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143 lines
16 KiB
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<document ID-CLB-Dataset="275944" ID-DOI="http://dx.doi.org/10.3897/contrib.entomol.73.e110258" ID-GBIF-Dataset="895a3095-4fbb-4161-9ed8-1484d8354f7d" ID-Pensoft-Pub="2511-6428-2-167" ID-Pensoft-UUID="10AD65E0AA955F9284FBFA1EAF1286C6" ID-ZooBank="EC125FAC97844575A6912FA6AEE0AA3D" ModsDocID="2511-6428-73-2-167" checkinTime="1698951195197" checkinUser="pensoft" docAuthor="Wichard, Wilfried" docDate="2023" docId="720B8A80172E507E96EBA3C0A85CF063" docLanguage="en" docName="ContribEntomol 73(2): 167-179" docOrigin="Contributions to Entomology 73 (2)" docPubDate="2023-11-02" docSource="http://dx.doi.org/10.3897/contrib.entomol.73.e110258" docTitle="Paduniella cretacea Wichard 2023, sp. nov." docType="treatment" docUuid="8CE659BD-2BCD-4080-ABDC-41B6DF7DCB17" docUuidSource="ZooBank" docVersion="2" id="10AD65E0AA955F9284FBFA1EAF1286C6" lastPageNumber="167" masterDocId="10AD65E0AA955F9284FBFA1EAF1286C6" masterDocTitle="Fossil Trichoptera embedded in mid-Cretaceous Burmese amber" masterLastPageNumber="179" masterPageNumber="167" pageNumber="167" updateTime="1698998955493" updateUser="ExternalLinkService">
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<mods:mods id="1DB95D0CE9D076CF5EA0A540E64A5880" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="0FA94C681EC96848DAE9D7D7F8A25937">
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<mods:title id="D0616C0B6CFD9754F58190FC5602F982">Fossil Trichoptera embedded in mid-Cretaceous Burmese amber</mods:title>
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<mods:name id="1BAAF0634E63DD5A798C78217D185325" type="personal">
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<mods:role id="FE0E61DA7811B68372D5906BCF7D09BF">
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<mods:roleTerm id="5FF0E262D91016B984BA102F54DCED45">Author</mods:roleTerm>
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<mods:namePart id="63335BC3751A0B6C3DA0500174D3CD30">Wichard, Wilfried</mods:namePart>
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<mods:nameIdentifier id="6A3986D11808AF01A05B7F7757E13C49" type="ORCID">https://orcid.org/0000-0002-5933-6292</mods:nameIdentifier>
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<mods:affiliation id="EE3FF9AFE01CFAF12034AFF4753A514A">University of Koeln, Institute of Biology, Koeln, Germany</mods:affiliation>
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<mods:nameIdentifier id="426731B3867AD5E5777BB02F106E04CB" type="email">wichard@uni-koeln.de</mods:nameIdentifier>
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<mods:typeOfResource id="FFF6DBA31E67537913C2254B0F19A91F">text</mods:typeOfResource>
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<mods:title id="C6412808DC83EC9B1C1DAE100C831926">Contributions to Entomology</mods:title>
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</mods:titleInfo>
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<mods:part id="3C17D08E5883A4D3D8D281D665ED5D11">
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<mods:date id="45025C3EA1A7DBD333E579975429C912">2023</mods:date>
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<mods:number id="03F8557AA44A0BA7BDD1C0F50D765451">2023-11-02</mods:number>
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<mods:detail id="34B5442E29598A17BA9016E58CAB6CFA" type="volume">
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<mods:number id="5CD151A0798717768952D35CA7ECF95C">73</mods:number>
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<mods:detail id="FD846E381E0782CC4FAC4860DFB8392E" type="issue">
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<mods:number id="2FD6CD53B92B115EFB394BC7C90CBB9A">2</mods:number>
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<mods:start id="2B3562732253EF091D5599ADEDCC8844">167</mods:start>
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<mods:url id="BE18F920DA9E8D2612BB08DC1C99F034">http://dx.doi.org/10.3897/contrib.entomol.73.e110258</mods:url>
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<mods:classification id="CC64FAE41110DA18824D533CA0472D50">journal article</mods:classification>
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<mods:identifier id="2057BA8718201159B08DB601875EC0B4" type="DOI">http://dx.doi.org/10.3897/contrib.entomol.73.e110258</mods:identifier>
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<treatment id="720B8A80172E507E96EBA3C0A85CF063" ID-GBIF-Taxon="216248513" LSID="urn:lsid:zoobank.org:act:8CE659BD-2BCD-4080-ABDC-41B6DF7DCB17" httpUri="http://treatment.plazi.org/id/720B8A80172E507E96EBA3C0A85CF063" lastPageNumber="167" pageId="0" pageNumber="167">
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<subSubSection id="B97861E1FD13F9305379D4E6185CD432" pageId="0" pageNumber="167" type="nomenclature">
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<paragraph id="CB5A78E7E91B605A4B1409E29746DF97" pageId="0" pageNumber="167">
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<taxonomicName id="0891F54A080482B76E865E3C7E8D07D8" LSID="https://zoobank.org/8CE659BD-2BCD-4080-ABDC-41B6DF7DCB17" authority="Wichard, 2023" authorityName="Wichard" authorityYear="2023" class="Insecta" family="Psychomyiidae" genus="Paduniella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paduniella cretacea" order="Trichoptera" pageId="0" pageNumber="167" phylum="Arthropoda" rank="species" species="cretacea" status="sp. nov.">Paduniella cretacea</taxonomicName>
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<taxonomicNameLabel id="722E8514A4ABDA476175B7F092F8C90A" pageId="0" pageNumber="167">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="5F2C5532153D17F773679F07A2B322AF" pageId="0" pageNumber="167" type="description">
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<paragraph id="9CD6B8DD5C46A665A4FBDD4119957734" pageId="0" pageNumber="167">
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<figureCitation id="B0DDD48AD569EB89B77F12BB8D03198D" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Paduniella cretacea sp. nov. in mid-Cretaceous Burmese amber, male holotype (Inventory no.: ZFMK-TRI 0008359. A. Male in ventral view; B. Paired bifurcated inferior appendages in ventral view; C. Paired 6 - segmented maxillary palps; D. Drawing of bifurcated inferior appendages (inf app) and median process (med pro) in lateral view." figureDoi="10.3897/contrib.entomol.73.e110258.figure2" httpUri="https://binary.pensoft.net/fig/928760" pageId="0" pageNumber="167">Fig. 2</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="AFE3514E59BCB613F1C81196C49774B3" pageId="0" pageNumber="167" type="holotype">
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<paragraph id="7B396D72C5FDCAD840251185904A686B" pageId="0" pageNumber="167">Holotype.</paragraph>
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<paragraph id="3EA295ECBC4FA86B1FCCF4EC62F4697D" pageId="0" pageNumber="167">
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Male-specimen deposited Zoological Research Museum Alexander Koenig, Bonn, Germany, Inventory no.: ZFMK-TRI000835 (ex coll. Patrick
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<normalizedToken id="97F4FA7612B3AA30992E905796308197" originalValue="Müller">Mueller</normalizedToken>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="1BA83C8704D8FB9C54BF47F397D0B36F" pageId="0" pageNumber="167" type="preservation">
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<paragraph id="118A65DF3618756D23BA795ECCC795B3" pageId="0" pageNumber="167">Preservation.</paragraph>
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<paragraph id="9ABDDA9F9A6D9C951D6213932C2F6EDB" pageId="0" pageNumber="167">The fossil male is well preserved in amber. Antennae present, as well as the six-segmented maxillary palps. The labial palps are not recognizable. The forewings show the venation well, the hind wings are unfortunately hidden. The inferior appendages of the male genital are visible from ventral.</paragraph>
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</subSubSection>
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<subSubSection id="28E77024EB648A6F0FB133A3D7D73856" pageId="0" pageNumber="167" type="etymology">
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<paragraph id="226E6A0C1FE0F24390DF9D6C4F6AA848" pageId="0" pageNumber="167">Etymology.</paragraph>
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<paragraph id="8700EB687636E951FD58D5351C863A7A" pageId="0" pageNumber="167">
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The extinct
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<taxonomicName id="09CCE7A82F5CCC5BB5B3E7F1146ED475" class="Insecta" family="Psychomyiidae" genus="Paduniella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paduniella" order="Trichoptera" pageId="0" pageNumber="167" phylum="Arthropoda" rank="genus">
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<emphasis id="EABBCB8A17B7A50AB066985C4706E47C" italics="true" pageId="0" pageNumber="167">Paduniella</emphasis>
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</taxonomicName>
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species is named after its geological age of the Cretaceous period (latin: Cretaceum).
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</paragraph>
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</subSubSection>
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<subSubSection id="578382E61F003549361C0A2054C01D85" pageId="0" pageNumber="167" type="description">
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<paragraph id="F8D58AA661B2FD8AFB69264AF36E83DF" pageId="0" pageNumber="167">Description.</paragraph>
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<paragraph id="5F23EDE77DC59C8CA35B67C25D660AE4" pageId="0" pageNumber="167">
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Male, forewings ca. 2.8 mm long, antennae about half as long as forewing. Maxillary palps each six-segmented (Fig.
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<figureCitation id="1DEACC93761E4E724CC4D93F7B63EE52" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Paduniella cretacea sp. nov. in mid-Cretaceous Burmese amber, male holotype (Inventory no.: ZFMK-TRI 0008359. A. Male in ventral view; B. Paired bifurcated inferior appendages in ventral view; C. Paired 6 - segmented maxillary palps; D. Drawing of bifurcated inferior appendages (inf app) and median process (med pro) in lateral view." figureDoi="10.3897/contrib.entomol.73.e110258.figure2" httpUri="https://binary.pensoft.net/fig/928760" pageId="0" pageNumber="167">2C</figureCitation>
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); labial palps not visible. Each forewing has forks II, III, IV, and V; hindwings not visible. In ventral view (Fig.
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<figureCitation id="2DC36E14EF6335C47BEA6E39B5E8A355" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Paduniella cretacea sp. nov. in mid-Cretaceous Burmese amber, male holotype (Inventory no.: ZFMK-TRI 0008359. A. Male in ventral view; B. Paired bifurcated inferior appendages in ventral view; C. Paired 6 - segmented maxillary palps; D. Drawing of bifurcated inferior appendages (inf app) and median process (med pro) in lateral view." figureDoi="10.3897/contrib.entomol.73.e110258.figure2" httpUri="https://binary.pensoft.net/fig/928760" pageId="0" pageNumber="167">2B</figureCitation>
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), male genitalia have deep, bifurcated inferior processes each with an elongate, overhanging process that is tapered and slightly curved apically. In lateral view (Fig.
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<figureCitation id="6D2CC82810FB88FC3EA533C83616126A" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Paduniella cretacea sp. nov. in mid-Cretaceous Burmese amber, male holotype (Inventory no.: ZFMK-TRI 0008359. A. Male in ventral view; B. Paired bifurcated inferior appendages in ventral view; C. Paired 6 - segmented maxillary palps; D. Drawing of bifurcated inferior appendages (inf app) and median process (med pro) in lateral view." figureDoi="10.3897/contrib.entomol.73.e110258.figure2" httpUri="https://binary.pensoft.net/fig/928760" pageId="0" pageNumber="167">2D</figureCitation>
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), a slender median process is visible.
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</paragraph>
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<caption id="10E0B7D0F27CBC7A11FC00A8CB5475A9" doi="10.3897/contrib.entomol.73.e110258.figure2" httpUri="https://binary.pensoft.net/fig/928760" pageId="0" pageNumber="167" start="Figure 2" startId="F2">
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<paragraph id="19E1C964DFC87E6A1A58F742511A6C25" pageId="0" pageNumber="167">
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<emphasis id="8582C960ABC10BA9D8CE68477086549A" bold="true" pageId="0" pageNumber="167">Figure 2.</emphasis>
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<taxonomicName id="B841C2367E1FCC7666C382FB80A3A4DE" authorityName="Wichard" authorityYear="2023" class="Insecta" family="Psychomyiidae" genus="Paduniella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paduniella cretacea" order="Trichoptera" pageId="0" pageNumber="167" phylum="Arthropoda" rank="species" species="cretacea">
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<emphasis id="CA4A5941BAABA6A594556021169035C3" italics="true" pageId="0" pageNumber="167">Paduniella cretacea</emphasis>
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</taxonomicName>
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sp. nov. in mid-Cretaceous Burmese amber, male holotype (Inventory no.: ZFMK-TRI0008359.
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<emphasis id="C5FB1C57390811F0B5B921538125463D" bold="true" pageId="0" pageNumber="167">A.</emphasis>
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Male in ventral view;
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<emphasis id="18E193808E05075582646EE9F9ED7382" bold="true" pageId="0" pageNumber="167">B.</emphasis>
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Paired bifurcated inferior appendages in ventral view;
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<emphasis id="13134AB90295AA80B43839D11BA01946" bold="true" pageId="0" pageNumber="167">C.</emphasis>
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Paired 6-segmented maxillary palps;
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<emphasis id="1EB9066FC0FCB1B6EE691D619F1DF91E" bold="true" pageId="0" pageNumber="167">D.</emphasis>
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Drawing of bifurcated inferior appendages (inf app) and median process (med pro) in lateral view.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection id="BB34824035366F3B691666EE79A19E3F" pageId="0" pageNumber="167" type="diagnosis">
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<paragraph id="86EC21EA9F2771D680F1FA9842C7BAFC" pageId="0" pageNumber="167">Diagnosis.</paragraph>
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<paragraph id="98F4A59E2945F7AC618BF6A7728D3372" pageId="0" pageNumber="167">
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The male genitalia of
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<taxonomicName id="B4F53C7A71EB059F18BDB1EE1F2DE6AD" class="Insecta" family="Psychomyiidae" genus="Paduniella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paduniella" order="Trichoptera" pageId="0" pageNumber="167" phylum="Arthropoda" rank="genus">
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<emphasis id="E5CBB66232F0B7C199DC3BCDA7BA141E" italics="true" pageId="0" pageNumber="167">Paduniella</emphasis>
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</taxonomicName>
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species have inferior appendages whose apices are either bifurcated or unbifurcated (
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<bibRefCitation id="E79DB9E443091A61A5B7C0D9D036787B" author="Li, YJ" journalOrPublisher="Transactions of the Linnean Society of London, Zoology" pageId="0" pageNumber="167" publicationUrl="https://digitalcommons.unl.edu/insectamundi/276" refId="B12" refString="Li, YJ, Morse, JC, 1997. The Paduniella (Trichoptera: Psychomyiidae) of China, with a phylogeny of the World species. Insecta Mundi 276. https://digitalcommons.unl.edu/insectamundi/276" title="The Paduniella (Trichoptera: Psychomyiidae) of China, with a phylogeny of the World species. Insecta Mundi 276." url="https://digitalcommons.unl.edu/insectamundi/276" year="1997">Li and Morse 1997</bibRefCitation>
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). The extinct
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<taxonomicName id="922B9067571D25BA13042192011BE055" authorityName="Wichard" authorityYear="2023" class="Insecta" family="Psychomyiidae" genus="Paduniella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paduniella cretacea" order="Trichoptera" pageId="0" pageNumber="167" phylum="Arthropoda" rank="species" species="cretacea">
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<emphasis id="E3D509C640F5710ACC6349202B35A963" italics="true" pageId="0" pageNumber="167">Paduniella cretacea</emphasis>
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</taxonomicName>
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sp. nov. has similar to the extant species,
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<taxonomicName id="C28ECF873EBABD7AF353F53644BB2A59" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="P. tanidai" order="Trichoptera" pageId="0" pageNumber="167" phylum="Arthropoda" rank="species" species="tanidai">
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<emphasis id="217B7736B96B48CC0E3AF88E761555FB" italics="true" pageId="0" pageNumber="167">P. tanidai</emphasis>
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</taxonomicName>
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from Japan (Nishimoto, 2011) and
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<taxonomicName id="1C8A74B0025698E7B41D03331C2C8595" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="P. burmana" order="Trichoptera" pageId="0" pageNumber="167" phylum="Arthropoda" rank="species" species="burmana">
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<emphasis id="773E14E0E9D644EDE352C495AD63D91B" italics="true" pageId="0" pageNumber="167">P. burmana</emphasis>
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</taxonomicName>
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(Johanson & Olah, 2010) from the Oriental region, bifurcated inferior appendages that are deeply incised at the apex rather than superficially notched. The fossil species is distinguished from the two extant species in the deeply bifurcated apex of the inferior appendage because one fork is needle-like, acuminate, and curved and clearly protrudes above the other stouter fork.
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<taxonomicName id="4C1A664B49DBCFEAE9E5A4B10E2C150E" authorityName="Wichard" authorityYear="2023" class="Insecta" family="Psychomyiidae" genus="Paduniella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paduniella cretacea" order="Trichoptera" pageId="0" pageNumber="167" phylum="Arthropoda" rank="species" species="cretacea">
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<emphasis id="CA24F57CD14CEC8FF016E197763D994F" italics="true" pageId="0" pageNumber="167">Paduniella cretacea</emphasis>
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</taxonomicName>
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<emphasis id="41558A4873EB3B96A7298574CE67069E" bold="true" pageId="0" pageNumber="167">sp. nov.</emphasis>
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is the first fossil
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<taxonomicName id="E08B96C9DADD97F17802C24E59AE9C9E" class="Insecta" family="Psychomyiidae" genus="Paduniella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paduniella" order="Trichoptera" pageId="0" pageNumber="167" phylum="Arthropoda" rank="genus">
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<emphasis id="5AAFDD8A1B42E11DA74396858EEE1B59" italics="true" pageId="0" pageNumber="167">Paduniella</emphasis>
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</taxonomicName>
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species found in ca. 100 million year old mid-Cretaceous Burmese amber.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |