273 lines
36 KiB
XML
273 lines
36 KiB
XML
<document id="5852BB5BEB029BABB74A8544AA2883E7" ID-DOI="10.5281/zenodo.169958" ID-GBIF-Dataset="c3f67774-ce96-432d-a2ab-9c176d7e3219" ID-ISSN="11755326" ID-Zenodo-Dep="169958" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459749106037" checkinUser="plazi" docAuthor="Pitts, James P." docDate="2005" docId="FE5B5F28FFF0B722FEEDF6C0FB8DFC4B" docLanguage="en" docName="zt01050p043.pdf" docOrigin="Zootaxa 1050" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Sphaeropthalma nocticaro Pitts 2005, New Species" docType="treatment" docVersion="11" lastPageNumber="43" masterDocId="02622750FFF1B726FFE5F241FF80FFC5" masterDocTitle="Description of a new species in the Sphaeropthalma pensylvanica speciesgroup (Hymenoptera: Mutillidae)" masterLastPageNumber="43" masterPageNumber="39" pageNumber="40" updateTime="1698211302435" updateUser="plazi">
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<mods:title id="A4BAEBB9D281518032A4F24FA5601302">Description of a new species in the Sphaeropthalma pensylvanica speciesgroup (Hymenoptera: Mutillidae)</mods:title>
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<mods:namePart id="50CB48F80B3F664DDADD992472C38404">Pitts, James P.</mods:namePart>
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<mods:typeOfResource id="18D54149ADBD29BD698D1EC804F9E378">text</mods:typeOfResource>
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<mods:title id="3BCBBC47C47F29ADA5416C46CDC2A945">Zootaxa</mods:title>
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<mods:part id="11FEB4E21BC4394D128F8054126EB612">
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<mods:date id="8A80DAA58E46EE1D9A7C2B02B215165E">2005</mods:date>
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<mods:detail id="A01D065FC008FA88097596034709E1F2" type="pubDate">
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<mods:number id="93FEE42B6A34485996157DB464F10D14">2005-09-16</mods:number>
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<mods:number id="CDA23902434E06D6DC0F9E7B9D26D1D6">1050</mods:number>
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<mods:identifier id="106D2E7478565BFDAC576EDDBCA7CE15" type="DOI">10.5281/zenodo.169958</mods:identifier>
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<treatment id="FE5B5F28FFF0B722FEEDF6C0FB8DFC4B" ID-DOI="http://doi.org/10.5281/zenodo.6267301" ID-GBIF-Taxon="119337288" ID-Zenodo-Dep="6267301" LSID="urn:lsid:plazi:treatment:FE5B5F28FFF0B722FEEDF6C0FB8DFC4B" httpUri="http://treatment.plazi.org/id/FE5B5F28FFF0B722FEEDF6C0FB8DFC4B" lastPageId="4" lastPageNumber="43" pageId="1" pageNumber="40">
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<subSubSection id="3EE8BDB5FFF0B727FEEDF6C0FC3CFB5E" box="[264,956,1153,1179]" pageId="1" pageNumber="40" type="nomenclature">
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<paragraph id="764DEE3EFFF0B727FEEDF6C0FC3CFB5E" blockId="1.[264,956,1153,1179]" box="[264,956,1153,1179]" pageId="1" pageNumber="40">
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<heading id="2D055952FFF0B727FEEDF6C0FC3CFB5E" bold="true" box="[264,956,1153,1179]" fontSize="11" level="1" pageId="1" pageNumber="40" reason="1">
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<emphasis id="4486322CFFF0B727FEEDF6C0FC3CFB5E" bold="true" box="[264,956,1153,1179]" pageId="1" pageNumber="40">
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<taxonomicName id="B1F295BDFFF0B727FEEDF6C0FD0CFB5E" authority="Pitts" authorityName="Pitts" authorityYear="2005" box="[264,652,1153,1179]" class="Insecta" family="Mutillidae" genus="Sphaeropthalma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="40" phylum="Arthropoda" rank="species" species="nocticaro" status="sp. nov.">
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<emphasis id="4486322CFFF0B727FEEDF6C0FDCAFB5E" bold="true" box="[264,586,1153,1179]" italics="true" pageId="1" pageNumber="40">Sphaeropthalma nocticaro</emphasis>
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Pitts
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</taxonomicName>
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,
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<taxonomicNameLabel id="5FB58F57FFF0B727FD7CF6C0FCB3FB5E" box="[665,819,1153,1179]" pageId="1" pageNumber="40" rank="species">New Species</taxonomicNameLabel>
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(
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<figureCitation id="EEC9F2BBFFF0B727FCA7F6C0FC34FB5E" box="[834,948,1153,1179]" captionStart-0="FIGURE 1 – 3" captionStart-1="FIGURE 4 – 5" captionStartId-0="2.[264,367,1260,1284]" captionStartId-1="3.[264,367,954,978]" captionTargetBox-0="[264,1322,473,1246]" captionTargetBox-1="[537,1080,443,917]" captionTargetId-0="figure@2.[264,1323,472,1248]" captionTargetId-1="figure@3.[486,1101,441,928]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURE 1 – 3. Holotype of Sphaeropthalma nocticaro, sp. nov., 1. Habitus, lateral view; 2. Head, dorsal view; 3. Head, frontal view." captionText-1="FIGURE 4 – 5. Holotype of Sphaeropthalma nocticaro, sp. nov., genitalia. 4. Dorsal view, right, ventral view, left; 5. Lateral view." httpUri-0="https://zenodo.org/record/169959/files/figure.png" httpUri-1="https://zenodo.org/record/169960/files/figure.png" pageId="1" pageNumber="40">Figs. 1–5</figureCitation>
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)
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="3EE8BDB5FFF0B727FEEDF68DFB3EFAF3" pageId="1" pageNumber="40" type="diagnosis">
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<paragraph id="764DEE3EFFF0B727FEEDF68DFB3EFAF3" blockId="1.[264,1323,1228,1894]" pageId="1" pageNumber="40">
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<emphasis id="4486322CFFF0B727FEEDF68DFE07FB20" box="[264,391,1228,1253]" italics="true" pageId="1" pageNumber="40">Diagnosis.</emphasis>
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This species is easily differentiated from other members of the
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<taxonomicName id="B1F295BDFFF0B727FB91F68DFAAAFB20" box="[1140,1322,1228,1253]" class="Insecta" family="Mutillidae" genus="Sphaeropthalma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="40" phylum="Arthropoda" rank="species" species="pensylvanica">
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<emphasis id="4486322CFFF0B727FB91F68DFAAAFB20" box="[1140,1322,1228,1253]" italics="true" pageId="1" pageNumber="40">S. pensylvanica</emphasis>
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</taxonomicName>
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speciesgroup by the piceous integumental coloration (
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<figureCitation id="EEC9F2BBFFF0B727FC7CF6B5FC60FACB" box="[921,992,1268,1294]" captionStart="FIGURE 1 – 3" captionStartId="2.[264,367,1260,1284]" captionTargetBox="[264,1322,473,1246]" captionTargetId="figure@2.[264,1323,472,1248]" captionTargetPageId="2" captionText="FIGURE 1 – 3. Holotype of Sphaeropthalma nocticaro, sp. nov., 1. Habitus, lateral view; 2. Head, dorsal view; 3. Head, frontal view." httpUri="https://zenodo.org/record/169959/files/figure.png" pageId="1" pageNumber="40">Fig. 1</figureCitation>
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), the white brachyplumose setae, and T3 being completely covered in distinct, white, plumose setae (
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<figureCitation id="EEC9F2BBFFF0B727FB8DF75DFB2DFAF3" box="[1128,1197,1308,1334]" captionStart="FIGURE 1 – 3" captionStartId="2.[264,367,1260,1284]" captionTargetBox="[264,1322,473,1246]" captionTargetId="figure@2.[264,1323,472,1248]" captionTargetPageId="2" captionText="FIGURE 1 – 3. Holotype of Sphaeropthalma nocticaro, sp. nov., 1. Habitus, lateral view; 2. Head, dorsal view; 3. Head, frontal view." httpUri="https://zenodo.org/record/169959/files/figure.png" pageId="1" pageNumber="40">Fig. 1</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="3EE8BDB5FFF0B725FEDDF705FD99FA12" lastPageId="3" lastPageNumber="42" pageId="1" pageNumber="40" type="description">
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<paragraph id="764DEE3EFFF0B727FEDDF705FB6DF98B" blockId="1.[264,1323,1228,1894]" pageId="1" pageNumber="40">
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<emphasis id="4486322CFFF0B727FEDDF705FBA9FA98" box="[312,1065,1348,1373]" italics="true" pageId="1" pageNumber="40">
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Description of Male
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<typeStatus id="A949509CFFF0B727FDD9F705FD2BFA98" box="[572,683,1348,1373]" pageId="1" pageNumber="40" type="holotype">Holotype</typeStatus>
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. Coloration and Setal Pattern.
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</emphasis>
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Integument piceous, flagellum black (
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<figureCitation id="EEC9F2BBFFF0B727FE35F72DFD96FA43" box="[464,534,1388,1414]" captionStart="FIGURE 1 – 3" captionStartId="2.[264,367,1260,1284]" captionTargetBox="[264,1322,473,1246]" captionTargetId="figure@2.[264,1323,472,1248]" captionTargetPageId="2" captionText="FIGURE 1 – 3. Holotype of Sphaeropthalma nocticaro, sp. nov., 1. Habitus, lateral view; 2. Head, dorsal view; 3. Head, frontal view." httpUri="https://zenodo.org/record/169959/files/figure.png" pageId="1" pageNumber="40">Fig. 1</figureCitation>
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). Body covered in sparse erect white brachyplumose setae, except front also with indistinct piceous brachyplumose setae present, and mesonotum, tibiae, tarsi, disk of T2, T4–T7 and S6–S7 also with piceous brachyplumose setae (
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<figureCitation id="EEC9F2BBFFF0B727FB65F7FDFB45FA13" box="[1152,1221,1468,1494]" captionStart="FIGURE 1 – 3" captionStartId="2.[264,367,1260,1284]" captionTargetBox="[264,1322,473,1246]" captionTargetId="figure@2.[264,1323,472,1248]" captionTargetPageId="2" captionText="FIGURE 1 – 3. Holotype of Sphaeropthalma nocticaro, sp. nov., 1. Habitus, lateral view; 2. Head, dorsal view; 3. Head, frontal view." httpUri="https://zenodo.org/record/169959/files/figure.png" pageId="1" pageNumber="40">Fig. 1</figureCitation>
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). Fringe of T1–T3 with dense distinct white plumose setae (
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<figureCitation id="EEC9F2BBFFF0B727FC62F7A5FC51FA3B" box="[903,977,1508,1534]" captionStart="FIGURE 1 – 3" captionStartId="2.[264,367,1260,1284]" captionTargetBox="[264,1322,473,1246]" captionTargetId="figure@2.[264,1323,472,1248]" captionTargetPageId="2" captionText="FIGURE 1 – 3. Holotype of Sphaeropthalma nocticaro, sp. nov., 1. Habitus, lateral view; 2. Head, dorsal view; 3. Head, frontal view." httpUri="https://zenodo.org/record/169959/files/figure.png" pageId="1" pageNumber="40">Fig. 1</figureCitation>
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). T3 disk covered in dense plumose white setae obscuring sculpture; some plumose setae yellow tinged (
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<figureCitation id="EEC9F2BBFFF0B727FB28F44DFA97F9E3" box="[1229,1303,1548,1574]" captionStart="FIGURE 1 – 3" captionStartId="2.[264,367,1260,1284]" captionTargetBox="[264,1322,473,1246]" captionTargetId="figure@2.[264,1323,472,1248]" captionTargetPageId="2" captionText="FIGURE 1 – 3. Holotype of Sphaeropthalma nocticaro, sp. nov., 1. Habitus, lateral view; 2. Head, dorsal view; 3. Head, frontal view." httpUri="https://zenodo.org/record/169959/files/figure.png" pageId="1" pageNumber="40">Fig. 1</figureCitation>
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). Wings infuscate with simple brown setae (
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<figureCitation id="EEC9F2BBFFF0B727FD1DF475FCBDF98B" box="[760,829,1588,1614]" captionStart="FIGURE 1 – 3" captionStartId="2.[264,367,1260,1284]" captionTargetBox="[264,1322,473,1246]" captionTargetId="figure@2.[264,1323,472,1248]" captionTargetPageId="2" captionText="FIGURE 1 – 3. Holotype of Sphaeropthalma nocticaro, sp. nov., 1. Habitus, lateral view; 2. Head, dorsal view; 3. Head, frontal view." httpUri="https://zenodo.org/record/169959/files/figure.png" pageId="1" pageNumber="40">Fig. 1</figureCitation>
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). Pterostigma and veins dark brown.
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</paragraph>
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<paragraph id="764DEE3EFFF0B724FEDDF41DFC08FE43" blockId="1.[264,1323,1228,1894]" lastBlockId="2.[264,1323,284,390]" lastPageId="2" lastPageNumber="41" pageId="1" pageNumber="40">
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<emphasis id="4486322CFFF0B727FEDDF41DFEFFF9B0" box="[312,383,1628,1653]" italics="true" pageId="1" pageNumber="40">Head.</emphasis>
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Subquadrate posteriorly; parallel behind eyes; moderately punctate throughout (
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<figureCitation id="EEC9F2BBFFF0B727FEF4F4C5FE03F95B" box="[273,387,1668,1694]" captionStart="FIGURE 1 – 3" captionStartId="2.[264,367,1260,1284]" captionTargetBox="[264,1322,473,1246]" captionTargetId="figure@2.[264,1323,472,1248]" captionTargetPageId="2" captionText="FIGURE 1 – 3. Holotype of Sphaeropthalma nocticaro, sp. nov., 1. Habitus, lateral view; 2. Head, dorsal view; 3. Head, frontal view." httpUri="https://zenodo.org/record/169959/files/figure.png" pageId="1" pageNumber="40">Figs. 2, 3</figureCitation>
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). Eye small, barely protruding (
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<figureCitation id="EEC9F2BBFFF0B727FD1AF4C5FCF1F95B" box="[767,881,1668,1694]" captionStart="FIGURE 1 – 3" captionStartId="2.[264,367,1260,1284]" captionTargetBox="[264,1322,473,1246]" captionTargetId="figure@2.[264,1323,472,1248]" captionTargetPageId="2" captionText="FIGURE 1 – 3. Holotype of Sphaeropthalma nocticaro, sp. nov., 1. Habitus, lateral view; 2. Head, dorsal view; 3. Head, frontal view." httpUri="https://zenodo.org/record/169959/files/figure.png" pageId="1" pageNumber="40">Figs. 2, 3</figureCitation>
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). Ocellus small, interocellar distance ~2.75X lateral ocellus diameter, ocellocular distance ~4.5X lateral ocellus diameter (
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<figureCitation id="EEC9F2BBFFF0B727FB1EF4EDFE99F92B" captionStart="FIGURE 1 – 3" captionStartId="2.[264,367,1260,1284]" captionTargetBox="[264,1322,473,1246]" captionTargetId="figure@2.[264,1323,472,1248]" captionTargetPageId="2" captionText="FIGURE 1 – 3. Holotype of Sphaeropthalma nocticaro, sp. nov., 1. Habitus, lateral view; 2. Head, dorsal view; 3. Head, frontal view." httpUri="https://zenodo.org/record/169959/files/figure.png" pageId="1" pageNumber="40">Fig. 2</figureCitation>
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). Glabrous triangular region present anterior to median ocellus. Antennal scrobe carinate dorsally; tubercle present. Mandible tridentate (wellworn in
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<typeStatus id="A949509CFFF0B727FC32F4BDFBC2F8D3" box="[983,1090,1788,1814]" pageId="1" pageNumber="40" type="holotype">holotype</typeStatus>
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), broad and vertical throughout (
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<figureCitation id="EEC9F2BBFFF0B727FE79F565FE66F8FB" box="[412,486,1828,1854]" captionStart="FIGURE 1 – 3" captionStartId="2.[264,367,1260,1284]" captionTargetBox="[264,1322,473,1246]" captionTargetId="figure@2.[264,1323,472,1248]" captionTargetPageId="2" captionText="FIGURE 1 – 3. Holotype of Sphaeropthalma nocticaro, sp. nov., 1. Habitus, lateral view; 2. Head, dorsal view; 3. Head, frontal view." httpUri="https://zenodo.org/record/169959/files/figure.png" pageId="1" pageNumber="40">Fig. 3</figureCitation>
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). Mandible with slight ventral excision resulting in basal angulation. Clypeus truncate apically, distinctly curved dorsally, with 2 small tubercles set back from corners of truncation. Clypeus greatly depressed below dorsal margin of mandibles (
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<figureCitation id="EEC9F2BBFFF3B724FB3AF35DFE98FE9B" captionStart="FIGURE 1 – 3" captionStartId="2.[264,367,1260,1284]" captionTargetBox="[264,1322,473,1246]" captionTargetId="figure@2.[264,1323,472,1248]" captionTargetPageId="2" captionText="FIGURE 1 – 3. Holotype of Sphaeropthalma nocticaro, sp. nov., 1. Habitus, lateral view; 2. Head, dorsal view; 3. Head, frontal view." httpUri="https://zenodo.org/record/169959/files/figure.png" pageId="2" pageNumber="41">Fig. 2, 3</figureCitation>
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). Malar space ~
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<date id="024CC8FEFFF3B724FE00F305FD89FE9B" box="[485,521,324,350]" pageId="2" pageNumber="41">2X</date>
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observed length of pedicel. First flagellomere ~2.25X length of pedicle. Second flagellomere slightly longer than first.
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</paragraph>
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<caption id="228DBEB6FFF3B724FEEDF6ADFDF9FAE3" httpUri="https://zenodo.org/record/169959/files/figure.png" pageId="2" pageNumber="41" targetBox="[264,1322,473,1246]" targetPageId="2">
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<paragraph id="764DEE3EFFF3B724FEEDF6ADFDF9FAE3" blockId="2.[264,1322,1260,1318]" pageId="2" pageNumber="41">
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<emphasis id="4486322CFFF3B724FEEDF6ADFE26FAC1" bold="true" box="[264,422,1260,1284]" pageId="2" pageNumber="41">FIGURE 1–3.</emphasis>
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Holotype of
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<taxonomicName id="B1F295BDFFF3B724FDDDF6ACFCD4FAC1" authority="Pitts, 2005" authorityName="Pitts" authorityYear="2005" box="[568,852,1261,1284]" class="Insecta" family="Mutillidae" genus="Sphaeropthalma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="41" phylum="Arthropoda" rank="species" species="nocticaro" status="sp. nov.">
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<emphasis id="4486322CFFF3B724FDDDF6ACFCD4FAC1" box="[568,852,1261,1284]" italics="true" pageId="2" pageNumber="41">Sphaeropthalma nocticaro</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="5FB58F57FFF3B724FC87F6ADFC38FAC6" box="[866,952,1260,1283]" pageId="2" pageNumber="41" rank="species">sp. nov.</taxonomicNameLabel>
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, 1. Habitus, lateral view; 2. Head, dorsal view; 3. Head, frontal view.
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</paragraph>
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</caption>
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<paragraph id="764DEE3EFFF3B724FEDDF7C3FC0CF949" blockId="2.[264,1325,1410,1876]" pageId="2" pageNumber="41">
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<emphasis id="4486322CFFF3B724FEDDF7C3FE3DFA5E" box="[312,445,1410,1435]" italics="true" pageId="2" pageNumber="41">
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<taxonomicName id="B1F295BDFFF3B724FEDDF7C3FE39FA5E" box="[312,441,1410,1435]" class="Crustacea" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="41" phylum="Arthropoda" rank="genus">Mesosoma</taxonomicName>
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.
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</emphasis>
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Moderately punctate. Notali complete. Parapsidal furrows present on posterior ~0.66 of mesonotum. Pronotum nitid anterior to epaulets, area almost reaching anterior margin. Propodeum coarsely punctate dorsally, vertical and lateral faces reticulate, becoming coarsely punctate ventrally. Dorsal face with median area longitudinally concave. Mesosternum unarmed. Metasternum truncate with small median tooth present. Coxae and trochanters unarmed. Marginal cell ~
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<date id="024CC8FEFFF3B724FBE9F40BFBB1F9A1" box="[1036,1073,1610,1636]" pageId="2" pageNumber="41">2X</date>
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length of stigma as measured along costa. Wings with 3 submarginal cells.
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</paragraph>
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<paragraph id="764DEE3EFFF3B725FEDDF4DBFBE5FE9B" blockId="2.[264,1325,1410,1876]" lastBlockId="3.[264,1322,284,350]" lastPageId="3" lastPageNumber="42" pageId="2" pageNumber="41">
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<emphasis id="4486322CFFF3B724FEDDF4DBFE3AF976" box="[312,442,1690,1715]" italics="true" pageId="2" pageNumber="41">Metasoma.</emphasis>
|
||
T1 nodose with T2. T1 moderately punctate laterally and apically, otherwise sparsely punctate with interstitial area nitid. T2 with small dense punctures; interstitial width less than diameter of punctures, nitid. T3–T6 micropunctate, appearing granulate. T7 wider than long; nitid, except for apical margin with setigerous punctures bearing fringe of short hooked setae. S2 felt line absent, although micropunctures present in felt line area. Punctation of sternites similar to respective tergites, except S7 with small punctations. S7 wider than long. Apical margin of S7 broadly triangulate.
|
||
</paragraph>
|
||
<caption id="228DBEB6FFF2B725FEEDF1FBFDF0FC32" httpUri="https://zenodo.org/record/169960/files/figure.png" pageId="3" pageNumber="42" targetBox="[537,1080,443,917]" targetPageId="3">
|
||
<paragraph id="764DEE3EFFF2B725FEEDF1FBFDF0FC32" blockId="3.[264,1322,952,1015]" pageId="3" pageNumber="42">
|
||
<emphasis id="4486322CFFF2B725FEEDF1FBFE2AFC17" bold="true" box="[264,426,952,978]" pageId="3" pageNumber="42">FIGURE 4–5.</emphasis>
|
||
Holotype of
|
||
<taxonomicName id="B1F295BDFFF2B725FDA5F1FBFCDEFC14" authority="Pitts, 2005" authorityName="Pitts" authorityYear="2005" box="[576,862,954,977]" class="Insecta" family="Mutillidae" genus="Sphaeropthalma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="42" phylum="Arthropoda" rank="species" species="nocticaro" status="sp. nov.">
|
||
<emphasis id="4486322CFFF2B725FDA5F1FBFCDEFC14" box="[576,862,954,977]" italics="true" pageId="3" pageNumber="42">Sphaeropthalma nocticaro</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="5FB58F57FFF2B725FC8AF1FBFC47FC14" box="[879,967,954,977]" pageId="3" pageNumber="42" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, genitalia. 4. Dorsal view, right, ventral view, left; 5. Lateral view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="764DEE3EFFF2B725FEDDF617FB82FA42" blockId="3.[264,1323,1109,1895]" pageId="3" pageNumber="42">
|
||
<emphasis id="4486322CFFF2B725FEDDF617FE2EFBAA" box="[312,430,1110,1135]" italics="true" pageId="3" pageNumber="42">Genitalia.</emphasis>
|
||
Paramere arcuate, stout at base, tapering toward apex, curved dorsally (
|
||
<figureCitation id="EEC9F2BBFFF2B725FEF4F63CFE0EFB52" box="[273,398,1149,1175]" captionStart="FIGURE 4 – 5" captionStartId="3.[264,367,954,978]" captionTargetBox="[537,1080,443,917]" captionTargetId="figure@3.[486,1101,441,928]" captionTargetPageId="3" captionText="FIGURE 4 – 5. Holotype of Sphaeropthalma nocticaro, sp. nov., genitalia. 4. Dorsal view, right, ventral view, left; 5. Lateral view." httpUri="https://zenodo.org/record/169960/files/figure.png" pageId="3" pageNumber="42">Figs. 4, 5</figureCitation>
|
||
). Dense setae present on ventral margin of paramere (
|
||
<figureCitation id="EEC9F2BBFFF2B725FBA4F63CFB3EFB52" box="[1089,1214,1149,1175]" captionStart="FIGURE 4 – 5" captionStartId="3.[264,367,954,978]" captionTargetBox="[537,1080,443,917]" captionTargetId="figure@3.[486,1101,441,928]" captionTargetPageId="3" captionText="FIGURE 4 – 5. Holotype of Sphaeropthalma nocticaro, sp. nov., genitalia. 4. Dorsal view, right, ventral view, left; 5. Lateral view." httpUri="https://zenodo.org/record/169960/files/figure.png" pageId="3" pageNumber="42">Figs. 4, 5</figureCitation>
|
||
). Cuspis elongate, 0.8X free length of paramere, basal portion cylindrical, dilated and flatted towards apex, inner and outer margin with long dense setae, tips of setae plumose with many branches, basal portion with plumose tipped setae ventrally, small basal pit present with long simple to brachyplumose setae, dorsum asetose (
|
||
<figureCitation id="EEC9F2BBFFF2B725FC23F75CFBBAFAF2" box="[966,1082,1309,1335]" captionStart="FIGURE 4 – 5" captionStartId="3.[264,367,954,978]" captionTargetBox="[537,1080,443,917]" captionTargetId="figure@3.[486,1101,441,928]" captionTargetPageId="3" captionText="FIGURE 4 – 5. Holotype of Sphaeropthalma nocticaro, sp. nov., genitalia. 4. Dorsal view, right, ventral view, left; 5. Lateral view." httpUri="https://zenodo.org/record/169960/files/figure.png" pageId="3" pageNumber="42">Figs. 4, 5</figureCitation>
|
||
). Basal lobe present on cuspis, setose (
|
||
<figureCitation id="EEC9F2BBFFF2B725FE01F704FDD9FA9A" box="[484,601,1349,1375]" captionStart="FIGURE 4 – 5" captionStartId="3.[264,367,954,978]" captionTargetBox="[537,1080,443,917]" captionTargetId="figure@3.[486,1101,441,928]" captionTargetPageId="3" captionText="FIGURE 4 – 5. Holotype of Sphaeropthalma nocticaro, sp. nov., genitalia. 4. Dorsal view, right, ventral view, left; 5. Lateral view." httpUri="https://zenodo.org/record/169960/files/figure.png" pageId="3" pageNumber="42">Figs. 4, 5</figureCitation>
|
||
). Digitus cylindrical, short, minutely pubescent (
|
||
<figureCitation id="EEC9F2BBFFF2B725FB43F704FA9CFA9A" box="[1190,1308,1349,1375]" captionStart="FIGURE 4 – 5" captionStartId="3.[264,367,954,978]" captionTargetBox="[537,1080,443,917]" captionTargetId="figure@3.[486,1101,441,928]" captionTargetPageId="3" captionText="FIGURE 4 – 5. Holotype of Sphaeropthalma nocticaro, sp. nov., genitalia. 4. Dorsal view, right, ventral view, left; 5. Lateral view." httpUri="https://zenodo.org/record/169960/files/figure.png" pageId="3" pageNumber="42">Figs. 4, 5</figureCitation>
|
||
). Penis valve bidentate ventrally, not angulate dorsally (
|
||
<figureCitation id="EEC9F2BBFFF2B725FC66F72CFC73FA42" box="[899,1011,1389,1415]" captionStart="FIGURE 4 – 5" captionStartId="3.[264,367,954,978]" captionTargetBox="[537,1080,443,917]" captionTargetId="figure@3.[486,1101,441,928]" captionTargetPageId="3" captionText="FIGURE 4 – 5. Holotype of Sphaeropthalma nocticaro, sp. nov., genitalia. 4. Dorsal view, right, ventral view, left; 5. Lateral view." httpUri="https://zenodo.org/record/169960/files/figure.png" pageId="3" pageNumber="42">Figs. 4, 5</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="764DEE3EFFF2B725FEDDF7D7FD81FA6A" blockId="3.[264,1323,1109,1895]" box="[312,513,1429,1455]" pageId="3" pageNumber="42">
|
||
<emphasis id="4486322CFFF2B725FEDDF7D7FE11FA6A" box="[312,401,1430,1455]" italics="true" pageId="3" pageNumber="42">Length.</emphasis>
|
||
~
|
||
<quantity id="B10A43DBFFF2B725FE42F7D4FE7DFA6A" box="[423,509,1429,1455]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" pageId="3" pageNumber="42" unit="mm" value="14.0">14 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="764DEE3EFFF2B725FEDDF7FFFD99FA12" blockId="3.[264,1323,1109,1895]" box="[312,537,1469,1495]" pageId="3" pageNumber="42">
|
||
<emphasis id="4486322CFFF2B725FEDDF7FFFE17FA12" box="[312,407,1470,1495]" italics="true" pageId="3" pageNumber="42">Female.</emphasis>
|
||
Unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="3EE8BDB5FFF2B725FEDDF7A7FC2DF98A" pageId="3" pageNumber="42" type="materials_examined">
|
||
<paragraph id="764DEE3EFFF2B725FEDDF7A7FC2DF98A" blockId="3.[264,1323,1109,1895]" pageId="3" pageNumber="42">
|
||
<emphasis id="4486322CFFF2B725FEDDF7A7FE6AFA3A" box="[312,490,1510,1535]" italics="true" pageId="3" pageNumber="42">
|
||
<typeStatus id="A949509CFFF2B725FEDDF7A7FEEFFA3A" box="[312,367,1510,1535]" pageId="3" pageNumber="42">Type</typeStatus>
|
||
Material.
|
||
</emphasis>
|
||
<emphasis id="4486322CFFF2B725FE10F7A4FDF1FA3A" bold="true" box="[501,625,1509,1535]" pageId="3" pageNumber="42">
|
||
<typeStatus id="A949509CFFF2B725FE10F7A4FDEBFA3A" box="[501,619,1509,1535]" pageId="3" pageNumber="42" type="holotype">Holotype</typeStatus>
|
||
:
|
||
</emphasis>
|
||
“Temascal, Oax[aca], Mex[ico],
|
||
<date id="024CC8FEFFF2B725FBE8F7A4FBFCFA3A" box="[1037,1148,1509,1535]" pageId="3" pageNumber="42" value="1964-06-22">VI2264</date>
|
||
, D.H. Janzen, Collector”. Deposited into
|
||
<collectionCode id="10E376FBFFF2B725FDA8F44CFD12F9E2" box="[589,658,1549,1575]" pageId="3" pageNumber="42">CISC</collectionCode>
|
||
.
|
||
<typeStatus id="A949509CFFF2B725FD44F44CFC93F9E2" box="[673,787,1549,1575]" pageId="3" pageNumber="42" type="paratype">
|
||
<emphasis id="4486322CFFF2B725FD44F44CFC93F9E2" bold="true" box="[673,787,1549,1575]" pageId="3" pageNumber="42">Paratype</emphasis>
|
||
</typeStatus>
|
||
: “
|
||
<collectingCountry id="0EE5AEAEFFF2B725FCD5F44CFC35F9E2" box="[816,949,1549,1575]" name="Guatemala" pageId="3" pageNumber="42">Guatemala</collectingCountry>
|
||
, Petén, Ruinas de Tikal,
|
||
<date id="024CC8FEFFF2B725FB0DF44CFEDFF98A" pageId="3" pageNumber="42" value="1927-08-29">27/29 August</date>
|
||
, 1974, E.M. Fisher, collr.” Deposited into
|
||
<collectionCode id="10E376FBFFF2B725FCB5F474FC29F98A" LSID="urn:lsid:biocol.org:col:34802" box="[848,937,1589,1615]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="3" pageNumber="42">LACM</collectionCode>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="3EE8BDB5FFF2B725FEDDF41FFD24F95A" pageId="3" pageNumber="42" type="etymology">
|
||
<paragraph id="764DEE3EFFF2B725FEDDF41FFD24F95A" blockId="3.[264,1323,1109,1895]" pageId="3" pageNumber="42">
|
||
<emphasis id="4486322CFFF2B725FEDDF41FFE3BF9B2" box="[312,443,1630,1655]" italics="true" pageId="3" pageNumber="42">Etymology.</emphasis>
|
||
From the Latin
|
||
<emphasis id="4486322CFFF2B725FD9DF41FFD39F9B2" box="[632,697,1630,1655]" italics="true" pageId="3" pageNumber="42">noctu</emphasis>
|
||
“by night” and Latin
|
||
<emphasis id="4486322CFFF2B725FC5CF41FFC6EF9B2" box="[953,1006,1630,1655]" italics="true" pageId="3" pageNumber="42">caro</emphasis>
|
||
“flesh”, in reference to the black coloration of the integument.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="3EE8BDB5FFF2B722FEDDF4EFFB8DFC4B" lastPageId="4" lastPageNumber="43" pageId="3" pageNumber="42" type="discussion">
|
||
<paragraph id="764DEE3EFFF2B722FEDDF4EFFCD2FE13" blockId="3.[264,1323,1109,1895]" lastBlockId="4.[264,1323,284,910]" lastPageId="4" lastPageNumber="43" pageId="3" pageNumber="42">
|
||
<emphasis id="4486322CFFF2B725FEDDF4EFFE26F902" box="[312,422,1710,1735]" italics="true" pageId="3" pageNumber="42">Remarks.</emphasis>
|
||
This species easily keys out to the subgenus
|
||
<taxonomicName id="B1F295BDFFF2B725FC31F4EFFB15F902" box="[980,1173,1710,1735]" class="Insecta" family="Mutillidae" genus="Sphaeropthalma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="42" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="4486322CFFF2B725FC31F4EFFB15F902" box="[980,1173,1710,1735]" italics="true" pageId="3" pageNumber="42">Sphaeropthalma</emphasis>
|
||
</taxonomicName>
|
||
(now the
|
||
<taxonomicName id="B1F295BDFFF2B725FAF1F4EFFE21F92A" class="Insecta" family="Mutillidae" genus="Sphaeropthalma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="42" phylum="Arthropoda" rank="species" species="pensylvanica">
|
||
<emphasis id="4486322CFFF2B725FAF1F4EFFE21F92A" italics="true" pageId="3" pageNumber="42">S. pensylvanica</emphasis>
|
||
</taxonomicName>
|
||
speciesgroup; see
|
||
<bibRefCitation id="126393CFFFF2B725FD6EF494FCCAF92A" author="Pitts" box="[651,842,1749,1775]" pageId="3" pageNumber="43" refString="Pitts, J. P., Parker, F. D. & Pitts-Singer, T. L. (2004) A review of the Sphaeropthalma uro speciesgroup (Hymenoptera: Mutillidae), with taxonomic changes. Journal of the Kansas Entomological Society, 77, 223 - 234." type="journal article" year="2004">
|
||
Pitts
|
||
<emphasis id="4486322CFFF2B725FD2CF497FC85F92A" box="[713,773,1750,1775]" italics="true" pageId="3" pageNumber="42">et al.</emphasis>
|
||
2004
|
||
</bibRefCitation>
|
||
) in
|
||
<bibRefCitation id="126393CFFFF2B725FC9EF494FB1FF92A" author="Manley" box="[891,1183,1749,1775]" pageId="3" pageNumber="43" refString="Manley, D. G. & Pitts, J. P. (2002) Key to the genera and subgenera of Mutillidae of America North of Mexico. Journal of Hymenoptera Research, 11, 72 - 101." type="journal article" year="2002">Manley and Pitts (2002)</bibRefCitation>
|
||
. In general, this species is quite similar morphologically to other species in this speciesgroup, except for the coloration and white plumose setal mentioned previously. This species is also probably diurnal like the other members of this group as deduced from the very small ocelli and dark integumental coloration. The genitalia also differ from the other species. For
|
||
<taxonomicName id="B1F295BDFFF5B722FEDDF305FE44FE98" authorityName="Pitts" authorityYear="2005" box="[312,452,324,349]" class="Insecta" family="Mutillidae" genus="Sphaeropthalma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="43" phylum="Arthropoda" rank="species" species="nocticaro">
|
||
<emphasis id="4486322CFFF5B722FEDDF305FE44FE98" box="[312,452,324,349]" italics="true" pageId="4" pageNumber="43">S. nocticaro</emphasis>
|
||
</taxonomicName>
|
||
, the paramere has ventral setae present much further towards the apex, the basal portion of cuspis is not as densely setose ventrally, and the digitus is much larger in comparison to the cuspis. There is also a slight difference in the shape of the apex of the cuspis between
|
||
<taxonomicName id="B1F295BDFFF5B722FE27F3FDFDCEFE10" authorityName="Pitts" authorityYear="2005" box="[450,590,444,469]" class="Insecta" family="Mutillidae" genus="Sphaeropthalma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="43" phylum="Arthropoda" rank="species" species="nocticaro">
|
||
<emphasis id="4486322CFFF5B722FE27F3FDFDCEFE10" box="[450,590,444,469]" italics="true" pageId="4" pageNumber="43">S. nocticaro</emphasis>
|
||
</taxonomicName>
|
||
and the other species.
|
||
</paragraph>
|
||
<paragraph id="764DEE3EFFF5B722FEDDF3A5FB8DFC4B" blockId="4.[264,1323,284,910]" pageId="4" pageNumber="43">
|
||
As with most organisms, morphological variation has been documented for many of the
|
||
<taxonomicName id="B1F295BDFFF5B722FED0F04DFE76FDE0" box="[309,502,524,549]" class="Insecta" family="Mutillidae" genus="Sphaeropthalma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="43" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="4486322CFFF5B722FED0F04DFE76FDE0" box="[309,502,524,549]" italics="true" pageId="4" pageNumber="43">Sphaeropthalma</emphasis>
|
||
</taxonomicName>
|
||
species. Because only two specimens are available, variation cannot be adequately addressed for
|
||
<taxonomicName id="B1F295BDFFF5B722FD8EF075FD7CFD88" authorityName="Pitts" authorityYear="2005" box="[619,764,564,589]" class="Insecta" family="Mutillidae" genus="Sphaeropthalma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="43" phylum="Arthropoda" rank="species" species="nocticaro">
|
||
<emphasis id="4486322CFFF5B722FD8EF075FD7CFD88" box="[619,764,564,589]" italics="true" pageId="4" pageNumber="43">S. nocticaro</emphasis>
|
||
</taxonomicName>
|
||
at this time. Furthermore, the
|
||
<typeStatus id="A949509CFFF5B722FB68F075FB74FD8B" box="[1165,1268,564,590]" pageId="4" pageNumber="43" type="holotype">holotype</typeStatus>
|
||
and
|
||
<typeStatus id="A949509CFFF5B722FEEDF01DFEEDFDB3" box="[264,365,604,630]" pageId="4" pageNumber="43" type="paratype">paratype</typeStatus>
|
||
are quite similar morphologically, differing only in wear of the mandibular teeth. Normal variation with male
|
||
<taxonomicName id="B1F295BDFFF5B722FDB9F0C5FC9DFD58" box="[604,797,644,669]" class="Insecta" family="Mutillidae" genus="Sphaeropthalma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="43" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="4486322CFFF5B722FDB9F0C5FC9DFD58" box="[604,797,644,669]" italics="true" pageId="4" pageNumber="43">Sphaeropthalma</emphasis>
|
||
</taxonomicName>
|
||
usually includes a large range in body size. For smaller individuals, the diameter and coarseness of punctations are normally reduced, and the integumental coloration is lighter (
|
||
<bibRefCitation id="126393CFFFF5B722FCFEF095FC5DFD2B" author="Ferguson" box="[795,989,724,750]" pageId="4" pageNumber="43" refString="Ferguson, W. E. (1962) Biological characteristics of the mutillid subgenus Photopsis Blake and their systematic values (Hymenoptera). University of California Publications in Entomology, 27, 1 - 92." type="journal article" year="1962">Ferguson, 1962</bibRefCitation>
|
||
). Variation of this
|
||
<typeStatus id="A949509CFFF5B722FB3DF095FA8BFD2B" box="[1240,1291,724,750]" pageId="4" pageNumber="43">type</typeStatus>
|
||
is expected for
|
||
<taxonomicName id="B1F295BDFFF5B722FE4EF0BDFDBBFCD0" authorityName="Pitts" authorityYear="2005" box="[427,571,764,789]" class="Insecta" family="Mutillidae" genus="Sphaeropthalma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="43" phylum="Arthropoda" rank="species" species="nocticaro">
|
||
<emphasis id="4486322CFFF5B722FE4EF0BDFDBBFCD0" box="[427,571,764,789]" italics="true" pageId="4" pageNumber="43">S. nocticaro</emphasis>
|
||
</taxonomicName>
|
||
once more specimens are found. The diagnostic characters, however, used to differentiate this species from other members of the
|
||
<taxonomicName id="B1F295BDFFF5B722FB8FF165FE21FCA0" class="Insecta" family="Mutillidae" genus="Sphaeropthalma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="43" phylum="Arthropoda" rank="species" species="pensylvanica">
|
||
<emphasis id="4486322CFFF5B722FB8FF165FE21FCA0" italics="true" pageId="4" pageNumber="43">Sphaeropthalma pensylvanica</emphasis>
|
||
</taxonomicName>
|
||
speciesgroup are unlikely to vary given the normal variation seen in other members of this group, as well as for
|
||
<taxonomicName id="B1F295BDFFF5B722FD27F135FC03FC48" box="[706,899,884,909]" class="Insecta" family="Mutillidae" genus="Sphaeropthalma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="43" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="4486322CFFF5B722FD27F135FC03FC48" box="[706,899,884,909]" italics="true" pageId="4" pageNumber="43">Sphaeropthalma</emphasis>
|
||
</taxonomicName>
|
||
as a whole.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |