197 lines
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197 lines
17 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.403.7145" ID-GBIF-Dataset="a6fda06f-575a-4ae4-8d40-ae927a01f959" ID-PMC="PMC4023238" ID-Pensoft-Pub="1313-2970-403-15" ID-PubMed="24843265" ID-ZBK="328E01EFBF324352AD7DBE989A3D716B" ModsDocAuthor="" ModsDocDate="2014" ModsDocID="1313-2970-403-15" ModsDocOrigin="ZooKeys 403" ModsDocTitle="A revision of Japanese species of the genus Psammoecus Latreille (Coleoptera, Silvanidae)" checkinTime="1451245966222" checkinUser="pensoft" docAuthor="Yoshida, Takahiro & Hirowatari, Toshiya" docDate="2014" docId="A5E95FDD6379BC13F049A90B86B4B3EF" docLanguage="en" docName="ZooKeys 403: 15-45" docOrigin="ZooKeys 403" docSource="http://dx.doi.org/10.3897/zookeys.403.7145" docTitle="Psammoecus fasciatus Reitter 1874" docType="treatment" docVersion="3" lastPageNumber="36" masterDocId="F8057B36FF80C436875CFFBFFFE6FF8F" masterDocTitle="A revision of Japanese species of the genus Psammoecus Latreille (Coleoptera, Silvanidae)" masterLastPageNumber="45" masterPageNumber="15" pageNumber="34" updateTime="1668158244412" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A revision of Japanese species of the genus Psammoecus Latreille (Coleoptera, Silvanidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Yoshida, Takahiro</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Hirowatari, Toshiya</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2014</mods:date>
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<mods:detail type="volume">
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<mods:number>403</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>15</mods:start>
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<mods:end>45</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.403.7145</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.403.7145</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-403-15</mods:identifier>
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<mods:identifier type="ZBK">328E01EFBF324352AD7DBE989A3D716B</mods:identifier>
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<mods:identifier type="ZooBank">328E01EFBF324352AD7DBE989A3D716B</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152052615" LSID="urn:lsid:plazi:treatment:A5E95FDD6379BC13F049A90B86B4B3EF" httpUri="http://treatment.plazi.org/id/A5E95FDD6379BC13F049A90B86B4B3EF" lastPageId="21" lastPageNumber="36" pageId="19" pageNumber="34">
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<subSubSection pageId="19" pageNumber="34" type="nomenclature">
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<paragraph pageId="19" pageNumber="34">
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<taxonomicName LSID="http://species-id.net/wiki/Psammoecus_fasciatus" authority="Reitter, 1874" authorityName="Reitter" authorityYear="1874" class="Insecta" family="Silvanidae" genus="Psammoecus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Psammoecus fasciatus" order="Coleoptera" pageId="19" pageNumber="34" phylum="Arthropoda" rank="species" species="fasciatus">Psammoecus fasciatus Reitter, 1874</taxonomicName>
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Figs 2D, 11and 14
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<normalizedToken originalValue="M–O">M-O</normalizedToken>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="19" pageNumber="34" type="reference_group">
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<paragraph pageId="19" pageNumber="34">
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<taxonomicName genus="Psamoecus" lsidName="Psamoecus" pageId="19" pageNumber="34" rank="genus">Psamoecus</taxonomicName>
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[sic.] fasciatus Reitter, 1874: 525. Type locality: Japan; Type deposition: the Natural History Museum, London; Type examined. (misspelling)
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</paragraph>
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<paragraph pageId="19" pageNumber="34">
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<taxonomicName class="Insecta" family="Silvanidae" genus="Psammoecus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Psammoecus fasciatus" order="Coleoptera" pageId="19" pageNumber="34" phylum="Arthropoda" rank="species" species="fasciatus">Psammoecus fasciatus</taxonomicName>
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:
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<bibRefCitation author="Hetschko, A" journalOrPublisher="W. Junk, Berlin" pageId="27" pageNumber="42" pagination="1 - 90" title="Coleopterorum Catalogus Pars 109" year="1930">Hetschko 1930</bibRefCitation>
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: 82. (catalogue) -
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<bibRefCitation author="Nakane, T" editor="Nakane, T" journalOrPublisher="Hokuryukan, Tokyo" pageId="27" pageNumber="42" pagination="195 - 196" title="Iconographia Insectorum Japonicorum Colore naturali edita" year="1963">Nakane 1963</bibRefCitation>
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: 196, fig. 16 in pl. 98. -
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<bibRefCitation author="Hisamatsu, S" journalOrPublisher="Hiroshima Mushi-no-kai Kaiho" pageId="27" pageNumber="42" pagination="207 - 209" title="Some notes of clavicorn beetles from Hiroshima Prefecture (Hiroshima-ken san kyuukakugun kochu note)." volume="16" year="1977">Hisamatsu 1977</bibRefCitation>
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: 21. -
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<bibRefCitation author="Sasaji, H" editor="Kurosawa, Y" journalOrPublisher="Hoikusha Publishing, Osaka" pageId="28" pageNumber="43" pagination="202 - 205" title="The Coleoptera of Japan in Color" year="1985">Sasaji 1985</bibRefCitation>
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: 204, fig. 34 in pl. 32. -
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<bibRefCitation author="Sato, M" editor="Hirashima, Y" journalOrPublisher="Entomological Laboratory, Faculty of Agriculture, Kyushu University, Fukuoka" pageId="28" pageNumber="43" pagination="197 - 538" title="Coleoptera." volumeTitle="A Check List of Japanese Insects" year="1989">Sato 1989</bibRefCitation>
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: 377. -
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<bibRefCitation author="Halstead, DGH" editor="Loebl, I" journalOrPublisher="Apollo Books, Stenstrup" pageId="27" pageNumber="42" pagination="496 - 501" title="Silvanidae." volumeTitle="Catalogue of Palaearctic Coleoptera" year="2007">Halstead et al. 2007</bibRefCitation>
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-
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<bibRefCitation author="Hirano, Y" journalOrPublisher="Kanagawa-chuho" pageId="27" pageNumber="42" pagination="57 - 83" title="Notes on Japanese Silvanidae (Nihonsan hosohiratamushi-ka ni tsuite)." volume="168" year="2009">Hirano 2009</bibRefCitation>
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: 64. -
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<bibRefCitation author="Hirano, Y" journalOrPublisher="Roppon-Ashi Entomological Books, Tokyo" pageId="27" pageNumber="42" title="Cucujoidea of Japan Vol. 2 Silvanidae, Byturidae, Biphyllidae." year="2010">Hirano 2010</bibRefCitation>
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: 12, 13. -
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<bibRefCitation author="Yoshida, T" journalOrPublisher="Japanese Journal of Systematic Entomology" pageId="28" pageNumber="43" pagination="85 - 90" title="A New Species of the Genus Psammoecus (Coleoptera, Silvanidae) from the Nansei Islands, Japan." volume="19" year="2013">Yoshida and Hirowatari 2013</bibRefCitation>
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: 5, 11, 16, 17.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="19" pageNumber="34" type="diagnosis">
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<paragraph pageId="19" pageNumber="34">Diagnosis.</paragraph>
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<paragraph pageId="19" pageNumber="34">
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This species is closely similar to
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<taxonomicName class="Insecta" family="Silvanidae" genus="Psammoecus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Psammoecus hiranoi" order="Coleoptera" pageId="19" pageNumber="34" phylum="Arthropoda" rank="species" species="hiranoi">Psammoecus hiranoi</taxonomicName>
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. It can be distinguished by the longer parameres, the shallower incision of the phallobase on the anterior 1/4 and the shorter teeth on the lateral margins of the pronotum.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="20" lastPageNumber="35" pageId="19" pageNumber="34" type="description">
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<paragraph pageId="19" pageNumber="34">Description.</paragraph>
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<paragraph pageId="19" pageNumber="34">Body length. 2.75-3.50 mm (n=49).</paragraph>
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<paragraph pageId="19" pageNumber="34">Coloration (Fig. 2D). Head and pronotum yellowish-brown to reddish-brown. Elytra with a variable horizontal dark band at middle, this band in lighter color specimens thinner towards elytral suture, lateral margins, apical portion and posterior elytral suture dark. Antennae brown or reddish-brown basally, 7th and 8th antennomeres darker, 9th and 10th almost black, 11th (apex) lighter than basal ones. Antennae of some specimens almost unicolorous.</paragraph>
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<paragraph pageId="19" pageNumber="34">Head (Fig. 11A, B, C). Broad, HW/HL 1.40-2.08; IE/HL 0.94-1.38. Temples enlarged behind eyes, narrowed gradually toward posterior margin. Eyes large, slightly prominent. Ventral surface with rough irregular punctuation, sparser punctuation on dorsal surface. Antennae 1.38-1.57 mm, 2nd antennomere short; covered with thick medium length pubescence and some long erect pubescence on each antennomere; approximate ratio of lectotype as follows: 2.5: 1.0: 1.3: 1.3: 1.3: 1.3: 1.3: 1.3: 1.3: 1.2: 1.8 (Fig. 11A).</paragraph>
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<paragraph pageId="19" pageNumber="34">Pronotum (Fig. 11B, C). Broad, PW/PL 0.69-1.52. Punctuation on dorsal surface very dense, but no punctures around posterior margin, punctuation on ventral surface sparser than dorsal surface, some of them in rows especially on posterior half. Pubescence composed of frequent short setae on dorsal surface and a long seta on each tooth on lateral margins and at anterior and posterior angles. Each anterior angle with a distinct group of a few very small teeth, each lateral margin with four teeth; tooth I comparatively small, teeth II, III and IV almost the same moderate size, each posterior angle with a comparatively small tooth.</paragraph>
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<paragraph pageId="20" pageNumber="35">
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<pageBreakToken pageId="20" pageNumber="35" start="start">Elytra</pageBreakToken>
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(Fig. 11E). Elongate-oval, EW/BL 0.41-0.47. Punctures wider than interstices. Pubescence composed of semi-erect medium length setae and some long erect setae in a row around lateral margins, longer toward anterior portion.
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</paragraph>
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<paragraph pageId="20" pageNumber="35">9th abdominal sternite (Fig. 14M). Strut cut at around half, diverging widely around the posterior base, ends of each branch curved inwards. Lateral sclerites triangular, flat.</paragraph>
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<paragraph pageId="20" pageNumber="35">Aedeagus (Figs 14N and O). Parameres simple stick-shaped, elongate, slightly curved inwards, punctuated sparsely except base, a few semi-long setae around apical portion. Phallobase diverging gradually around anterior 1/4, a little bulging along inner margins of each branch, lateral margins broadening. Penis also elongate, punctuated on posterior 1/6, more densely on ventral portion, especially densely on apical portion.</paragraph>
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</subSubSection>
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<subSubSection pageId="20" pageNumber="35" type="type series">
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<paragraph pageId="20" pageNumber="35">Type series.</paragraph>
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<paragraph pageId="20" pageNumber="35">Lectotype: male, Mt. Maya-san, Kobe City, Hyogo Prefecture, Japan, 1871, G. Lewis leg. (BMNH). Paralectotype: 1 female, same locality, 1871, G. Lewis leg. (BMNH). (here designated)</paragraph>
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</subSubSection>
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<subSubSection lastPageId="21" lastPageNumber="36" pageId="20" pageNumber="35" type="specimens examined">
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<paragraph pageId="20" pageNumber="35">Specimens examined.</paragraph>
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<paragraph lastPageId="21" lastPageNumber="36" pageId="20" pageNumber="35">
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Russia: [Primorsky Krai] 1 ex., Chuguevka Village, 27
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<normalizedToken originalValue="–VI–">-VI-</normalizedToken>
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1999, Y. Notsu leg. (EUMJ). JAPAN: [Niigata Pref.] 1 male & 1 ex., Renge-Onsen, Itoigawa City, 14
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<normalizedToken originalValue="–VIII–">-VIII-</normalizedToken>
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2003, I. Tanaka leg. (ITC). [Tokyo Pref.] 7 exs.,
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<pageBreakToken pageId="21" pageNumber="36" start="start">Mikura</pageBreakToken>
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Island, Mikurajima Village, 11
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<normalizedToken originalValue="–VI–">-VI-</normalizedToken>
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1977, T. Nakane leg. (HUSE). [Nagano Pref.] 1 ex.,
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<normalizedToken originalValue="Togakushi-Kôgen">Togakushi-Kogen</normalizedToken>
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, Nagano City, 5
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<normalizedToken originalValue="–6–VII–">-6-VII-</normalizedToken>
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2008, K. Mizuno leg. (KMC); 2 ex.,
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<normalizedToken originalValue="Ôbora">Obora</normalizedToken>
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, Ueda City, 24
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<normalizedToken originalValue="–VII–">-VII-</normalizedToken>
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2013, T. Yoshida leg. (ELKU). [Kyoto Pref.] 1 ex., Kifune, Kyoto City, 6
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<normalizedToken originalValue="–IV–">-IV-</normalizedToken>
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1988, I. Tanaka leg. (ITC). [Hyogo Pref.] 1 ex., Itoi, Asago City, 26
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<normalizedToken originalValue="–IX–">-IX-</normalizedToken>
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2004, I. Tanaka leg. (ITC); 5 exs., Sakanotani,
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<normalizedToken originalValue="Shisô">Shiso</normalizedToken>
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City, 19
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<normalizedToken originalValue="–VII–">-VII-</normalizedToken>
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2009, K.
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<normalizedToken originalValue="Itô">Ito</normalizedToken>
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leg. (KMC); 1 ex., Onzui,
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<normalizedToken originalValue="Shisô">Shiso</normalizedToken>
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City, 19
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<normalizedToken originalValue="–VII–">-VII-</normalizedToken>
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2009, K.
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<normalizedToken originalValue="Utô">Uto</normalizedToken>
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leg. (KMC); 1 ex., Mt. Yuzuruha-san, Nantan Town, Awaji Island, 18
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<normalizedToken originalValue="–X–">-X-</normalizedToken>
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2003, I. Tanaka leg. (ITC). [Nara Pref.] 1 ex., Mt. Kasuga-yama, Nara City, 5
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<normalizedToken originalValue="–V–">-V-</normalizedToken>
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1985, I. Tanaka leg. (ITC); 2 exs., same locality, 5
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<normalizedToken originalValue="–V–">-V-</normalizedToken>
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2012, K. Ikeuchi leg. (ELKU); 1 ex., same locality, 31
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<normalizedToken originalValue="–VII–">-VII-</normalizedToken>
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2012, K. Matsuda leg. (ELKU). [Tokushima Pref.] 1 ex., Mt. Shibakoya-yama, Kamiyama Town, 31
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<normalizedToken originalValue="–VII–">-VII-</normalizedToken>
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1975, M. Yoshida leg. (EUMJ). [Ehime Pref.] 1 ex., Nishidani, Yanadani Village, 15
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<normalizedToken originalValue="–16–VII–">-16-VII-</normalizedToken>
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1994, K. Aita leg. (EUMJ). [Kochi Pref.] 1 ex., Muroto Cape, Muroto City, 7
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<normalizedToken originalValue="–VI–">-VI-</normalizedToken>
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1959, S. Hisamatsu leg. (EUMJ). [Nagasaki Pref.] 1 ex., Mt. Tatera-san, Tsushima Island 16
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<normalizedToken originalValue="–IX–">-IX-</normalizedToken>
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1995, N. Narukawa leg. (ELKU); 3 exs., Midake, 12
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<normalizedToken originalValue="–X–">-X-</normalizedToken>
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1977, Y. Notsu leg. (EUMJ). [Kagoshima Pref.] 1 male & 4 exs., Kirishima, 29
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<normalizedToken originalValue="–VII–">-VII-</normalizedToken>
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1971, T. Nakane leg. (HUSE); 2 exs.,
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<normalizedToken originalValue="Shiratani-unsuikyô">Shiratani-unsuikyo</normalizedToken>
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, Yakushima Island, 15
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<normalizedToken originalValue="–VIII–">-VIII-</normalizedToken>
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2005, J. Ogawa leg. (EUMJ); 1 male & 1 ex., Onoaida
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<normalizedToken originalValue="Hodô">Hodo</normalizedToken>
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, Yakushima Island, 14
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<normalizedToken originalValue="–15–IX–">-15-IX-</normalizedToken>
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2002, N. Ohbayashi leg. (EUMJ); 2 exs., Kosugidani, Yakushima Island, 2
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<normalizedToken originalValue="–V–">-V-</normalizedToken>
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1984, K. Mizuno leg. (OMNH); 4 exs.,
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<normalizedToken originalValue="Anbô">Anbo</normalizedToken>
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, Yakushima Island, 16
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<normalizedToken originalValue="–VII–">-VII-</normalizedToken>
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1989, N. Narukawa leg. (ELKU); 5 exs., Nogi, Nishino-omote, Tanegashima Island, 10
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<normalizedToken originalValue="–VII–">-VII-</normalizedToken>
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1974, S. Hisamatsu leg. (EUMJ).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="21" pageNumber="36" type="distribution">
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<paragraph pageId="21" pageNumber="36">Distribution.</paragraph>
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<paragraph pageId="21" pageNumber="36">JAPAN: Hokkaido, Honshu, Shikoku, Kyushu, Mikura (Izu Islands), Awaji, Yakushima and Tanegashima Islands; Burma; Korea; Russia.</paragraph>
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</subSubSection>
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<subSubSection pageId="21" pageNumber="36" type="biological notes">
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<paragraph pageId="21" pageNumber="36">Biological notes.</paragraph>
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<paragraph pageId="21" pageNumber="36">The first author (Yoshida) collected this species from various kinds of dead leaves from evergreen trees, deciduous trees and bamboo.</paragraph>
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</subSubSection>
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<subSubSection pageId="21" pageNumber="36" type="remarks">
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<paragraph pageId="21" pageNumber="36">Remarks.</paragraph>
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<paragraph pageId="21" pageNumber="36">
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The syntypes of this species consist of three specimens: two of them belong to this species, but the remaining one represents
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<taxonomicName class="Insecta" family="Silvanidae" genus="Psammoecus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Psammoecus trimaculatus" order="Coleoptera" pageId="21" pageNumber="36" phylum="Arthropoda" rank="species" species="trimaculatus">Psammoecus trimaculatus</taxonomicName>
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or
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<taxonomicName class="Insecta" family="Silvanidae" genus="Psammoecus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Psammoecus triguttatus" order="Coleoptera" pageId="21" pageNumber="36" phylum="Arthropoda" rank="species" species="triguttatus">Psammoecus triguttatus</taxonomicName>
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. We designate a male specimen as lectotype, and a female specimen as paralectotype. The type specimens were collected from Mt. Maya-san, Kobe City, Hyogo.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |