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<document id="7C5650407CB02F04EE78F8B7BA73FC28" ID-CLB-Dataset="58772" ID-DOI="10.11646/zootaxa.5244.3.3" ID-GBIF-Dataset="3a2ab5b9-fb66-4c1f-802e-cf519c36f6b8" ID-ISSN="1175-5326" ID-Zenodo-Dep="7656289" ID-ZooBank="73D7B5AC-32F8-4C7D-85CD-960816BB22EA" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1676882860034" checkinUser="plazi" docAuthor="Santos-Silva, Antonio &amp; Monné, Miguel A." docDate="2023" docId="1754AB31FF978528FF7A4650FEE91467" docLanguage="en" docName="zootaxa.5244.3.3.pdf" docOrigin="Zootaxa 5244 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5244.3.3" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Estola nearnsi Santos-Silva &amp; Monné 2023, sp. nov." docType="treatment" docVersion="9" lastPageNumber="259" masterDocId="EB6DD349FF9B8527FFED4217FFD7175F" masterDocTitle="The incredible biodiversity of American Lamiinae (Coleoptera, Cerambycidae) description of four new species, synonymy, and taxonomical notes" masterLastPageNumber="260" masterPageNumber="244" pageNumber="256" updateTime="1699475472773" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="061C9C012D0E9B6E668D1B23F2459CCE">The incredible biodiversity of American Lamiinae (Coleoptera, Cerambycidae) description of four new species, synonymy, and taxonomical notes</mods:title>
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<mods:namePart id="7440FF4EEBCAACAF772C0B42F6EB0E2A">Santos-Silva, Antonio</mods:namePart>
<mods:affiliation id="5A0443243725EA79B56B6ED9864AD764">Museu de Zoologia, Universidade de São Paulo, São Paulo, SP, Brazil.</mods:affiliation>
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<mods:namePart id="04333BEA9DA636B1E6F2A7B94080D2A9">Monné, Miguel A.</mods:namePart>
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<mods:affiliation id="F8B29AA75D96EE8DC6EDD1E58586C38E">Museu Nacional, Universidade Federal do Rio de Janeiro, Quinta da Boa Vista, São Cristóvão, 20940.040, Rio de Janeiro, RJ, Brazil. monne @ uol. com. br; https: // orcid. org / 0000 - 0001 - 8825 - 3122</mods:affiliation>
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<paragraph id="9F421A27FF97852BFF7A4650FE7C133D" blockId="12.[151,427,1095,1158]" box="[151,427,1095,1122]" pageId="12" pageNumber="256">
<heading id="C40AAD4BFF97852BFF7A4650FE7C133D" bold="true" box="[151,427,1095,1122]" fontSize="11" level="1" pageId="12" pageNumber="256" reason="1">
<emphasis id="AD89C635FF97852BFF7A4650FE7C133D" bold="true" box="[151,427,1095,1122]" pageId="12" pageNumber="256">
<taxonomicName id="58FD61A4FF97852BFF7A4650FE91133E" ID-CoL="BRHB3" authority="Santos-Silva &amp; Monné, 2023" authorityName="Santos-Silva &amp; Monné" authorityYear="2023" box="[151,326,1095,1122]" class="Insecta" family="Cerambycidae" genus="Estola" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="256" phylum="Arthropoda" rank="species" species="nearnsi" status="sp. nov.">
<emphasis id="AD89C635FF97852BFF7A4650FE91133E" bold="true" box="[151,326,1095,1122]" italics="true" pageId="12" pageNumber="256">Estola nearnsi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B6BA7B4EFF97852BFEA3465FFE7C133D" box="[334,427,1096,1122]" pageId="12" pageNumber="256" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="D7E749ACFF978529FF7A467CFD031021" lastPageId="14" lastPageNumber="258" pageId="12" pageNumber="256" type="description">
<paragraph id="9F421A27FF97852BFF7A467CFEF013D9" blockId="12.[151,427,1095,1158]" box="[151,295,1131,1158]" pageId="12" pageNumber="256">
(
<figureCitation id="07C606A2FF97852BFF72467CFEF713D9" box="[159,288,1131,1158]" captionStart="FIGURES 28–33" captionStartId="14.[152,270,1713,1737]" captionTargetBox="[151,1436,181,1689]" captionTargetId="figure-21@14.[151,1436,181,1689]" captionTargetPageId="14" captionText="FIGURES 28–33. Estola nearnsi sp. nov. 28–32) Holotype female: 28) Dorsal habitus; 29) Ventral habitus; 30) Lateral habitus; 31) Head, frontal view; 32) Protarsal claws. 33) Paratype female, dorsal habitus." figureDoi="http://doi.org/10.5281/zenodo.7656309" httpUri="https://zenodo.org/record/7656309/files/figure.png" pageId="12" pageNumber="256">Figs 28–33</figureCitation>
)
</paragraph>
<paragraph id="9F421A27FF97852BFF7A46A4FC031122" blockId="12.[151,1437,1203,2022]" pageId="12" pageNumber="256">
<emphasis id="AD89C635FF97852BFF7A46A4FE2E1392" bold="true" box="[151,505,1203,1230]" pageId="12" pageNumber="256">
Description. 
<typeStatus id="4046A485FF97852BFEDE46A3FE771391" box="[307,416,1204,1230]" pageId="12" pageNumber="256" type="holotype">Holotype</typeStatus>
female
</emphasis>
(
<figureCitation id="07C606A2FF97852BFDE646A4FD451391" box="[523,658,1203,1230]" captionStart="FIGURES 28–33" captionStartId="14.[152,270,1713,1737]" captionTargetBox="[151,1436,181,1689]" captionTargetId="figure-21@14.[151,1436,181,1689]" captionTargetPageId="14" captionText="FIGURES 28–33. Estola nearnsi sp. nov. 28–32) Holotype female: 28) Dorsal habitus; 29) Ventral habitus; 30) Lateral habitus; 31) Head, frontal view; 32) Protarsal claws. 33) Paratype female, dorsal habitus." figureDoi="http://doi.org/10.5281/zenodo.7656309" httpUri="https://zenodo.org/record/7656309/files/figure.png" pageId="12" pageNumber="256">Figs 28–32</figureCitation>
). Head capsule dark brown dorsally and laterally, brown, gradually lighter toward prothorax ventrally; ventral mouthparts reddish brown, except yellowish-brown apex of palpomeres; anteclypeus and labrum brown posteriorly, fulvous anteriorly; scape orangish-brown on basal third, brownish centrally, dark brown on apical third; pedicel brown on basal third, dark brown on posterior 2/3; antennomeres III–IV orangish about basal 2/3, dark brown on remaining surface; antennomere V orangish on basal half, dark brown on apical half; antennomere VI orangish on basal 3/4, dark brown on apical quarter; antennomeres VII and IX reddish brown about basal half, dark brown on apical half; antennomere VIII and X reddish brown except dark brown apex; antennomere XI reddish brown, slightly brownish about middle. Prothorax and ventral surface of meso- and metathorax black. Elytra blackish basally, dark brown with irregular blackish areas interspersed on remaining anterior 2/3, slightly lighter on posterior third. Basal quarter of tibiae brown dorsally and laterally, blackish ventrally, remaining surface orangish, except dark ring after middle on protibia, ring lighter dorsally, and dark ring about middle on meso- and metatibiae, ring blackish dorsally. Tarsi mostly reddish brown, with irregular orangish areas. Ventrites 1–4 black, except dark reddish brown apex; ventrite 5 mostly dark brown.
</paragraph>
<paragraph id="9F421A27FF97852AFF2A449FFD9814FA" blockId="12.[151,1437,1203,2022]" lastBlockId="13.[151,1437,151,2013]" lastPageId="13" lastPageNumber="257" pageId="12" pageNumber="256">
<emphasis id="AD89C635FF97852BFF2A449FFED911FD" bold="true" box="[199,270,1672,1698]" pageId="12" pageNumber="256">Head.</emphasis>
Frons abundantly, coarsely punctate; with dense straw-colored pubescence, pubescence more yellowish-white close to eyes, except dense, large light brown pubescent macula on each side of median groove close to clypeus, and one dense, large light brown pubescent macula on each side close to antennal tubercles; with long, erect yellowish-white setae interspersed. Area between antennal tubercles with two convergent straw-colored pubescent bands centrally, continuing the frontal pubescence, except glabrous median groove, and dense light brown pubescence on sides; remaining surface of vertex with straw-colored pubescence close to eyes and centrally close to prothorax, and dense light yellowish-brown pubescence, with short straw colored setae interspersed on remaining surface; with long, erect, both yellowish-white and brownish setae interspersed. Area behind eyes with abundant fulvous pubescence partially obscuring integument. Genae 0.3 times lower eye lobe length; with abundant yellowish-white pubescence with light brown pubescence interspersed, except glabrous apex, and central area with sparse light brown pubescence and short, decumbent whitish setae interspersed. Wide central area of postclypeus with dense yellowish-white pubescence, pubescence slightly yellower laterally, and long, somewhat abundant yellowish-white setae directed forward interspersed. Sides of postclypeus glabrous. Labrum with dense yellowishwhite pubescence partially obscuring integument on posterior 2/3, with long, erect setae of same color interspersed, glabrous on anterior third, except anterior margin with fringe of yellowish setae. Gulamentum smooth, glabrous, except intermaxillary process slightly depressed, finely punctate, and with straw-colored pubescence not obscuring integument, and a few somewhat long, erect yellowish-brown setae interspersed. Antennal tubercles with dense light brown pubescence and yellowish-white pubescence interspersed, except frontal region mostly with dense yellowish-white pubescence. Distance between upper eye lobes 0.34 times distance between outer margins of eyes; in frontal view, distance between lower eye lobes 0.62 times distance between outer margins of eyes. Antennae 1.55 times elytral length, reaching elytral apex near apex of antennomere VIII. Scape with abundant yellowish-white pubescence partially obscuring integument on basal 2/3, somewhat abundant brownish pubescence on dorsal surface of apical third, with yellowish-white pubescence interspersed on some areas, and abundant whitish pubescence on apical third of sides and ventral surface; with a few long, erect yellowish-white setae near apex of ventral surface. Pedicel with somewhat abundant yellowish-white pubescence on base of dorsal surface, almost entire sides, and entire ventral surface, and somewhat sparse brownish pubescence on remaining surface; with a few long, erect yellowish-white setae ventrally. Antennomeres III–XI with abundant yellowish-white pubescence almost obscuring integument on light area, and somewhat sparse brownish pubescence on dark area; antennomeres III–X with sparse, long, erect yellowish-white setae ventrally, setae gradually shorter and sparser toward X, setae dark on apical region of some antennomeres; antennomeres V–XI with short, erect yellowish-white setae interspersed. Antennal formula based on length of antennomere III: scape = 1.10; pedicel = 0.29; IV = 1.15; V = 1.02; VI = 0.94; VII = 0.85; VIII = 0.81; IX = 0.72; X = 0.64; XI = 0.68.
</paragraph>
<paragraph id="9F421A27FF96852AFF2A41B8FDDF1012" blockId="13.[151,1437,151,2013]" pageId="13" pageNumber="257">
<emphasis id="AD89C635FF96852AFF2A41B8FEF01496" bold="true" box="[199,295,943,969]" pageId="13" pageNumber="257">Thorax.</emphasis>
Prothorax wider than long; lateral tubercle acute, moderately large, located slightly after middle; sides divergent from anterolateral angles to lateral tubercles, then convergent toward posterolateral angles. Pronotum abundantly, coarsely punctate; with dense brownish-yellow pubescence with part of central anterior half with light brown pubescence, area close to anterior margin and sides of anterior quarter with light brown pubescence, sides with light brown pubescence close to lateral tubercles of prothorax, and wide central area close to posterior margin with both, yellowish and brownish pubescence; sides with a few long, erect dark setae, and posterior margin with sparse, long yellowish setae directed backward. Sides of prothorax with dense brownish-yellow pubescence, except superior region of anterior third with dense light brown pubescence. Prosternum with abundant brownish-yellow pubescence partially obscuring integument, with whitish setae interspersed. Prosternal process with pubescence as on prosternum; narrowest area 0.12 times procoxal width. Mesoventrite with somewhat abundant yellowish-white pubescence not obscuring integument centrally, and dense brownish-yellow pubescence laterally. Mesanepisternum and mesepimeron with dense brownish-yellow pubescence, except yellower pubescence close to elytra. Mesoventral process with dense yellowish pubescence; narrowest area 0.46 times mesocoxal width. Metanepisternum and sides of metaventrite with dense yellowish-brown pubescence; remaining surface of metaventrite with abundant yellowish-white pubescence not obscuring integument. Scutellum with dense brownish-yellow pubescence. 
<emphasis id="AD89C635FF96852AFAA447B0FA4B129E" bold="true" box="[1353,1436,1447,1473]" pageId="13" pageNumber="257">Elytra.</emphasis>
Abundantly, coarsely punctate on anterior third, punctures gradually finer, sparer toward apex; with dense brownish-yellow pubescence and abundant brown pubescent areas interspersed, except dense, bristly yellowish-brown pubescence on anterior quarter of suture; with somewhat abundant, long, suberect dark brown setae throughout, and sparse, long, erect yellowish setae on sides and apex. 
<emphasis id="AD89C635FF96852AFD154420FCE0110E" bold="true" box="[760,823,1591,1617]" pageId="13" pageNumber="257">Legs.</emphasis>
Femora with dense brownish-yellow pubescence, and a few long, erect setae of same color on dorsal apex. Tibiae with moderately abundant yellowish-white pubescence not obscuring integument, except inner margin of dark area of protibiae with dense, bristly dark-brown setae, dorsal surface of dark ring on meso- and metatibiae with abundant, short, erect black setae, and remaining surface of dark rings with somewhat sparse brownish pubescence; all tibiae with long, erect yellowish-white setae interspersed, setae distinctly denser on dorsal surface of meso- and metatibiae. Dorsal surface of tarsi with abundant whitish pubescence not obscuring integument, except basal 2/3 of tarsomeres I with yellowish pubescence; metatarsomere I about as long as II–III together.
</paragraph>
<paragraph id="9F421A27FF96852AFF2A4541FE7C10E6" blockId="13.[151,1437,151,2013]" pageId="13" pageNumber="257">
<emphasis id="AD89C635FF96852AFF2A4541FE94102F" bold="true" box="[199,323,1878,1904]" pageId="13" pageNumber="257">Abdomen.</emphasis>
Ventrites with dense brownish-yellow pubescence, and long, erect setae of same color interspersed, erect setae slightly more abundant laterally. Ventrite 5 triangularly depressed centrally on posterior third, and apical margin slightly concave.
</paragraph>
<paragraph id="9F421A27FF96852AFF2A45D5FA4A1082" blockId="13.[151,1437,151,2013]" box="[199,1437,1986,2013]" pageId="13" pageNumber="257">
<emphasis id="AD89C635FF96852AFF2A45D5FEE81083" bold="true" box="[199,319,1986,2012]" pageId="13" pageNumber="257">Variation.</emphasis>
Brownish-yellow pubescence in the 
<typeStatus id="4046A485FF96852AFD3845D4FCE01082" box="[725,823,1987,2013]" pageId="13" pageNumber="257" type="holotype">holotype</typeStatus>
differs by yellowish-white pubescence on the 
<typeStatus id="4046A485FF96852AFAD845D4FA4E1082" box="[1333,1433,1987,2013]" pageId="13" pageNumber="257" type="paratype">paratype</typeStatus>
.
</paragraph>
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<emphasis id="AD89C635FF958529FF7544A6FE8C1196" bold="true" box="[152,347,1713,1737]" pageId="14" pageNumber="258">FIGURES 28–33.</emphasis>
<taxonomicName id="58FD61A4FF958529FE8C44A6FE231196" authority="Santos-Silva &amp; Monné, 2023" authorityName="Santos-Silva &amp; Monné" authorityYear="2023" box="[353,500,1713,1737]" class="Insecta" family="Cerambycidae" genus="Estola" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="258" phylum="Arthropoda" rank="species" species="nearnsi" status="sp. nov.">
<emphasis id="AD89C635FF958529FE8C44A6FE231196" box="[353,500,1713,1737]" italics="true" pageId="14" pageNumber="258">Estola nearnsi</emphasis>
</taxonomicName>
<emphasis id="AD89C635FF958529FE1644A6FD4C1196" bold="true" box="[507,667,1713,1737]" pageId="14" pageNumber="258">
<taxonomicNameLabel id="B6BA7B4EFF958529FE1644A6FD9B1196" box="[507,588,1713,1737]" pageId="14" pageNumber="258" rank="species">sp. nov.</taxonomicNameLabel>
28–32)
</emphasis>
Holotype female: 
<emphasis id="AD89C635FF958529FCB444A6FCAC1196" bold="true" box="[857,891,1713,1737]" pageId="14" pageNumber="258">28)</emphasis>
Dorsal habitus; 
<emphasis id="AD89C635FF958529FBCE44A6FB931196" bold="true" box="[1059,1092,1713,1737]" pageId="14" pageNumber="258">29)</emphasis>
Ventral habitus; 
<emphasis id="AD89C635FF958529FB1C44A6FAC41196" bold="true" box="[1265,1299,1713,1737]" pageId="14" pageNumber="258">30)</emphasis>
Lateral habitus; 
<emphasis id="AD89C635FF958529FF2F44C2FF3411B2" bold="true" box="[194,227,1749,1773]" pageId="14" pageNumber="258">31)</emphasis>
Head, frontal view; 
<emphasis id="AD89C635FF958529FE5844C2FE0111B2" bold="true" box="[437,470,1749,1773]" pageId="14" pageNumber="258">32)</emphasis>
Protarsal claws. 
<emphasis id="AD89C635FF958529FD6944C2FD7211B2" bold="true" box="[644,677,1749,1773]" pageId="14" pageNumber="258">33)</emphasis>
Paratype female, dorsal habitus.
</paragraph>
</caption>
<paragraph id="9F421A27FF958529FF2A450CFD031021" blockId="14.[151,1436,1819,2026]" pageId="14" pageNumber="258">
<emphasis id="AD89C635FF958529FF2A450CFE4C106A" bold="true" box="[199,411,1819,1845]" pageId="14" pageNumber="258">Dimensions (mm)</emphasis>
(
<typeStatus id="4046A485FF958529FE46450CFDC2106A" box="[427,533,1819,1845]" pageId="14" pageNumber="258" type="holotype">Holotype</typeStatus>
female/ 
<typeStatus id="4046A485FF958529FD9F450CFD05106A" box="[626,722,1819,1845]" pageId="14" pageNumber="258" type="paratype">paratype</typeStatus>
female). Total length, 8.10/7.65; prothoracic length, 1.60/1.55; anterior prothoracic width, 1.75/1.55; posterior prothoracic width, 1.75/1.70; maximum prothoracic width, 2.35/2.15; humeral width, 2.80/2.65; elytral length, 6.00/5.65.
</paragraph>
</subSubSection>
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<paragraph id="9F421A27FF958529FF2A459FFBF510B5" blockId="14.[151,1436,1819,2026]" pageId="14" pageNumber="258">
<emphasis id="AD89C635FF958529FF2A459FFEAF10FE" bold="true" box="[199,376,1927,1954]" pageId="14" pageNumber="258">Type material.</emphasis>
<materialsCitation id="2F95107AFF958529FE6F4590FC131099" ID-GBIF-Occurrence="4037947304" collectingDate="2011-10-13" collectingDateMax="2011-10-16" collectingDateMin="2011-10-13" collectionCode="FSCA" collectorName="Wappes &amp; Skillman" country="Bolivia" elevation="385" latitude="-17.666666" location="Snake Farm" longLatPrecision="1277" longitude="-63.45" municipality="Potrerillo del Guenda" pageId="14" pageNumber="258" specimenCount="1" stateProvince="Santa Cruz" typeStatus="holotype">
<typeStatus id="4046A485FF958529FE6F4590FE3D10FE" box="[386,490,1927,1953]" pageId="14" pageNumber="258" type="holotype">Holotype</typeStatus>
female from 
<collectingCountry id="E7EA5AB7FF958529FD61459FFCD510FD" box="[652,770,1928,1954]" name="Bolivia" pageId="14" pageNumber="258">BOLIVIA</collectingCountry>
, 
<emphasis id="AD89C635FF958529FCFD459FFC4710FD" box="[784,912,1928,1954]" italics="true" pageId="14" pageNumber="258">
<collectingRegion id="5D39D4C5FF958529FCFD459FFC4710FD" box="[784,912,1928,1954]" country="Bolivia" name="Santa Cruz" pageId="14" pageNumber="258">Santa Cruz</collectingRegion>
</emphasis>
: 
<collectingMunicipality id="7F26805DFF958529FC4F4590FB4A10FE" box="[930,1181,1927,1953]" pageId="14" pageNumber="258">Potrerillo del Guendá</collectingMunicipality>
, 
<location id="9A224CFCFF958529FB464590FAEF10FD" LSID="urn:lsid:plazi:treatment:1754AB31FF978528FF7A4650FEE91467:9A224CFCFF958529FB464590FAEF10FD" box="[1195,1336,1927,1954]" country="Bolivia" latitude="-17.666666" longLatPrecision="1277" longitude="-63.45" municipality="Potrerillo del Guenda" name="Snake Farm" pageId="14" pageNumber="258" stateProvince="Santa Cruz">Snake Farm</location>
, 
<geoCoordinate id="FAC97CE0FF958529FAAB459EFA4B10FB" box="[1350,1436,1929,1956]" degrees="17" direction="south" minutes="40" orientation="latitude" pageId="14" pageNumber="258" precision="925" value="-17.666666">17º40′S</geoCoordinate>
<geoCoordinate id="FAC97CE0FF958529FF7A45BAFF2E1097" box="[151,249,1965,1992]" degrees="63" direction="west" minutes="27" orientation="longitude" pageId="14" pageNumber="258" precision="925" value="-63.45">63º27′W</geoCoordinate>
, 
<quantity id="5805B7C2FF958529FEEB45BBFE5E1099" box="[262,393,1964,1990]" metricMagnitude="2" metricUnit="m" metricValue="3.85" metricValueMax="4.0" metricValueMin="3.7" pageId="14" pageNumber="258" unit="m" value="385.0" valueMax="400.0" valueMin="370.0">
<elevation id="14D0FD14FF958529FEEB45BBFE5E1099" box="[262,393,1964,1990]" metricMagnitude="2" metricUnit="m" metricValue="3.85" metricValueMax="4.0" metricValueMin="3.7" pageId="14" pageNumber="258" unit="m" value="385.0" valueMax="400.0" valueMin="370.0">370–400 m</elevation>
</quantity>
, 
<date id="EB433CE7FF958529FE7A45BCFDEE109A" box="[407,569,1963,1989]" pageId="14" pageNumber="258" value="2011-10-13" valueMax="2011-10-16" valueMin="2011-10-13">
<collectingDate id="FB07C50FFF958529FE7A45BCFDEE109A" box="[407,569,1963,1989]" pageId="14" pageNumber="258" value="2011-10-13" valueMax="2011-10-16" valueMin="2011-10-13">13–16.X.2011</collectingDate>
</date>
, 
<collectorName id="32087FF1FF958529FDAB45BBFD491099" box="[582,670,1964,1990]" pageId="14" pageNumber="258">Wappes</collectorName>
&amp; 
<collectorName id="32087FF1FF958529FD2A45BCFCFB109A" box="[711,812,1963,1989]" pageId="14" pageNumber="258">Skillman</collectorName>
leg. (
<collectionCode id="F9EC82E2FF958529FC8245BBFC6C1099" box="[879,955,1964,1990]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="14" pageNumber="258" type="Museum">FSCA</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="2F95107AFF958529FC3D45BBFBF510B5" ID-GBIF-Occurrence="4037947309" collectingDate="2011-11-15" collectingDateMax="2011-11-22" collectingDateMin="2011-11-15" collectionCode="MZSP, FSCA" collectorName="Bettela; Bonaso &amp; Romero" country="Bolivia" elevation="385" latitude="-17.666666" location="Snake Farm" longLatPrecision="1277" longitude="-63.45" municipality="Potrerillo del Guenda" pageId="14" pageNumber="258" specimenCount="1" stateProvince="Santa Cruz" typeStatus="paratype">
<typeStatus id="4046A485FF958529FC3D45BBFBE51099" box="[976,1074,1964,1990]" pageId="14" pageNumber="258" type="paratype">Paratype</typeStatus>
female, same data as holotype, except 
<date id="EB433CE7FF958529FF0A45D8FE4510B5" box="[231,402,1999,2026]" pageId="14" pageNumber="258" value="2011-11-15" valueMax="2011-11-22" valueMin="2011-11-15">
<collectingDate id="FB07C50FFF958529FF0A45D8FE4510B5" box="[231,402,1999,2026]" pageId="14" pageNumber="258" value="2011-11-15" valueMax="2011-11-22" valueMin="2011-11-15">15–22.XI.2011</collectingDate>
</date>
, 
<collectorName id="32087FF1FF958529FE7345D8FE2710B6" box="[414,496,1999,2025]" pageId="14" pageNumber="258">Bettela</collectorName>
, 
<collectorName id="32087FF1FF958529FE1645C7FD9810B5" box="[507,591,2000,2026]" pageId="14" pageNumber="258">Bonaso</collectorName>
&amp; 
<collectorName id="32087FF1FF958529FD9E45C7FD1A10B5" box="[627,717,2000,2026]" pageId="14" pageNumber="258">Romero</collectorName>
leg. (
<collectionCode id="F9EC82E2FF958529FCE145C7FC8010B5" box="[780,855,2000,2026]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="14" pageNumber="258" type="Museum">MZSP</collectionCode>
, formerly 
<collectionCode id="F9EC82E2FF958529FC2645C7FBC110B5" box="[971,1046,2000,2026]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="14" pageNumber="258" type="Museum">FSCA</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="D7E749ACFF948528FF2A4281FC4B17A7" pageId="15" pageNumber="259" type="etymology">
<paragraph id="9F421A27FF948528FF2A4281FC4B17A7" blockId="15.[151,1437,150,824]" pageId="15" pageNumber="259">
<emphasis id="AD89C635FF948528FF2A4281FE9917EF" bold="true" box="[199,334,150,176]" pageId="15" pageNumber="259">Etymology.</emphasis>
We are happy to name this species in honor of Eugenio H. Nearns (National Museum of Natural History, Smithsonian Institution, 
<collectingCountry id="E7EA5AB7FF948528FDE242ADFCF9178B" box="[527,814,186,212]" name="United States of America" pageId="15" pageNumber="259">United States of America</collectingCountry>
) for his friendship and for his great contribution to the knowledge of the Onciderini (
<taxonomicName id="58FD61A4FF948528FE0A42C9FDB017A7" authorityName="Linnaeus" authorityYear="1758" box="[487,615,222,248]" class="Magnoliopsida" higherTaxonomySource="GBIF" kingdom="Plantae" order="Coleoptera" pageId="15" pageNumber="259" phylum="Tracheophyta" rank="order">Coleoptera</taxonomicName>
, 
<taxonomicName id="58FD61A4FF948528FD9F42C9FCC017A7" authorityName="Latreille" authorityYear="1802" box="[626,791,222,248]" class="Insecta" family="Cerambycidae" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="259" phylum="Arthropoda" rank="family">Cerambycidae</taxonomicName>
, Lamiinae).
</paragraph>
</subSubSection>
<subSubSection id="D7E749ACFF948528FF2A4315FEE91467" pageId="15" pageNumber="259" type="discussion">
<paragraph id="9F421A27FF948528FF2A4315FEE91467" blockId="15.[151,1437,150,824]" pageId="15" pageNumber="259">
<emphasis id="AD89C635FF948528FF2A4315FEEC1643" bold="true" box="[199,315,258,284]" pageId="15" pageNumber="259">Remarks.</emphasis>
<taxonomicName id="58FD61A4FF948528FEAF4315FE321643" authority="Santos-Silva &amp; Monné, 2023" authorityName="Santos-Silva &amp; Monné" authorityYear="2023" box="[322,485,258,284]" class="Insecta" family="Cerambycidae" genus="Estola" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="259" phylum="Arthropoda" rank="species" species="nearnsi" status="sp. nov.">
<emphasis id="AD89C635FF948528FEAF4315FE321643" box="[322,485,258,284]" italics="true" pageId="15" pageNumber="259">Estola nearnsi</emphasis>
</taxonomicName>
<emphasis id="AD89C635FF948528FE004315FD901643" bold="true" box="[493,583,258,284]" pageId="15" pageNumber="259">
<taxonomicNameLabel id="B6BA7B4EFF948528FE004315FD901643" box="[493,583,258,284]" pageId="15" pageNumber="259" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is similar to 
<taxonomicName id="58FD61A4FF948528FD374314FBCA1643" authority="Breuning, 1940" authorityName="Breuning" authorityYear="1940" box="[730,1053,258,284]" class="Insecta" family="Cerambycidae" genus="Estola" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="259" phylum="Arthropoda" rank="species" species="annulipes">
<emphasis id="AD89C635FF948528FD374314FCB01643" box="[730,871,258,284]" italics="true" pageId="15" pageNumber="259">E. annulipes</emphasis>
Breuning, 1940
</taxonomicName>
(see photographs on 
<bibRefCitation id="FB6C67D6FF948528FAE34315FA4B1643" author="Bezark, L. G." box="[1294,1436,258,284]" pageId="15" pageNumber="259" refId="ref10171" refString="Bezark, L. G. (2023) A photographic Catalog of the Cerambycidae of the World. New World Cerambycidae Catalog. Available from: http: // bezbycids. com / byciddb / wdefault. asp? w = n / (accessed 3 January 2023)" type="url" year="2023">Bezark 2023</bibRefCitation>
and 
<bibRefCitation id="FB6C67D6FF948528FF2B4331FE03161F" author="Galileo, M. H. M. &amp; Martins, U. R." box="[198,468,294,320]" pageId="15" pageNumber="259" pagination="65 - 309" refId="ref10344" refString="Galileo, M. H. M. &amp; Martins, U. R. (2006) Inventario descritivo. In: Galileo, M. H. M. &amp; Martins, U. R. (Orgs.), Cerambycidae (Coleoptera, Insecta) do Parque Copesul de Protecao Ambiental, Triunfo, Rio Grande do Sul, Brasil. Museu de Ciencias Naturais da Fundac &quot; o Zoobotanica do RS, Porto Alegre, pp. 65 - 309." type="book chapter" year="2006">Galileo &amp; Martins 2006</bibRefCitation>
), but differs by the lower eye lobes distinctly longer than genae (according to 
<bibRefCitation id="FB6C67D6FF948528FAD94331FF33163B" author="Breuning, S." pageId="15" pageNumber="259" pagination="1 - 210" refId="ref10221" refString="Breuning, S. (1974) Revision des Rhodopinini americains. Studia Entomologica, 17 (1 - 4), 1 - 210." type="journal article" year="1974">Breuning (1974)</bibRefCitation>
, lower eye lobes half as long as genae in 
<taxonomicName id="58FD61A4FF948528FD43435CFCED163B" authorityName="Breuning" authorityYear="1940" box="[686,826,330,356]" class="Insecta" family="Cerambycidae" genus="Estola" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="259" phylum="Arthropoda" rank="species" species="annulipes">
<emphasis id="AD89C635FF948528FD43435CFCED163B" box="[686,826,330,356]" italics="true" pageId="15" pageNumber="259">E. annulipes</emphasis>
</taxonomicName>
), frons distinctly wider than long (according to 
<bibRefCitation id="FB6C67D6FF948528FAA3435CFED816D7" author="Breuning, S." pageId="15" pageNumber="259" pagination="1 - 210" refId="ref10221" refString="Breuning, S. (1974) Revision des Rhodopinini americains. Studia Entomologica, 17 (1 - 4), 1 - 210." type="journal article" year="1974">Breuning (1974)</bibRefCitation>
, frons as wide as long), and pronotum without oblique yellowish band basally (present in 
<taxonomicName id="58FD61A4FF948528FAED4378FA5B16D7" authorityName="Breuning" authorityYear="1940" box="[1280,1420,366,392]" class="Insecta" family="Cerambycidae" genus="Estola" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="259" phylum="Arthropoda" rank="species" species="annulipes">
<emphasis id="AD89C635FF948528FAED4378FA5B16D7" box="[1280,1420,366,392]" italics="true" pageId="15" pageNumber="259">E. annulipes</emphasis>
</taxonomicName>
). The new species differs from 
<taxonomicName id="58FD61A4FF948528FE0F4384FCB916F3" authority="Breuning, 1940" authorityName="Breuning" authorityYear="1940" box="[482,878,402,428]" class="Insecta" family="Cerambycidae" genus="Estola" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="259" phylum="Arthropoda" rank="species" species="fuscomarmorata">
<emphasis id="AD89C635FF948528FE0F4384FD6D16F3" box="[482,698,402,428]" italics="true" pageId="15" pageNumber="259">E. fuscomarmorata</emphasis>
Breuning, 1940
</taxonomicName>
(see photographs on 
<bibRefCitation id="FB6C67D6FF948528FBB14385FB3D16F3" author="Bezark, L. G." box="[1116,1258,402,428]" pageId="15" pageNumber="259" refId="ref10171" refString="Bezark, L. G. (2023) A photographic Catalog of the Cerambycidae of the World. New World Cerambycidae Catalog. Available from: http: // bezbycids. com / byciddb / wdefault. asp? w = n / (accessed 3 January 2023)" type="url" year="2023">Bezark 2023</bibRefCitation>
and 
<bibRefCitation id="FB6C67D6FF948528FACD4385FEC6168F" author="Lingafelter, S. &amp; Monne, M. A. &amp; Nearns, E. H." pageId="15" pageNumber="259" refId="ref10521" refString="Lingafelter, S. W, Monne, M. A. &amp; Nearns, E. H. (2023) Online Image Database of Cerambycoid Primary Types of Smithsonian Intitution. Available from: http: // SmithsonianCerambycidae. com / (accessed 3 January 2023)" type="url" year="2023">
Lingafelter 
<emphasis id="AD89C635FF948528FF7A43A0FF07168F" box="[151,208,438,464]" italics="true" pageId="15" pageNumber="259">et al.</emphasis>
2023
</bibRefCitation>
) by the body stouter (slender in 
<taxonomicName id="58FD61A4FF948528FD9043A0FC82168F" authorityName="Breuning" authorityYear="1940" box="[637,853,438,464]" class="Insecta" family="Cerambycidae" genus="Estola" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="259" phylum="Arthropoda" rank="species" species="fuscomarmorata">
<emphasis id="AD89C635FF948528FD9043A0FC82168F" box="[637,853,438,464]" italics="true" pageId="15" pageNumber="259">E. fuscomarmorata</emphasis>
</taxonomicName>
), antennomeres III and VI bicolorous (entirely dark in 
<taxonomicName id="58FD61A4FF948528FF5B43CCFE4616AB" authorityName="Breuning" authorityYear="1940" box="[182,401,474,500]" class="Insecta" family="Cerambycidae" genus="Estola" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="259" phylum="Arthropoda" rank="species" species="fuscomarmorata">
<emphasis id="AD89C635FF948528FF5B43CCFE4616AB" box="[182,401,474,500]" italics="true" pageId="15" pageNumber="259">E. fuscomarmorata</emphasis>
</taxonomicName>
), and elytral pubescence not forming oblique band on anterior third (forming in 
<taxonomicName id="58FD61A4FF948528FADB43CCFEC21547" authorityName="Breuning" authorityYear="1940" class="Insecta" family="Cerambycidae" genus="Estola" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="259" phylum="Arthropoda" rank="species" species="fuscomarmorata">
<emphasis id="AD89C635FF948528FADB43CCFEC21547" italics="true" pageId="15" pageNumber="259">E. fuscomarmorata</emphasis>
</taxonomicName>
). 
<taxonomicName id="58FD61A4FF948528FEC343E9FE041547" authority="Santos-Silva &amp; Monné, 2023" authorityName="Santos-Silva &amp; Monné" authorityYear="2023" box="[302,467,510,536]" class="Insecta" family="Cerambycidae" genus="Estola" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="259" phylum="Arthropoda" rank="species" species="nearnsi" status="sp. nov.">
<emphasis id="AD89C635FF948528FEC343E9FE041547" box="[302,467,510,536]" italics="true" pageId="15" pageNumber="259">Estola nearnsi</emphasis>
</taxonomicName>
<emphasis id="AD89C635FF948528FE3143E9FDE01547" bold="true" box="[476,567,510,536]" pageId="15" pageNumber="259">
<taxonomicNameLabel id="B6BA7B4EFF948528FE3143E9FDE01547" box="[476,567,510,536]" pageId="15" pageNumber="259" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be separated from 
<taxonomicName id="58FD61A4FF948528FCAB43E8FB791547" authority="Breuning, 1940" authorityName="Breuning" authorityYear="1940" box="[838,1198,510,536]" class="Insecta" family="Cerambycidae" genus="Estola" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="259" phylum="Arthropoda" rank="species" species="nigrosignata">
<emphasis id="AD89C635FF948528FCAB43E8FC221547" box="[838,1013,511,536]" italics="true" pageId="15" pageNumber="259">E. nigrosignata</emphasis>
Breuning, 1940
</taxonomicName>
(see photographs on 
<bibRefCitation id="FB6C67D6FF948528FF7A4035FEFF1563" author="Bezark, L. G." box="[151,296,546,572]" pageId="15" pageNumber="259" refId="ref10171" refString="Bezark, L. G. (2023) A photographic Catalog of the Cerambycidae of the World. New World Cerambycidae Catalog. Available from: http: // bezbycids. com / byciddb / wdefault. asp? w = n / (accessed 3 January 2023)" type="url" year="2023">Bezark 2023</bibRefCitation>
) and 
<taxonomicName id="58FD61A4FF948528FE8A4034FD7A1563" authority="(Fabricius, 1793)" baseAuthorityName="Fabricius" baseAuthorityYear="1793" box="[359,685,546,572]" class="Insecta" family="Cerambycidae" genus="Estola" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="259" phylum="Arthropoda" rank="species" species="obscura">
<emphasis id="AD89C635FF948528FE8A4034FE361563" box="[359,481,546,572]" italics="true" pageId="15" pageNumber="259">E. obscura</emphasis>
(Fabricius, 1793)
</taxonomicName>
, by the elytra without grayish-white or whitish pubescent maculae interspersed (grayish-white or whitish pubescence present on the elytra of 
<taxonomicName id="58FD61A4FF948528FC044050FB40153F" authorityName="Breuning" authorityYear="1940" box="[1001,1175,583,608]" class="Insecta" family="Cerambycidae" genus="Estola" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="259" phylum="Arthropoda" rank="species" species="nigrosignata">
<emphasis id="AD89C635FF948528FC044050FB40153F" box="[1001,1175,583,608]" italics="true" pageId="15" pageNumber="259">E. nigrosignata</emphasis>
</taxonomicName>
and 
<taxonomicName id="58FD61A4FF948528FB3C4050FA9B153F" baseAuthorityName="Fabricius" baseAuthorityYear="1793" box="[1233,1356,582,608]" class="Insecta" family="Cerambycidae" genus="Estola" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="259" phylum="Arthropoda" rank="species" species="obscura">
<emphasis id="AD89C635FF948528FB3C4050FA9B153F" box="[1233,1356,582,608]" italics="true" pageId="15" pageNumber="259">E. obscura</emphasis>
</taxonomicName>
); from 
<taxonomicName id="58FD61A4FF948528FF7A407CFE0815DB" authority="Breuning, 1940" authorityName="Breuning" authorityYear="1940" box="[151,479,618,644]" class="Insecta" family="Cerambycidae" genus="Estola" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="259" phylum="Arthropoda" rank="species" species="compacta">
<emphasis id="AD89C635FF948528FF7A407CFEF115DB" box="[151,294,619,644]" italics="true" pageId="15" pageNumber="259">E. compacta</emphasis>
Breuning, 1940
</taxonomicName>
(see photographs on 
<bibRefCitation id="FB6C67D6FF948528FD3A407DFCBB15DB" author="Bezark, L. G." box="[727,876,618,644]" pageId="15" pageNumber="259" refId="ref10171" refString="Bezark, L. G. (2023) A photographic Catalog of the Cerambycidae of the World. New World Cerambycidae Catalog. Available from: http: // bezbycids. com / byciddb / wdefault. asp? w = n / (accessed 3 January 2023)" type="url" year="2023">Bezark 2023</bibRefCitation>
), by the elytra without wide dark pubescent area on anterior half (present in 
<taxonomicName id="58FD61A4FF948528FE244098FD8115F7" authorityName="Breuning" authorityYear="1940" box="[457,598,655,680]" class="Insecta" family="Cerambycidae" genus="Estola" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="259" phylum="Arthropoda" rank="species" species="compacta">
<emphasis id="AD89C635FF948528FE244098FD8115F7" box="[457,598,655,680]" italics="true" pageId="15" pageNumber="259">E. compacta</emphasis>
</taxonomicName>
); from 
<taxonomicName id="58FD61A4FF948528FD464098FC1B15F7" authority="Lameere, 1893" authorityName="Lameere" authorityYear="1893" box="[683,972,654,681]" class="Insecta" family="Cerambycidae" genus="Estola" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="259" phylum="Arthropoda" rank="species" species="hispida">
<emphasis id="AD89C635FF948528FD464098FCCB15F7" box="[683,796,654,680]" italics="true" pageId="15" pageNumber="259">E. hispida</emphasis>
Lameere, 1893
</taxonomicName>
(see photograph on 
<bibRefCitation id="FB6C67D6FF948528FB5C4099FA9415F7" author="Bezark, L. G." box="[1201,1347,654,680]" pageId="15" pageNumber="259" refId="ref10171" refString="Bezark, L. G. (2023) A photographic Catalog of the Cerambycidae of the World. New World Cerambycidae Catalog. Available from: http: // bezbycids. com / byciddb / wdefault. asp? w = n / (accessed 3 January 2023)" type="url" year="2023">Bezark 2023</bibRefCitation>
), by the body stouter (slender in 
<taxonomicName id="58FD61A4FF948528FE4540A4FDCE1593" authorityName="Lameere" authorityYear="1893" box="[424,537,690,716]" class="Insecta" family="Cerambycidae" genus="Estola" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="259" phylum="Arthropoda" rank="species" species="hispida">
<emphasis id="AD89C635FF948528FE4540A4FDCE1593" box="[424,537,690,716]" italics="true" pageId="15" pageNumber="259">E. hispida</emphasis>
</taxonomicName>
), and elytral punctures not distinctly contrasting with the pubescence (distinctly contrasting in 
<taxonomicName id="58FD61A4FF948528FED640C0FE7915AF" authorityName="Lameere" authorityYear="1893" box="[315,430,726,752]" class="Insecta" family="Cerambycidae" genus="Estola" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="259" phylum="Arthropoda" rank="species" species="hispida">
<emphasis id="AD89C635FF948528FED640C0FE7915AF" box="[315,430,726,752]" italics="true" pageId="15" pageNumber="259">E. hispida</emphasis>
</taxonomicName>
); and from 
<taxonomicName id="58FD61A4FF948528FDD540C0FC3315AF" authority="Martins &amp; Galileo, 2006" authorityName="Martins &amp; Galileo" authorityYear="2006" box="[568,996,726,752]" class="Insecta" family="Cerambycidae" genus="Estola" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="259" phylum="Arthropoda" rank="species" species="timbauba">
<emphasis id="AD89C635FF948528FDD540C0FD1415AF" box="[568,707,726,752]" italics="true" pageId="15" pageNumber="259">E. timbauba</emphasis>
Martins &amp; 
<bibRefCitation id="FB6C67D6FF948528FCA740C1FC3315AF" author="Galileo, M. H. M. &amp; Martins, U. R." box="[842,996,726,752]" pageId="15" pageNumber="259" pagination="65 - 309" refId="ref10344" refString="Galileo, M. H. M. &amp; Martins, U. R. (2006) Inventario descritivo. In: Galileo, M. H. M. &amp; Martins, U. R. (Orgs.), Cerambycidae (Coleoptera, Insecta) do Parque Copesul de Protecao Ambiental, Triunfo, Rio Grande do Sul, Brasil. Museu de Ciencias Naturais da Fundac &quot; o Zoobotanica do RS, Porto Alegre, pp. 65 - 309." type="book chapter" year="2006">Galileo, 2006</bibRefCitation>
</taxonomicName>
(see photographs on 
<bibRefCitation id="FB6C67D6FF948528FB3640C1FABC15AF" author="Bezark, L. G." box="[1243,1387,726,752]" pageId="15" pageNumber="259" refId="ref10171" refString="Bezark, L. G. (2023) A photographic Catalog of the Cerambycidae of the World. New World Cerambycidae Catalog. Available from: http: // bezbycids. com / byciddb / wdefault. asp? w = n / (accessed 3 January 2023)" type="url" year="2023">Bezark 2023</bibRefCitation>
and original description), by the pronotal and elytral pubescence distinctly denser (sparser in 
<taxonomicName id="58FD61A4FF948528FB9940ECFB2A144B" authorityName="Martins &amp; Galileo" authorityYear="2006" box="[1140,1277,762,788]" class="Insecta" family="Cerambycidae" genus="Estola" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="259" phylum="Arthropoda" rank="species" species="timbauba">
<emphasis id="AD89C635FF948528FB9940ECFB2A144B" box="[1140,1277,762,788]" italics="true" pageId="15" pageNumber="259">E. timbauba</emphasis>
</taxonomicName>
, especially on the pronotum).
</paragraph>
</subSubSection>
</treatment>
</document>