan Lisbon. Photograph taken on 28 September 2014.
<emphasis id="0FFC5965455FFFDE9D5CF8E0FB670B5F" bold="true" box="[1148,1172,1870,1891]" pageId="4" pageNumber="39">D.</emphasis>
Close-up of an inflorescence that carries salmon-orange flowers. The racemes are distinctly capitate and flowers have prominent basal swellings. All photographs by Gideon F. Smith.
</paragraph>
</caption>
<caption id="69F7D5FF455EFFDF99A8FADEFE340A62" ID-DOI="http://doi.org/10.5281/zenodo.10254761" ID-Zenodo-Dep="10254761" httpUri="https://zenodo.org/record/10254761/files/figure.png" pageId="5" pageNumber="40" startId="5.[136,229,1392,1414]" targetBox="[178,1413,217,1332]" targetPageId="5" targetType="figure">
<paragraph id="3D378577455EFFDF99A8FADEFE340A62" blockId="5.[136,1452,1392,1630]" pageId="5" pageNumber="40">
<emphasis id="0FFC5965455EFFDF99A8FADEFF0C09B9" bold="true" box="[136,255,1392,1414]" pageId="5" pageNumber="40">FIGURE 3.</emphasis>
Two nothospecies with which
<taxonomicName id="FA88FEF4455EFFDF9B0EFADEFD4909B9" authorityName="Allioni" authorityYear="1773" box="[558,698,1392,1413]" class="Liliopsida" family="Asphodelaceae" genus="Aloe" kingdom="Plantae" order="Asparagales" pageId="5" pageNumber="40" phylum="Tracheophyta" rank="species" species="maculata">
<emphasis id="0FFC5965455EFFDF9B0EFADEFD4909B9" box="[558,698,1392,1413]" italics="true" pageId="5" pageNumber="40">Aloe maculata</emphasis>
</taxonomicName>
can be confused.
<emphasis id="0FFC5965455EFFDF9A4BFADEFC7009B9" bold="true" box="[875,899,1392,1413]" pageId="5" pageNumber="40">A.</emphasis>
<taxonomicName id="FA88FEF4455EFFDF9AAAFADEFBDA09B9" authorityName="Todaro" authorityYear="1878" box="[906,1065,1392,1414]" class="Liliopsida" family="Asphodelaceae" genus="Aloe" isHybrid="true" kingdom="Plantae" order="Asparagales" pageId="5" pageNumber="40" phylum="Tracheophyta" rank="species" species="schimperi">
<emphasis id="0FFC5965455EFFDF9AAAFADEFC4509B9" box="[906,950,1392,1413]" italics="true" pageId="5" pageNumber="40">Aloe</emphasis>
×
<emphasis id="0FFC5965455EFFDF9AEAFADEFBDA09B9" box="[970,1065,1392,1413]" italics="true" pageId="5" pageNumber="40">schimperi</emphasis>
</taxonomicName>
[
<taxonomicName id="FA88FEF4455EFFDF9D18FADFFB5E09B9" authorityName="Allioni" authorityYear="1773" box="[1080,1197,1392,1414]" class="Liliopsida" family="Asphodelaceae" genus="Aloe" kingdom="Plantae" order="Asparagales" pageId="5" pageNumber="40" phylum="Tracheophyta" rank="species" species="maculata">
<emphasis id="0FFC5965455EFFDF9D18FADFFB5E09B9" box="[1080,1197,1392,1414]" italics="true" pageId="5" pageNumber="40">A. maculata</emphasis>
</taxonomicName>
×
<taxonomicName id="FA88FEF4455EFFDF9DE9FADFFAD709BA" authorityName="Haworth" authorityYear="1804" box="[1225,1316,1393,1414]" class="Liliopsida" family="Asphodelaceae" genus="Aloe" kingdom="Plantae" order="Asparagales" pageId="5" pageNumber="40" phylum="Tracheophyta" rank="species" species="striata">
<emphasis id="0FFC5965455EFFDF9DE9FADFFAD709BA" box="[1225,1316,1393,1414]" italics="true" pageId="5" pageNumber="40">A. striata</emphasis>
</taxonomicName>
] cultivated in the Botanical Garden of the University of Porto, Porto. Note the distinctly pinkish leaf margins, a character inherited from
<taxonomicName id="FA88FEF4455EFFDF9C6AFA3BFA550996" authorityName="Haworth" authorityYear="1804" box="[1354,1446,1429,1450]" class="Liliopsida" family="Asphodelaceae" genus="Aloe" kingdom="Plantae" order="Asparagales" pageId="5" pageNumber="40" phylum="Tracheophyta" rank="species" species="striata">
<emphasis id="0FFC5965455EFFDF9C6AFA3BFA550996" box="[1354,1446,1429,1450]" italics="true" pageId="5" pageNumber="40">A. striata</emphasis>
</taxonomicName>
.
<emphasis id="0FFC5965455EFFDF99A8FA17FF6D09F2" bold="true" box="[136,158,1465,1486]" pageId="5" pageNumber="40">B.</emphasis>
The flowers of
<taxonomicName id="FA88FEF4455EFFDF981EFA17FE3409F1" authorityName="Todaro" authorityYear="1878" box="[318,455,1464,1486]" class="Liliopsida" family="Asphodelaceae" genus="Aloe" isHybrid="true" kingdom="Plantae" order="Asparagales" pageId="5" pageNumber="40" phylum="Tracheophyta" rank="species" species="schimperi">
<emphasis id="0FFC5965455EFFDF981EFA17FEBE09F2" box="[318,333,1465,1486]" italics="true" pageId="5" pageNumber="40">A</emphasis>
. ×
<emphasis id="0FFC5965455EFFDF9848FA16FE3409F1" box="[360,455,1464,1485]" italics="true" pageId="5" pageNumber="40">schimperi</emphasis>
</taxonomicName>
lack the prominent basal swellings present in
<taxonomicName id="FA88FEF4455EFFDF9AB0FA17FBF509F1" authorityName="Allioni" authorityYear="1773" box="[912,1030,1464,1486]" class="Liliopsida" family="Asphodelaceae" genus="Aloe" kingdom="Plantae" order="Asparagales" pageId="5" pageNumber="40" phylum="Tracheophyta" rank="species" species="maculata">
<emphasis id="0FFC5965455EFFDF9AB0FA17FBF509F1" box="[912,1030,1464,1486]" italics="true" pageId="5" pageNumber="40">A. maculata</emphasis>
</taxonomicName>
. Both photographs A and B were taken on 22 December 2017.
<emphasis id="0FFC5965455EFFDF986BFA72FE9009CE" bold="true" box="[331,355,1500,1522]" pageId="5" pageNumber="40">C.</emphasis>
The rosettes of
<taxonomicName id="FA88FEF4455EFFDF98DEFA73FD6209CE" box="[510,657,1501,1522]" class="Liliopsida" family="Asphodelaceae" genus="Aloe" isHybrid="true" kingdom="Plantae" order="Asparagales" pageId="5" pageNumber="40" phylum="Tracheophyta" rank="species" species="commutata">
<emphasis id="0FFC5965455EFFDF98DEFA73FDFE09CE" box="[510,525,1501,1522]" italics="true" pageId="5" pageNumber="40">A</emphasis>
. ×
<emphasis id="0FFC5965455EFFDF9B06FA73FD6209CE" box="[550,657,1501,1522]" italics="true" pageId="5" pageNumber="40">commutata</emphasis>
</taxonomicName>
[
<taxonomicName id="FA88FEF4455EFFDF9BBFFA73FCB109CD" box="[671,834,1500,1522]" class="Liliopsida" family="Asphodelaceae" genus="Aloe" kingdom="Plantae" order="Asparagales" pageId="5" pageNumber="40" phylum="Tracheophyta" rank="species" species="grandidentata">
<emphasis id="0FFC5965455EFFDF9BBFFA73FCB109CD" box="[671,834,1500,1522]" italics="true" pageId="5" pageNumber="40">A. grandidentata</emphasis>
</taxonomicName>
×
<taxonomicName id="FA88FEF4455EFFDF9A7CFA73FC2309CD" authorityName="Allioni" authorityYear="1773" box="[860,976,1500,1522]" class="Liliopsida" family="Asphodelaceae" genus="Aloe" kingdom="Plantae" order="Asparagales" pageId="5" pageNumber="40" phylum="Tracheophyta" rank="species" species="maculata">
<emphasis id="0FFC5965455EFFDF9A7CFA73FC2309CD" box="[860,976,1500,1522]" italics="true" pageId="5" pageNumber="40">A. maculata</emphasis>
</taxonomicName>
] resemble those of
<taxonomicName id="FA88FEF4455EFFDF9DAEFA73FAF109CD" authorityName="Allioni" authorityYear="1773" box="[1166,1282,1500,1522]" class="Liliopsida" family="Asphodelaceae" genus="Aloe" kingdom="Plantae" order="Asparagales" pageId="5" pageNumber="40" phylum="Tracheophyta" rank="species" species="maculata">
<emphasis id="0FFC5965455EFFDF9DAEFA73FAF109CD" box="[1166,1282,1500,1522]" italics="true" pageId="5" pageNumber="40">A. maculata</emphasis>
</taxonomicName>
, but, as shown in
<emphasis id="0FFC5965455EFFDF99A8F9AFFF6A0A2A" bold="true" box="[136,153,1537,1558]" pageId="5" pageNumber="40">D</emphasis>
, its inflorescences are elongated (not capitate) and the flowers of this nothospecies, too, lack basal swellings, as found in
<taxonomicName id="FA88FEF4455EFFDF9C25F9AFFA560A29" box="[1285,1445,1536,1558]" class="Liliopsida" family="Asphodelaceae" genus="Aloe" kingdom="Plantae" order="Asparagales" pageId="5" pageNumber="40" phylum="Tracheophyta" rank="species" species="grandidentata">
<emphasis id="0FFC5965455EFFDF9C25F9AFFA560A29" box="[1285,1445,1536,1558]" italics="true" pageId="5" pageNumber="40">A. grandidentata</emphasis>
</taxonomicName>
. Both photographs C and D were taken in the Huntington Botanical Gardens, San Marino, California, U.S.A., on 27 April 2011. All photographs by Gideon F. Smith.
</paragraph>
</caption>
<subSubSection id="7592D6FC455EFFDC999DF922FEEC0DAD" lastPageId="6" lastPageNumber="41" pageId="5" pageNumber="40" type="distribution">
<paragraph id="3D378577455EFFDF999DF922FC4F0B2F" blockId="5.[136,1452,1676,2063]" pageId="5" pageNumber="40">
<emphasis id="0FFC5965455EFFDF999DF922FE500A9B" bold="true" box="[189,419,1676,1703]" pageId="5" pageNumber="40">Localities recorded</emphasis>
:—
<bibRefCitation id="5919F886455EFFDF98E4F923FD5E0A9B" author="Almeida" box="[452,685,1677,1703]" firstAuthor="Almeida" page="209" pageId="5" pageNumber="40" refId="ref8656" refString="Almeida, J. D. de. (2012) Flora exotica subespontanea de Portugal continental (plantas vasculares). 5. a edicao. M. Sc. thesis. University of Coimbra, Coimbra, 281 pp. Available from: https: // docplayer. com. br / 48643620 - Flora-exotica-subespontanea-de-portugalcontinental-plantas-vasculares. html (accessed 1 December 2023)." type="book" year="2012">Almeida (2012: 209</bibRefCitation>
, as “
<taxonomicName id="FA88FEF4455EFFDF9BC4F923FC620A9B" authorityName="Haworth" authorityYear="1804" baseAuthorityName="Aiton" baseAuthorityYear="1789" box="[740,913,1677,1703]" class="Liliopsida" family="Asphodelaceae" genus="Aloe" kingdom="Plantae" order="Asparagales" pageId="5" pageNumber="40" phylum="Tracheophyta" rank="species" species="saponaria">
<emphasis id="0FFC5965455EFFDF9BC4F923FC620A9B" box="[740,913,1677,1703]" italics="true" pageId="5" pageNumber="40">Aloe saponaria</emphasis>
</taxonomicName>
”) recorded it as casual in BL (Alhadas) and Ag (Sagres).
<bibRefCitation id="5919F886455EFFDF99D3F91FFE2A0AF7" author="Guemes" box="[243,473,1713,1739]" firstAuthor="Guemes" page="314" pageId="5" pageNumber="40" pagination="308 - 314" refId="ref10660" refString="Guemes, J. (2013) Aloe L. In: E. Rico, M. B. Crespo, A. Quintanar, A. Herrero &amp; C. Aedo (Eds.) Flora iberica 20. Real Jardin Botanico, CSIC, Madrid, pp. 308 - 314. Available from: http: // www. floraiberica. es / floraiberica / texto / pdfs / 20 _ 183 _ 32 _ Aloe. pdf (accessed 1 December 2023)." type="book chapter" year="2013">Güemes (2013: 314)</bibRefCitation>
recorded it for E (Serra dEl Rei), BAl, and Ag.
<bibRefCitation id="5919F886455EFFDF9ADAF91FFAE10AF7" author="Silva" box="[1018,1298,1713,1740]" etAl="et al." firstAuthor="Silva" page="69 - 70" pageId="5" pageNumber="40" pagination="58 - 81" refId="ref13229" refString="Silva, V., Figueiredo, E. &amp; Smith, G. F. (2015) Alien succulents naturalised and cultivated on the central west coast of Portugal. Bradleya 33: 58 - 81. https: // doi. org / 10.25223 / brad. n 33.2015. a 10" type="journal article" year="2015">
Silva
<emphasis id="0FFC5965455EFFDF9D1AF91CFB9F0AF7" box="[1082,1132,1713,1739]" italics="true" pageId="5" pageNumber="40">et al</emphasis>
. (2015: 6970
</bibRefCitation>
, 75) recorded it as casual and escaped at Cascais, Oeiras, north of
<collectingRegion id="FF4C4B95455EFFDF9BF2F97BFCD00AD3" box="[722,803,1749,1775]" country="Portugal" name="Lisboa" pageId="5" pageNumber="40">Lisbon</collectingRegion>
. It is here recorded at the following further localities; São João do Estoril (E); Almograve,
<collectingRegion id="FF4C4B95455EFFDF98D4F957FDC20B2F" box="[500,561,1785,1811]" country="Portugal" name="Porto" pageId="5" pageNumber="40">Porto</collectingRegion>
Covo (BAl),
<collectingRegion id="FF4C4B95455EFFDF9BEBF957FCF50B2F" box="[715,774,1785,1811]" country="South Africa" name="Western Cape" pageId="5" pageNumber="40">Cabo</collectingRegion>
de Sagres (Ag).
</paragraph>
<paragraph id="3D378577455EFFDF999DF8B2FE4E0B43" blockId="5.[136,1452,1676,2063]" pageId="5" pageNumber="40">
<emphasis id="0FFC5965455EFFDF999DF8B2FDF00B0A" bold="true" box="[189,515,1820,1846]" pageId="5" pageNumber="40">Country / region of origin</emphasis>
:—Southern Africa. It occurs from the
<collectingRegion id="FF4C4B95455EFFDF9AEFF8B3FBFA0B0B" box="[975,1033,1821,1847]" country="South Africa" name="Western Cape" pageId="5" pageNumber="40">Cape</collectingRegion>
Peninsula through the provinces of Western and
<collectingRegion id="FF4C4B95455EFFDF983AF8EFFE420B67" box="[282,433,1857,1883]" country="South Africa" name="Eastern Cape" pageId="5" pageNumber="40">Eastern Cape</collectingRegion>
, into the eastern
<collectingRegion id="FF4C4B95455EFFDF9B4EF8EFFD110B67" box="[622,738,1857,1883]" country="South Africa" name="Free State" pageId="5" pageNumber="40">Free State</collectingRegion>
, through
<collectingRegion id="FF4C4B95455EFFDF9A6BF8EFFC080B67" box="[843,1019,1857,1883]" country="South Africa" name="KwaZulu-Natal" pageId="5" pageNumber="40">KwaZulu-Natal</collectingRegion>
to
<collectingRegion id="FF4C4B95455EFFDF9D3FF8EFFB450B67" box="[1055,1206,1857,1883]" country="South Africa" name="Mpumalanga" pageId="5" pageNumber="40">Mpumalanga</collectingRegion>
,
<collectingCountry id="459FC5E7455EFFDF9DE0F8EFFAA20B67" box="[1216,1361,1857,1883]" name="South Africa" pageId="5" pageNumber="40">South Africa</collectingCountry>
, as well as in
<collectingCountry id="459FC5E7455EFFDF99E3F8CBFEEE0B43" box="[195,285,1893,1919]" name="Lesotho" pageId="5" pageNumber="40">Lesotho</collectingCountry>
and
<collectingCountry id="459FC5E7455EFFDF9874F8CBFE490B43" box="[340,442,1893,1919]" name="Swaziland" pageId="5" pageNumber="40">Eswatini</collectingCountry>
.
</paragraph>
<paragraph id="3D378577455EFFDC999DF826FE870D18" blockId="5.[136,1452,1676,2063]" lastBlockId="6.[136,1452,158,1301]" lastPageId="6" lastPageNumber="41" pageId="5" pageNumber="40">
<emphasis id="0FFC5965455EFFDF999DF826FDD00B9F" bold="true" box="[189,547,1928,1955]" pageId="5" pageNumber="40">Secondary distribution range</emphasis>
:—
<taxonomicName id="FA88FEF4455EFFDF9B68F827FD1D0B9F" authorityName="Allioni" authorityYear="1773" box="[584,750,1929,1955]" class="Liliopsida" family="Asphodelaceae" genus="Aloe" kingdom="Plantae" order="Asparagales" pageId="5" pageNumber="40" phylum="Tracheophyta" rank="species" species="maculata">
<emphasis id="0FFC5965455EFFDF9B68F827FD1D0B9F" box="[584,750,1929,1955]" italics="true" pageId="5" pageNumber="40">Aloe maculata</emphasis>
</taxonomicName>
is very sparingly naturalised in areas with a Mediterranean climate. It is reportedly known as such from the western Mediterranean area:
<collectingCountry id="459FC5E7455EFFDF9D25F803FBA20BFB" box="[1029,1105,1965,1991]" name="France" pageId="5" pageNumber="40">France</collectingCountry>
(Provence and
<collectingCountry id="459FC5E7455EFFDF9C27F803FA930BFB" box="[1287,1376,1965,1991]" name="France" pageId="5" pageNumber="40">Corsica</collectingCountry>
;
<bibRefCitation id="5919F886455EFFDF9C4DF803FE810BD7" author="Tison &amp; de Foucault" firstAuthor="Tison" pageId="5" pageNumber="40" pagination="299 - 301" refId="ref14566" refString="Tison, J. - M. &amp; de Foucault, B. (2014) Xanthorrhoeaceae. Flora Gallica. Flore de France. Biotope Editions, Meze, pp. 299 - 301, whole work 1195 pp." type="book chapter" year="2014">Tison &amp; de Foucault 2014</bibRefCitation>
),
<collectingCountry id="459FC5E7455EFFDF98A8F87FFE3B0BD7" box="[392,456,2001,2027]" name="Spain" pageId="5" pageNumber="40">Spain</collectingCountry>
(
<bibRefCitation id="5919F886455EFFDF98F8F87FFD070BD7" author="Guillot Ortiz" box="[472,756,2001,2027]" etAl="et al." firstAuthor="Guillot Ortiz" pageId="5" pageNumber="40" pagination="1 - 58" refId="ref10880" refString="Guillot Ortiz, D., Laguna Lumbreras, E. &amp; Rossello Picornell, J. A. (2008) La familia Aloaceae en la flora aloctona valenciana. Bouteloua 6: 1 - 58. Available from: http: // www. floramontiberica. org / Bouteloua / MonogBouteloua _ 06 _ Aloaceae. pdf (accessed 1 December 2023)." type="journal article" year="2008">
Guillot Ortiz
<emphasis id="0FFC5965455EFFDF9B55F87CFD5A0BD7" box="[629,681,2001,2027]" italics="true" pageId="5" pageNumber="40">et al</emphasis>
. 2008
</bibRefCitation>
,
<bibRefCitation id="5919F886455EFFDF9BDFF87FFBFC0BD7" author="Aymerich &amp; Saez" box="[767,1039,2001,2027]" firstAuthor="Aymerich" pageId="5" pageNumber="40" pagination="215 - 242" refId="ref8816" refString="Aymerich, P. &amp; Saez, L. (2019) Checklist of the vascular alien flora of Catalonia (northeastern Iberian Peninsula, Spain). Mediterranean Botany 40 (2): 215 - 242. https: // doi. org / 10.5209 / mbot. 63608" type="journal article" year="2019">Aymerich &amp; Sáez 2019</bibRefCitation>
),
<collectingCountry id="459FC5E7455EFFDF9D05F87FFB790BD7" box="[1061,1162,2001,2027]" name="Gibraltar" pageId="5" pageNumber="40">Gibraltar</collectingCountry>
(
<bibRefCitation id="5919F886455EFFDF9DBAF87FFAD00BD7" author="Lamb" box="[1178,1315,2001,2027]" firstAuthor="Lamb" pageId="5" pageNumber="40" pagination="267 - 272" refId="ref11505" refString="Lamb, B. M. (1996) The genus Aloe (L.) in Gibraltar. Almoraima 15: 267 - 272." type="journal article" year="1996">Lamb 1996</bibRefCitation>
), and
<collectingCountry id="459FC5E7455EFFDF9C4AF87FFA580BD7" box="[1386,1451,2001,2027]" name="Malta" pageId="5" pageNumber="40">Malta</collectingCountry>
(
<bibRefCitation id="5919F886455EFFDF99AFF85BFED60433" author="Mifsud" box="[143,293,2037,2063]" firstAuthor="Mifsud" pageId="5" pageNumber="40" refId="ref11882" refString="Mifsud, S. (2022) Malta wild plants. Available online at: https: // maltawildplants. com (accessed November 2022)." type="url" year="2022">Mifsud 2022</bibRefCitation>
; erroneously as
<taxonomicName id="FA88FEF4455EFFDF98C3F85BFC9E0433" box="[483,877,2037,2063]" class="Liliopsida" family="Asphodelaceae" genus="Aloe" kingdom="Plantae" order="Asparagales" pageId="5" pageNumber="40" phylum="Tracheophyta" rank="subSpecies" species="microstigma" subSpecies="microstigma">
<emphasis id="0FFC5965455EFFDF98C3F85BFD7E0433" box="[483,653,2037,2063]" italics="true" pageId="5" pageNumber="40">A. microstigma</emphasis>
subsp.
<emphasis id="0FFC5965455EFFDF9BC2F858FC9E0433" box="[738,877,2038,2063]" italics="true" pageId="5" pageNumber="40">microstigma</emphasis>
</taxonomicName>
). In
<collectingCountry id="459FC5E7455EFFDF9A8CF85BFC130433" box="[940,992,2037,2063]" name="Italy" pageId="5" pageNumber="40">Italy</collectingCountry>
, it is a mere casual (
<bibRefCitation id="5919F886455EFFDF9DE3F85BFA530433" author="Galasso, G. &amp; Conti, F. &amp; Peruzzi, L. &amp; Ardenghi, N. M. G. &amp; Roma-Marzio, F. &amp; Rosati, L. &amp; Santangelo, A. &amp; Scoppola, A. &amp; Scortegagna, S. &amp; Selvaggi, A. &amp; Selvi, F. &amp; Soldano, A. &amp; Stinca, A. &amp; Wagensommer, R. P. &amp; Wilhalm, T. &amp; Bartolucci, F." box="[1219,1440,2037,2063]" journalOrPublisher="Plant Biosystems" pageId="5" pageNumber="40" pagination="556 - 592" part="152" refId="ref10101" refString="Galasso, G., Conti, F., Peruzzi, L., Ardenghi, N. M. G., Banfi, E., Celesti-Grapow, L., Albano, A., Alessandrini, A., Bacchetta, G., Ballelli, S., Bandini Mazzanti, M., Barberis, G., Bernardo, L., Blasi, C., Bouvet, D., Bovio, M., Cecchi, L., Del Guacchio, E., Domina, G., Fascetti, S., Gallo, L., Gubellini, L., Guiggi, A., Iamonico, D., Iberite, M., Jimenez-Mejias, P., Lattanzi, E., Marchetti, D., Martinetto, E., Masin, R. R., Medagli, P., Passalacqua, N. G., Peccenini, S., Pennesi, R., Pierini, B., Podda, L., Poldini, L., Prosser, F., Raimondo, F. M., Roma-Marzio, F., Rosati, L., Santangelo, A., Scoppola, A., Scortegagna, S., Selvaggi, A., Selvi, F., Soldano, A., Stinca, A., Wagensommer, R. P., Wilhalm, T. &amp; Bartolucci, F. (2018) An updated checklist of the vascular flora alien to Italy. Plant Biosystems 152 (3): 556 - 592. https: // doi. org / 10.1080 / 11263504.2018.1441197" title="An updated checklist of the vascular flora alien to Italy" type="journal article" year="2018">
Galasso
<emphasis id="0FFC5965455EFFDF9C04F858FAA60433" box="[1316,1365,2037,2063]" italics="true" pageId="5" pageNumber="40">et al</emphasis>
. 2018
</bibRefCitation>
). It is furthermore locally naturalised in the Canary Islands (unpublished records by FV) and in
<collectingCountry id="459FC5E7455DFFDC9DC9FF30FACE0C84" box="[1257,1341,158,184]" name="Algeria" pageId="6" pageNumber="41">Algeria</collectingCountry>
in North Africa (
<bibRefCitation id="5919F886455DFFDC99C1FF6CFE2C0CE0" author="Sakhraoui" box="[225,479,194,221]" etAl="et al." firstAuthor="Sakhraoui" pageId="6" pageNumber="41" pagination="225 - 229" refId="ref12871" refString="Sakhraoui, N., Verloove, F. &amp; Smith, G. F. (2023) Aloe maculata All. (Asphodelaceae subfam. Alooideae): a new addition to the alien flora of Algeria and North Africa. Bradleya 41: 225 - 229. https: // doi. org / 10.25223 / brad. n 41.2023. a 16" type="journal article" year="2023">
Sakhraoui
<emphasis id="0FFC5965455DFFDC987EFF6DFE620CE0" box="[350,401,194,220]" italics="true" pageId="6" pageNumber="41">et al</emphasis>
. 2023
</bibRefCitation>
). It is also naturalised in parts of
<taxonomicName id="FA88FEF4455DFFDC9A45FF6CFC3F0CE0" baseAuthorityName="Randall" baseAuthorityYear="2007" box="[869,972,194,220]" class="Liliopsida" family="Orchidaceae" genus="Australia" kingdom="Plantae" order="Asparagales" pageId="6" pageNumber="50" phylum="Tracheophyta" rank="genus">Australia</taxonomicName>
(
<bibRefCitation id="5919F886455DFFDC9AFCFF6CFB890CE0" author="Randall" box="[988,1146,194,220]" firstAuthor="Randall" pageId="6" pageNumber="41" refId="ref12359" refString="Randall, R. P. (2007 [September]) The introduced flora of Australia and its weed status. CRC for Australian Weed Management, Department of Agriculture and Food, Western Australia [Waite Campus, University of Adelaide, PMB 1, Glen Osmond, SA 5064 Australia.], 524 pp. Available from: https: // www. aabr. org. au / images / stories / resources / weeds / intro _ flora _ australia. pdf (accessed 1 December 2023)." type="book" year="2007">Randall 2007</bibRefCitation>
,
<bibRefCitation id="5919F886455DFFDC9DA7FF6CFA570CE0" author="Weeds of Australia" box="[1159,1444,194,220]" firstAuthor="Weeds of Australia" pageId="6" pageNumber="41" refId="ref14723" refString="Weeds of Australia (2023) Aloe arborescens Mill. Available from: https: // keyserver. lucidcentral. org / weeds / data / media / Html / aloe _ arborescens. htm (accessed 31 January 2023)." type="url" year="2023">
Weeds of
<taxonomicName id="FA88FEF4455DFFDC9DDAFF6CFA920CE0" baseAuthorityName="Randall" baseAuthorityYear="2007" box="[1274,1377,194,220]" class="Liliopsida" family="Orchidaceae" genus="Australia" kingdom="Plantae" order="Asparagales" pageId="6" pageNumber="50" phylum="Tracheophyta" rank="genus">Australia</taxonomicName>
2023
</bibRefCitation>
) and
<collectingCountry id="459FC5E7455DFFDC9998FF49FEBD0D3C" box="[184,334,230,257]" name="New Zealand" pageId="6" pageNumber="41">New Zealand</collectingCountry>
(
<bibRefCitation id="5919F886455DFFDC987DFF48FD9D0D3C" author="Schonberger" box="[349,622,230,257]" etAl="et al." firstAuthor="Schonberger" pageId="6" pageNumber="41" refId="ref13080" refString="Schonberger, I., Wilton, A. D., Boardman, K. F., Breitwieser, I., De Lange, P. J., De Pauw, B., Ford, K. A., Gibb, E. S., Glenny, D. S., Greer, P. A., Heenan, P. B., Korver, M. A., Maule, H. G., Novis, P. M., Prebble, J. M., Smissen, R. D. &amp; Tawiri, K. (2021) Checklist of the New Zealand Flora - Seed Plants. Manaaki Whenua-Landcare Research, Lincoln, 412 pp. https: // doi. org / 10.26065 / ax 7 t- 8 y 85" type="book" year="2021">
Schönberger
<emphasis id="0FFC5965455DFFDC98D3FF49FDD70D3C" box="[499,548,230,256]" italics="true" pageId="6" pageNumber="41">et al</emphasis>
. 2021
</bibRefCitation>
). In some areas of its secondary range,
<taxonomicName id="FA88FEF4455DFFDC9D09FF49FB430D3C" authorityName="Allioni" authorityYear="1773" box="[1065,1200,230,257]" class="Liliopsida" family="Asphodelaceae" genus="Aloe" kingdom="Plantae" order="Asparagales" pageId="6" pageNumber="41" phylum="Tracheophyta" rank="species" species="maculata">
<emphasis id="0FFC5965455DFFDC9D09FF49FB430D3C" box="[1065,1200,230,257]" italics="true" pageId="6" pageNumber="41">A. maculata</emphasis>
</taxonomicName>
is considered to be an environmental weed.
</paragraph>
<paragraph id="3D378577455DFFDC999DFE80FEEC0DAD" blockId="6.[136,1452,158,1301]" pageId="6" pageNumber="41">
<emphasis id="0FFC5965455DFFDC999DFE80FE600D74" bold="true" box="[189,403,302,328]" pageId="6" pageNumber="41">
Status in
<collectingCountry id="459FC5E7455DFFDC980BFE80FE600D74" box="[299,403,302,328]" name="Portugal" pageId="6" pageNumber="41">Portugal</collectingCountry>
</emphasis>
:—Naturalised. It was previously recorded in a list of “more or less naturalised” taxa (
<bibRefCitation id="5919F886455DFFDC9C69FE80FE990D50" author="Almeida &amp; Freitas" firstAuthor="Almeida" page="118" pageId="6" pageNumber="41" pagination="117 - 130" refId="ref8722" refString="Almeida, J. D. de &amp; Freitas, H. (2006) Exotic naturalized flora of continental Portugal. A reassessment. Botanica Complutensis 30: 117 - 130." type="journal article" year="2006">Almeida &amp; Freitas 2006: 118</bibRefCitation>
), and later recorded as casual (
<bibRefCitation id="5919F886455DFFDC9BE1FEFCFC6E0D50" author="Almeida" box="[705,925,338,364]" firstAuthor="Almeida" page="209" pageId="6" pageNumber="41" refId="ref8656" refString="Almeida, J. D. de. (2012) Flora exotica subespontanea de Portugal continental (plantas vasculares). 5. a edicao. M. Sc. thesis. University of Coimbra, Coimbra, 281 pp. Available from: https: // docplayer. com. br / 48643620 - Flora-exotica-subespontanea-de-portugalcontinental-plantas-vasculares. html (accessed 1 December 2023)." type="book" year="2012">Almeida 2012: 209</bibRefCitation>
) and as “Casual. Cultivated, escaped” (
<bibRefCitation id="5919F886455DFFDC9C75FEFCFEE00DAD" author="Silva" etAl="et al." firstAuthor="Silva" page="75" pageId="6" pageNumber="41" pagination="58 - 81" refId="ref13229" refString="Silva, V., Figueiredo, E. &amp; Smith, G. F. (2015) Alien succulents naturalised and cultivated on the central west coast of Portugal. Bradleya 33: 58 - 81. https: // doi. org / 10.25223 / brad. n 33.2015. a 10" type="journal article" year="2015">
Silva
<emphasis id="0FFC5965455DFFDC9CB7FEFDFF6D0DAC" italics="true" pageId="6" pageNumber="41">et al</emphasis>
. 2015: 75
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="7592D6FC455DFFDC999DFE34FBC80928" pageId="6" pageNumber="41" type="discussion">
<paragraph id="3D378577455DFFDC999DFE34FB120D89" blockId="6.[136,1452,158,1301]" box="[189,1249,410,437]" pageId="6" pageNumber="41">
<emphasis id="0FFC5965455DFFDC999DFE34FECC0D88" bold="true" box="[189,319,410,436]" pageId="6" pageNumber="41">References</emphasis>
:—
<bibRefCitation id="5919F886455DFFDC9840FE34FD7A0D88" author="Almeida &amp; Freitas" box="[352,649,410,436]" firstAuthor="Almeida" pageId="6" pageNumber="41" pagination="117 - 130" refId="ref8722" refString="Almeida, J. D. de &amp; Freitas, H. (2006) Exotic naturalized flora of continental Portugal. A reassessment. Botanica Complutensis 30: 117 - 130." type="journal article" year="2006">Almeida &amp; Freitas (2006)</bibRefCitation>
;
<bibRefCitation id="5919F886455DFFDC9BB3FE34FCBB0D88" author="Almeida" box="[659,840,410,436]" firstAuthor="Almeida" pageId="6" pageNumber="41" refId="ref8656" refString="Almeida, J. D. de. (2012) Flora exotica subespontanea de Portugal continental (plantas vasculares). 5. a edicao. M. Sc. thesis. University of Coimbra, Coimbra, 281 pp. Available from: https: // docplayer. com. br / 48643620 - Flora-exotica-subespontanea-de-portugalcontinental-plantas-vasculares. html (accessed 1 December 2023)." type="book" year="2012">Almeida (2012)</bibRefCitation>
;
<bibRefCitation id="5919F886455DFFDC9A73FE34FBF70D88" author="Guemes" box="[851,1028,410,436]" firstAuthor="Guemes" pageId="6" pageNumber="41" pagination="308 - 314" refId="ref10660" refString="Guemes, J. (2013) Aloe L. In: E. Rico, M. B. Crespo, A. Quintanar, A. Herrero &amp; C. Aedo (Eds.) Flora iberica 20. Real Jardin Botanico, CSIC, Madrid, pp. 308 - 314. Available from: http: // www. floraiberica. es / floraiberica / texto / pdfs / 20 _ 183 _ 32 _ Aloe. pdf (accessed 1 December 2023)." type="book chapter" year="2013">Güemes (2013)</bibRefCitation>
;
<bibRefCitation id="5919F886455DFFDC9D30FE34FB2E0D89" author="Silva" box="[1040,1245,410,437]" etAl="et al." firstAuthor="Silva" pageId="6" pageNumber="41" pagination="58 - 81" refId="ref13229" refString="Silva, V., Figueiredo, E. &amp; Smith, G. F. (2015) Alien succulents naturalised and cultivated on the central west coast of Portugal. Bradleya 33: 58 - 81. https: // doi. org / 10.25223 / brad. n 33.2015. a 10" type="journal article" year="2015">
Silva
<emphasis id="0FFC5965455DFFDC9D70FE35FB720D88" box="[1104,1153,410,436]" italics="true" pageId="6" pageNumber="41">et al</emphasis>
. (2015)
</bibRefCitation>
.
</paragraph>
<paragraph id="3D378577455DFFDC999DFE10FD810E78" blockId="6.[136,1452,158,1301]" pageId="6" pageNumber="41">
<emphasis id="0FFC5965455DFFDC999DFE10FEF30DE4" bold="true" box="[189,256,446,472]" pageId="6" pageNumber="41">Notes</emphasis>
:—
<taxonomicName id="FA88FEF4455DFFDC9804FE10FE360DE4" ID-CoL="C3TJ" authorityName="Allioni" authorityYear="1773" box="[292,453,446,472]" class="Liliopsida" family="Asphodelaceae" genus="Aloe" kingdom="Plantae" order="Asparagales" pageId="6" pageNumber="41" phylum="Tracheophyta" rank="species" species="maculata">
<emphasis id="0FFC5965455DFFDC9804FE10FE360DE4" box="[292,453,446,472]" italics="true" pageId="6" pageNumber="41">Aloe maculata</emphasis>
</taxonomicName>
has been recorded as naturalised in the Iberian Peninsula, in
<collectingCountry id="459FC5E7455DFFDC9D48FE10FB5B0DE4" box="[1128,1192,446,472]" name="Spain" pageId="6" pageNumber="41">Spain</collectingCountry>
and
<collectingCountry id="459FC5E7455DFFDC9DFDFE10FACF0DE4" box="[1245,1340,446,472]" name="Portugal" pageId="6" pageNumber="41">Portugal</collectingCountry>
(
<bibRefCitation id="5919F886455DFFDC9C69FE10FE950DC0" author="Almeida &amp; Freitas" firstAuthor="Almeida" page="119" pageId="6" pageNumber="41" pagination="117 - 130" refId="ref8722" refString="Almeida, J. D. de &amp; Freitas, H. (2006) Exotic naturalized flora of continental Portugal. A reassessment. Botanica Complutensis 30: 117 - 130." type="journal article" year="2006">Almeida &amp; Freitas 2006: 119</bibRefCitation>
,
<bibRefCitation id="5919F886455DFFDC9850FE4CFD3C0DC0" author="Guillot Ortiz" box="[368,719,482,509]" etAl="et al." firstAuthor="Guillot Ortiz" page="28 - 32" pageId="6" pageNumber="41" pagination="1 - 58" refId="ref10880" refString="Guillot Ortiz, D., Laguna Lumbreras, E. &amp; Rossello Picornell, J. A. (2008) La familia Aloaceae en la flora aloctona valenciana. Bouteloua 6: 1 - 58. Available from: http: // www. floramontiberica. org / Bouteloua / MonogBouteloua _ 06 _ Aloaceae. pdf (accessed 1 December 2023)." type="journal article" year="2008">
Guillot Ortiz
<emphasis id="0FFC5965455DFFDC9B25FE4DFDC60DC0" box="[517,565,482,508]" italics="true" pageId="6" pageNumber="41">et al</emphasis>
. 2008: 2832
</bibRefCitation>
,
<bibRefCitation id="5919F886455DFFDC9BF7FE4CFC430DC0" author="Almeida" box="[727,944,482,508]" firstAuthor="Almeida" page="209" pageId="6" pageNumber="41" refId="ref8656" refString="Almeida, J. D. de. (2012) Flora exotica subespontanea de Portugal continental (plantas vasculares). 5. a edicao. M. Sc. thesis. University of Coimbra, Coimbra, 281 pp. Available from: https: // docplayer. com. br / 48643620 - Flora-exotica-subespontanea-de-portugalcontinental-plantas-vasculares. html (accessed 1 December 2023)." type="book" year="2012">Almeida 2012: 209</bibRefCitation>
,
<bibRefCitation id="5919F886455DFFDC9A9AFE4CFB7C0DC0" author="Guemes" box="[954,1167,482,508]" firstAuthor="Guemes" page="312" pageId="6" pageNumber="41" pagination="308 - 314" refId="ref10660" refString="Guemes, J. (2013) Aloe L. In: E. Rico, M. B. Crespo, A. Quintanar, A. Herrero &amp; C. Aedo (Eds.) Flora iberica 20. Real Jardin Botanico, CSIC, Madrid, pp. 308 - 314. Available from: http: // www. floraiberica. es / floraiberica / texto / pdfs / 20 _ 183 _ 32 _ Aloe. pdf (accessed 1 December 2023)." type="book chapter" year="2013">Güemes 2013: 312</bibRefCitation>
). In places in
<collectingCountry id="459FC5E7455DFFDC9C05FE4CFA770DC0" box="[1317,1412,482,508]" name="Portugal" pageId="6" pageNumber="41">Portugal</collectingCountry>
the species has become naturalised through self-replacing populations and is now spreading without human aid. At some locations
<taxonomicName id="FA88FEF4455DFFDC99D4FD85FE880E78" ID-CoL="C3TJ" authorityName="Allioni" authorityYear="1773" box="[244,379,554,581]" class="Liliopsida" family="Asphodelaceae" genus="Aloe" kingdom="Plantae" order="Asparagales" pageId="6" pageNumber="41" phylum="Tracheophyta" rank="species" species="maculata">
<emphasis id="0FFC5965455DFFDC99D4FD85FE880E78" box="[244,379,554,581]" italics="true" pageId="6" pageNumber="41">A. maculata</emphasis>
</taxonomicName>
has become invasive.
</paragraph>
<paragraph id="3D378577455DFFDC999DFDE0FC900EE8" blockId="6.[136,1452,158,1301]" pageId="6" pageNumber="41">
<taxonomicName id="FA88FEF4455DFFDC999DFDE0FEAC0E54" ID-CoL="C3TJ" authorityName="Allioni" authorityYear="1773" box="[189,351,590,616]" class="Liliopsida" family="Asphodelaceae" genus="Aloe" kingdom="Plantae" order="Asparagales" pageId="6" pageNumber="41" phylum="Tracheophyta" rank="species" species="maculata">
<emphasis id="0FFC5965455DFFDC999DFDE0FEAC0E54" box="[189,351,590,616]" italics="true" pageId="6" pageNumber="41">Aloe maculata</emphasis>
</taxonomicName>
is a distinctive species that can be easily identified based on the adaxial leaf surface that is spotted, often densely so, with whitish maculations (
<figureCitation id="A5B399F2455DFFDC9B5FFDDCFD0A0EB0" box="[639,761,626,652]" captionStart="FIGURE 2" captionStartId="4.[136,229,1798,1820]" captionTargetBox="[171,1417,844,1755]" captionTargetId="figure-263@4.[151,1436,821,1773]" captionTargetPageId="4" captionText="FIGURE 2. Aloe maculata in Portugal. A. Naturalised at Almograve along the Baixo Alentejo coast. Photograph taken on 29 December 2017. B. Naturalised at Porto Covo along the Alentejo Litoral coast.Photograph taken on 30 December 2017. C. Naturalised at Malveira da Serra, Almoinhas Velhas, in Cascais, near metropolitan Lisbon. Photograph taken on 28 September 2014. D. Close-up of an inflorescence that carries salmon-orange flowers. The racemes are distinctly capitate and flowers have prominent basal swellings. All photographs by Gideon F. Smith." figureDoi="http://doi.org/10.5281/zenodo.10254759" httpUri="https://zenodo.org/record/10254759/files/figure.png" pageId="6" pageNumber="41">Fig. 2AC</figureCitation>
) and its capitate inflorescences (
<figureCitation id="A5B399F2455DFFDC9D4FFDDCFB3F0EB0" box="[1135,1228,626,652]" captionStart="FIGURE 2" captionStartId="4.[136,229,1798,1820]" captionTargetBox="[171,1417,844,1755]" captionTargetId="figure-263@4.[151,1436,821,1773]" captionTargetPageId="4" captionText="FIGURE 2. Aloe maculata in Portugal. A. Naturalised at Almograve along the Baixo Alentejo coast. Photograph taken on 29 December 2017. B. Naturalised at Porto Covo along the Alentejo Litoral coast.Photograph taken on 30 December 2017. C. Naturalised at Malveira da Serra, Almoinhas Velhas, in Cascais, near metropolitan Lisbon. Photograph taken on 28 September 2014. D. Close-up of an inflorescence that carries salmon-orange flowers. The racemes are distinctly capitate and flowers have prominent basal swellings. All photographs by Gideon F. Smith." figureDoi="http://doi.org/10.5281/zenodo.10254759" httpUri="https://zenodo.org/record/10254759/files/figure.png" pageId="6" pageNumber="41">Fig. 2D</figureCitation>
). Depending on the provenance of material of this species, flowering can take place at virtually any time of the year. Thus far only summer-flowering material of
<taxonomicName id="FA88FEF4455DFFDC985DFD15FDF70EE8" ID-CoL="C3TJ" authorityName="Allioni" authorityYear="1773" box="[381,516,698,725]" class="Liliopsida" family="Asphodelaceae" genus="Aloe" kingdom="Plantae" order="Asparagales" pageId="6" pageNumber="41" phylum="Tracheophyta" rank="species" species="maculata">
<emphasis id="0FFC5965455DFFDC985DFD15FDF70EE8" box="[381,516,698,725]" italics="true" pageId="6" pageNumber="41">A. maculata</emphasis>
</taxonomicName>
has been observed in
<collectingCountry id="459FC5E7455DFFDC9BDCFD14FCAC0EE8" box="[764,863,698,724]" name="Portugal" pageId="6" pageNumber="41">Portugal</collectingCountry>
.
</paragraph>
<paragraph id="3D378577455DFFDC999DFD70FBE70FB5" blockId="6.[136,1452,158,1301]" pageId="6" pageNumber="41">
The only alooid material cultivated in
<collectingCountry id="459FC5E7455DFFDC9B4BFD70FD390EC4" box="[619,714,734,760]" name="Portugal" pageId="6" pageNumber="41">Portugal</collectingCountry>
with which
<taxonomicName id="FA88FEF4455DFFDC9A75FD71FC2F0EC4" ID-CoL="C3TJ" authorityName="Allioni" authorityYear="1773" box="[853,988,734,761]" class="Liliopsida" family="Asphodelaceae" genus="Aloe" kingdom="Plantae" order="Asparagales" pageId="6" pageNumber="41" phylum="Tracheophyta" rank="species" species="maculata">
<emphasis id="0FFC5965455DFFDC9A75FD71FC2F0EC4" box="[853,988,734,761]" italics="true" pageId="6" pageNumber="41">A. maculata</emphasis>
</taxonomicName>
can be confused is
<taxonomicName id="FA88FEF4455DFFDC9D99FD71FF0B0F20" ID-CoL="668G6" authority="Todaro (1878: 70" authorityName="Todaro" authorityYear="1878" class="Liliopsida" family="Asphodelaceae" genus="Aloe" isHybrid="true" kingdom="Plantae" order="Asparagales" pageId="6" pageNumber="41" phylum="Tracheophyta" rank="species" species="schimperi">
<emphasis id="0FFC5965455DFFDC9D99FD71FB390EC5" box="[1209,1226,735,761]" italics="true" pageId="6" pageNumber="41">A</emphasis>
. ×
<emphasis id="0FFC5965455DFFDC9DC7FD70FAA60EC4" box="[1255,1365,734,760]" italics="true" pageId="6" pageNumber="41">schimperi</emphasis>
<bibRefCitation id="5919F886455DFFDC9C7CFD70FF0B0F20" author="Todaro" firstAuthor="Todaro" page="70" pageId="6" pageNumber="41" refId="ref14607" refString="Todaro, A. (1878 [1876 -]) Hortus Botanicus Panormitanus. Vol. 1. Cyri Visconti &amp; Francisci Lao, Palermo, 64 pp. + XL plates." type="book" year="1878">Todaro (1878: 70</bibRefCitation>
</taxonomicName>
, plate 16) [
<taxonomicName id="FA88FEF4455DFFDC9858FCADFDF30F20" ID-CoL="C3TJ" authorityName="Allioni" authorityYear="1773" box="[376,512,770,797]" class="Liliopsida" family="Asphodelaceae" genus="Aloe" kingdom="Plantae" order="Asparagales" pageId="6" pageNumber="41" phylum="Tracheophyta" rank="species" species="maculata">
<emphasis id="0FFC5965455DFFDC9858FCADFDF30F20" box="[376,512,770,797]" italics="true" pageId="6" pageNumber="41">A. maculata</emphasis>
</taxonomicName>
×
<taxonomicName id="FA88FEF4455DFFDC9B3FFCADFC810F20" ID-CoL="C44T" authority="Haworth (1804: 18)" authorityName="Haworth" authorityPageNumber="18" authorityYear="1804" box="[543,882,770,797]" class="Liliopsida" family="Asphodelaceae" genus="Aloe" kingdom="Plantae" order="Asparagales" pageId="6" pageNumber="41" phylum="Tracheophyta" rank="species" species="striata">
<emphasis id="0FFC5965455DFFDC9B3FFCADFD7B0F20" box="[543,648,771,797]" italics="true" pageId="6" pageNumber="41">A. striata</emphasis>
<bibRefCitation id="5919F886455DFFDC9BB0FCACFC810F20" author="Haworth" box="[656,882,770,796]" firstAuthor="Haworth" page="18" pageId="6" pageNumber="41" pagination="1 - 28" refId="ref10946" refString="Haworth, A. H. (1804) A new arrangement of the genus Aloe, with a chronological sketch of the progressive knowledge of that genus, and of other succulent genera. Transactions of the Linnean Society of London 7 (1): 1 - 28. https: // doi. org / 10.1111 / j. 1096 - 3642.1804. tb 00276. x" type="journal article" year="1804">Haworth (1804: 18)</bibRefCitation>
</taxonomicName>
] (
<figureCitation id="A5B399F2455DFFDC9AA9FCACFBF60F20" box="[905,1029,770,796]" captionStart="FIGURE 3" captionStartId="5.[136,229,1392,1414]" captionTargetBox="[178,1413,217,1332]" captionTargetId="figure-17@5.[151,1436,190,1368]" captionTargetPageId="5" captionText="FIGURE 3. Two nothospecies with which Aloe maculata can be confused. A. Aloe ×schimperi [A. maculata × A. striata] cultivated in the Botanical Garden of the University of Porto, Porto. Note the distinctly pinkish leaf margins, a character inherited from A. striata. B. The flowers of A. ×schimperi lack the prominent basal swellings present in A. maculata. Both photographs A and B were taken on 22 December 2017. C. The rosettes of A. ×commutata [A. grandidentata × A. maculata] resemble those of A. maculata, but, as shown in D, itsinflorescences are elongated (not capitate) and the flowers of this nothospecies, too, lack basal swellings,as found in A.grandidentata. Both photographs C and D were taken in the Huntington Botanical Gardens, San Marino, California, U.S.A., on 27 April 2011. All photographs by Gideon F. Smith." figureDoi="http://doi.org/10.5281/zenodo.10254761" httpUri="https://zenodo.org/record/10254761/files/figure.png" pageId="6" pageNumber="41">Fig. 3AB</figureCitation>
), of which the arguably better known name
<taxonomicName id="FA88FEF4455DFFDC99EEFC89FD9E0F7C" ID-CoL="C43G" authority="Baker (1902: 430)" authorityName="Baker" authorityPageNumber="430" authorityYear="1902" box="[206,621,806,833]" class="Liliopsida" family="Asphodelaceae" genus="Aloe" isHybrid="true" kingdom="Plantae" order="Asparagales" pageId="6" pageNumber="41" phylum="Tracheophyta" rank="species" species="schoenlandii">
<emphasis id="0FFC5965455DFFDC99EEFC89FF2C0F7D" box="[206,223,807,833]" italics="true" pageId="6" pageNumber="41">A</emphasis>
. ×
<emphasis id="0FFC5965455DFFDC99DFFC88FE7D0F7C" box="[255,398,806,832]" italics="true" pageId="6" pageNumber="41">schoenlandii</emphasis>
<bibRefCitation id="5919F886455DFFDC98B8FC88FD9E0F7C" author="Baker" box="[408,621,806,832]" firstAuthor="Baker" page="430" pageId="6" pageNumber="41" pagination="429 - 430" refId="ref8867" refString="Baker, J. G. (1902) New or noteworthy plants. The Gardeners' Chronicle ser. 3, 32: 429 - 430. Available from: https: // www. biodiversitylibrary. org / item / 83820 # page / 494 / mode / 1 up (accessed 1 December 2023)." type="journal article" year="1902">Baker (1902: 430)</bibRefCitation>
</taxonomicName>
, as
<emphasis id="0FFC5965455DFFDC9B85FC88FCD00F7C" box="[677,803,806,832]" italics="true" pageId="6" pageNumber="41">Schönlandi</emphasis>
,
<emphasis id="0FFC5965455DFFDC9A1CFC89FC760F7C" box="[828,901,807,832]" italics="true" pageId="6" pageNumber="41">pro sp</emphasis>
., is a synonym (
<bibRefCitation id="5919F886455DFFDC9D66FC88FA6D0F7C" author="Figueiredo &amp; Smith" box="[1094,1438,806,832]" firstAuthor="Figueiredo" page="14" pageId="6" pageNumber="41" pagination="9 - 17" refId="ref9680" refString="Figueiredo, E. &amp; Smith, G. F. (2016) Aloe × schimperi Tod. (Aloe maculata All. × Aloe striata Haw.), the earliest name applicable to the common and invasive nothospecies known as Aloe × schoenlandii Baker (Asphodelaceae: Alooideae). Haseltonia 22: 9 - 17. https: // doi. org / 10.2985 / 026.022.0103" type="journal article" year="2016">Figueiredo &amp; Smith 2016: 14</bibRefCitation>
). However, the leaf margins of
<taxonomicName id="FA88FEF4455DFFDC98C4FCE5FD700F58" ID-CoL="668G6" authorityName="Todaro" authorityYear="1878" box="[484,643,842,869]" class="Liliopsida" family="Asphodelaceae" genus="Aloe" isHybrid="true" kingdom="Plantae" order="Asparagales" pageId="6" pageNumber="41" phylum="Tracheophyta" rank="species" species="schimperi">
<emphasis id="0FFC5965455DFFDC98C4FCE5FE060F59" box="[484,501,843,869]" italics="true" pageId="6" pageNumber="41">A</emphasis>
. ×
<emphasis id="0FFC5965455DFFDC9B35FCE4FD700F58" box="[533,643,842,868]" italics="true" pageId="6" pageNumber="41">schimperi</emphasis>
</taxonomicName>
are distinctly pinkish—a character inherited from
<taxonomicName id="FA88FEF4455DFFDC9DF0FCE5FAC80F58" ID-CoL="C44T" authorityName="Haworth" authorityYear="1804" box="[1232,1339,843,869]" class="Liliopsida" family="Asphodelaceae" genus="Aloe" kingdom="Plantae" order="Asparagales" pageId="6" pageNumber="41" phylum="Tracheophyta" rank="species" species="striata">
<emphasis id="0FFC5965455DFFDC9DF0FCE5FAC80F58" box="[1232,1339,843,869]" italics="true" pageId="6" pageNumber="41">A. striata</emphasis>
</taxonomicName>
—and the flowers lack the prominent basal swellings evident in the flowers of
<taxonomicName id="FA88FEF4455DFFDC9AA6FCC1FBFE0FB4" ID-CoL="C3TJ" authorityName="Allioni" authorityYear="1773" box="[902,1037,878,905]" class="Liliopsida" family="Asphodelaceae" genus="Aloe" kingdom="Plantae" order="Asparagales" pageId="6" pageNumber="41" phylum="Tracheophyta" rank="species" species="maculata">
<emphasis id="0FFC5965455DFFDC9AA6FCC1FBFE0FB4" box="[902,1037,878,905]" italics="true" pageId="6" pageNumber="41">A. maculata</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="3D378577455DFFDC999DFC3CFB360894" blockId="6.[136,1452,158,1301]" pageId="6" pageNumber="41">
<taxonomicName id="FA88FEF4455DFFDC999DFC3CFE290F90" ID-CoL="C3HX" authority="Todaro" authorityName="Todaro" box="[189,474,914,941]" class="Liliopsida" family="Asphodelaceae" genus="Aloe" isHybrid="true" kingdom="Plantae" order="Asparagales" pageId="6" pageNumber="41" phylum="Tracheophyta" rank="species" species="commutata">
<emphasis id="0FFC5965455DFFDC999DFC3CFF030F90" box="[189,240,914,940]" italics="true" pageId="6" pageNumber="41">Aloe</emphasis>
×
<emphasis id="0FFC5965455DFFDC9828FC3DFE770F90" box="[264,388,915,940]" italics="true" pageId="6" pageNumber="41">commutata</emphasis>
Todaro
</taxonomicName>
(1876: 75, t. 18) (
<figureCitation id="A5B399F2455DFFDC9B88FC3CFCD20F90" box="[680,801,914,940]" captionStart="FIGURE 3" captionStartId="5.[136,229,1392,1414]" captionTargetBox="[178,1413,217,1332]" captionTargetId="figure-17@5.[151,1436,190,1368]" captionTargetPageId="5" captionText="FIGURE 3. Two nothospecies with which Aloe maculata can be confused. A. Aloe ×schimperi [A. maculata × A. striata] cultivated in the Botanical Garden of the University of Porto, Porto. Note the distinctly pinkish leaf margins, a character inherited from A. striata. B. The flowers of A. ×schimperi lack the prominent basal swellings present in A. maculata. Both photographs A and B were taken on 22 December 2017. C. The rosettes of A. ×commutata [A. grandidentata × A. maculata] resemble those of A. maculata, but, as shown in D, itsinflorescences are elongated (not capitate) and the flowers of this nothospecies, too, lack basal swellings,as found in A.grandidentata. Both photographs C and D were taken in the Huntington Botanical Gardens, San Marino, California, U.S.A., on 27 April 2011. All photographs by Gideon F. Smith." figureDoi="http://doi.org/10.5281/zenodo.10254761" httpUri="https://zenodo.org/record/10254761/files/figure.png" pageId="6" pageNumber="41">Fig. 3CD</figureCitation>
) (see
<bibRefCitation id="5919F886455DFFDC9A45FC3CFB7A0F90" author="Figueiredo &amp; Smith" box="[869,1161,914,940]" firstAuthor="Figueiredo" pageId="6" pageNumber="41" pagination="25 - 32" refId="ref9620" refString="Figueiredo, E. &amp; Smith, G. F. (2012) Clarifying the application of the long-confused name Aloe commutata, and the establishment of Aloe × commutata Tod. (Asphodelaceae). Bradleya 30: 25 - 32. https: // doi. org / 10.25223 / brad. n 30.2012. a 7" type="journal article" year="2012">Figueiredo &amp; Smith 2012</bibRefCitation>
), of which the parents are
<taxonomicName id="FA88FEF4455DFFDC99A8FC19FE230FEC" ID-CoL="C3NQ" authority="Salm-Dyck" authorityName="Salm-Dyck" box="[136,464,950,977]" class="Liliopsida" family="Asphodelaceae" genus="Aloe" kingdom="Plantae" order="Asparagales" pageId="6" pageNumber="41" phylum="Tracheophyta" rank="species" species="grandidentata">
<emphasis id="0FFC5965455DFFDC99A8FC19FEBB0FEC" box="[136,328,950,977]" italics="true" pageId="6" pageNumber="41">A. grandidentata</emphasis>
Salm-Dyck
</taxonomicName>
(1822: 3 [species no. 2]) and
<taxonomicName id="FA88FEF4455DFFDC9A08FC19FC430FEC" ID-CoL="C3TJ" authorityName="Allioni" authorityYear="1773" box="[808,944,950,977]" class="Liliopsida" family="Asphodelaceae" genus="Aloe" kingdom="Plantae" order="Asparagales" pageId="6" pageNumber="41" phylum="Tracheophyta" rank="species" species="maculata">
<emphasis id="0FFC5965455DFFDC9A08FC19FC430FEC" box="[808,944,950,977]" italics="true" pageId="6" pageNumber="41">A. maculata</emphasis>
</taxonomicName>
, has a similar, typically maculate aloe-like vegetative morphological appearance as
<taxonomicName id="FA88FEF4455DFFDC9B75FC75FD2D0FC8" ID-CoL="C3TJ" authorityName="Allioni" authorityYear="1773" box="[597,734,986,1013]" class="Liliopsida" family="Asphodelaceae" genus="Aloe" kingdom="Plantae" order="Asparagales" pageId="6" pageNumber="41" phylum="Tracheophyta" rank="species" species="maculata">
<emphasis id="0FFC5965455DFFDC9B75FC75FD2D0FC8" box="[597,734,986,1013]" italics="true" pageId="6" pageNumber="41">A. maculata</emphasis>
</taxonomicName>
. However,
<taxonomicName id="FA88FEF4455DFFDC9A42FC75FBFC0FC8" ID-CoL="C3HX" box="[866,1039,987,1013]" class="Liliopsida" family="Asphodelaceae" genus="Aloe" isHybrid="true" kingdom="Plantae" order="Asparagales" pageId="6" pageNumber="41" phylum="Tracheophyta" rank="species" species="commutata">
<emphasis id="0FFC5965455DFFDC9A42FC75FC800FC9" box="[866,883,987,1013]" italics="true" pageId="6" pageNumber="41">A</emphasis>
. ×
<emphasis id="0FFC5965455DFFDC9AB3FC75FBFC0FC8" box="[915,1039,987,1012]" italics="true" pageId="6" pageNumber="41">commutata</emphasis>
</taxonomicName>
has elongated racemes (not capitate ones) and clavate (not basally inflated) flowers, and the leaf margins are concolorous, unlike those of
<taxonomicName id="FA88FEF4455DFFDC9C2DFC51FA580824" ID-CoL="668G6" authorityName="Todaro" authorityYear="1878" box="[1293,1451,1022,1049]" class="Liliopsida" family="Asphodelaceae" genus="Aloe" isHybrid="true" kingdom="Plantae" order="Asparagales" pageId="6" pageNumber="41" phylum="Tracheophyta" rank="species" species="schimperi">
<emphasis id="0FFC5965455DFFDC9C2DFC51FAED0825" box="[1293,1310,1023,1049]" italics="true" pageId="6" pageNumber="41">A</emphasis>
. ×
<emphasis id="0FFC5965455DFFDC9C1DFC50FA580824" box="[1341,1451,1022,1048]" italics="true" pageId="6" pageNumber="41">schimperi</emphasis>
</taxonomicName>
that are pinkish. Thus far we have not observed
<taxonomicName id="FA88FEF4455DFFDC9BE7FB8DFC850800" ID-CoL="C3HX" box="[711,886,1059,1085]" class="Liliopsida" family="Asphodelaceae" genus="Aloe" isHybrid="true" kingdom="Plantae" order="Asparagales" pageId="6" pageNumber="41" phylum="Tracheophyta" rank="species" species="commutata">
<emphasis id="0FFC5965455DFFDC9BE7FB8DFD2B0801" box="[711,728,1059,1085]" italics="true" pageId="6" pageNumber="41">A</emphasis>
. ×
<emphasis id="0FFC5965455DFFDC9BDAFB8DFC850800" box="[762,886,1059,1084]" italics="true" pageId="6" pageNumber="41">commutata</emphasis>
</taxonomicName>
in cultivation anywhere in continental
<collectingCountry id="459FC5E7455DFFDC9C65FB8CFA540800" box="[1349,1447,1058,1084]" name="Portugal" pageId="6" pageNumber="41">Portugal</collectingCountry>
, although it is popular as a garden subject in some parts of the world, for example in the Boyce Thompson Arboretum, in Superior,
<collectingRegion id="FF4C4B95455DFFDC9834FBC4FE8208B8" box="[276,369,1130,1156]" country="United States of America" name="Arizona" pageId="6" pageNumber="41">Arizona</collectingRegion>
,
<collectingCountry id="459FC5E7455DFFDC985DFBC4FE3908B8" box="[381,458,1130,1156]" name="United States of America" pageId="6" pageNumber="41">U.S.A.</collectingCountry>
(see
<bibRefCitation id="5919F886455DFFDC9B28FBC4FD7D08B9" author="Smith" box="[520,654,1130,1157]" firstAuthor="Smith" pageId="6" pageNumber="41" pagination="28 - 34" refId="ref13346" refString="Smith, G. F. (2005) The Boyce Thompson Arboretum near Superior, Arizona: a stunning botanical and succulent garden in cowboy country. Aloe 42 (1 &amp; 2): 28 - 34." type="journal article" year="2005">Smith 2005</bibRefCitation>
) and in the Huntington Botanical Gardens, in
<collectingCountry id="459FC5E7455DFFDC9DBEFBC4FAD508B8" box="[1182,1318,1130,1156]" name="San Marino" pageId="6" pageNumber="41">San Marino</collectingCountry>
,
<collectingRegion id="FF4C4B95455DFFDC9C13FBC4FA5408B8" box="[1331,1447,1130,1156]" country="United States of America" name="California" pageId="6" pageNumber="41">California</collectingRegion>
,
<collectingCountry id="459FC5E7455DFFDC99A8FB20FF260894" box="[136,213,1166,1192]" name="United States of America" pageId="6" pageNumber="41">U.S.A.</collectingCountry>
(see
<bibRefCitation id="5919F886455DFFDC9830FB20FE650894" author="Smith" box="[272,406,1166,1192]" firstAuthor="Smith" pageId="6" pageNumber="41" pagination="90 - 94" refId="ref13307" refString="Smith, G. F. (1997) The Desert Garden of the Huntington Botanical Gardens: one of the great gardens of the world. Aloe 34 (3 &amp; 4): 90 - 94." type="journal article" year="1997">Smith 1997</bibRefCitation>
), but, interestingly, not in
<collectingCountry id="459FC5E7455DFFDC9B9BFB20FCBF0894" box="[699,844,1166,1192]" name="South Africa" pageId="6" pageNumber="41">South Africa</collectingCountry>
, from where its parents originate.
</paragraph>
<paragraph id="3D378577455DFFDC999DFB1DFBC80928" blockId="6.[136,1452,158,1301]" pageId="6" pageNumber="41">
In
<taxonomicName id="FA88FEF4455DFFDC99FDFB1DFE9608F0" ID-CoL="C3TJ" authorityName="Allioni" authorityYear="1773" box="[221,357,1202,1229]" class="Liliopsida" family="Asphodelaceae" genus="Aloe" kingdom="Plantae" order="Asparagales" pageId="6" pageNumber="41" phylum="Tracheophyta" rank="species" species="maculata">
<emphasis id="0FFC5965455DFFDC99FDFB1DFE9608F0" box="[221,357,1202,1229]" italics="true" pageId="6" pageNumber="41">A. maculata</emphasis>
</taxonomicName>
, one other subspecies,
<taxonomicName id="FA88FEF4455DFFDC9B4DFB1DFEF208CC" ID-CoL="5FHYN" authority="(Reynolds 1937: 148) Gideon F. Sm. &amp; Figueiredo" authorityName="Gideon F. Sm. &amp; Figueiredo" baseAuthorityName="Reynolds" baseAuthorityPageNumber="148" baseAuthorityYear="1937" class="Liliopsida" family="Asphodelaceae" genus="Aloe" kingdom="Plantae" order="Asparagales" pageId="6" pageNumber="41" phylum="Tracheophyta" rank="subSpecies" species="maculata" subSpecies="ficksburgensis">
<emphasis id="0FFC5965455DFFDC9B4DFB1DFD0608F0" box="[621,757,1202,1229]" italics="true" pageId="6" pageNumber="41">A. maculata</emphasis>
subsp.
<emphasis id="0FFC5965455DFFDC9A6DFB1CFC1F08F0" box="[845,1004,1202,1228]" italics="true" pageId="6" pageNumber="41">ficksburgensis</emphasis>
(
<bibRefCitation id="5919F886455DFFDC9ADBFB1CFB1108F0" author="Reynolds" box="[1019,1250,1202,1228]" firstAuthor="Reynolds" page="148" pageId="6" pageNumber="41" pagination="143 - 150" refId="ref12576" refString="Reynolds, G. W. (1937) A new Aloe from South-West Africa, together with new varieties from the Transvaal and Orange Free State. Journal of South African Botany 3: 143 - 150." type="journal article" year="1937">Reynolds 1937: 148</bibRefCitation>
) Gideon F.Sm. &amp; Figueiredo
</taxonomicName>
in
<bibRefCitation id="5919F886455DFFDC9806FB78FDC508CD" author="Smith" box="[294,566,1238,1265]" etAl="et al." firstAuthor="Smith" page="15" pageId="6" pageNumber="41" pagination="13 - 18" refId="ref13771" refString="Smith, G. F., Figueiredo, E., Klopper, R. R. &amp; Crouch, N. R. (2012 b) Aloe maculata All. (Asphodelaceae) in the Free State province, South Africa, and resurrection of ' var. ficksburgensis'. Bradleya 30: 13 - 18. https: // doi. org / 10.25223 / brad. n 30.2012. a 5" type="journal article" year="2012" yearSuffix="b">
Smith
<emphasis id="0FFC5965455DFFDC9850FB79FE5A08CC" box="[368,425,1238,1264]" italics="true" pageId="6" pageNumber="41">et al.</emphasis>
(2012b: 15)
</bibRefCitation>
, is recognised, which is only known from the eastern
<collectingRegion id="FF4C4B95455DFFDC9DB6FB79FAF908CC" box="[1174,1290,1238,1265]" country="South Africa" name="Free State" pageId="6" pageNumber="41">Free State</collectingRegion>
,
<collectingCountry id="459FC5E7455DFFDC9C36FB78FA5408CC" box="[1302,1447,1238,1264]" name="South Africa" pageId="6" pageNumber="41">South Africa</collectingCountry>
, and western
<collectingCountry id="459FC5E7455DFFDC9836FB54FE870928" box="[278,372,1274,1300]" name="Lesotho" pageId="6" pageNumber="41">Lesotho</collectingCountry>
. This subspecies has not been observed in continental
<collectingCountry id="459FC5E7455DFFDC9AF4FB54FBC40928" box="[980,1079,1274,1300]" name="Portugal" pageId="6" pageNumber="41">Portugal</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>