625 lines
74 KiB
XML
625 lines
74 KiB
XML
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<mods:title id="3A180EFEE9113565DC41016B0D5B7334">Allopatric cryptic diversity in the alpine species complex Phtheochroa frigidana s. lat. (Lepidoptera: Tortricidae)</mods:title>
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<mods:namePart id="FA1141B09A3E8DABD4949EF7A25F8E99">Zlatkov, Boyan</mods:namePart>
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<mods:affiliation id="0768FE19B5138D6C3D73249BDA7ECC2C">Institute of Biodiversity and Ecosystem Research, Bulgarian Academy of Sciences, 1 Tsar Osvoboditel Blvd, 1000 Sofia, Bulgaria. & National Museum of Natural History, Bulgarian Academy of Sciences, 1 Tsar Osvoboditel Blvd, 1000 Sofia, Bulgaria. & urn: lsid: zoobank. org: author: 3538 D 3 F 5 - 6822 - 403 F-B 9 E 6 - E 8 A 95 E 884404 & Corresponding author: bzlatkov @ gmail. com</mods:affiliation>
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<mods:nameIdentifier id="0B672D69DED974CFA5221626B685EEE9" type="email">bzlatkov@gmail.com</mods:nameIdentifier>
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<mods:namePart id="975FC7D8BCD7F649E8C2FFF4B2DE6981">Huemer, Peter</mods:namePart>
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<mods:affiliation id="13D941C99D886FE0039EAC66422BFCBA">Tiroler Landesmuseen Betriebsgesellschaft m. b. H., Sammlungs- und Forschungszentrum, Naturwissenschaftliche Sammlungen, Krajnc-Str. 1, A- 6060 Hall in Tirol, Austria. & Email: p. huemer @ tiroler-landesmuseen. at & urn: lsid: zoobank. org: author: 4 E 6 F 001 E-DB 9 D- 460 C-AB 39 - 0 CE 7 CC 773 B 59</mods:affiliation>
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<mods:title id="59604D5478E5CF3E34FE22AEAE0841D7">European Journal of Taxonomy</mods:title>
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<mods:part id="D0E1430D5CC2DCC7FC5A4C3183E0C6C3">
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<mods:date id="95C5E294FE51A91873389F0DF86A0E36">2017</mods:date>
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<mods:number id="6BBB939B73FB87853805B099B976EDA4">2017-11-21</mods:number>
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<subSubSection id="C35A7BF1144D4E7FB6986A7C8012DABD" box="[488,1100,1645,1672]" pageId="2" pageNumber="3" type="nomenclature">
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<paragraph id="8BFF287A144D4E7FB6986A7C8012DABD" blockId="2.[474,1113,1645,1707]" box="[488,1100,1645,1672]" pageId="2" pageNumber="3">
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<heading id="D0B79F16144D4E7FB6986A7C8012DABD" box="[488,1100,1645,1672]" centered="true" fontSize="11" level="2" pageId="2" pageNumber="3" reason="2">
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<taxonomicName id="4C4053F9144D4E7FB6986A7C879CDAB2" ID-CoL="92YKZ" authority="(Rebel, 1908)" authorityName="Zlatkov & Huemer" authorityYear="2017" baseAuthorityName="Rebel" baseAuthorityYear="1908" box="[488,962,1645,1672]" class="Insecta" family="Tortricidae" genus="Phtheochroa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="schawerdae" status="comb. nov.">
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<emphasis id="B934F468144D4E7FB6986A7C8744DABD" bold="true" box="[488,794,1645,1672]" italics="true" pageId="2" pageNumber="3">Phtheochroa schawerdae</emphasis>
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(Rebel, 1908)
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</taxonomicName>
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<taxonomicNameLabel id="A2074913144D4E7FB4BA6A7C8012DABD" box="[970,1100,1645,1672]" pageId="2" pageNumber="3" rank="species">comb. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C35A7BF1144D4E7FB6AA6A818007DA9F" box="[474,1113,1680,1707]" pageId="2" pageNumber="3" type="description">
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<paragraph id="8BFF287A144D4E7FB6AA6A818007DA9F" blockId="2.[474,1113,1645,1707]" box="[474,1113,1680,1707]" pageId="2" pageNumber="3">
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<figureCitation id="137B34FF144D4E7FB6AA6A818640DA9F" box="[474,542,1680,1706]" captionStart="Fig" captionStartId="4.[189,232,1824,1850]" captionTargetBox="[230,1357,265,1782]" captionTargetId="figure@4.[230,1357,265,1782]" captionTargetPageId="4" captionText="Fig. 1. Adults of Phtheochroa spp. A–D. P. schawerdae (Rebel, 1908) comb. nov. A. ♂, Bulgaria, Pirin Mts. B. ♂, Bulgaria, Rila Mts. C. ♀, Bulgaria, Rila Mts. D. ♂, Republic of Macedonia, Korab Mts. – E–F. P. alpinana sp. nov., France, Alpes Maritimes. E. Holotype, ♂. F. Paratype, ♀. – G. P. apenninana sp. nov., holotype, ♂, Italy, Gran Sasso National Park. – H. P. frigidana (Guenée, 1845) stat. rev., ♂, neotype of Eupoecilia frigidana, Andorra, Pyrenees. – I–J. P. cantabriana sp. nov., Spain, Picos de Europa National Park. I. Holotype, ♂. J. Paratype, ♂. Scale bar = 5 mm, all to scale." figureDoi="http://doi.org/10.5281/zenodo.3838414" httpUri="https://zenodo.org/record/3838414/files/figure.png" pageId="2" pageNumber="3">Figs 1</figureCitation>
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A–D, 2A–B, 3A–B, 4A–C, 5, 8A, 9, 10;
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<tableCitation id="C6C21DC1144D4E7FB3726A818007DA9F" box="[1026,1113,1680,1706]" captionStart="Table 1" captionStartId="7.[189,258,268,294]" captionTargetPageId="7" captionText="Table 1. Minimal uncorrected COI p-distances between different taxa of the P. frigidana s. lat. species complex." httpUri="http://table.plazi.org/id/DF3F78F214484E7AB7CD6D1D8574DD7D" pageId="2" pageNumber="3" tableUuid="DF3F78F214484E7AB7CD6D1D8574DD7D">Table 1</tableCitation>
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</paragraph>
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<subSubSection id="C35A7BF1144D4E7FB7CD6AC38772DBED" pageId="2" pageNumber="3" type="reference_group">
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<paragraph id="8BFF287A144D4E7FB7CD6AC3871DDAD8" blockId="2.[189,835,1746,1807]" box="[189,835,1746,1773]" pageId="2" pageNumber="3">
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<treatmentCitation id="0AE10E6B144D4E7FB7CD6AC38761DAD8" author="Schawerda K." box="[189,831,1746,1773]" page="255" pageId="2" pageNumber="3" year="1908">
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<taxonomicName id="4C4053F9144D4E7FB7CD6AC38761DAD8" ID-CoL="92YKZ" authority="Rebel in Schawerda, 1908: 255" authorityName="Rebel in Schawerda" authorityPageNumber="255" authorityYear="1908" box="[189,831,1746,1773]" class="Insecta" family="Tortricidae" genus="Conchylis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="schawerdae">
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<emphasis id="B934F468144D4E7FB7CD6AC3859BDAD9" box="[189,453,1746,1772]" italics="true" pageId="2" pageNumber="3">Conchylis schawerdae</emphasis>
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Rebel in
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<bibRefCitation id="EFD1558B144D4E7FB5466AC38761DAD8" author="Schawerda K." box="[566,831,1746,1773]" pageId="2" pageNumber="3" pagination="250 - 273" refId="ref12480" refString="Schawerda K. 1908. Bericht der Sektion fur Lepidopterologie. Verhandlungen der Kaiserlich- Koniglichen Zoologisch-Botanischen Gesellschaft in Wien 58: 250 - 273." type="journal article" year="1908">Schawerda, 1908: 255</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8BFF287A144D4E7FB7CD6AE4875EDB3A" blockId="2.[189,835,1746,1807]" box="[189,768,1781,1807]" pageId="2" pageNumber="3">
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<treatmentCitationGroup id="AB500F54144D4E7FB7CD6AE4875EDB3A" box="[189,768,1781,1807]" pageId="2" pageNumber="3">
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<treatmentCitation id="0AE10E6B144D4E7FB7CD6AE486DADB3A" author="Rebel H." box="[189,644,1781,1807]" page="42" pageId="2" pageNumber="3" year="1916">
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<taxonomicName id="4C4053F9144D4E7FB7CD6AE486DADB3A" ID-CoL="95B8C" authority="Rebel, 1916: 42" authorityName="Rebel" authorityPageNumber="42" authorityYear="1916" box="[189,644,1781,1807]" class="Insecta" family="Tortricidae" genus="Euxanthis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="drenowskyi">
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<emphasis id="B934F468144D4E7FB7CD6AE485E1DB3A" box="[189,447,1781,1807]" italics="true" pageId="2" pageNumber="3">Euxanthis drenowskyi</emphasis>
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||
<bibRefCitation id="EFD1558B144D4E7FB6B56AE486DADB3A" author="Rebel H." box="[453,644,1781,1807]" pageId="2" pageNumber="3" pagination="36 - 42" refId="ref12430" refString="Rebel H. 1916. Beitrag zur Lepidopterenfauna Bulgariens. Verhandlungen der Kaiserlich-Koniglichen Zoologisch-Botanischen Gesellschaft in Wien 66: 36 - 42." type="journal article" year="1916">Rebel, 1916: 42</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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.
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<emphasis id="B934F468144D4E7FB5FF6AE4875EDB3A" bold="true" box="[655,768,1781,1807]" pageId="2" pageNumber="3">Syn. nov.</emphasis>
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8BFF287A144D4E7FB7CD6B2680FCDB64" blockId="2.[189,1186,1846,1908]" box="[189,1186,1846,1873]" pageId="2" pageNumber="3">
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<treatmentCitationGroup id="AB500F54144D4E7FB7CD6B2680FCDB64" box="[189,1186,1846,1873]" pageId="2" pageNumber="3">
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<taxonomicName id="4C4053F9144D4E7FB7CD6B268599DB65" box="[189,455,1846,1872]" class="Insecta" family="Tortricidae" genus="Hysterosia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="drenowskyi">
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<emphasis id="B934F468144D4E7FB7CD6B268599DB65" box="[189,455,1846,1872]" italics="true" pageId="2" pageNumber="3">Hysterosia drenowskyi</emphasis>
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</taxonomicName>
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–
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<treatmentCitation id="0AE10E6B144D4E7FB6946B278697DB64" author="Razowski J." box="[484,713,1846,1873]" page="72" pageId="2" pageNumber="3" year="1970">
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<bibRefCitation id="EFD1558B144D4E7FB6946B278697DB64" author="Razowski J." box="[484,713,1846,1873]" pageId="2" pageNumber="3" pagination="3" refId="ref12243" refString="Razowski J. 1970. Cochylidae. In: Amsel H. G., Gregor F. & Reisser H. (eds) Microlepidoptera Palaearctica 3. Georg Fromme, Vienna, Austria." type="book chapter" year="1970">Razowski 1970: 72</bibRefCitation>
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</treatmentCitation>
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, pl. 1, fig. 2, pl. 29, fig. 2, pl. 112, fig. 2.
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</treatmentCitationGroup>
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</paragraph>
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||
<paragraph id="8BFF287A144D4E7FB7CD6B4B86B8DB41" blockId="2.[189,1186,1846,1908]" box="[189,742,1882,1908]" pageId="2" pageNumber="3">
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<treatmentCitationGroup id="AB500F54144D4E7FB7CD6B4B86B8DB41" box="[189,742,1882,1908]" pageId="2" pageNumber="3">
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<taxonomicName id="4C4053F9144D4E7FB7CD6B4B85BEDB41" ID-CoL="4GX8X" baseAuthorityName="Rebel" baseAuthorityYear="1916" box="[189,480,1882,1908]" class="Insecta" family="Tortricidae" genus="Phtheochroa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="drenowskyi">
|
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<emphasis id="B934F468144D4E7FB7CD6B4B85BEDB41" box="[189,480,1882,1908]" italics="true" pageId="2" pageNumber="3">Phtheochroa drenowskyi</emphasis>
|
||
</taxonomicName>
|
||
–
|
||
<treatmentCitation id="0AE10E6B144D4E7FB68D6B4B86BDDB41" author="Razowski J." box="[509,739,1882,1908]" page="38" pageId="2" pageNumber="3" year="2002">
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<bibRefCitation id="EFD1558B144D4E7FB68D6B4B86BDDB41" author="Razowski J." box="[509,739,1882,1908]" pageId="2" pageNumber="3" refId="ref12358" refString="Razowski J. 2002. Tortricidae of Europe. Volume 1. Tortricinae and Chlidanothinae. Frantisek Slamka, Bratislava, Slovakia." type="book" year="2002">Razowski 2002: 38</bibRefCitation>
|
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</treatmentCitation>
|
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.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="8BFF287A144D4E7FB7CD6B8D8771DB80" blockId="2.[189,815,1947,2009]" box="[189,815,1947,1974]" pageId="2" pageNumber="3">
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<treatmentCitationGroup id="AB500F54144D4E7FB7CD6B8D8771DB80" box="[189,815,1947,1974]" pageId="2" pageNumber="3">
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non
|
||
<taxonomicName id="4C4053F9144D4E7FB7806B8A8652DB80" box="[240,524,1947,1973]" class="Insecta" family="Tortricidae" genus="Phtheochroa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="drenovskyi">
|
||
<emphasis id="B934F468144D4E7FB7806B8A8652DB80" box="[240,524,1947,1973]" italics="true" pageId="2" pageNumber="3">Phtheochroa drenovskyi</emphasis>
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||
</taxonomicName>
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||
–
|
||
<treatmentCitation id="0AE10E6B144D4E7FB5596B8A8775DB80" author="Razowski J." box="[553,811,1947,1973]" page="103" pageId="2" pageNumber="3" year="1991">
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<bibRefCitation id="EFD1558B144D4E7FB5596B8A8775DB80" author="Razowski J." box="[553,811,1947,1973]" pageId="2" pageNumber="3" pagination="99 - 162" refId="ref12279" refString="Razowski J. 1991 a. The catalogue of the species of Tortricidae (Lepidoptera). Part I: Palaearctic Chlidanothinae and Tortricinae: Cochylini, Tortricini, Ceracini, Cnephasiini. Acta Zoologica Cracoviensia 34 (1): 99 - 162." type="journal article" year="1991">Razowski 1991a: 103</bibRefCitation>
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</treatmentCitation>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="8BFF287A144D4E7FB7CD6BAE8772DBED" blockId="2.[189,815,1947,2009]" box="[189,812,1982,2009]" pageId="2" pageNumber="3">
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<treatmentCitationGroup id="AB500F54144D4E7FB7CD6BAE8772DBED" box="[189,812,1982,2009]" pageId="2" pageNumber="3">
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non
|
||
<taxonomicName id="4C4053F9144D4E7FB7806BAF8659DBED" box="[240,519,1982,2008]" class="Insecta" family="Tortricidae" genus="Phtheochroa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="drenovskii">
|
||
<emphasis id="B934F468144D4E7FB7806BAF8659DBED" box="[240,519,1982,2008]" italics="true" pageId="2" pageNumber="3">Phtheochroa drenovskii</emphasis>
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||
</taxonomicName>
|
||
–
|
||
<treatmentCitation id="0AE10E6B144D4E7FB5556BAF8779DBED" author="Razowski" box="[549,807,1982,2008]" page="166" pageId="2" pageNumber="3" year="1991">
|
||
<bibRefCitation id="EFD1558B144D4E7FB5556BAF8779DBED" author="Razowski" box="[549,807,1982,2008]" pageId="2" pageNumber="3" pagination="163 - 187" refId="ref12324" refString="Razowski 1991 b. Notes on Cochylinae genus Phtheochroa Stephens (Lepidoptera: Tortricidae) with descriptions of new American species. Acta Zoologica Cracoviensia 34 (1): 163 - 187." type="journal article" year="1991">Razowski 1991b: 166</bibRefCitation>
|
||
</treatmentCitation>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35A7BF1144C4E7EB7CD6D1A86D0DE44" pageId="3" pageNumber="4" type="diagnosis">
|
||
<paragraph id="8BFF287A144C4E7EB7CD6D1A8568DD10" blockId="3.[189,310,267,293]" box="[189,310,267,293]" pageId="3" pageNumber="4">
|
||
<heading id="D0B79F16144C4E7EB7CD6D1A8568DD10" bold="true" box="[189,310,267,293]" fontSize="11" level="3" pageId="3" pageNumber="4" reason="3">
|
||
<emphasis id="B934F468144C4E7EB7CD6D1A8568DD10" bold="true" box="[189,310,267,293]" pageId="3" pageNumber="4">Diagnosis</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BFF287A144C4E7EB7CD6D2C86D0DE44" blockId="3.[189,1399,317,625]" pageId="3" pageNumber="4">
|
||
Externally, this species resembles
|
||
<taxonomicName id="4C4053F9144C4E7EB5256D2F86A0DD62" authority="Zlatkov & Huemer, 2017" authorityName="Zlatkov & Huemer" authorityYear="2017" box="[597,766,318,343]" class="Insecta" family="Tortricidae" genus="Phtheochroa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="apenninana" status="sp. nov.">
|
||
<emphasis id="B934F468144C4E7EB5256D2F86A0DD62" box="[597,766,318,343]" italics="true" pageId="3" pageNumber="4">P. apenninana</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2074913144C4E7EB4766D2F873BDD6D" box="[774,869,318,344]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, with a deep yellow forewing upperside and a dark grey hindwing. The most conspicuous difference can be seen in the vesica: the axes of the right diverticulum and the sclerotized phallus form an angle of 100–120° in
|
||
<taxonomicName id="4C4053F9144C4E7EB3766D9580F1DDAB" authorityName="Rebel in Schawerda" authorityYear="1908" box="[1030,1199,388,414]" class="Insecta" family="Tortricidae" genus="Conchylis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="schawerdae">
|
||
<emphasis id="B934F468144C4E7EB3766D9580F1DDAB" box="[1030,1199,388,414]" italics="true" pageId="3" pageNumber="4">P. schawerdae</emphasis>
|
||
</taxonomicName>
|
||
and a maximum of 90° in
|
||
<taxonomicName id="4C4053F9144C4E7EB6406DB98584DDF4" authority="Zlatkov & Huemer, 2017" authorityName="Zlatkov & Huemer" authorityYear="2017" box="[304,474,424,449]" class="Insecta" family="Tortricidae" genus="Phtheochroa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="apenninana" status="sp. nov.">
|
||
<emphasis id="B934F468144C4E7EB6406DB98584DDF4" box="[304,474,424,449]" italics="true" pageId="3" pageNumber="4">P. apenninana</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2074913144C4E7EB6926DB6861CDDF4" box="[482,578,423,449]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
|
||
; the right part of the vesica is larger in
|
||
<taxonomicName id="4C4053F9144C4E7EB36D6DB98098DDF4" authority="Zlatkov & Huemer, 2017" authorityName="Zlatkov & Huemer" authorityYear="2017" box="[1053,1222,424,449]" class="Insecta" family="Tortricidae" genus="Phtheochroa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="apenninana" status="sp. nov.">
|
||
<emphasis id="B934F468144C4E7EB36D6DB98098DDF4" box="[1053,1222,424,449]" italics="true" pageId="3" pageNumber="4">P. apenninana</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2074913144C4E7EB3BF6DB68175DDF4" box="[1231,1323,423,449]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
|
||
Other characters of less importance are the shape of the valva (wider in
|
||
<taxonomicName id="4C4053F9144C4E7EB4876DDA80FDDDD1" authorityName="Rebel in Schawerda" authorityYear="1908" box="[1015,1187,458,484]" class="Insecta" family="Tortricidae" genus="Conchylis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="schawerdae">
|
||
<emphasis id="B934F468144C4E7EB4876DDA80FDDDD1" box="[1015,1187,458,484]" italics="true" pageId="3" pageNumber="4">P. schawerdae</emphasis>
|
||
</taxonomicName>
|
||
, more slender in
|
||
<taxonomicName id="4C4053F9144C4E7EB7CD6DFF8539DE32" authority="Zlatkov & Huemer, 2017" authorityName="Zlatkov & Huemer" authorityYear="2017" box="[189,359,494,519]" class="Insecta" family="Tortricidae" genus="Phtheochroa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="apenninana" status="sp. nov.">
|
||
<emphasis id="B934F468144C4E7EB7CD6DFF8539DE32" box="[189,359,494,519]" italics="true" pageId="3" pageNumber="4">P. apenninana</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2074913144C4E7EB61F6DFF8593DE3D" box="[367,461,494,520]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
|
||
) and transtilla (short, with wide base in
|
||
<taxonomicName id="4C4053F9144C4E7EB4C26DFF8002DE32" authority="Zlatkov & Huemer, 2017" authorityName="Zlatkov & Huemer" authorityYear="2017" box="[946,1116,494,519]" class="Insecta" family="Tortricidae" genus="Phtheochroa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="apenninana" status="sp. nov.">
|
||
<emphasis id="B934F468144C4E7EB4C26DFF8002DE32" box="[946,1116,494,519]" italics="true" pageId="3" pageNumber="4">P. apenninana</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2074913144C4E7EB3146DFF809FDE3D" box="[1124,1217,494,520]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
|
||
but variable in
|
||
<taxonomicName id="4C4053F9144C4E7EB7CD6E00853ADE1F" authorityName="Rebel in Schawerda" authorityYear="1908" box="[189,356,528,554]" class="Insecta" family="Tortricidae" genus="Conchylis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="schawerdae">
|
||
<emphasis id="B934F468144C4E7EB7CD6E00853ADE1F" box="[189,356,528,554]" italics="true" pageId="3" pageNumber="4">P. schawerdae</emphasis>
|
||
</taxonomicName>
|
||
). The ventral phallic process is strongly curved to the right, in contrast to all other species of the group. The upperside wing colour easily distinguishes
|
||
<taxonomicName id="4C4053F9144C4E7EB4E36E258062DE7B" authorityName="Rebel in Schawerda" authorityYear="1908" box="[915,1084,564,590]" class="Insecta" family="Tortricidae" genus="Conchylis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="schawerdae">
|
||
<emphasis id="B934F468144C4E7EB4E36E258062DE7B" box="[915,1084,564,590]" italics="true" pageId="3" pageNumber="4">P. schawerdae</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<taxonomicName id="4C4053F9144C4E7EB3F56E258155DE7B" authority="Zlatkov & Huemer, 2017" authorityName="Zlatkov & Huemer" authorityYear="2017" box="[1157,1291,564,590]" class="Insecta" family="Tortricidae" genus="Phtheochroa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="alpinana" status="sp. nov.">
|
||
<emphasis id="B934F468144C4E7EB3F56E258155DE7B" box="[1157,1291,564,590]" italics="true" pageId="3" pageNumber="4">P. alpinana</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2074913144C4E7EB2646E25812CDE7B" box="[1300,1394,564,590]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="4C4053F9144C4E7EB7CD6E498519DE44" authorityName="Zlatkov & Huemer" authorityYear="2017" baseAuthorityName="Guenee" baseAuthorityYear="1845" box="[189,327,599,625]" class="Insecta" family="Tortricidae" genus="Phtheochroa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="frigidana">
|
||
<emphasis id="B934F468144C4E7EB7CD6E498519DE44" box="[189,327,599,625]" italics="true" pageId="3" pageNumber="4">P. frigidana</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C4053F9144C4E7EB6F06E498672DE44" authority="Zlatkov & Huemer, 2017" authorityName="Zlatkov & Huemer" authorityYear="2017" box="[384,556,599,625]" class="Insecta" family="Tortricidae" genus="Phtheochroa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="cantabriana" status="sp. nov.">
|
||
<emphasis id="B934F468144C4E7EB6F06E498672DE44" box="[384,556,599,625]" italics="true" pageId="3" pageNumber="4">P. cantabriana</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2074913144C4E7EB5446E4686D0DE44" box="[564,654,599,625]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35A7BF1144C4E7EB7CD6EB180BFD87A" pageId="3" pageNumber="4" type="materials_examined">
|
||
<paragraph id="8BFF287A144C4E7EB7CD6EB185F4DE8F" blockId="3.[189,426,672,698]" box="[189,426,672,698]" pageId="3" pageNumber="4">
|
||
<heading id="D0B79F16144C4E7EB7CD6EB185F4DE8F" bold="true" box="[189,426,672,698]" fontSize="11" level="3" pageId="3" pageNumber="4" reason="3">
|
||
<emphasis id="B934F468144C4E7EB7CD6EB185F4DE8F" bold="true" box="[189,426,672,698]" pageId="3" pageNumber="4">Material examined</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BFF287A144C4E7EB7CD6EC38701DFD6" blockId="3.[189,1406,721,995]" pageId="3" pageNumber="4">
|
||
<collectingCountry id="F35768EA144C4E7EB7CD6EC38507DED9" box="[189,345,722,748]" name="Bulgaria" pageId="3" pageNumber="4">BULGARIA</collectingCountry>
|
||
:
|
||
<specimenCount id="9D46E3F3144C4E7EB6116EC385D3DED8" box="[353,397,722,749]" pageId="3" pageNumber="4" type="male">1 ♂</specimenCount>
|
||
, Rila Mts, Yurushki chal [Ovcharets] summit,
|
||
<quantity id="4CB8859F144C4E7EB4EC6EC387ACDED8" box="[924,1010,722,749]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="3" pageNumber="4" unit="m" value="2100.0">
|
||
<date id="FFFE0EBA144C4E7EB4EC6EC38789DED9" box="[924,983,722,748]" pageId="3" pageNumber="4" value="2100">2100</date>
|
||
m
|
||
</quantity>
|
||
a.s.l.,
|
||
<date id="FFFE0EBA144C4E7EB3496EC38098DED9" box="[1081,1222,722,748]" pageId="3" pageNumber="4" value="1907-07-24">
|
||
24 Jul.
|
||
<date id="FFFE0EBA144C4E7EB3F96EC38098DED9" box="[1161,1222,722,748]" pageId="3" pageNumber="4" value="1907" year="1907">1907</date>
|
||
</date>
|
||
,
|
||
<collectionCode id="ED51B0BF144C4E7EB3BE6EC380B9DED9" box="[1230,1255,722,748]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15406" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="3" pageNumber="4" type="Herbarium">A</collectionCode>
|
||
. Drenowsky leg. (
|
||
<collectionCode id="ED51B0BF144C4E7EB7886EE4853CDF3A" box="[248,354,757,783]" pageId="3" pageNumber="4">NMNHS</collectionCode>
|
||
);
|
||
<specimenCount id="9D46E3F3144C4E7EB60A6EE485F8DF25" box="[378,422,757,784]" pageId="3" pageNumber="4" type="male">1 ♂</specimenCount>
|
||
, Rila Mts, Nalbant [Kovach] summit,
|
||
<quantity id="4CB8859F144C4E7EB4116EE487E6DF25" box="[865,952,757,784]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="3" pageNumber="4" unit="m" value="2000.0">
|
||
<date id="FFFE0EBA144C4E7EB4116EE487C2DF3A" box="[865,924,757,783]" pageId="3" pageNumber="4" value="2000">2000</date>
|
||
m
|
||
</quantity>
|
||
a.s.l.,
|
||
<date id="FFFE0EBA144C4E7EB3726EE480CCDF3A" box="[1026,1170,757,783]" pageId="3" pageNumber="4" value="1907-08-01">
|
||
1 Aug.
|
||
<date id="FFFE0EBA144C4E7EB3256EE480CCDF3A" box="[1109,1170,757,783]" pageId="3" pageNumber="4" value="1907" year="1907">1907</date>
|
||
</date>
|
||
,
|
||
<collectionCode id="ED51B0BF144C4E7EB3EB6EE480EADF3A" box="[1179,1204,757,783]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15406" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="3" pageNumber="4" type="Herbarium">A</collectionCode>
|
||
. Drenowsky leg. (
|
||
<collectionCode id="ED51B0BF144C4E7EB7B76F08856FDF06" box="[199,305,793,819]" pageId="3" pageNumber="4">NMNHS</collectionCode>
|
||
);
|
||
<specimenCount id="9D46E3F3144C4E7EB63A6F0885D3DF06" box="[330,397,793,819]" pageId="3" pageNumber="4" type="male">3 ♂♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D46E3F3144C4E7EB6EA6F088598DF06" box="[410,454,793,819]" pageId="3" pageNumber="4" type="female">1 ♀</specimenCount>
|
||
, Rila Mts, Tiha Rila area,
|
||
<quantity id="4CB8859F144C4E7EB58B6F09870DDF06" box="[763,851,792,819]" metricMagnitude="3" metricUnit="m" metricValue="1.85" pageId="3" pageNumber="4" unit="m" value="1850.0">
|
||
<date id="FFFE0EBA144C4E7EB58B6F098768DF06" box="[763,822,792,819]" pageId="3" pageNumber="4" value="1850">1850</date>
|
||
m
|
||
</quantity>
|
||
a.s.l.,
|
||
<geoCoordinate id="EE744EBD144C4E7EB4EF6F06806BDF06" box="[927,1077,791,819]" degrees="42" direction="north" minutes="08" orientation="latitude" pageId="3" pageNumber="4" precision="15" seconds="27" value="42.140835">
|
||
42°08′27′′
|
||
<collectionCode id="ED51B0BF144C4E7EB36D6F08806BDF06" box="[1053,1077,793,819]" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13092" lsid="urn:lsid:biocol.org:col:13092" name="Nanjing University" pageId="3" pageNumber="4" type="Herbarium">N</collectionCode>
|
||
</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE744EBD144C4E7EB3316F06808DDF06" box="[1089,1235,791,819]" degrees="23" direction="east" minutes="26" orientation="longitude" pageId="3" pageNumber="4" precision="15" seconds="42" value="23.445">
|
||
23°26′42′′
|
||
<collectionCode id="ED51B0BF144C4E7EB3CF6F08808DDF06" box="[1215,1235,793,819]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15670" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="3" pageNumber="4" type="Herbarium">E</collectionCode>
|
||
</geoCoordinate>
|
||
,
|
||
<date id="FFFE0EBA144C4E7EB3AF6F08812FDF06" box="[1247,1393,793,819]" pageId="3" pageNumber="4" value="2013-08-02">
|
||
2 Aug.
|
||
<date id="FFFE0EBA144C4E7EB2446F08812FDF06" box="[1332,1393,793,819]" pageId="3" pageNumber="4" value="2013" year="2013">2013</date>
|
||
</date>
|
||
,
|
||
<collectionCode id="ED51B0BF144C4E7EB7CD6F2D848ADF63" box="[189,212,828,854]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15534" lsid="urn:lsid:biocol.org:col:15534" name="Botanischer Garten und Botanisches Museum Berlin-Dahlem, Zentraleinrichtung der Freien Universitaet" pageId="3" pageNumber="4" type="Herbarium">B</collectionCode>
|
||
. Zlatkov leg. (
|
||
<collectionCode id="ED51B0BF144C4E7EB60B6F2D8593DF63" box="[379,461,828,854]" country="Austria" httpUri="http://grbio.org/cool/7159-6y1v" name="Tiroler Landesmuseum Ferdinandeum" pageId="3" pageNumber="4">TLMF</collectionCode>
|
||
) (genitalia slide,
|
||
<specimenCount id="9D46E3F3144C4E7EB5E76F2D86F3DF63" box="[663,685,828,854]" pageId="3" pageNumber="4" type="female">♀</specimenCount>
|
||
, 1/
|
||
<date id="FFFE0EBA144C4E7EB5BE6F2D8763DF63" box="[718,829,828,854]" pageId="3" pageNumber="4" value="2013-08-02">2.8.2013</date>
|
||
);
|
||
<specimenCount id="9D46E3F3144C4E7EB4216F2D87CBDF63" box="[849,917,828,854]" pageId="3" pageNumber="4" type="male">2 ♂♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D46E3F3144C4E7EB4D16F2D8792DF63" box="[929,972,828,854]" pageId="3" pageNumber="4" type="female">1 ♀</specimenCount>
|
||
, Rila Mts, Kalin Dam,
|
||
<geoCoordinate id="EE744EBD144C4E7EB3AD6F2B812CDF63" box="[1245,1394,826,854]" degrees="42" direction="north" minutes="11" orientation="latitude" pageId="3" pageNumber="4" precision="15" seconds="03" value="42.184166">
|
||
42°11′03′′
|
||
<collectionCode id="ED51B0BF144C4E7EB22A6F2D812CDF63" box="[1370,1394,828,854]" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13092" lsid="urn:lsid:biocol.org:col:13092" name="Nanjing University" pageId="3" pageNumber="4" type="Herbarium">N</collectionCode>
|
||
</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE744EBD144C4E7EB7CD6F4C8510DF4C" box="[189,334,861,889]" degrees="23" direction="east" minutes="15" orientation="longitude" pageId="3" pageNumber="4" precision="15" seconds="42" value="23.261667">
|
||
23°15′42′′
|
||
<collectionCode id="ED51B0BF144C4E7EB64A6F4E8510DF4C" box="[314,334,863,889]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15670" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="3" pageNumber="4" type="Herbarium">E</collectionCode>
|
||
</geoCoordinate>
|
||
,
|
||
<quantity id="4CB8859F144C4E7EB62A6F4E85ECDF4C" box="[346,434,863,890]" metricMagnitude="3" metricUnit="m" metricValue="2.45" pageId="3" pageNumber="4" unit="m" value="2450.0">
|
||
<date id="FFFE0EBA144C4E7EB62A6F4E85CBDF4F" box="[346,405,863,890]" pageId="3" pageNumber="4" value="2450">2450</date>
|
||
m
|
||
</quantity>
|
||
a.s.l.,
|
||
<date id="FFFE0EBA144C4E7EB68D6F4E86C3DF4C" box="[509,669,863,889]" pageId="3" pageNumber="4" value="2013-08-20">
|
||
20 Aug.
|
||
<date id="FFFE0EBA144C4E7EB5106F4E86C3DF4C" box="[608,669,863,889]" pageId="3" pageNumber="4" value="2013" year="2013">2013</date>
|
||
</date>
|
||
,
|
||
<collectionCode id="ED51B0BF144C4E7EB5D86F4E86E1DF4C" box="[680,703,863,889]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15534" lsid="urn:lsid:biocol.org:col:15534" name="Botanischer Garten und Botanisches Museum Berlin-Dahlem, Zentraleinrichtung der Freien Universitaet" pageId="3" pageNumber="4" type="Herbarium">B</collectionCode>
|
||
. Zlatkov leg. (
|
||
<collectionCode id="ED51B0BF144C4E7EB4166F4E87ECDF4C" box="[870,946,863,889]" country="Bulgaria" httpUri="http://grbio.org/cool/7wb6-ejfj" name="University of Sofia, Biology Faculty" pageId="3" pageNumber="4">BFUS</collectionCode>
|
||
) (genitalia slide,
|
||
<specimenCount id="9D46E3F3144C4E7EB30B6F4E80CDDF4C" box="[1147,1171,863,889]" pageId="3" pageNumber="4" type="male">♂</specimenCount>
|
||
, 1/
|
||
<date id="FFFE0EBA144C4E7EB3C26F4E816EDF4C" box="[1202,1328,863,889]" pageId="3" pageNumber="4" value="2013-08-20">20.8.2013</date>
|
||
);
|
||
<specimenCount id="9D46E3F3144C4E7EB2346F4E8129DF4C" box="[1348,1399,863,889]" pageId="3" pageNumber="4" type="male">1 ♂</specimenCount>
|
||
, Rila Mts, Granchar Chalet,
|
||
<geoCoordinate id="EE744EBD144C4E7EB5756F9186C2DFA8" box="[517,668,896,925]" degrees="42" direction="north" minutes="07" orientation="latitude" pageId="3" pageNumber="4" precision="15" seconds="18" value="42.121666">
|
||
42°07′18′′
|
||
<collectionCode id="ED51B0BF144C4E7EB5F46F9286C2DFA8" box="[644,668,899,925]" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13092" lsid="urn:lsid:biocol.org:col:13092" name="Nanjing University" pageId="3" pageNumber="4" type="Herbarium">N</collectionCode>
|
||
</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE744EBD144C4E7EB5D96F918762DFA8" box="[681,828,896,925]" degrees="23" direction="east" minutes="35" orientation="longitude" pageId="3" pageNumber="4" precision="15" seconds="36" value="23.593334">
|
||
23°35′36′′
|
||
<collectionCode id="ED51B0BF144C4E7EB4586F928762DFA8" box="[808,828,899,925]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15670" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="3" pageNumber="4" type="Herbarium">E</collectionCode>
|
||
</geoCoordinate>
|
||
,
|
||
<quantity id="4CB8859F144C4E7EB4396F9387FDDFA8" box="[841,931,898,925]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="3" pageNumber="4" unit="m" value="2200.0">
|
||
<date id="FFFE0EBA144C4E7EB4396F9387DADFA9" box="[841,900,898,924]" pageId="3" pageNumber="4" value="2200">2200</date>
|
||
m
|
||
</quantity>
|
||
a.s.l.,
|
||
<date id="FFFE0EBA144C4E7EB4816F9380D8DFA9" box="[1009,1158,898,924]" pageId="3" pageNumber="4" value="2013-08-06">
|
||
6 Aug.
|
||
<date id="FFFE0EBA144C4E7EB3396F9380D8DFA9" box="[1097,1158,898,924]" pageId="3" pageNumber="4" value="2013" year="2013">2013</date>
|
||
</date>
|
||
,
|
||
<collectionCode id="ED51B0BF144C4E7EB3E36F9280F4DFA8" box="[1171,1194,899,925]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15534" lsid="urn:lsid:biocol.org:col:15534" name="Botanischer Garten und Botanisches Museum Berlin-Dahlem, Zentraleinrichtung der Freien Universitaet" pageId="3" pageNumber="4" type="Herbarium">B</collectionCode>
|
||
. Zlatkov (
|
||
<collectionCode id="ED51B0BF144C4E7EB2526F938130DFA9" box="[1314,1390,898,924]" country="Bulgaria" httpUri="http://grbio.org/cool/7wb6-ejfj" name="University of Sofia, Biology Faculty" pageId="3" pageNumber="4">BFUS</collectionCode>
|
||
) (genitalia slide, 1/
|
||
<date id="FFFE0EBA144C4E7EB6E06FB485A1DFF5" box="[400,511,933,960]" pageId="3" pageNumber="4" value="2013-08-06">6.8.2013</date>
|
||
);
|
||
<specimenCount id="9D46E3F3144C4E7EB5666FB48602DFF5" box="[534,604,933,960]" pageId="3" pageNumber="4" type="male">2 ♂♂</specimenCount>
|
||
, Pirin Mts, Suhodolski preval,
|
||
<geoCoordinate id="EE744EBD144C4E7EB4A36FB28035DFF5" box="[979,1131,931,960]" degrees="41" direction="north" minutes="47" orientation="latitude" pageId="3" pageNumber="4" precision="15" seconds="57" value="41.799168">
|
||
41°47′57′′
|
||
<collectionCode id="ED51B0BF144C4E7EB3236FB78035DFF5" box="[1107,1131,934,960]" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13092" lsid="urn:lsid:biocol.org:col:13092" name="Nanjing University" pageId="3" pageNumber="4" type="Herbarium">N</collectionCode>
|
||
</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE744EBD144C4E7EB3096FB28153DFF5" box="[1145,1293,931,960]" degrees="23" direction="east" minutes="21" orientation="longitude" pageId="3" pageNumber="4" precision="15" seconds="34" value="23.359446">
|
||
23°21′34′′
|
||
<collectionCode id="ED51B0BF144C4E7EB3896FB78153DFF5" box="[1273,1293,934,960]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15670" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="3" pageNumber="4" type="Herbarium">E</collectionCode>
|
||
</geoCoordinate>
|
||
,
|
||
<quantity id="4CB8859F144C4E7EB26C6FB48129DFF5" box="[1308,1399,933,960]" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="3" pageNumber="4" unit="m" value="2600.0">
|
||
<date id="FFFE0EBA144C4E7EB26C6FB48109DF8A" box="[1308,1367,933,959]" pageId="3" pageNumber="4" value="2600">2600</date>
|
||
m
|
||
</quantity>
|
||
a.s.l.,
|
||
<date id="FFFE0EBA144C4E7EB6726FD885D9DFD6" box="[258,391,968,995]" pageId="3" pageNumber="4" value="2012-07-09">
|
||
9 Jul.
|
||
<date id="FFFE0EBA144C4E7EB6396FD885D9DFD6" box="[329,391,969,995]" pageId="3" pageNumber="4" value="2012" year="2012">2012</date>
|
||
</date>
|
||
,
|
||
<collectionCode id="ED51B0BF144C4E7EB6E26FD885F7DFD6" box="[402,425,969,995]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15534" lsid="urn:lsid:biocol.org:col:15534" name="Botanischer Garten und Botanisches Museum Berlin-Dahlem, Zentraleinrichtung der Freien Universitaet" pageId="3" pageNumber="4" type="Herbarium">B</collectionCode>
|
||
. Zlatkov and
|
||
<collectionCode id="ED51B0BF144C4E7EB5396FD8863FDFD6" box="[585,609,969,995]" country="Norway" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13093" lsid="urn:lsid:biocol.org:col:13093" name="Botanical Museum - University of Oslo" pageId="3" pageNumber="4" type="Herbarium">O</collectionCode>
|
||
. Sivilov leg. (
|
||
<collectionCode id="ED51B0BF144C4E7EB4756FD88710DFD6" box="[773,846,969,995]" country="Bulgaria" httpUri="http://grbio.org/cool/7wb6-ejfj" name="University of Sofia, Biology Faculty" pageId="3" pageNumber="4">BFUS</collectionCode>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BFF287A144C4E7EB7CD680380BFD87A" blockId="3.[189,1398,1040,1103]" pageId="3" pageNumber="4">
|
||
<collectingCountry id="F35768EA144C4E7EB7CD6803852BD819" box="[189,373,1042,1068]" name="Macedonia" pageId="3" pageNumber="4">MACEDONIA</collectingCountry>
|
||
:
|
||
<specimenCount id="9D46E3F3144C4E7EB6F0680385F0D819" box="[384,430,1042,1068]" pageId="3" pageNumber="4" type="male">
|
||
1
|
||
<specimenCount id="9D46E3F3144C4E7EB6E6680385F0D819" box="[406,430,1042,1068]" pageId="3" pageNumber="4">♂</specimenCount>
|
||
</specimenCount>
|
||
, Mavrovo-Korab, eastern ridge,
|
||
<geoCoordinate id="EE744EBD144C4E7EB45D6801879DD819" box="[813,963,1040,1068]" degrees="41" direction="north" minutes="47" orientation="latitude" pageId="3" pageNumber="4" precision="15" seconds="08" value="41.785557">
|
||
41°47′08′′
|
||
<collectionCode id="ED51B0BF144C4E7EB4DB6803879DD819" box="[939,963,1042,1068]" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13092" lsid="urn:lsid:biocol.org:col:13092" name="Nanjing University" pageId="3" pageNumber="4" type="Herbarium">N</collectionCode>
|
||
</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE744EBD144C4E7EB4BF6801803FD819" box="[975,1121,1040,1068]" degrees="20" direction="east" minutes="34" orientation="longitude" pageId="3" pageNumber="4" precision="15" seconds="46" value="20.579445">
|
||
20°34′46′′
|
||
<collectionCode id="ED51B0BF144C4E7EB33D6803803FD819" box="[1101,1121,1042,1068]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15670" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="3" pageNumber="4" type="Herbarium">E</collectionCode>
|
||
</geoCoordinate>
|
||
, ca
|
||
<quantity id="4CB8859F144C4E7EB3FF6800816FD819" box="[1167,1329,1041,1068]" metricMagnitude="3" metricUnit="m" metricValue="2.3625" metricValueMax="2.4" metricValueMin="2.325" pageId="3" pageNumber="4" unit="m" value="2362.5" valueMax="2400.0" valueMin="2325.0">
|
||
<date id="FFFE0EBA144C4E7EB3FF6800814DD819" box="[1167,1299,1041,1068]" pageId="3" pageNumber="4" value="2325" valueMax="2400">2325–2400</date>
|
||
m
|
||
</quantity>
|
||
a.s.l.,
|
||
<date id="FFFE0EBA144C4E7EB7CD682485F5D87A" box="[189,427,1077,1103]" pageId="3" pageNumber="4" value="2011-07-28" valueMax="2011-08-01" valueMin="2011-07-28">
|
||
<date id="FFFE0EBA144C4E7EB7CD68248536D87A" box="[189,360,1077,1103]" day="28" last_day="01" last_month="08" month="07" monthName="Jul" pageId="3" pageNumber="4">
|
||
<date id="FFFE0EBA144C4E7EB7CD68248577D87A" box="[189,297,1077,1103]" day="01" month="07" pageId="3" pageNumber="4" partOrder="mdy" value="1928-07-01" year="1928">
|
||
<date id="FFFE0EBA144C4E7EB7CD68248553D87A" box="[189,269,1077,1103]" day="28" month="07" monthName="Jul" pageId="3" pageNumber="4" year="1928">28 Jul.</date>
|
||
–
|
||
<date id="FFFE0EBA144C4E7EB66F68248577D87A" box="[287,297,1077,1103]" day="01" month="08" monthName="Aug" pageId="3" pageNumber="4" partOrder="dmy" value="2011-08-01" year="2011">1</date>
|
||
</date>
|
||
Aug.
|
||
</date>
|
||
2011
|
||
</date>
|
||
, Huemer and Tarmann leg. (
|
||
<collectionCode id="ED51B0BF144C4E7EB58868248714D87A" box="[760,842,1077,1103]" country="Austria" httpUri="http://grbio.org/cool/7159-6y1v" name="Tiroler Landesmuseum Ferdinandeum" pageId="3" pageNumber="4">TLMF</collectionCode>
|
||
) (genitalia slide, 1/28.7-
|
||
<date id="FFFE0EBA144C4E7EB3186824808AD87A" box="[1128,1236,1077,1103]" pageId="3" pageNumber="4" value="2011-08-01">1.8.2011</date>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35A7BF1144C4E7BB7CD686C8027DFAA" lastPageId="6" lastPageNumber="7" pageId="3" pageNumber="4" type="description">
|
||
<paragraph id="8BFF287A144C4E7EB7CD686C8535D8A2" blockId="3.[189,363,1149,1175]" box="[189,363,1149,1175]" pageId="3" pageNumber="4">
|
||
<heading id="D0B79F16144C4E7EB7CD686C8535D8A2" bold="true" box="[189,363,1149,1175]" fontSize="11" level="3" pageId="3" pageNumber="4" reason="3">
|
||
<emphasis id="B934F468144C4E7EB7CD686C8535D8A2" bold="true" box="[189,363,1149,1175]" pageId="3" pageNumber="4">Redescription</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BFF287A144C4E7EB79A68A185BBD8FE" blockId="3.[234,485,1200,1227]" box="[234,485,1200,1227]" pageId="3" pageNumber="4">
|
||
<emphasis id="B934F468144C4E7EB79A68A18574D8FF" bold="true" box="[234,298,1200,1226]" pageId="3" pageNumber="4">Male</emphasis>
|
||
(
|
||
<figureCitation id="137B34FF144C4E7EB64A68A18525D8FF" box="[314,379,1200,1226]" captionStart="Fig" captionStartId="4.[189,232,1824,1850]" captionTargetBox="[230,1357,265,1782]" captionTargetId="figure@4.[230,1357,265,1782]" captionTargetPageId="4" captionText="Fig. 1. Adults of Phtheochroa spp. A–D. P. schawerdae (Rebel, 1908) comb. nov. A. ♂, Bulgaria, Pirin Mts. B. ♂, Bulgaria, Rila Mts. C. ♀, Bulgaria, Rila Mts. D. ♂, Republic of Macedonia, Korab Mts. – E–F. P. alpinana sp. nov., France, Alpes Maritimes. E. Holotype, ♂. F. Paratype, ♀. – G. P. apenninana sp. nov., holotype, ♂, Italy, Gran Sasso National Park. – H. P. frigidana (Guenée, 1845) stat. rev., ♂, neotype of Eupoecilia frigidana, Andorra, Pyrenees. – I–J. P. cantabriana sp. nov., Spain, Picos de Europa National Park. I. Holotype, ♂. J. Paratype, ♂. Scale bar = 5 mm, all to scale." figureDoi="http://doi.org/10.5281/zenodo.3838414" httpUri="https://zenodo.org/record/3838414/files/figure.png" pageId="3" pageNumber="4">Fig. 1</figureCitation>
|
||
A–B, D)
|
||
</paragraph>
|
||
<paragraph id="8BFF287A144C4E7EB7CD68F2811ED92A" blockId="3.[189,1398,1250,1312]" pageId="3" pageNumber="4">HEAD. Frons and vertex rust yellow. Labial palps pointed anterad, relatively long, dorsally rust yellow, laterally rust. Scapus and pedicellus dorsally rusty, ventrally creamy, flagellum with yellow scales.</paragraph>
|
||
<paragraph id="8BFF287A144C4E7EB7CD695E80F6D9C3" blockId="3.[189,1399,1358,1531]" pageId="3" pageNumber="4">
|
||
THORAX. Dorsally, including tegulae, rusty yellow, ventrally, mid- and hindlegs creamy, forelegs rusty. Forewing length
|
||
<quantity id="4CB8859F144C4E7EB6FD69638641D9B9" box="[397,543,1394,1420]" metricMagnitude="-3" metricUnit="m" metricValue="8.45" metricValueMax="9.6" metricValueMin="7.3" pageId="3" pageNumber="4" unit="mm" value="8.45" valueMax="9.6" valueMin="7.3">7.3–9.6 mm</quantity>
|
||
(x =8.1, n =11), upperside deep yellow, rarely rust, with more or less pronounced dorsal blotch, cilia concolourous with wings, underside dark grey, with creamy line in costal area. Hindwing upperside dark grey, cilia pale grey, with darker basal part, underside dark grey, with creamy costal area, sometimes with short white longitudinal line near end of vein M
|
||
<subScript id="17C42A3F144C4E7EB3E869FC80FFD9CE" attach="left" box="[1176,1185,1517,1531]" fontSize="6" pageId="3" pageNumber="4">2</subScript>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BFF287A144C4E7EB7CD6A3586E3DA0B" blockId="3.[189,701,1572,1599]" box="[189,701,1572,1599]" pageId="3" pageNumber="4">ABDOMEN. Grey with scattered white scales.</paragraph>
|
||
<paragraph id="8BFF287A144C4E78B7CD6A7C8128DD59" blockId="3.[189,1399,1645,2024]" lastBlockId="5.[189,1399,268,365]" lastPageId="5" lastPageNumber="6" pageId="3" pageNumber="4">
|
||
GENITALIA (
|
||
<figureCitation id="137B34FF144C4E7EB6366A7C85CCDAB2" box="[326,402,1645,1671]" captionStart="Fig" captionStartId="5.[189,232,1813,1839]" captionTargetBox="[230,1355,468,1774]" captionTargetId="figure@5.[189,1398,417,1778]" captionTargetPageId="5" captionText="Fig. 2. Male genitalia (without phalli) of Phtheochroa spp. A–B. P. schawerdae (Rebel, 1908) comb. nov. A. Bulgaria, Rila Mts. B. Republic of Macedonia, Korab Mts. – C. P. alpinana sp. nov., France, Alpes Maritimes, holotype. – D. P. apenninana sp. nov., Italy, Gran Sasso National Park, holotype. – E. P. frigidana (Guenée, 1845) stat. rev., Andorra, Pyrenees, neotype of Eupoecilia frigidana. – F. P. cantabriana sp. nov., Spain, Picos de Europa National Park, holotype. Scale bar = 250 µm, all to scale." figureDoi="http://doi.org/10.5281/zenodo.3838416" httpUri="https://zenodo.org/record/3838416/files/figure.png" pageId="3" pageNumber="4">Fig. 2</figureCitation>
|
||
A–B). Valva with weak dorsal curvature, round apically, with parallel costa and sacculus, both moderately sclerotized. Valva may appear wider or narrower depending on preparation. Uncus relatively well sclerotized, tapered apically. Socii rounded, convergent and connected medially with membrane. Transtilla (
|
||
<figureCitation id="137B34FF144C4E7EB57B6AC6860CDAC4" box="[523,594,1751,1777]" captionStart="Fig" captionStartId="6.[188,231,1807,1833]" captionTargetBox="[191,1389,1348,1764]" captionTargetId="figure@6.[188,1397,1344,1765]" captionTargetPageId="6" captionText="Fig. 3. Transtilla of Phtheochroa spp. A–B. P. schawerdae (Rebel, 1908) comb. nov., Bulgaria, Rila Mts. – C. P. alpinana sp. nov., France, Alpes Maritimes, holotype. – D. P. apenninana sp. nov., Italy, Gran Sasso National Park, holotype. – E. P. frigidana (Guenée, 1845) stat. rev., Andorra, Pyrenees, neotype of Eupoecilia frigidana. – F. P. cantabriana sp. nov., Spain, Picos de Europa National Park, paratype. Scale bar = 250 µm, all to scale." figureDoi="http://doi.org/10.5281/zenodo.3838418" httpUri="https://zenodo.org/record/3838418/files/figure.png" pageId="3" pageNumber="4">Fig. 3</figureCitation>
|
||
A–B) dorsally spinulous, variably shaped in specimens from same population: apically rounded or relatively angled, wider or narrower. Phallus (
|
||
<figureCitation id="137B34FF144C4E7EB3196AEB80EFDB20" box="[1129,1201,1786,1813]" captionStart="Fig" captionStartId="9.[189,232,1856,1882]" captionTargetBox="[199,1390,280,1807]" captionTargetId="figure@9.[189,1398,265,1814]" captionTargetPageId="9" captionText="Fig. 5. Phalli with vesica everted, Phtheochroa schawerdae (Rebel, 1908) comb. nov. A–B. Bulgaria, Rila Mts. –C–D. Republic of Macedonia, Korab Mts.A, C: left.B, D: dorsal.Abbreviations:gs = gonopore sclerotization; ld = left diverticulum; rd = right diverticulum; svd = small ventral diverticulum; vpp = ventral phallic process. Scale bar = 250 µm." figureDoi="http://doi.org/10.5281/zenodo.3838422" httpUri="https://zenodo.org/record/3838422/files/figure.png" pageId="3" pageNumber="4">Fig. 5</figureCitation>
|
||
) slightly curved ventrally, medioventrally with large, slender, sharp-tipped process (
|
||
<figureCitation id="137B34FF144C4E7EB4A06B0C8048DB02" box="[976,1046,1821,1847]" captionStart="Fig" captionStartId="8.[189,232,1824,1850]" captionTargetBox="[189,1398,1117,1787]" captionTargetId="figure@8.[189,1398,1117,1787]" captionTargetPageId="8" captionText="Fig. 4. Ventral phallic process of Phtheochroa spp., ventral view. A–C. P. schawerdae (Rebel, 1908) comb. nov. A–B. Bulgaria, Rila Mts. C. Republic of Macedonia, Korab Mts. – D–E. P. alpinana sp. nov., France, Alpes Maritimes. D. Holotype. E. Paratype. – F–G. P. apenninana sp. nov., Italy, Gran Sasso National Park. F. Paratype. G. Holotype. – H. P. frigidana (Guenée, 1845) stat. rev., Andorra, Pyrenees, neotype of Eupoecilia frigidana. – I–J. P. cantabriana sp. nov., Spain, Picos de Europa National Park. I. Holotype. J. Paratype. Scale bar = 100 µm, all to scale." figureDoi="http://doi.org/10.5281/zenodo.3838420" httpUri="https://zenodo.org/record/3838420/files/figure.png" pageId="3" pageNumber="4">Fig. 4</figureCitation>
|
||
A–C), with apex curved to the right at nearly right angle. Vesica with large main part protruded at right from sclerotized phallus and two large diverticula bearing long, robust cornuti. Small conical, undulating structure (diverticulum) emerges ventrally from main part. Gonopore located dorsally, surrounded by semicylindrical sclerotized plate with minute lentiform tubercles. Main part medially constricted by circular furrow in which gonopore is sunk. Diverticula emerge close to each other from right side, right one straight and directed posterolaterally, left one curved and pointed laterally. In lateral left view, main part of vesica looks ovoid or subspherical. Diverticula may vary in width, but their length and orientation are constant. In dorsal view, axes of right diverticulum and sclerotized phallus form angle of 100–120°. Cornuti aciculate,
|
||
</paragraph>
|
||
<caption id="DF3F78F2144B4E79B7CD6B3180C1DBDF" ID-DOI="http://doi.org/10.5281/zenodo.3838414" ID-Zenodo-Dep="3838414" httpUri="https://zenodo.org/record/3838414/files/figure.png" pageId="4" pageNumber="5" startId="4.[189,232,1824,1850]" targetBox="[230,1357,265,1782]" targetPageId="4">
|
||
<paragraph id="8BFF287A144B4E79B7CD6B3180C1DBDF" blockId="4.[189,1399,1824,2027]" pageId="4" pageNumber="5">
|
||
<emphasis id="B934F468144B4E79B7CD6B318554DB0F" bold="true" box="[189,266,1824,1850]" pageId="4" pageNumber="5">Fig. 1.</emphasis>
|
||
Adults of
|
||
<taxonomicName id="4C4053F9144B4E79B6F56B308645DB0E" authorityName="Stephens" authorityYear="1829" box="[389,539,1825,1851]" class="Insecta" family="Tortricidae" genus="Phtheochroa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B934F468144B4E79B6F56B308645DB0E" box="[389,539,1825,1851]" italics="true" pageId="4" pageNumber="5">Phtheochroa</emphasis>
|
||
</taxonomicName>
|
||
spp.
|
||
<emphasis id="B934F468144B4E79B52A6B3186CDDB0F" bold="true" box="[602,659,1824,1850]" pageId="4" pageNumber="5">A–D</emphasis>
|
||
.
|
||
<taxonomicName id="4C4053F9144B4E79B5D26B3087AFDB0E" authority="(Rebel, 1908)" authorityName="Zlatkov & Huemer" authorityYear="2017" baseAuthorityName="Rebel" baseAuthorityYear="1908" box="[674,1009,1824,1851]" class="Insecta" family="Tortricidae" genus="Conchylis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="schawerdae" status="comb. nov.">
|
||
<emphasis id="B934F468144B4E79B5D26B308717DB0E" box="[674,841,1825,1851]" italics="true" pageId="4" pageNumber="5">P. schawerdae</emphasis>
|
||
(Rebel, 1908)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2074913144B4E79B4896B318025DB0E" box="[1017,1147,1824,1851]" pageId="4" pageNumber="5" rank="species">comb. nov.</taxonomicNameLabel>
|
||
<emphasis id="B934F468144B4E79B3F16B3180C8DB0F" bold="true" box="[1153,1174,1824,1850]" pageId="4" pageNumber="5">A</emphasis>
|
||
. ♂, Bulgaria, Pirin Mts.
|
||
<emphasis id="B934F468144B4E79B78C6B55854EDB6B" bold="true" box="[252,272,1860,1886]" pageId="4" pageNumber="5">B</emphasis>
|
||
. ♂, Bulgaria, Rila Mts.
|
||
<emphasis id="B934F468144B4E79B5486B558613DB6B" bold="true" box="[568,589,1860,1886]" pageId="4" pageNumber="5">C</emphasis>
|
||
. ♀, Bulgaria, Rila Mts.
|
||
<emphasis id="B934F468144B4E79B4066B5587D5DB6B" bold="true" box="[886,907,1860,1886]" pageId="4" pageNumber="5">D</emphasis>
|
||
. ♂, Republic of Macedonia, Korab Mts. –
|
||
<emphasis id="B934F468144B4E79B7A16B76855BDBB4" bold="true" box="[209,261,1895,1921]" pageId="4" pageNumber="5">E–F</emphasis>
|
||
.
|
||
<taxonomicName id="4C4053F9144B4E79B6626B7985CBDBB4" authority="Zlatkov & Huemer, 2017" authorityName="Zlatkov & Huemer" authorityYear="2017" box="[274,405,1895,1921]" class="Insecta" family="Tortricidae" genus="Phtheochroa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="alpinana" status="sp. nov.">
|
||
<emphasis id="B934F468144B4E79B6626B7985CBDBB4" box="[274,405,1895,1921]" italics="true" pageId="4" pageNumber="5">P. alpinana</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2074913144B4E79B6EA6B7685ABDBB4" box="[410,501,1895,1921]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, France, Alpes Maritimes.
|
||
<emphasis id="B934F468144B4E79B4596B768763DBB4" bold="true" box="[809,829,1895,1921]" pageId="4" pageNumber="5">E</emphasis>
|
||
. Holotype, ♂.
|
||
<emphasis id="B934F468144B4E79B4956B7687A9DBB4" bold="true" box="[997,1015,1895,1921]" pageId="4" pageNumber="5">F</emphasis>
|
||
. Paratype, ♀. –
|
||
<emphasis id="B934F468144B4E79B3DE6B76809BDBB4" bold="true" box="[1198,1221,1895,1921]" pageId="4" pageNumber="5">G</emphasis>
|
||
.
|
||
<taxonomicName id="4C4053F9144B4E79B3A16B798129DBB4" authority="Zlatkov & Huemer, 2017" authorityName="Zlatkov & Huemer" authorityYear="2017" box="[1233,1399,1896,1921]" class="Insecta" family="Tortricidae" genus="Phtheochroa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="apenninana" status="sp. nov.">
|
||
<emphasis id="B934F468144B4E79B3A16B798129DBB4" box="[1233,1399,1896,1921]" italics="true" pageId="4" pageNumber="5">P. apenninana</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2074913144B4E79B7CD6B9A8542DB90" box="[189,284,1931,1957]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype, ♂, Italy, Gran Sasso National Park. –
|
||
<emphasis id="B934F468144B4E79B4176B9B8720DB91" bold="true" box="[871,894,1930,1956]" pageId="4" pageNumber="5">H</emphasis>
|
||
.
|
||
<taxonomicName id="4C4053F9144B4E79B4FE6B9A8085DB90" authority="(Guenee, 1845)" authorityName="Zlatkov & Huemer" authorityYear="2017" baseAuthorityName="Guenee" baseAuthorityYear="1845" box="[910,1243,1930,1957]" class="Insecta" family="Tortricidae" genus="Phtheochroa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="frigidana" status="stat. nov.">
|
||
<emphasis id="B934F468144B4E79B4FE6B9A8044DB91" box="[910,1050,1930,1956]" italics="true" pageId="4" pageNumber="5">P. frigidana</emphasis>
|
||
(Guenée, 1845)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2074913144B4E79B3956B9A8112DB90" box="[1253,1356,1931,1957]" pageId="4" pageNumber="5">stat. rev.</taxonomicNameLabel>
|
||
, ♂, neotype of
|
||
<taxonomicName id="4C4053F9144B4E79B6386BBC8660DBF2" authorityName="Guenee, 1845 Zlatkov & Huemer" authorityYear="2017" baseAuthorityName="Guenee" baseAuthorityYear="1845" box="[328,574,1965,1991]" class="Insecta" family="Tortricidae" genus="Eupoecilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="frigidana">
|
||
<emphasis id="B934F468144B4E79B6386BBC8660DBF2" box="[328,574,1965,1991]" italics="true" pageId="4" pageNumber="5">Eupoecilia frigidana</emphasis>
|
||
</taxonomicName>
|
||
, Andorra, Pyrenees. –
|
||
<emphasis id="B934F468144B4E79B4286BBF87DFDBFD" bold="true" box="[856,897,1966,1992]" pageId="4" pageNumber="5">I–J</emphasis>
|
||
.
|
||
<taxonomicName id="4C4053F9144B4E79B4E26BBF801FDBF2" authority="Zlatkov & Huemer, 2017" authorityName="Zlatkov & Huemer" authorityYear="2017" box="[914,1089,1965,1991]" class="Insecta" family="Tortricidae" genus="Phtheochroa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="cantabriana" status="sp. nov.">
|
||
<emphasis id="B934F468144B4E79B4E26BBF801FDBF2" box="[914,1089,1965,1991]" italics="true" pageId="4" pageNumber="5">P. cantabriana</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2074913144B4E79B33C6BBF80F3DBFD" box="[1100,1197,1966,1992]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, Spain, Picos de Europa National Park.
|
||
<emphasis id="B934F468144B4E79B6BA6BC0858BDBDE" bold="true" box="[458,469,2001,2027]" pageId="4" pageNumber="5">I</emphasis>
|
||
. Holotype, ♂.
|
||
<emphasis id="B934F468144B4E79B5F46BC086CDDBDE" bold="true" box="[644,659,2001,2027]" pageId="4" pageNumber="5">J</emphasis>
|
||
. Paratype, ♂. Scale bar = 5 mm, all to scale.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF3F78F2144A4E78B7CD6B048543DBEA" ID-DOI="http://doi.org/10.5281/zenodo.3838416" ID-Zenodo-Dep="3838416" httpUri="https://zenodo.org/record/3838416/files/figure.png" pageId="5" pageNumber="6" startId="5.[189,232,1813,1839]" targetBox="[230,1355,468,1774]" targetPageId="5">
|
||
<paragraph id="8BFF287A144A4E78B7CD6B048543DBEA" blockId="5.[189,1399,1813,2016]" pageId="5" pageNumber="6">
|
||
<emphasis id="B934F468144A4E78B7CD6B048552DB1A" bold="true" box="[189,268,1813,1839]" pageId="5" pageNumber="6">Fig. 2.</emphasis>
|
||
Male genitalia (without phalli) of
|
||
<taxonomicName id="4C4053F9144A4E78B5DD6B04871DDB1A" authorityName="Stephens" authorityYear="1829" box="[685,835,1813,1839]" class="Insecta" family="Tortricidae" genus="Phtheochroa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B934F468144A4E78B5DD6B04871DDB1A" box="[685,835,1813,1839]" italics="true" pageId="5" pageNumber="6">Phtheochroa</emphasis>
|
||
</taxonomicName>
|
||
spp.
|
||
<emphasis id="B934F468144A4E78B4F66B0487E3DB1A" bold="true" box="[902,957,1813,1839]" pageId="5" pageNumber="6">A–B</emphasis>
|
||
.
|
||
<taxonomicName id="4C4053F9144A4E78B4BE6B07817ADB1A" authority="(Rebel, 1908)" authorityName="Zlatkov & Huemer" authorityYear="2017" baseAuthorityName="Rebel" baseAuthorityYear="1908" box="[974,1316,1813,1839]" class="Insecta" family="Tortricidae" genus="Conchylis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="schawerdae" status="comb. nov.">
|
||
<emphasis id="B934F468144A4E78B4BE6B078026DB1A" box="[974,1144,1813,1839]" italics="true" pageId="5" pageNumber="6">P. schawerdae</emphasis>
|
||
(Rebel, 1908)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2074913144A4E78B25E6B0484B1DB66" pageId="5" pageNumber="6" rank="species">comb. nov.</taxonomicNameLabel>
|
||
<emphasis id="B934F468144A4E78B7876B298552DB67" bold="true" box="[247,268,1848,1874]" pageId="5" pageNumber="6">A</emphasis>
|
||
. Bulgaria, Rila Mts.
|
||
<emphasis id="B934F468144A4E78B57A6B288640DB66" bold="true" box="[522,542,1849,1875]" pageId="5" pageNumber="6">B</emphasis>
|
||
. Republic of Macedonia, Korab Mts. –
|
||
<emphasis id="B934F468144A4E78B48E6B29804DDB67" bold="true" box="[1022,1043,1848,1874]" pageId="5" pageNumber="6">C</emphasis>
|
||
.
|
||
<taxonomicName id="4C4053F9144A4E78B3536B2880F4DB67" authority="Zlatkov & Huemer, 2017" authorityName="Zlatkov & Huemer" authorityYear="2017" box="[1059,1194,1848,1874]" class="Insecta" family="Tortricidae" genus="Phtheochroa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="alpinana" status="sp. nov.">
|
||
<emphasis id="B934F468144A4E78B3536B2880F4DB67" box="[1059,1194,1848,1874]" italics="true" pageId="5" pageNumber="6">P. alpinana</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2074913144A4E78B3C36B28814CDB66" box="[1203,1298,1849,1875]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, France, Alpes Maritimes, holotype. –
|
||
<emphasis id="B934F468144A4E78B55A6B4D8661DB43" bold="true" box="[554,575,1884,1910]" pageId="5" pageNumber="6">D</emphasis>
|
||
.
|
||
<taxonomicName id="4C4053F9144A4E78B5216B4D86A3DB40" authority="Zlatkov & Huemer, 2017" authorityName="Zlatkov & Huemer" authorityYear="2017" box="[593,765,1884,1909]" class="Insecta" family="Tortricidae" genus="Phtheochroa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="apenninana" status="sp. nov.">
|
||
<emphasis id="B934F468144A4E78B5216B4D86A3DB40" box="[593,765,1884,1909]" italics="true" pageId="5" pageNumber="6">P. apenninana</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2074913144A4E78B4776B4D8736DB43" box="[775,872,1884,1910]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, Italy, Gran Sasso National Park, holotype. –
|
||
<emphasis id="B934F468144A4E78B7AA6B6E84B0DBAC" bold="true" box="[218,238,1919,1945]" pageId="5" pageNumber="6">E</emphasis>
|
||
.
|
||
<taxonomicName id="4C4053F9144A4E78B6736B91863EDBAF" authority="(Guenee, 1845)" authorityName="Guenee, 1845 Zlatkov & Huemer" authorityYear="2017" baseAuthorityName="Guenee" baseAuthorityYear="1845" box="[259,608,1919,1946]" class="Insecta" family="Tortricidae" genus="Eupoecilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="frigidana" status="stat. nov.">
|
||
<emphasis id="B934F468144A4E78B6736B9185CADBAC" box="[259,404,1919,1945]" italics="true" pageId="5" pageNumber="6">P. frigidana</emphasis>
|
||
(Guenée, 1845)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2074913144A4E78B51E6B6E8684DBAC" box="[622,730,1919,1945]" pageId="5" pageNumber="6">stat. rev.</taxonomicNameLabel>
|
||
, Andorra, Pyrenees, neotype of
|
||
<taxonomicName id="4C4053F9144A4E78B3046B6E8131DBAC" authorityName="Guenee, 1845 Zlatkov & Huemer" authorityYear="2017" baseAuthorityName="Guenee" baseAuthorityYear="1845" box="[1140,1391,1919,1945]" class="Insecta" family="Tortricidae" genus="Eupoecilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="frigidana">
|
||
<emphasis id="B934F468144A4E78B3046B6E8131DBAC" box="[1140,1391,1919,1945]" italics="true" pageId="5" pageNumber="6">Eupoecilia frigidana</emphasis>
|
||
</taxonomicName>
|
||
. –
|
||
<emphasis id="B934F468144A4E78B7A46BB384B8DB89" bold="true" box="[212,230,1954,1980]" pageId="5" pageNumber="6">F</emphasis>
|
||
.
|
||
<taxonomicName id="4C4053F9144A4E78B7866BB285FDDB89" authority="Zlatkov & Huemer, 2017" authorityName="Zlatkov & Huemer" authorityYear="2017" box="[246,419,1954,1980]" class="Insecta" family="Tortricidae" genus="Phtheochroa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="cantabriana" status="sp. nov.">
|
||
<emphasis id="B934F468144A4E78B7866BB285FDDB89" box="[246,419,1954,1980]" italics="true" pageId="5" pageNumber="6">P. cantabriana</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2074913144A4E78B6DC6BB28654DB88" box="[428,522,1955,1981]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, Spain, Picos de Europa National Park, holotype. Scale bar = 250 µm, all to scale.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BFF287A14494E7BB7CD6D1A8697DD7D" blockId="6.[189,1398,267,329]" pageId="6" pageNumber="7">capitate, of equal length, left one slightly bent. Barely discernible acanthae present on left portion of main part of vesica and at end of diverticula.</paragraph>
|
||
<paragraph id="8BFF287A14494E7BB79A6D6985E7DDA7" blockId="6.[234,441,375,402]" box="[234,441,375,402]" pageId="6" pageNumber="7">
|
||
<emphasis id="B934F46814494E7BB79A6D69851BDDA7" bold="true" box="[234,325,376,402]" pageId="6" pageNumber="7">Female</emphasis>
|
||
(
|
||
<figureCitation id="137B34FF14494E7BB6266D6685F0DDA7" box="[342,430,375,402]" captionStart="Fig" captionStartId="4.[189,232,1824,1850]" captionTargetBox="[230,1357,265,1782]" captionTargetId="figure@4.[230,1357,265,1782]" captionTargetPageId="4" captionText="Fig. 1. Adults of Phtheochroa spp. A–D. P. schawerdae (Rebel, 1908) comb. nov. A. ♂, Bulgaria, Pirin Mts. B. ♂, Bulgaria, Rila Mts. C. ♀, Bulgaria, Rila Mts. D. ♂, Republic of Macedonia, Korab Mts. – E–F. P. alpinana sp. nov., France, Alpes Maritimes. E. Holotype, ♂. F. Paratype, ♀. – G. P. apenninana sp. nov., holotype, ♂, Italy, Gran Sasso National Park. – H. P. frigidana (Guenée, 1845) stat. rev., ♂, neotype of Eupoecilia frigidana, Andorra, Pyrenees. – I–J. P. cantabriana sp. nov., Spain, Picos de Europa National Park. I. Holotype, ♂. J. Paratype, ♂. Scale bar = 5 mm, all to scale." figureDoi="http://doi.org/10.5281/zenodo.3838414" httpUri="https://zenodo.org/record/3838414/files/figure.png" pageId="6" pageNumber="7">Fig. 1C</figureCitation>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="8BFF287A14494E7BB7CD6DBB809CDEE8" blockId="6.[189,1399,426,734]" pageId="6" pageNumber="7">
|
||
Generally resembles male, but forewing length
|
||
<quantity id="4CB8859F14494E7BB5806DBB8720DDF1" box="[752,894,426,452]" metricMagnitude="-3" metricUnit="m" metricValue="8.25" metricValueMax="9.2" metricValueMin="7.3" pageId="6" pageNumber="7" unit="mm" value="8.25" valueMax="9.2" valueMin="7.3">7.3–9.2 mm</quantity>
|
||
(n= 2), underside with larger creamy area, hindwing underside creamy or whitish. Female genitalia (
|
||
<figureCitation id="137B34FF14494E7BB4296DDC87EADDD2" box="[857,948,461,487]" captionStart="Fig" captionStartId="16.[189,232,1930,1956]" captionTargetBox="[188,1400,281,1861]" captionTargetId="figure@16.[180,1408,269,1871]" captionTargetPageId="16" captionText="Fig. 8. Female genitalia of Phtheochroa spp. A. P. schawerdae (Rebel, 1908) comb. nov., Bulgaria, Rila Mts. – B. P. alpinana sp. nov., France, Alpes Maritimes, paratype. Arrow: ventral diverticulum of ductus bursae. Scale bar = 250 µm." figureDoi="http://doi.org/10.5281/zenodo.3838428" httpUri="https://zenodo.org/record/3838428/files/figure.png" pageId="6" pageNumber="7">Fig. 8A</figureCitation>
|
||
) with tergum 8 medially membranous, sterigma with two lateral protuberances, antrum nearly rectangular, with round anterior angles. Apophyses anteriores longer than apophyses posteriores. Ductus bursae ventrally membranous, with conical diverticulum of thick cuticle curved to the left and triangular sclerite dorsally. Corpus bursae with large sclerite on ventral side forming several (5–6) folds at the right, then expanding medially and dorsally on left side, reaching dorsal area of ductus bursae. Right side of corpus bursae with extensively folded membranous area. Ductus of accessory bursa emerges medioventrally, folded cuticle and small spines present to the left of its emerging area. Ductus seminalis inserted on ventral side.
|
||
</paragraph>
|
||
<paragraph id="8BFF287A14494E7BB7CD6F1C861FDF12" blockId="6.[189,577,780,807]" box="[189,577,780,807]" pageId="6" pageNumber="7">
|
||
<emphasis id="B934F46814494E7BB7CD6F1C8525DF12" bold="true" box="[189,379,781,807]" pageId="6" pageNumber="7">
|
||
<heading id="D0B79F1614494E7BB7CD6F1C8563DF12" box="[189,317,781,807]" pageId="6" pageNumber="7">Molecular</heading>
|
||
data
|
||
</emphasis>
|
||
(
|
||
<tableCitation id="C6C21DC114494E7BB6FB6F1D85BADF12" box="[395,484,780,807]" captionStart="Table 1" captionStartId="7.[189,258,268,294]" captionTargetPageId="7" captionText="Table 1. Minimal uncorrected COI p-distances between different taxa of the P. frigidana s. lat. species complex." httpUri="http://table.plazi.org/id/DF3F78F214484E7AB7CD6D1D8574DD7D" pageId="6" pageNumber="7" tableUuid="DF3F78F214484E7AB7CD6D1D8574DD7D">Table 1</tableCitation>
|
||
,
|
||
<figureCitation id="137B34FF14494E7BB6826F1D8669DF12" box="[498,567,780,807]" captionStart="Fig" captionStartId="18.[189,232,1930,1956]" captionTargetBox="[190,1397,1281,1884]" captionTargetId="figure@18.[172,1412,1251,1902]" captionTargetPageId="18" captionText="Fig. 9. Maximum likelihood tree (built with MEGA6) of cytochrome c oxidase subunit I (COI) barcode fragments. Values at the nodes are bootstrap support values based on 500 replicates." figureDoi="http://doi.org/10.5281/zenodo.3838430" httpUri="https://zenodo.org/record/3838430/files/figure.png" pageId="6" pageNumber="7">Fig. 9</figureCitation>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="8BFF287A14494E7BB7CD6F2E8027DFAA" blockId="6.[189,1399,831,928]" pageId="6" pageNumber="7">
|
||
BIN URI: BOLD:ABA3262. The maximum intraspecific divergence of the barcode region is considerable with 1.59% (n=4), supporting two geographically separated clusters in the Dinaric Mts. and Rila Mts. The minimum distance to the nearest neighbour,
|
||
<taxonomicName id="4C4053F914494E7BB58B6F9787FDDFAA" authority="Zlatkov & Huemer, 2017" authorityName="Zlatkov & Huemer" authorityYear="2017" box="[763,931,902,927]" class="Insecta" family="Tortricidae" genus="Phtheochroa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="apenninana" status="sp. nov.">
|
||
<emphasis id="B934F46814494E7BB58B6F9787FDDFAA" box="[763,931,902,927]" italics="true" pageId="6" pageNumber="7">P. apenninana</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A207491314494E7BB4DA6F97805ADF95" box="[938,1028,902,928]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is 3.69%.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35A7BF114494E7BB7CD6FDF8601D854" pageId="6" pageNumber="7" type="biology_ecology">
|
||
<paragraph id="8BFF287A14494E7BB7CD6FDF857EDFDD" blockId="6.[189,288,974,1000]" box="[189,288,974,1000]" pageId="6" pageNumber="7">
|
||
<heading id="D0B79F1614494E7BB7CD6FDF857EDFDD" bold="true" box="[189,288,974,1000]" fontSize="11" level="3" pageId="6" pageNumber="7" reason="3">
|
||
<emphasis id="B934F46814494E7BB7CD6FDF857EDFDD" bold="true" box="[189,288,974,1000]" pageId="6" pageNumber="7">Ecology</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BFF287A14494E7BB7CD68118601D854" blockId="6.[189,1398,1024,1121]" pageId="6" pageNumber="7">
|
||
The moths fly in July–August at daytime, in the morning and late afternoon until twilight, in sunny, calm weather. Preimaginal stages and larval host plant are unknown. Habitat: humid subalpine meadows at altitudes from
|
||
<quantity id="4CB8859F14494E7BB6186856867FD854" box="[360,545,1095,1121]" metricMagnitude="3" metricUnit="m" metricValue="2.3" metricValueMax="2.6" metricValueMin="2.0" pageId="6" pageNumber="7" unit="m" value="2300.0" valueMax="2600.0" valueMin="2000.0">
|
||
<date id="FFFE0EBA14494E7BB6186856865DD854" box="[360,515,1095,1121]" pageId="6" pageNumber="7" value="2000" valueMax="2600">2000 to 2600</date>
|
||
m
|
||
</quantity>
|
||
a.s.l.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35A7BF114494E7BB7CD68818691D935" pageId="6" pageNumber="7" type="distribution">
|
||
<paragraph id="8BFF287A14494E7BB7CD6881859AD89F" blockId="6.[189,452,1168,1194]" box="[189,452,1168,1194]" pageId="6" pageNumber="7">
|
||
<heading id="D0B79F1614494E7BB7CD68818508D89F" box="[189,342,1168,1194]" pageId="6" pageNumber="7">
|
||
<emphasis id="B934F46814494E7BB7CD68818508D89F" bold="true" box="[189,342,1168,1194]" pageId="6" pageNumber="7">Distribution</emphasis>
|
||
</heading>
|
||
(
|
||
<figureCitation id="137B34FF14494E7BB616688185E5D89F" box="[358,443,1168,1194]" captionStart="Fig" captionStartId="20.[189,232,1990,2016]" captionTargetBox="[189,1398,1296,1948]" captionTargetId="figure@20.[183,1404,1290,1954]" captionTargetPageId="20" captionText="Fig. 10. Distribution of the Phtheochroa frigidana s. lat. species complex based on examined material." figureDoi="http://doi.org/10.5281/zenodo.3838432" httpUri="https://zenodo.org/record/3838432/files/figure.png" pageId="6" pageNumber="7">Fig. 10</figureCitation>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="8BFF287A14494E7BB7CD68D28691D935" blockId="6.[189,1398,1218,1280]" pageId="6" pageNumber="7">Known from the highest Balkan mountain systems: Dinaric Mts, Rila Mts, Pirin Mts, and most probably Carpathian Mts (material not checked by us).</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |