treatments-xml/data/D9/6C/87/D96C878CFFCEFF81FDF7FB60B233FAAB.xml

<document id="593EBEE1D7CEAD77BFFC01446D781F38" ID-CLB-Dataset="299624" ID-DOI="10.5852/ejt.2024.943.2589" ID-GBIF-Dataset="e774f8bb-1116-4e58-9f9e-ad33196cfaeb" ID-ISSN="2118-9773" ID-Zenodo-Dep="12656822" ID-ZooBank="DAE3F640-EF0C-409A-B93F-FA90223653F7" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="karina" IM.metadata_approvedBy="karina" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="karina" checkinTime="1720095579889" checkinUser="plazi" docAuthor="Lv, Sha-Sha, Long, Jian-Kun, Yang, Lin &amp; Chen, Xiang-Sheng" docDate="2024" docId="D96C878CFFCEFF81FDF7FB60B233FAAB" docLanguage="en" docName="EJT.2024.943.127-143.pdf" docOrigin="European Journal of Taxonomy 943" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2589/11801" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Catonidia uncinata Lv, Long, Yang &amp; Chen, 2024, sp. nov." docType="treatment" docUuid="AED15F3E-EFB4-48FD-B4A0-8F6AFCA6CC0A" docUuidSource="ZooBank" docVersion="8" lastPageNumber="141" masterDocId="2555FFF4FFC5FF8FFF84FFB8B619FF83" masterDocTitle="Four new species of the planthopper genus Catonidia Uhler, 1896 from China (Hemiptera: Fulgoromorpha: Achilidae)" masterLastPageNumber="143" masterPageNumber="127" pageNumber="138" updateTime="1720613427385" updateUser="karina" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="F7B3365754C135BFFCCFCCA66032B964">Four new species of the planthopper genus Catonidia Uhler, 1896 from China (Hemiptera: Fulgoromorpha: Achilidae)</mods:title>
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<mods:namePart id="EE98E88316724753D689AB009A662C74">Lv, Sha-Sha</mods:namePart>
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<mods:affiliation id="0EA586FB216E10F31A51BBA6680EA82E">Institute of Entomology, Guizhou University, Guiyang, Guizhou 550025, P. R. China. &amp; The Provincial Special Key Laboratory for Development and Utilization of Insect Resources, Guizhou University, Guiyang, Guizhou 550025, P. R. China.</mods:affiliation>
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<mods:namePart id="FB17781D5470FABE8C1C28F5FD0E56ED">Long, Jian-Kun</mods:namePart>
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<mods:affiliation id="9A3853F6B98A90C6E47F476D78F1ADB6">Institute of Entomology, Guizhou University, Guiyang, Guizhou 550025, P. R. China. &amp; The Provincial Special Key Laboratory for Development and Utilization of Insect Resources, Guizhou University, Guiyang, Guizhou 550025, P. R. China.</mods:affiliation>
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<mods:namePart id="06060E21840077B1384BD2A9BF174FFC">Yang, Lin</mods:namePart>
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<mods:affiliation id="076A79B452BD1A543532BBEF47E0EF64">Institute of Entomology, Guizhou University, Guiyang, Guizhou 550025, P. R. China. &amp; The Provincial Special Key Laboratory for Development and Utilization of Insect Resources, Guizhou University, Guiyang, Guizhou 550025, P. R. China.</mods:affiliation>
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<mods:namePart id="ED0DC754E79473E18602A38D8DAA79E2">Chen, Xiang-Sheng</mods:namePart>
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<mods:affiliation id="DF76122A2B46024FB5317052CD3AC322">Institute of Entomology, Guizhou University, Guiyang, Guizhou 550025, P. R. China. &amp; The Provincial Special Key Laboratory for Development and Utilization of Insect Resources, Guizhou University, Guiyang, Guizhou 550025, P. R. China.</mods:affiliation>
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<heading id="0A3281F6FFCEFF84FDF7FB60B5D8FB70" box="[627,961,1240,1267]" centered="true" fontSize="11" level="2" pageId="11" pageNumber="138" reason="2">
<taxonomicName id="96C54D19FFCEFF84FDF7FB60B546FB70" authorityName="Lv &amp; Long &amp; Yang &amp; Chen" authorityYear="2024" box="[627,863,1240,1267]" class="Insecta" family="Achilidae" genus="Catonidia" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="138" phylum="Arthropoda" rank="species" species="uncinata" status="sp. nov.">
<emphasis id="63B1EA88FFCEFF84FDF7FB60B546FB70" bold="true" box="[627,863,1240,1267]" italics="true" pageId="11" pageNumber="138">Catonidia uncinata</emphasis>
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<taxonomicNameLabel id="788257F3FFCEFF84FCE3FB61B5D8FB70" box="[871,961,1241,1267]" pageId="11" pageNumber="138" rank="species">sp. nov.</taxonomicNameLabel>
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urn:lsid:zoobank.org:act: 
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<paragraph id="517A369AFFCEFF84FD5CFAA7B542FABA" blockId="11.[349,1240,1240,1337]" box="[728,859,1311,1337]" pageId="11" pageNumber="138">
<figureCitation id="C9FE2A1FFFCEFF84FD5CFAA7B542FABA" box="[728,859,1311,1337]" captionStart-0="Figs 36–46" captionStart-1="Fig" captionStartId-0="12.[189,242,1889,1915]" captionStartId-1="13.[189,232,1998,2024]" captionTargetBox-0="[209,1391,374,1834]" captionTargetBox-1="[192,1388,1015,1936]" captionTargetId-0="figure-19@12.[189,1398,365,1851]" captionTargetId-1="figure-224@13.[189,1398,996,1960]" captionTargetPageId-0="12" captionTargetPageId-1="13" captionText-0="Figs 36–46. Catonidia uncinata sp. nov., ♂, holotype (IEGU). 36. Habitus, dorsal view. 37. Habitus, lateral view. 38. Head and thorax, dorsal view. 39. Frons, ventral view. 40. Forewing. 41. Hindwing. 42. Pygofer, lateral view. 43. Pygofer, ventral view. 44. Left gonostyli, inner surface. 45. Aedeagus, left lateral view. 46. Pygofer, dorsal view. Scale bars = 0.5 mm." captionText-1="Fig. 47. Geographic distributions of species of Catonidia Uhler, 1896." figureDoi-0="http://doi.org/10.5281/zenodo.12656832" figureDoi-1="http://doi.org/10.5281/zenodo.12656834" httpUri-0="https://zenodo.org/record/12656832/files/figure.png" httpUri-1="https://zenodo.org/record/12656834/files/figure.png" pageId="11" pageNumber="138">Figs 36–47</figureCitation>
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<emphasis id="63B1EA88FFCEFF84FF39FAD8B72EFAF9" bold="true" box="[189,311,1376,1402]" pageId="11" pageNumber="138">Diagnosis</emphasis>
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Salient features of new species include: forewings (
<figureCitation id="C9FE2A1FFFCEFF84FC9AFA37B5BCFA2A" box="[798,933,1423,1449]" captionStart="Figs 36–46" captionStartId="12.[189,242,1889,1915]" captionTargetBox="[209,1391,374,1834]" captionTargetId="figure-19@12.[189,1398,365,1851]" captionTargetPageId="12" captionText="Figs 36–46. Catonidia uncinata sp. nov., ♂, holotype (IEGU). 36. Habitus, dorsal view. 37. Habitus, lateral view. 38. Head and thorax, dorsal view. 39. Frons, ventral view. 40. Forewing. 41. Hindwing. 42. Pygofer, lateral view. 43. Pygofer, ventral view. 44. Left gonostyli, inner surface. 45. Aedeagus, left lateral view. 46. Pygofer, dorsal view. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12656832" httpUri="https://zenodo.org/record/12656832/files/figure.png" pageId="11" pageNumber="138">Figs 36–37</figureCitation>
) transverse veins of costal area grayish white, radial and medial areas with 6 clear brown spots; dorsal lobe of phallobase (
<figureCitation id="C9FE2A1FFFCEFF84FB14FA0AB2FFFA4E" box="[1168,1254,1458,1485]" captionStart="Figs 36–46" captionStartId="12.[189,242,1889,1915]" captionTargetBox="[209,1391,374,1834]" captionTargetId="figure-19@12.[189,1398,365,1851]" captionTargetPageId="12" captionText="Figs 36–46. Catonidia uncinata sp. nov., ♂, holotype (IEGU). 36. Habitus, dorsal view. 37. Habitus, lateral view. 38. Head and thorax, dorsal view. 39. Frons, ventral view. 40. Forewing. 41. Hindwing. 42. Pygofer, lateral view. 43. Pygofer, ventral view. 44. Left gonostyli, inner surface. 45. Aedeagus, left lateral view. 46. Pygofer, dorsal view. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12656832" httpUri="https://zenodo.org/record/12656832/files/figure.png" pageId="11" pageNumber="138">Fig. 45</figureCitation>
) long spiny, lateral lobes transected at apex, curled into inverted hooked process along ventral margin at end, ventral lobe short, middle to apex knife shaped, apical margin blunt, lateral sides with a lamellar process at the basal ⅓.
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<subSubSection id="19DF6511FFCEFF84FF39F9E6B2C0F949" pageId="11" pageNumber="138" type="etymology">
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<emphasis id="63B1EA88FFCEFF84FF39F9E6B75CF9FB" bold="true" box="[189,325,1630,1656]" pageId="11" pageNumber="138">Etymology</emphasis>
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The specific epithet is derived from the Latin word ‘ 
<emphasis id="63B1EA88FFCEFF84FCC6F935B5ACF925" box="[834,949,1677,1702]" italics="true" pageId="11" pageNumber="138">uncinatus</emphasis>
’, referring to the lateral lobes of the phallobase being curled into an inverted hooked process along the ventral margin at apex.
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<paragraph id="517A369AFFCEFF84FF39F949B774F889" blockId="11.[189,365,1776,1803]" box="[189,365,1776,1803]" pageId="11" pageNumber="138">
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<emphasis id="63B1EA88FFCEFF84FF39F949B774F889" bold="true" box="[189,365,1776,1803]" pageId="11" pageNumber="138">Type material</emphasis>
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<emphasis id="63B1EA88FFCEFF84FF6EF8A7B744F8BA" bold="true" box="[234,349,1823,1849]" pageId="11" pageNumber="138">
<typeStatus id="8E7E8838FFCEFF84FF6EF8A7B744F8BA" box="[234,349,1823,1849]" pageId="11" pageNumber="138" type="holotype">Holotype</typeStatus>
</emphasis>
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<materialsCitation id="E1AD3CC7FFCEFF84FF39F8FAB496F803" ID-GBIF-Occurrence="4903684304" collectingDate="2012-07-28" collectionCode="IEGU" collectorName="Wei-Cheng Yang" country="China" latitude="22.033333" location="Xishuangbanna National Natural Reserve" longitude="100.9" pageId="11" pageNumber="138" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan Province" typeStatus="holotype">
<collectingCountry id="29D2760AFFCEFF84FF39F8FAB701F8DF" box="[189,280,1858,1884]" name="China" pageId="11" pageNumber="138">CHINA</collectingCountry>
– 
<emphasis id="63B1EA88FFCEFF84FEBFF8FAB40CF8DF" bold="true" box="[315,533,1858,1884]" pageId="11" pageNumber="138">
<collectingRegion id="9301F878FFCEFF84FEBFF8FAB40CF8DF" box="[315,533,1858,1884]" country="China" name="Yunnan" pageId="11" pageNumber="138">Yunnan Province</collectingRegion>
</emphasis>
• 
<specimenCount id="47C3FD13FFCEFF84FDB4F8FBB450F8DE" box="[560,585,1859,1885]" count="1" pageId="11" pageNumber="138" type="male">♂</specimenCount>
; 
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; 
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<geoCoordinate id="34F1505DFFCEFF84FB55F8FAB352F8DE" box="[1233,1355,1857,1885]" degrees="22" direction="north" minutes="02" orientation="latitude" pageId="11" pageNumber="138" precision="925" value="22.033333">100°90′ E</geoCoordinate>
; 
<date id="257B105AFFCEFF84FADDF8FAB735F803" pageId="11" pageNumber="138" value="2012-07-28">
<collectingDate id="353FE9B2FFCEFF84FADDF8FAB735F803" pageId="11" pageNumber="138" value="2012-07-28">28 Jul. 2012</collectingDate>
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; 
<collectorName id="FC30534CFFCEFF84FEB3F8DDB419F803" box="[311,512,1893,1920]" pageId="11" pageNumber="138">Wei-Cheng Yang</collectorName>
leg.; 
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.
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<emphasis id="63B1EA88FFC9FF83FF39F8D9B755F8FF" bold="true" box="[189,332,1889,1916]" pageId="12" pageNumber="139">Figs 36–46.</emphasis>
<taxonomicName id="96C54D19FFC9FF83FEDCF8DAB459F8FF" authorityName="Lv &amp; Long &amp; Yang &amp; Chen" authorityYear="2024" box="[344,576,1890,1916]" class="Insecta" family="Achilidae" genus="Catonidia" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="139" phylum="Arthropoda" rank="species" species="uncinata" status="sp. nov.">
<emphasis id="63B1EA88FFC9FF83FEDCF8DAB459F8FF" box="[344,576,1890,1916]" italics="true" pageId="12" pageNumber="139">Catonidia uncinata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="788257F3FFC9FF83FDC3F8DAB4BFF8FF" box="[583,678,1890,1916]" pageId="12" pageNumber="139" rank="species">sp. nov.</taxonomicNameLabel>
, ♂, holotype (IEGU). 
<emphasis id="63B1EA88FFC9FF83FC30F8DAB5C8F8FF" bold="true" box="[948,977,1890,1916]" pageId="12" pageNumber="139">36</emphasis>
. Habitus, dorsal view. 
<emphasis id="63B1EA88FFC9FF83FB6CF8DAB31DF8FF" bold="true" box="[1256,1284,1890,1916]" pageId="12" pageNumber="139">37</emphasis>
. Habitus, lateral view. 
<emphasis id="63B1EA88FFC9FF83FEDEF83DB76CF81C" bold="true" box="[346,373,1925,1951]" pageId="12" pageNumber="139">38</emphasis>
. Head and thorax, dorsal view. 
<emphasis id="63B1EA88FFC9FF83FD7DF83DB50DF81C" bold="true" box="[761,788,1925,1951]" pageId="12" pageNumber="139">39</emphasis>
. Frons, ventral view. 
<emphasis id="63B1EA88FFC9FF83FB9AF83DB220F81C" bold="true" box="[1054,1081,1925,1951]" pageId="12" pageNumber="139">40</emphasis>
. Forewing. 
<emphasis id="63B1EA88FFC9FF83FB49F83DB2F1F81C" bold="true" box="[1229,1256,1925,1951]" pageId="12" pageNumber="139">41</emphasis>
. Hindwing. 
<emphasis id="63B1EA88FFC9FF83FF3BF810B6C3F841" bold="true" box="[191,218,1960,1986]" pageId="12" pageNumber="139">42</emphasis>
. Pygofer, lateral view. 
<emphasis id="63B1EA88FFC9FF83FE6DF810B41FF841" bold="true" box="[489,518,1960,1986]" pageId="12" pageNumber="139">43</emphasis>
. Pygofer, ventral view. 
<emphasis id="63B1EA88FFC9FF83FC99F810B523F841" bold="true" box="[797,826,1960,1986]" pageId="12" pageNumber="139">44</emphasis>
. Left gonostyli, inner surface. 
<emphasis id="63B1EA88FFC9FF83FB27F810B2D9F841" bold="true" box="[1187,1216,1960,1986]" pageId="12" pageNumber="139">45</emphasis>
. Aedeagus, left lateral view. 
<emphasis id="63B1EA88FFC9FF83FED7F873B769F866" bold="true" box="[339,368,1995,2021]" pageId="12" pageNumber="139">46</emphasis>
. Pygofer, dorsal view. Scale bars = 0.5 mm.
</paragraph>
</caption>
<paragraph id="517A369AFFC8FF82FF6EFEB1B770FEA0" blockId="13.[189,1398,265,361]" box="[234,361,265,291]" pageId="13" pageNumber="140">
<emphasis id="63B1EA88FFC8FF82FF6EFEB1B770FEA0" bold="true" box="[234,361,265,291]" pageId="13" pageNumber="140">
<typeStatus id="8E7E8838FFC8FF82FF6EFEB1B770FEA0" box="[234,361,265,291]" pageId="13" pageNumber="140" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="517A369AFFC8FF82FF39FE94B5BDFEEA" blockId="13.[189,1398,265,361]" pageId="13" pageNumber="140">
<materialsCitation id="E1AD3CC7FFC8FF82FF39FE94B5B9FEEA" ID-GBIF-Occurrence="4903684303" collectingDate="2012-07-28" collectionCode="IEGU" collectorName="Wei-Cheng Yang &amp; Zhi-Min Chang" country="China" latitude="22.033333" location="Xishuangbanna National Natural Reserve" longitude="100.9" pageId="13" pageNumber="140" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Yunnan Province" typeStatus="paratype">
<collectingCountry id="29D2760AFFC8FF82FF39FE94B701FEC5" box="[189,280,300,326]" name="China" pageId="13" pageNumber="140">CHINA</collectingCountry>
– 
<emphasis id="63B1EA88FFC8FF82FEB7FE94B413FEC6" bold="true" box="[307,522,299,326]" pageId="13" pageNumber="140">
<collectingRegion id="9301F878FFC8FF82FEB7FE94B413FEC6" box="[307,522,299,326]" country="China" name="Yunnan" pageId="13" pageNumber="140">Yunnan Province</collectingRegion>
</emphasis>
• 
<specimenCount id="47C3FD13FFC8FF82FDA4FE94B457FEC5" box="[544,590,300,326]" count="1" pageId="13" pageNumber="140" type="male">1 ♂</specimenCount>
, 
<specimenCount id="47C3FD13FFC8FF82FDDDFE94B49CFEC5" box="[601,645,300,326]" count="1" pageId="13" pageNumber="140" type="female">1 ♀</specimenCount>
; 
<location id="541A6041FFC8FF82FD16FE94B263FEC5" LSID="urn:lsid:plazi:treatment:D96C878CFFCEFF81FDF7FB60B233FAAB:541A6041FFC8FF82FD16FE94B263FEC5" box="[658,1146,300,326]" country="China" latitude="22.033333" name="Xishuangbanna National Natural Reserve" pageId="13" pageNumber="140" stateProvince="Yunnan Province">Xishuangbanna National Natural Reserve</location>
; 
<geoCoordinate id="34F1505DFFC8FF82FB01FE92B2ECFEC5" box="[1157,1269,298,326]" degrees="22" direction="north" minutes="02" orientation="latitude" pageId="13" pageNumber="140" precision="925" value="22.033333">22°02′ N,</geoCoordinate>
<geoCoordinate id="34F1505DFFC8FF82FB7FFE92B368FEC5" box="[1275,1393,298,326]" degrees="22" direction="north" minutes="02" orientation="latitude" pageId="13" pageNumber="140" precision="925" value="22.033333">100°90′ E</geoCoordinate>
; 
<date id="257B105AFFC8FF82FF39FEF7B748FEEA" box="[189,337,335,361]" pageId="13" pageNumber="140" value="2012-07-28">
<collectingDate id="353FE9B2FFC8FF82FF39FEF7B748FEEA" box="[189,337,335,361]" pageId="13" pageNumber="140" value="2012-07-28">28 Jul. 2012</collectingDate>
</date>
; 
<collectorName id="FC30534CFFC8FF82FED8FEF7B43DFEEA" box="[348,548,335,361]" pageId="13" pageNumber="140">Wei-Cheng Yang</collectorName>
and 
<collectorName id="FC30534CFFC8FF82FDD9FEF7B50CFEEA" box="[605,789,335,361]" pageId="13" pageNumber="140">Zhi-Min Chang</collectorName>
leg.; 
<collectionCode id="37D4AE5FFFC8FF82FCD2FEF7B5B9FEEA" box="[854,928,335,361]" pageId="13" pageNumber="140">IEGU</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="19DF6511FFC8FF81FF39FE28B56BFC1B" lastPageId="14" lastPageNumber="141" pageId="13" pageNumber="140" type="description">
<paragraph id="517A369AFFC8FF82FF39FE28B756FE29" blockId="13.[189,335,400,426]" box="[189,335,400,426]" pageId="13" pageNumber="140">
<heading id="0A3281F6FFC8FF82FF39FE28B756FE29" bold="true" box="[189,335,400,426]" fontSize="11" level="3" pageId="13" pageNumber="140" reason="3">
<emphasis id="63B1EA88FFC8FF82FF39FE28B756FE29" bold="true" box="[189,335,400,426]" pageId="13" pageNumber="140">Description</emphasis>
</heading>
</paragraph>
<paragraph id="517A369AFFC8FF82FF39FE07B34DFE5A" blockId="13.[189,1364,447,473]" box="[189,1364,447,473]" pageId="13" pageNumber="140">
<smallCapsWord id="579CA046FFC8FF82FF39FE07B777FE54" baselines="466,466" box="[189,366,447,473]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Measurements" pageId="13" pageNumber="140">MEASUREMENTS</smallCapsWord>
. Body length including forewing: male 
<quantity id="963D9B7FFFC8FF82FCC1FE07B5CBFE5A" box="[837,978,447,473]" metricMagnitude="-3" metricUnit="m" metricValue="8.7" metricValueMax="9.3" metricValueMin="8.1" pageId="13" pageNumber="140" unit="mm" value="8.7" valueMax="9.3" valueMin="8.1">8.1–9.3 mm</quantity>
(N = 2), female 
<quantity id="963D9B7FFFC8FF82FB11FE07B2F6FE5A" box="[1173,1263,447,473]" metricMagnitude="-3" metricUnit="m" metricValue="8.7" pageId="13" pageNumber="140" unit="mm" value="8.7">8.7 mm</quantity>
(N = 1).
</paragraph>
<paragraph id="517A369AFFC8FF82FF39FDB8B790FD49" blockId="13.[189,1401,512,714]" pageId="13" pageNumber="140">
<smallCapsWord id="579CA046FFC8FF82FF39FDB8B751FD9A" baselines="531,532" box="[189,328,512,538]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Coloration" pageId="13" pageNumber="140">COLORATION</smallCapsWord>
. General color yellowish brown (
<figureCitation id="C9FE2A1FFFC8FF82FD57FDB8B545FD99" box="[723,860,512,538]" captionStart="Figs 36–46" captionStartId="12.[189,242,1889,1915]" captionTargetBox="[209,1391,374,1834]" captionTargetId="figure-19@12.[189,1398,365,1851]" captionTargetPageId="12" captionText="Figs 36–46. Catonidia uncinata sp. nov., ♂, holotype (IEGU). 36. Habitus, dorsal view. 37. Habitus, lateral view. 38. Head and thorax, dorsal view. 39. Frons, ventral view. 40. Forewing. 41. Hindwing. 42. Pygofer, lateral view. 43. Pygofer, ventral view. 44. Left gonostyli, inner surface. 45. Aedeagus, left lateral view. 46. Pygofer, dorsal view. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12656832" httpUri="https://zenodo.org/record/12656832/files/figure.png" pageId="13" pageNumber="140">Figs 36–37</figureCitation>
). Vertex, base of frons and pronotum fuscous. Eyes reddish brown, ocelli light red. Posterior of pronotum milk-white. Tegula light yellowish brown. Forewings yellowish brown, large number of unclear grayish brown and dark spots distributed along veins, transverse veins of costal area grayish white, radial and medial areas with 6 clear brown spots, veins yellowish brown. Hindwings nearly hyaline, veins pale grayish brown. Legs and abdomen yellowish brown.
</paragraph>
<paragraph id="517A369AFFC8FF81FF39FD4AB5CDFE50" blockId="13.[188,1401,754,921]" lastBlockId="14.[189,1400,265,468]" lastPageId="14" lastPageNumber="141" pageId="13" pageNumber="140">
<smallCapsWord id="579CA046FFC8FF82FF39FD4AB6E5FC89" baselines="773,773" box="[189,252,754,780]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Head" pageId="13" pageNumber="140">HEAD</smallCapsWord>
<smallCapsWord id="579CA046FFC8FF82FE82FD4FB72AFC89" baselines="773" box="[262,307,759,778]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="and" pageId="13" pageNumber="140">AND</smallCapsWord>
<smallCapsWord id="579CA046FFC8FF82FEB8FD4AB781FC89" baselines="773,773" box="[316,408,754,780]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Thorax" pageId="13" pageNumber="140">THORAX</smallCapsWord>
. Head (
<figureCitation id="C9FE2A1FFFC8FF82FE7DFD4AB49BFC8F" box="[505,642,754,780]" captionStart="Figs 36–46" captionStartId="12.[189,242,1889,1915]" captionTargetBox="[209,1391,374,1834]" captionTargetId="figure-19@12.[189,1398,365,1851]" captionTargetPageId="12" captionText="Figs 36–46. Catonidia uncinata sp. nov., ♂, holotype (IEGU). 36. Habitus, dorsal view. 37. Habitus, lateral view. 38. Head and thorax, dorsal view. 39. Frons, ventral view. 40. Forewing. 41. Hindwing. 42. Pygofer, lateral view. 43. Pygofer, ventral view. 44. Left gonostyli, inner surface. 45. Aedeagus, left lateral view. 46. Pygofer, dorsal view. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12656832" httpUri="https://zenodo.org/record/12656832/files/figure.png" pageId="13" pageNumber="140">Figs 36, 38</figureCitation>
) including eyes narrower than pronotum (0.6: 1). Vertex wider than long in middle line (2.7: 1), apex concave, median carina absent, anterior margin angular, posterior margin subangular concave, lateral margins ridged, nearly parallel. Frons (
<figureCitation id="C9FE2A1FFFC8FF82FBC6FC80B282FCD1" box="[1090,1179,824,850]" captionStart="Figs 36–46" captionStartId="12.[189,242,1889,1915]" captionTargetBox="[209,1391,374,1834]" captionTargetId="figure-19@12.[189,1398,365,1851]" captionTargetPageId="12" captionText="Figs 36–46. Catonidia uncinata sp. nov., ♂, holotype (IEGU). 36. Habitus, dorsal view. 37. Habitus, lateral view. 38. Head and thorax, dorsal view. 39. Frons, ventral view. 40. Forewing. 41. Hindwing. 42. Pygofer, lateral view. 43. Pygofer, ventral view. 44. Left gonostyli, inner surface. 45. Aedeagus, left lateral view. 46. Pygofer, dorsal view. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12656832" httpUri="https://zenodo.org/record/12656832/files/figure.png" pageId="13" pageNumber="140">Fig. 39</figureCitation>
) longer in middle line than maximum width (1.5: 1), width at base narrower than width at apex (0.6: 1), median carina distinct, lateral margins concave. Postclypeus (
<figureCitation id="C9FE2A1FFFC8FF82FD63FCC6B527FC1A" box="[743,830,894,921]" captionStart="Figs 36–46" captionStartId="12.[189,242,1889,1915]" captionTargetBox="[209,1391,374,1834]" captionTargetId="figure-19@12.[189,1398,365,1851]" captionTargetPageId="12" captionText="Figs 36–46. Catonidia uncinata sp. nov., ♂, holotype (IEGU). 36. Habitus, dorsal view. 37. Habitus, lateral view. 38. Head and thorax, dorsal view. 39. Frons, ventral view. 40. Forewing. 41. Hindwing. 42. Pygofer, lateral view. 43. Pygofer, ventral view. 44. Left gonostyli, inner surface. 45. Aedeagus, left lateral view. 46. Pygofer, dorsal view. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12656832" httpUri="https://zenodo.org/record/12656832/files/figure.png" pageId="13" pageNumber="140">Fig. 39</figureCitation>
) shorter in middle line than frons (about 0.7: 1), median carina distinct, lateral margins excavate. Pronotum (
<figureCitation id="C9FE2A1FFFCBFF81FC13FEB1B23DFEA0" box="[919,1060,265,291]" captionStart="Figs 36–46" captionStartId="12.[189,242,1889,1915]" captionTargetBox="[209,1391,374,1834]" captionTargetId="figure-19@12.[189,1398,365,1851]" captionTargetPageId="12" captionText="Figs 36–46. Catonidia uncinata sp. nov., ♂, holotype (IEGU). 36. Habitus, dorsal view. 37. Habitus, lateral view. 38. Head and thorax, dorsal view. 39. Frons, ventral view. 40. Forewing. 41. Hindwing. 42. Pygofer, lateral view. 43. Pygofer, ventral view. 44. Left gonostyli, inner surface. 45. Aedeagus, left lateral view. 46. Pygofer, dorsal view. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12656832" httpUri="https://zenodo.org/record/12656832/files/figure.png" pageId="14" pageNumber="141">Figs 36, 38</figureCitation>
) longer in middle line than vertex (2.4: 1), median carina distinct, lateral carinae raised, not extending to posterior margin, confluent with midline at anterior margin to form angular process. Mesonotum (
<figureCitation id="C9FE2A1FFFCBFF81FC7FFEF7B29BFEEA" box="[1019,1154,335,361]" captionStart="Figs 36–46" captionStartId="12.[189,242,1889,1915]" captionTargetBox="[209,1391,374,1834]" captionTargetId="figure-19@12.[189,1398,365,1851]" captionTargetPageId="12" captionText="Figs 36–46. Catonidia uncinata sp. nov., ♂, holotype (IEGU). 36. Habitus, dorsal view. 37. Habitus, lateral view. 38. Head and thorax, dorsal view. 39. Frons, ventral view. 40. Forewing. 41. Hindwing. 42. Pygofer, lateral view. 43. Pygofer, ventral view. 44. Left gonostyli, inner surface. 45. Aedeagus, left lateral view. 46. Pygofer, dorsal view. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12656832" httpUri="https://zenodo.org/record/12656832/files/figure.png" pageId="14" pageNumber="141">Figs 36, 38</figureCitation>
) about 3.7 × as long as pronotum in midline, longer than pronotum and vertex combined (2.4: 1), basal of median carina present clearly, lateral carinae unclear. Forewings (
<figureCitation id="C9FE2A1FFFCBFF81FCA6FE2EB562FE33" box="[802,891,406,432]" captionStart="Figs 36–46" captionStartId="12.[189,242,1889,1915]" captionTargetBox="[209,1391,374,1834]" captionTargetId="figure-19@12.[189,1398,365,1851]" captionTargetPageId="12" captionText="Figs 36–46. Catonidia uncinata sp. nov., ♂, holotype (IEGU). 36. Habitus, dorsal view. 37. Habitus, lateral view. 38. Head and thorax, dorsal view. 39. Frons, ventral view. 40. Forewing. 41. Hindwing. 42. Pygofer, lateral view. 43. Pygofer, ventral view. 44. Left gonostyli, inner surface. 45. Aedeagus, left lateral view. 46. Pygofer, dorsal view. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12656832" httpUri="https://zenodo.org/record/12656832/files/figure.png" pageId="14" pageNumber="141">Fig. 40</figureCitation>
) broadened apically, longer than maximal width (2.1: 1). Hindwings (
<figureCitation id="C9FE2A1FFFCBFF81FE79FE01B44BFE50" box="[509,594,441,467]" captionStart="Figs 36–46" captionStartId="12.[189,242,1889,1915]" captionTargetBox="[209,1391,374,1834]" captionTargetId="figure-19@12.[189,1398,365,1851]" captionTargetPageId="12" captionText="Figs 36–46. Catonidia uncinata sp. nov., ♂, holotype (IEGU). 36. Habitus, dorsal view. 37. Habitus, lateral view. 38. Head and thorax, dorsal view. 39. Frons, ventral view. 40. Forewing. 41. Hindwing. 42. Pygofer, lateral view. 43. Pygofer, ventral view. 44. Left gonostyli, inner surface. 45. Aedeagus, left lateral view. 46. Pygofer, dorsal view. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12656832" httpUri="https://zenodo.org/record/12656832/files/figure.png" pageId="14" pageNumber="141">Fig. 41</figureCitation>
) longer than widest part (1.5: 1).
</paragraph>
<caption id="05BA6612FFC8FF82FF39F876B5E0F86A" ID-DOI="http://doi.org/10.5281/zenodo.12656834" ID-Zenodo-Dep="12656834" box="[189,1017,1998,2025]" httpUri="https://zenodo.org/record/12656834/files/figure.png" pageId="13" pageNumber="140" startId="13.[189,232,1998,2024]" targetBox="[192,1388,1015,1936]" targetPageId="13">
<paragraph id="517A369AFFC8FF82FF39F876B5E0F86A" blockId="13.[189,1017,1998,2025]" box="[189,1017,1998,2025]" pageId="13" pageNumber="140">
<emphasis id="63B1EA88FFC8FF82FF39F876B702F86B" bold="true" box="[189,283,1998,2024]" pageId="13" pageNumber="140">Fig. 47.</emphasis>
Geographic distributions of species of 
<taxonomicName id="96C54D19FFC8FF82FD63F876B5EBF86B" authority="Uhler, 1896" authorityName="Uhler" authorityYear="1896" box="[743,1010,1998,2024]" class="Insecta" family="Achilidae" genus="Catonidia" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="140" phylum="Arthropoda" rank="genus">
<emphasis id="63B1EA88FFC8FF82FD63F876B547F86B" box="[743,862,1998,2024]" italics="true" pageId="13" pageNumber="140">Catonidia</emphasis>
<bibRefCitation id="35544B6BFFC8FF82FCE2F876B5EBF86B" author="Uhler P. R." box="[870,1010,1998,2024]" pageId="13" pageNumber="140" pagination="255 - 297" refId="ref7987" refString="Uhler P. R. 1896. Summary of the Hemiptera of Japan, presented to the United States National Museum by Professor Mitzukuri. Proceedings of the United States National Museum 19 (1108): 255 - 297. https: // doi. org / 10.5479 / si. 00963801.1108.255" type="journal article" year="1896">Uhler, 1896</bibRefCitation>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="517A369AFFCBFF81FF39FE43B56BFC1B" blockId="14.[187,1399,506,920]" pageId="14" pageNumber="141">
<smallCapsWord id="579CA046FFCBFF81FF39FE43B6E6FD90" baselines="526,526" box="[189,255,507,533]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Male" pageId="14" pageNumber="141">MALE</smallCapsWord>
<smallCapsWord id="579CA046FFCBFF81FE8FFDB8B760FD90" baselines="526" box="[267,377,512,531]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="genitalia" pageId="14" pageNumber="141">GENITALIA</smallCapsWord>
. Pygofer (
<figureCitation id="C9FE2A1FFFCBFF81FE7FFE42B4ADFD97" box="[507,692,506,533]" captionStart="Figs 36–46" captionStartId="12.[189,242,1889,1915]" captionTargetBox="[209,1391,374,1834]" captionTargetId="figure-19@12.[189,1398,365,1851]" captionTargetPageId="12" captionText="Figs 36–46. Catonidia uncinata sp. nov., ♂, holotype (IEGU). 36. Habitus, dorsal view. 37. Habitus, lateral view. 38. Head and thorax, dorsal view. 39. Frons, ventral view. 40. Forewing. 41. Hindwing. 42. Pygofer, lateral view. 43. Pygofer, ventral view. 44. Left gonostyli, inner surface. 45. Aedeagus, left lateral view. 46. Pygofer, dorsal view. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12656832" httpUri="https://zenodo.org/record/12656832/files/figure.png" pageId="14" pageNumber="141">Figs 42–43, 46</figureCitation>
) in lateral view with dorsal margin distinctly shorter than ventral margin, ⅓ of posterior margin strongly angulated caudad; in ventral view, apical margin of medioventral process concave medially. Anal tube (
<figureCitation id="C9FE2A1FFFCBFF81FC9FFDF9B5B8FDD8" box="[795,929,577,603]" captionStart="Figs 36–46" captionStartId="12.[189,242,1889,1915]" captionTargetBox="[209,1391,374,1834]" captionTargetId="figure-19@12.[189,1398,365,1851]" captionTargetPageId="12" captionText="Figs 36–46. Catonidia uncinata sp. nov., ♂, holotype (IEGU). 36. Habitus, dorsal view. 37. Habitus, lateral view. 38. Head and thorax, dorsal view. 39. Frons, ventral view. 40. Forewing. 41. Hindwing. 42. Pygofer, lateral view. 43. Pygofer, ventral view. 44. Left gonostyli, inner surface. 45. Aedeagus, left lateral view. 46. Pygofer, dorsal view. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12656832" httpUri="https://zenodo.org/record/12656832/files/figure.png" pageId="14" pageNumber="141">Figs 42, 46</figureCitation>
) longer than widest part by about 1.5: 
<quantity id="963D9B7FFFCBFF81FAECFDF9B6CDFDFD" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="14" pageNumber="141" unit="in" value="1.0">1 in</quantity>
dorsal view, apical margin broadly rounded, basal margin narrow, slightly constricted near base. Aedeagus (
<figureCitation id="C9FE2A1FFFCBFF81FEC0FD3FB787FD21" box="[324,414,647,674]" captionStart="Figs 36–46" captionStartId="12.[189,242,1889,1915]" captionTargetBox="[209,1391,374,1834]" captionTargetId="figure-19@12.[189,1398,365,1851]" captionTargetPageId="12" captionText="Figs 36–46. Catonidia uncinata sp. nov., ♂, holotype (IEGU). 36. Habitus, dorsal view. 37. Habitus, lateral view. 38. Head and thorax, dorsal view. 39. Frons, ventral view. 40. Forewing. 41. Hindwing. 42. Pygofer, lateral view. 43. Pygofer, ventral view. 44. Left gonostyli, inner surface. 45. Aedeagus, left lateral view. 46. Pygofer, dorsal view. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12656832" httpUri="https://zenodo.org/record/12656832/files/figure.png" pageId="14" pageNumber="141">Fig. 45</figureCitation>
) symmetrical, in lateral view, phallobase narrow at base, dorsal lobe long spiny; lateral lobes transected at apex, curled into inverted hooked process along ventral margin at apex, ventral margin of hook process with small process in the middle; ventral lobe shorter, basal part constricted into stalk, middle to end knife shaped, apical margin blunt, lateral sides with lamellar processes at basal ⅓; apical margin of phallobasal conjunctival processes wide and rounded, lateral sides each with long and large lamellar process along dorsal and ventral margin at subapical area. Gonostyli (
<figureCitation id="C9FE2A1FFFCBFF81FB12FC8FB305FCD2" box="[1174,1308,823,849]" captionStart="Figs 36–46" captionStartId="12.[189,242,1889,1915]" captionTargetBox="[209,1391,374,1834]" captionTargetId="figure-19@12.[189,1398,365,1851]" captionTargetPageId="12" captionText="Figs 36–46. Catonidia uncinata sp. nov., ♂, holotype (IEGU). 36. Habitus, dorsal view. 37. Habitus, lateral view. 38. Head and thorax, dorsal view. 39. Frons, ventral view. 40. Forewing. 41. Hindwing. 42. Pygofer, lateral view. 43. Pygofer, ventral view. 44. Left gonostyli, inner surface. 45. Aedeagus, left lateral view. 46. Pygofer, dorsal view. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12656832" httpUri="https://zenodo.org/record/12656832/files/figure.png" pageId="14" pageNumber="141">Figs 42, 44</figureCitation>
) longer than wide, base narrow, apical margin broadly rounded, dorsal margin with stout twisted process near middle, inner side of base with slender, finger-like process.
</paragraph>
</subSubSection>
<subSubSection id="19DF6511FFCBFF81FF39FC07B442FB8B" box="[189,603,959,1032]" pageId="14" pageNumber="141" type="distribution">
<paragraph id="517A369AFFCBFF81FF39FC07B741FC5A" blockId="14.[189,344,959,985]" box="[189,344,959,985]" pageId="14" pageNumber="141">
<heading id="0A3281F6FFCBFF81FF39FC07B741FC5A" bold="true" box="[189,344,959,985]" fontSize="11" level="3" pageId="14" pageNumber="141" reason="3">
<emphasis id="63B1EA88FFCBFF81FF39FC07B741FC5A" bold="true" box="[189,344,959,985]" pageId="14" pageNumber="141">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="517A369AFFCBFF81FF39FC55B442FB8B" blockId="14.[189,603,1005,1032]" box="[189,603,1005,1032]" pageId="14" pageNumber="141">
<collectingCountry id="29D2760AFFCBFF81FF39FC55B71AFB84" box="[189,259,1005,1031]" name="China" pageId="14" pageNumber="141">China</collectingCountry>
(
<collectingRegion id="9301F878FFCBFF81FE96FC56B7C7FB84" box="[274,478,1005,1032]" country="China" name="Yunnan" pageId="14" pageNumber="141">Yunnan province</collectingRegion>
) (
<figureCitation id="C9FE2A1FFFCBFF81FE72FC55B454FB8B" box="[502,589,1005,1032]" captionStart="Fig" captionStartId="13.[189,232,1998,2024]" captionTargetBox="[192,1388,1015,1936]" captionTargetId="figure-224@13.[189,1398,996,1960]" captionTargetPageId="13" captionText="Fig. 47. Geographic distributions of species of Catonidia Uhler, 1896." figureDoi="http://doi.org/10.5281/zenodo.12656834" httpUri="https://zenodo.org/record/12656834/files/figure.png" pageId="14" pageNumber="141">Fig. 47</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="19DF6511FFCBFF81FF39FB97B233FAAB" pageId="14" pageNumber="141" type="discussion">
<paragraph id="517A369AFFCBFF81FF39FB97B729FBCA" blockId="14.[189,304,1071,1097]" box="[189,304,1071,1097]" pageId="14" pageNumber="141">
<heading id="0A3281F6FFCBFF81FF39FB97B729FBCA" bold="true" box="[189,304,1071,1097]" fontSize="11" level="3" pageId="14" pageNumber="141" reason="3">
<emphasis id="63B1EA88FFCBFF81FF39FB97B729FBCA" bold="true" box="[189,304,1071,1097]" pageId="14" pageNumber="141">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="517A369AFFCBFF81FF39FBE5B233FAAB" blockId="14.[189,1401,1117,1320]" pageId="14" pageNumber="141">
This species is similar to 
<taxonomicName id="96C54D19FFCBFF81FE6BFBE5B203FBFB" authority="Wang &amp; Huang, 1990" authorityName="Wang &amp; Huang" authorityYear="1990" box="[495,1050,1117,1144]" class="Insecta" family="Achilidae" genus="Catonidia" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="141" phylum="Arthropoda" rank="species" species="wuyishanana">
<emphasis id="63B1EA88FFCBFF81FE6BFBE5B512FBF4" box="[495,779,1117,1143]" italics="true" pageId="14" pageNumber="141">Catonidia wuyishanana</emphasis>
<bibRefCitation id="35544B6BFFCBFF81FC96FBE6B203FBFB" author="Wang S. Z. &amp; Huang J. &amp; Song J. Z." box="[786,1050,1118,1144]" pageId="14" pageNumber="141" pagination="120 - 123" refId="ref8040" refString="Wang S. Z., Huang J. &amp; Song J. Z. 1990. Notes on the Catonidia from China (Homoptera: Achilidae). Acta Agriculturae Boreali-Sinica 5 (4): 120 - 123." type="journal article" year="1990">Wang &amp; Huang, 1990</bibRefCitation>
</taxonomicName>
, but differs from the latter as follows: (1) radial and medial areas of forewing with 6 clear brown spots (radial and medial areas of forewing without 6 clear brown spots in 
<taxonomicName id="96C54D19FFCBFF81FD2CFB1CB57EFB3D" authorityName="Wang &amp; Huang" authorityYear="1990" box="[680,871,1188,1214]" class="Insecta" family="Achilidae" genus="Catonidia" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="141" phylum="Arthropoda" rank="species" species="wuyishanana">
<emphasis id="63B1EA88FFCBFF81FD2CFB1CB57EFB3D" box="[680,871,1188,1214]" italics="true" pageId="14" pageNumber="141">C. wuyishanana</emphasis>
</taxonomicName>
); (2) lateral lobes transected at apex, curled into an inverted hooked process along the ventral margin at apex (lateral lobes acute at apex, without inverted hooked process along the ventral margin at apex in 
<taxonomicName id="96C54D19FFCBFF81FC20FB53B27FFA87" authorityName="Wang &amp; Huang" authorityYear="1990" box="[932,1126,1258,1285]" class="Insecta" family="Achilidae" genus="Catonidia" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="141" phylum="Arthropoda" rank="species" species="wuyishanana">
<emphasis id="63B1EA88FFCBFF81FC20FB53B27FFA87" box="[932,1126,1258,1285]" italics="true" pageId="14" pageNumber="141">C. wuyishanana</emphasis>
</taxonomicName>
); (3) apical margin of ventral lobe blunt (apical margin of ventral lobe acute in 
<taxonomicName id="96C54D19FFCBFF81FCDEFAB6B200FAA4" authorityName="Wang &amp; Huang" authorityYear="1990" box="[858,1049,1293,1320]" class="Insecta" family="Achilidae" genus="Catonidia" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="141" phylum="Arthropoda" rank="species" species="wuyishanana">
<emphasis id="63B1EA88FFCBFF81FCDEFAB6B200FAA4" box="[858,1049,1293,1320]" italics="true" pageId="14" pageNumber="141">C. wuyishanana</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>