158 lines
16 KiB
XML
158 lines
16 KiB
XML
<document id="6B11CA04E22B90E3C7F1280B41B44120" ID-DOI="10.5281/zenodo.158726" ID-GBIF-Dataset="346a7f7d-c951-4756-acdb-09c0a30d52cf" ID-ISSN="11755326" ID-Zenodo-Dep="158726" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461038753034" checkinUser="plazi" docAuthor="Erwin, Terry L." docDate="2004" docId="325787BEFF80FFF16C0FFEC7FB1986C1" docLanguage="en" docName="zt00790.pdf" docOrigin="Zootaxa 790" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Epikastea mancocapac Erwin, new species" docType="treatment" docVersion="9" lastPageNumber="16" masterDocId="CE6EFFC6FF8EFFFE6D07FFDBFFFE8717" masterDocTitle="The beetle family Carabidae of Costa Rica: The genus Epikastea Liebke of the Plochonida Group, with new Neotropical species and notes on their way of life (Insecta: Coleoptera, Lebiini, Agrina)" masterLastPageNumber="20" masterPageNumber="1" pageNumber="15" updateTime="1698575098462" updateUser="plazi">
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<mods:titleInfo id="65D99CC6235FB5318DEF28A5EB45E4F7">
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<mods:title id="F29A5E1557F6C33A1B923DCC59C5BEF8">The beetle family Carabidae of Costa Rica: The genus Epikastea Liebke of the Plochonida Group, with new Neotropical species and notes on their way of life (Insecta: Coleoptera, Lebiini, Agrina)</mods:title>
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</mods:titleInfo>
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<mods:name id="BAE75351F123DF551B33E74B3AF70A35" type="personal">
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<mods:role id="80AEB659C8D3026A381A892FB2AF668C">
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<mods:roleTerm id="828D5D84C68075492B9E3754EDAE517C">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="8ADE18E18E89836F9965544229661DDA">Erwin, Terry L.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="F40A5D535BAC1891613E4CB8876780D2">text</mods:typeOfResource>
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<mods:titleInfo id="981EFC8710C7C8374CA194AD1DC19DE7">
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<mods:title id="2F952E50C33B94B45A64071AEF6506D6">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="8290F061BAA6BD4E775E366911D3671D">
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<mods:date id="47730A5673286F7804E421846376EB98">2004</mods:date>
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<mods:detail id="37A56C217EB9AAF865D88D0F36626458" type="volume">
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<mods:number id="1A4B482EECD25B635A7DF8F73A868619">790</mods:number>
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</mods:detail>
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<mods:extent id="228C8371AAFD4AEDC52C2BB61F5D0891" unit="page">
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<mods:start id="B3D78E9A8FCE1B8C66F1369DFC436B87">1</mods:start>
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<mods:end id="C1C53DB3B7055F8B3A5A3C9D97B99D2C">20</mods:end>
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<mods:classification id="4C9BBDE532C31F18D1FF335A973DF41F">journal article</mods:classification>
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<mods:identifier id="EE715B36DB1D9467879D4C7C7A96F013" type="DOI">10.5281/zenodo.158726</mods:identifier>
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<mods:identifier id="089362653277AC6E57B1B3940695FC0B" type="GBIF-Dataset">346a7f7d-c951-4756-acdb-09c0a30d52cf</mods:identifier>
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<mods:identifier id="E39D0D4BEA76E662FCFCD2FF6CC4FD93" type="ISSN">11755326</mods:identifier>
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<mods:identifier id="005A12619267806B3E6460A00E87D481" type="Zenodo-Dep">158726</mods:identifier>
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<mods:identifier id="EA8022435D727C416909096033930F03" type="ZooBank">E1878D62-01DE-4BB2-8960-23085056D27F</mods:identifier>
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</mods:mods>
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<treatment id="325787BEFF80FFF16C0FFEC7FB1986C1" ID-DOI="http://doi.org/10.5281/zenodo.6270972" ID-GBIF-Taxon="119609496" ID-Zenodo-Dep="6270972" LSID="urn:lsid:plazi:treatment:325787BEFF80FFF16C0FFEC7FB1986C1" httpUri="http://treatment.plazi.org/id/325787BEFF80FFF16C0FFEC7FB1986C1" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15">
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<subSubSection id="F2E46523FF80FFF06C0FFEC7FE75864E" pageId="14" pageNumber="15" type="nomenclature">
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<paragraph id="BA4136A8FF80FFF06C0FFEC7FCEF8622" blockId="14.[264,785,283,345]" box="[264,785,283,310]" pageId="14" pageNumber="15">
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<heading id="E10981C4FF80FFF06C0FFEC7FCEF8622" bold="true" box="[264,785,283,310]" fontSize="11" level="1" pageId="14" pageNumber="15" reason="1">
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<emphasis id="888AEABAFF80FFF06C0FFEC7FCEF8622" bold="true" box="[264,785,283,310]" pageId="14" pageNumber="15">
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<taxonomicName id="7DFE4D2BFF80FFF06C0FFEC7FD8B8622" ID-CoL="6FTCK" authority="Erwin" authorityName="Erwin" box="[264,629,283,310]" class="Insecta" family="Carabidae" genus="Epikastea" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="mancocapac" status="sp. nov.">
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<emphasis id="888AEABAFF80FFF06C0FFEC7FDE38621" bold="true" box="[264,541,284,310]" italics="true" pageId="14" pageNumber="15">Epikastea mancocapac</emphasis>
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Erwin
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</taxonomicName>
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,
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<taxonomicNameLabel id="93B957C1FF80FFF06F85FEC0FCEF8622" box="[642,785,283,309]" pageId="14" pageNumber="15" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="BA4136A8FF80FFF06C0FFEE4FE75864E" blockId="14.[264,785,283,345]" box="[264,395,319,345]" pageId="14" pageNumber="15">(Fig. 4, 13)</paragraph>
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</subSubSection>
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<subSubSection id="F2E46523FF80FFF06C0FFE52FC6E841C" pageId="14" pageNumber="15" type="materials_examined">
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<paragraph id="BA4136A8FF80FFF06C0FFE52FBCF86DC" blockId="14.[264,1323,393,1899]" pageId="14" pageNumber="15">
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<emphasis id="888AEABAFF80FFF06C0FFE52FE7F86B4" bold="true" box="[264,385,393,419]" pageId="14" pageNumber="15">
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<typeStatus id="6545880AFF80FFF06C0FFE52FE8286B4" box="[264,380,393,419]" pageId="14" pageNumber="15" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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Male,
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<collectingCountry id="C2E97638FF80FFF06CDEFE52FDD986B4" box="[473,551,393,419]" name="Peru" pageId="14" pageNumber="15">PERÚ</collectingCountry>
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, LORETO, Samiria River, Camp Manco Capac, 04º 43’ 0 S, 074º 18’ 0 W, June (T.L. Erwin, G.P. Servat, et al.)(
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<collectionCode id="DCEFAE6DFF80FFF06E19FE6AFC7C86DC" box="[798,898,433,459]" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="14" pageNumber="15">NMNH</collectionCode>
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: ADP093696).
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</paragraph>
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<paragraph id="BA4136A8FF80FFF06C3FFE02FC6E841C" blockId="14.[264,1323,393,1899]" pageId="14" pageNumber="15">
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<emphasis id="888AEABAFF80FFF06C3FFE02FD6686E4" bold="true" box="[312,664,473,499]" pageId="14" pageNumber="15">Derivation of specific name.</emphasis>
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The specific epithet,
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<emphasis id="888AEABAFF80FFF06E9CFE02FBC886E5" box="[923,1078,473,498]" italics="true" pageId="14" pageNumber="15">
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<taxonomicName id="7DFE4D2BFF80FFF06E9CFE02FBCC86E5" box="[923,1074,473,498]" class="Insecta" family="Carabidae" genus="Epikastea" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="mancocapac">mancocapac</taxonomicName>
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,
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</emphasis>
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is used as a noun in apposition based on the name of the founder of the Inca Empire, Manco Capac, who was the Sun's son, and the first Inca. He came from Lake Titicaca and had the mission of looking for a good place to found his empire that he did with the help of a silver stick. When this silver stick could penetrate the soil it indicated fertility and it did so at Cerro Huanacaure in Cusco,
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<collectingCountry id="C2E97638FF80FFF06CCBFD7AFDFB85AC" box="[460,517,673,699]" name="Peru" pageId="14" pageNumber="15">Perú</collectingCountry>
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. Although our camp in the lowlands had nothing to do with Cusco and the Sun’s son, the locals saw fit to name the area Manco Capac, so I use their custom and name this species after the name of the local camp.
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</paragraph>
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</subSubSection>
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<subSubSection id="F2E46523FF80FFF06C3FFCC2FC8A8424" box="[312,884,793,819]" pageId="14" pageNumber="15" type="description">
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<paragraph id="BA4136A8FF80FFF06C3FFCC2FC8A8424" blockId="14.[264,1323,393,1899]" box="[312,884,793,819]" pageId="14" pageNumber="15">
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<emphasis id="888AEABAFF80FFF06C3FFCC2FE008424" bold="true" box="[312,510,793,819]" pageId="14" pageNumber="15">Common name.</emphasis>
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Inca bigheaded carabid beetle.
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</paragraph>
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</subSubSection>
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<subSubSection id="F2E46523FF80FFF06C3FFC9AFC2A84BC" pageId="14" pageNumber="15" type="diagnosis">
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<paragraph id="BA4136A8FF80FFF06C3FFC9AFC2A84BC" blockId="14.[264,1323,393,1899]" pageId="14" pageNumber="15">
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<emphasis id="888AEABAFF80FFF06C3FFC9AFE46844C" bold="true" box="[312,440,833,859]" pageId="14" pageNumber="15">Diagnosis.</emphasis>
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With the attributes of the genus as described above and pronotum unicolorous, rufous; elytron without preapical spot. Head with course granulate isodiametric microscupture, surface dull. Size moderately large for genus.
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</paragraph>
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</subSubSection>
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<subSubSection id="F2E46523FF80FFF16C3FFC62FE5C8649" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15" type="description">
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<paragraph id="BA4136A8FF80FFF06C3FFC62FC23816C" blockId="14.[264,1323,393,1899]" pageId="14" pageNumber="15">
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<emphasis id="888AEABAFF80FFF06C3FFC62FE3184C4" bold="true" box="[312,463,953,979]" pageId="14" pageNumber="15">Description.</emphasis>
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(Fig. 4). Size medium: ABL =
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<quantity id="7D069B4DFF80FFF06E4BFC62FBF884C4" box="[844,1030,953,979]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" metricValueMax="10.2" metricValueMin="7.8" pageId="14" pageNumber="15" unit="mm" value="9.0" valueMax="10.2" valueMin="7.8">7.8 to 10.2 mm</quantity>
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, SBL =
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<quantity id="7D069B4DFF80FFF0696BFC62FAD884C4" box="[1132,1318,953,979]" metricMagnitude="-3" metricUnit="m" metricValue="7.475" metricValueMax="7.5" metricValueMin="7.45" pageId="14" pageNumber="15" unit="mm" value="7.475" valueMax="7.5" valueMin="7.45">7.45 to 7.5 mm</quantity>
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, TW = 3.0 to
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<quantity id="7D069B4DFF80FFF06CB3FC3AFDE884EC" box="[436,534,993,1019]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="14" pageNumber="15" unit="mm" value="3.2">3.2 mm</quantity>
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.
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<emphasis id="888AEABAFF80FFF06F20FC3AFD8B84ED" box="[551,629,993,1018]" italics="true" pageId="14" pageNumber="15">Color:</emphasis>
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Dorsum of head, prothorax and entire venter rufescent; mouthparts, antennomeres 1–4 and legs rufotestaceous; antennomeres testaceous. Metathorasic wings clear.
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<emphasis id="888AEABAFF80FFF06F19FBEAFD8A835D" box="[542,628,1073,1098]" italics="true" pageId="14" pageNumber="15">Luster:</emphasis>
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Semimatte on head; pronotum and elytron moderately shiny.
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<emphasis id="888AEABAFF80FFF06C53FB82FDEF8365" box="[340,529,1113,1138]" italics="true" pageId="14" pageNumber="15">Microsculpture:</emphasis>
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Isodiametric meshes throughout, these granular on head and pronotum; slightly transverse on elytra.
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<emphasis id="888AEABAFF80FFF06F9EFB5AFD1C838D" box="[665,738,1153,1178]" italics="true" pageId="14" pageNumber="15">Head:</emphasis>
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Subequal across eyes to that of pronotum; frontal furrows extended to level of anterior supraorbital seta, moderately impressed proximal to clypeus, margined laterally by well developed carina, surface near carina slightly rugose; eyes large, produced, gena absent; frons and occiput slightly convex; labrum moderately emarginate; labial palpomere 4 depressed, subequal in size to maxillary palpomere 4.
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<emphasis id="888AEABAFF80FFF069F1FAFAFEA18275" italics="true" pageId="14" pageNumber="15">Prothorax:</emphasis>
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Pronotum cordiform, moderately convex, depressed along midline, side margins broadly explanate, markedly sinuate anterior to slightly acute hind angle, base slightly rounded, not lobed posteriorly.
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<emphasis id="888AEABAFF80FFF06F7CFA42FCEC82A5" box="[635,786,1433,1458]" italics="true" pageId="14" pageNumber="15">Pterothorax:</emphasis>
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Normal for Agrina, fully winged.
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<emphasis id="888AEABAFF80FFF069ADFA42FB1582A5" box="[1194,1259,1433,1458]" italics="true" pageId="14" pageNumber="15">Legs:</emphasis>
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Normal for Agrina.
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<emphasis id="888AEABAFF80FFF06CDFFA1AFDB982CD" box="[472,583,1473,1498]" italics="true" pageId="14" pageNumber="15">Abdomen</emphasis>
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: Sterna normal for Agrina, glabrous, except normal paired ambulatory setae on sterna III–V; males with two and females with 3 pairs long setae on sternum VI.
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<emphasis id="888AEABAFF80FFF06CAFF9CAFD9B813D" box="[424,613,1553,1578]" italics="true" pageId="14" pageNumber="15">Male genitalia:</emphasis>
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Phallus (
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<figureCitation id="22C52A2DFF80FFF06FD8F9CAFCC5813C" box="[735,827,1553,1579]" captionStart="FIGURE 13" captionStartId="11.[264,368,995,1019]" captionTargetBox="[411,1169,289,978]" captionTargetId="figure@11.[409,1183,285,983]" captionTargetPageId="11" captionText="FIGURE 13. Phallus, dorsal, ventral, left lateral aspects of Epikastea mancocapac Erwin n. sp., specimen # ADP 0093696." httpUri="https://zenodo.org/record/158728/files/figure.png" pageId="14" pageNumber="15">Fig. 13</figureCitation>
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) with ostium of 1/4 its length, slightly catopic, apex slightly elongate, narrowly rounded; endophallus without sclerotized attributes. Parameres asymmetric, right very small, left larger.
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</paragraph>
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<paragraph id="BA4136A8FF80FFF06C3FF952FBFF81E4" blockId="14.[264,1323,393,1899]" pageId="14" pageNumber="15">
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<emphasis id="888AEABAFF80FFF06C3FF952FDD781B4" bold="true" box="[312,553,1673,1699]" pageId="14" pageNumber="15">Dispersal potential.</emphasis>
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These beetles have been fogged from the canopy and from a cluster of suspended dry leaves
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<quantity id="7D069B4DFF80FFF06F57F96AFD8B81DC" box="[592,629,1713,1739]" metricMagnitude="0" metricUnit="m" metricValue="4.0" pageId="14" pageNumber="15" unit="m" value="4.0">4m</quantity>
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off the ground in a tree and these beetles are fully winged, thus they are likely good dispersers, as are most arboreal beetles.
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</paragraph>
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<paragraph id="BA4136A8FF80FFF16C3FF8DAFE5C8649" blockId="14.[264,1323,393,1899]" lastBlockId="15.[264,1323,284,470]" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15">
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<emphasis id="888AEABAFF80FFF06C3FF8DAFE3B800C" bold="true" box="[312,453,1793,1819]" pageId="14" pageNumber="15">Way of life.</emphasis>
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See under genus. One adult of this species was fogged from the canopy of a of a
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<taxonomicName id="7DFE4D2BFF80FFF06C48F8F2FDE08055" box="[335,542,1833,1858]" class="Gastropoda" family="Cypraeidae" genus="Mauritia" kingdom="Animalia" order="Littorinimorpha" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="flexuosa">
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<emphasis id="888AEABAFF80FFF06C48F8F2FDE08055" box="[335,542,1833,1858]" italics="true" pageId="14" pageNumber="15">
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<collectingCountry id="C2E97638FF80FFF06C48F8F2FE498055" box="[335,439,1833,1858]" name="Mauritius" pageId="14" pageNumber="15">Mauritia</collectingCountry>
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flexuosa
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</emphasis>
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</taxonomicName>
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L. palm and another from a cluster of suspended dry leaves
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<quantity id="7D069B4DFF80FFF069DAF8F2FAFC8054" box="[1245,1282,1833,1859]" metricMagnitude="0" metricUnit="m" metricValue="4.0" pageId="14" pageNumber="15" unit="m" value="4.0">4m</quantity>
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off the ground in a broad leaf tree in
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<collectingCountry id="C2E97638FF80FFF06F8BF88AFD38807C" box="[652,710,1873,1899]" name="Peru" pageId="14" pageNumber="15">Perú</collectingCountry>
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. The known altitudinal range of this species is
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<quantity id="7D069B4DFF80FFF069EFF88AFAD5807C" box="[1256,1323,1873,1899]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="14" pageNumber="15" unit="m" value="100.0">100m</quantity>
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above sea level. Adults have been obtained in June; hence, there is activity by them in the rainy season.
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</paragraph>
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</subSubSection>
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<subSubSection id="F2E46523FF81FFF16C3FFEB0FB1986C1" pageId="15" pageNumber="16" type="materials_examined">
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<paragraph id="BA4136A8FF81FFF16C3FFEB0FB1986C1" blockId="15.[264,1323,284,470]" pageId="15" pageNumber="16">
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<emphasis id="888AEABAFF81FFF16C3FFEB0FD6F8692" bold="true" box="[312,657,363,389]" pageId="15" pageNumber="16">Other specimens examined.</emphasis>
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<typeStatus id="6545880AFF81FFF16F9FFEB7FCFC8691" box="[664,770,364,390]" pageId="15" pageNumber="16" type="paratype">Paratype</typeStatus>
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. Female,
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<collectingCountry id="C2E97638FF81FFF16E74FEB7FC3F8691" box="[883,961,364,390]" name="Peru" pageId="15" pageNumber="16">PERÚ</collectingCountry>
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, LORETO, Samiria River, Camp Manco Capac, 04º 43’ 0 S, 074º 18’ 0 W, June (T.L. Erwin, et al.)(
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<collectionCode id="DCEFAE6DFF81FFF16909FE4FFB8F86B9" box="[1038,1137,404,430]" pageId="15" pageNumber="16">MUSM</collectionCode>
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: ADP066767).
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<emphasis id="888AEABAFF81FFF16C3FFE60FD9286C2" bold="true" box="[312,620,443,469]" pageId="15" pageNumber="16">Geographic distribution.</emphasis>
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Known only from the Samiria River watershed,
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<collectingCountry id="C2E97638FF81FFF169AEFE67FB1D86C1" box="[1193,1251,444,470]" name="Peru" pageId="15" pageNumber="16">Perú</collectingCountry>
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.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |