174 lines
21 KiB
XML
174 lines
21 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.238.3624" ID-PMC="PMC3496945" ID-Pensoft-Pub="1313-2970-238-57" ID-Pensoft-UUID="FFCDE657FFBEFFAEFF94FFFDA240FFB3" ID-PubMed="23226708" ID-Zenodo-Dep="577750" ID-ZooBank="928C2017C15A4F67956EB5747F3BF6D8" ModsDocID="1313-2970-238-57" checkinTime="1451248584044" checkinUser="pensoft" docAuthor="Zhang, Junfeng" docDate="2012" docId="4CCCE7414E4E50CF8A227315A835C099" docLanguage="en" docName="ZooKeys 238: 57-76" docOrigin="ZooKeys 238" docPubDate="2012-11-06" docSource="http://dx.doi.org/10.3897/zookeys.238.3624" docTitle="Orientisargus Zhang 2012" docType="treatment" docVersion="3" id="FFCDE657FFBEFFAEFF94FFFDA240FFB3" lastPageNumber="61" masterDocId="FFCDE657FFBEFFAEFF94FFFDA240FFB3" masterDocTitle="Orientisargidae fam. n., a new Jurassic family of Archisargoidea (Diptera, Brachycera), with review of Archisargidae from China" masterLastPageNumber="76" masterPageNumber="57" pageNumber="59" updateTime="1668154673256" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Orientisargidae fam. n., a new Jurassic family of Archisargoidea (Diptera, Brachycera), with review of Archisargidae from China</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Zhang, Junfeng</mods:namePart>
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<mods:affiliation>College of Palaeontology, Shenyang Normal University, Shenyang, 110034, P. R. China & Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing 210008, P. R. China</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2012</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2012-11-06</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>238</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>57</mods:start>
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<mods:end>76</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.238.3624</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.238.3624</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-238-57</mods:identifier>
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<mods:identifier type="ZooBank">928C2017C15A4F67956EB5747F3BF6D8</mods:identifier>
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<mods:identifier type="Pensoft-UUID">FFCDE657FFBEFFAEFF94FFFDA240FFB3</mods:identifier>
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<mods:identifier type="Zenodo-Dep">577750</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="182235765" LSID="urn:lsid:plazi:treatment:4CCCE7414E4E50CF8A227315A835C099" httpUri="http://treatment.plazi.org/id/4CCCE7414E4E50CF8A227315A835C099" lastPageId="4" lastPageNumber="61" pageId="2" pageNumber="59">
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<subSubSection pageId="2" pageNumber="59" type="nomenclature">
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<paragraph pageId="2" pageNumber="59">
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Family
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<taxonomicName LSID="4CCCE741-4E4E-50CF-8A22-7315A835C099" authority="Rohdendorf, 1962" authorityName="Zhang" authorityYear="2012" class="Insecta" family="Archisargidae" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Diptera" pageId="2" pageNumber="59" phylum="Arthropoda" rank="family">Archisargidae Rohdendorf, 1962</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="2" pageNumber="59" type="reference_group">
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<paragraph pageId="2" pageNumber="59">
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<taxonomicName authorityName="Zhang" authorityYear="2012" class="Insecta" family="Uranorhagionidae" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Diptera" pageId="2" pageNumber="59" phylum="Arthropoda" rank="family">Uranorhagionidae</taxonomicName>
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KY Zhang, Yang et Ren, 2010, p. 564, syn. n.
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</paragraph>
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<paragraph pageId="2" pageNumber="59">
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<taxonomicName class="Insecta" family="Origoasilidae" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Diptera" pageId="2" pageNumber="59" phylum="Arthropoda" rank="family">Origoasilidae</taxonomicName>
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KY Zhang, Yang et Ren, 2011, p. 995, syn. n.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="2" pageNumber="59" type="type genus">
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<paragraph pageId="2" pageNumber="59">Type genus.</paragraph>
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<paragraph pageId="2" pageNumber="59">
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<taxonomicName authorityName="Rohdendorf" authorityYear="1938" class="Insecta" family="Archisargidae" genus="Archisargus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Diptera" pageId="2" pageNumber="59" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="2" pageNumber="59">Archisargus</emphasis>
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</taxonomicName>
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Rohdendorf, 1938
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="3" lastPageNumber="60" pageId="2" pageNumber="59" type="included subfamilies">
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<paragraph pageId="2" pageNumber="59">Included subfamilies.</paragraph>
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<paragraph lastPageId="3" lastPageNumber="60" pageId="2" pageNumber="59">
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<taxonomicName authorityName="Rohdendorf" authorityYear="1962" lsidName="" pageId="2" pageNumber="59" rank="subFamily" subFamily="Archisarginae">Archisarginae</taxonomicName>
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Rohdendorf, 1962 and
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<taxonomicName lsidName="" pageId="2" pageNumber="59" rank="subFamily" subFamily="Uranorhagioninae">Uranorhagioninae</taxonomicName>
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KY Zhang, Yang et Ren, 2010, stat. n. (=
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<taxonomicName authorityName="Zhang" authorityYear="2012" lsidName="" pageId="2" pageNumber="59" rank="subFamily" subFamily="Mostovskisarginae">Mostovskisarginae</taxonomicName>
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JF Zhang, 2010, syn. n.)
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<pageBreakToken pageId="3" pageNumber="60" start="start">.</pageBreakToken>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="3" pageNumber="60" type="redefinition">
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<paragraph pageId="3" pageNumber="60">Redefinition.</paragraph>
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<paragraph pageId="3" pageNumber="60">
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Moderate- to large-sized flies. Body robust but usually narrow and long, strongly pubescent but devoid of bristles; first flagellomere of antenna unsegmented, arista well developed at tip of first flagellomere; hind legs stout and long, femora clavate; tibial spurs and pulvilliform empodium well developed; wing narrow and long, subpetiolate, alula absent; all longitudinal veins well developed, ending at wing margin; C running around entire wing margin although thinned near to, or beyond, wing tip, C and R strong, Sc and R1 long, R4+5 bifurcated, R2+3 usually straight and long but in some specific members R2+3 short and significantly curved [see
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Daohugosargus eximius (KY Zhang, Yang et Ren, 2008) comb. n. (originally Sharasargus eximius KY Zhang, Yang et Ren, 2008; after KY Zhang et al. 2008)." httpUri="https://binary.pensoft.net/fig/23016" pageId="3" pageNumber="60">Figure 4</figureCitation>
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,
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<taxonomicName class="Insecta" family="Archisargidae" genus="Daohugosargus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Diptera" pageId="3" pageNumber="60" phylum="Arthropoda" rank="species" species="eximius">
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<emphasis italics="true" pageId="3" pageNumber="60">Daohugosargus eximius</emphasis>
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</taxonomicName>
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(KY Zhang et al., 2008) comb. n., originally
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<taxonomicName class="Insecta" family="Archisargidae" genus="Sharasargus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Diptera" pageId="3" pageNumber="60" phylum="Arthropoda" rank="species" species="eximius">
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<emphasis italics="true" pageId="3" pageNumber="60">Sharasargus eximius</emphasis>
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</taxonomicName>
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KY Zhang et al., 2008], in most representatives crossvein r-m meeting R4+5 distad to Rs fork but in some specific members far basad to Rs fork (see
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Daohugosargus eximius (KY Zhang, Yang et Ren, 2008) comb. n. (originally Sharasargus eximius KY Zhang, Yang et Ren, 2008; after KY Zhang et al. 2008)." httpUri="https://binary.pensoft.net/fig/23016" pageId="3" pageNumber="60">Figure 4</figureCitation>
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,
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<taxonomicName class="Insecta" family="Archisargidae" genus="Daohugosargus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Diptera" pageId="3" pageNumber="60" phylum="Arthropoda" rank="species" species="eximius">
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<emphasis italics="true" pageId="3" pageNumber="60">Daohugosargus eximius</emphasis>
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</taxonomicName>
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), origin of Rs usually basad to, but in some specific members distad to, d base (see
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Daohugosargus eximius (KY Zhang, Yang et Ren, 2008) comb. n. (originally Sharasargus eximius KY Zhang, Yang et Ren, 2008; after KY Zhang et al. 2008)." httpUri="https://binary.pensoft.net/fig/23016" pageId="3" pageNumber="60">Figure 4</figureCitation>
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,
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<taxonomicName class="Insecta" family="Archisargidae" genus="Daohugosargus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Diptera" pageId="3" pageNumber="60" phylum="Arthropoda" rank="species" species="eximius">
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<emphasis italics="true" pageId="3" pageNumber="60">Daohugosargus eximius</emphasis>
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</taxonomicName>
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), discoidal cell shifted distally of wing midpoint.
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</paragraph>
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<caption httpUri="https://binary.pensoft.net/fig/23016" pageId="3" pageNumber="60" start="Figure 4" startId="F4">
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<paragraph pageId="3" pageNumber="60">
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<emphasis bold="true" pageId="3" pageNumber="60">Figure 4.</emphasis>
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<taxonomicName class="Insecta" family="Archisargidae" genus="Daohugosargus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Diptera" pageId="3" pageNumber="60" phylum="Arthropoda" rank="species" species="eximius">
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<emphasis italics="true" pageId="3" pageNumber="60">Daohugosargus eximius</emphasis>
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</taxonomicName>
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(KY Zhang, Yang et Ren, 2008) comb. n. (originally
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<taxonomicName class="Insecta" family="Archisargidae" genus="Sharasargus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Diptera" pageId="3" pageNumber="60" phylum="Arthropoda" rank="species" species="eximius">
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<emphasis italics="true" pageId="3" pageNumber="60">Sharasargus eximius</emphasis>
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</taxonomicName>
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KY Zhang, Yang et Ren, 2008; after KY Zhang et al. 2008).
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection lastPageId="4" lastPageNumber="61" pageId="3" pageNumber="60" type="remarks">
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<paragraph pageId="3" pageNumber="60">Remarks.</paragraph>
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<paragraph lastPageId="4" lastPageNumber="61" pageId="3" pageNumber="60">
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<bibRefCitation author="Rohdendorf, BB" journalOrPublisher="Seismic Publishing House, Beijing" pageId="12" pageNumber="69" refId="B9" refString="Rohdendorf, BB, 1962. Fundamentals of paleontology. Vol. 9. Arthropoda, Tracheata and Chelicerata. Academy of Sciences USSR Press, Moscow, 307-344." title="Fundamentals of paleontology. Vol. 9. Arthropoda, Tracheata and Chelicerata. Academy of Sciences USSR Press, Moscow, 307 - 344." year="1962">Rohdendorf (1962)</bibRefCitation>
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defined the family
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<taxonomicName class="Insecta" family="Archisargidae" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Diptera" pageId="3" pageNumber="60" phylum="Arthropoda" rank="family">Archisargidae</taxonomicName>
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based on a single poorly preserved wing from the Callovian-Oxfordian Karabastau Formation.
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<bibRefCitation author="Kovalev, VG" journalOrPublisher="Paleontological Journal" pageId="12" pageNumber="69" pagination="84 - 100" refId="B3" refString="Kovalev, VG, 1981. The oldest representatives of the Diptera with short antennae from the Jurassic in Siberia. Paleontological Journal 3: 84 - 100" title="The oldest representatives of the Diptera with short antennae from the Jurassic in Siberia." volume="3" year="1981">Kovalev (1981)</bibRefCitation>
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argued that this family was described from very poor material and thus nothing definite can be said about its systematic position. However, the type genus
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<taxonomicName authorityName="Rohdendorf" authorityYear="1938" class="Insecta" family="Archisargidae" genus="Archisargus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Diptera" pageId="3" pageNumber="60" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="3" pageNumber="60">Archisargus</emphasis>
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</taxonomicName>
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Rohdendorf, 1938, the only representative of the
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<taxonomicName class="Insecta" family="Archisargidae" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Diptera" pageId="3" pageNumber="60" phylum="Arthropoda" rank="family">Archisargidae</taxonomicName>
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, clearly has little in common with the Jurassic
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<taxonomicName class="Hexapoda" family="Rhagionidae" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Diptera" pageId="3" pageNumber="60" phylum="Arthropoda" rank="family">Rhagionidae</taxonomicName>
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: it is a large fly with a long (16 mm) narrow wing.
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<bibRefCitation author="Mostovski, MB" journalOrPublisher="Russian Entomological Journal" pageId="12" pageNumber="69" pagination="117 - 124" refId="B4" refString="Mostovski, MB, 1996a. To the knowledge of Archisargoidea (Diptera, Brachycera). Families Eremochaetidae and Archisargidae. Russian Entomological Journal 5: 117 - 124" title="To the knowledge of Archisargoidea (Diptera, Brachycera). Families Eremochaetidae and Archisargidae." volume="5" year="1996 a">Mostovski (1996a</bibRefCitation>
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,
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<bibRefCitation author="Mostovski, MB" journalOrPublisher="Museum of Northern Arizona Bulletin" pageId="12" pageNumber="69" pagination="329 - 332" refId="B5" refString="Mostovski, MB, 1996b. New species of the genus Mesosolva Hong, 1983 (Diptera, Archisargidae) from the Jurassic of Kazakhstan and Mongolia. Museum of Northern Arizona Bulletin 60: 329 - 332" title="New species of the genus Mesosolva Hong, 1983 (Diptera, Archisargidae) from the Jurassic of Kazakhstan and Mongolia." volume="60" year="1996 b">1996b</bibRefCitation>
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) described many new species referred, respectively, to some new genera or the known genera and assigned these to
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<taxonomicName class="Insecta" family="Archisargidae" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Diptera" pageId="3" pageNumber="60" phylum="Arthropoda" rank="family">Archisargidae</taxonomicName>
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. Immediately after these, he re-described the type species of type genus based on the holotype; and a redefinition of the
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<taxonomicName class="Insecta" family="Archisargidae" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Diptera" pageId="3" pageNumber="60" phylum="Arthropoda" rank="family">Archisargidae</taxonomicName>
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was proposed: wing venation not, or slightly, costalized; C running around entire wing margin although thinned beyond wing tip
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<pageBreakToken pageId="4" pageNumber="61" start="start">or</pageBreakToken>
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R4 end; R1 long; R4+5 bifurcated; crossvein r-m meeting R4+5 distad to Rs fork; base of discoidal cell distad to origin of Rs from R; M4, if present, connecting with discoidal cell (
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<bibRefCitation author="Mostovski, MB" journalOrPublisher="Paleontologicheskiy Zhurnal" pageId="12" pageNumber="69" pagination="72 - 77" refId="B6" refString="Mostovski, MB, 1997. To the knowledge of fossil dipterans of superfamily Archisargoidea (Diptera, Brachycera). Paleontologicheskiy Zhurnal 1: 72 - 77" title="To the knowledge of fossil dipterans of superfamily Archisargoidea (Diptera, Brachycera)." volume="1" year="1997">Mostovski 1997</bibRefCitation>
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).
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</paragraph>
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<paragraph pageId="4" pageNumber="61">
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Recently, numerous well preserved archisargid flies have been recovered from the Daohugou biota, China (
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<bibRefCitation author="Zhang, JF" journalOrPublisher="Paleontological Journal" pageId="13" pageNumber="70" pagination="409 - 412" refId="B19" refString="Zhang, JF, Zhang, HC, 2003. Two new species of archisargids (Insecta: Diptera: Archisargidae) from the Upper Jurassic Daohugou Formation (Inner Mongolia, Northeastern China). Paleontological Journal 37: 409 - 412" title="Two new species of archisargids (Insecta: Diptera: Archisargidae) from the Upper Jurassic Daohugou Formation (Inner Mongolia, Northeastern China)." volume="37" year="2003">JF Zhang and HC Zhang 2003</bibRefCitation>
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;
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<bibRefCitation author="Zhang, JF" journalOrPublisher="Palaeontology" pageId="12" pageNumber="69" pagination="307 - 317" publicationUrl="10.1111/j.1475-4983.2010.00934.x" refId="B12" refString="Zhang, JF, 2010a. Records of bizarre Jurassic brachycerans in the Daohugou biota, China (Diptera, Brachycera, Archisargidae and Rhagionemestriidae). Palaeontology 53 (2): 307 - 317, 10.1111/j.1475-4983.2010.00934.x" title="Records of bizarre Jurassic brachycerans in the Daohugou biota, China (Diptera, Brachycera, Archisargidae and Rhagionemestriidae)." url="10.1111/j.1475-4983.2010.00934.x" volume="53" year="2010 a">JF Zhang 2010a</bibRefCitation>
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,
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<bibRefCitation author="Zhang, JF" journalOrPublisher="Journal of Paleontology" pageId="12" pageNumber="69" pagination="1154 - 1159" refId="B16" refString="Zhang, JF, 2012b. Distinct but rare archisargid flies from Jurassic of China (Diptera, Archisargidae), with discussion of the systematic position of Origoasilus pingquanensis Zhang et al., 2011. Journal of Paleontology 85 (6): 1154 - 1159" title="Distinct but rare archisargid flies from Jurassic of China (Diptera, Archisargidae), with discussion of the systematic position of Origoasilus pingquanensis Zhang et al., 2011." volume="85" year="2012 b">2012b</bibRefCitation>
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;
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<bibRefCitation author="Zhang, KY" journalOrPublisher="Zootaxa" pageId="13" pageNumber="70" pagination="1 - 17" refId="B21" refString="Zhang, KY, Yang, D, Ren, D, Shih, CK, 2007a. The oldest Calosargus Mostovski, 1997 from the Middle Jurassic of China (Diptera: Brachycera: Archisargidae). Zootaxa 1645: 1 - 17" title="The oldest Calosargus Mostovski, 1997 from the Middle Jurassic of China (Diptera: Brachycera: Archisargidae)." volume="1645" year="2007 a">KY Zhang et al. 2007a</bibRefCitation>
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,
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<bibRefCitation author="Zhang, KY" journalOrPublisher="Annales Zoologici (Warszawa)" pageId="13" pageNumber="70" pagination="827 - 832" refId="B22" refString="Zhang, KY, Yang, D, Ren, D, Shih, CK, 2007b. The earliest species of the extinct genus Archisargus from China (Diptera: Brachycera: Archisargidae). Annales Zoologici (Warszawa) 57: 827 - 832" title="The earliest species of the extinct genus Archisargus from China (Diptera: Brachycera: Archisargidae)." volume="57" year="2007 b">2007b</bibRefCitation>
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,
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<bibRefCitation author="Zhang, KY" journalOrPublisher="Entomological Science" pageId="13" pageNumber="70" pagination="269 - 272" publicationUrl="10.1111/j.1479-8298.2008.00271.x" refId="B23" refString="Zhang, KY, Yang, D, Ren, D, 2008a. Middle Jurassic fossils of the genus Sharasargus from Inner Mongolia, China (Diptera: Archisargidae). Entomological Science 11: 269 - 272, 10.1111/j.1479-8298.2008.00271.x" title="Middle Jurassic fossils of the genus Sharasargus from Inner Mongolia, China (Diptera: Archisargidae)." url="10.1111/j.1479-8298.2008.00271.x" volume="11" year="2008 a">2008a</bibRefCitation>
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,
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<bibRefCitation author="Zhang, KY" journalOrPublisher="Zootaxa" pageId="13" pageNumber="70" pagination="61 - 65" refId="B27" refString="Zhang, KY, Li, JH, Yang, D, Ren, D, 2009. A new species of Archirhagio Rohdendorf, 1938 from the Middle Jurassic of Inner Mongolia of China (Diptera: Archisargidae). Zootaxa 1984: 61 - 65" title="A new species of Archirhagio Rohdendorf, 1938 from the Middle Jurassic of Inner Mongolia of China (Diptera: Archisargidae)." volume="1984" year="2009">2009</bibRefCitation>
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,
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<bibRefCitation author="Zhang, KY" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="13" pageNumber="70" pagination="563 - 572" publicationUrl="10.1111/j.1096-3642.2009.00552.x" refId="B28" refString="Zhang, KY, Yang, D, Ren, D, Shih, CK, 2010a. An evolutional special case in the lower Orthorrhapha: some attractive fossil flies from the Middle Jurassic of China (Insecta: Diptera: Brachycera). Zoological Journal of the Linnean Society 158: 563 - 572, 10.1111/j.1096-3642.2009.00552.x" title="An evolutional special case in the lower Orthorrhapha: some attractive fossil flies from the Middle Jurassic of China (Insecta: Diptera: Brachycera)." url="10.1111/j.1096-3642.2009.00552.x" volume="158" year="2010 a">2010a</bibRefCitation>
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,
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<bibRefCitation author="Zhang, KY" journalOrPublisher="Entomological science" pageId="13" pageNumber="70" pagination="75 - 80" publicationUrl="10.1111/j.1479-8298.2009.00350.x" refId="B29" refString="Zhang, KY, Yang, D, Ren, D, Shih, CK, 2010b. New archisargids from China (Insecta: Diptera). Entomological science 13: 75 - 80, 10.1111/j.1479-8298.2009.00350.x" title="New archisargids from China (Insecta: Diptera)." url="10.1111/j.1479-8298.2009.00350.x" volume="13" year="2010 b">2010b</bibRefCitation>
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). The familial diagnosis may be further supplemented based on information derived from these new results.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |