229 lines
30 KiB
XML
229 lines
30 KiB
XML
<document id="1CFCE271018C05B11A32D80327F8CE7F" ID-DOI="10.11646/zootaxa.4020.1.7" ID-GBIF-Dataset="82b2f539-5d25-42d6-95c5-e8b16a353e12" ID-ISSN="1175-5326" ID-Zenodo-Dep="237970" ID-ZooBank="BD8D1880-79A9-4672-99AF-5B3D1BEFAFEE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461269068152" checkinUser="plazi" docAuthor="Bickel, Daniel J." docDate="2015" docId="172D87FDFFBFFF8CFF3DA51301C5FAFE" docLanguage="en" docName="zt04020p182.pdf" docOrigin="Zootaxa 4020 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Systenus" docType="key" docVersion="8" lastPageNumber="171" masterDocId="EB14FF85FFBDFF8EFFAAA70A0459FFCC" masterDocTitle="The Costa Rican Systenus Loew (Diptera: Dolichopodidae): rich local sympatry in an otherwise rare genus" masterLastPageNumber="182" masterPageNumber="169" pageNumber="171" updateTime="1698611248703" updateUser="plazi">
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<mods:title id="7F0D763AAD0464329E0EF51056219107">The Costa Rican Systenus Loew (Diptera: Dolichopodidae): rich local sympatry in an otherwise rare genus</mods:title>
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<mods:name id="7CCA6BCF99B2F05CCFF17CBAD6685C75" type="personal">
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<mods:namePart id="250CDABCDCA89B05E7F84BA0F32C1FC4">Bickel, Daniel J.</mods:namePart>
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<mods:title id="17F6CC423AB09CC4873F727CFC933B6B">Zootaxa</mods:title>
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<mods:part id="C73648C5690C7340D34DB73CFE7771AB">
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<mods:date id="4A63DF804263A93D25E4CA596F02CA9C">2015</mods:date>
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<mods:number id="6734144FDDC8B15E5FB0FF947F1C83FB">4020</mods:number>
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<mods:identifier id="E9FBB408884325CDDFBC0AD7B4B9F4A5" type="DOI">10.11646/zootaxa.4020.1.7</mods:identifier>
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<treatment id="172D87FDFFBFFF8CFF3DA51301C5FAFE" ID-DOI="http://doi.org/10.5281/zenodo.6121795" ID-GBIF-Taxon="119638513" ID-Zenodo-Dep="6121795" LSID="urn:lsid:plazi:treatment:172D87FDFFBFFF8CFF3DA51301C5FAFE" httpUri="http://treatment.plazi.org/id/172D87FDFFBFFF8CFF3DA51301C5FAFE" lastPageNumber="171" pageId="2" pageNumber="171">
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<subSubSection id="D79E6560FFBFFF8CFF3DA51306C4FDFF" box="[151,669,537,563]" pageId="2" pageNumber="171" type="nomenclature">
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<paragraph id="9F3B36EBFFBFFF8CFF3DA51306C4FDFF" blockId="2.[151,669,537,563]" box="[151,669,537,563]" pageId="2" pageNumber="171">
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<heading id="C4738187FFBFFF8CFF3DA51306C4FDFF" bold="true" box="[151,669,537,563]" fontSize="11" level="1" pageId="2" pageNumber="171" reason="1">
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<emphasis id="ADF0EAF9FFBFFF8CFF3DA51306C4FDFF" bold="true" box="[151,669,537,563]" pageId="2" pageNumber="171">
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Key to males of
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<taxonomicName id="58844D68FFBFFF8CFEF5A5130590FDFF" ID-CoL="7RNG" ID-ENA="1000637" box="[351,457,537,563]" class="Insecta" family="Dolichopodidae" genus="Systenus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="171" phylum="Arthropoda" rank="genus">
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<emphasis id="ADF0EAF9FFBFFF8CFEF5A5130590FDFF" bold="true" box="[351,457,537,563]" italics="true" pageId="2" pageNumber="171">Systenus</emphasis>
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</taxonomicName>
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from
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<collectingCountry id="E793767BFFBFFF8CFDBFA51306C4FDFF" box="[533,669,537,563]" name="Costa Rica" pageId="2" pageNumber="171">Costa Rica</collectingCountry>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="D79E6560FFBFFF8CFF3DA55001C5FAFE" pageId="2" pageNumber="171" type="key">
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<keyStep id="24702E4EFFBFFF8CFF3DA55001C2FD40" pageId="2" pageNumber="171">
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<paragraph id="9F3B36EBFFBFFF8CFF3DA55001CCFDA3" blockId="2.[151,1436,602,1330]" box="[151,1429,602,623]" pageId="2" pageNumber="171">
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<keyLead id="247595DEFFBFFF8CFF3DA55001CCFDA3" box="[151,1429,602,623]" pageId="2" pageNumber="171">1 All femora yellow.................................................................................... 2</keyLead>
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</paragraph>
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<paragraph id="9F3B36EBFFBFFF8CFF3DA57D01C2FD40" blockId="2.[151,1436,602,1330]" box="[151,1435,631,652]" pageId="2" pageNumber="171">
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<keyLead id="247595DEFFBFFF8CFF3DA57D01C2FD40" box="[151,1435,631,652]" pageId="2" pageNumber="171">- At least one femur with some brown cuticle................................................................. 4</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="24702E4EFFBFFF8CFF3DA59F01C5FD28" pageId="2" pageNumber="171">
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<paragraph id="9F3B36EBFFBFFF8CFF3DA59F01C5FD04" blockId="2.[151,1436,602,1330]" pageId="2" pageNumber="171">
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<keyLead id="247595DEFFBFFF8CFF3DA59F01C5FD04" pageId="2" pageNumber="171">
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2 Cercus short, wide and subrectangular (
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<figureCitation id="07BF2A6EFFBFFF8CFDCBA59F06F9FD66" box="[609,672,661,682]" captionStart="FIGURE 4" captionStartId="11.[151,250,1770,1792]" captionTargetBox="[160,1436,628,1748]" captionTargetId="figure@11.[151,1436,628,1749]" captionTargetPageId="11" captionText="FIGURE 4. Systenus eboritibia sp. nov. (holotype): a. male habitus, left lateral; b. hypopygium, left lateral. S. parkeri sp. nov.: c. male habitus, left lateral; d. hypopygium, left lateral; e. male postabdomen, right lateral." httpUri="https://zenodo.org/record/237974/files/figure.png" pageId="2" pageNumber="171">Figs 4</figureCitation>
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d, e); tibiae yellow with brownish vestiture; postpedicel basally subrectangular and yellow, and abruptly narrowed to elongate dark brown tapering point distally (
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<figureCitation id="07BF2A6EFFBFFF8CFC5DA5B80068FD0B" box="[1015,1073,690,711]" captionStart="FIGURE 4" captionStartId="11.[151,250,1770,1792]" captionTargetBox="[160,1436,628,1748]" captionTargetId="figure@11.[151,1436,628,1749]" captionTargetPageId="11" captionText="FIGURE 4. Systenus eboritibia sp. nov. (holotype): a. male habitus, left lateral; b. hypopygium, left lateral. S. parkeri sp. nov.: c. male habitus, left lateral; d. hypopygium, left lateral; e. male postabdomen, right lateral." httpUri="https://zenodo.org/record/237974/files/figure.png" pageId="2" pageNumber="171">Fig. 4</figureCitation>
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c).............
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<taxonomicName id="58844D68FFBFFF8CFB41A5BB0110FD0B" ID-CoL="9HYXF" box="[1259,1353,689,711]" class="Insecta" family="Dolichopodidae" genus="Systenus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="171" phylum="Arthropoda" rank="species" species="parkeri" status="sp. nov.">
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<emphasis id="ADF0EAF9FFBFFF8CFB41A5BB0110FD0B" box="[1259,1353,689,711]" italics="true" pageId="2" pageNumber="171">S. parkeri</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="B6C35782FFBFFF8CFAE5A5B801C5FD04" box="[1359,1436,690,712]" pageId="2" pageNumber="171" rank="species">
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<emphasis id="ADF0EAF9FFBFFF8CFAE5A5B801C5FD04" bold="true" box="[1359,1436,690,712]" pageId="2" pageNumber="171">sp. nov.</emphasis>
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</taxonomicNameLabel>
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</keyLead>
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</paragraph>
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<paragraph id="9F3B36EBFFBFFF8CFF3DA5DA01C5FD28" blockId="2.[151,1436,602,1330]" box="[151,1436,719,741]" pageId="2" pageNumber="171">
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<keyLead id="247595DEFFBFFF8CFF3DA5DA01C5FD28" box="[151,1436,719,741]" pageId="2" pageNumber="171">
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<emphasis id="ADF0EAF9FFBFFF8CFF3DA5DA04C6FD29" box="[151,159,720,741]" italics="true" pageId="2" pageNumber="171">-</emphasis>
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Cercus elongate and digitiform (
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<figureCitation id="07BF2A6EFFBFFF8CFDBDA5C50608FD28" box="[535,593,719,740]" captionStart="FIGURE 1" captionStartId="4.[151,250,1872,1894]" captionTargetBox="[238,1352,195,1838]" captionTargetId="figure@4.[234,1352,195,1839]" captionTargetPageId="4" captionText="FIGURE 1. Systenus tenorio sp. nov. (holotype): a. male habitus, left lateral (terminalia removed); b. hypopygium, left lateral. S. flavifemoratus sp. nov. (holotype): c. male habitus, right lateral; d. hypopygium, left lateral. S. maculipennis sp. nov.: e. male habitus, left lateral (terminalia removed); f. hypopygium, left lateral. Abbreviations: cer, cercus; epan, epandrium; hyp, hypandrium; ph, phallus; st, sternite; sur, surstylus; tg, tergite; 6, 7, 8, abdominal segments." httpUri="https://zenodo.org/record/237971/files/figure.png" pageId="2" pageNumber="171">Figs 1</figureCitation>
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d, f); tibia I white with ivory colored vestiture; postpedicel various.............. 3
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</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="24702E4EFFBFFF8CFF3DA5E701CFFCF1" pageId="2" pageNumber="171">
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<paragraph id="9F3B36EBFFBFFF8CFF3DA5E701CFFCEC" blockId="2.[151,1436,602,1330]" pageId="2" pageNumber="171">
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<keyLead id="247595DEFFBFFF8CFF3DA5E701CFFCEC" pageId="2" pageNumber="171">
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3 Wing apex brown, and posterior membrane reduced near apex so vein M forms the posterior margin (
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<figureCitation id="07BF2A6EFFBFFF8CFB0AA5E7008FFCCE" box="[1184,1238,749,770]" captionStart="FIGURE 1" captionStartId="4.[151,250,1872,1894]" captionTargetBox="[238,1352,195,1838]" captionTargetId="figure@4.[234,1352,195,1839]" captionTargetPageId="4" captionText="FIGURE 1. Systenus tenorio sp. nov. (holotype): a. male habitus, left lateral (terminalia removed); b. hypopygium, left lateral. S. flavifemoratus sp. nov. (holotype): c. male habitus, right lateral; d. hypopygium, left lateral. S. maculipennis sp. nov.: e. male habitus, left lateral (terminalia removed); f. hypopygium, left lateral. Abbreviations: cer, cercus; epan, epandrium; hyp, hypandrium; ph, phallus; st, sternite; sur, surstylus; tg, tergite; 6, 7, 8, abdominal segments." httpUri="https://zenodo.org/record/237971/files/figure.png" pageId="2" pageNumber="171">Fig. 1</figureCitation>
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e); CuAx ratio> 1.0.................................................................................
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<taxonomicName id="58844D68FFBFFF8CFB01A403011AFCD3" ID-CoL="9HWXR" box="[1195,1347,777,799]" class="Insecta" family="Dolichopodidae" genus="Systenus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="171" phylum="Arthropoda" rank="species" species="maculipennis" status="sp. nov.">
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<emphasis id="ADF0EAF9FFBFFF8CFB01A403011AFCD3" box="[1195,1347,777,799]" italics="true" pageId="2" pageNumber="171">S. maculipennis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="B6C35782FFBFFF8CFAE3A40001CFFCEC" box="[1353,1430,778,800]" pageId="2" pageNumber="171" rank="species">
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<emphasis id="ADF0EAF9FFBFFF8CFAE3A40001CFFCEC" bold="true" box="[1353,1430,778,800]" pageId="2" pageNumber="171">sp. nov.</emphasis>
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</taxonomicNameLabel>
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</keyLead>
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</paragraph>
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<paragraph id="9F3B36EBFFBFFF8CFF3DA42D01CFFCF1" blockId="2.[151,1436,602,1330]" box="[151,1430,807,829]" pageId="2" pageNumber="171">
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<keyLead id="247595DEFFBFFF8CFF3DA42D01CFFCF1" box="[151,1430,807,829]" pageId="2" pageNumber="171">
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- Wing hyaline, unmodified (
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<figureCitation id="07BF2A6EFFBFFF8CFE43A42D0679FCF0" box="[489,544,807,828]" captionStart="FIGURE 1" captionStartId="4.[151,250,1872,1894]" captionTargetBox="[238,1352,195,1838]" captionTargetId="figure@4.[234,1352,195,1839]" captionTargetPageId="4" captionText="FIGURE 1. Systenus tenorio sp. nov. (holotype): a. male habitus, left lateral (terminalia removed); b. hypopygium, left lateral. S. flavifemoratus sp. nov. (holotype): c. male habitus, right lateral; d. hypopygium, left lateral. S. maculipennis sp. nov.: e. male habitus, left lateral (terminalia removed); f. hypopygium, left lateral. Abbreviations: cer, cercus; epan, epandrium; hyp, hypandrium; ph, phallus; st, sternite; sur, surstylus; tg, tergite; 6, 7, 8, abdominal segments." httpUri="https://zenodo.org/record/237971/files/figure.png" pageId="2" pageNumber="171">Fig. 1</figureCitation>
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c); CuAx ratio <0.7....................................
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<taxonomicName id="58844D68FFBFFF8CFB0AA42D011BFCF1" ID-CoL="9HTL3" box="[1184,1346,807,829]" class="Insecta" family="Dolichopodidae" genus="Systenus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="171" phylum="Arthropoda" rank="species" species="flavifemoratus" status="sp. nov.">
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<emphasis id="ADF0EAF9FFBFFF8CFB0AA42D011BFCF1" box="[1184,1346,807,829]" italics="true" pageId="2" pageNumber="171">S. flavifemoratus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="B6C35782FFBFFF8CFAE3A42D01CFFCF1" box="[1353,1430,807,829]" pageId="2" pageNumber="171" rank="species">
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<emphasis id="ADF0EAF9FFBFFF8CFAE3A42D01CFFCF1" bold="true" box="[1353,1430,807,829]" pageId="2" pageNumber="171">sp. nov.</emphasis>
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</taxonomicNameLabel>
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</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="24702E4EFFBFFF8CFF3DA44F01CCFCBB" pageId="2" pageNumber="171">
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<paragraph id="9F3B36EBFFBFFF8CFF3DA44F01CCFC96" blockId="2.[151,1436,602,1330]" box="[151,1429,837,858]" pageId="2" pageNumber="171">
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<keyLead id="247595DEFFBFFF8CFF3DA44F01CCFC96" box="[151,1429,837,858]" pageId="2" pageNumber="171">4 All femora with at least some brown cuticle; postpedicel elongate, triangular and tapering gradually.................. 5</keyLead>
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</paragraph>
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<paragraph id="9F3B36EBFFBFFF8CFF3DA46801CCFCBB" blockId="2.[151,1436,602,1330]" box="[151,1429,866,887]" pageId="2" pageNumber="171">
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<keyLead id="247595DEFFBFFF8CFF3DA46801CCFCBB" box="[151,1429,866,887]" pageId="2" pageNumber="171">- Femora I and II yellow, and only femur III with brown cuticle distally; postpedicel various.......................... 6</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="24702E4EFFBFFF8CFF3DA47501C5FC21" pageId="2" pageNumber="171">
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<paragraph id="9F3B36EBFFBFFF8CFF3DA47501C5FC7E" blockId="2.[151,1436,602,1330]" pageId="2" pageNumber="171">
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<keyLead id="247595DEFFBFFF8CFF3DA47501C5FC7E" pageId="2" pageNumber="171">
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5 Both tibiae I and III with white cuticle and short ivory colored vestiture; tibia II yellow with brownish vestiture (
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<figureCitation id="07BF2A6EFFBFFF8CFAB4A4750101FC58" box="[1310,1368,895,916]" captionStart="FIGURE 4" captionStartId="11.[151,250,1770,1792]" captionTargetBox="[160,1436,628,1748]" captionTargetId="figure@11.[151,1436,628,1749]" captionTargetPageId="11" captionText="FIGURE 4. Systenus eboritibia sp. nov. (holotype): a. male habitus, left lateral; b. hypopygium, left lateral. S. parkeri sp. nov.: c. male habitus, left lateral; d. hypopygium, left lateral; e. male postabdomen, right lateral." httpUri="https://zenodo.org/record/237974/files/figure.png" pageId="2" pageNumber="171">Fig. 4</figureCitation>
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a); cer- cus elongate and projecting well beyond surstylus (
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<figureCitation id="07BF2A6EFFBFFF8CFD66A497075CFC7E" box="[716,773,925,946]" captionStart="FIGURE 4" captionStartId="11.[151,250,1770,1792]" captionTargetBox="[160,1436,628,1748]" captionTargetId="figure@11.[151,1436,628,1749]" captionTargetPageId="11" captionText="FIGURE 4. Systenus eboritibia sp. nov. (holotype): a. male habitus, left lateral; b. hypopygium, left lateral. S. parkeri sp. nov.: c. male habitus, left lateral; d. hypopygium, left lateral; e. male postabdomen, right lateral." httpUri="https://zenodo.org/record/237974/files/figure.png" pageId="2" pageNumber="171">Fig. 4</figureCitation>
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b)....................................
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<taxonomicName id="58844D68FFBFFF8CFB79A4960113FC7E" ID-CoL="9HVQV" box="[1235,1354,924,946]" class="Insecta" family="Dolichopodidae" genus="Systenus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="171" phylum="Arthropoda" rank="species" species="eboritibia" status="sp. nov.">
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<emphasis id="ADF0EAF9FFBFFF8CFB79A4960113FC7E" box="[1235,1354,924,946]" italics="true" pageId="2" pageNumber="171">S. eboritibia</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="B6C35782FFBFFF8CFAE5A49601C5FC7E" box="[1359,1436,924,946]" pageId="2" pageNumber="171" rank="species">
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<emphasis id="ADF0EAF9FFBFFF8CFAE5A49601C5FC7E" bold="true" box="[1359,1436,924,946]" pageId="2" pageNumber="171">sp. nov.</emphasis>
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</taxonomicNameLabel>
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</keyLead>
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</paragraph>
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<paragraph id="9F3B36EBFFBFFF8CFF3DA4B001C5FC21" blockId="2.[151,1436,602,1330]" pageId="2" pageNumber="171">
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<keyLead id="247595DEFFBFFF8CFF3DA4B001C5FC21" pageId="2" pageNumber="171">
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- Tibia I only with white cuticle and ivory colored vestiture; tibia II yellow and tibia III brownish (
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<figureCitation id="07BF2A6EFFBFFF8CFB2AA4B000EFFC03" box="[1152,1206,954,975]" captionStart="FIGURE 1" captionStartId="4.[151,250,1872,1894]" captionTargetBox="[238,1352,195,1838]" captionTargetId="figure@4.[234,1352,195,1839]" captionTargetPageId="4" captionText="FIGURE 1. Systenus tenorio sp. nov. (holotype): a. male habitus, left lateral (terminalia removed); b. hypopygium, left lateral. S. flavifemoratus sp. nov. (holotype): c. male habitus, right lateral; d. hypopygium, left lateral. S. maculipennis sp. nov.: e. male habitus, left lateral (terminalia removed); f. hypopygium, left lateral. Abbreviations: cer, cercus; epan, epandrium; hyp, hypandrium; ph, phallus; st, sternite; sur, surstylus; tg, tergite; 6, 7, 8, abdominal segments." httpUri="https://zenodo.org/record/237971/files/figure.png" pageId="2" pageNumber="171">Fig. 1</figureCitation>
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a); cercus and short and digitiform, not extending beyond surstylus (
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<figureCitation id="07BF2A6EFFBFFF8CFD2CA4DD06E5FC20" box="[646,700,983,1004]" captionStart="FIGURE 1" captionStartId="4.[151,250,1872,1894]" captionTargetBox="[238,1352,195,1838]" captionTargetId="figure@4.[234,1352,195,1839]" captionTargetPageId="4" captionText="FIGURE 1. Systenus tenorio sp. nov. (holotype): a. male habitus, left lateral (terminalia removed); b. hypopygium, left lateral. S. flavifemoratus sp. nov. (holotype): c. male habitus, right lateral; d. hypopygium, left lateral. S. maculipennis sp. nov.: e. male habitus, left lateral (terminalia removed); f. hypopygium, left lateral. Abbreviations: cer, cercus; epan, epandrium; hyp, hypandrium; ph, phallus; st, sternite; sur, surstylus; tg, tergite; 6, 7, 8, abdominal segments." httpUri="https://zenodo.org/record/237971/files/figure.png" pageId="2" pageNumber="171">Fig. 1</figureCitation>
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b)............................................
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<taxonomicName id="58844D68FFBFFF8CFB46A4DD0110FC21" ID-CoL="9HZLJ" box="[1260,1353,983,1005]" class="Insecta" family="Dolichopodidae" genus="Systenus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="171" phylum="Arthropoda" rank="species" species="tenorio" status="sp. nov.">
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<emphasis id="ADF0EAF9FFBFFF8CFB46A4DD0110FC21" box="[1260,1353,983,1005]" italics="true" pageId="2" pageNumber="171">S. tenorio</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="B6C35782FFBFFF8CFAE5A4DD01C5FC21" box="[1359,1436,983,1005]" pageId="2" pageNumber="171" rank="species">
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<emphasis id="ADF0EAF9FFBFFF8CFAE5A4DD01C5FC21" bold="true" box="[1359,1436,983,1005]" pageId="2" pageNumber="171">sp. nov.</emphasis>
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</taxonomicNameLabel>
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</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="24702E4EFFBFFF8CFF3DA4FF01CCFB88" pageId="2" pageNumber="171">
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<paragraph id="9F3B36EBFFBFFF8CFF3DA4FF01C5FBE4" blockId="2.[151,1436,602,1330]" pageId="2" pageNumber="171">
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<keyLead id="247595DEFFBFFF8CFF3DA4FF01C5FBE4" pageId="2" pageNumber="171">
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6 Vein M apically bent and forming sharp point on wing margin (
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<figureCitation id="07BF2A6EFFBFFF8CFCE2A4FF07D8FBC6" box="[840,897,1013,1034]" captionStart="FIGURE 3" captionStartId="8.[151,250,1858,1880]" captionTargetBox="[166,1436,813,1836]" captionTargetId="figure@8.[151,1436,813,1836]" captionTargetPageId="8" captionText="FIGURE 3. Systenus divericatus sp. nov. (holotype): a. male habitus, left lateral; b. hypopygium, left lateral. S. zurqui sp. nov.: c. male habitus, left lateral; d. hypopygium, left lateral." httpUri="https://zenodo.org/record/237973/files/figure.png" pageId="2" pageNumber="171">Fig. 3</figureCitation>
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a); tibia III entirely yellow; postpedicel gradually taper- ing; tibia and tarsus I with ivory colored vestiture; cercus flagellate with long setae (
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<figureCitation id="07BF2A6EFFBFFF8CFBA2A3180066FBEB" box="[1032,1087,1042,1063]" captionStart="FIGURE 3" captionStartId="8.[151,250,1858,1880]" captionTargetBox="[166,1436,813,1836]" captionTargetId="figure@8.[151,1436,813,1836]" captionTargetPageId="8" captionText="FIGURE 3. Systenus divericatus sp. nov. (holotype): a. male habitus, left lateral; b. hypopygium, left lateral. S. zurqui sp. nov.: c. male habitus, left lateral; d. hypopygium, left lateral." httpUri="https://zenodo.org/record/237973/files/figure.png" pageId="2" pageNumber="171">Fig. 3</figureCitation>
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b)........
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<taxonomicName id="58844D68FFBFFF8CFB6CA31B0111FBEB" ID-CoL="9HYSV" box="[1222,1352,1041,1063]" class="Insecta" family="Dolichopodidae" genus="Systenus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="171" phylum="Arthropoda" rank="species" species="divericatus" status="sp. nov.">
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<emphasis id="ADF0EAF9FFBFFF8CFB6CA31B0111FBEB" box="[1222,1352,1041,1063]" italics="true" pageId="2" pageNumber="171">S. divericatus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="B6C35782FFBFFF8CFAE5A31801C5FBE4" box="[1359,1436,1042,1064]" pageId="2" pageNumber="171" rank="species">
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<emphasis id="ADF0EAF9FFBFFF8CFAE5A31801C5FBE4" bold="true" box="[1359,1436,1042,1064]" pageId="2" pageNumber="171">sp. nov.</emphasis>
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</taxonomicNameLabel>
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</keyLead>
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</paragraph>
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<paragraph id="9F3B36EBFFBFFF8CFF3DA32501CCFB88" blockId="2.[151,1436,602,1330]" box="[151,1429,1071,1096]" pageId="2" pageNumber="171">
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<keyLead id="247595DEFFBFFF8CFF3DA32501CCFB88" box="[151,1429,1071,1096]" pageId="2" pageNumber="171">- Vein M and R4+5 subparallel and wing margin entire; tibia III with some brown cuticle; other features various........... 7</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="24702E4EFFBFFF8CFF3DA34501C2FB70" pageId="2" pageNumber="171">
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<paragraph id="9F3B36EBFFBFFF8CFF3DA34501C5FB4E" blockId="2.[151,1436,602,1330]" pageId="2" pageNumber="171">
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<keyLead id="247595DEFFBFFF8CFF3DA34501C5FB4E" pageId="2" pageNumber="171">
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7 Tibia I with normal short black vestiture; segment 7 (hypopygial peduncle) elongate, almost twice length of epandrium (
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<figureCitation id="07BF2A6EFFBFFF8CFADFA34504B4FB4D" captionStart="FIGURE 3" captionStartId="8.[151,250,1858,1880]" captionTargetBox="[166,1436,813,1836]" captionTargetId="figure@8.[151,1436,813,1836]" captionTargetPageId="8" captionText="FIGURE 3. Systenus divericatus sp. nov. (holotype): a. male habitus, left lateral; b. hypopygium, left lateral. S. zurqui sp. nov.: c. male habitus, left lateral; d. hypopygium, left lateral." httpUri="https://zenodo.org/record/237973/files/figure.png" pageId="2" pageNumber="171">Fig. 3</figureCitation>
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c), cercus elongate and clavate with subapical field of strong pale setae (
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<figureCitation id="07BF2A6EFFBFFF8CFC08A3660783FB4D" box="[930,986,1132,1153]" captionStart="FIGURE 3" captionStartId="8.[151,250,1858,1880]" captionTargetBox="[166,1436,813,1836]" captionTargetId="figure@8.[151,1436,813,1836]" captionTargetPageId="8" captionText="FIGURE 3. Systenus divericatus sp. nov. (holotype): a. male habitus, left lateral; b. hypopygium, left lateral. S. zurqui sp. nov.: c. male habitus, left lateral; d. hypopygium, left lateral." httpUri="https://zenodo.org/record/237973/files/figure.png" pageId="2" pageNumber="171">Fig. 3</figureCitation>
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d)....................
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<taxonomicName id="58844D68FFBFFF8CFB5FA3660113FB4E" ID-CoL="9HVW9" box="[1269,1354,1132,1154]" class="Insecta" family="Dolichopodidae" genus="Systenus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="171" phylum="Arthropoda" rank="species" species="zurqui" status="sp. nov.">
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<emphasis id="ADF0EAF9FFBFFF8CFB5FA3660113FB4E" box="[1269,1354,1132,1154]" italics="true" pageId="2" pageNumber="171">S. zurqui</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="B6C35782FFBFFF8CFAE5A36601C5FB4E" box="[1359,1436,1132,1154]" pageId="2" pageNumber="171" rank="species">
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<emphasis id="ADF0EAF9FFBFFF8CFAE5A36601C5FB4E" bold="true" box="[1359,1436,1132,1154]" pageId="2" pageNumber="171">sp. nov.</emphasis>
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</taxonomicNameLabel>
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</keyLead>
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</paragraph>
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<paragraph id="9F3B36EBFFBFFF8CFF3DA38001C2FB70" blockId="2.[151,1436,602,1330]" pageId="2" pageNumber="171">
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<keyLead id="247595DEFFBFFF8CFF3DA38001C2FB70" pageId="2" pageNumber="171">- Tibia I with some ivory colored vestiture, segment 7 subequal in length to epandrium; cercus elongate with only short pale setae............................................................................................... 8</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="24702E4EFFBFFF8CFF3DA3CE01C5FAFE" pageId="2" pageNumber="171">
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<paragraph id="9F3B36EBFFBFFF8CFF3DA3CE01C5FB34" blockId="2.[151,1436,602,1330]" pageId="2" pageNumber="171">
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<keyLead id="247595DEFFBFFF8CFF3DA3CE01C5FB34" pageId="2" pageNumber="171">
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8 Tibia I only with ivory-colored vestiture on anterior face; tibia III with brown cuticle only on distal quarter (
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<figureCitation id="07BF2A6EFFBFFF8CFB45A3CE0171FB15" box="[1263,1320,1220,1241]" captionStart="FIGURE 2" captionStartId="6.[151,250,1973,1995]" captionTargetBox="[167,1436,373,1951]" captionTargetId="figure@6.[151,1436,373,1952]" captionTargetPageId="6" captionText="FIGURE 2. Systenus emusorum sp. nov. (holotype): a. male habitus, left lateral; b. hypopygium, left lateral. S. naranjensis sp. nov. (holotype): c. male habitus, left lateral; d. hypopygium, left lateral; e. male head, anterior." httpUri="https://zenodo.org/record/237972/files/figure.png" pageId="2" pageNumber="171">Fig. 2</figureCitation>
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a), surstylus squarish with long dorsoapical projection (
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<figureCitation id="07BF2A6EFFBFFF8CFDD1A3E806EDFB3B" box="[635,692,1250,1271]" captionStart="FIGURE 2" captionStartId="6.[151,250,1973,1995]" captionTargetBox="[167,1436,373,1951]" captionTargetId="figure@6.[151,1436,373,1952]" captionTargetPageId="6" captionText="FIGURE 2. Systenus emusorum sp. nov. (holotype): a. male habitus, left lateral; b. hypopygium, left lateral. S. naranjensis sp. nov. (holotype): c. male habitus, left lateral; d. hypopygium, left lateral; e. male head, anterior." httpUri="https://zenodo.org/record/237972/files/figure.png" pageId="2" pageNumber="171">Fig. 2</figureCitation>
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b)..........................................
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<taxonomicName id="58844D68FFBFFF8CFB67A3EB0110FB3B" ID-CoL="9HXT9" box="[1229,1353,1249,1271]" class="Insecta" family="Dolichopodidae" genus="Systenus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="171" phylum="Arthropoda" rank="species" species="emusorum" status="sp. nov.">
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<emphasis id="ADF0EAF9FFBFFF8CFB67A3EB0110FB3B" box="[1229,1353,1249,1271]" italics="true" pageId="2" pageNumber="171">S. emusorum</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="B6C35782FFBFFF8CFAE5A3E801C5FB34" box="[1359,1436,1250,1272]" pageId="2" pageNumber="171" rank="species">
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<emphasis id="ADF0EAF9FFBFFF8CFAE5A3E801C5FB34" bold="true" box="[1359,1436,1250,1272]" pageId="2" pageNumber="171">sp. nov.</emphasis>
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</taxonomicNameLabel>
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</keyLead>
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</paragraph>
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<paragraph id="9F3B36EBFFBFFF8CFF3DA3F501C5FAFE" blockId="2.[151,1436,602,1330]" pageId="2" pageNumber="171">
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<keyLead id="247595DEFFBFFF8CFF3DA3F501C5FAFE" pageId="2" pageNumber="171">
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- Femur, tibia and tarsus I almost entirely covered with short ivory-colored vestiture; tibia III with brown cuticle on basal quarter and distal quarter (
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<figureCitation id="07BF2A6EFFBFFF8CFE3EA2160597FAFD" box="[404,462,1308,1329]" captionStart="FIGURE 2" captionStartId="6.[151,250,1973,1995]" captionTargetBox="[167,1436,373,1951]" captionTargetId="figure@6.[151,1436,373,1952]" captionTargetPageId="6" captionText="FIGURE 2. Systenus emusorum sp. nov. (holotype): a. male habitus, left lateral; b. hypopygium, left lateral. S. naranjensis sp. nov. (holotype): c. male habitus, left lateral; d. hypopygium, left lateral; e. male head, anterior." httpUri="https://zenodo.org/record/237972/files/figure.png" pageId="2" pageNumber="171">Fig. 2</figureCitation>
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c), surstylus elongate and curved (
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<figureCitation id="07BF2A6EFFBFFF8CFCBFA2160717FAFD" box="[789,846,1308,1329]" captionStart="FIGURE 2" captionStartId="6.[151,250,1973,1995]" captionTargetBox="[167,1436,373,1951]" captionTargetId="figure@6.[151,1436,373,1952]" captionTargetPageId="6" captionText="FIGURE 2. Systenus emusorum sp. nov. (holotype): a. male habitus, left lateral; b. hypopygium, left lateral. S. naranjensis sp. nov. (holotype): c. male habitus, left lateral; d. hypopygium, left lateral; e. male head, anterior." httpUri="https://zenodo.org/record/237972/files/figure.png" pageId="2" pageNumber="171">Fig. 2</figureCitation>
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d).............................
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<taxonomicName id="58844D68FFBFFF8CFB17A2160110FAFE" ID-CoL="9HSKT" box="[1213,1353,1308,1330]" class="Insecta" family="Dolichopodidae" genus="Systenus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="171" phylum="Arthropoda" rank="species" species="naranjensis" status="sp. nov.">
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<emphasis id="ADF0EAF9FFBFFF8CFB17A2160110FAFE" box="[1213,1353,1308,1330]" italics="true" pageId="2" pageNumber="171">S. naranjensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="B6C35782FFBFFF8CFAE5A21601C5FAFE" box="[1359,1436,1308,1330]" pageId="2" pageNumber="171" rank="species">
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<emphasis id="ADF0EAF9FFBFFF8CFAE5A21601C5FAFE" bold="true" box="[1359,1436,1308,1330]" pageId="2" pageNumber="171">sp. nov.</emphasis>
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</taxonomicNameLabel>
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</keyLead>
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</paragraph>
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</keyStep>
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</subSubSection>
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</treatment>
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</document> |