470 lines
75 KiB
XML
470 lines
75 KiB
XML
<document ID-DOI="10.3897/zookeys.19.221" ID-GBIF-Dataset="e6ea8eea-7156-4785-8313-f4bd88dadad1" ID-ISSN="1313–2970" ID-Zenodo-Dep="576503" ID-ZooBank="C473F9F6-1AE7-4B3F-B17F-CA1C2709010C" checkinTime="1587956031210" checkinUser="plazi" docAuthor="Wesener, Thomas, Enghoff, Henrik & Sierwald, Petra" docDate="2009" docId="1772122E784BFF96FF013BD7AE50EC4F" docLanguage="en" docName="ZK_article_2037.pdf" docOrigin="ZooKeys 19 (19)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@11b73717" docStyleName="zookeys.2008.journal_article" docTitle="Alluviobolus Wesener & Enghoff & Sierwald 2009, gen. n." docType="treatment" docUuid="3190519A-4F6B-45C8-BF88-209D0E41A9C8" docUuidSource="ZooBank" docVersion="6" lastPageNumber="81" masterDocId="EB4B6A567807FFC6FF8D3F74AF59EA51" masterDocTitle="Review of the Spirobolida on Madagascar, with descriptions of twelve new genera, including three genera of ' fire millipedes' (Diplopoda)" masterLastPageNumber="128" masterPageNumber="1" pageNumber="77" updateTime="1636630017160" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Review of the Spirobolida on Madagascar, with descriptions of twelve new genera, including three genera of ' fire millipedes' (Diplopoda)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Wesener, Thomas</mods:namePart>
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<mods:nameIdentifier type="LSID">urn:lsid:zoobank.org:author:</mods:nameIdentifier>
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<mods:affiliation>Field Museum of Natural History, 1400 S. Lake Shore Drive, Chicago, IL 60605, U. S. A. & Zoologisches Forschungsmuseum Alexander Koenig, Museumsmeile Bonn, Adenauerallee 160, D- 53113 Bonn, Germany. & B 54 D 3623309</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Enghoff, Henrik</mods:namePart>
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<mods:nameIdentifier type="LSID">urn:lsid:zoobank.org:author:</mods:nameIdentifier>
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<mods:affiliation>Natural History Museum of Denmark, Universitetsparken 15, DK- 2100 Copenhagen, Denmark</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Sierwald, Petra</mods:namePart>
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<mods:nameIdentifier type="LSID">urn:lsid:zoobank.org:author:</mods:nameIdentifier>
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<mods:affiliation>Field Museum of Natural History, 1400 S. Lake Shore Drive, Chicago, IL 60605, U. S. A.</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2009</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2009-09-04</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>19</mods:number>
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||
</mods:detail>
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||
<mods:detail type="issue">
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<mods:number>19</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>128</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.3897/zookeys.19.221</mods:identifier>
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<mods:identifier type="GBIF-Dataset">e6ea8eea-7156-4785-8313-f4bd88dadad1</mods:identifier>
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<mods:identifier type="ISSN">1313–2970</mods:identifier>
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<mods:identifier type="Zenodo-Dep">576503</mods:identifier>
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<mods:identifier type="ZooBank">C473F9F6-1AE7-4B3F-B17F-CA1C2709010C</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.3791439" ID-GBIF-Taxon="164184221" ID-Zenodo-Dep="3791439" LSID="urn:lsid:zoobank.org:act:3190519A-4F6B-45C8-BF88-209D0E41A9C8" httpUri="http://treatment.plazi.org/id/1772122E784BFF96FF013BD7AE50EC4F" lastPageId="80" lastPageNumber="81" pageId="76" pageNumber="77">
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<subSubSection box="[140,489,1187,1214]" pageId="76" pageNumber="77" type="nomenclature">
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<paragraph blockId="76.[140,965,1187,1249]" box="[140,489,1187,1214]" pageId="76" pageNumber="77">
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<heading box="[140,489,1187,1214]" fontSize="11" level="2" pageId="76" pageNumber="77" reason="3">
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<emphasis bold="true" box="[140,489,1187,1214]" pageId="76" pageNumber="77">
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<taxonomicName authority="Wesener" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[140,396,1187,1214]" class="Diplopoda" family="Pachybolidae" genus="Alluviobolus" kingdom="Animalia" order="Spirobolida" pageId="76" pageNumber="77" phylum="Arthropoda" rank="genus" status="gen. n.">
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<emphasis bold="true" box="[140,288,1187,1214]" italics="true" pageId="76" pageNumber="77">Alluviobolus</emphasis>
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||
Wesener
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</taxonomicName>
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||
,
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||
<taxonomicNameLabel box="[408,489,1187,1213]" pageId="76" pageNumber="77" rank="genus">gen. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection box="[140,965,1222,1249]" pageId="76" pageNumber="77" type="description">
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<paragraph blockId="76.[140,965,1187,1249]" box="[140,965,1222,1249]" pageId="76" pageNumber="77">
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<uri box="[140,965,1222,1249]" pageId="76" pageNumber="77">
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urn:lsid:zoobank.org:act:
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<uuid box="[421,965,1222,1249]" pageId="76" pageNumber="77">3190519A-4F6B-45C8-BF88-209D0E41A9C8</uuid>
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</uri>
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</paragraph>
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</subSubSection>
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<subSection lastPageId="80" lastPageNumber="81" pageId="76" pageNumber="77" type="multiple">
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<subSubSection box="[140,628,1293,1320]" pageId="76" pageNumber="77" type="type_taxon">
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<paragraph blockId="76.[140,628,1293,1320]" box="[140,628,1293,1320]" pageId="76" pageNumber="77">
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<emphasis bold="true" box="[140,287,1293,1319]" pageId="76" pageNumber="77">
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<typeStatus box="[140,198,1293,1319]" pageId="76" pageNumber="77">Type</typeStatus>
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species
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</emphasis>
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:
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<taxonomicName authority="Wesener & Enghoff & Sierwald, 2009" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[302,553,1293,1319]" class="Diplopoda" family="Pachybolidae" genus="Alluviobolus" kingdom="Animalia" order="Spirobolida" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" status="sp. n.">
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<emphasis box="[302,553,1293,1319]" italics="true" pageId="76" pageNumber="77">Alluviobolus laticlavius</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel box="[567,628,1294,1320]" pageId="76" pageNumber="77" rank="species">sp. n.</taxonomicNameLabel>
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||
</paragraph>
|
||
</subSubSection>
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||
<subSubSection pageId="76" pageNumber="77" type="reference_group">
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||
<paragraph blockId="76.[140,419,1363,1460]" box="[140,419,1363,1390]" pageId="76" pageNumber="77">
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||
<emphasis bold="true" box="[140,413,1363,1389]" pageId="76" pageNumber="77">
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||
<emphasis box="[140,212,1363,1389]" inLineHeading="true" pageId="76" pageNumber="77" reason="1">Other</emphasis>
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species included
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</emphasis>
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||
:
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||
</paragraph>
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||
<paragraph blockId="76.[140,419,1363,1460]" box="[140,341,1399,1425]" pageId="76" pageNumber="77">
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||
<taxonomicName authority="Wesener & Enghoff & Sierwald, 2009" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[140,273,1399,1425]" class="Diplopoda" family="Pachybolidae" genus="Alluviobolus" kingdom="Animalia" order="Spirobolida" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" status="sp. n.">
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<emphasis box="[140,273,1399,1425]" italics="true" pageId="76" pageNumber="77">A. tsimelahy</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[280,341,1399,1425]" pageId="76" pageNumber="77" rank="species">sp. n.</taxonomicNameLabel>
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||
</paragraph>
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||
<paragraph blockId="76.[140,419,1363,1460]" box="[140,334,1434,1460]" pageId="76" pageNumber="77">
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||
<taxonomicName authority="Wesener & Enghoff & Sierwald, 2009" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[140,266,1434,1460]" class="Diplopoda" family="Pachybolidae" genus="Alluviobolus" kingdom="Animalia" order="Spirobolida" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" status="sp. n.">
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<emphasis box="[140,266,1434,1460]" italics="true" pageId="76" pageNumber="77">A. antanosy</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[273,334,1434,1460]" pageId="76" pageNumber="77" rank="species">sp. n.</taxonomicNameLabel>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection lastPageId="77" lastPageNumber="78" pageId="76" pageNumber="77" type="diagnosis">
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||
<paragraph blockId="76.[140,1108,1504,1672]" pageId="76" pageNumber="77">
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<emphasis bold="true" box="[140,264,1504,1530]" inLineHeading="true" pageId="76" pageNumber="77" reason="1">Diagnosis</emphasis>
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: anterior (
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<figureCitation box="[389,493,1504,1530]" captionStart="Figure 44" captionStartId="78.[140,212,695,717]" captionTargetBox="[140,1108,165,676]" captionTargetId="figure@78.[140,1109,165,676]" captionTargetPageId="78" captionText="Figure 44. Alluviobolus laticlavius sp. n. male holotype. A head, lateral view B legs 1–7, ventral view C telson D gnathochilarium, ventral view E anterior gonopod, anterior view F right posterior gonopod, anterior view G anterior gonopod, posterior view H right posterior gonopod, telopodite, posterior view I posterior gonopods, coxite and sternite, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; L = ledge; Md = mandible; Me = mentum; Pre = preanal ring; St = sternite; ST = stipites; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3820827" httpUri="https://zenodo.org/record/3820827/files/figure.png" pageId="76" pageNumber="77">Fig. 44E</figureCitation>
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) and posterior gonopods (
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<figureCitation box="[803,899,1504,1531]" captionStart="Figure 45" captionStartId="79.[140,212,1462,1484]" captionTargetBox="[140,1108,165,1295]" captionTargetId="figure@79.[140,1108,165,887]" captionTargetPageId="79" captionText="Figure 45. Alluviobolus tsimelahy sp. n. A–C, F–I male holotype, D, E female paratype. A head, lateral view B legs 1–7, ventral view C telson D vulva and sterno-coxite of second leg, posterior view E left mandible, mesal view F anterior gonopod, anterior view G right posterior gonopod, anterior view H anterior gonopod, posterior view I right posterior gonopod, posterior view. 3iT = 3-combed internal tooth; Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; eT = external tooth; Gn = gnathochilarium; LT = lateral tooth; Md = mandible; mp = molar plate; O = operculum; pL = pectinate lamella; Pre = preanal ring; pv = posterior valve; St = sternite; St-Cx = coxosternite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767332" httpUri="https://zenodo.org/record/3767332/files/figure.png" pageId="76" pageNumber="77">Fig. 45I</figureCitation>
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) uniquely shaped for Malagasy
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<taxonomicName authorityName="Cook" authorityYear="1895" box="[304,445,1539,1565]" class="Diplopoda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirobolida" pageId="76" pageNumber="77" phylum="Arthropoda" rank="order">Spirobolida</taxonomicName>
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: mesal process of anterior gonopod coxite absent (
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||
<figureCitation captionStart="Figure 44" captionStartId="78.[140,212,695,717]" captionTargetBox="[140,1108,165,676]" captionTargetId="figure@78.[140,1109,165,676]" captionTargetPageId="78" captionText="Figure 44. Alluviobolus laticlavius sp. n. male holotype. A head, lateral view B legs 1–7, ventral view C telson D gnathochilarium, ventral view E anterior gonopod, anterior view F right posterior gonopod, anterior view G anterior gonopod, posterior view H right posterior gonopod, telopodite, posterior view I posterior gonopods, coxite and sternite, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; L = ledge; Md = mandible; Me = mentum; Pre = preanal ring; St = sternite; ST = stipites; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3820827" httpUri="https://zenodo.org/record/3820827/files/figure.png" pageId="76" pageNumber="77">Fig. 44E</figureCitation>
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); telopodite mesally protruding into slender process (
|
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<figureCitation box="[844,997,1575,1601]" captionStart="Figure 44" captionStartId="78.[140,212,695,717]" captionTargetBox="[140,1108,165,676]" captionTargetId="figure@78.[140,1109,165,676]" captionTargetPageId="78" captionText="Figure 44. Alluviobolus laticlavius sp. n. male holotype. A head, lateral view B legs 1–7, ventral view C telson D gnathochilarium, ventral view E anterior gonopod, anterior view F right posterior gonopod, anterior view G anterior gonopod, posterior view H right posterior gonopod, telopodite, posterior view I posterior gonopods, coxite and sternite, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; L = ledge; Md = mandible; Me = mentum; Pre = preanal ring; St = sternite; ST = stipites; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3820827" httpUri="https://zenodo.org/record/3820827/files/figure.png" pageId="76" pageNumber="77">Figs 44E, G</figureCitation>
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), process apically with a sharp-edged retrorse projection (
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<figureCitation box="[712,825,1610,1636]" captionStart="Figure 44" captionStartId="78.[140,212,695,717]" captionTargetBox="[140,1108,165,676]" captionTargetId="figure@78.[140,1109,165,676]" captionTargetPageId="78" captionText="Figure 44. Alluviobolus laticlavius sp. n. male holotype. A head, lateral view B legs 1–7, ventral view C telson D gnathochilarium, ventral view E anterior gonopod, anterior view F right posterior gonopod, anterior view G anterior gonopod, posterior view H right posterior gonopod, telopodite, posterior view I posterior gonopods, coxite and sternite, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; L = ledge; Md = mandible; Me = mentum; Pre = preanal ring; St = sternite; ST = stipites; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3820827" httpUri="https://zenodo.org/record/3820827/files/figure.png" pageId="76" pageNumber="77">Fig. 44G</figureCitation>
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). Coxite and telopodite of posterior gonopods not coaxial, telopodite slightly bended posteriorly (
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<figureCitation box="[1061,1108,1645,1671]" captionStart="Figure 42" captionStartId="77.[139,211,599,621]" captionTargetBox="[139,1107,165,580]" captionTargetId="figure@77.[139,1108,165,580]" captionTargetPageId="77" captionText="Figure 42. Alluviobolus tsimelahy sp. n., A living male from Tsimelahy B male and female specimen from Ebosika." figureDoi="http://doi.org/10.5281/zenodo.3767330" httpUri="https://zenodo.org/record/3767330/files/figure.png" pageId="76" pageNumber="77">Fig.</figureCitation>
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</paragraph>
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<caption ID-DOI="http://doi.org/10.5281/zenodo.3767330" ID-Zenodo-Dep="3767330" httpUri="https://zenodo.org/record/3767330/files/figure.png" pageId="77" pageNumber="78" startId="77.[139,211,599,621]" targetBox="[139,1107,165,580]" targetPageId="77">
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<paragraph blockId="77.[139,1107,563,654]" pageId="77" pageNumber="78">
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<emphasis bold="true" box="[139,253,599,621]" pageId="77" pageNumber="78">Figure 42.</emphasis>
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<taxonomicName authority="Wesener & Enghoff & Sierwald, 2009" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[261,461,600,621]" class="Diplopoda" family="Pachybolidae" genus="Alluviobolus" kingdom="Animalia" order="Spirobolida" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" status="sp. n.">
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<emphasis box="[261,461,600,621]" italics="true" pageId="77" pageNumber="78">Alluviobolus tsimelahy</emphasis>
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</taxonomicName>
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||
<taxonomicNameLabel box="[470,524,600,621]" pageId="77" pageNumber="78" rank="species">sp. n.</taxonomicNameLabel>
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,
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<emphasis bold="true" box="[537,556,599,621]" pageId="77" pageNumber="78">A</emphasis>
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living male from Tsimelahy
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<emphasis bold="true" box="[832,849,599,621]" pageId="77" pageNumber="78">B</emphasis>
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male and female specimen from Ebosika.
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</paragraph>
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</caption>
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<caption ID-DOI="http://doi.org/10.5281/zenodo.3820825" ID-Zenodo-Dep="3820825" box="[139,723,1099,1121]" httpUri="https://zenodo.org/record/3820825/files/figure.png" pageId="77" pageNumber="78" startId="77.[139,211,1099,1121]" targetBox="[139,1111,687,1080]" targetPageId="77">
|
||
<paragraph box="[139,723,1099,1121]" pageId="77" pageNumber="78">
|
||
<emphasis bold="true" box="[139,250,1099,1121]" pageId="77" pageNumber="78">Figure 43.</emphasis>
|
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Distribution map of the genus
|
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<emphasis box="[544,653,1099,1120]" italics="true" pageId="77" pageNumber="78">Alluviobolus</emphasis>
|
||
gen. n.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="77.[140,1108,1187,1672]" pageId="77" pageNumber="78">
|
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45I). Telopodites with a torsion, opening of sperm canal discharging apico-laterally (
|
||
<figureCitation box="[221,324,1222,1249]" captionStart="Figure 45" captionStartId="79.[140,212,1462,1484]" captionTargetBox="[140,1108,165,1295]" captionTargetId="figure@79.[140,1108,165,887]" captionTargetPageId="79" captionText="Figure 45. Alluviobolus tsimelahy sp. n. A–C, F–I male holotype, D, E female paratype. A head, lateral view B legs 1–7, ventral view C telson D vulva and sterno-coxite of second leg, posterior view E left mandible, mesal view F anterior gonopod, anterior view G right posterior gonopod, anterior view H anterior gonopod, posterior view I right posterior gonopod, posterior view. 3iT = 3-combed internal tooth; Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; eT = external tooth; Gn = gnathochilarium; LT = lateral tooth; Md = mandible; mp = molar plate; O = operculum; pL = pectinate lamella; Pre = preanal ring; pv = posterior valve; St = sternite; St-Cx = coxosternite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767332" httpUri="https://zenodo.org/record/3767332/files/figure.png" pageId="77" pageNumber="78">Figs 45I</figureCitation>
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,
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<figureCitation box="[338,396,1223,1249]" captionStart="Figure 47" captionStartId="85.[140,212,825,847]" captionTargetBox="[140,1108,165,794]" captionTargetId="figure@85.[140,1108,465,804]" captionTargetPageId="85" captionText="Figure 47. Alluviobolus antanosy sp. n., male holotype. A head, lateral view B legs 1–7, ventral view C telson D anterior gonopod, anterior view E left posterior gonopod, anterior view F anterior gonopod, posterior view G left posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; IL = incisura lateralis; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767336" httpUri="https://zenodo.org/record/3767336/files/figure.png" pageId="77" pageNumber="78">47G</figureCitation>
|
||
). Natural colour varies between species from brown, red to pitch-black (
|
||
<figureCitation box="[292,404,1258,1284]" captionStart="Figure 42" captionStartId="77.[139,211,599,621]" captionTargetBox="[139,1107,165,580]" captionTargetId="figure@77.[139,1108,165,580]" captionTargetPageId="77" captionText="Figure 42. Alluviobolus tsimelahy sp. n., A living male from Tsimelahy B male and female specimen from Ebosika." figureDoi="http://doi.org/10.5281/zenodo.3767330" httpUri="https://zenodo.org/record/3767330/files/figure.png" pageId="77" pageNumber="78">Figs 42A</figureCitation>
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,
|
||
<figureCitation box="[419,474,1258,1284]" captionStart="Figure 44" captionStartId="78.[140,212,695,717]" captionTargetBox="[140,1108,165,676]" captionTargetId="figure@78.[140,1109,165,676]" captionTargetPageId="78" captionText="Figure 44. Alluviobolus laticlavius sp. n. male holotype. A head, lateral view B legs 1–7, ventral view C telson D gnathochilarium, ventral view E anterior gonopod, anterior view F right posterior gonopod, anterior view G anterior gonopod, posterior view H right posterior gonopod, telopodite, posterior view I posterior gonopods, coxite and sternite, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; L = ledge; Md = mandible; Me = mentum; Pre = preanal ring; St = sternite; ST = stipites; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3820827" httpUri="https://zenodo.org/record/3820827/files/figure.png" pageId="77" pageNumber="78">44C</figureCitation>
|
||
), dorsally at least on mesozonites with an orange or red stripe of species-specific width. A similar stripe is also present in
|
||
<taxonomicName authorityName="de Saussure & Zehntner" authorityYear="1901" box="[965,1101,1293,1319]" class="Diplopoda" family="Pachybolidae" genus="Spiromimus" kingdom="Animalia" order="Spirobolida" pageId="77" pageNumber="78" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[965,1101,1293,1319]" italics="true" pageId="77" pageNumber="78">Spiromimus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[140,268,1328,1354]" class="Diplopoda" family="Pachybolidae" genus="Caprobolus" kingdom="Animalia" order="Spirobolida" pageId="77" pageNumber="78" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[140,268,1328,1354]" italics="true" pageId="77" pageNumber="78">Caprobolus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName authority="Wesener & Enghoff & Sierwald, 2009" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[330,463,1328,1354]" class="Diplopoda" family="Pachybolidae" genus="Ostinobolus" kingdom="Animalia" order="Spirobolida" pageId="77" pageNumber="78" phylum="Arthropoda" rank="genus" status="gen. n.">
|
||
<emphasis box="[330,463,1328,1354]" italics="true" pageId="77" pageNumber="78">Ostinobolus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[473,557,1329,1355]" pageId="77" pageNumber="78" rank="genus">gen. n.</taxonomicNameLabel>
|
||
species.
|
||
<taxonomicName authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[664,805,1328,1354]" class="Diplopoda" family="Pachybolidae" genus="Alluviobolus" kingdom="Animalia" order="Spirobolida" pageId="77" pageNumber="78" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[664,805,1328,1354]" italics="true" pageId="77" pageNumber="78">Alluviobolus</emphasis>
|
||
</taxonomicName>
|
||
species share the habitus and colour pattern with species of the sympatric living genus
|
||
<taxonomicName authority="Wesener & Enghoff & Sierwald, 2009" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[882,1015,1363,1389]" class="Diplopoda" family="Pachybolidae" genus="Ostinobolus" kingdom="Animalia" order="Spirobolida" pageId="77" pageNumber="78" phylum="Arthropoda" rank="genus" status="gen. n.">
|
||
<emphasis box="[882,1015,1363,1389]" italics="true" pageId="77" pageNumber="78">Ostinobolus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1025,1107,1364,1390]" pageId="77" pageNumber="78" rank="genus">gen. n.</taxonomicNameLabel>
|
||
Both genera, however, possess completely different anterior and posterior gonopods. Gnathochilarium with a subdivided mentum and a single sclerotized ledge on each stipites (
|
||
<figureCitation box="[343,457,1469,1495]" captionStart="Figure 44" captionStartId="78.[140,212,695,717]" captionTargetBox="[140,1108,165,676]" captionTargetId="figure@78.[140,1109,165,676]" captionTargetPageId="78" captionText="Figure 44. Alluviobolus laticlavius sp. n. male holotype. A head, lateral view B legs 1–7, ventral view C telson D gnathochilarium, ventral view E anterior gonopod, anterior view F right posterior gonopod, anterior view G anterior gonopod, posterior view H right posterior gonopod, telopodite, posterior view I posterior gonopods, coxite and sternite, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; L = ledge; Md = mandible; Me = mentum; Pre = preanal ring; St = sternite; ST = stipites; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3820827" httpUri="https://zenodo.org/record/3820827/files/figure.png" pageId="77" pageNumber="78">Fig. 44D</figureCitation>
|
||
), like in
|
||
<taxonomicName authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[565,711,1469,1495]" class="Diplopoda" family="Pachybolidae" genus="Flagellobolus" kingdom="Animalia" order="Spirobolida" pageId="77" pageNumber="78" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[565,711,1469,1495]" italics="true" pageId="77" pageNumber="78">Flagellobolus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[728,879,1469,1495]" class="Diplopoda" family="Pachybolidae" genus="Riotintobolus" kingdom="Animalia" order="Spirobolida" pageId="77" pageNumber="78" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[728,879,1469,1495]" italics="true" pageId="77" pageNumber="78">Riotintobolus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[896,1099,1469,1495]" class="Diplopoda" family="Pachybolidae" genus="Pseudocentrobolus" kingdom="Animalia" order="Spirobolida" pageId="77" pageNumber="78" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[896,1099,1469,1495]" italics="true" pageId="77" pageNumber="78">Pseudocentrobolus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[140,287,1504,1530]" class="Diplopoda" family="Pachybolidae" genus="Granitobolus" kingdom="Animalia" order="Spirobolida" pageId="77" pageNumber="78" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[140,287,1504,1530]" italics="true" pageId="77" pageNumber="78">Granitobolus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[306,434,1504,1530]" class="Diplopoda" family="Pachybolidae" genus="Caprobolus" kingdom="Animalia" order="Spirobolida" pageId="77" pageNumber="78" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[306,434,1504,1530]" italics="true" pageId="77" pageNumber="78">Caprobolus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authority="Wesener & Enghoff & Sierwald, 2009" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[453,586,1504,1530]" class="Diplopoda" family="Pachybolidae" genus="Ostinobolus" kingdom="Animalia" order="Spirobolida" pageId="77" pageNumber="78" phylum="Arthropoda" rank="genus" status="gen. n.">
|
||
<emphasis box="[453,586,1504,1530]" italics="true" pageId="77" pageNumber="78">Ostinobolus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[598,686,1504,1531]" pageId="77" pageNumber="78" rank="genus">
|
||
<emphasis bold="true" box="[598,678,1504,1530]" pageId="77" pageNumber="78">gen. n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
A torsion of the posterior gonopods (
|
||
<figureCitation box="[198,295,1539,1566]" captionStart="Figure 45" captionStartId="79.[140,212,1462,1484]" captionTargetBox="[140,1108,165,1295]" captionTargetId="figure@79.[140,1108,165,887]" captionTargetPageId="79" captionText="Figure 45. Alluviobolus tsimelahy sp. n. A–C, F–I male holotype, D, E female paratype. A head, lateral view B legs 1–7, ventral view C telson D vulva and sterno-coxite of second leg, posterior view E left mandible, mesal view F anterior gonopod, anterior view G right posterior gonopod, anterior view H anterior gonopod, posterior view I right posterior gonopod, posterior view. 3iT = 3-combed internal tooth; Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; eT = external tooth; Gn = gnathochilarium; LT = lateral tooth; Md = mandible; mp = molar plate; O = operculum; pL = pectinate lamella; Pre = preanal ring; pv = posterior valve; St = sternite; St-Cx = coxosternite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767332" httpUri="https://zenodo.org/record/3767332/files/figure.png" pageId="77" pageNumber="78">Fig. 45I</figureCitation>
|
||
) is also present in
|
||
<taxonomicName authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[521,695,1539,1565]" class="Diplopoda" family="Pachybolidae" genus="Zehntnerobolus" kingdom="Animalia" order="Spirobolida" pageId="77" pageNumber="78" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[521,695,1539,1565]" italics="true" pageId="77" pageNumber="78">Zehntnerobolus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation box="[713,823,1539,1566]" captionStart="Figure 22" captionStartId="38.[140,212,1526,1548]" captionTargetBox="[167,1080,166,1506]" captionTargetId="figure@38.[168,1080,165,1506]" captionTargetPageId="38" captionText="Figure 22. Zehntnerobolus rubripes comb. n., A, C–G male lectotype, B female paratype, A telson B, C right midbody leg D anterior gonopod, anterior view E anterior gonopod, posterior view F left posterior gonopod, anterior view G left posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Cx = coxite; Gr = groove; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite; Tp = telopodite process. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767298" httpUri="https://zenodo.org/record/3767298/files/figure.png" pageId="77" pageNumber="78">Fig. 22G</figureCitation>
|
||
) and
|
||
<taxonomicName authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[893,1044,1539,1565]" class="Diplopoda" family="Pachybolidae" genus="Riotintobolus" kingdom="Animalia" order="Spirobolida" pageId="77" pageNumber="78" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[893,1044,1539,1565]" italics="true" pageId="77" pageNumber="78">Riotintobolus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation captionStart="Figure 26" captionStartId="46.[140,212,1398,1420]" captionTargetBox="[184,1064,165,1374]" captionTargetId="figure@46.[183,1065,165,1100]" captionTargetPageId="46" captionText="Figure 26. Riotintobolus mandenensis sp. n., A, B, I–M male holotype, C–H female paratype. A head, lateral view B telson C head, detail of incisura lateralis D head, frontal view E second leg with vulva F gnathochilarium, overview G gnathochilarium, detail of mentum H left mandible, mesal view I anterior gonopod, anterior view Į left posterior gonopod, anterior view K anterior gonopod, posterior view L telopodite of left posterior gonopod, posterior view M left posterior gonopod, overview, posterior view. 3iT = 3-combed internal tooth; av = anal valves; Co = collum; Cx = coxite; eT = external tooth; Gn = gnathochilarium; Gr = groove; IL = incisura lateralis; L = ledge; LL = lamella linguales; LT = lateral tooth; Md = mandible; Me = mentum; mp = molar plate; pL = pectinate lamella; Pre = preanal ring; sf = setiferous foveolae; St = sternite; ST = stipites; sub = subanal scale; T = telopodite; Vu = vulva. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767304" httpUri="https://zenodo.org/record/3767304/files/figure.png" pageId="77" pageNumber="78">Fig. 26M</figureCitation>
|
||
). A strange similarity can be observed in the posterior gonopods of
|
||
<taxonomicName authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" class="Diplopoda" family="Pachybolidae" genus="Alluviobolus" kingdom="Animalia" order="Spirobolida" pageId="77" pageNumber="78" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="77" pageNumber="78">Alluviobolus</emphasis>
|
||
</taxonomicName>
|
||
with drawings of
|
||
<taxonomicName authority="Attems, 1915" authorityName="Attems" authorityYear="1915" box="[381,788,1610,1637]" class="Diplopoda" family="Trigoniulidae" genus="Trigoniulus" kingdom="Animalia" order="Spirobolida" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species">
|
||
<emphasis box="[381,622,1610,1636]" italics="true" pageId="77" pageNumber="78">Trigoniulus castaneus</emphasis>
|
||
<bibRefCitation author="Attems" box="[629,788,1610,1637]" firstAuthor="Attems" pageId="77" pageNumber="78" pagination="1 - 38" refId="ref56727" refString="Attems C (1915) Myriopoden von Neu Guinea. II. Nova Guinea. Resultats de l'expedition scientifique neerlandaise a la Nouvelle Guinee en 1912 et 1913 sous les auspices de A Fransen 13: 1 - 38." type="journal article" year="1915">Attems, 1915</bibRefCitation>
|
||
</taxonomicName>
|
||
from New
|
||
<collectingCountry box="[921,1008,1610,1636]" name="Guinea-Bissau" pageId="77" pageNumber="78">Guinea</collectingCountry>
|
||
(
|
||
<bibRefCitation author="Attems" firstAuthor="Attems" pageId="77" pageNumber="78" pagination="1 - 38" refId="ref56727" refString="Attems C (1915) Myriopoden von Neu Guinea. II. Nova Guinea. Resultats de l'expedition scientifique neerlandaise a la Nouvelle Guinee en 1912 et 1913 sous les auspices de A Fransen 13: 1 - 38." type="journal article" year="1915">Attems 1915</bibRefCitation>
|
||
: pl. I, fig. 13).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.3820827" ID-Zenodo-Dep="3820827" httpUri="https://zenodo.org/record/3820827/files/figure.png" pageId="78" pageNumber="79" startId="78.[140,212,695,717]" targetBox="[140,1108,165,676]" targetPageId="78">
|
||
<paragraph pageId="78" pageNumber="79">
|
||
<emphasis bold="true" box="[140,253,695,717]" pageId="78" pageNumber="79">Figure 44.</emphasis>
|
||
<emphasis box="[261,468,695,716]" italics="true" pageId="78" pageNumber="79">Alluviobolus laticlavius</emphasis>
|
||
sp. n. male holotype.
|
||
<emphasis bold="true" box="[681,700,695,717]" pageId="78" pageNumber="79">A</emphasis>
|
||
head, lateral view
|
||
<emphasis bold="true" box="[879,896,695,717]" pageId="78" pageNumber="79">B</emphasis>
|
||
legs 1–7, ventral view
|
||
<emphasis bold="true" box="[140,159,727,749]" pageId="78" pageNumber="79">C</emphasis>
|
||
telson
|
||
<emphasis bold="true" box="[226,245,727,749]" pageId="78" pageNumber="79">D</emphasis>
|
||
gnathochilarium, ventral view
|
||
<emphasis bold="true" box="[535,550,727,749]" pageId="78" pageNumber="79">E</emphasis>
|
||
anterior gonopod, anterior view
|
||
<emphasis bold="true" box="[858,872,727,749]" pageId="78" pageNumber="79">F</emphasis>
|
||
right posterior gonopod, anterior view
|
||
<emphasis bold="true" box="[268,288,759,781]" pageId="78" pageNumber="79">G</emphasis>
|
||
anterior gonopod, posterior view
|
||
<emphasis bold="true" box="[608,628,759,781]" pageId="78" pageNumber="79">H</emphasis>
|
||
right posterior gonopod, telopodite, posterior view
|
||
<emphasis bold="true" box="[140,148,791,813]" pageId="78" pageNumber="79">I</emphasis>
|
||
posterior gonopods, coxite and sternite, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; L = ledge; Md = mandible; Me = mentum; Pre = preanal ring; St = sternite; ST = stipites; sub = subanal scale; T = telopodite. Scale bars = 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection pageId="78" pageNumber="79" type="distribution">
|
||
<paragraph blockId="78.[140,1108,941,1672]" pageId="78" pageNumber="79">
|
||
<emphasis bold="true" box="[185,486,941,967]" pageId="78" pageNumber="79">
|
||
<emphasis box="[185,336,941,967]" inLineHeading="true" pageId="78" pageNumber="79" reason="1">Distribution</emphasis>
|
||
and ecology
|
||
</emphasis>
|
||
: species of
|
||
<taxonomicName authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[615,749,941,967]" class="Diplopoda" family="Pachybolidae" genus="Alluviobolus" kingdom="Animalia" order="Spirobolida" pageId="78" pageNumber="79" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[615,749,941,967]" italics="true" pageId="78" pageNumber="79">Alluviobolus</emphasis>
|
||
</taxonomicName>
|
||
were found in spiny, subhumid and humid forests or rainforests in southeast
|
||
<collectingCountry box="[656,789,976,1002]" name="Madagascar" pageId="78" pageNumber="79">Madagascar</collectingCountry>
|
||
(
|
||
<figureCitation box="[805,890,976,1002]" captionStart="Figure 43" captionStartId="77.[139,211,1099,1121]" captionTargetBox="[139,1111,687,1080]" captionTargetId="figure@77.[139,1108,688,1080]" captionTargetPageId="77" captionText="Figure 43. Distribution map of the genus Alluviobolus gen. n." figureDoi="http://doi.org/10.5281/zenodo.3820825" httpUri="https://zenodo.org/record/3820825/files/figure.png" pageId="78" pageNumber="79">Fig. 43</figureCitation>
|
||
). The records from the spiny forest are, however, restricted to the wet alluvial of rivers and small streams, and the same species also occurs in the rainforest of Ebosika.
|
||
<taxonomicName authority="Wesener & Enghoff & Sierwald, 2009" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[821,962,1047,1073]" class="Diplopoda" family="Pachybolidae" genus="Alluviobolus" kingdom="Animalia" order="Spirobolida" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" status="sp. n.">
|
||
<emphasis box="[821,962,1047,1073]" italics="true" pageId="78" pageNumber="79">A. laticlavius</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[968,1029,1047,1073]" pageId="78" pageNumber="79" rank="species">sp. n.</taxonomicNameLabel>
|
||
occurs in the semi-dry forest of Petriky. All species of
|
||
<taxonomicName authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[665,799,1082,1108]" class="Diplopoda" family="Pachybolidae" genus="Alluviobolus" kingdom="Animalia" order="Spirobolida" pageId="78" pageNumber="79" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[665,799,1082,1108]" italics="true" pageId="78" pageNumber="79">Alluviobolus</emphasis>
|
||
</taxonomicName>
|
||
were collected on or inside the first few centimetres of leaf litter in the forests.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="80" lastPageNumber="81" pageId="78" pageNumber="79" type="description">
|
||
<paragraph blockId="78.[140,1108,941,1672]" pageId="78" pageNumber="79">
|
||
<emphasis bold="true" box="[185,335,1152,1178]" pageId="78" pageNumber="79">
|
||
<emphasis box="[185,330,1152,1178]" inLineHeading="true" pageId="78" pageNumber="79" reason="1">Description</emphasis>
|
||
.
|
||
</emphasis>
|
||
<emphasis box="[342,406,1152,1178]" italics="true" pageId="78" pageNumber="79">Males</emphasis>
|
||
: length up to
|
||
<quantity box="[567,653,1152,1179]" metricMagnitude="-2" metricUnit="m" metricValue="4.7" pageId="78" pageNumber="79" unit="mm" value="47.0">47 mm</quantity>
|
||
, diameter up to
|
||
<quantity box="[839,933,1152,1179]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" pageId="78" pageNumber="79" unit="mm" value="4.3">4.3 mm</quantity>
|
||
. 43–49 podous rings. Females: length up to
|
||
<quantity box="[461,546,1187,1214]" metricMagnitude="-2" metricUnit="m" metricValue="6.2" pageId="78" pageNumber="79" unit="mm" value="62.0">62 mm</quantity>
|
||
, diameter up to 6.0 mm. 43–49 podous rings.
|
||
</paragraph>
|
||
<paragraph blockId="78.[140,1108,941,1672]" box="[185,687,1223,1249]" pageId="78" pageNumber="79">Apodous rings absent in all adult specimens.</paragraph>
|
||
<paragraph blockId="78.[140,1108,941,1672]" pageId="78" pageNumber="79">
|
||
<emphasis box="[185,260,1258,1284]" italics="true" pageId="78" pageNumber="79">Colour</emphasis>
|
||
species-specific. Dorsally at least on mesozonites always with a red or orange stripe.
|
||
</paragraph>
|
||
<paragraph blockId="78.[140,1108,941,1672]" pageId="78" pageNumber="79">
|
||
<emphasis box="[185,244,1328,1354]" italics="true" pageId="78" pageNumber="79">Head</emphasis>
|
||
: each eye with circa 22–25 ocelli arranged in 4 or 5 vertical rows (
|
||
<figureCitation box="[998,1103,1328,1354]" captionStart="Figure 44" captionStartId="78.[140,212,695,717]" captionTargetBox="[140,1108,165,676]" captionTargetId="figure@78.[140,1109,165,676]" captionTargetPageId="78" captionText="Figure 44. Alluviobolus laticlavius sp. n. male holotype. A head, lateral view B legs 1–7, ventral view C telson D gnathochilarium, ventral view E anterior gonopod, anterior view F right posterior gonopod, anterior view G anterior gonopod, posterior view H right posterior gonopod, telopodite, posterior view I posterior gonopods, coxite and sternite, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; L = ledge; Md = mandible; Me = mentum; Pre = preanal ring; St = sternite; ST = stipites; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3820827" httpUri="https://zenodo.org/record/3820827/files/figure.png" pageId="78" pageNumber="79">Figs 44A</figureCitation>
|
||
, 45A, 47A). Labrum with standard three irregular teeth and one row of 10–12 stout marginal setae. Clypeus with two setiferous foveolae on each side. Antennae of medium length, reaching back to ring 5 (
|
||
<figureCitation box="[590,696,1434,1460]" captionStart="Figure 44" captionStartId="78.[140,212,695,717]" captionTargetBox="[140,1108,165,676]" captionTargetId="figure@78.[140,1109,165,676]" captionTargetPageId="78" captionText="Figure 44. Alluviobolus laticlavius sp. n. male holotype. A head, lateral view B legs 1–7, ventral view C telson D gnathochilarium, ventral view E anterior gonopod, anterior view F right posterior gonopod, anterior view G anterior gonopod, posterior view H right posterior gonopod, telopodite, posterior view I posterior gonopods, coxite and sternite, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; L = ledge; Md = mandible; Me = mentum; Pre = preanal ring; St = sternite; ST = stipites; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3820827" httpUri="https://zenodo.org/record/3820827/files/figure.png" pageId="78" pageNumber="79">Fig. 44A</figureCitation>
|
||
). Relative lengths of antennomeres: 1<<2=3=4=5<6 (
|
||
<figureCitation box="[334,440,1469,1495]" captionStart="Figure 44" captionStartId="78.[140,212,695,717]" captionTargetBox="[140,1108,165,676]" captionTargetId="figure@78.[140,1109,165,676]" captionTargetPageId="78" captionText="Figure 44. Alluviobolus laticlavius sp. n. male holotype. A head, lateral view B legs 1–7, ventral view C telson D gnathochilarium, ventral view E anterior gonopod, anterior view F right posterior gonopod, anterior view G anterior gonopod, posterior view H right posterior gonopod, telopodite, posterior view I posterior gonopods, coxite and sternite, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; L = ledge; Md = mandible; Me = mentum; Pre = preanal ring; St = sternite; ST = stipites; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3820827" httpUri="https://zenodo.org/record/3820827/files/figure.png" pageId="78" pageNumber="79">Fig. 44A</figureCitation>
|
||
). Terminal antennomere with four large sensory cones located together inside a membranous area. Antennomere 5 latero-apically with four rows, antennomere 6 with two rows of sensilla basiconica.
|
||
</paragraph>
|
||
<paragraph blockId="78.[140,1108,941,1672]" lastBlockId="80.[140,1108,167,1566]" lastPageId="80" lastPageNumber="81" pageId="78" pageNumber="79">
|
||
<emphasis box="[185,373,1575,1601]" italics="true" pageId="78" pageNumber="79">Gnathochilarium</emphasis>
|
||
unusual (
|
||
<figureCitation box="[489,599,1575,1601]" captionStart="Figure 44" captionStartId="78.[140,212,695,717]" captionTargetBox="[140,1108,165,676]" captionTargetId="figure@78.[140,1109,165,676]" captionTargetPageId="78" captionText="Figure 44. Alluviobolus laticlavius sp. n. male holotype. A head, lateral view B legs 1–7, ventral view C telson D gnathochilarium, ventral view E anterior gonopod, anterior view F right posterior gonopod, anterior view G anterior gonopod, posterior view H right posterior gonopod, telopodite, posterior view I posterior gonopods, coxite and sternite, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; L = ledge; Md = mandible; Me = mentum; Pre = preanal ring; St = sternite; ST = stipites; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3820827" httpUri="https://zenodo.org/record/3820827/files/figure.png" pageId="78" pageNumber="79">Fig. 44D</figureCitation>
|
||
). Lamellae linguales each with two standard setae located behind one another. Stipites each with three apical setae. Mentum basally subdivided by a well-developed suture (
|
||
<figureCitation box="[594,740,1645,1672]" captionStart="Figure 46" captionStartId="83.[140,212,1014,1036]" captionTargetBox="[140,1108,165,990]" captionTargetId="figure@83.[140,1108,165,667]" captionTargetPageId="83" captionText="Figure 46. Alluviobolus tsimelahy sp. n. specimens from Ebosika, A–G, I–M male, H female. A head, lateral view B legs 1–7, ventral view C telson D gnathochilarium, ventral view E gnathochilarium, mentum F gnathochilarium, dorsal view on endochilarium G, H left midbody legs I anterior gonopod, anterior view Į right posterior gonopod, anterior view K anterior gonopod, posterior view L right posterior gonopod, telopodite, posterior view M right posterior gonopods, coxite and sternite, posterior view. Apo = apodeme; av = anal valves; Co = collum; cp = central pads; Cx = coxite; Endo = endochilarium; Gn = gnathochilarium; L = ledge; Md = mandible; Me = mentum; Pre = preanal ring; St = sternite; ST = stipites; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767334" httpUri="https://zenodo.org/record/3767334/files/figure.png" pageId="78" pageNumber="79">Figs 46D, E</figureCitation>
|
||
). Stipites each towards mentum with a wide sclerotized ledge (
|
||
<figureCitation box="[499,605,167,194]" captionStart="Figure 46" captionStartId="83.[140,212,1014,1036]" captionTargetBox="[140,1108,165,990]" captionTargetId="figure@83.[140,1108,165,667]" captionTargetPageId="83" captionText="Figure 46. Alluviobolus tsimelahy sp. n. specimens from Ebosika, A–G, I–M male, H female. A head, lateral view B legs 1–7, ventral view C telson D gnathochilarium, ventral view E gnathochilarium, mentum F gnathochilarium, dorsal view on endochilarium G, H left midbody legs I anterior gonopod, anterior view Į right posterior gonopod, anterior view K anterior gonopod, posterior view L right posterior gonopod, telopodite, posterior view M right posterior gonopods, coxite and sternite, posterior view. Apo = apodeme; av = anal valves; Co = collum; cp = central pads; Cx = coxite; Endo = endochilarium; Gn = gnathochilarium; L = ledge; Md = mandible; Me = mentum; Pre = preanal ring; St = sternite; ST = stipites; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767334" httpUri="https://zenodo.org/record/3767334/files/figure.png" pageId="80" pageNumber="81">Fig. 46E</figureCitation>
|
||
). Palpi of gnathochilarium with numerous sensilla. Hypopharyngeal crest with a field of spine-like structures (
|
||
<figureCitation box="[884,984,202,229]" captionStart="Figure 46" captionStartId="83.[140,212,1014,1036]" captionTargetBox="[140,1108,165,990]" captionTargetId="figure@83.[140,1108,165,667]" captionTargetPageId="83" captionText="Figure 46. Alluviobolus tsimelahy sp. n. specimens from Ebosika, A–G, I–M male, H female. A head, lateral view B legs 1–7, ventral view C telson D gnathochilarium, ventral view E gnathochilarium, mentum F gnathochilarium, dorsal view on endochilarium G, H left midbody legs I anterior gonopod, anterior view Į right posterior gonopod, anterior view K anterior gonopod, posterior view L right posterior gonopod, telopodite, posterior view M right posterior gonopods, coxite and sternite, posterior view. Apo = apodeme; av = anal valves; Co = collum; cp = central pads; Cx = coxite; Endo = endochilarium; Gn = gnathochilarium; L = ledge; Md = mandible; Me = mentum; Pre = preanal ring; St = sternite; ST = stipites; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767334" httpUri="https://zenodo.org/record/3767334/files/figure.png" pageId="80" pageNumber="81">Fig. 46F</figureCitation>
|
||
). Endochilarium apically with 4–6, towards hypopharyngeal crest with>20 sensilla.
|
||
</paragraph>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.3767332" ID-Zenodo-Dep="3767332" httpUri="https://zenodo.org/record/3767332/files/figure.png" pageId="79" pageNumber="80" startId="79.[140,212,1462,1484]" targetBox="[140,1108,165,1295]" targetPageId="79">
|
||
<paragraph blockId="79.[140,1108,1462,1676]" pageId="79" pageNumber="80">
|
||
<emphasis bold="true" box="[140,251,1462,1484]" pageId="79" pageNumber="80">Figure 45.</emphasis>
|
||
<taxonomicName authority="Wesener & Enghoff & Sierwald, 2009" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[258,456,1462,1483]" class="Diplopoda" family="Pachybolidae" genus="Alluviobolus" kingdom="Animalia" order="Spirobolida" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" status="sp. n.">
|
||
<emphasis box="[258,456,1462,1483]" italics="true" pageId="79" pageNumber="80">Alluviobolus tsimelahy</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[463,514,1463,1484]" pageId="79" pageNumber="80" rank="species">sp. n.</taxonomicNameLabel>
|
||
A–C, F–I male holotype, D, E female paratype.
|
||
<emphasis bold="true" box="[970,989,1462,1484]" pageId="79" pageNumber="80">A</emphasis>
|
||
head, lateral view
|
||
<emphasis bold="true" box="[187,204,1494,1516]" pageId="79" pageNumber="80">B</emphasis>
|
||
legs 1–7, ventral view
|
||
<emphasis bold="true" box="[412,431,1494,1516]" pageId="79" pageNumber="80">C</emphasis>
|
||
telson
|
||
<emphasis bold="true" box="[496,515,1494,1516]" pageId="79" pageNumber="80">D</emphasis>
|
||
vulva and sterno-coxite of second leg, posterior view
|
||
<emphasis bold="true" box="[1003,1018,1494,1516]" pageId="79" pageNumber="80">E</emphasis>
|
||
left mandible, mesal view
|
||
<emphasis bold="true" box="[303,317,1526,1548]" pageId="79" pageNumber="80">F</emphasis>
|
||
anterior gonopod, anterior view
|
||
<emphasis bold="true" box="[623,643,1526,1548]" pageId="79" pageNumber="80">G</emphasis>
|
||
right posterior gonopod, anterior view
|
||
<emphasis bold="true" box="[1009,1029,1526,1548]" pageId="79" pageNumber="80">H</emphasis>
|
||
anterior gonopod, posterior view
|
||
<emphasis bold="true" box="[373,381,1558,1580]" pageId="79" pageNumber="80">I</emphasis>
|
||
right posterior gonopod, posterior view. 3iT = 3-combed internal tooth; Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; eT = external tooth; Gn = gnathochilarium; LT = lateral tooth; Md = mandible; mp = molar plate; O = operculum; pL = pectinate lamella; Pre = preanal ring; pv = posterior valve; St = sternite; St-Cx = coxosternite; T = telopodite. Scale bars = 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="80.[140,1108,167,1566]" pageId="80" pageNumber="81">
|
||
<emphasis box="[185,291,272,298]" italics="true" pageId="80" pageNumber="81">Mandible</emphasis>
|
||
basal joints swollen (
|
||
<figureCitation box="[540,645,272,299]" captionStart="Figure 45" captionStartId="79.[140,212,1462,1484]" captionTargetBox="[140,1108,165,1295]" captionTargetId="figure@79.[140,1108,165,887]" captionTargetPageId="79" captionText="Figure 45. Alluviobolus tsimelahy sp. n. A–C, F–I male holotype, D, E female paratype. A head, lateral view B legs 1–7, ventral view C telson D vulva and sterno-coxite of second leg, posterior view E left mandible, mesal view F anterior gonopod, anterior view G right posterior gonopod, anterior view H anterior gonopod, posterior view I right posterior gonopod, posterior view. 3iT = 3-combed internal tooth; Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; eT = external tooth; Gn = gnathochilarium; LT = lateral tooth; Md = mandible; mp = molar plate; O = operculum; pL = pectinate lamella; Pre = preanal ring; pv = posterior valve; St = sternite; St-Cx = coxosternite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767332" httpUri="https://zenodo.org/record/3767332/files/figure.png" pageId="80" pageNumber="81">Fig. 45A</figureCitation>
|
||
). External tooth simple, rounded; mesal tooth with three cusps (
|
||
<figureCitation box="[403,505,307,334]" captionStart="Figure 45" captionStartId="79.[140,212,1462,1484]" captionTargetBox="[140,1108,165,1295]" captionTargetId="figure@79.[140,1108,165,887]" captionTargetPageId="79" captionText="Figure 45. Alluviobolus tsimelahy sp. n. A–C, F–I male holotype, D, E female paratype. A head, lateral view B legs 1–7, ventral view C telson D vulva and sterno-coxite of second leg, posterior view E left mandible, mesal view F anterior gonopod, anterior view G right posterior gonopod, anterior view H anterior gonopod, posterior view I right posterior gonopod, posterior view. 3iT = 3-combed internal tooth; Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; eT = external tooth; Gn = gnathochilarium; LT = lateral tooth; Md = mandible; mp = molar plate; O = operculum; pL = pectinate lamella; Pre = preanal ring; pv = posterior valve; St = sternite; St-Cx = coxosternite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767332" httpUri="https://zenodo.org/record/3767332/files/figure.png" pageId="80" pageNumber="81">Fig. 45E</figureCitation>
|
||
). Four or five pectinate lamellae. Molar plate with few (circa 5) transverse furrows, anterior three furrows greatly enlarged, posterior furrows minute (
|
||
<figureCitation box="[237,339,378,405]" captionStart="Figure 45" captionStartId="79.[140,212,1462,1484]" captionTargetBox="[140,1108,165,1295]" captionTargetId="figure@79.[140,1108,165,887]" captionTargetPageId="79" captionText="Figure 45. Alluviobolus tsimelahy sp. n. A–C, F–I male holotype, D, E female paratype. A head, lateral view B legs 1–7, ventral view C telson D vulva and sterno-coxite of second leg, posterior view E left mandible, mesal view F anterior gonopod, anterior view G right posterior gonopod, anterior view H anterior gonopod, posterior view I right posterior gonopod, posterior view. 3iT = 3-combed internal tooth; Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; eT = external tooth; Gn = gnathochilarium; LT = lateral tooth; Md = mandible; mp = molar plate; O = operculum; pL = pectinate lamella; Pre = preanal ring; pv = posterior valve; St = sternite; St-Cx = coxosternite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767332" httpUri="https://zenodo.org/record/3767332/files/figure.png" pageId="80" pageNumber="81">Fig. 45E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="80.[140,1108,167,1566]" box="[185,1108,413,440]" pageId="80" pageNumber="81">
|
||
<emphasis box="[185,274,413,439]" italics="true" pageId="80" pageNumber="81">Collum:</emphasis>
|
||
smooth, inconspicuous. Ventrally not protruding as far as ring 2 (
|
||
<figureCitation box="[994,1095,413,440]" captionStart="Figure 45" captionStartId="79.[140,212,1462,1484]" captionTargetBox="[140,1108,165,1295]" captionTargetId="figure@79.[140,1108,165,887]" captionTargetPageId="79" captionText="Figure 45. Alluviobolus tsimelahy sp. n. A–C, F–I male holotype, D, E female paratype. A head, lateral view B legs 1–7, ventral view C telson D vulva and sterno-coxite of second leg, posterior view E left mandible, mesal view F anterior gonopod, anterior view G right posterior gonopod, anterior view H anterior gonopod, posterior view I right posterior gonopod, posterior view. 3iT = 3-combed internal tooth; Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; eT = external tooth; Gn = gnathochilarium; LT = lateral tooth; Md = mandible; mp = molar plate; O = operculum; pL = pectinate lamella; Pre = preanal ring; pv = posterior valve; St = sternite; St-Cx = coxosternite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767332" httpUri="https://zenodo.org/record/3767332/files/figure.png" pageId="80" pageNumber="81">Fig. 45A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="80.[140,1108,167,1566]" pageId="80" pageNumber="81">
|
||
<emphasis box="[185,307,448,474]" italics="true" pageId="80" pageNumber="81">Body rings:</emphasis>
|
||
dorsally and laterally leather-like, meso- and metazona ventrally with numerous transverse impressions. Ozopores starting at ring 6, touching suture between mesozona and metazona (
|
||
<figureCitation box="[504,642,519,545]" captionStart="Figure 42" captionStartId="77.[139,211,599,621]" captionTargetBox="[139,1107,165,580]" captionTargetId="figure@77.[139,1108,165,580]" captionTargetPageId="77" captionText="Figure 42. Alluviobolus tsimelahy sp. n., A living male from Tsimelahy B male and female specimen from Ebosika." figureDoi="http://doi.org/10.5281/zenodo.3767330" httpUri="https://zenodo.org/record/3767330/files/figure.png" pageId="80" pageNumber="81">Figs 42A, B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="80.[140,1108,167,1566]" pageId="80" pageNumber="81">
|
||
<emphasis box="[185,251,554,580]" italics="true" pageId="80" pageNumber="81">Telson</emphasis>
|
||
: preanal process absent. Anal valves with lips and micropunctation, slightly elongated posteriorly. Subanal scale inconspicuous (
|
||
<figureCitation box="[722,828,589,615]" captionStart="Figure 44" captionStartId="78.[140,212,695,717]" captionTargetBox="[140,1108,165,676]" captionTargetId="figure@78.[140,1109,165,676]" captionTargetPageId="78" captionText="Figure 44. Alluviobolus laticlavius sp. n. male holotype. A head, lateral view B legs 1–7, ventral view C telson D gnathochilarium, ventral view E anterior gonopod, anterior view F right posterior gonopod, anterior view G anterior gonopod, posterior view H right posterior gonopod, telopodite, posterior view I posterior gonopods, coxite and sternite, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; L = ledge; Md = mandible; Me = mentum; Pre = preanal ring; St = sternite; ST = stipites; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3820827" httpUri="https://zenodo.org/record/3820827/files/figure.png" pageId="80" pageNumber="81">Fig. 44C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="80.[140,1108,167,1566]" pageId="80" pageNumber="81">
|
||
<emphasis box="[185,231,625,650]" italics="true" pageId="80" pageNumber="81">Legs</emphasis>
|
||
: coxae 1 and 2 elongated and fused with sternum, podomeres from prefemur to tarsus in both sexes each with 4–10 ventral/mesal setae. Length of midbody legs circa 1.1 times body diameter in males and 0.7 times body diameter in females. Each podomere with an apical, ventral seta (
|
||
<figureCitation box="[658,809,730,756]" captionStart="Figure 46" captionStartId="83.[140,212,1014,1036]" captionTargetBox="[140,1108,165,990]" captionTargetId="figure@83.[140,1108,165,667]" captionTargetPageId="83" captionText="Figure 46. Alluviobolus tsimelahy sp. n. specimens from Ebosika, A–G, I–M male, H female. A head, lateral view B legs 1–7, ventral view C telson D gnathochilarium, ventral view E gnathochilarium, mentum F gnathochilarium, dorsal view on endochilarium G, H left midbody legs I anterior gonopod, anterior view Į right posterior gonopod, anterior view K anterior gonopod, posterior view L right posterior gonopod, telopodite, posterior view M right posterior gonopods, coxite and sternite, posterior view. Apo = apodeme; av = anal valves; Co = collum; cp = central pads; Cx = coxite; Endo = endochilarium; Gn = gnathochilarium; L = ledge; Md = mandible; Me = mentum; Pre = preanal ring; St = sternite; ST = stipites; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767334" httpUri="https://zenodo.org/record/3767334/files/figure.png" pageId="80" pageNumber="81">Figs 46G, H</figureCitation>
|
||
). Coxae 3 and beyond of cylindrical shape (
|
||
<figureCitation box="[346,494,765,792]" captionStart="Figure 46" captionStartId="83.[140,212,1014,1036]" captionTargetBox="[140,1108,165,990]" captionTargetId="figure@83.[140,1108,165,667]" captionTargetPageId="83" captionText="Figure 46. Alluviobolus tsimelahy sp. n. specimens from Ebosika, A–G, I–M male, H female. A head, lateral view B legs 1–7, ventral view C telson D gnathochilarium, ventral view E gnathochilarium, mentum F gnathochilarium, dorsal view on endochilarium G, H left midbody legs I anterior gonopod, anterior view Į right posterior gonopod, anterior view K anterior gonopod, posterior view L right posterior gonopod, telopodite, posterior view M right posterior gonopods, coxite and sternite, posterior view. Apo = apodeme; av = anal valves; Co = collum; cp = central pads; Cx = coxite; Endo = endochilarium; Gn = gnathochilarium; L = ledge; Md = mandible; Me = mentum; Pre = preanal ring; St = sternite; ST = stipites; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767334" httpUri="https://zenodo.org/record/3767334/files/figure.png" pageId="80" pageNumber="81">Figs 46G, H</figureCitation>
|
||
). Tarsus with a stout dorso-apical seta and three pairs of ventral setae.
|
||
</paragraph>
|
||
<paragraph blockId="80.[140,1108,167,1566]" pageId="80" pageNumber="81">
|
||
<emphasis box="[185,425,835,861]" italics="true" pageId="80" pageNumber="81">Male sexual characters</emphasis>
|
||
: coxae
|
||
<quantity box="[509,583,835,862]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" metricValueMax="1.778" metricValueMin="0.762" pageId="80" pageNumber="81" unit="in" value="5.0" valueMax="7.0" valueMin="3.0">3–7 in</quantity>
|
||
some species modified (
|
||
<figureCitation box="[856,960,835,862]" captionStart="Figure 44" captionStartId="78.[140,212,695,717]" captionTargetBox="[140,1108,165,676]" captionTargetId="figure@78.[140,1109,165,676]" captionTargetPageId="78" captionText="Figure 44. Alluviobolus laticlavius sp. n. male holotype. A head, lateral view B legs 1–7, ventral view C telson D gnathochilarium, ventral view E anterior gonopod, anterior view F right posterior gonopod, anterior view G anterior gonopod, posterior view H right posterior gonopod, telopodite, posterior view I posterior gonopods, coxite and sternite, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; L = ledge; Md = mandible; Me = mentum; Pre = preanal ring; St = sternite; ST = stipites; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3820827" httpUri="https://zenodo.org/record/3820827/files/figure.png" pageId="80" pageNumber="81">Figs 44B</figureCitation>
|
||
, 45B). Tarsal pads absent.
|
||
</paragraph>
|
||
<paragraph blockId="80.[140,1108,167,1566]" pageId="80" pageNumber="81">
|
||
<emphasis box="[185,379,906,932]" italics="true" pageId="80" pageNumber="81">Anterior gonopods</emphasis>
|
||
: median sternal projection triangular and elongated with a wide tip (
|
||
<figureCitation box="[187,289,941,967]" captionStart="Figure 44" captionStartId="78.[140,212,695,717]" captionTargetBox="[140,1108,165,676]" captionTargetId="figure@78.[140,1109,165,676]" captionTargetPageId="78" captionText="Figure 44. Alluviobolus laticlavius sp. n. male holotype. A head, lateral view B legs 1–7, ventral view C telson D gnathochilarium, ventral view E anterior gonopod, anterior view F right posterior gonopod, anterior view G anterior gonopod, posterior view H right posterior gonopod, telopodite, posterior view I posterior gonopods, coxite and sternite, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; L = ledge; Md = mandible; Me = mentum; Pre = preanal ring; St = sternite; ST = stipites; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3820827" httpUri="https://zenodo.org/record/3820827/files/figure.png" pageId="80" pageNumber="81">Fig. 44E</figureCitation>
|
||
). Sternite always longer than coxite and telopodite, except for telopodite process. Coxite process absent. Coxite in anterior view well-visible, in posterior view only laterally present (
|
||
<figureCitation box="[393,500,1011,1038]" captionStart="Figure 44" captionStartId="78.[140,212,695,717]" captionTargetBox="[140,1108,165,676]" captionTargetId="figure@78.[140,1109,165,676]" captionTargetPageId="78" captionText="Figure 44. Alluviobolus laticlavius sp. n. male holotype. A head, lateral view B legs 1–7, ventral view C telson D gnathochilarium, ventral view E anterior gonopod, anterior view F right posterior gonopod, anterior view G anterior gonopod, posterior view H right posterior gonopod, telopodite, posterior view I posterior gonopods, coxite and sternite, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; L = ledge; Md = mandible; Me = mentum; Pre = preanal ring; St = sternite; ST = stipites; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3820827" httpUri="https://zenodo.org/record/3820827/files/figure.png" pageId="80" pageNumber="81">Fig. 44G</figureCitation>
|
||
). Telopodite mesally always with a long slender process. Tip with a sharp-edged, retrorse process (
|
||
<figureCitation box="[649,756,1047,1073]" captionStart="Figure 44" captionStartId="78.[140,212,695,717]" captionTargetBox="[140,1108,165,676]" captionTargetId="figure@78.[140,1109,165,676]" captionTargetPageId="78" captionText="Figure 44. Alluviobolus laticlavius sp. n. male holotype. A head, lateral view B legs 1–7, ventral view C telson D gnathochilarium, ventral view E anterior gonopod, anterior view F right posterior gonopod, anterior view G anterior gonopod, posterior view H right posterior gonopod, telopodite, posterior view I posterior gonopods, coxite and sternite, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; L = ledge; Md = mandible; Me = mentum; Pre = preanal ring; St = sternite; ST = stipites; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3820827" httpUri="https://zenodo.org/record/3820827/files/figure.png" pageId="80" pageNumber="81">Fig. 44G</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="80.[140,1108,167,1566]" pageId="80" pageNumber="81">
|
||
<emphasis box="[185,469,1082,1108]" italics="true" pageId="80" pageNumber="81">Posterior gonopods in situ</emphasis>
|
||
almost completely covered by coxite and telopodite of anterior gonopod. Coxite and telopodite of posterior gonopod clearly divided (
|
||
<figureCitation captionStart="Figure 45" captionStartId="79.[140,212,1462,1484]" captionTargetBox="[140,1108,165,1295]" captionTargetId="figure@79.[140,1108,165,887]" captionTargetPageId="79" captionText="Figure 45. Alluviobolus tsimelahy sp. n. A–C, F–I male holotype, D, E female paratype. A head, lateral view B legs 1–7, ventral view C telson D vulva and sterno-coxite of second leg, posterior view E left mandible, mesal view F anterior gonopod, anterior view G right posterior gonopod, anterior view H anterior gonopod, posterior view I right posterior gonopod, posterior view. 3iT = 3-combed internal tooth; Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; eT = external tooth; Gn = gnathochilarium; LT = lateral tooth; Md = mandible; mp = molar plate; O = operculum; pL = pectinate lamella; Pre = preanal ring; pv = posterior valve; St = sternite; St-Cx = coxosternite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767332" httpUri="https://zenodo.org/record/3767332/files/figure.png" pageId="80" pageNumber="81">Figs 45I</figureCitation>
|
||
,
|
||
<figureCitation box="[196,253,1152,1178]" captionStart="Figure 47" captionStartId="85.[140,212,825,847]" captionTargetBox="[140,1108,165,794]" captionTargetId="figure@85.[140,1108,465,804]" captionTargetPageId="85" captionText="Figure 47. Alluviobolus antanosy sp. n., male holotype. A head, lateral view B legs 1–7, ventral view C telson D anterior gonopod, anterior view E left posterior gonopod, anterior view F anterior gonopod, posterior view G left posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; IL = incisura lateralis; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767336" httpUri="https://zenodo.org/record/3767336/files/figure.png" pageId="80" pageNumber="81">47G</figureCitation>
|
||
). Sternite sclerotized and well-visible (
|
||
<figureCitation box="[698,797,1152,1178]" captionStart="Figure 44" captionStartId="78.[140,212,695,717]" captionTargetBox="[140,1108,165,676]" captionTargetId="figure@78.[140,1109,165,676]" captionTargetPageId="78" captionText="Figure 44. Alluviobolus laticlavius sp. n. male holotype. A head, lateral view B legs 1–7, ventral view C telson D gnathochilarium, ventral view E anterior gonopod, anterior view F right posterior gonopod, anterior view G anterior gonopod, posterior view H right posterior gonopod, telopodite, posterior view I posterior gonopods, coxite and sternite, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; L = ledge; Md = mandible; Me = mentum; Pre = preanal ring; St = sternite; ST = stipites; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3820827" httpUri="https://zenodo.org/record/3820827/files/figure.png" pageId="80" pageNumber="81">Figs 44I</figureCitation>
|
||
, 45I). Coxite mesally with a single groove. Joint protruding into a short stem towards telopodite, latter as long and wide as coxite, curved posteriorly (
|
||
<figureCitation box="[610,711,1222,1249]" captionStart="Figure 45" captionStartId="79.[140,212,1462,1484]" captionTargetBox="[140,1108,165,1295]" captionTargetId="figure@79.[140,1108,165,887]" captionTargetPageId="79" captionText="Figure 45. Alluviobolus tsimelahy sp. n. A–C, F–I male holotype, D, E female paratype. A head, lateral view B legs 1–7, ventral view C telson D vulva and sterno-coxite of second leg, posterior view E left mandible, mesal view F anterior gonopod, anterior view G right posterior gonopod, anterior view H anterior gonopod, posterior view I right posterior gonopod, posterior view. 3iT = 3-combed internal tooth; Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; eT = external tooth; Gn = gnathochilarium; LT = lateral tooth; Md = mandible; mp = molar plate; O = operculum; pL = pectinate lamella; Pre = preanal ring; pv = posterior valve; St = sternite; St-Cx = coxosternite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767332" httpUri="https://zenodo.org/record/3767332/files/figure.png" pageId="80" pageNumber="81">Fig. 45I</figureCitation>
|
||
). Telopodites arranged parallel to one another. A torsion present in telopodites, sperm canal discharging apico-laterally (
|
||
<figureCitation box="[148,261,1293,1319]" captionStart="Figure 44" captionStartId="78.[140,212,695,717]" captionTargetBox="[140,1108,165,676]" captionTargetId="figure@78.[140,1109,165,676]" captionTargetPageId="78" captionText="Figure 44. Alluviobolus laticlavius sp. n. male holotype. A head, lateral view B legs 1–7, ventral view C telson D gnathochilarium, ventral view E anterior gonopod, anterior view F right posterior gonopod, anterior view G anterior gonopod, posterior view H right posterior gonopod, telopodite, posterior view I posterior gonopods, coxite and sternite, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; L = ledge; Md = mandible; Me = mentum; Pre = preanal ring; St = sternite; ST = stipites; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3820827" httpUri="https://zenodo.org/record/3820827/files/figure.png" pageId="80" pageNumber="81">Fig. 44H</figureCitation>
|
||
). Tip of telopodite mesally with a sclerite of species-specific shape (
|
||
<emphasis box="[1062,1074,1294,1319]" italics="true" pageId="80" pageNumber="81">x</emphasis>
|
||
in
|
||
<figureCitation box="[140,251,1328,1354]" captionStart="Figure 44" captionStartId="78.[140,212,695,717]" captionTargetBox="[140,1108,165,676]" captionTargetId="figure@78.[140,1109,165,676]" captionTargetPageId="78" captionText="Figure 44. Alluviobolus laticlavius sp. n. male holotype. A head, lateral view B legs 1–7, ventral view C telson D gnathochilarium, ventral view E anterior gonopod, anterior view F right posterior gonopod, anterior view G anterior gonopod, posterior view H right posterior gonopod, telopodite, posterior view I posterior gonopods, coxite and sternite, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; L = ledge; Md = mandible; Me = mentum; Pre = preanal ring; St = sternite; ST = stipites; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3820827" httpUri="https://zenodo.org/record/3820827/files/figure.png" pageId="80" pageNumber="81">Figs 44H</figureCitation>
|
||
, 45I, 47G).
|
||
</paragraph>
|
||
<paragraph blockId="80.[140,1108,167,1566]" pageId="80" pageNumber="81">
|
||
<emphasis box="[185,454,1363,1389]" italics="true" pageId="80" pageNumber="81">Female sexual characters:</emphasis>
|
||
vulva simple, bivalve-like, with a small, poorly sclerotized operculum at base (
|
||
<figureCitation box="[358,465,1398,1425]" captionStart="Figure 45" captionStartId="79.[140,212,1462,1484]" captionTargetBox="[140,1108,165,1295]" captionTargetId="figure@79.[140,1108,165,887]" captionTargetPageId="79" captionText="Figure 45. Alluviobolus tsimelahy sp. n. A–C, F–I male holotype, D, E female paratype. A head, lateral view B legs 1–7, ventral view C telson D vulva and sterno-coxite of second leg, posterior view E left mandible, mesal view F anterior gonopod, anterior view G right posterior gonopod, anterior view H anterior gonopod, posterior view I right posterior gonopod, posterior view. 3iT = 3-combed internal tooth; Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; eT = external tooth; Gn = gnathochilarium; LT = lateral tooth; Md = mandible; mp = molar plate; O = operculum; pL = pectinate lamella; Pre = preanal ring; pv = posterior valve; St = sternite; St-Cx = coxosternite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767332" httpUri="https://zenodo.org/record/3767332/files/figure.png" pageId="80" pageNumber="81">Fig. 45D</figureCitation>
|
||
). Posterior valve apically overlapping part of anterior one. Basally on each valve towards opening with two or three rows of setae (
|
||
<figureCitation box="[940,1048,1434,1461]" captionStart="Figure 45" captionStartId="79.[140,212,1462,1484]" captionTargetBox="[140,1108,165,1295]" captionTargetId="figure@79.[140,1108,165,887]" captionTargetPageId="79" captionText="Figure 45. Alluviobolus tsimelahy sp. n. A–C, F–I male holotype, D, E female paratype. A head, lateral view B legs 1–7, ventral view C telson D vulva and sterno-coxite of second leg, posterior view E left mandible, mesal view F anterior gonopod, anterior view G right posterior gonopod, anterior view H anterior gonopod, posterior view I right posterior gonopod, posterior view. 3iT = 3-combed internal tooth; Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; eT = external tooth; Gn = gnathochilarium; LT = lateral tooth; Md = mandible; mp = molar plate; O = operculum; pL = pectinate lamella; Pre = preanal ring; pv = posterior valve; St = sternite; St-Cx = coxosternite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767332" httpUri="https://zenodo.org/record/3767332/files/figure.png" pageId="80" pageNumber="81">Fig. 45D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="80" pageNumber="81" type="etymology">
|
||
<paragraph blockId="80.[140,1108,167,1566]" pageId="80" pageNumber="81">
|
||
<emphasis bold="true" box="[185,319,1469,1495]" inLineHeading="true" pageId="80" pageNumber="81" reason="1">Etymology</emphasis>
|
||
:
|
||
<taxonomicName authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[338,477,1469,1495]" class="Diplopoda" family="Pachybolidae" genus="Alluviobolus" kingdom="Animalia" order="Spirobolida" pageId="80" pageNumber="81" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[338,477,1469,1495]" italics="true" pageId="80" pageNumber="81">Alluviobolus</emphasis>
|
||
</taxonomicName>
|
||
, masculine, Latin, consists of Alluvium referring to the alluvial of a river where large numbers of
|
||
<taxonomicName authority="Wesener & Enghoff & Sierwald, 2009" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[671,810,1504,1531]" class="Diplopoda" family="Pachybolidae" genus="Alluviobolus" kingdom="Animalia" order="Spirobolida" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" status="sp. n.">
|
||
<emphasis box="[671,810,1504,1531]" italics="true" pageId="80" pageNumber="81">A. tsimelahy</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[817,881,1505,1531]" pageId="80" pageNumber="81" rank="species">sp. n.</taxonomicNameLabel>
|
||
could be collected, and
|
||
<emphasis box="[191,258,1539,1565]" italics="true" pageId="80" pageNumber="81">-bolus</emphasis>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</subSection>
|
||
</treatment>
|
||
</document> |