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<document id="57479D011E6F9BDAA9127C4CA78B87F4" ID-CLB-Dataset="141497" ID-DOI="10.11646/zootaxa.5187.1.4" ID-GBIF-Dataset="ad986c9b-507a-40ee-a021-f609269230de" ID-ISSN="1175-5326" ID-Zenodo-Dep="7078226" ID-ZooBank="E2A407A8-BED4-4C26-BE5C-B3579CB9B038" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1663144208795" checkinUser="plazi" docAuthor="Bayartogtokh, Badamdorj, Ermilov, Sergey G. &amp; Joharchi, Omid" docDate="2022" docId="642887FA63716700FF24FA7E5686FDED" docLanguage="en" docName="zootaxa.5187.1.4.pdf" docOrigin="Zootaxa 5187 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Lepidacarus maafushiensis Bayartogtokh &amp; Ermilov &amp; Joharchi 2022, sp. nov." docType="treatment" docVersion="5" lastPageNumber="16" masterDocId="9811FF826373670AFFB3FFFC5229FFD0" masterDocTitle="Ontogenetic instars of Lepidacarus maafushiensis sp. nov. from the Maldives, with remarks on morphological ontogeny of Lohmanniidae (Acari, Oribatida)" masterLastPageNumber="29" masterPageNumber="7" pageNumber="9" updateTime="1699380459367" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="F8A20F5359A0FD9EBC5D66B608CE2D95">Ontogenetic instars of Lepidacarus maafushiensis sp. nov. from the Maldives, with remarks on morphological ontogeny of Lohmanniidae (Acari, Oribatida)</mods:title>
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<mods:namePart id="A49FA46C02C8A94A23A5AE53F267B73A">Bayartogtokh, Badamdorj</mods:namePart>
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<mods:affiliation id="11BFCF95E73E8AB4D9FC01A07BB2A4CB">Department of Biology, School of Arts and Sciences, National University of Mongolia, Ulaanbaatar, Mongolia. bayartogtokh @ num. edu. mn; https: // orcid. org / 0000 - 0002 - 6370 - 5759</mods:affiliation>
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<mods:namePart id="47CECF28464F1734AF58B888EF351532">Ermilov, Sergey G.</mods:namePart>
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<mods:affiliation id="44D5BDA2843058E1925A984CA2ACE86F">Institute of Environmental and Agricultural Biology (X-BIO), Tyumen State University, Tyumen, Russia. ermilovacari @ yandex. ru; https: // orcid. org / 0000 - 0002 - 0913 - 131 X</mods:affiliation>
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<mods:namePart id="00D6AD27A6676188D7F4BA9279371071">Joharchi, Omid</mods:namePart>
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<mods:affiliation id="9B69A45D2E4B7AF844FF719630F423F1">Institute of Environmental and Agricultural Biology (X-BIO), Tyumen State University, Tyumen, Russia. ermilovacari @ yandex. ru; https: // orcid. org / 0000 - 0002 - 0913 - 131 X</mods:affiliation>
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<taxonomicName id="2B814D6F63716708FF24FA7E53CBFA4D" authority="Bayartogtokh &amp; Ermilov &amp; Joharchi, 2022" authorityName="Bayartogtokh &amp; Ermilov &amp; Joharchi" authorityYear="2022" box="[151,482,1410,1437]" class="Arachnida" family="Lohmanniidae" genus="Lepidacarus" kingdom="Animalia" order="Sarcoptiformes" pageId="2" pageNumber="9" phylum="Arthropoda" rank="species" species="maafushiensis" status="sp. nov.">
<emphasis id="DEF5EAFE63716708FF24FA7E53CBFA4D" bold="true" box="[151,482,1410,1437]" italics="true" pageId="2" pageNumber="9">Lepidacarus maafushiensis</emphasis>
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(
<figureCitation id="74BA2A6963716708FF2CFA5B5338FA11" box="[159,273,1447,1473]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-10="FIGURE 11" captionStart-11="FIGURE 12" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStart-6="FIGURE 7" captionStart-7="FIGURE 8" captionStart-8="FIGURE 9" captionStart-9="FIGURE 10" captionStartId-0="3.[151,250,1564,1588]" captionStartId-1="4.[151,250,1504,1528]" captionStartId-10="14.[151,250,1643,1667]" captionStartId-11="15.[151,250,1869,1893]" captionStartId-2="5.[151,250,1665,1689]" captionStartId-3="7.[151,250,1783,1807]" captionStartId-4="8.[151,250,1639,1663]" captionStartId-5="10.[151,250,1965,1989]" captionStartId-6="11.[151,250,1436,1460]" captionStartId-7="12.[151,250,1028,1052]" captionStartId-8="12.[151,250,1970,1994]" captionStartId-9="13.[151,250,1728,1752]" captionTargetBox-0="[167,1413,390,1520]" captionTargetBox-1="[179,1423,234,1438]" captionTargetBox-10="[151,1436,181,1615]" captionTargetBox-11="[151,1436,181,1845]" captionTargetBox-2="[165,1421,199,1593]" captionTargetBox-3="[151,1436,309,1744]" captionTargetBox-4="[151,1436,181,1615]" captionTargetBox-5="[207,1368,631,1889]" captionTargetBox-6="[175,1422,201,1381]" captionTargetBox-7="[350,1247,206,982]" captionTargetBox-8="[294,1309,1157,1931]" captionTargetBox-9="[160,1430,181,1676]" captionTargetId-0="figure-107@3.[151,1436,378,1537]" captionTargetId-1="figure-17@4.[151,1436,181,1481]" captionTargetId-10="figure-17@14.[151,1436,181,1619]" captionTargetId-11="figure-17@15.[151,1436,181,1845]" captionTargetId-2="figure-17@5.[151,1436,181,1641]" captionTargetId-3="figure-79@7.[151,1436,309,1760]" captionTargetId-4="figure-17@8.[151,1436,181,1615]" captionTargetId-5="figure-267@10.[151,1436,597,1942]" captionTargetId-6="figure-17@11.[151,1436,181,1412]" captionTargetId-7="figure-17@12.[325,1262,181,1004]" captionTargetId-8="figure-41@12.[273,1314,1112,1946]" captionTargetId-9="figure-17@13.[151,1436,181,1704]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-10="14" captionTargetPageId-11="15" captionTargetPageId-2="5" captionTargetPageId-3="7" captionTargetPageId-4="8" captionTargetPageId-5="10" captionTargetPageId-6="11" captionTargetPageId-7="12" captionTargetPageId-8="12" captionTargetPageId-9="13" captionText-0="FIGURE 1. Lepidacarus maafushiensis sp. nov., adult: A—dorsal view; B—ventral view (gnathosoma and legs except trochanters not shown). Scale bar 100 μm." captionText-1="FIGURE 2. Lepidacarus maafushiensis sp. nov., adult (A, C, E) and larva (B, D, F): A, B—subcapitulum, ventral view; C, D—palp, left, paraxial view; E—chelicera (basal part not shown), left, paraxial view; F—chelicera, left, paraxial view. Scale bar 20 μm (A–C, E, F), 10 μm (D)." captionText-10="FIGURE 11. Lepidacarus maafushiensis sp. nov., tritonymph, SEM micrographs: A—dorsal view; B—part of gastronotic region, dorsal view; C—ventral view; D—anterodorsal view; E—part of prodorsum, dorsal view." captionText-11="FIGURE 12. Lepidacarus maafushiensis sp. nov., deuto- (A–D; F) and tritonymph (E), microscope images: A—part of prodorsum, lateral view; B—part of ventral side, ventral view; C—subcapitulum and palp, ventral view; D—part of epimeral region, ventral view; E—genital plate, right, ventral view; F—leg III, right, antiaxial view." captionText-2="FIGURE 3. Lepidacarus maafushiensis sp. nov., adult:A—leg I, without trochanter, right, antiaxial view; B—femur, genu and tibia of leg II, right, antiaxial view; C—femur, genu and tibia of leg III, left, antiaxial view; D—leg IV, without trochanter, left, antiaxial view. Scale bar 20 μm." captionText-3="FIGURE 4. Lepidacarus maafushiensis sp. nov., adult, SEM micrographs: A—dorsal view; B—ventral view; C—ventrolateral view; D—anterodorsal view." captionText-4="FIGURE 5. Lepidacarus maafushiensis sp. nov., adult, SEM micrographs: A—posterior view; B—gnathosoma, lateral view; C—genital region, ventrolateral view; D—anoadanal region, lateral view." captionText-5="FIGURE 6. Lepidacarus maafushiensis sp. nov., larva: A—dorsal view; B—ventral view (gnathosoma and legs except trochanters not shown); C—part of prodorsum and notogaster, lateral view. Scale bar 50 μm." captionText-6="FIGURE 7. Lepidacarus maafushiensis sp. nov., dorsal view of juvenile instars: A –protonymph; B—tritonymph. Scale bar 50 μm (A), 100 μm (B)." captionText-7="FIGURE 8. Lepidacarus maafushiensis sp. nov., epimeral region of juvenile instars: A –protonymph; B—deutonymph; C— tritonymph. Scale bar 50 μm." captionText-8="FIGURE 9. Lepidacarus maafushiensis sp. nov., anogenital region of juvenile instars: A –protonymph; B—deutonymph; C— tritonymph. Scale bar 50 μm." captionText-9="FIGURE 10. Lepidacarus maafushiensis sp. nov., larva (A–C) and protonymph (D):A—leg I, without trochanter, left, paraxial view; B—leg II, without trochanter, left, paraxial view; C—leg III, without trochanter, left, antiaxial view; D—leg IV, without trochanter, right, antiaxial view. Scale bar 20 μm." figureDoi-0="http://doi.org/10.5281/zenodo.7078228" figureDoi-1="http://doi.org/10.5281/zenodo.7078230" figureDoi-10="http://doi.org/10.5281/zenodo.7078252" figureDoi-11="http://doi.org/10.5281/zenodo.7078254" figureDoi-2="http://doi.org/10.5281/zenodo.7078232" figureDoi-3="http://doi.org/10.5281/zenodo.7078234" figureDoi-4="http://doi.org/10.5281/zenodo.7078236" figureDoi-5="http://doi.org/10.5281/zenodo.7078242" figureDoi-6="http://doi.org/10.5281/zenodo.7078244" figureDoi-7="http://doi.org/10.5281/zenodo.7078246" figureDoi-8="http://doi.org/10.5281/zenodo.7078248" figureDoi-9="http://doi.org/10.5281/zenodo.7078250" httpUri-0="https://zenodo.org/record/7078228/files/figure.png" httpUri-1="https://zenodo.org/record/7078230/files/figure.png" httpUri-10="https://zenodo.org/record/7078252/files/figure.png" httpUri-11="https://zenodo.org/record/7078254/files/figure.png" httpUri-2="https://zenodo.org/record/7078232/files/figure.png" httpUri-3="https://zenodo.org/record/7078234/files/figure.png" httpUri-4="https://zenodo.org/record/7078236/files/figure.png" httpUri-5="https://zenodo.org/record/7078242/files/figure.png" httpUri-6="https://zenodo.org/record/7078244/files/figure.png" httpUri-7="https://zenodo.org/record/7078246/files/figure.png" httpUri-8="https://zenodo.org/record/7078248/files/figure.png" httpUri-9="https://zenodo.org/record/7078250/files/figure.png" pageId="2" pageNumber="9">Figs 1–12</figureCitation>
)
</paragraph>
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<subSubSection id="A49B656763716709FF24FA12530BFEB5" lastPageId="3" lastPageNumber="10" pageId="2" pageNumber="9" type="diagnosis">
<paragraph id="EC3E36EC63716709FF24FA12530BFEB5" blockId="2.[151,1437,1518,2013]" lastBlockId="3.[151,1437,151,357]" lastPageId="3" lastPageNumber="10" pageId="2" pageNumber="9">
<emphasis id="DEF5EAFE63716708FF24FA12533BF9D8" bold="true" box="[151,274,1518,1544]" pageId="2" pageNumber="9">Diagnosis.</emphasis>
<emphasis id="DEF5EAFE63716708FEA9FA13537EF9D9" box="[282,343,1519,1545]" italics="true" pageId="2" pageNumber="9">Adult</emphasis>
. Body length: 531–581. Prodorsum, notogaster, partially subcapitular mentum and epimeres III, IV heavily tuberculate. Rostrum broadly concave, with median triangular tooth. Rostral, lamellar, interlamellar, anterior exobothridial, posterior exobothridial, notogastral, and adanal setae widely phylliform, rounded distally, heavily barbed. Bothridial seta long, narrowly elongate fusiform, with sparsely ciliate one side and densely serrate opposite side. Notogaster with numerous oval depressions in medioposterior part. Eighteen pairs of notogastral setae present including two pairs of additional setae. Dorsal transverse bands present. Subcapitulum with four pairs of widely phylliform, rounded distally, ciliate setae (
<emphasis id="DEF5EAFE63716708FD41F93B512FF930" box="[754,774,1735,1760]" italics="true" pageId="2" pageNumber="9">m</emphasis>
<subScript id="700534A963716708FCB5F9255127F937" attach="left" box="[774,782,1753,1767]" fontSize="6" pageId="2" pageNumber="9">2</subScript>
, 
<emphasis id="DEF5EAFE63716708FCADF93B511BF930" box="[798,818,1735,1760]" italics="true" pageId="2" pageNumber="9">m</emphasis>
<subScript id="700534A963716708FC81F9255113F937" attach="left" box="[818,826,1753,1767]" fontSize="6" pageId="2" pageNumber="9">3</subScript>
, 
<emphasis id="DEF5EAFE63716708FCF8F93B5176F930" box="[843,863,1735,1760]" italics="true" pageId="2" pageNumber="9">m</emphasis>
<subScript id="700534A963716708FCECF925514EF937" attach="left" box="[863,871,1753,1767]" fontSize="6" pageId="2" pageNumber="9">4</subScript>
, and 
<emphasis id="DEF5EAFE63716708FC1AF93B519EF931" box="[937,951,1735,1761]" italics="true" pageId="2" pageNumber="9">h</emphasis>
). Palp setation: 0–1–0–2–9(+ω). Epimeral setal formula: 10–7–3–4; all setae widely phylliform, rounded distally, ciliate. One pair of genital setae widely phylliform, rounded distally, ciliate; other setae as well as anal setae setiform, shortly ciliate. Tarsus II with two solenidia. Many leg setae widely phylliform, rounded distally, ciliate. 
<emphasis id="DEF5EAFE63716708FC10F8CF567BF89C" box="[931,1106,1843,1869]" italics="true" pageId="2" pageNumber="9">Juvenile instars</emphasis>
. Prodorsum and gastronotum sparsely microtuberculate. Rostrum broadly concave, with median triangular tooth. Rostral, lamellar, interlamellar, anterior exobothridial, posterior exobothridial, true gastronotic, and true adanal setae widely phylliform, rounded distally, heavily barbed. Bothridial seta long, narrowly elongate fusiform, with sparsely ciliate one side and densely serrate opposite side. Gastronotum with numerous oval depressions in medioposterior part. Larva with 16 pairs of gastronotic setae including three pairs of 
<emphasis id="DEF5EAFE63706709FD2DFF6B5085FF60" box="[670,684,151,176]" italics="true" pageId="3" pageNumber="10">p</emphasis>
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– 
<emphasis id="DEF5EAFE63706709FD71FF6B50F9FF60" box="[706,720,151,176]" italics="true" pageId="3" pageNumber="10">p</emphasis>
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on pseudoanal plates; protonymph with 17 pairs (added 
<emphasis id="DEF5EAFE63706709FAC5FF6B57ADFF60" box="[1398,1412,151,176]" italics="true" pageId="3" pageNumber="10">n</emphasis>
<subScript id="700534A963706709FA37FF5557A5FF67" attach="left" box="[1412,1420,169,183]" fontSize="6" pageId="3" pageNumber="10">1</subScript>
), deuto- (added 
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<subScript id="700534A963706709FEF5FF315367FF0B" attach="left" box="[326,334,205,219]" fontSize="6" pageId="3" pageNumber="10">2</subScript>
) and tritonymph with 18 pairs of gastronotic setae. Palp setation of larva and protonymph: 0–0–0– 2–9(+ω); seta on femur added in deutonymph. Setal formulas for epimeres: larva: 3–1–2; protonymph: 4–2–2–1; deutonymph: 6–4–3–3, tritonymph: 8–5–3–4; seta 
<emphasis id="DEF5EAFE63706709FD7AFEFF50CAFECD" box="[713,739,259,285]" italics="true" pageId="3" pageNumber="10">1c</emphasis>
in larva setiform, shortly ciliate; other setae widely phylliform, rounded distally, ciliate. Deutonymph and tritonymph with one pair of lateral, widely phylliform, rounded distally, ciliate setae.
</paragraph>
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<caption id="B8FE666463706709FF24F9E0506CF988" ID-DOI="http://doi.org/10.5281/zenodo.7078228" ID-Zenodo-Dep="7078228" httpUri="https://zenodo.org/record/7078228/files/figure.png" pageId="3" pageNumber="10" startId="3.[151,250,1564,1588]" targetBox="[167,1413,390,1520]" targetPageId="3">
<paragraph id="EC3E36EC63706709FF24F9E0506CF988" blockId="3.[151,1436,1564,1624]" pageId="3" pageNumber="10">
<emphasis id="DEF5EAFE63706709FF24F9E05331F9E4" bold="true" box="[151,280,1564,1588]" pageId="3" pageNumber="10">FIGURE 1.</emphasis>
<taxonomicName id="2B814D6F63706709FE97F9E05017F9E4" authority="Bayartogtokh &amp; Ermilov &amp; Joharchi, 2022" authorityName="Bayartogtokh &amp; Ermilov &amp; Joharchi" authorityYear="2022" box="[292,574,1564,1588]" class="Arachnida" family="Lohmanniidae" genus="Lepidacarus" kingdom="Animalia" order="Sarcoptiformes" pageId="3" pageNumber="10" phylum="Arthropoda" rank="species" species="maafushiensis" status="sp. nov.">
<emphasis id="DEF5EAFE63706709FE97F9E05017F9E4" box="[292,574,1564,1588]" italics="true" pageId="3" pageNumber="10">Lepidacarus maafushiensis</emphasis>
</taxonomicName>
<emphasis id="DEF5EAFE63706709FDFAF9E050B6F9E4" bold="true" box="[585,671,1564,1588]" pageId="3" pageNumber="10">
<taxonomicNameLabel id="C5C6578563706709FDFAF9E050B6F9E4" box="[585,671,1564,1588]" pageId="3" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, adult: A—dorsal view; B—ventral view (gnathosoma and legs except trochanters not shown). Scale bar 100 μm.
</paragraph>
</caption>
<subSubSection id="A49B656763706702FF24F956505DF813" lastPageId="8" lastPageNumber="15" pageId="3" pageNumber="10" type="description">
<paragraph id="EC3E36EC63706709FF24F95652F6F914" blockId="3.[151,267,1706,1769]" box="[151,223,1706,1732]" pageId="3" pageNumber="10">
<heading id="B776818063706709FF24F95652F6F914" bold="true" box="[151,223,1706,1732]" fontSize="11" level="1" pageId="3" pageNumber="10" reason="1">
<emphasis id="DEF5EAFE63706709FF24F95652F6F914" bold="true" box="[151,223,1706,1732]" pageId="3" pageNumber="10">Adult</emphasis>
</heading>
</paragraph>
<paragraph id="EC3E36EC63706709FF24F9325322F939" blockId="3.[151,267,1706,1769]" box="[151,267,1742,1769]" pageId="3" pageNumber="10">
(
<figureCitation id="74BA2A6963706709FF2CF932532BF939" box="[159,258,1742,1769]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStartId-0="3.[151,250,1564,1588]" captionStartId-1="4.[151,250,1504,1528]" captionStartId-2="5.[151,250,1665,1689]" captionStartId-3="7.[151,250,1783,1807]" captionStartId-4="8.[151,250,1639,1663]" captionTargetBox-0="[167,1413,390,1520]" captionTargetBox-1="[179,1423,234,1438]" captionTargetBox-2="[165,1421,199,1593]" captionTargetBox-3="[151,1436,309,1744]" captionTargetBox-4="[151,1436,181,1615]" captionTargetId-0="figure-107@3.[151,1436,378,1537]" captionTargetId-1="figure-17@4.[151,1436,181,1481]" captionTargetId-2="figure-17@5.[151,1436,181,1641]" captionTargetId-3="figure-79@7.[151,1436,309,1760]" captionTargetId-4="figure-17@8.[151,1436,181,1615]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="7" captionTargetPageId-4="8" captionText-0="FIGURE 1. Lepidacarus maafushiensis sp. nov., adult: A—dorsal view; B—ventral view (gnathosoma and legs except trochanters not shown). Scale bar 100 μm." captionText-1="FIGURE 2. Lepidacarus maafushiensis sp. nov., adult (A, C, E) and larva (B, D, F): A, B—subcapitulum, ventral view; C, D—palp, left, paraxial view; E—chelicera (basal part not shown), left, paraxial view; F—chelicera, left, paraxial view. Scale bar 20 μm (A–C, E, F), 10 μm (D)." captionText-2="FIGURE 3. Lepidacarus maafushiensis sp. nov., adult:A—leg I, without trochanter, right, antiaxial view; B—femur, genu and tibia of leg II, right, antiaxial view; C—femur, genu and tibia of leg III, left, antiaxial view; D—leg IV, without trochanter, left, antiaxial view. Scale bar 20 μm." captionText-3="FIGURE 4. Lepidacarus maafushiensis sp. nov., adult, SEM micrographs: A—dorsal view; B—ventral view; C—ventrolateral view; D—anterodorsal view." captionText-4="FIGURE 5. Lepidacarus maafushiensis sp. nov., adult, SEM micrographs: A—posterior view; B—gnathosoma, lateral view; C—genital region, ventrolateral view; D—anoadanal region, lateral view." figureDoi-0="http://doi.org/10.5281/zenodo.7078228" figureDoi-1="http://doi.org/10.5281/zenodo.7078230" figureDoi-2="http://doi.org/10.5281/zenodo.7078232" figureDoi-3="http://doi.org/10.5281/zenodo.7078234" figureDoi-4="http://doi.org/10.5281/zenodo.7078236" httpUri-0="https://zenodo.org/record/7078228/files/figure.png" httpUri-1="https://zenodo.org/record/7078230/files/figure.png" httpUri-2="https://zenodo.org/record/7078232/files/figure.png" httpUri-3="https://zenodo.org/record/7078234/files/figure.png" httpUri-4="https://zenodo.org/record/7078236/files/figure.png" pageId="3" pageNumber="10">Figs 1–5</figureCitation>
)
</paragraph>
<paragraph id="EC3E36EC63706709FF24F8EA56E2F884" blockId="3.[151,1436,1814,2020]" pageId="3" pageNumber="10">
<emphasis id="DEF5EAFE63706709FF24F8EA5300F8E0" bold="true" box="[151,297,1814,1840]" pageId="3" pageNumber="10">Description.</emphasis>
<emphasis id="DEF5EAFE63706709FE82F8EB53FAF8E0" box="[305,467,1815,1840]" italics="true" pageId="3" pageNumber="10">Measurements</emphasis>
. Medium sized species as compared to other species of the genus. Body length: 564 (
<typeStatus id="333A884E63706709FF2CF8C6532EF884" box="[159,263,1850,1876]" pageId="3" pageNumber="10" type="holotype">holotype</typeStatus>
), 531–581 (
<specimenCount id="FA87FD6563706709FE38F8C6500CF884" box="[395,549,1850,1876]" pageId="3" pageNumber="10" type="generic" typeStatus="paratypes">six paratypes</specimenCount>
); notogaster width: 282 (
<typeStatus id="333A884E63706709FC89F8C6518BF884" box="[826,930,1850,1876]" pageId="3" pageNumber="10" type="holotype">holotype</typeStatus>
), 249–298 (
<specimenCount id="FA87FD6563706709FB95F8C656E9F884" box="[1062,1216,1850,1876]" pageId="3" pageNumber="10" type="generic" typeStatus="paratypes">six paratypes</specimenCount>
).
</paragraph>
<paragraph id="EC3E36EC63706709FF74F8A3530CF834" blockId="3.[151,1436,1814,2020]" pageId="3" pageNumber="10">
<emphasis id="DEF5EAFE63706709FF74F8A3536CF8A8" box="[199,325,1887,1912]" italics="true" pageId="3" pageNumber="10">Integument</emphasis>
. Body colour yellowish-brown. Surface of body and legs densely microreticulate (micropolygonal ornamentation).Also, prodorsum, notogaster, partially subcapitular mentum and epimeres III, IV heavily tuberculate (diameter of tubercle up to 12) and partially densely microgranulate. Macroreticulate (macropolygonal) pattern on body absent.
</paragraph>
<caption id="B8FE66646377670EFF24FA1C53D2F991" ID-DOI="http://doi.org/10.5281/zenodo.7078230" ID-Zenodo-Dep="7078230" httpUri="https://zenodo.org/record/7078230/files/figure.png" pageId="4" pageNumber="11" startId="4.[151,250,1504,1528]" targetBox="[179,1423,234,1438]" targetPageId="4">
<paragraph id="EC3E36EC6377670EFF24FA1C53D2F991" blockId="4.[151,1436,1504,1601]" pageId="4" pageNumber="11">
<emphasis id="DEF5EAFE6377670EFF24FA1C533CFA28" bold="true" box="[151,277,1504,1528]" pageId="4" pageNumber="11">FIGURE 2.</emphasis>
<taxonomicName id="2B814D6F6377670EFEAEFA1C501DFA28" authority="Bayartogtokh &amp; Ermilov &amp; Joharchi, 2022" authorityName="Bayartogtokh &amp; Ermilov &amp; Joharchi" authorityYear="2022" box="[285,564,1504,1528]" class="Arachnida" family="Lohmanniidae" genus="Lepidacarus" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="11" phylum="Arthropoda" rank="species" species="maafushiensis" status="sp. nov.">
<emphasis id="DEF5EAFE6377670EFEAEFA1C501DFA28" box="[285,564,1504,1528]" italics="true" pageId="4" pageNumber="11">Lepidacarus maafushiensis</emphasis>
</taxonomicName>
<emphasis id="DEF5EAFE6377670EFD88FA1D50A7FA29" bold="true" box="[571,654,1505,1529]" pageId="4" pageNumber="11">
<taxonomicNameLabel id="C5C657856377670EFD88FA1D50A7FA29" box="[571,654,1505,1529]" pageId="4" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, adult (A, C, E) and larva (B, D, F): A, B—subcapitulum, ventral view; C, D—palp, left, paraxial view; E—chelicera (basal part not shown), left, paraxial view; F—chelicera, left, paraxial view. Scale bar 20 μm (A–C, E, F), 10 μm (D).
</paragraph>
</caption>
<paragraph id="EC3E36EC6377670EFF74F993535FF8B1" blockId="4.[151,1437,1647,2033]" pageId="4" pageNumber="11">
<emphasis id="DEF5EAFE6377670EFF74F993536DF959" box="[199,324,1647,1673]" italics="true" pageId="4" pageNumber="11">Prodorsum</emphasis>
. Roughly triangular in dorsal view, occupying about 2/3 of dorsal length. Rostrum broadly concave, with median triangular tooth. Lateral side of rostrum with some slight barbs and partially undulate. Rostral, lamellar, interlamellar, anterior exobothridial, and posterior exobothridial setae widely phylliform, rounded distally, heavily barbed. Bothridial seta (77–82) with short stalk and narrowly elongate fusiform head having sparsely ciliate one side and densely serrate opposite side. Lateral tubercle of prodorsum (close and anterolateral to anterior exobothridial seta) well developed. Region between lamellar and interlamellar setae with some depressions. Postbothridial band present, very broad.
</paragraph>
<paragraph id="EC3E36EC6377670EFF74F890573AF821" blockId="4.[151,1437,1647,2033]" pageId="4" pageNumber="11">
<emphasis id="DEF5EAFE6377670EFF74F890536AF855" box="[199,323,1900,1925]" italics="true" pageId="4" pageNumber="11">Notogaster</emphasis>
. With numerous oval depressions in medioposterior part. Eighteen pairs of notogastral setae present (sixteen pairs of basic setae and two pairs— 
<emphasis id="DEF5EAFE6377670EFDC9F86C50A1F879" box="[634,648,1936,1961]" italics="true" pageId="4" pageNumber="11">n</emphasis>
<subScript id="700534A96377670EFD3BF85D50B9F87F" attach="left" box="[648,656,1953,1967]" fontSize="6" pageId="4" pageNumber="11">1</subScript>
, 
<emphasis id="DEF5EAFE6377670EFD2FF86C5083F879" box="[668,682,1936,1961]" italics="true" pageId="4" pageNumber="11">n</emphasis>
<subScript id="700534A96377670EFD19F85D509BF87F" attach="left" box="[682,690,1953,1967]" fontSize="6" pageId="4" pageNumber="11">2</subScript>
—of additional setae); all setae widely phylliform, rounded distally, heavily barbed. Five transverse bands (others not visible) present dorsally; 
<emphasis id="DEF5EAFE6377670EFC53F84F51C7F81D" box="[992,1006,1971,1997]" italics="true" pageId="4" pageNumber="11">S</emphasis>
<subScript id="700534A96377670EFC5DF83951DFF803" attach="left" box="[1006,1014,1989,2003]" fontSize="6" pageId="4" pageNumber="11">2</subScript>
complete, 
<emphasis id="DEF5EAFE6377670EFBC1F84F56A9F81D" box="[1138,1152,1971,1997]" italics="true" pageId="4" pageNumber="11">S</emphasis>
<subScript id="700534A96377670EFB33F83956A1F803" attach="left" box="[1152,1160,1989,2003]" fontSize="6" pageId="4" pageNumber="11">3</subScript>
– 
<emphasis id="DEF5EAFE6377670EFB25F84F568DF81D" box="[1174,1188,1971,1997]" italics="true" pageId="4" pageNumber="11">S</emphasis>
<subScript id="700534A96377670EFB17F8395685F803" attach="left" box="[1188,1196,1989,2003]" fontSize="6" pageId="4" pageNumber="11">6</subScript>
interrupted medially. Lyrifissures 
<emphasis id="DEF5EAFE6377670EFE91F8245311F821" box="[290,312,2008,2033]" italics="true" pageId="4" pageNumber="11">ia</emphasis>
, 
<emphasis id="DEF5EAFE6377670EFEF5F824534BF821" box="[326,354,2008,2033]" italics="true" pageId="4" pageNumber="11">im</emphasis>
, 
<emphasis id="DEF5EAFE6377670EFEC3F82B53AFF821" box="[368,390,2007,2033]" italics="true" pageId="4" pageNumber="11">ih</emphasis>
, and 
<emphasis id="DEF5EAFE6377670EFE70F82453F0F821" box="[451,473,2008,2033]" italics="true" pageId="4" pageNumber="11">ip</emphasis>
distinct; 
<emphasis id="DEF5EAFE6377670EFDF2F824504BF821" box="[577,610,2008,2033]" italics="true" pageId="4" pageNumber="11">ips</emphasis>
not observed. Ventral bands close to adanal region indistinct.
</paragraph>
<caption id="B8FE66646376670FFF24F97D53F5F931" ID-DOI="http://doi.org/10.5281/zenodo.7078232" ID-Zenodo-Dep="7078232" httpUri="https://zenodo.org/record/7078232/files/figure.png" pageId="5" pageNumber="12" startId="5.[151,250,1665,1689]" targetBox="[165,1421,199,1593]" targetPageId="5">
<paragraph id="EC3E36EC6376670FFF24F97D53F5F931" blockId="5.[151,1437,1665,1762]" pageId="5" pageNumber="12">
<emphasis id="DEF5EAFE6376670FFF24F97D533BF949" bold="true" box="[151,274,1665,1689]" pageId="5" pageNumber="12">FIGURE 3.</emphasis>
<taxonomicName id="2B814D6F6376670FFEABF97D5004F949" authority="Bayartogtokh &amp; Ermilov &amp; Joharchi, 2022" authorityName="Bayartogtokh &amp; Ermilov &amp; Joharchi" authorityYear="2022" box="[280,557,1665,1689]" class="Arachnida" family="Lohmanniidae" genus="Lepidacarus" kingdom="Animalia" order="Sarcoptiformes" pageId="5" pageNumber="12" phylum="Arthropoda" rank="species" species="maafushiensis" status="sp. nov.">
<emphasis id="DEF5EAFE6376670FFEABF97D5004F949" box="[280,557,1665,1689]" italics="true" pageId="5" pageNumber="12">Lepidacarus maafushiensis</emphasis>
</taxonomicName>
<emphasis id="DEF5EAFE6376670FFD81F97D50ABF949" bold="true" box="[562,642,1665,1689]" pageId="5" pageNumber="12">
<taxonomicNameLabel id="C5C657856376670FFD81F97D50ABF949" box="[562,642,1665,1689]" pageId="5" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, adult: A—leg I, without trochanter, right, antiaxial view; B—femur, genu and tibia of leg II, right, antiaxial view; C—femur, genu and tibia of leg III, left, antiaxial view; D—leg IV, without trochanter, left, antiaxial view. Scale bar 20 μm.
</paragraph>
</caption>
<paragraph id="EC3E36EC6376670FFF74F8F15177F828" blockId="5.[151,1437,1805,2040]" pageId="5" pageNumber="12">
<emphasis id="DEF5EAFE6376670FFF74F8F1537FF8F7" box="[199,342,1805,1831]" italics="true" pageId="5" pageNumber="12">Gnathosoma</emphasis>
. Subcapitulum size: 139–147 × 110–114. Six pairs of subcapitular setae developed including four pairs of 
<emphasis id="DEF5EAFE6376670FFF47F8CC5321F899" box="[244,264,1840,1865]" italics="true" pageId="5" pageNumber="12">m</emphasis>
; 
<emphasis id="DEF5EAFE6376670FFEABF8CC530FF899" box="[280,294,1840,1865]" italics="true" pageId="5" pageNumber="12">a</emphasis>
(41–45) setiform, smooth; 
<emphasis id="DEF5EAFE6376670FFDE8F8CC5046F899" box="[603,623,1840,1865]" italics="true" pageId="5" pageNumber="12">m</emphasis>
<subScript id="700534A96376670FFDC3F8BE5051F880" attach="left" box="[624,632,1858,1872]" fontSize="6" pageId="5" pageNumber="12">1</subScript>
(49–53) setiform, slightly barbed; 
<emphasis id="DEF5EAFE6376670FFBB0F8CC563EF899" box="[1027,1047,1840,1865]" italics="true" pageId="5" pageNumber="12">m</emphasis>
<subScript id="700534A96376670FFBA4F8BE5636F880" attach="left" box="[1047,1055,1858,1872]" fontSize="6" pageId="5" pageNumber="12">2</subScript>
, 
<emphasis id="DEF5EAFE6376670FFB9DF8CC566BF899" box="[1070,1090,1840,1865]" italics="true" pageId="5" pageNumber="12">m</emphasis>
<subScript id="700534A96376670FFBF1F8BE5663F880" attach="left" box="[1090,1098,1858,1872]" fontSize="6" pageId="5" pageNumber="12">3</subScript>
, 
<emphasis id="DEF5EAFE6376670FFBEAF8CC5644F899" box="[1113,1133,1840,1865]" italics="true" pageId="5" pageNumber="12">m</emphasis>
<subScript id="700534A96376670FFBDEF8BE565CF880" attach="left" box="[1133,1141,1858,1872]" fontSize="6" pageId="5" pageNumber="12">4</subScript>
, and 
<emphasis id="DEF5EAFE6376670FFB07F8D356EBF899" box="[1204,1218,1839,1865]" italics="true" pageId="5" pageNumber="12">h</emphasis>
widely phylliform, rounded distally, ciliate. Adoral seta 
<emphasis id="DEF5EAFE6376670FFDFAF8AF504BF8BC" box="[585,610,1875,1900]" italics="true" pageId="5" pageNumber="12">or</emphasis>
<subScript id="700534A96376670FFDD1F8985043F8A2" attach="left" box="[610,618,1892,1906]" fontSize="6" pageId="5" pageNumber="12">1</subScript>
lobe-shaped, slightly roughened distally; 
<emphasis id="DEF5EAFE6376670FFBEBF8AF5658F8BC" box="[1112,1137,1875,1900]" italics="true" pageId="5" pageNumber="12">or</emphasis>
<subScript id="700534A96376670FFBC2F8985650F8A2" attach="left" box="[1137,1145,1892,1906]" fontSize="6" pageId="5" pageNumber="12">2</subScript>
(28–32) setiform, bluntended, with one small barb in distal part; 
<emphasis id="DEF5EAFE6376670FFD32F88A50B3F85F" box="[641,666,1910,1935]" italics="true" pageId="5" pageNumber="12">or</emphasis>
<subScript id="700534A96376670FFD29F87B508BF845" attach="left" box="[666,674,1927,1941]" fontSize="6" pageId="5" pageNumber="12">3</subScript>
(22–24) setiform, with attenuate tip, smooth. Palp length: 49–53; palp setation: 0–1–0–2–9(+ω); second tarsal—posterior—seta 
<emphasis id="DEF5EAFE6376670FFCD8F86B51A3F861" box="[875,906,1943,1969]" italics="true" pageId="5" pageNumber="12">lt”</emphasis>
absent; three distal setae of tarsus— 
<emphasis id="DEF5EAFE6376670FFA8DF86B5774F861" box="[1342,1373,1943,1969]" italics="true" pageId="5" pageNumber="12">ul’</emphasis>
, 
<emphasis id="DEF5EAFE6376670FFAC3F86B57BFF861" box="[1392,1430,1943,1969]" italics="true" pageId="5" pageNumber="12">ul”</emphasis>
, 
<emphasis id="DEF5EAFE6376670FFF24F8475299F804" box="[151,176,1979,2004]" italics="true" pageId="5" pageNumber="12">su</emphasis>
—connected basally. Postpalpal seta (12) stick-like, roughened. Chelicera length: 168–176. Cheliceral seta 
<emphasis id="DEF5EAFE6376670FFAC7F84657B5F804" box="[1396,1436,1978,2004]" italics="true" pageId="5" pageNumber="12">cha</emphasis>
(10–12) needleform, smooth; 
<emphasis id="DEF5EAFE6376670FFE55F8225027F828" box="[486,526,2014,2040]" italics="true" pageId="5" pageNumber="12">chb</emphasis>
(36–41) setiform, roughened.
</paragraph>
<paragraph id="EC3E36EC6375670CFF74FF6B538CFF05" blockId="6.[151,1437,151,609]" pageId="6" pageNumber="13">
<emphasis id="DEF5EAFE6375670CFF74FF6B5088FF60" box="[199,673,151,177]" italics="true" pageId="6" pageNumber="13">Epimeral and lateral podosomal regions</emphasis>
. Epimeral setal formula: 10–7–3–4. All setae widely phylliform, rounded distally, ciliate.
</paragraph>
<paragraph id="EC3E36EC6375670CFF74FF23532DFE59" blockId="6.[151,1437,151,609]" pageId="6" pageNumber="13">
<emphasis id="DEF5EAFE6375670CFF74FF2353BBFF28" box="[199,402,223,249]" italics="true" pageId="6" pageNumber="13">Anogenital region</emphasis>
. Lateral pair of genital setae larger than others, widely phylliform, rounded distally, ciliate; other setae setiform, shortly ciliate (of these, five pairs of medial setae shorter, 20–24; four pairs of mediolateral setae longer, 28–32). Transverse furrow of genital plate distinct. Two pairs of anal setae (28–32) setiform, shortly ciliate. Four pairs of adanal setae widely phylliform, rounded distally, barbed. Anal and adanal lyrifissures not observed.
</paragraph>
<paragraph id="EC3E36EC6375670CFF74FE6F536BFDB1" blockId="6.[151,1437,151,609]" pageId="6" pageNumber="13">
<emphasis id="DEF5EAFE6375670CFF74FE6F52D5FE7C" box="[199,252,403,428]" italics="true" pageId="6" pageNumber="13">Legs</emphasis>
. Claw of each leg smooth on dorsal side, with tooth proximoventrally. Formulas of leg setation and solenidia: leg I (0–5–3–5–17) [2–1–2], leg II (0–5–3–5–13) [1–1–2], leg III (2–4–2–3–10) [1–1–0], leg IV (2–3–2– 3–10) [1–0–0]; homology of setae and solenidia indicated in 
<tableCitation id="A10303576375670CFCFAFE2751B7FE25" box="[841,926,475,501]" captionStart="TABLE 1" captionStartId="9.[151,239,151,175]" captionTargetPageId="9" captionText="TABLE 1. Development of leg setation of Lepidacarus maafushiensis sp. nov." httpUri="http://table.plazi.org/id/B8FE6664637A6703FF24FF6B51C8FF7F" pageId="6" pageNumber="13" tableUuid="B8FE6664637A6703FF24FF6B51C8FF7F">Table 1</tableCitation>
. Solenidion ω 
<subScript id="700534A96375670CFB8CFE11566EFE2B" attach="left" box="[1087,1095,493,507]" fontSize="6" pageId="6" pageNumber="13">1</subScript>
on tarsus I, ω 
<subScript id="700534A96375670CFB50FE1156C2FE2B" attach="left" box="[1251,1259,493,507]" fontSize="6" pageId="6" pageNumber="13">1</subScript>
, ω 
<subScript id="700534A96375670CFABEFE11573CFE2B" attach="left" box="[1293,1301,493,507]" fontSize="6" pageId="6" pageNumber="13">2</subScript>
on tarsus II and φ on tibia III bacilliform; other solenidia setiform. Solenidion ω 
<subScript id="700534A96375670CFC25FDED51B7FDCF" attach="left" box="[918,926,529,543]" fontSize="6" pageId="6" pageNumber="13">2</subScript>
and seta 
<emphasis id="DEF5EAFE6375670CFBBBFE03560EFDC9" box="[1032,1063,511,537]" italics="true" pageId="6" pageNumber="13">ft”</emphasis>
on tarsus I inserted close to each other. Famulus tubercle-like, inserted anterolateral to solenidion ω 
<subScript id="700534A96375670CFC22FDC951B0FD93" attach="left" box="[913,921,565,579]" fontSize="6" pageId="6" pageNumber="13">1</subScript>
. Some leg setae widely phylliform, rounded distally, ciliate.
</paragraph>
<paragraph id="EC3E36EC6375670CFF24FD4E5375FD1C" blockId="6.[151,348,690,753]" box="[151,348,690,716]" pageId="6" pageNumber="13">
<heading id="B77681806375670CFF24FD4E5375FD1C" bold="true" box="[151,348,690,716]" fontSize="11" level="1" pageId="6" pageNumber="13" reason="1">
<emphasis id="DEF5EAFE6375670CFF24FD4E5375FD1C" bold="true" box="[151,348,690,716]" pageId="6" pageNumber="13">Juvenile instars</emphasis>
</heading>
</paragraph>
<paragraph id="EC3E36EC6375670CFF24FD2B531CFD21" blockId="6.[151,348,690,753]" box="[151,309,727,753]" pageId="6" pageNumber="13">
(
<figureCitation id="74BA2A696375670CFF2CFD2B52C1FD21" box="[159,232,727,753]" captionStart="FIGURE 2" captionStartId="4.[151,250,1504,1528]" captionTargetBox="[179,1423,234,1438]" captionTargetId="figure-17@4.[151,1436,181,1481]" captionTargetPageId="4" captionText="FIGURE 2. Lepidacarus maafushiensis sp. nov., adult (A, C, E) and larva (B, D, F): A, B—subcapitulum, ventral view; C, D—palp, left, paraxial view; E—chelicera (basal part not shown), left, paraxial view; F—chelicera, left, paraxial view. Scale bar 20 μm (A–C, E, F), 10 μm (D)." figureDoi="http://doi.org/10.5281/zenodo.7078230" httpUri="https://zenodo.org/record/7078230/files/figure.png" pageId="6" pageNumber="13">Figs 2</figureCitation>
, 
<figureCitation id="74BA2A696375670CFF47FD2B5305FD21" box="[244,300,727,753]" captionStart-0="FIGURE 6" captionStart-1="FIGURE 7" captionStart-2="FIGURE 8" captionStart-3="FIGURE 9" captionStart-4="FIGURE 10" captionStart-5="FIGURE 11" captionStart-6="FIGURE 12" captionStartId-0="10.[151,250,1965,1989]" captionStartId-1="11.[151,250,1436,1460]" captionStartId-2="12.[151,250,1028,1052]" captionStartId-3="12.[151,250,1970,1994]" captionStartId-4="13.[151,250,1728,1752]" captionStartId-5="14.[151,250,1643,1667]" captionStartId-6="15.[151,250,1869,1893]" captionTargetBox-0="[207,1368,631,1889]" captionTargetBox-1="[175,1422,201,1381]" captionTargetBox-2="[350,1247,206,982]" captionTargetBox-3="[294,1309,1157,1931]" captionTargetBox-4="[160,1430,181,1676]" captionTargetBox-5="[151,1436,181,1615]" captionTargetBox-6="[151,1436,181,1845]" captionTargetId-0="figure-267@10.[151,1436,597,1942]" captionTargetId-1="figure-17@11.[151,1436,181,1412]" captionTargetId-2="figure-17@12.[325,1262,181,1004]" captionTargetId-3="figure-41@12.[273,1314,1112,1946]" captionTargetId-4="figure-17@13.[151,1436,181,1704]" captionTargetId-5="figure-17@14.[151,1436,181,1619]" captionTargetId-6="figure-17@15.[151,1436,181,1845]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionTargetPageId-2="12" captionTargetPageId-3="12" captionTargetPageId-4="13" captionTargetPageId-5="14" captionTargetPageId-6="15" captionText-0="FIGURE 6. Lepidacarus maafushiensis sp. nov., larva: A—dorsal view; B—ventral view (gnathosoma and legs except trochanters not shown); C—part of prodorsum and notogaster, lateral view. Scale bar 50 μm." captionText-1="FIGURE 7. Lepidacarus maafushiensis sp. nov., dorsal view of juvenile instars: A –protonymph; B—tritonymph. Scale bar 50 μm (A), 100 μm (B)." captionText-2="FIGURE 8. Lepidacarus maafushiensis sp. nov., epimeral region of juvenile instars: A –protonymph; B—deutonymph; C— tritonymph. Scale bar 50 μm." captionText-3="FIGURE 9. Lepidacarus maafushiensis sp. nov., anogenital region of juvenile instars: A –protonymph; B—deutonymph; C— tritonymph. Scale bar 50 μm." captionText-4="FIGURE 10. Lepidacarus maafushiensis sp. nov., larva (A–C) and protonymph (D):A—leg I, without trochanter, left, paraxial view; B—leg II, without trochanter, left, paraxial view; C—leg III, without trochanter, left, antiaxial view; D—leg IV, without trochanter, right, antiaxial view. Scale bar 20 μm." captionText-5="FIGURE 11. Lepidacarus maafushiensis sp. nov., tritonymph, SEM micrographs: A—dorsal view; B—part of gastronotic region, dorsal view; C—ventral view; D—anterodorsal view; E—part of prodorsum, dorsal view." captionText-6="FIGURE 12. Lepidacarus maafushiensis sp. nov., deuto- (A–D; F) and tritonymph (E), microscope images: A—part of prodorsum, lateral view; B—part of ventral side, ventral view; C—subcapitulum and palp, ventral view; D—part of epimeral region, ventral view; E—genital plate, right, ventral view; F—leg III, right, antiaxial view." figureDoi-0="http://doi.org/10.5281/zenodo.7078242" figureDoi-1="http://doi.org/10.5281/zenodo.7078244" figureDoi-2="http://doi.org/10.5281/zenodo.7078246" figureDoi-3="http://doi.org/10.5281/zenodo.7078248" figureDoi-4="http://doi.org/10.5281/zenodo.7078250" figureDoi-5="http://doi.org/10.5281/zenodo.7078252" figureDoi-6="http://doi.org/10.5281/zenodo.7078254" httpUri-0="https://zenodo.org/record/7078242/files/figure.png" httpUri-1="https://zenodo.org/record/7078244/files/figure.png" httpUri-2="https://zenodo.org/record/7078246/files/figure.png" httpUri-3="https://zenodo.org/record/7078248/files/figure.png" httpUri-4="https://zenodo.org/record/7078250/files/figure.png" httpUri-5="https://zenodo.org/record/7078252/files/figure.png" httpUri-6="https://zenodo.org/record/7078254/files/figure.png" pageId="6" pageNumber="13">6–12</figureCitation>
)
</paragraph>
<paragraph id="EC3E36EC6375670CFF24FCE25360FC75" blockId="6.[151,1437,798,2013]" pageId="6" pageNumber="13">
<emphasis id="DEF5EAFE6375670CFF24FCE25300FCE8" bold="true" box="[151,297,798,824]" pageId="6" pageNumber="13">Description.</emphasis>
<emphasis id="DEF5EAFE6375670CFE9FFCE353E7FCE8" box="[300,462,799,824]" italics="true" pageId="6" pageNumber="13">Measurements</emphasis>
. Body length:larva:265–273(
<specimenCount id="FA87FD656375670CFCA8FCE351F8FCE9" box="[795,977,799,825]" pageId="6" pageNumber="13" type="generic">three specimens</specimenCount>
), protonymph:315–365(
<specimenCount id="FA87FD656375670CFB51FCE357B9FCE9" box="[1250,1424,799,825]" pageId="6" pageNumber="13" type="generic">four specimens</specimenCount>
); deutonymph: 415–431 (
<specimenCount id="FA87FD656375670CFE1BFCBF506FFC8D" box="[424,582,835,861]" pageId="6" pageNumber="13" type="generic">11 specimens</specimenCount>
), tritonymph: 464–514 (
<specimenCount id="FA87FD656375670CFCEFFCBF51D5FC8D" box="[860,1020,835,861]" pageId="6" pageNumber="13" type="generic">14 specimens</specimenCount>
). Body width: larva: 149–166 (
<specimenCount id="FA87FD656375670CFAD1FCBF5339FC51" pageId="6" pageNumber="13" type="generic">three specimens</specimenCount>
), protonymph: 182–199 (
<specimenCount id="FA87FD656375670CFD9FFC9B50F6FC51" box="[556,735,871,897]" pageId="6" pageNumber="13" type="generic">four specimens</specimenCount>
); deutonymph: 215–232 (
<specimenCount id="FA87FD656375670CFC4CFC9B56B2FC51" box="[1023,1179,871,897]" pageId="6" pageNumber="13" type="generic">11 specimens</specimenCount>
), tritonymph: 249–282 (
<specimenCount id="FA87FD656375670CFF13FC775317FC75" box="[160,318,907,933]" pageId="6" pageNumber="13" type="generic">14 specimens</specimenCount>
).
</paragraph>
<paragraph id="EC3E36EC6375670CFF74FC535044FBE5" blockId="6.[151,1437,798,2013]" pageId="6" pageNumber="13">
<emphasis id="DEF5EAFE6375670CFF74FC53536CFC18" box="[199,325,943,968]" italics="true" pageId="6" pageNumber="13">Integument</emphasis>
. General body cuticle colourless, rarely slightly yellowish in some nymphs. Surface of body and legs densely microreticulate (micropolygonal ornamentation). Also, prodorsum and gastronotum sparsely microtuberculate in larva, proto- and deutonymph versus tubercles usually large in tritonymph. Macroreticulate (macropolygonal) pattern on body absent.
