treatments-xml/data/8A/41/78/8A4178BFD3C45D99B1E677B0BB43D178.xml

<document ID-CLB-Dataset="21464" ID-DOI="http://dx.doi.org/10.3897/zookeys.949.51269" ID-GBIF-Dataset="e4d2c425-4596-487c-8c4f-b09330b48203" ID-PMC="PMC7381721" ID-Pensoft-Pub="1313-2970-949-1" ID-Pensoft-UUID="303D64593B3B550DBB7CE01AA547409D" ID-PubMed="32765176" ID-ZooBank="93BFA448BB7343CDBC4EA86090DA63D5" ModsDocID="1313-2970-949-1" checkinTime="1594832489974" checkinUser="pensoft" docAuthor="Salata, Sebastian &amp; Fisher, Brian L." docDate="2020" docId="8A4178BFD3C45D99B1E677B0BB43D178" docLanguage="en" docName="ZooKeys 949: 1-185" docOrigin="ZooKeys 949" docSource="http://dx.doi.org/10.3897/zookeys.949.51269" docTitle="Pheidole hazo Salata &amp; Fisher 2020, sp. nov." docType="treatment" docUuid="A8CA690D-98D2-4DC9-8B8A-CB60BE22F0ED" docUuidSource="ZooBank" docVersion="6" id="303D64593B3B550DBB7CE01AA547409D" lastPageNumber="1" masterDocId="303D64593B3B550DBB7CE01AA547409D" masterDocTitle="Taxonomic revision of the Pheidole sikorae species group (Hymenoptera, Formicidae) from Madagascar" masterLastPageNumber="185" masterPageNumber="1" pageNumber="1" updateTime="1732651982216" updateUser="ExternalLinkService">
<mods:mods id="82A10AB69E987BD5B86D5A4D95400A6C" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="D17650CC37E94DECDB6674CCFF136D07">
<mods:title id="62C6A7700AB3C922FF72C4124E16408A">Taxonomic revision of the Pheidole sikorae species group (Hymenoptera, Formicidae) from Madagascar</mods:title>
</mods:titleInfo>
<mods:name id="9923C681FE74BA5CAF108F587861B371" type="personal">
<mods:role id="866907EDD457575E6C870273669D528D">
<mods:roleTerm id="9FABC24BBA8238BEF62EA5293F43F827">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D48D3B9ECD15235058710ED88E6DCCB8">Salata, Sebastian</mods:namePart>
<mods:affiliation id="0F01EEAB53788DB2E836860B37680FF1">Department of Entomology, California Academy of Sciences, San Francisco, CA 94118, USA</mods:affiliation>
<mods:nameIdentifier id="05A4F9970FB6E50B396FDBC0D8799B62" type="orcid">https://orcid.org/0000-0003-0811-2309</mods:nameIdentifier>
<mods:nameIdentifier id="9833F0C424EA323D06FF3A812C6E51F4" type="email">sdsalata@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="A2CEFD0FF119E706CA41D19100FAFA05" type="personal">
<mods:role id="520F20FBB5EE48D871D63CD08BEE51D9">
<mods:roleTerm id="8721BC5F1337256640864A1D44D188B4">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C859EA9B251E3B2A5E624548EB70A8D0">Fisher, Brian L.</mods:namePart>
<mods:affiliation id="900E23858F6806650F6943F10F56E8F5">Department of Entomology, California Academy of Sciences, San Francisco, CA 94118, USA</mods:affiliation>
<mods:nameIdentifier id="889A9B77DE53715ED45E02D53F5A4E7E" type="orcid">https://orcid.org/0000-0002-4653-3270</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="69D73B346CCD3CDCF3A2C66B03A282FB">text</mods:typeOfResource>
<mods:relatedItem id="B97AAC9E6F12D423092458D6E7D242C9" type="host">
<mods:titleInfo id="A517E55ED800A5F445D1AF8B2648544A">
<mods:title id="89BBCD058EA80A4EF80EFB32F8938991">ZooKeys</mods:title>
</mods:titleInfo>
<mods:part id="82631843D0E532FCD024E4942C89E8BB">
<mods:date id="83BC373BAF02A32114AC81B760B5B93F">2020</mods:date>
<mods:detail id="20F6410C43BE305EA45B1E6535F2DEE6" type="volume">
<mods:number id="12951B23AB349E3333CC4BBDD76D9D47">949</mods:number>
</mods:detail>
<mods:extent id="FD7FB6E1C26E7ABEAF16CA9F757598EC" unit="page">
<mods:start id="C5C924AD68B701FF0857CD03B2BCAD82">1</mods:start>
<mods:end id="EDF2678652EA7337057EBA898EF8FAAD">185</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="2088689880557216D1011CDF0D155B75">
<mods:url id="077C268007E8C3C22A4198BEAE4F5658">http://dx.doi.org/10.3897/zookeys.949.51269</mods:url>
</mods:location>
<mods:classification id="018A403343D2F5F713D0E0A7DEF67675">journal article</mods:classification>
<mods:identifier id="1B8447B858BC44BDC6DF5E989CB0E868" type="DOI">http://dx.doi.org/10.3897/zookeys.949.51269</mods:identifier>
<mods:identifier id="F074DD27B54261848474EA63F17627B6" type="Pensoft-Pub">1313-2970-949-1</mods:identifier>
<mods:identifier id="B8BF6EA36D699495964B426659A083FD" type="ZooBank">93BFA448BB7343CDBC4EA86090DA63D5</mods:identifier>
<mods:identifier id="DDE09E3F45171027726F24C1CEC64734" type="Pensoft-UUID">303D64593B3B550DBB7CE01AA547409D</mods:identifier>
</mods:mods>
<treatment id="8A4178BFD3C45D99B1E677B0BB43D178" ID-GBIF-Taxon="165090173" LSID="urn:lsid:zoobank.org:act:A8CA690D-98D2-4DC9-8B8A-CB60BE22F0ED" httpUri="http://treatment.plazi.org/id/8A4178BFD3C45D99B1E677B0BB43D178" lastPageNumber="1" pageId="0" pageNumber="1" scope_family="Formicidae" scope_order="Hymenoptera">
<subSubSection id="6EC8F2B5D89EDAF00EACA9C1DEEECE61" pageId="0" pageNumber="1" type="nomenclature">
<paragraph id="FCCC7084D4CC5EDE875B62A44361B419" pageId="0" pageNumber="1">
<taxonomicName id="E9A403075884232AA6A250F27EFFACDC" ID-CoL="CML8T" LSID="8A4178BF-D3C4-5D99-B1E6-77B0BB43D178" authority="Salata &amp; Fisher, 2020" authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole hazo" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="hazo" status="sp. nov.">Pheidole hazo</taxonomicName>
<taxonomicNameLabel id="27CC0188464C71E7EE15C5107664BFD2" pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
<figureCitation id="1183A3E04D4F08987D751160B1FA1F18" captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Pheidole hazo sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0427762) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0427790)." figureDoi="10.3897/zookeys.949.51269.figure33" httpUri="https://binary.pensoft.net/fig/432202" pageId="0" pageNumber="1">Figs 33A-F</figureCitation>
<figureCitation id="AE91A750C8ADED78A75073340D26F38A" captionStart="Figure 63" captionStartId="F63" captionText="Figure 63. Major worker, hypostomal teeth. Pheidole alina sp. nov. (A). Pheidole ambohimanga sp. nov. (B). Pheidole analavelona sp. nov. (C). Pheidole andohahela sp. nov. (D). Pheidole anomala sp. nov. (E). Pheidole anosyenne sp. nov. (F). Pheidole antranohofa sp. nov. (G). Pheidole beanka sp. nov. (H). Pheidole befotaka sp. nov. (I). Pheidole dasos sp. nov. (J). Pheidole flavominuta sp. nov. (K). Pheidole gracilis sp. nov. (L). Pheidole haboka sp. nov. (M). Pheidole havoana sp. nov. (N). Pheidole hazo sp. nov. (O). Pheidole itremo sp. nov. (P). Pheidole joffreville sp. nov. (Q). Pheidole kely sp. nov. (R). Pheidole lavasoa sp. nov. (S). Pheidole litigiosa Forel (T). Pheidole mahamavo sp. nov. (U). Pheidole mainty sp. nov. (V). Pheidole mamiratra sp. nov. (W). Pheidole manantenina sp. nov. (X)." figureDoi="10.3897/zookeys.949.51269.figure63" httpUri="https://binary.pensoft.net/fig/432232" pageId="0" pageNumber="1">, 63O</figureCitation>
<figureCitation id="92CC35DB220BCD586822B76BF4351EA3" captionStart="Figure 65" captionStartId="F65" captionText="Figure 65. Distribution. Pheidole alina sp. nov. (A). Pheidole ambohimanga sp. nov. (B). Pheidole analavelona sp. nov. (C). Pheidole andohahela sp. nov. (D). Pheidole anomala sp. nov. (E). Pheidole anosyenne sp. nov. (F). Pheidole antranohofa sp. nov. (G). Pheidole beanka sp. nov. (H). Pheidole befotaka sp. nov. (I). Pheidole dasos sp. nov. (J). Pheidole flavominuta sp. nov. (K). Pheidole gracilis sp. nov. (L). Pheidole haboka sp. nov. (M). Pheidole havoana sp. nov. (N). Pheidole hazo sp. nov. (O). Pheidole itremo sp. nov. (P). Pheidole joffreville sp. nov. (Q). Pheidole kely sp. nov. (R). Pheidole lavasoa sp. nov. (S). Pheidole litigiosa Forel (T)." figureDoi="10.3897/zookeys.949.51269.figure65" httpUri="https://binary.pensoft.net/fig/432234" pageId="0" pageNumber="1">, 65O</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="E4B5A77B55C61DD4A4B46FDAB1B73675" pageId="0" pageNumber="1" type="type material">
<paragraph id="A4B232AB9CCE644DD83E7EFB48FDFB58" pageId="0" pageNumber="1">Type material.</paragraph>
<paragraph id="ED6E4AE6B41441F49F627D4E8F1D3891" pageId="0" pageNumber="1">
<emphasis id="D75DDBEC90750059282AFAED2C5AD7DA" bold="true" italics="true" pageId="0" pageNumber="1">Holotype.</emphasis>
Madagascar. • 1 major worker; Antananarivo; 3 km 41°NE Andranomay, 11.5 km 147°SSE Anjozorobe; -18.47333, 47.96; alt. 1300 m; 5 Dec 2000; Fisher et al. leg.; montane rainforest, ex dead branch above ground; BLF02441; CASENT0427790, bottom specimen on the pin (CASC). 
<emphasis id="1B4FC761D28090BB539F2E59E97380FB" bold="true" italics="true" pageId="0" pageNumber="1">Paratypes.</emphasis>
• 5w., 8s.; same data as for holotype; CASENT0427762, CASENT00872251, CASENT0413711, CASENT0413712, CASENT0427765 (CASC, MHNG, PBZT).
</paragraph>
<caption id="B6E1B3D5708AC89F1630A3599E6FC6BC" doi="10.3897/zookeys.949.51269.figure33" httpUri="https://binary.pensoft.net/fig/432202" pageId="0" pageNumber="1" start="Figure 33" startId="F33">
<paragraph id="8D3C1F28DD1B976CC869DA179F507780" pageId="0" pageNumber="1">
<emphasis id="AAC7BDEB5AB28810F453007778599287" bold="true" pageId="0" pageNumber="1">Figure 33.</emphasis>
<taxonomicName id="7E01466CE7ACC3FCE59D0DEAFDF18AE9" authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole hazo" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="hazo">
<emphasis id="850763BE71B569268556FCF2702B8428" italics="true" pageId="0" pageNumber="1">Pheidole hazo</emphasis>
</taxonomicName>
sp. nov., full-face view (
<emphasis id="7C04CD421C281B8BA810B2F1AC83DD5B" bold="true" pageId="0" pageNumber="1">A</emphasis>
), profile (
<emphasis id="3B0337648F4C4A0C5F36E3D00D7C7751" bold="true" pageId="0" pageNumber="1">C</emphasis>
), and dorsal view (
<emphasis id="A0795D93CF04CC16BAC3B4688E6CEAE4" bold="true" pageId="0" pageNumber="1">E</emphasis>
) of paratype minor worker (CASENT0427762) and full-face view (
<emphasis id="2B54AA1037E93903FCBB055AFF076BD7" bold="true" pageId="0" pageNumber="1">B</emphasis>
), profile (
<emphasis id="047A23BFBF4956827D7F43A022339A1D" bold="true" pageId="0" pageNumber="1">D</emphasis>
), and dorsal view (
<emphasis id="534CEE77104B5D86F156D376AEF746A1" bold="true" pageId="0" pageNumber="1">F</emphasis>
) of holotype major worker (CASENT0427790).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="4639B138EB1240E84E175DDA63904B70" pageId="0" pageNumber="1" type="diagnosis">
<paragraph id="B01CE0CD896066ED3BDB0EB955799111" pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph id="B16866D4EC97821AB5D49F638D450A1C" pageId="0" pageNumber="1">
Moderately large species. 
<emphasis id="02657E21E11E98F442B03DE5298FC54C" bold="true" italics="true" pageId="0" pageNumber="1">Major workers.</emphasis>
Head in full-face view sub-oval, not widening posteriorly, with anterior and posterior sides relatively straight, in lateral view sub-oval; ventral and dorsal faces convex; sides of the head with dense, long, suberect to erect pilosity; medial part of frons with moderately dense, thin, longitudinal rugae, rugae in posteromedial portion directed slightly outward, interspaces shiny with sparse and distinct rugofoveolae; lateral sides with longitudinal, thicker, and relatively sparser rugae; interspaces shiny with relatively dense and distinct rugofoveolae; occipital lobes, and area posterolateral from eyes without smooth notches; scape, when laid back, exceeding the midlength of head by two-fifths of its length; inner hypostomal teeth distinct, large, closely spaced, triangular, with rounded apex directed slightly outward; outer hypostomal teeth lobe-like, wider than inner hypostomal teeth and approximately the same height, apex directed outward; inner and outer hypostomal teeth closely spaced and connected by concavity; mesosoma with thick and sparse foveolae; pronotum with additional transverse to irregular, thin rugae; katepisternum with smooth notch; gaster smooth; body orange. 
<emphasis id="9AE7C6900C2779D63F8E905432B4776C" bold="true" italics="true" pageId="0" pageNumber="1">Minor workers.</emphasis>
Head foveolate with additional longitudinal to irregular, interrupted, and moderately thick rugae; area posterolateral from eyes with weaker foveolae and smooth notches; scape, when laid back, surpassing the posterior head margin by one-fifth of its length; promesonotum moderately low and moderately long, arched; promesonotal groove absent; propodeal spines minute and triangular; mesosoma with thick foveolae; lateral sides of propodeum, katepisternum, and anepisternum with smooth notches; body yellow.
</paragraph>
</subSubSection>
<subSubSection id="F7D8CD4B9AB7946D622C777C06045DD7" pageId="0" pageNumber="1" type="description">
<paragraph id="5EFCE535DD05D38DF0E17CAD3BCBED53" pageId="0" pageNumber="1">Description.</paragraph>
<paragraph id="A3DF136CF1D833402AC3E48157E16DDA" pageId="0" pageNumber="1">
<emphasis id="855567C283CD9F8F889B0E4E7F1162A9" bold="true" pageId="0" pageNumber="1">Major workers.</emphasis>
Measurements (
<emphasis id="07B28AA971D3397FBD8656E68B218165" italics="true" pageId="0" pageNumber="1">N</emphasis>
= 9): HL: 1.06-1.14 (1.09); HW: 1.02-1.13 (1.08); SL: 0.63-0.66 (0.64); EL: 0.14-0.17 (0.15); WL: 0.98-1.08 (1.03); PSL: 0.16-0.18 (0.17); MTL: 0.63-0.65 (0.64); PNW: 0.41-0.52 (0.47); PTW: 0.13-0.15 (0.14); PPW: 0.25-0.3 (0.27); CI: 97.6-104.3 (101.0); SI: 56.4-63.3 (59.7); PSLI: 15.1-17.1 (16.0); PPI: 47.3-55.8 (51.5); PNI: 39.2-47.0 (43.3); MTI: 57.1-63.9 (59.9).
</paragraph>
<paragraph id="1C74EA4B7BC716A21AA51F73122C500D" pageId="0" pageNumber="1">
<emphasis id="1F56DCCE62E87EFC2FEE2942A6C63B10" bold="true" italics="true" pageId="0" pageNumber="1">Head.</emphasis>
In full-face view sub-oval, not widening posteriorly, with anterior and posterior sides relatively straight (Fig. 
<figureCitation id="4407D514C14C541510F4C9195B2A1D3F" captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Pheidole hazo sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0427762) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0427790)." figureDoi="10.3897/zookeys.949.51269.figure33" httpUri="https://binary.pensoft.net/fig/432202" pageId="0" pageNumber="1">33B</figureCitation>
). In lateral view sub-oval; ventral and dorsal faces convex; inner hypostomal teeth visible. Sides of the head with dense, long, suberect to erect pilosity; whole head with dense, long, decumbent to erect pilosity. Medial part of frons with moderately dense, thin, longitudinal rugae, rugae in posteromedial directed slightly outward, interspaces shiny with sparse and distinct rugofoveolae; lateral sides with longitudinal, thicker, and relatively sparser rugae; interspaces shiny with relatively dense and distinct rugofoveolae. Occipital lobes with sparse, thick, and longitudinal to irregular rugae; interspaces rugofoveolate. Gena with relatively dense and thick, longitudinal rugae and rugofoveolate interspaces. Area posterolateral from eyes shiny, with dense, longitudinal rugofoveolae. Centre of clypeus smooth and shiny, lateral sides with indistinct rugulae; median notch present, moderately wide, and shallow; median longitudinal carina absent; lateral longitudinal carinae absent. Scape, when laid back, exceeding the midlength of head by two-fifths of its length; pilosity subdecumbent to erect (Fig. 
<figureCitation id="2574260BC681262C7013F058EEC9B1FB" captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Pheidole hazo sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0427762) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0427790)." figureDoi="10.3897/zookeys.949.51269.figure33" httpUri="https://binary.pensoft.net/fig/432202" pageId="0" pageNumber="1">33B, D</figureCitation>
). Inner hypostomal teeth distinct, large, closely spaced, triangular, with rounded apex directed slightly outward; outer hypostomal teeth lobe-like, wider than inner hypostomal teeth and approximately the same height, apex directed outward; inner and outer hypostomal teeth closely spaced and connected by concavity (Fig. 
<figureCitation id="7BC1084F416D128EAD2CC6BDFFC1098E" captionStart="Figure 63" captionStartId="F63" captionText="Figure 63. Major worker, hypostomal teeth. Pheidole alina sp. nov. (A). Pheidole ambohimanga sp. nov. (B). Pheidole analavelona sp. nov. (C). Pheidole andohahela sp. nov. (D). Pheidole anomala sp. nov. (E). Pheidole anosyenne sp. nov. (F). Pheidole antranohofa sp. nov. (G). Pheidole beanka sp. nov. (H). Pheidole befotaka sp. nov. (I). Pheidole dasos sp. nov. (J). Pheidole flavominuta sp. nov. (K). Pheidole gracilis sp. nov. (L). Pheidole haboka sp. nov. (M). Pheidole havoana sp. nov. (N). Pheidole hazo sp. nov. (O). Pheidole itremo sp. nov. (P). Pheidole joffreville sp. nov. (Q). Pheidole kely sp. nov. (R). Pheidole lavasoa sp. nov. (S). Pheidole litigiosa Forel (T). Pheidole mahamavo sp. nov. (U). Pheidole mainty sp. nov. (V). Pheidole mamiratra sp. nov. (W). Pheidole manantenina sp. nov. (X)." figureDoi="10.3897/zookeys.949.51269.figure63" httpUri="https://binary.pensoft.net/fig/432232" pageId="0" pageNumber="1">63O</figureCitation>
). 
<emphasis id="E11747712417094CA0DA9048C95C620A" bold="true" italics="true" pageId="0" pageNumber="1">Mesosoma.</emphasis>
In lateral view, promesonotum short, angular, and moderately low, posterior mesonotum moderately steep, mesonotal process indistinct, tubercle-like; promesonotal groove absent; metanotal groove absent; propodeal spines moderately long, moderately narrow, with acute apex; humeral area laterally weakly produced (Fig. 
<figureCitation id="299C286709CF8BB528F53552EE9DE79F" captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Pheidole hazo sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0427762) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0427790)." figureDoi="10.3897/zookeys.949.51269.figure33" httpUri="https://binary.pensoft.net/fig/432202" pageId="0" pageNumber="1">33D</figureCitation>
). Surface shiny with thick and sparse foveolae; pronotum with additional transverse to irregular, thin rugae; katepisternum with smooth notch. Pilosity sparse, long, and erect (Fig. 
<figureCitation id="E8C03396BCFA09970165BABB00B320AC" captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Pheidole hazo sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0427762) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0427790)." figureDoi="10.3897/zookeys.949.51269.figure33" httpUri="https://binary.pensoft.net/fig/432202" pageId="0" pageNumber="1">33D, F</figureCitation>
). 
<emphasis id="42477527B64B21950EC1C2AE0DDD5376" bold="true" italics="true" pageId="0" pageNumber="1">Petiole.</emphasis>
Shiny with fine and sparse foveolae; node smooth to finely foveolate, low, triangular, with rounded and thin apex, in rear view node dorsoventrally straight to slightly convex; pilosity moderately sparse and erect (Fig. 
<figureCitation id="4EDCA506CCC8133EEC2694DAB407FCA0" captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Pheidole hazo sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0427762) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0427790)." figureDoi="10.3897/zookeys.949.51269.figure33" httpUri="https://binary.pensoft.net/fig/432202" pageId="0" pageNumber="1">33D, F</figureCitation>
). 
<emphasis id="A71C912E376BFF439A6D291D24F4EA9F" bold="true" italics="true" pageId="0" pageNumber="1">Postpetiole.</emphasis>
Shiny and smooth; in dorsal view oval, lateral margins medially with two dentate projections; pilosity long, moderately sparse, and erect (Fig. 
<figureCitation id="EE9D64887EF42EF7B30BCFC3C5FC5E0E" captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Pheidole hazo sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0427762) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0427790)." figureDoi="10.3897/zookeys.949.51269.figure33" httpUri="https://binary.pensoft.net/fig/432202" pageId="0" pageNumber="1">33D, F</figureCitation>
). 
<emphasis id="D4380BDD269566699C6A5E2643791795" bold="true" italics="true" pageId="0" pageNumber="1">Gaster.</emphasis>
Shiny and smooth; pilosity moderately dense, long, and erect (Fig. 
<figureCitation id="4D627185DAB91DB2D6B37D8AAC94A83D" captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Pheidole hazo sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0427762) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0427790)." figureDoi="10.3897/zookeys.949.51269.figure33" httpUri="https://binary.pensoft.net/fig/432202" pageId="0" pageNumber="1">33D, F</figureCitation>
). 
<emphasis id="DECC76AAD800EE509D53DF8EFC82BCDF" bold="true" italics="true" pageId="0" pageNumber="1">Colour.</emphasis>
Dark orange; mandibles slightly darker; legs yellow (Fig. 
<figureCitation id="EFEE760C3FD1D45980A708EC7EC92D7B" captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Pheidole hazo sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0427762) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0427790)." figureDoi="10.3897/zookeys.949.51269.figure33" httpUri="https://binary.pensoft.net/fig/432202" pageId="0" pageNumber="1">33D, F</figureCitation>
).
</paragraph>
<paragraph id="EA322A95246E6BA7D7A3BAEAF4E975AC" pageId="0" pageNumber="1">
<emphasis id="2AEBC4743AA9E0639EDD0A5F92091C91" bold="true" pageId="0" pageNumber="1">Minor workers.</emphasis>
Measurements (
<emphasis id="88446C42A40C984777386AF62FD41221" italics="true" pageId="0" pageNumber="1">N</emphasis>
= 5): HL: 0.6-0.63 (0.61); HW: 0.5-0.53 (0.52); SL: 0.61-0.64 (0.62); EL: 0.11-0.12 (0.12); WL: 0.76-0.82 (0.79); PSL: 0.08-0.09 (0.09); MTL: 0.52-0.55 (0.54); PNW: 0.33-0.35 (0.34); PTW: 0.09-0.1 (0.09); PPW: 0.14-0.15 (0.15); CI: 114.8-121.9 (118.8); SI: 115.4-123.9 (120.9); PSLI: 12.9-14.6 (13.9); PPI: 57.3-70.4 (62.7); PNI: 63.4-67.4 (66.0); MTI: 101.5-106.2 (103.8).
</paragraph>
<paragraph id="5FDD74890659227E91545AE86BE3958B" pageId="0" pageNumber="1">
<emphasis id="912F5297AF5A33E08B6839AE74D815CB" bold="true" italics="true" pageId="0" pageNumber="1">Head.</emphasis>
Cephalic margin indistinctly convex or straight (Fig. 
<figureCitation id="E933F6B7767DA8D824E828A4D04D3DAE" captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Pheidole hazo sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0427762) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0427790)." figureDoi="10.3897/zookeys.949.51269.figure33" httpUri="https://binary.pensoft.net/fig/432202" pageId="0" pageNumber="1">33A</figureCitation>
). Pilosity relatively sparse, long, decumbent to suberect. Sculpture foveolate with additional longitudinal to irregular, interrupted, and moderately thick rugae; area posterolateral from eyes with weaker foveolae and smooth notches. Clypeus with median longitudinal carina absent; two lateral longitudinal carinae absent. Scape, when laid back, surpassing the posterior head margin by one-fifth of its length; pilosity dense, subdecumbent to erect (Fig. 
<figureCitation id="2B1E00753D8EEA931908EC56D240B5D3" captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Pheidole hazo sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0427762) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0427790)." figureDoi="10.3897/zookeys.949.51269.figure33" httpUri="https://binary.pensoft.net/fig/432202" pageId="0" pageNumber="1">33A, C</figureCitation>
). 
<emphasis id="67CB60889B2D7349DFEFA59C77848000" bold="true" italics="true" pageId="0" pageNumber="1">Mesosoma.</emphasis>
In lateral view, promesonotum moderately low and moderately long, arched; promesonotal groove absent; metanotal groove distinct; propodeal spines minute and triangular (Fig. 
<figureCitation id="933E916BBF72AA198EBAD7DEC25209AE" captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Pheidole hazo sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0427762) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0427790)." figureDoi="10.3897/zookeys.949.51269.figure33" httpUri="https://binary.pensoft.net/fig/432202" pageId="0" pageNumber="1">33C</figureCitation>
). Sculpture shiny and with thick foveolae; lateral sides of propodeum, katepisternum, and anepisternum with smooth notches. Pilosity very sparse, moderately long, and erect (Fig. 
<figureCitation id="B36BA49ABE12D9A36F3585BFA2BA887D" captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Pheidole hazo sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0427762) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0427790)." figureDoi="10.3897/zookeys.949.51269.figure33" httpUri="https://binary.pensoft.net/fig/432202" pageId="0" pageNumber="1">33C, E</figureCitation>
). 
<emphasis id="A054EA476F19458CC66B2A1DE7A8DC8A" bold="true" italics="true" pageId="0" pageNumber="1">Postpetiole.</emphasis>
Short, low, and relatively flat; with few short, erect setae (Fig. 
<figureCitation id="072C69EA9CF56844AF585DF4462BA33A" captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Pheidole hazo sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0427762) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0427790)." figureDoi="10.3897/zookeys.949.51269.figure33" httpUri="https://binary.pensoft.net/fig/432202" pageId="0" pageNumber="1">33C, E</figureCitation>
). 
<emphasis id="28B7C6BF94BEF5C0FC0521A160519FD4" bold="true" italics="true" pageId="0" pageNumber="1">Gaster.</emphasis>
With sparse, erect pilosity (Fig. 
<figureCitation id="36C50772456EA946BCDA80017174A4AB" captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Pheidole hazo sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0427762) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0427790)." figureDoi="10.3897/zookeys.949.51269.figure33" httpUri="https://binary.pensoft.net/fig/432202" pageId="0" pageNumber="1">33C, E</figureCitation>
). 
<emphasis id="6C6176AF438BD12C17A9034EFF615A1A" bold="true" italics="true" pageId="0" pageNumber="1">Colour.</emphasis>
Yellow, vertex slightly darker (Fig. 
<figureCitation id="2345A0CE54F407F71DB8A3FF8761B2A9" captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Pheidole hazo sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0427762) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0427790)." figureDoi="10.3897/zookeys.949.51269.figure33" httpUri="https://binary.pensoft.net/fig/432202" pageId="0" pageNumber="1">33C, E</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="3EB000AC546BFC91AA9BA299F7D41228" pageId="0" pageNumber="1" type="etymology">
<paragraph id="72A5E2B82AC91A6EEAACBEA81871BF88" pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph id="E42CB74774F221FB959F3E739426F16C" pageId="0" pageNumber="1">Malagasy for tree in reference to arboreal nesting preferences of the species.</paragraph>
</subSubSection>
<subSubSection id="E5ED39BB5B71FA1FF7C648E7B31A77A8" pageId="0" pageNumber="1" type="biology_ecology">
<paragraph id="C78614F21788CFA0B985A396C5F1D150" pageId="0" pageNumber="1">Biology.</paragraph>
<paragraph id="3CE44E26EAE444927262A54C277CF1FF" pageId="0" pageNumber="1">The species was collected at 1300 m in elevation, in montane rainforest. Nest was located in dead branch above ground.</paragraph>
</subSubSection>
<subSubSection id="C55AEFE7ACA0940CE98B96754E947324" pageId="0" pageNumber="1" type="comments">
<paragraph id="F08547F60E4629758001E5E22DFAFD61" pageId="0" pageNumber="1">Comments.</paragraph>
<paragraph id="5D38CC318402EE26E2C62D862C846D4B" pageId="0" pageNumber="1">
<taxonomicName id="F49563B5636870E859A1DF0CF15F8B88" authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole hazo" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="hazo">
<emphasis id="D4F063D24B447B40CDAA004A53F99603" italics="true" pageId="0" pageNumber="1">Pheidole hazo</emphasis>
</taxonomicName>
sp. nov., described from the vicinity of Andranoma in Antananarivo, is most similar to 
<taxonomicName id="5DFBD141208381EDCAA6B4A896643902" lsidName="P. dasos" pageId="0" pageNumber="1" rank="species" species="dasos">
<emphasis id="A84143EB0214D25F372686B33D6AFBEC" italics="true" pageId="0" pageNumber="1">P. dasos</emphasis>
</taxonomicName>
sp. nov., known from Makirovana forest in Antsiranana, and 
<taxonomicName id="3CF08C39054F56FA37E1A58C424F5B54" lsidName="P. sparsa" pageId="0" pageNumber="1" rank="species" species="sparsa">
<emphasis id="8DBA438BA4D7E048651B3CD62309F2AE" italics="true" pageId="0" pageNumber="1">P. sparsa</emphasis>
</taxonomicName>
sp. nov., recorded so far only from Bemanevika in Mahajanga. Majors of 
<taxonomicName id="81194B424F1D38DA2641C348718C22A2" lsidName="P. hazo" pageId="0" pageNumber="1" rank="species" species="hazo">
<emphasis id="D46B815749F7B4EA82557875CC7FC7A6" italics="true" pageId="0" pageNumber="1">P. hazo</emphasis>
</taxonomicName>
sp. nov. can be distinguished from both taxa by medial part of frons with moderately dense rugae with interspaces with distinct rugofoveolae, and pronotum lacking smooth notches; minors of 
<taxonomicName id="DC63EACF2A0698AB507E7930063DCB7B" lsidName="P. hazo" pageId="0" pageNumber="1" rank="species" species="hazo">
<emphasis id="DB51CF155B5BADF840088A01B392F362" italics="true" pageId="0" pageNumber="1">P. hazo</emphasis>
</taxonomicName>
sp. nov. can be distinguished from 
<taxonomicName id="FA04A50054BD7C4D0C3827D9172627E6" lsidName="P. sparsa" pageId="0" pageNumber="1" rank="species" species="sparsa">
<emphasis id="B218704819F5A80F80B38AD6EEDAF36B" italics="true" pageId="0" pageNumber="1">P. sparsa</emphasis>
</taxonomicName>
sp. nov. by yellow body colouration, and mesosoma predominantly with dense foveolae, and from 
<taxonomicName id="5E793E79155E67C4E82005901FA57569" lsidName="P. dasos" pageId="0" pageNumber="1" rank="species" species="dasos">
<emphasis id="42DE2DC652540CE31C00D0725D71D111" italics="true" pageId="0" pageNumber="1">P. dasos</emphasis>
</taxonomicName>
sp. nov. by yellow body colouration, shorter scape surpassing the posterior head margin by one-fifth of its length, and pronotum with distinct and dense foveolae.
</paragraph>
</subSubSection>
</treatment>
</document>