</paragraph>
<paragraph id="EC3E36EC6375670CFF74FBC35362FA85" blockId="6.[151,1437,798,2013]" pageId="6" pageNumber="13">
<emphasis id="DEF5EAFE6375670CFF74FBC3536DFB89" box="[199,324,1087,1113]" italics="true" pageId="6" pageNumber="13">Prodorsum</emphasis>
. Roughly triangular in dorsal view, occupying about 2/3 of dorsal length. Rostrum broadly concave, with median triangular tooth. Lateral side of rostrum with some slight barbs and partially undulate. Rostral, lamellar, interlamellar, anterior exobothridial, and posterior exobothridial setae widely phylliform, rounded distally, heavily barbed. Bothridial seta (
<emphasis id="DEF5EAFE6375670CFE02FB5753E6FB14" box="[433,463,1195,1220]" italics="true" pageId="6" pageNumber="13">La</emphasis>
: 49–57; 
<emphasis id="DEF5EAFE6375670CFD89FB575070FB14" box="[570,601,1195,1220]" italics="true" pageId="6" pageNumber="13">Pn</emphasis>
: 53–61; 
<emphasis id="DEF5EAFE6375670CFD77FB5750CFFB14" box="[708,742,1195,1220]" italics="true" pageId="6" pageNumber="13">Dn</emphasis>
: 61–69; 
<emphasis id="DEF5EAFE6375670CFCE2FB575146FB14" box="[849,879,1195,1220]" italics="true" pageId="6" pageNumber="13">Tn</emphasis>
: 69–77) with short stalk and narrowly elongate fusiform head having sparsely ciliate one side and densely serrate opposite side. Lateral tubercle of prodorsum (close and anterolateral to anterior exobothridial seta) well developed. Region between lamellar and interlamellar setae with some depressions. Postbothridial band distinct in larva and protonymph versus slightly visible in deuto- and tritonymph.
</paragraph>
<paragraph id="EC3E36EC6375670CFF74FAA35180F9FD" blockId="6.[151,1437,798,2013]" pageId="6" pageNumber="13">
<emphasis id="DEF5EAFE6375670CFF74FAA35388FAA8" box="[199,417,1375,1401]" italics="true" pageId="6" pageNumber="13">Gastronotic region</emphasis>
. With numerous oval depressions in medioposterior part (poorly visible in larva and protonymph versus clearer visible in deuto- and tritonymph). Larva with 16 pairs of gastronotic setae including three pairs of 
<emphasis id="DEF5EAFE6375670CFE81FA5B5369FA10" box="[306,320,1447,1472]" italics="true" pageId="6" pageNumber="13">p</emphasis>
<subScript id="700534A96375670CFEF3FA455361FA17" attach="left" box="[320,328,1465,1479]" fontSize="6" pageId="6" pageNumber="13">1</subScript>
– 
<emphasis id="DEF5EAFE6375670CFEE5FA5B534DFA10" box="[342,356,1447,1472]" italics="true" pageId="6" pageNumber="13">p</emphasis>
<subScript id="700534A96375670CFED7FA455345FA17" attach="left" box="[356,364,1465,1479]" fontSize="6" pageId="6" pageNumber="13">3</subScript>
on pseudoanal plates; seta 
<emphasis id="DEF5EAFE6375670CFD2EFA5B5082FA11" box="[669,683,1447,1473]" italics="true" pageId="6" pageNumber="13">h</emphasis>
<subScript id="700534A96375670CFD18FA45509AFA17" attach="left" box="[683,691,1465,1479]" fontSize="6" pageId="6" pageNumber="13">4</subScript>
not developed. Protonymph with 17 pairs (added 
<emphasis id="DEF5EAFE6375670CFB50FA5B56D8FA10" box="[1251,1265,1447,1472]" italics="true" pageId="6" pageNumber="13">n</emphasis>
<subScript id="700534A96375670CFB42FA4556D0FA17" attach="left" box="[1265,1273,1465,1479]" fontSize="6" pageId="6" pageNumber="13">1</subScript>
), deutonymph with 18 pairs (added 
<emphasis id="DEF5EAFE6375670CFE22FA3753B6FA34" box="[401,415,1483,1508]" italics="true" pageId="6" pageNumber="13">n</emphasis>
<subScript id="700534A96375670CFE2CFA21538EFA3B" attach="left" box="[415,423,1501,1515]" fontSize="6" pageId="6" pageNumber="13">2</subScript>
), tritonymph with 18 pairs of gastronotic setae. All gastronotic setae—except setiform, smooth setae 
<emphasis id="DEF5EAFE6375670CFE85FA13536DF9D9" box="[310,324,1519,1545]" italics="true" pageId="6" pageNumber="13">h</emphasis>
<subScript id="700534A96375670CFEF7F9FD5365F9DF" attach="left" box="[324,332,1537,1551]" fontSize="6" pageId="6" pageNumber="13">3</subScript>
(12–14) and 
<emphasis id="DEF5EAFE6375670CFE5EFA1353D2F9D8" box="[493,507,1519,1544]" italics="true" pageId="6" pageNumber="13">p</emphasis>
<subScript id="700534A96375670CFE48F9FD502AF9DF" attach="left" box="[507,515,1537,1551]" fontSize="6" pageId="6" pageNumber="13">1</subScript>
– 
<emphasis id="DEF5EAFE6375670CFDA2FA135036F9D8" box="[529,543,1519,1544]" italics="true" pageId="6" pageNumber="13">p</emphasis>
<subScript id="700534A96375670CFDACF9FD500EF9DF" attach="left" box="[543,551,1537,1551]" fontSize="6" pageId="6" pageNumber="13">3</subScript>
(8–10) in larva—widely phylliform, rounded distally, heavily barbed. Some transverse bands present dorsally, usually slightly visible or indistinct.
</paragraph>
<paragraph id="EC3E36EC6375670CFF74F9CB576CF949" blockId="6.[151,1437,798,2013]" pageId="6" pageNumber="13">
<emphasis id="DEF5EAFE6375670CFF74F9CB537FF981" box="[199,342,1591,1617]" italics="true" pageId="6" pageNumber="13">Gnathosoma</emphasis>
. Generally, similar in adult. Larva with three pairs of subcapitular setae (
<emphasis id="DEF5EAFE6375670CFB10F9CB5698F980" box="[1187,1201,1591,1616]" italics="true" pageId="6" pageNumber="13">a</emphasis>
, 
<emphasis id="DEF5EAFE6375670CFB72F9CB56FCF980" box="[1217,1237,1591,1616]" italics="true" pageId="6" pageNumber="13">m</emphasis>
<subScript id="700534A96375670CFB66F9B556F4F987" attach="left" box="[1237,1245,1609,1623]" fontSize="6" pageId="6" pageNumber="13">1</subScript>
, 
<emphasis id="DEF5EAFE6375670CFB5EF9CB56D2F981" box="[1261,1275,1591,1617]" italics="true" pageId="6" pageNumber="13">h</emphasis>
); 
<emphasis id="DEF5EAFE6375670CFAA6F9CB5700F980" box="[1301,1321,1591,1616]" italics="true" pageId="6" pageNumber="13">m</emphasis>
<subScript id="700534A96375670CFA9AF9B55718F987" attach="left" box="[1321,1329,1609,1623]" fontSize="6" pageId="6" pageNumber="13">2</subScript>
added in protonymph, 
<emphasis id="DEF5EAFE6375670CFE82F9A7536CF9A4" box="[305,325,1627,1652]" italics="true" pageId="6" pageNumber="13">m</emphasis>
<quantity id="2B799B096375670CFEF6F9915344F9A5" box="[325,365,1627,1659]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="6" pageNumber="13" unit="in" value="3.0">
<subScript id="700534A96375670CFEF6F9915364F9AB" attach="left" box="[325,333,1645,1659]" fontSize="6" pageId="6" pageNumber="13">3</subScript>
in
</quantity>
deutonymph, 
<emphasis id="DEF5EAFE6375670CFDA7F9A75001F9A4" box="[532,552,1627,1652]" italics="true" pageId="6" pageNumber="13">m</emphasis>
<quantity id="2B799B096375670CFD9AF9915078F9A5" box="[553,593,1627,1659]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="6" pageNumber="13" unit="in" value="4.0">
<subScript id="700534A96375670CFD9AF9915018F9AB" attach="left" box="[553,561,1645,1659]" fontSize="6" pageId="6" pageNumber="13">4</subScript>
in
</quantity>
tritonymph. Larva with two pairs of adoral setae (
<emphasis id="DEF5EAFE6375670CFB13F9A75690F9A4" box="[1184,1209,1627,1652]" italics="true" pageId="6" pageNumber="13">or</emphasis>
<subScript id="700534A96375670CFB0AF99156E8F9AB" attach="left" box="[1209,1217,1645,1659]" fontSize="6" pageId="6" pageNumber="13">1</subScript>
, 
<emphasis id="DEF5EAFE6375670CFB61F9A756C2F9A4" box="[1234,1259,1627,1652]" italics="true" pageId="6" pageNumber="13">or</emphasis>
<subScript id="700534A96375670CFB58F99156DAF9AB" attach="left" box="[1259,1267,1645,1659]" fontSize="6" pageId="6" pageNumber="13">2</subScript>
); 
<emphasis id="DEF5EAFE6375670CFABDF9A7570EF9A4" box="[1294,1319,1627,1652]" italics="true" pageId="6" pageNumber="13">or</emphasis>
<subScript id="700534A96375670CFA94F9915706F9AB" attach="left" box="[1319,1327,1645,1659]" fontSize="6" pageId="6" pageNumber="13">3</subScript>
added in protonymph. Palp setation of larva and protonymph: 0–0–0–2–9(+ω); seta on femur added in deutonymph.
</paragraph>
<paragraph id="EC3E36EC6375670CFF74F95F503CF8D5" blockId="6.[151,1437,798,2013]" pageId="6" pageNumber="13">
<emphasis id="DEF5EAFE6375670CFF74F95F53ABF96C" box="[199,386,1699,1725]" italics="true" pageId="6" pageNumber="13">Epimeral region</emphasis>
. Setal formulas for epimeres: larva: 3–1–2; protonymph: 4–2–2–1; deutonymph: 6–4–3–3, tritonymph: 8–5–3–4. Seta 
<emphasis id="DEF5EAFE6375670CFE67F93B53C7F931" box="[468,494,1735,1761]" italics="true" pageId="6" pageNumber="13">1c</emphasis>
in larva setiform, shortly ciliate; other setae widely phylliform, rounded distally, ciliate. Claparède’s organ distinct.
</paragraph>
<paragraph id="EC3E36EC6375670DFF74F8F356CFFECD" blockId="6.[151,1437,798,2013]" lastBlockId="7.[151,1437,151,285]" lastPageId="7" lastPageNumber="14" pageId="6" pageNumber="13">
<emphasis id="DEF5EAFE6375670CFF74F8F353B8F8F8" box="[199,401,1807,1833]" italics="true" pageId="6" pageNumber="13">Anogenital region</emphasis>
. Genital valves divided in deuto- and tritonymph (undivided in protonymph). Larva without genital, anal and adanal setae; protonymph without anal setae, but with one pair of genital (18–20; setiform, shortly ciliate) and four pairs of adanal (8–10, setiform, smooth) setae; deutonymph with five pairs of genital (four pairs of medial setae: 12, setiform, shortly ciliate; one pair of lateral setae: widely phylliform, rounded distally, ciliate), two pairs of anal (12, setiform, smooth) and four pairs of adanal (widely phylliform, rounded distally, heavily barbed) setae; tritonymph with eight pairs of genital (three pairs of medial setae: 12, setiform, shortly ciliate; four pairs of mediolateral setae: 20, setiform, shortly ciliate; one pair of lateral setae: widely phylliform, rounded distally, ciliate), two pairs of anal (20–24, setiform, shortly ciliate) and four pairs of adanal (widely phylliform, rounded distally, heavily barbed) setae. Pseudoadanal (in protonymph) and pseudoanal (in deutonymph) plates with paraproctal setae. Cupules 
<emphasis id="DEF5EAFE6374670DFF49FEFF5339FECD" box="[250,272,259,285]" italics="true" pageId="7" pageNumber="14">ih</emphasis>
and 
<emphasis id="DEF5EAFE6374670DFEF5FEFF534EFECC" box="[326,359,259,284]" italics="true" pageId="7" pageNumber="14">ips</emphasis>
appearing in normal ontogenetic pattern; adanal and anal cupules not observed.
</paragraph>
<caption id="B8FE66646374670DFF24F90B5395F8E3" ID-DOI="http://doi.org/10.5281/zenodo.7078234" ID-Zenodo-Dep="7078234" httpUri="https://zenodo.org/record/7078234/files/figure.png" pageId="7" pageNumber="14" startId="7.[151,250,1783,1807]" targetBox="[151,1436,309,1744]" targetPageId="7">
<paragraph id="EC3E36EC6374670DFF24F90B5395F8E3" blockId="7.[151,1437,1783,1843]" pageId="7" pageNumber="14">
<emphasis id="DEF5EAFE6374670DFF24F90B533BF8DF" bold="true" box="[151,274,1783,1807]" pageId="7" pageNumber="14">FIGURE 4.</emphasis>
<taxonomicName id="2B814D6F6374670DFEABF90B5005F8DF" authority="Bayartogtokh &amp; Ermilov &amp; Joharchi, 2022" authorityName="Bayartogtokh &amp; Ermilov &amp; Joharchi" authorityYear="2022" box="[280,556,1783,1807]" class="Arachnida" family="Lohmanniidae" genus="Lepidacarus" kingdom="Animalia" order="Sarcoptiformes" pageId="7" pageNumber="14" phylum="Arthropoda" rank="species" species="maafushiensis" status="sp. nov.">
<emphasis id="DEF5EAFE6374670DFEABF90B5005F8DF" box="[280,556,1783,1807]" italics="true" pageId="7" pageNumber="14">Lepidacarus maafushiensis</emphasis>
</taxonomicName>
<emphasis id="DEF5EAFE6374670DFD81F90B50ABF8DF" bold="true" box="[562,642,1783,1807]" pageId="7" pageNumber="14">
<taxonomicNameLabel id="C5C657856374670DFD81F90B50ABF8DF" box="[562,642,1783,1807]" pageId="7" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, adult, SEM micrographs: A—dorsal view; B—ventral view; C—ventrolateral view; D—anterodorsal view.
</paragraph>
</caption>
<caption id="B8FE6664637B6702FF24F99B51ABF973" ID-DOI="http://doi.org/10.5281/zenodo.7078236" ID-Zenodo-Dep="7078236" httpUri="https://zenodo.org/record/7078236/files/figure.png" pageId="8" pageNumber="15" startId="8.[151,250,1639,1663]" targetBox="[151,1436,181,1615]" targetPageId="8">
<paragraph id="EC3E36EC637B6702FF24F99B51ABF973" blockId="8.[151,1436,1639,1699]" pageId="8" pageNumber="15">
<emphasis id="DEF5EAFE637B6702FF24F99B533DF9AF" bold="true" box="[151,276,1639,1663]" pageId="8" pageNumber="15">FIGURE 5.</emphasis>
<taxonomicName id="2B814D6F637B6702FEA8F99B5018F9AF" authority="Bayartogtokh &amp; Ermilov &amp; Joharchi, 2022" authorityName="Bayartogtokh &amp; Ermilov &amp; Joharchi" authorityYear="2022" box="[283,561,1639,1663]" class="Arachnida" family="Lohmanniidae" genus="Lepidacarus" kingdom="Animalia" order="Sarcoptiformes" pageId="8" pageNumber="15" phylum="Arthropoda" rank="species" species="maafushiensis" status="sp. nov.">
<emphasis id="DEF5EAFE637B6702FEA8F99B5018F9AF" box="[283,561,1639,1663]" italics="true" pageId="8" pageNumber="15">Lepidacarus maafushiensis</emphasis>
</taxonomicName>
<emphasis id="DEF5EAFE637B6702FD8BF99B50A0F9AF" bold="true" box="[568,649,1639,1663]" pageId="8" pageNumber="15">
<taxonomicNameLabel id="C5C65785637B6702FD8BF99B50A0F9AF" box="[568,649,1639,1663]" pageId="8" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, adult, SEM micrographs: A—posterior view; B—gnathosoma, lateral view; C—genital region, ventrolateral view; D—anoadanal region, lateral view.
</paragraph>
</caption>
<paragraph id="EC3E36EC637B6702FF74F90A505DF813" blockId="8.[151,1437,1781,1988]" pageId="8" pageNumber="15">
<emphasis id="DEF5EAFE637B6702FF74F90A52D5F8DF" box="[199,252,1782,1807]" italics="true" pageId="8" pageNumber="15">Legs</emphasis>
. Claw of each leg smooth on dorsal side, with tooth proximoventrally. Formulas of leg setation and solenidia: larva: I (0–2–3–4–15) [2–1–1], II (0–2–3–3–13) [1–1–1], III (0–2–2–3–10) [1–1–0]; protonymph: I (0– 2–3–4–16) [2–1–2], II (0–3–3–3–13) [1–1–1], III (1–3–2–3–10) [1–1–0], IV (0–0–0–0–6) [0–0–0]; deutonymph: I (0–5–3–4–16) [2–1–2], II (0–5–3–3–13) [1–1–2], III (2–3–2–3–10) [1–1–0], IV (1–2–2–2–10) [1–0–0]; tritonymph: I (0–5–3–5–17) [2–1–2], II (0–5–3–5–13) [1–1–2], III (2–4–2–3–10) [1–1–0], IV (2–3–2–3–10) [1–0–0]; homology of setae and solenidia indicated in 
<tableCitation id="A1030357637B6702FDA9F8555047F813" box="[538,622,1961,1987]" captionStart="TABLE 1" captionStartId="9.[151,239,151,175]" captionTargetPageId="9" captionText="TABLE 1. Development of leg setation of Lepidacarus maafushiensis sp. nov." httpUri="http://table.plazi.org/id/B8FE6664637A6703FF24FF6B51C8FF7F" pageId="8" pageNumber="15" tableUuid="B8FE6664637A6703FF24FF6B51C8FF7F">Table 1</tableCitation>
.
</paragraph>
</subSubSection>
<caption id="B8FE6664637A6703FF24FF6B51C8FF7F" ID-Table-UUID="B8FE6664637A6703FF24FF6B51C8FF7F" box="[151,993,151,175]" httpUri="http://table.plazi.org/id/B8FE6664637A6703FF24FF6B51C8FF7F" pageId="9" pageNumber="16" startId="9.[151,239,151,175]" targetBox="[163,1293,192,1141]" targetIsTable="true" targetPageId="9">
<paragraph id="EC3E36EC637A6703FF24FF6B51C8FF7F" blockId="9.[151,993,151,175]" box="[151,993,151,175]" pageId="9" pageNumber="16">
<emphasis id="DEF5EAFE637A6703FF24FF6B532AFF7F" bold="true" box="[151,259,151,175]" pageId="9" pageNumber="16">TABLE 1</emphasis>
. Development of leg setation of 
<taxonomicName id="2B814D6F637A6703FDD1FF6B51ACFF7F" authority="Bayartogtokh &amp; Ermilov &amp; Joharchi, 2022" authorityName="Bayartogtokh &amp; Ermilov &amp; Joharchi" authorityYear="2022" box="[610,901,151,175]" class="Arachnida" family="Lohmanniidae" genus="Lepidacarus" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="16" phylum="Arthropoda" rank="species" species="maafushiensis" status="sp. nov.">
<emphasis id="DEF5EAFE637A6703FDD1FF6B51ACFF7F" box="[610,901,151,175]" italics="true" pageId="9" pageNumber="16">Lepidacarus maafushiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C5C65785637A6703FC3FFF6B51C8FF7F" box="[908,993,151,175]" pageId="9" pageNumber="16" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="EC3E36EC637A6703FF10FF3C518FFBA5" pageId="9" pageNumber="16">
<table id="9E81C44C637A98F5FF10FF3C5724FBA5" box="[163,1293,192,1141]" gridcols="6" gridrows="24" pageId="9" pageNumber="16">
<tr id="52B134AE637A98F5FF10FF3C5724FF07" box="[163,1293,192,215]" gridrow="0" pageId="9" pageNumber="16">
<th id="11605DD2637A98F5FF10FF3C52F4FF07" box="[163,221,192,215]" gridcol="0" gridrow="0" pageId="9" pageNumber="16">Instar</th>
<th id="11605DD2637A98F5FE81FF3C5360FF07" box="[306,329,192,215]" gridcol="1" gridrow="0" pageId="9" pageNumber="16">
<emphasis id="DEF5EAFE637A6703FE81FF3C5360FF07" box="[306,329,192,215]" italics="true" pageId="9" pageNumber="16">Tr</emphasis>
</th>
<th id="11605DD2637A98F5FE37FF3C53EAFF07" box="[388,451,192,215]" gridcol="2" gridrow="0" pageId="9" pageNumber="16">
<emphasis id="DEF5EAFE637A6703FE37FF3C53B6FF07" box="[388,415,192,215]" italics="true" pageId="9" pageNumber="16">Fe</emphasis>
</th>
<th id="11605DD2637A98F5FDBEFF3C50AEFF07" box="[525,647,192,215]" gridcol="3" gridrow="0" pageId="9" pageNumber="16">
<emphasis id="DEF5EAFE637A6703FDBEFF3C5002FF07" box="[525,555,192,215]" italics="true" pageId="9" pageNumber="16">Ge</emphasis>
</th>
<th id="11605DD2637A98F5FD6DFF3C5147FF07" box="[734,878,192,215]" gridcol="4" gridrow="0" pageId="9" pageNumber="16">
<emphasis id="DEF5EAFE637A6703FD6DFF3C50DBFF07" box="[734,754,192,215]" italics="true" pageId="9" pageNumber="16">Ti</emphasis>
</th>
<th id="11605DD2637A98F5FC2AFF3C5724FF07" box="[921,1293,192,215]" gridcol="5" gridrow="0" pageId="9" pageNumber="16">
<emphasis id="DEF5EAFE637A6703FC2AFF3C5198FF07" box="[921,945,192,215]" italics="true" pageId="9" pageNumber="16">Ta</emphasis>
</th>
</tr>
<tr id="52B134AE637A98F5FF10FF145724FED0" box="[163,1293,232,256]" gridrow="1" pageId="9" pageNumber="16">
<th id="11605DD2637A98F5FF10FF145724FED0" box="[163,1293,232,256]" colspan="6" colspanRight="5" gridcol="0" gridrow="1" pageId="9" pageNumber="16">Leg I</th>
</tr>
<tr id="52B134AE637A98F5FF10FEEC5724FEFE" box="[163,1293,272,302]" gridrow="2" pageId="9" pageNumber="16">
<th id="11605DD2637A98F5FF10FEEC52F4FEFE" box="[163,221,272,302]" gridcol="0" gridrow="2" pageId="9" pageNumber="16">
<emphasis id="DEF5EAFE637A6703FF10FEED5297FEF8" box="[163,190,273,296]" italics="true" pageId="9" pageNumber="16">La</emphasis>
</th>
<td id="11605DD2637A98F5FE81FEEC5360FEFE" box="[306,329,272,302]" gridcol="1" gridrow="2" pageId="9" pageNumber="16">
<emphasis id="DEF5EAFE637A6703FE81FEEC5316FEF7" box="[306,319,272,295]" italics="true" pageId="9" pageNumber="16">–</emphasis>
</td>
<td id="11605DD2637A98F5FE37FEEC53EAFEFE" box="[388,451,272,302]" gridcol="2" gridrow="2" pageId="9" pageNumber="16">
<emphasis id="DEF5EAFE637A6703FE37FEEC53B8FEF8" box="[388,401,272,296]" italics="true" pageId="9" pageNumber="16">d</emphasis>
, 
<emphasis id="DEF5EAFE637A6703FE2EFEEC53EAFEF8" box="[413,451,272,296]" italics="true" pageId="9" pageNumber="16">bv”</emphasis>
</td>
<td id="11605DD2637A98F5FDBEFEEC50AEFEFE" box="[525,647,272,302]" gridcol="3" gridrow="2" pageId="9" pageNumber="16">
<emphasis id="DEF5EAFE637A6703FDBEFEEC5033FEF8" box="[525,538,272,296]" italics="true" pageId="9" pageNumber="16">d</emphasis>
, 
<emphasis id="DEF5EAFE637A6703FD94FEEC5017FEF8" box="[551,574,272,296]" italics="true" pageId="9" pageNumber="16">(l)</emphasis>
, σ’, σ”
</td>
<td id="11605DD2637A98F5FD6DFEEC5147FEFE" box="[734,878,272,302]" gridcol="4" gridrow="2" pageId="9" pageNumber="16">
<emphasis id="DEF5EAFE637A6703FD6DFEEC50C2FEF8" box="[734,747,272,296]" italics="true" pageId="9" pageNumber="16">d</emphasis>
, 
<emphasis id="DEF5EAFE637A6703FD44FEEC511BFEF8" box="[759,818,272,296]" italics="true" pageId="9" pageNumber="16">xt1, l’</emphasis>
, 
<emphasis id="DEF5EAFE637A6703FC8CFEEC517AFEF7" box="[831,851,272,295]" italics="true" pageId="9" pageNumber="16">v’</emphasis>
, φ
</td>
<td id="11605DD2637A98F5FC2AFEEC5724FEFE" box="[921,1293,272,302]" gridcol="5" gridrow="2" pageId="9" pageNumber="16">
<emphasis id="DEF5EAFE637A6703FC2AFEEC519EFEF8" box="[921,951,272,296]" italics="true" pageId="9" pageNumber="16">(ft)</emphasis>
, 
<emphasis id="DEF5EAFE637A6703FC77FEEC51CEFEF7" box="[964,999,272,295]" italics="true" pageId="9" pageNumber="16">(tc)</emphasis>
, 
<emphasis id="DEF5EAFE637A6703FC47FEEC5638FEF7" box="[1012,1041,272,295]" italics="true" pageId="9" pageNumber="16">(p)</emphasis>
, 
<emphasis id="DEF5EAFE637A6703FBADFEEC5615FEF7" box="[1054,1084,272,295]" italics="true" pageId="9" pageNumber="16">(u)</emphasis>
, 
<emphasis id="DEF5EAFE637A6703FBFBFEEC564FFEF7" box="[1096,1126,272,295]" italics="true" pageId="9" pageNumber="16">(a)</emphasis>
, 
<emphasis id="DEF5EAFE637A6703FBC0FEED5654FEF8" box="[1139,1149,273,296]" italics="true" pageId="9" pageNumber="16">s</emphasis>
, 
<emphasis id="DEF5EAFE637A6703FB3AFEED56B2FEF8" box="[1161,1179,273,296]" italics="true" pageId="9" pageNumber="16">m</emphasis>
, 
<emphasis id="DEF5EAFE637A6703FB1BFEEC56F8FEF7" box="[1192,1233,272,295]" italics="true" pageId="9" pageNumber="16">(pv)</emphasis>
, ε, ω 1
</td>
</tr>
<tr id="52B134AE637A98F5FF10FEC55724FE86" box="[163,1293,313,342]" gridrow="3" pageId="9" pageNumber="16">
<th id="11605DD2637A98F5FF10FEC552F4FE86" box="[163,221,313,342]" gridcol="0" gridrow="3" pageId="9" pageNumber="16">
<emphasis id="DEF5EAFE637A6703FF10FEC55296FE80" box="[163,191,313,336]" italics="true" pageId="9" pageNumber="16">Pn</emphasis>
</th>
<td id="11605DD2637A98F5FE81FEC55360FE86" box="[306,329,313,342]" gridcol="1" gridrow="3" pageId="9" pageNumber="16">–</td>
<td id="11605DD2637A98F5FE37FEC553EAFE86" box="[388,451,313,342]" gridcol="2" gridrow="3" pageId="9" pageNumber="16">–</td>
<td id="11605DD2637A98F5FDBEFEC550AEFE86" box="[525,647,313,342]" gridcol="3" gridrow="3" pageId="9" pageNumber="16">–</td>
<td id="11605DD2637A98F5FD6DFEC55147FE86" box="[734,878,313,342]" gridcol="4" gridrow="3" pageId="9" pageNumber="16">–</td>
<td id="11605DD2637A98F5FC2AFEC55724FE86" box="[921,1293,313,342]" gridcol="5" gridrow="3" pageId="9" pageNumber="16">
<emphasis id="DEF5EAFE637A6703FC2AFEC5518FFE80" box="[921,934,313,336]" italics="true" pageId="9" pageNumber="16">n</emphasis>
, ω2
</td>
</tr>
<tr id="52B134AE637A98F5FF10FE9C5724FEA8" box="[163,1293,352,376]" gridrow="4" pageId="9" pageNumber="16">
<th id="11605DD2637A98F5FF10FE9C52F4FEA8" box="[163,221,352,376]" gridcol="0" gridrow="4" pageId="9" pageNumber="16">
<emphasis id="DEF5EAFE637A6703FF10FE9D52EBFEA8" box="[163,194,353,376]" italics="true" pageId="9" pageNumber="16">Dn</emphasis>
</th>
<td id="11605DD2637A98F5FE81FE9C5360FEA8" box="[306,329,352,376]" gridcol="1" gridrow="4" pageId="9" pageNumber="16">
<emphasis id="DEF5EAFE637A6703FE81FE9D5316FEA8" box="[306,319,353,376]" italics="true" pageId="9" pageNumber="16">–</emphasis>
</td>
<td id="11605DD2637A98F5FE37FE9C53EAFEA8" box="[388,451,352,376]" gridcol="2" gridrow="4" pageId="9" pageNumber="16">
<emphasis id="DEF5EAFE637A6703FE37FE9C53B2FEA8" box="[388,411,352,376]" italics="true" pageId="9" pageNumber="16">(l)</emphasis>
, 
<emphasis id="DEF5EAFE637A6703FE1BFE9D53E8FEA8" box="[424,449,353,376]" italics="true" pageId="9" pageNumber="16">v”</emphasis>
</td>
<td id="11605DD2637A98F5FDBEFE9C50AEFEA8" box="[525,647,352,376]" gridcol="3" gridrow="4" pageId="9" pageNumber="16">–</td>
<td id="11605DD2637A98F5FD6DFE9C5147FEA8" box="[734,878,352,376]" gridcol="4" gridrow="4" pageId="9" pageNumber="16">–</td>
<td id="11605DD2637A98F5FC2AFE9C5724FEA8" box="[921,1293,352,376]" gridcol="5" gridrow="4" pageId="9" pageNumber="16">–</td>
</tr>
<tr id="52B134AE637A98F5FF10FE755724FE70" box="[163,1293,393,416]" gridrow="5" pageId="9" pageNumber="16">
<th id="11605DD2637A98F5FF10FE7552F4FE70" box="[163,221,393,416]" gridcol="0" gridrow="5" pageId="9" pageNumber="16">
<emphasis id="DEF5EAFE637A6703FF10FE755297FE70" box="[163,190,393,416]" italics="true" pageId="9" pageNumber="16">Tn</emphasis>
</th>
<td id="11605DD2637A98F5FE81FE755360FE70" box="[306,329,393,416]" gridcol="1" gridrow="5" pageId="9" pageNumber="16">–</td>
<td id="11605DD2637A98F5FE37FE7553EAFE70" box="[388,451,393,416]" gridcol="2" gridrow="5" pageId="9" pageNumber="16">–</td>
<td id="11605DD2637A98F5FDBEFE7550AEFE70" box="[525,647,393,416]" gridcol="3" gridrow="5" pageId="9" pageNumber="16">–</td>
<td id="11605DD2637A98F5FD6DFE755147FE70" box="[734,878,393,416]" gridcol="4" gridrow="5" pageId="9" pageNumber="16">
<emphasis id="DEF5EAFE637A6703FD6DFE7550D7FE70" box="[734,766,393,416]" italics="true" pageId="9" pageNumber="16">xt’’</emphasis>
</td>
<td id="11605DD2637A98F5FC2AFE755724FE70" box="[921,1293,393,416]" gridcol="5" gridrow="5" pageId="9" pageNumber="16">
<emphasis id="DEF5EAFE637A6703FC2AFE75518FFE70" box="[921,934,393,416]" italics="true" pageId="9" pageNumber="16">–</emphasis>
</td>
</tr>
<tr id="52B134AE637A98F5FF10FE4D5724FE19" box="[163,1293,433,457]" gridrow="6" pageId="9" pageNumber="16">
<th id="11605DD2637A98F5FF10FE4D52F4FE19" box="[163,221,433,457]" gridcol="0" gridrow="6" pageId="9" pageNumber="16">
<emphasis id="DEF5EAFE637A6703FF10FE4D5296FE19" box="[163,191,433,457]" italics="true" pageId="9" pageNumber="16">Ad</emphasis>
</th>
<td id="11605DD2637A98F5FE81FE4D5360FE19" box="[306,329,433,457]" gridcol="1" gridrow="6" pageId="9" pageNumber="16">–</td>
<td id="11605DD2637A98F5FE37FE4D53EAFE19" box="[388,451,433,457]" gridcol="2" gridrow="6" pageId="9" pageNumber="16">–</td>
<td id="11605DD2637A98F5FDBEFE4D50AEFE19" box="[525,647,433,457]" gridcol="3" gridrow="6" pageId="9" pageNumber="16">–</td>
<td id="11605DD2637A98F5FD6DFE4D5147FE19" box="[734,878,433,457]" gridcol="4" gridrow="6" pageId="9" pageNumber="16">–</td>
<td id="11605DD2637A98F5FC2AFE4D5724FE19" box="[921,1293,433,457]" gridcol="5" gridrow="6" pageId="9" pageNumber="16">–</td>
</tr>
<tr id="52B134AE637A98F5FF10FE265724FE22" box="[163,1293,474,498]" gridrow="7" pageId="9" pageNumber="16">
<th id="11605DD2637A98F5FF10FE265724FE22" box="[163,1293,474,498]" colspan="6" colspanRight="5" gridcol="0" gridrow="7" pageId="9" pageNumber="16">Leg II</th>
</tr>
<tr id="52B134AE637A98F5FF10FDFD5724FDCF" box="[163,1293,513,543]" gridrow="8" pageId="9" pageNumber="16">
<th id="11605DD2637A98F5FF10FDFD52F4FDCF" box="[163,221,513,543]" gridcol="0" gridrow="8" pageId="9" pageNumber="16">
<emphasis id="DEF5EAFE637A6703FF10FDFE5297FDC9" box="[163,190,514,537]" italics="true" pageId="9" pageNumber="16">La</emphasis>
</th>
<td id="11605DD2637A98F5FE81FDFD5360FDCF" box="[306,329,513,543]" gridcol="1" gridrow="8" pageId="9" pageNumber="16">
<emphasis id="DEF5EAFE637A6703FE81FDFE5316FDC9" box="[306,319,514,537]" italics="true" pageId="9" pageNumber="16">–</emphasis>
</td>
<td id="11605DD2637A98F5FE37FDFD53EAFDCF" box="[388,451,513,543]" gridcol="2" gridrow="8" pageId="9" pageNumber="16">
<emphasis id="DEF5EAFE637A6703FE37FDFD53B8FDC9" box="[388,401,513,537]" italics="true" pageId="9" pageNumber="16">d</emphasis>
, 
<emphasis id="DEF5EAFE637A6703FE2EFDFD53EAFDC9" box="[413,451,513,537]" italics="true" pageId="9" pageNumber="16">bv”</emphasis>
</td>
<td id="11605DD2637A98F5FDBEFDFD50AEFDCF" box="[525,647,513,543]" gridcol="3" gridrow="8" pageId="9" pageNumber="16">
<emphasis id="DEF5EAFE637A6703FDBEFDFD5033FDC9" box="[525,538,513,537]" italics="true" pageId="9" pageNumber="16">d</emphasis>
, 
<emphasis id="DEF5EAFE637A6703FD94FDFD5017FDC9" box="[551,574,513,537]" italics="true" pageId="9" pageNumber="16">(l)</emphasis>
, σ
</td>
<td id="11605DD2637A98F5FD6DFDFD5147FDCF" box="[734,878,513,543]" gridcol="4" gridrow="8" pageId="9" pageNumber="16">
<emphasis id="DEF5EAFE637A6703FD6DFDFD50C2FDC9" box="[734,747,513,537]" italics="true" pageId="9" pageNumber="16">d</emphasis>
, 
<emphasis id="DEF5EAFE637A6703FD44FDFE511BFDC9" box="[759,818,513,538]" italics="true" pageId="9" pageNumber="16">xt1, l’</emphasis>
, φ
</td>
<td id="11605DD2637A98F5FC2AFDFD5724FDCF" box="[921,1293,513,543]" gridcol="5" gridrow="8" pageId="9" pageNumber="16">
<emphasis id="DEF5EAFE637A6703FC2AFDFE519EFDCA" box="[921,951,514,538]" italics="true" pageId="9" pageNumber="16">(ft)</emphasis>
, 
<emphasis id="DEF5EAFE637A6703FC77FDFE51CEFDC9" box="[964,999,514,537]" italics="true" pageId="9" pageNumber="16">(tc)</emphasis>
, 
<emphasis id="DEF5EAFE637A6703FC47FDFE5638FDC9" box="[1012,1041,514,537]" italics="true" pageId="9" pageNumber="16">(p)</emphasis>
, 
<emphasis id="DEF5EAFE637A6703FBADFDFE5615FDC9" box="[1054,1084,514,537]" italics="true" pageId="9" pageNumber="16">(u)</emphasis>
, 
<emphasis id="DEF5EAFE637A6703FBFBFDFE564FFDC9" box="[1096,1126,514,537]" italics="true" pageId="9" pageNumber="16">(a)</emphasis>
, 
<emphasis id="DEF5EAFE637A6703FBC0FDFE5654FDC9" box="[1139,1149,514,537]" italics="true" pageId="9" pageNumber="16">s</emphasis>
, 
<emphasis id="DEF5EAFE637A6703FB3AFDFE569BFDC9" box="[1161,1202,514,537]" italics="true" pageId="9" pageNumber="16">(pv)</emphasis>
, ω1
</td>
</tr>
<tr id="52B134AE637A98F5FF10FDD65724FD92" box="[163,1293,554,578]" gridrow="9" pageId="9" pageNumber="16">
<th id="11605DD2637A98F5FF10FDD652F4FD92" box="[163,221,554,578]" gridcol="0" gridrow="9" pageId="9" pageNumber="16">
<emphasis id="DEF5EAFE637A6703FF10FDD65296FD91" box="[163,191,554,577]" italics="true" pageId="9" pageNumber="16">Pn</emphasis>
</th>
<td id="11605DD2637A98F5FE81FDD65360FD92" box="[306,329,554,578]" gridcol="1" gridrow="9" pageId="9" pageNumber="16">
<emphasis id="DEF5EAFE637A6703FE81FDD65316FD91" box="[306,319,554,577]" italics="true" pageId="9" pageNumber="16">–</emphasis>
</td>
<td id="11605DD2637A98F5FE37FDD653EAFD92" box="[388,451,554,578]" gridcol="2" gridrow="9" pageId="9" pageNumber="16">
<emphasis id="DEF5EAFE637A6703FE37FDD653B0FD92" box="[388,409,554,578]" italics="true" pageId="9" pageNumber="16">l”</emphasis>
</td>
<td id="11605DD2637A98F5FDBEFDD650AEFD92" box="[525,647,554,578]" gridcol="3" gridrow="9" pageId="9" pageNumber="16">–</td>
<td id="11605DD2637A98F5FD6DFDD65147FD92" box="[734,878,554,578]" gridcol="4" gridrow="9" pageId="9" pageNumber="16">–</td>
<td id="11605DD2637A98F5FC2AFDD65724FD92" box="[921,1293,554,578]" gridcol="5" gridrow="9" pageId="9" pageNumber="16">–</td>
</tr>
<tr id="52B134AE637A98F5FF10FDAE5724FDA0" box="[163,1293,594,624]" gridrow="10" pageId="9" pageNumber="16">
<th id="11605DD2637A98F5FF10FDAE52F4FDA0" box="[163,221,594,624]" gridcol="0" gridrow="10" pageId="9" pageNumber="16">
<emphasis id="DEF5EAFE637A6703FF10FDAF52EBFDBA" box="[163,194,595,618]" italics="true" pageId="9" pageNumber="16">Dn</emphasis>
</th>
<td id="11605DD2637A98F5FE81FDAE5360FDA0" box="[306,329,594,624]" gridcol="1" gridrow="10" pageId="9" pageNumber="16">
<emphasis id="DEF5EAFE637A6703FE81FDAE5316FDB9" box="[306,319,594,617]" italics="true" pageId="9" pageNumber="16">–</emphasis>
</td>
<td id="11605DD2637A98F5FE37FDAE53EAFDA0" box="[388,451,594,624]" gridcol="2" gridrow="10" pageId="9" pageNumber="16">
<emphasis id="DEF5EAFE637A6703FE37FDAE53BAFDBA" box="[388,403,594,618]" italics="true" pageId="9" pageNumber="16">l’</emphasis>
, 
<emphasis id="DEF5EAFE637A6703FE13FDAE5390FDB9" box="[416,441,594,617]" italics="true" pageId="9" pageNumber="16">v”</emphasis>
</td>
<td id="11605DD2637A98F5FDBEFDAE50AEFDA0" box="[525,647,594,624]" gridcol="3" gridrow="10" pageId="9" pageNumber="16">
<emphasis id="DEF5EAFE637A6703FDBEFDAE5033FDB9" box="[525,538,594,617]" italics="true" pageId="9" pageNumber="16">–</emphasis>
</td>
<td id="11605DD2637A98F5FD6DFDAE5147FDA0" box="[734,878,594,624]" gridcol="4" gridrow="10" pageId="9" pageNumber="16">
<emphasis id="DEF5EAFE637A6703FD6DFDAF50C2FDBA" box="[734,747,595,618]" italics="true" pageId="9" pageNumber="16">–</emphasis>
</td>
<td id="11605DD2637A98F5FC2AFDAE5724FDA0" box="[921,1293,594,624]" gridcol="5" gridrow="10" pageId="9" pageNumber="16">ω 2</td>
</tr>
<tr id="52B134AE637A98F5FF10FD865724FD42" box="[163,1293,634,658]" gridrow="11" pageId="9" pageNumber="16">
<th id="11605DD2637A98F5FF10FD8652F4FD42" box="[163,221,634,658]" gridcol="0" gridrow="11" pageId="9" pageNumber="16">
<emphasis id="DEF5EAFE637A6703FF10FD875297FD42" box="[163,190,635,658]" italics="true" pageId="9" pageNumber="16">Tn</emphasis>
</th>
<td id="11605DD2637A98F5FE81FD865360FD42" box="[306,329,634,658]" gridcol="1" gridrow="11" pageId="9" pageNumber="16">–</td>
<td id="11605DD2637A98F5FE37FD8653EAFD42" box="[388,451,634,658]" gridcol="2" gridrow="11" pageId="9" pageNumber="16">–</td>
<td id="11605DD2637A98F5FDBEFD8650AEFD42" box="[525,647,634,658]" gridcol="3" gridrow="11" pageId="9" pageNumber="16">–</td>
<td id="11605DD2637A98F5FD6DFD865147FD42" box="[734,878,634,658]" gridcol="4" gridrow="11" pageId="9" pageNumber="16">
<emphasis id="DEF5EAFE637A6703FD6DFD865134FD42" box="[734,797,634,658]" italics="true" pageId="9" pageNumber="16">xt2, v’</emphasis>
</td>
<td id="11605DD2637A98F5FC2AFD865724FD42" box="[921,1293,634,658]" gridcol="5" gridrow="11" pageId="9" pageNumber="16">
<emphasis id="DEF5EAFE637A6703FC2AFD87518FFD42" box="[921,934,635,658]" italics="true" pageId="9" pageNumber="16">–</emphasis>
</td>
</tr>
<tr id="52B134AE637A98F5FF10FD5E5724FD6A" box="[163,1293,674,698]" gridrow="12" pageId="9" pageNumber="16">
<th id="11605DD2637A98F5FF10FD5E52F4FD6A" box="[163,221,674,698]" gridcol="0" gridrow="12" pageId="9" pageNumber="16">
<emphasis id="DEF5EAFE637A6703FF10FD5E5296FD6A" box="[163,191,674,698]" italics="true" pageId="9" pageNumber="16">Ad</emphasis>
</th>
<td id="11605DD2637A98F5FE81FD5E5360FD6A" box="[306,329,674,698]" gridcol="1" gridrow="12" pageId="9" pageNumber="16">–</td>
<td id="11605DD2637A98F5FE37FD5E53EAFD6A" box="[388,451,674,698]" gridcol="2" gridrow="12" pageId="9" pageNumber="16">–</td>
<td id="11605DD2637A98F5FDBEFD5E50AEFD6A" box="[525,647,674,698]" gridcol="3" gridrow="12" pageId="9" pageNumber="16">–</td>
<td id="11605DD2637A98F5FD6DFD5E5147FD6A" box="[734,878,674,698]" gridcol="4" gridrow="12" pageId="9" pageNumber="16">–</td>
<td id="11605DD2637A98F5FC2AFD5E5724FD6A" box="[921,1293,674,698]" gridcol="5" gridrow="12" pageId="9" pageNumber="16">–</td>
</tr>
<tr id="52B134AE637A98F5FF10FD375724FD33" box="[163,1293,715,739]" gridrow="13" pageId="9" pageNumber="16">
<th id="11605DD2637A98F5FF10FD375724FD33" box="[163,1293,715,739]" colspan="6" colspanRight="5" gridcol="0" gridrow="13" pageId="9" pageNumber="16">Leg III</th>
</tr>
<tr id="52B134AE637A98F5FF10FD0F5724FCDC" box="[163,1293,755,780]" gridrow="14" pageId="9" pageNumber="16">
<th id="11605DD2637A98F5FF10FD0F52F4FCDC" box="[163,221,755,780]" gridcol="0" gridrow="14" pageId="9" pageNumber="16">
<emphasis id="DEF5EAFE637A6703FF10FD085297FCDB" box="[163,190,756,779]" italics="true" pageId="9" pageNumber="16">La</emphasis>
</th>
<td id="11605DD2637A98F5FE81FD0F5360FCDC" box="[306,329,755,780]" gridcol="1" gridrow="14" pageId="9" pageNumber="16">
<emphasis id="DEF5EAFE637A6703FE81FD0F5316FCDA" box="[306,319,755,778]" italics="true" pageId="9" pageNumber="16">–</emphasis>
</td>
<td id="11605DD2637A98F5FE37FD0F53EAFCDC" box="[388,451,755,780]" gridcol="2" gridrow="14" pageId="9" pageNumber="16">
<emphasis id="DEF5EAFE637A6703FE37FD0F53B8FCDB" box="[388,401,755,779]" italics="true" pageId="9" pageNumber="16">d</emphasis>
, 
<emphasis id="DEF5EAFE637A6703FE2EFD0F5395FCDA" box="[413,444,755,778]" italics="true" pageId="9" pageNumber="16">ev’</emphasis>
</td>
<td id="11605DD2637A98F5FDBEFD0F50AEFCDC" box="[525,647,755,780]" gridcol="3" gridrow="14" pageId="9" pageNumber="16">
<emphasis id="DEF5EAFE637A6703FDBEFD0F5033FCDB" box="[525,538,755,779]" italics="true" pageId="9" pageNumber="16">d</emphasis>
, 
<emphasis id="DEF5EAFE637A6703FD94FD0F501FFCDB" box="[551,566,755,779]" italics="true" pageId="9" pageNumber="16">l’</emphasis>
, σ
</td>
<td id="11605DD2637A98F5FD6DFD0F5147FCDC" box="[734,878,755,780]" gridcol="4" gridrow="14" pageId="9" pageNumber="16">
<emphasis id="DEF5EAFE637A6703FD6DFD0F50C2FCDB" box="[734,747,755,779]" italics="true" pageId="9" pageNumber="16">d</emphasis>
, 
<emphasis id="DEF5EAFE637A6703FD44FD0F512FFCDB" box="[759,774,755,779]" italics="true" pageId="9" pageNumber="16">l’</emphasis>
, 
<emphasis id="DEF5EAFE637A6703FCA0FD0F510EFCDA" box="[787,807,755,778]" italics="true" pageId="9" pageNumber="16">v’</emphasis>
, φ
</td>
<td id="11605DD2637A98F5FC2AFD0F5724FCDC" box="[921,1293,755,780]" gridcol="5" gridrow="14" pageId="9" pageNumber="16">
<emphasis id="DEF5EAFE637A6703FC2AFD0F519CFCDB" box="[921,949,755,779]" italics="true" pageId="9" pageNumber="16">ft”</emphasis>
, 
<emphasis id="DEF5EAFE637A6703FC72FD0F51CDFCDA" box="[961,996,755,778]" italics="true" pageId="9" pageNumber="16">(tc)</emphasis>
, 
<emphasis id="DEF5EAFE637A6703FC42FD0F562FFCDA" box="[1009,1030,755,778]" italics="true" pageId="9" pageNumber="16">p’</emphasis>
, 
<emphasis id="DEF5EAFE637A6703FBA0FD0F5619FCDA" box="[1043,1072,755,778]" italics="true" pageId="9" pageNumber="16">(u)</emphasis>
, 
<emphasis id="DEF5EAFE637A6703FB8EFD0F567BFCDA" box="[1085,1106,755,778]" italics="true" pageId="9" pageNumber="16">a’</emphasis>
, 
<emphasis id="DEF5EAFE637A6703FBECFD085640FCDB" box="[1119,1129,756,779]" italics="true" pageId="9" pageNumber="16">s</emphasis>
, 
<emphasis id="DEF5EAFE637A6703FBC5FD0F56B7FCDA" box="[1142,1182,755,778]" italics="true" pageId="9" pageNumber="16">(pv)</emphasis>
</td>
</tr>
<tr id="52B134AE637A98F5FF10FCE75724FCE9" box="[163,1293,795,825]" gridrow="15" pageId="9" pageNumber="16">
<th id="11605DD2637A98F5FF10FCE752F4FCE9" box="[163,221,795,825]" gridcol="0" gridrow="15" pageId="9" pageNumber="16">
<emphasis id="DEF5EAFE637A6703FF10FCE05296FCE3" box="[163,191,796,819]" italics="true" pageId="9" pageNumber="16">Pn</emphasis>
</th>
<td id="11605DD2637A98F5FE81FCE75360FCE9" box="[306,329,795,825]" gridcol="1" gridrow="15" pageId="9" pageNumber="16">
<emphasis id="DEF5EAFE637A6703FE81FCE0536FFCE3" box="[306,326,796,819]" italics="true" pageId="9" pageNumber="16">v’</emphasis>
</td>
<td id="11605DD2637A98F5FE37FCE753EAFCE9" box="[388,451,795,825]" gridcol="2" gridrow="15" pageId="9" pageNumber="16">
<emphasis id="DEF5EAFE637A6703FE37FCE753A2FCE3" box="[388,395,795,819]" italics="true" pageId="9" pageNumber="16">l</emphasis>
p 
<emphasis id="DEF5EAFE637A6703FE21FCE053B3FCE3" box="[402,410,796,819]" italics="true" pageId="9" pageNumber="16">’</emphasis>
</td>
<td id="11605DD2637A98F5FDBEFCE750AEFCE9" box="[525,647,795,825]" gridcol="3" gridrow="15" pageId="9" pageNumber="16">–</td>
<td id="11605DD2637A98F5FD6DFCE75147FCE9" box="[734,878,795,825]" gridcol="4" gridrow="15" pageId="9" pageNumber="16">–</td>
<td id="11605DD2637A98F5FC2AFCE75724FCE9" box="[921,1293,795,825]" gridcol="5" gridrow="15" pageId="9" pageNumber="16">–</td>
</tr>
<tr id="52B134AE637A98F5FF10FCBF5724FC8B" box="[163,1293,835,859]" gridrow="16" pageId="9" pageNumber="16">
<th id="11605DD2637A98F5FF10FCBF52F4FC8B" box="[163,221,835,859]" gridcol="0" gridrow="16" pageId="9" pageNumber="16">
<emphasis id="DEF5EAFE637A6703FF10FCB852EBFC8B" box="[163,194,836,859]" italics="true" pageId="9" pageNumber="16">Dn</emphasis>
</th>
<td id="11605DD2637A98F5FE81FCBF5360FC8B" box="[306,329,835,859]" gridcol="1" gridrow="16" pageId="9" pageNumber="16">
<emphasis id="DEF5EAFE637A6703FE81FCBF5368FC8B" box="[306,321,835,859]" italics="true" pageId="9" pageNumber="16">l’</emphasis>
</td>
<td id="11605DD2637A98F5FE37FCBF53EAFC8B" box="[388,451,835,859]" gridcol="2" gridrow="16" pageId="9" pageNumber="16">–</td>
<td id="11605DD2637A98F5FDBEFCBF50AEFC8B" box="[525,647,835,859]" gridcol="3" gridrow="16" pageId="9" pageNumber="16">–</td>
<td id="11605DD2637A98F5FD6DFCBF5147FC8B" box="[734,878,835,859]" gridcol="4" gridrow="16" pageId="9" pageNumber="16">–</td>
<td id="11605DD2637A98F5FC2AFCBF5724FC8B" box="[921,1293,835,859]" gridcol="5" gridrow="16" pageId="9" pageNumber="16">–</td>
</tr>
<tr id="52B134AE637A98F5FF10FC905724FC5A" box="[163,1293,876,906]" gridrow="17" pageId="9" pageNumber="16">
<th id="11605DD2637A98F5FF10FC9052F4FC5A" box="[163,221,876,906]" gridcol="0" gridrow="17" pageId="9" pageNumber="16">
<emphasis id="DEF5EAFE637A6703FF10FC915297FC54" box="[163,190,877,900]" italics="true" pageId="9" pageNumber="16">Tn</emphasis>
</th>
<td id="11605DD2637A98F5FE81FC905360FC5A" box="[306,329,876,906]" gridcol="1" gridrow="17" pageId="9" pageNumber="16">
<emphasis id="DEF5EAFE637A6703FE81FC905316FC53" box="[306,319,876,899]" italics="true" pageId="9" pageNumber="16">–</emphasis>
</td>
<td id="11605DD2637A98F5FE37FC9053EAFC5A" box="[388,451,876,906]" gridcol="2" gridrow="17" pageId="9" pageNumber="16">
<emphasis id="DEF5EAFE637A6703FE37FC9053A2FC54" box="[388,395,876,900]" italics="true" pageId="9" pageNumber="16">l</emphasis>
a 
<emphasis id="DEF5EAFE637A6703FE22FC9053B0FC53" box="[401,409,876,899]" italics="true" pageId="9" pageNumber="16">’</emphasis>
</td>
<td id="11605DD2637A98F5FDBEFC9050AEFC5A" box="[525,647,876,906]" gridcol="3" gridrow="17" pageId="9" pageNumber="16">–</td>
<td id="11605DD2637A98F5FD6DFC905147FC5A" box="[734,878,876,906]" gridcol="4" gridrow="17" pageId="9" pageNumber="16">–</td>
<td id="11605DD2637A98F5FC2AFC905724FC5A" box="[921,1293,876,906]" gridcol="5" gridrow="17" pageId="9" pageNumber="16">–</td>
</tr>
<tr id="52B134AE637A98F5FF10FC685724FC7C" box="[163,1293,916,940]" gridrow="18" pageId="9" pageNumber="16">
<th id="11605DD2637A98F5FF10FC6852F4FC7C" box="[163,221,916,940]" gridcol="0" gridrow="18" pageId="9" pageNumber="16">
<emphasis id="DEF5EAFE637A6703FF10FC685296FC7C" box="[163,191,916,940]" italics="true" pageId="9" pageNumber="16">Ad</emphasis>
</th>
<td id="11605DD2637A98F5FE81FC685360FC7C" box="[306,329,916,940]" gridcol="1" gridrow="18" pageId="9" pageNumber="16">–</td>
<td id="11605DD2637A98F5FE37FC6853EAFC7C" box="[388,451,916,940]" gridcol="2" gridrow="18" pageId="9" pageNumber="16">–</td>
<td id="11605DD2637A98F5FDBEFC6850AEFC7C" box="[525,647,916,940]" gridcol="3" gridrow="18" pageId="9" pageNumber="16">–</td>
<td id="11605DD2637A98F5FD6DFC685147FC7C" box="[734,878,916,940]" gridcol="4" gridrow="18" pageId="9" pageNumber="16">–</td>
<td id="11605DD2637A98F5FC2AFC685724FC7C" box="[921,1293,916,940]" gridcol="5" gridrow="18" pageId="9" pageNumber="16">–</td>
</tr>
<tr id="52B134AE637A98F5FF10FC415724FC05" box="[163,1293,957,981]" gridrow="19" pageId="9" pageNumber="16">
<th id="11605DD2637A98F5FF10FC415724FC05" box="[163,1293,957,981]" colspan="6" colspanRight="5" gridcol="0" gridrow="19" pageId="9" pageNumber="16">Leg IV</th>
</tr>
<tr id="52B134AE637A98F5FF10FC195724FC2D" box="[163,1293,997,1021]" gridrow="20" pageId="9" pageNumber="16">
<th id="11605DD2637A98F5FF10FC1952F4FC2D" box="[163,221,997,1021]" gridcol="0" gridrow="20" pageId="9" pageNumber="16">
<emphasis id="DEF5EAFE637A6703FF10FC195296FC2C" box="[163,191,997,1020]" italics="true" pageId="9" pageNumber="16">Pn</emphasis>
</th>
<td id="11605DD2637A98F5FE81FC195360FC2D" box="[306,329,997,1021]" gridcol="1" gridrow="20" pageId="9" pageNumber="16">–</td>
<td id="11605DD2637A98F5FE37FC1953EAFC2D" box="[388,451,997,1021]" gridcol="2" gridrow="20" pageId="9" pageNumber="16">–</td>
<td id="11605DD2637A98F5FDBEFC1950AEFC2D" box="[525,647,997,1021]" gridcol="3" gridrow="20" pageId="9" pageNumber="16">–</td>
<td id="11605DD2637A98F5FD6DFC195147FC2D" box="[734,878,997,1021]" gridcol="4" gridrow="20" pageId="9" pageNumber="16">–</td>
<td id="11605DD2637A98F5FC2AFC195724FC2D" box="[921,1293,997,1021]" gridcol="5" gridrow="20" pageId="9" pageNumber="16">
<emphasis id="DEF5EAFE637A6703FC2AFC19519CFC2D" box="[921,949,997,1021]" italics="true" pageId="9" pageNumber="16">ft”</emphasis>
, 
<emphasis id="DEF5EAFE637A6703FC72FC1951FFFC2C" box="[961,982,997,1020]" italics="true" pageId="9" pageNumber="16">p’</emphasis>
, 
<emphasis id="DEF5EAFE637A6703FC50FC195628FC2C" box="[995,1025,997,1020]" italics="true" pageId="9" pageNumber="16">(u)</emphasis>
, 
<emphasis id="DEF5EAFE637A6703FBBEFC19561FFC2C" box="[1037,1078,997,1020]" italics="true" pageId="9" pageNumber="16">(pv)</emphasis>
</td>
</tr>
<tr id="52B134AE637A98F5FF10FBF15724FBF5" box="[163,1293,1037,1061]" gridrow="21" pageId="9" pageNumber="16">
<th id="11605DD2637A98F5FF10FBF152F4FBF5" box="[163,221,1037,1061]" gridcol="0" gridrow="21" pageId="9" pageNumber="16">
<emphasis id="DEF5EAFE637A6703FF10FBF252EBFBF5" box="[163,194,1038,1061]" italics="true" pageId="9" pageNumber="16">Dn</emphasis>
</th>
<td id="11605DD2637A98F5FE81FBF15360FBF5" box="[306,329,1037,1061]" gridcol="1" gridrow="21" pageId="9" pageNumber="16">
<emphasis id="DEF5EAFE637A6703FE81FBF1536FFBF4" box="[306,326,1037,1060]" italics="true" pageId="9" pageNumber="16">v’</emphasis>
</td>
<td id="11605DD2637A98F5FE37FBF153EAFBF5" box="[388,451,1037,1061]" gridcol="2" gridrow="21" pageId="9" pageNumber="16">
<emphasis id="DEF5EAFE637A6703FE37FBF153B8FBF5" box="[388,401,1037,1061]" italics="true" pageId="9" pageNumber="16">d</emphasis>
, 
<emphasis id="DEF5EAFE637A6703FE2EFBF15395FBF4" box="[413,444,1037,1060]" italics="true" pageId="9" pageNumber="16">ev’</emphasis>
</td>
<td id="11605DD2637A98F5FDBEFBF150AEFBF5" box="[525,647,1037,1061]" gridcol="3" gridrow="21" pageId="9" pageNumber="16">
<emphasis id="DEF5EAFE637A6703FDBEFBF15033FBF5" box="[525,538,1037,1061]" italics="true" pageId="9" pageNumber="16">d</emphasis>
, 
<emphasis id="DEF5EAFE637A6703FD94FBF1501FFBF5" box="[551,566,1037,1061]" italics="true" pageId="9" pageNumber="16">l’</emphasis>
, σ
</td>
<td id="11605DD2637A98F5FD6DFBF15147FBF5" box="[734,878,1037,1061]" gridcol="4" gridrow="21" pageId="9" pageNumber="16">
<emphasis id="DEF5EAFE637A6703FD6DFBF150C2FBF5" box="[734,747,1037,1061]" italics="true" pageId="9" pageNumber="16">d</emphasis>
, 
<emphasis id="DEF5EAFE637A6703FD44FBF1512FFBF5" box="[759,774,1037,1061]" italics="true" pageId="9" pageNumber="16">l’</emphasis>
</td>
<td id="11605DD2637A98F5FC2AFBF15724FBF5" box="[921,1293,1037,1061]" gridcol="5" gridrow="21" pageId="9" pageNumber="16">
<emphasis id="DEF5EAFE637A6703FC2AFBF15195FBF4" box="[921,956,1037,1060]" italics="true" pageId="9" pageNumber="16">(tc)</emphasis>
, 
<emphasis id="DEF5EAFE637A6703FC7BFBF151F4FBF4" box="[968,989,1037,1060]" italics="true" pageId="9" pageNumber="16">a’</emphasis>
, 
<emphasis id="DEF5EAFE637A6703FC59FBF251DDFBF5" box="[1002,1012,1038,1061]" italics="true" pageId="9" pageNumber="16">s</emphasis>
</td>
</tr>
<tr id="52B134AE637A98F5FF10FBC95724FB9D" box="[163,1293,1077,1101]" gridrow="22" pageId="9" pageNumber="16">
<th id="11605DD2637A98F5FF10FBC952F4FB9D" box="[163,221,1077,1101]" gridcol="0" gridrow="22" pageId="9" pageNumber="16">
<emphasis id="DEF5EAFE637A6703FF10FBCA5297FB9D" box="[163,190,1078,1101]" italics="true" pageId="9" pageNumber="16">Tn</emphasis>
</th>
<td id="11605DD2637A98F5FE81FBC95360FB9D" box="[306,329,1077,1101]" gridcol="1" gridrow="22" pageId="9" pageNumber="16">
<emphasis id="DEF5EAFE637A6703FE81FBC95368FB9D" box="[306,321,1077,1101]" italics="true" pageId="9" pageNumber="16">l’</emphasis>
</td>
<td id="11605DD2637A98F5FE37FBC953EAFB9D" box="[388,451,1077,1101]" gridcol="2" gridrow="22" pageId="9" pageNumber="16">
<emphasis id="DEF5EAFE637A6703FE37FBC953BAFB9D" box="[388,403,1077,1101]" italics="true" pageId="9" pageNumber="16">l’</emphasis>
</td>
<td id="11605DD2637A98F5FDBEFBC950AEFB9D" box="[525,647,1077,1101]" gridcol="3" gridrow="22" pageId="9" pageNumber="16">
<emphasis id="DEF5EAFE637A6703FDBEFBCA5033FB9D" box="[525,538,1078,1101]" italics="true" pageId="9" pageNumber="16">–</emphasis>
</td>
<td id="11605DD2637A98F5FD6DFBC95147FB9D" box="[734,878,1077,1101]" gridcol="4" gridrow="22" pageId="9" pageNumber="16">
<emphasis id="DEF5EAFE637A6703FD6DFBCA50DBFB9D" box="[734,754,1078,1101]" italics="true" pageId="9" pageNumber="16">v’</emphasis>
</td>
<td id="11605DD2637A98F5FC2AFBC95724FB9D" box="[921,1293,1077,1101]" gridcol="5" gridrow="22" pageId="9" pageNumber="16">
<emphasis id="DEF5EAFE637A6703FC2AFBCA518FFB9D" box="[921,934,1078,1101]" italics="true" pageId="9" pageNumber="16">–</emphasis>
</td>
</tr>
<tr id="52B134AE637A98F5FF10FBA15724FBA5" box="[163,1293,1117,1141]" gridrow="23" pageId="9" pageNumber="16">
<th id="11605DD2637A98F5FF10FBA152F4FBA5" box="[163,221,1117,1141]" gridcol="0" gridrow="23" pageId="9" pageNumber="16">
<emphasis id="DEF5EAFE637A6703FF10FBA15296FBA5" box="[163,191,1117,1141]" italics="true" pageId="9" pageNumber="16">Ad</emphasis>
</th>
<td id="11605DD2637A98F5FE81FBA15360FBA5" box="[306,329,1117,1141]" gridcol="1" gridrow="23" pageId="9" pageNumber="16">–</td>
<td id="11605DD2637A98F5FE37FBA153EAFBA5" box="[388,451,1117,1141]" gridcol="2" gridrow="23" pageId="9" pageNumber="16">–</td>
<td id="11605DD2637A98F5FDBEFBA150AEFBA5" box="[525,647,1117,1141]" gridcol="3" gridrow="23" pageId="9" pageNumber="16">–</td>
<td id="11605DD2637A98F5FD6DFBA15147FBA5" box="[734,878,1117,1141]" gridcol="4" gridrow="23" pageId="9" pageNumber="16">–</td>
<td id="11605DD2637A98F5FC2AFBA15724FBA5" box="[921,1293,1117,1141]" gridcol="5" gridrow="23" pageId="9" pageNumber="16">–</td>
</tr>
</table>
</paragraph>
<paragraph id="EC3E36EC637A6703FF24FB7F56B8FB32" blockId="9.[151,1437,1155,1251]" pageId="9" pageNumber="16">
<tableNote id="11673762637A6703FF24FB7F56B8FB32" pageId="9" pageNumber="16" targetBox="[163,1293,192,1141]" targetPageId="9">
Note: Roman letters refer to normal setae, Greek letters refer to solenidia (except ɛ = famulus). Single quotation mark (
<emphasis id="DEF5EAFE637A6703FF13FB5B5280FB6E" box="[160,169,1191,1214]" italics="true" pageId="9" pageNumber="16">’</emphasis>
) designates setae on the anterior and double quotation mark (
<emphasis id="DEF5EAFE637A6703FCE1FB5B5148FB6E" box="[850,865,1191,1214]" italics="true" pageId="9" pageNumber="16">”</emphasis>
) setae on the posterior side of a given leg segment. Parentheses refer to a pair of setae. Setae are listed only for the stage in which they first appear.
</tableNote>
</paragraph>
<subSubSection id="A49B6567637A6703FF74FAED509AF9D3" pageId="9" pageNumber="16" type="materials_examined">
<paragraph id="EC3E36EC637A6703FF74FAED5638FAA3" blockId="9.[151,1437,1297,1791]" pageId="9" pageNumber="16">
<emphasis id="DEF5EAFE637A6703FF74FAED5399FAFB" bold="true" box="[199,432,1297,1323]" pageId="9" pageNumber="16">Material examined.</emphasis>
<materialsCitation id="5CE93CB1637A6703FE04FAED5036FAFB" ID-GBIF-Occurrence="3901404302" box="[439,543,1297,1323]" collectingDate="2021-12-25" collectorName="O. Joharchi" country="Maldives" latitude="3.9445834" location="Maafushi Island" longLatPrecision="1" longitude="73.49311" municipality="Maafushi Island" pageId="9" pageNumber="16" specimenCount="1" typeStatus="holotype">
<typeStatus id="333A884E637A6703FE04FAED5036FAFB" box="[439,543,1297,1323]" pageId="9" pageNumber="16" type="holotype">Holotype</typeStatus>
</materialsCitation>
and 
<materialsCitation id="5CE93CB1637A6703FDE6FAED50C5FAFB" ID-GBIF-Occurrence="3901404301" box="[597,748,1297,1323]" collectingDate="2021-12-25" collectorName="O. Joharchi" country="Maldives" latitude="3.9445834" location="Maafushi Island" longLatPrecision="1" longitude="73.49311" municipality="Maafushi Island" pageId="9" pageNumber="16" specimenCount="1" typeStatus="paratype">
<specimenCount id="FA87FD65637A6703FDE6FAED505FFAFB" box="[597,630,1297,1323]" pageId="9" pageNumber="16" type="generic">six</specimenCount>
<typeStatus id="333A884E637A6703FDCFFAED50C5FAFB" box="[636,748,1297,1323]" pageId="9" pageNumber="16" type="paratype">paratypes</typeStatus>
</materialsCitation>
: 
<collectingCountry id="9496767C637A6703FD45FAED5148FAFB" box="[758,865,1297,1323]" name="Maldives" pageId="9" pageNumber="16">Maldives</collectingCountry>
, 
<collectingMunicipality id="0C5AAC96637A6703FCDFFAED560AFAFB" box="[876,1059,1297,1323]" pageId="9" pageNumber="16">Maafushi Island</collectingMunicipality>
, 
<geoCoordinate id="89B5502B637A6703FB9DFAED5695FAFC" box="[1070,1212,1297,1324]" degrees="3" direction="north" minutes="56" orientation="latitude" pageId="9" pageNumber="16" precision="1" seconds="40.5" value="3.9445834">3°56'40.5&quot;N</geoCoordinate>
, 
<geoCoordinate id="89B5502B637A6703FB74FAED5776FAFC" box="[1223,1375,1297,1324]" degrees="73" direction="east" minutes="29" orientation="longitude" pageId="9" pageNumber="16" precision="1" seconds="35.2" value="73.49311">73°29'35.2&quot;E</geoCoordinate>
, soillitter samples under the Indian Almond Tree near the coast, 
<date id="983F102C637A6703FC9FFAC9519CFA80" box="[812,949,1333,1360]" pageId="9" pageNumber="16" value="2021-12-25">
<collectingDate id="887BE9C4637A6703FC9FFAC9519CFA80" box="[812,949,1333,1360]" pageId="9" pageNumber="16" value="2021-12-25">25.XII.2021</collectingDate>
</date>
(leg 
<collectorName id="4174533A637A6703FC5EFAC9565BFA9F" box="[1005,1138,1333,1359]" pageId="9" pageNumber="16">O. Joharchi</collectorName>
). Three larvae, four proto-, 11 deuto-, and 14 tritonymphs: the same data as for the 
<typeStatus id="333A884E637A6703FCB5FAA55141FAA3" box="[774,872,1369,1395]" pageId="9" pageNumber="16" type="holotype">holotype</typeStatus>
and 
<typeStatus id="333A884E637A6703FC2DFAA55624FAA3" box="[926,1037,1369,1395]" pageId="9" pageNumber="16" type="paratype">paratypes</typeStatus>
.
</paragraph>
<paragraph id="EC3E36EC637A6703FF74FA81509AF9D3" blockId="9.[151,1437,1297,1791]" pageId="9" pageNumber="16">
<emphasis id="DEF5EAFE637A6703FF74FA8153A5FA47" bold="true" box="[199,396,1405,1431]" pageId="9" pageNumber="16">Type deposition.</emphasis>
The 
<typeStatus id="333A884E637A6703FE74FA815000FA47" box="[455,553,1405,1431]" pageId="9" pageNumber="16" type="holotype">holotype</typeStatus>
is deposited in the collection of the Senckenberg Museum of Natural History, Görlitz, 
<collectingCountry id="9496767C637A6703FF4AFA5D534CFA6B" box="[249,357,1441,1467]" name="Germany" pageId="9" pageNumber="16">Germany</collectingCountry>
; 
<specimenCount id="FA87FD65637A6703FEC7FA5D5027FA6B" box="[372,526,1441,1467]" pageId="9" pageNumber="16" type="generic" typeStatus="paratypes">six paratypes</specimenCount>
, 
<specimenCount id="FA87FD65637A6703FDAEFA5D5081FA6B" box="[541,680,1441,1467]" pageId="9" pageNumber="16" type="larva">three larvae</specimenCount>
, four proto-, 11 deuto-, and 14 tritonymphs are deposited in the collection of the 
<collectingRegion id="2E45F80E637A6703FEE9FA3953DFFA0F" box="[346,502,1477,1503]" country="Russia" name="Tyumen'" pageId="9" pageNumber="16">Tyumen State</collectingRegion>
University Museum of Zoology, 
<collectingRegion id="2E45F80E637A6703FCC5FA3951FDFA0F" box="[886,980,1477,1503]" country="Russia" name="Tyumen'" pageId="9" pageNumber="16">Tyumen</collectingRegion>
, 
<collectingCountry id="9496767C637A6703FC51FA395619FA0F" box="[994,1072,1477,1503]" name="Russia" pageId="9" pageNumber="16">Russia</collectingCountry>
. All specimens are preserved in 70% solution of ethanol with a drop of glycerol.
</paragraph>
</subSubSection>
<subSubSection id="A49B6567637A6703FF74F9F156EAF9F7" box="[199,1219,1549,1575]" pageId="9" pageNumber="16" type="etymology">
<paragraph id="EC3E36EC637A6703FF74F9F156EAF9F7" blockId="9.[151,1437,1297,1791]" box="[199,1219,1549,1575]" pageId="9" pageNumber="16">
<emphasis id="DEF5EAFE637A6703FF74F9F15367F9F7" bold="true" box="[199,334,1549,1575]" pageId="9" pageNumber="16">Etymology.</emphasis>
The species name 
<taxonomicName id="2B814D6F637A6703FD90F9F150EBF9F7" authorityName="Bayartogtokh &amp; Ermilov &amp; Joharchi" authorityYear="2022" box="[547,706,1549,1575]" class="Arachnida" family="Lohmanniidae" genus="Lepidacarus" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="16" phylum="Arthropoda" rank="species" species="maafushiensis">
<emphasis id="DEF5EAFE637A6703FD90F9F150EBF9F7" box="[547,706,1549,1575]" italics="true" pageId="9" pageNumber="16">maafushiensis</emphasis>
</taxonomicName>
refers to the place of origin, Maafushi Island.
</paragraph>
</subSubSection>
<subSubSection id="A49B6567637A6703FF74F9CD577AF92F" pageId="9" pageNumber="16" type="diagnosis">
<paragraph id="EC3E36EC637A6703FF74F9CD577AF92F" blockId="9.[151,1437,1297,1791]" pageId="9" pageNumber="16">
<emphasis id="DEF5EAFE637A6703FF74F9CD53E4F99B" bold="true" box="[199,461,1585,1611]" pageId="9" pageNumber="16">Differential diagnosis.</emphasis>
In having elongate fusiform bothridial seta and numerous notogastral depressions, adult of 
<taxonomicName id="2B814D6F637A6703FF07F9A953CCF9BF" authority="Bayartogtokh &amp; Ermilov &amp; Joharchi, 2022" authorityName="Bayartogtokh &amp; Ermilov &amp; Joharchi" authorityYear="2022" box="[180,485,1621,1647]" class="Arachnida" family="Lohmanniidae" genus="Lepidacarus" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="16" phylum="Arthropoda" rank="species" species="maafushiensis" status="sp. nov.">
<emphasis id="DEF5EAFE637A6703FF07F9A953CCF9BF" box="[180,485,1621,1647]" italics="true" pageId="9" pageNumber="16">Lepidacarus maafushiensis</emphasis>
</taxonomicName>
<emphasis id="DEF5EAFE637A6703FE59F9A9506BF9BF" bold="true" box="[490,578,1621,1647]" pageId="9" pageNumber="16">
<taxonomicNameLabel id="C5C65785637A6703FE59F9A9506BF9BF" box="[490,578,1621,1647]" pageId="9" pageNumber="16" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is similar to 
<taxonomicName id="2B814D6F637A6703FD7CF9A95681F9BF" authority="Haq &amp; Ramani, 1997" authorityName="Haq &amp; Ramani" authorityYear="1997" box="[719,1192,1621,1647]" class="Arachnida" family="Lohmanniidae" genus="Lepidacarus" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="16" phylum="Arthropoda" rank="species" species="ennarpi">
<emphasis id="DEF5EAFE637A6703FD7CF9A9519EF9BF" box="[719,951,1621,1647]" italics="true" pageId="9" pageNumber="16">Lepidacarus ennarpi</emphasis>
<bibRefCitation id="88104B1D637A6703FC0EF9A95681F9BF" author="Haq, M. A. &amp; Ramani, N." box="[957,1192,1621,1647]" pageId="9" pageNumber="16" pagination="119 - 123" refId="ref12103" refString="Haq, M. A. &amp; Ramani, N. (1997) Oribatid mites from Lakshadweep - a new species of Lepidacarus Csiszar (Acari: Oribatei: Lohmanniidae). Entomon, 22 (2), 119 - 123." type="journal article" year="1997">Haq &amp; Ramani, 1997</bibRefCitation>
</taxonomicName>
from 
<collectingCountry id="9496767C637A6703FB59F9A9570EF9BF" box="[1258,1319,1621,1647]" name="India" pageId="9" pageNumber="16">India</collectingCountry>
. However, the new species can be distinguished from the later by the size of rostral setae (comparatively very large, connected medially versus medium-sized, distinctly separated), the morphology of rostrum (broadly concave, with median triangular tooth versus rounded) and bothridial setae (head sparsely ciliate on one side and densely serrate opposite side versus both sides of head ciliate), and the number of setae on epimere IV (four pairs versus three pairs).
</paragraph>
</subSubSection>
<subSubSection id="A49B6567637A6700FF24F8AD5686FDED" lastPageId="10" lastPageNumber="17" pageId="9" pageNumber="16" type="description">
<paragraph id="EC3E36EC637A6703FF24F8AD5029F8BB" blockId="9.[151,512,1873,1899]" box="[151,512,1873,1899]" pageId="9" pageNumber="16">
<heading id="B7768180637A6703FF24F8AD5029F8BB" bold="true" box="[151,512,1873,1899]" fontSize="11" level="1" pageId="9" pageNumber="16" reason="1">
<emphasis id="DEF5EAFE637A6703FF24F8AD5029F8BB" bold="true" box="[151,512,1873,1899]" pageId="9" pageNumber="16">Ontogenetic transformations</emphasis>
</heading>
</paragraph>
<paragraph id="EC3E36EC637A6700FF24F8655686FDED" blockId="9.[151,1437,1945,2043]" lastBlockId="10.[151,1437,151,573]" lastPageId="10" lastPageNumber="17" pageId="9" pageNumber="16">
The juveniles are colourless or slightly yellowish in some nymphs, adult of yellowish-brown colour. In both juveniles and adult, cuticle of body and legs densely microreticulate, prodorsum and gastronotum sparsely microtuberculate. Rostrum broadly concave, with median triangular tooth, in both juveniles and adult. Form of the prodorsal and gastronotic/notogastral setae is the same throughout ontogenetic instars: larva with 13 pairs of gastronotic setae (larval seta 
<emphasis id="DEF5EAFE63796700FEA8FF475300FF05" box="[283,297,187,213]" italics="true" pageId="10" pageNumber="17">h</emphasis>
<subScript id="700534A963796700FE9AFF315318FF0B" attach="left" box="[297,305,205,219]" fontSize="6" pageId="10" pageNumber="17">4</subScript>
absent), protonymph with 17 pairs (setae of 
<emphasis id="DEF5EAFE63796700FC9BFF47511FFF04" box="[808,822,187,212]" italics="true" pageId="10" pageNumber="17">p</emphasis>
-series newly appeared), deuto-, tritonymphs and adult with 18 pairs of gastronotic/notogastral setae. Larva with three pairs of subcapitular setae (
<emphasis id="DEF5EAFE63796700FBCEFF2356A2FF28" box="[1149,1163,223,248]" italics="true" pageId="10" pageNumber="17">a</emphasis>
, 
<emphasis id="DEF5EAFE63796700FB24FF235682FF28" box="[1175,1195,223,248]" italics="true" pageId="10" pageNumber="17">m</emphasis>
<subScript id="700534A963796700FB1FFF0D569DFF2F" attach="right" box="[1196,1204,241,255]" fontSize="6" pageId="10" pageNumber="17">1</subScript>
, 
<emphasis id="DEF5EAFE63796700FB73FF2356E7FF29" box="[1216,1230,223,249]" italics="true" pageId="10" pageNumber="17">h</emphasis>
), seta 
<emphasis id="DEF5EAFE63796700FAA6FF235700FF28" box="[1301,1321,223,248]" italics="true" pageId="10" pageNumber="17">m</emphasis>
<subScript id="700534A963796700FA99FF0D571BFF2F" attach="left" box="[1322,1330,241,255]" fontSize="6" pageId="10" pageNumber="17">2</subScript>
appeared in protonymph, 
<emphasis id="DEF5EAFE63796700FEFDFEFF534BFECC" box="[334,354,259,284]" italics="true" pageId="10" pageNumber="17">m</emphasis>
<quantity id="2B799B0963796700FED1FEE953A0FECD" box="[354,393,259,291]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="10" pageNumber="17" unit="in" value="3.0">
<subScript id="700534A963796700FED1FEE95343FEF3" attach="left" box="[354,362,277,291]" fontSize="6" pageId="10" pageNumber="17">3</subScript>
in
</quantity>
deutonymph and 
<emphasis id="DEF5EAFE63796700FDE4FEFF5042FECC" box="[599,619,259,284]" italics="true" pageId="10" pageNumber="17">m</emphasis>
<quantity id="2B799B0963796700FDD8FEE950BBFECD" box="[619,658,259,291]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="10" pageNumber="17" unit="in" value="4.0">
<subScript id="700534A963796700FDD8FEE9505AFEF3" attach="left" box="[619,627,277,291]" fontSize="6" pageId="10" pageNumber="17">4</subScript>
in
</quantity>
tritonymph. Larva with two pairs of adoral setae (
<emphasis id="DEF5EAFE63796700FB66FEFF56C7FECC" box="[1237,1262,259,284]" italics="true" pageId="10" pageNumber="17">or</emphasis>
<subScript id="700534A963796700FB5DFEE956DFFEF3" attach="left" box="[1262,1270,277,291]" fontSize="6" pageId="10" pageNumber="17">1</subScript>
, 
<emphasis id="DEF5EAFE63796700FAB5FEFF5736FECC" box="[1286,1311,259,284]" italics="true" pageId="10" pageNumber="17">or</emphasis>
<subScript id="700534A963796700FAACFEE9570EFEF3" attach="left" box="[1311,1319,277,291]" fontSize="6" pageId="10" pageNumber="17">2</subScript>
), and seta 
<emphasis id="DEF5EAFE63796700FF24FEDB5299FE90" box="[151,176,295,320]" italics="true" pageId="10" pageNumber="17">or</emphasis>
<subScript id="700534A963796700FF03FEC55291FE97" attach="left" box="[176,184,313,327]" fontSize="6" pageId="10" pageNumber="17">3</subScript>
added in protonymph. Palp femur with no seta in larva and protonymph, but femoral seta added in deutonymph. Larva with six pairs of epimeral setae (
<emphasis id="DEF5EAFE63796700FDE7FEB75059FEB5" box="[596,624,331,357]" italics="true" pageId="10" pageNumber="17">1a</emphasis>
, 
<emphasis id="DEF5EAFE63796700FDCCFEB750B5FEB5" box="[639,668,331,357]" italics="true" pageId="10" pageNumber="17">1b</emphasis>
, 
<emphasis id="DEF5EAFE63796700FD19FEB750ECFEB5" box="[682,709,331,357]" italics="true" pageId="10" pageNumber="17">1c</emphasis>
, 
<emphasis id="DEF5EAFE63796700FD67FEB750C6FEB5" box="[724,751,331,357]" italics="true" pageId="10" pageNumber="17">2a</emphasis>
, 
<emphasis id="DEF5EAFE63796700FD4CFEB75133FEB5" box="[767,794,331,357]" italics="true" pageId="10" pageNumber="17">3a</emphasis>
, 
<emphasis id="DEF5EAFE63796700FC99FEB7516FFEB5" box="[810,838,331,357]" italics="true" pageId="10" pageNumber="17">3b</emphasis>
), seta 
<emphasis id="DEF5EAFE63796700FC21FEB75184FEB5" box="[914,941,331,357]" italics="true" pageId="10" pageNumber="17">1d</emphasis>
, 
<emphasis id="DEF5EAFE63796700FC0EFEB751F1FEB5" box="[957,984,331,357]" italics="true" pageId="10" pageNumber="17">2b</emphasis>
and 
<emphasis id="DEF5EAFE63796700FBA2FEB75604FEB5" box="[1041,1069,331,357]" italics="true" pageId="10" pageNumber="17">4a</emphasis>
newly appeared in protonymph, while 
<emphasis id="DEF5EAFE63796700FF68FE9352DCFE59" box="[219,245,367,393]" italics="true" pageId="10" pageNumber="17">3c</emphasis>
, 
<emphasis id="DEF5EAFE63796700FEB2FE935337FE59" box="[257,286,367,393]" italics="true" pageId="10" pageNumber="17">4b</emphasis>
, 
<emphasis id="DEF5EAFE63796700FE9AFE93536DFE59" box="[297,324,367,393]" italics="true" pageId="10" pageNumber="17">4c</emphasis>
added in deutonymph, seta 
<emphasis id="DEF5EAFE63796700FDC6FE9350B8FE59" box="[629,657,367,393]" italics="true" pageId="10" pageNumber="17">4d</emphasis>
developed in tritonymph, seta 
<emphasis id="DEF5EAFE63796700FC50FE9351D7FE59" box="[995,1022,367,393]" italics="true" pageId="10" pageNumber="17">4e</emphasis>
added in adult. Number of anogenital setae gradually increasing from larva to adult: larva without anogenital setae; one pair of genital and four pairs of anal setae appeared in protonymph; four pairs of genital and two pairs of anal setae formed in deutonymph; three pairs of genital setae added in tritonymph, and one more pair of genital setae added in adult. Paraproctal valves of larva (segment PS) with three pairs of setae, protonymph (segment AD) without seta, and deutonymph (segment AN) with two pairs of setae. Leg setation increasing from instar to instar as shown in 
<tableCitation id="A103035763796700FBE6FDDF5680FDED" box="[1109,1193,547,573]" captionStart="TABLE 1" captionStartId="9.[151,239,151,175]" captionTargetPageId="9" captionText="TABLE 1. Development of leg setation of Lepidacarus maafushiensis sp. nov." httpUri="http://table.plazi.org/id/B8FE6664637A6703FF24FF6B51C8FF7F" pageId="10" pageNumber="17" tableUuid="B8FE6664637A6703FF24FF6B51C8FF7F">Table 1</tableCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>