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<document id="134FEEA01099885994DCF3DB808EC26B" ID-CLB-Dataset="303514" ID-DOI="10.50826/bnmnszool.48.2_35" ID-GBIF-Dataset="ae1af4e1-c35d-41db-99cc-8e05c3e3f510" ID-ISSN="2434-091X" ID-Zenodo-Dep="13824279" IM.bibliography_approvedBy="karina" IM.illustrations_approvedBy="karina" IM.materialsCitations_approvedBy="karina" IM.metadata_approvedBy="karina" IM.tables_approvedBy="karina" IM.taxonomicNames_approvedBy="karina" IM.treatments_approvedBy="karina" checkinTime="1718643474207" checkinUser="felipe" docAuthor="Manikandan, K, Megalaa, N, Valliappan, Subramanian, Nandini, K, Rani, Lourdu V, Dakshinamurthi, Senthil &amp; Nagappan, Nagappan" docDate="2022" docId="4B2487854234A523392EA49229CCF92B" docLanguage="en" docName="BullNatlMusNatSciSerA.48.2.35-83.pdf" docOrigin="Bulletin of the National Museum of Nature and Science. Series A, Zoology 48 (2)" docSource="http://dx.doi.org/10.4103/jpbs.jpbs_806_23" docStyle="DocumentStyle:A6FFB80BFE6276686563735C80A89639.2:BullNatlMusNatSciSerA.2021-.journal_article.english.affilindex" docStyleId="A6FFB80BFE6276686563735C80A89639" docStyleName="BullNatlMusNatSciSerA.2021-.journal_article.english.affilindex" docStyleVersion="2" docTitle="Naxioides sahulensis Manikandan &amp; Megalaa &amp; Valliappan &amp; Nandini &amp; Rani &amp; Dakshinamurthi &amp; Nagappan 2022" docType="treatment" docVersion="4" lastPageNumber="52" masterDocId="B71DFFFD423AA5323A05A1332814FFCE" masterDocTitle="Crabs (Crustacea, Decapoda) from the Seas of East and Southeast Asia Collected by the RV Hakuhō Maru (KH- 72 - 1 Cruise) 3. Sahul Shelf" masterLastPageNumber="83" masterPageNumber="35" pageNumber="49" updateTime="1726964216263" updateUser="karina" zenodo-license-document="CC-BY-4.0">
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<mods:title id="7D90707282226059D3D1A9CED0C500D9">Crabs (Crustacea, Decapoda) from the Seas of East and Southeast Asia Collected by the RV Hakuhō Maru (KH- 72 - 1 Cruise) 3. Sahul Shelf</mods:title>
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<mods:namePart id="98A8AE7426C5523063253F858F3C7C8F">Manikandan, K</mods:namePart>
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<mods:namePart id="A4D777C5323F8828AAE61A66195060EF">Megalaa, N</mods:namePart>
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<mods:namePart id="97B1E2FC946CEED50D4C65A2E8B70B92">Valliappan, Subramanian</mods:namePart>
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<mods:namePart id="0432B5E7405CEB74A482E48E02040BBA">Nandini, K</mods:namePart>
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<mods:namePart id="AAAC1B94AA926A079FBDBF47D505B808">Rani, Lourdu V</mods:namePart>
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<mods:namePart id="E9BA27599042006A914BEC62F6282938">Dakshinamurthi, Senthil</mods:namePart>
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<subSubSection id="8B9765184234A53C392EA4922C78FA77" box="[811,1132,1441,1465]" pageId="14" pageNumber="49" type="nomenclature">
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<taxonomicName id="048D4D104234A53C392EA4922C78FA77" ID-CoL="993X7" authority="Manikandan &amp; Megalaa &amp; Valliappan &amp; Nandini &amp; Rani &amp; Dakshinamurthi &amp; Nagappan, 2022" authorityName="Manikandan &amp; Megalaa &amp; Valliappan &amp; Nandini &amp; Rani &amp; Dakshinamurthi &amp; Nagappan" authorityYear="2022" box="[811,1132,1441,1465]" class="Malacostraca" family="Epialtidae" genus="Naxioides" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="49" phylum="Arthropoda" rank="species" species="sahulensis" status="sp. nov.">Naxioides sahulensis sp. nov.</taxonomicName>
</paragraph>
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<paragraph id="C33236934234A53C39ACA4FA2BF9FA11" blockId="14.[707,1236,1409,1799]" box="[937,1005,1481,1503]" pageId="14" pageNumber="49">
(
<figureCitation id="5BB62A164234A53C39B5A4FA2BF2FA11" box="[944,998,1481,1503]" captionStart="Fig" captionStartId="15.[163,196,1566,1588]" captionTargetBox="[140,1235,215,1532]" captionTargetId="figure-63@15.[138,1237,213,1533]" captionTargetPageId="15" captionText="Fig. 7.ɹNaxioides sahulensis sp. nov., ♀ (holotype, NSMT-Cr 30722; CB 10.6mm excluding lateral spines×PCL 13.8mm excluding intestinal spine) from sta. 32. A: Dorsal habitus. BD: Cephalothorx in dorsal (B), ventral (C) and lateral (D) views." figureDoi="http://doi.org/10.5281/zenodo.13824297" httpUri="https://zenodo.org/record/13824297/files/figure.png" pageId="14" pageNumber="49">Fig. 7</figureCitation>
)
</paragraph>
<paragraph id="C33236934234A53C3901A4DE2CBCF9EE" blockId="14.[707,1236,1409,1799]" pageId="14" pageNumber="49">
http://zoobank.org/
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urn:lsid:zoobank.org:act:
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<subSubSection id="8B9765184234A52238D8A75629CDFDCB" lastPageId="16" lastPageNumber="51" pageId="14" pageNumber="49" type="materials_examined">
<paragraph id="C33236934234A52238D8A75629CDFDCB" blockId="14.[707,1236,1409,1799]" lastBlockId="16.[140,670,215,1799]" lastPageId="16" lastPageNumber="51" pageId="14" pageNumber="49">
<emphasis id="F1F9EA814234A53C38D8A7562BABF9B3" box="[733,959,1637,1661]" italics="true" pageId="14" pageNumber="49">Material examined</emphasis>
.
<collectionCode id="A59CAE564234A53C39E0A7562C1EF9B2" box="[997,1034,1637,1660]" country="Italy" name="Collection of Leptospira Strains" pageId="14" pageNumber="49">RV</collectionCode>
<emphasis id="F1F9EA814234A53C3E23A7562CC0F9B2" box="[1062,1236,1637,1661]" italics="true" pageId="14" pageNumber="49">Hakuhō Maru</emphasis>
KH-72-1 cruise, sta. 32, 1 ♀(
<typeStatus id="1C3688314234A53C3E2DA7B42C9FF951" box="[1064,1163,1671,1695]" pageId="14" pageNumber="49" type="holotype">holotype</typeStatus>
), (
<collectionCode id="A59CAE564234A53C3EAAA7B42CC0F951" box="[1199,1236,1671,1695]" country="Australia" name="The CB Rhizobium Collection" pageId="14" pageNumber="49">CB</collectionCode>
<quantity id="04759B764234A53C38C6A7992B0AF90F" box="[707,798,1706,1730]" metricMagnitude="-2" metricUnit="m" metricValue="1.06" pageId="14" pageNumber="49" unit="mm" value="10.6">10.6 mm</quantity>
excluding lateral spines×
<collectionCode id="A59CAE564234A53C3E45A7992C64F90C" box="[1088,1136,1706,1730]" pageId="14" pageNumber="49">PCL</collectionCode>
<quantity id="04759B764234A53C3E7DA7992CC7F90F" box="[1144,1235,1706,1730]" metricMagnitude="-2" metricUnit="m" metricValue="1.38" pageId="14" pageNumber="49" unit="mm" value="13.8">13.8 mm</quantity>
excluding intestinal spine), NSMT-Cr 30722.
<emphasis id="F1F9EA814234A53C38D8A7C32C43F8C9" box="[733,1111,1775,1799]" italics="true" pageId="14" pageNumber="49">Comparative material examined.</emphasis>
<taxonomicName id="048D4D104234A53D3E6CA7DC29DDF90C" authority="(Haswell, 1880)" baseAuthorityName="Haswell" baseAuthorityYear="1880" class="Malacostraca" family="Epialtidae" genus="Naxioides" kingdom="Animalia" lastPageId="15" lastPageNumber="50" order="Decapoda" pageId="14" pageNumber="49" phylum="Arthropoda" rank="species" species="tenuirostris">
<emphasis id="F1F9EA814234A53D3E6CA7DC2912F90C" italics="true" lastPageId="15" lastPageNumber="50" pageId="14" pageNumber="49">Naxioides tenuirostris</emphasis>
(Haswell, 1880)
</taxonomicName>
.
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<collectingRegion id="0149F8714235A53D3BDCA7992A4EF90C" box="[473,602,1706,1730]" country="Australia" name="Queensland" pageId="15" pageNumber="50">Queensland</collectingRegion>
,
<collectingCountry id="BB9A76034235A53D386CA79928D2F92B" name="Australia" pageId="15" pageNumber="50">Australia</collectingCountry>
: N end
<collectorName id="6E7853454235A53D3B11A7FE29A8F92B" box="[276,444,1741,1765]" pageId="15" pageNumber="50">Albany Passage</collectorName>
,
<collectorName id="6E7853454235A53D3BC2A7FE2A2CF92B" box="[455,568,1741,1765]" pageId="15" pageNumber="50">Cape York</collectorName>
, dredged,
<date id="B73310534235A53D3A89A7DC298CF8C9" box="[140,408,1775,1799]" pageId="15" pageNumber="50" value="1928-08" valueMax="1928-09-31" valueMin="1928-08-00">
<collectingDate id="A777E9BB4235A53D3A89A7DC298CF8C9" box="[140,408,1775,1799]" pageId="15" pageNumber="50" value="1928-08" valueMax="1928-09-31" valueMin="1928-08-00">AugustSeptember 1928</collectingDate>
</date>
, 3 ƏƏ (8.5×
<quantity id="04759B764235A53D3830A7DC2A9BF8C9" box="[565,655,1775,1799]" metricMagnitude="-2" metricUnit="m" metricValue="1.18" pageId="15" pageNumber="50" unit="mm" value="11.8">11.8 mm</quantity>
14.7×
<quantity id="04759B764235A53D3909A7992B7FF90C" box="[780,875,1706,1730]" metricMagnitude="-2" metricUnit="m" metricValue="2.05" pageId="15" pageNumber="50" unit="mm" value="20.5">20.5 mm</quantity>
),
<specimenCount id="D58BFD1A4235A53D398DA7992BDBF90C" box="[904,975,1706,1730]" count="3" pageId="15" pageNumber="50" type="generic">3 juv.</specimenCount>
♀♀ (9.0×
<quantity id="04759B764235A53D3E6EA7992CD2F90F" box="[1131,1222,1706,1730]" metricMagnitude="-2" metricUnit="m" metricValue="1.3599999999999999" pageId="15" pageNumber="50" unit="mm" value="13.6">13.6 mm</quantity>
12.0×
<quantity id="04759B764235A53D3909A7FE2B7FF92B" box="[780,875,1741,1765]" metricMagnitude="-2" metricUnit="m" metricValue="1.77" pageId="15" pageNumber="50" unit="mm" value="17.7">17.7 mm</quantity>
), 1 spent ♀ (12.1×
<quantity id="04759B764235A53D3E57A7FE2CA5F92B" box="[1106,1201,1741,1765]" metricMagnitude="-2" metricUnit="m" metricValue="1.8399999999999999" pageId="15" pageNumber="50" unit="mm" value="18.4">18.4 mm</quantity>
), 1 ovig. ♀ (14.1×
<quantity id="04759B764235A53D397DA7DC2BC3F8C9" box="[888,983,1775,1799]" metricMagnitude="-2" metricUnit="m" metricValue="1.96" pageId="15" pageNumber="50" unit="mm" value="19.6">19.6 mm</quantity>
),
<collectionCode id="A59CAE564235A53D39EEA7DC2C02F8C9" box="[1003,1046,1775,1799]" country="Australia" name="Australian Museum" pageId="15" pageNumber="50" type="Museum">AM</collectionCode>
<specimenCode id="932B9EE84235A53D3E25A7DC2C61F8C9" box="[1056,1141,1775,1799]" collectionCode="AM" country="Australia" name="Australian Museum" pageId="15" pageNumber="50" type="Museum">P13491</specimenCode>
</materialsCitation>
;
<materialsCitation id="73E53CCE4235A5223E87A7DD290CFE99" collectingDate="1964-11-10" collectionCode="AM" collectorName="SE Gulf" country="Australia" elevation="29" lastPageId="16" lastPageNumber="51" latitude="-16.76111" longLatPrecision="21" longitude="140.22638" pageId="15" pageNumber="50" specimenCode="P18589" specimenCount="1" stateProvince="Queensland">
&quot;Rama&quot; trawl sta. 1541,
<date id="B7331053422AA5223B45A1E42A02FF21" box="[320,534,215,239]" pageId="16" pageNumber="51" value="1964-11-10">
<collectingDate id="A777E9BB422AA5223B45A1E42A02FF21" box="[320,534,215,239]" pageId="16" pageNumber="51" value="1964-11-10">10 November 1964</collectingDate>
</date>
,
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of Carpentaria,
<geoCoordinate id="A6B95054422AA5223B13A1C92984FEDC" box="[278,400,249,274]" degrees="16" direction="south" minutes="45" orientation="latitude" pageId="16" pageNumber="51" precision="15" seconds="40" value="-16.76111">16°4540″S</geoCoordinate>
,
<geoCoordinate id="A6B95054422AA5223B99A1C92A33FEDF" box="[412,551,249,274]" degrees="140" direction="east" minutes="13" orientation="longitude" pageId="16" pageNumber="51" precision="15" seconds="35" value="140.22638">140°1335″E</geoCoordinate>
,
<quantity id="04759B76422AA5223834A1C92A72FEDF" box="[561,614,250,274]" metricMagnitude="1" metricUnit="m" metricValue="2.9" pageId="16" pageNumber="51" unit="m" value="29.0">
<elevation id="48A0D1A0422AA5223834A1C92A72FEDF" box="[561,614,250,274]" metricMagnitude="1" metricUnit="m" metricValue="2.9" pageId="16" pageNumber="51" unit="m" value="29.0">29 m</elevation>
</quantity>
, 1Ə (13.0×
<quantity id="04759B76422AA5223ADBA02F2929FEFB" box="[222,317,284,309]" metricMagnitude="-2" metricUnit="m" metricValue="1.95" pageId="16" pageNumber="51" unit="mm" value="19.5">19.5 mm</quantity>
),
<specimenCount id="D58BFD1A422AA5223B5EA02E29B6FEFB" box="[347,418,285,309]" count="1" pageId="16" pageNumber="51" type="generic">1 juv.</specimenCount>
$ (14.7 ×
<quantity id="04759B76422AA5223837A02E2A85FEFB" box="[562,657,285,309]" metricMagnitude="-2" metricUnit="m" metricValue="2.17" pageId="16" pageNumber="51" unit="mm" value="21.7">21.7 mm</quantity>
),
<collectionCode id="A59CAE56422AA5223A89A00C28A3FE99" box="[140,183,319,343]" country="Australia" name="Australian Museum" pageId="16" pageNumber="51" type="Museum">AM</collectionCode>
<specimenCode id="932B9EE8422AA5223AC1A00C290CFE99" box="[196,280,319,343]" collectionCode="AM" country="Australia" name="Australian Museum" pageId="16" pageNumber="51" type="Museum">P18589</specimenCode>
</materialsCitation>
;
<materialsCitation id="73E53CCE422AA5223B2CA00C2A27FE53" collectingDate="1979-02-15" collectionCode="AM" collectorName="Cape York" country="Iceland" elevation="18" latitude="-10.916667" location="3 miles E of Turtle Head" longLatPrecision="1296" longitude="142.68333" pageId="16" pageNumber="51" specimenCode="P64814" specimenCount="1">
FNQ 79-49,
<location id="C6526048422AA5223BB8A00C28D3FEB4" LSID="urn:lsid:plazi:treatment:4B2487854234A523392EA49229CCF92B:C6526048422AA5223BB8A00C28D3FEB4" country="Iceland" latitude="-10.916667" longLatPrecision="1296" longitude="142.68333" name="3 miles E of Turtle Head" pageId="16" pageNumber="51">
<quantity id="04759B76422AA5223BB8A00C2A05FE99" box="[445,529,319,343]" metricMagnitude="3" metricUnit="m" metricValue="4.828032" pageId="16" pageNumber="51" unit="mi" value="3.0">3 miles</quantity>
E of Turtle Head
</location>
, Island,
<collectorName id="6E785345422AA5223B30A05129A0FEB4" box="[309,436,354,378]" pageId="16" pageNumber="51">Cape York</collectorName>
,
<geoCoordinate id="A6B95054422AA5223BCCA0512A09FEB4" box="[457,541,353,378]" degrees="10" direction="south" minutes="55" orientation="latitude" pageId="16" pageNumber="51" precision="925" value="-10.916667">10°55S</geoCoordinate>
,
<geoCoordinate id="A6B95054422AA5223831A0512A8DFEB7" box="[564,665,353,378]" degrees="142" direction="east" minutes="41" orientation="longitude" pageId="16" pageNumber="51" precision="925" value="142.68333">142°41E</geoCoordinate>
,
<quantity id="04759B76422AA5223A89A0B628D5FE52" box="[140,193,389,413]" metricMagnitude="1" metricUnit="m" metricValue="1.8" pageId="16" pageNumber="51" unit="m" value="18.0">
<elevation id="48A0D1A0422AA5223A89A0B628D5FE52" box="[140,193,389,413]" metricMagnitude="1" metricUnit="m" metricValue="1.8" pageId="16" pageNumber="51" unit="m" value="18.0">18 m</elevation>
</quantity>
,
<date id="B7331053422AA5223AD4A0B7298FFE53" box="[209,411,388,413]" pageId="16" pageNumber="51" value="1979-02-15">
<collectingDate id="A777E9BB422AA5223AD4A0B7298FFE53" box="[209,411,388,413]" pageId="16" pageNumber="51" value="1979-02-15">15 February 1979</collectingDate>
</date>
,
<collectionCode id="A59CAE56422AA5223BADA0B629C7FE53" box="[424,467,389,413]" country="Australia" name="Australian Museum" pageId="16" pageNumber="51" type="Museum">AM</collectionCode>
<specimenCode id="932B9EE8422AA5223BDAA0B62A27FE53" box="[479,563,389,413]" collectionCode="AM" country="Australia" name="Australian Museum" pageId="16" pageNumber="51" type="Museum">P64814</specimenCode>
</materialsCitation>
,
<materialsCitation id="73E53CCE422AA5223847A0B629C1FDCB" collectingDate="1979-02-15" collectionCode="AM" collectorName="Cape York" country="Iceland" elevation="20" latitude="-10.65" longLatPrecision="1297" longitude="142.5" pageId="16" pageNumber="51" specimenCode="P64816" specimenCount="1">
<specimenCount id="D58BFD1A422AA5223847A0B62A97FE53" box="[578,643,389,413]" count="1" pageId="16" pageNumber="51" type="generic">1 juv.</specimenCount>
$ (5.9×
<quantity id="04759B76422AA5223AD4A0942937FE71" box="[209,291,423,447]" metricMagnitude="-3" metricUnit="m" metricValue="8.9" pageId="16" pageNumber="51" unit="mm" value="8.9">8.9 mm</quantity>
); FNQ 79-50,
<collectorName id="6E785345422AA5223BC7A0942A23FE71" box="[450,567,423,447]" pageId="16" pageNumber="51">Cape York</collectorName>
,
<geoCoordinate id="A6B95054422AA5223846A0942A83FE71" box="[579,663,422,447]" degrees="10" direction="south" minutes="39" orientation="latitude" pageId="16" pageNumber="51" precision="925" value="-10.65">10°39S</geoCoordinate>
,
<geoCoordinate id="A6B95054422AA5223A89A0F928E5FE2F" box="[140,241,457,482]" degrees="142" direction="east" minutes="30" orientation="longitude" pageId="16" pageNumber="51" precision="925" value="142.5">142°30E</geoCoordinate>
,
<quantity id="04759B76422AA5223AF9A0F92925FE2F" box="[252,305,458,482]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="16" pageNumber="51" unit="m" value="20.0">
<elevation id="48A0D1A0422AA5223AF9A0F92925FE2F" box="[252,305,458,482]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="16" pageNumber="51" unit="m" value="20.0">20 m</elevation>
</quantity>
,
<date id="B7331053422AA5223B38A0F929EAFE2C" box="[317,510,458,482]" pageId="16" pageNumber="51" value="1979-02-15">
<collectingDate id="A777E9BB422AA5223B38A0F929EAFE2C" box="[317,510,458,482]" pageId="16" pageNumber="51" value="1979-02-15">15 February 1979</collectingDate>
</date>
, carapace only (10.3×
<quantity id="04759B76422AA5223ADBA0DF2929FDCB" box="[222,317,492,517]" metricMagnitude="-2" metricUnit="m" metricValue="1.52" pageId="16" pageNumber="51" unit="mm" value="15.2">15.2 mm</quantity>
),
<collectionCode id="A59CAE56422AA5223B4BA0DE296DFDCB" box="[334,377,493,517]" country="Australia" name="Australian Museum" pageId="16" pageNumber="51" type="Museum">AM</collectionCode>
<specimenCode id="932B9EE8422AA5223B85A0DE29C1FDCB" box="[384,469,493,517]" collectionCode="AM" country="Australia" name="Australian Museum" pageId="16" pageNumber="51" type="Museum">P64816</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="97F2661B4235A53D3AA6A72D29ADF9A1" ID-DOI="http://doi.org/10.5281/zenodo.13824297" ID-Zenodo-Dep="13824297" httpUri="https://zenodo.org/record/13824297/files/figure.png" pageId="15" pageNumber="50" startId="15.[163,196,1566,1588]" targetBox="[140,1235,215,1532]" targetPageId="15" targetType="figure">
<paragraph id="C33236934235A53D3AA6A72D29ADF9A1" blockId="15.[163,1213,1565,1647]" pageId="15" pageNumber="50">
Fig. 7.ɹ
<taxonomicName id="048D4D104235A53D3AF1A72D29A7F9FD" authority="Manikandan &amp; Megalaa &amp; Valliappan &amp; Nandini &amp; Rani &amp; Dakshinamurthi &amp; Nagappan, 2022" authorityName="Manikandan &amp; Megalaa &amp; Valliappan &amp; Nandini &amp; Rani &amp; Dakshinamurthi &amp; Nagappan" authorityYear="2022" box="[244,435,1566,1587]" class="Malacostraca" family="Epialtidae" genus="Naxioides" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="50" phylum="Arthropoda" rank="species" species="sahulensis" status="sp. nov.">
<emphasis id="F1F9EA814235A53D3AF1A72D29A7F9FD" box="[244,435,1566,1587]" italics="true" pageId="15" pageNumber="50">Naxioides sahulensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EACA57FA4235A53D3BBCA72C2A16F9FA" box="[441,514,1567,1588]" pageId="15" pageNumber="50" rank="species">sp. nov.</taxonomicNameLabel>
, ♀ (holotype, NSMT-Cr 30722; CB 10.6mm excluding lateral spines×PCL 13.8mm excluding intestinal spine) from sta. 32. A: Dorsal habitus. BD: Cephalothorx in dorsal (B), ventral (C) and lateral (D) views.
</paragraph>
</caption>
<subSubSection id="8B976518422AA5233AA2A32328DAFF21" lastPageId="17" lastPageNumber="52" pageId="16" pageNumber="51" type="description">
<paragraph id="C3323693422AA5223AA2A3232A8AFA3C" blockId="16.[140,670,215,1799]" pageId="16" pageNumber="51">
<emphasis id="F1F9EA81422AA5223AA2A32329B7FDE9" box="[167,419,527,551]" italics="true" pageId="16" pageNumber="51">
Description of
<typeStatus id="1C368831422AA5223B42A33C29B7FDE9" box="[327,419,527,551]" pageId="16" pageNumber="51" type="holotype">holotype</typeStatus>
</emphasis>
. Carapace (
<figureCitation id="5BB62A16422AA5223827A33C2A81FDE9" box="[546,661,527,551]" captionStart="Fig" captionStartId="15.[163,196,1566,1588]" captionTargetBox="[140,1235,215,1532]" captionTargetId="figure-63@15.[138,1237,213,1533]" captionTargetPageId="15" captionText="Fig. 7.ɹNaxioides sahulensis sp. nov., ♀ (holotype, NSMT-Cr 30722; CB 10.6mm excluding lateral spines×PCL 13.8mm excluding intestinal spine) from sta. 32. A: Dorsal habitus. BD: Cephalothorx in dorsal (B), ventral (C) and lateral (D) views." figureDoi="http://doi.org/10.5281/zenodo.13824297" httpUri="https://zenodo.org/record/13824297/files/figure.png" pageId="16" pageNumber="51">Fig. 7AB</figureCitation>
) pyriform, surface closely tomentose; gastric, cardiac, branchial, intestinal regions well-demarcated, moderately elevated. Gastric region low, bearing 4 long spines, 11 tubercles; anterior end with 2 acute tubercles just behind basis of pseudorostrum; anterior slope medially with slender spine, low spine on both sides; protogastric spines short, directed anterolaterally; mesogastric spine longest, upright, slightly curving anteriorly, with low acute tubercle on both sides, broad U-shaped row of 3 distinct tubercles anteriorly. Cardiac region conical, dorsally elongated into long slender spine, as high as mesogastric spine. Hepatic region inflated, dorso-subproximal surface bearing minute tubercles, lateral surface bearing 2 spines, upper spine longer than lower spine, weakly curved dorsally. Branchial region not markedly inflated, longitudinally with 2 long, slender mesobranchial spines of similar size, small spine anterior to anterior spine, anteromesially with large protuberance mounted with small tubercles; lateral subsurface with 3 short spines of similar size; metabranchial region with small medial spine just anterior to posterolateral carapace margin. Intestinal region elongated posteriorly into large, conical projection, length one-third of
<collectionCode id="A59CAE56422AA5223AEDA4842908FA01" box="[232,284,1463,1487]" pageId="16" pageNumber="51">PCL</collectionCode>
. Posterior carapace margin moderately convex, unarmed, not visible in dorsal view.
</paragraph>
<paragraph id="C3323693422AA5223AA2A4CE2BF0FD1B" blockId="16.[140,670,215,1799]" lastBlockId="16.[707,1236,215,1799]" pageId="16" pageNumber="51">
Pseudorostrum (
<figureCitation id="5BB62A16422AA5223B58A4CE29C3F9DB" box="[349,471,1533,1557]" captionStart="Fig" captionStartId="15.[163,196,1566,1588]" captionTargetBox="[140,1235,215,1532]" captionTargetId="figure-63@15.[138,1237,213,1533]" captionTargetPageId="15" captionText="Fig. 7.ɹNaxioides sahulensis sp. nov., ♀ (holotype, NSMT-Cr 30722; CB 10.6mm excluding lateral spines×PCL 13.8mm excluding intestinal spine) from sta. 32. A: Dorsal habitus. BD: Cephalothorx in dorsal (B), ventral (C) and lateral (D) views." figureDoi="http://doi.org/10.5281/zenodo.13824297" httpUri="https://zenodo.org/record/13824297/files/figure.png" pageId="16" pageNumber="51">Fig. 7AC</figureCitation>
) distinctly divergent (ca. 30°), straight in lateral view. Pseudorostral spine slender, length 0.3 of
<collectionCode id="A59CAE56422AA5223BEFA7712A0AF994" box="[490,542,1602,1626]" pageId="16" pageNumber="51">PCL</collectionCode>
, dorsal surface weakly ridged toward frontal region, bearing short, subdistal spine dorsomesially, tips tapered laterally. Supraorbital eave (
<figureCitation id="5BB62A16422AA522381BA7992A81F90C" box="[542,661,1706,1730]" captionStart="Fig" captionStartId="15.[163,196,1566,1588]" captionTargetBox="[140,1235,215,1532]" captionTargetId="figure-63@15.[138,1237,213,1533]" captionTargetPageId="15" captionText="Fig. 7.ɹNaxioides sahulensis sp. nov., ♀ (holotype, NSMT-Cr 30722; CB 10.6mm excluding lateral spines×PCL 13.8mm excluding intestinal spine) from sta. 32. A: Dorsal habitus. BD: Cephalothorx in dorsal (B), ventral (C) and lateral (D) views." figureDoi="http://doi.org/10.5281/zenodo.13824297" httpUri="https://zenodo.org/record/13824297/files/figure.png" pageId="16" pageNumber="51">Fig. 7AB</figureCitation>
) strongly expanded laterally; anterior angle produced into long, sharp preorbital spine projecting anterodorsally, almost parallel to anterior gastric spine, gently incurved, with pointed apex; antorbital angle produced into small, acute triangular tooth, directed laterally. Dorsal orbital hiatus rounded. Postocular cup apically bifid; upper postorbital margin generally concave (in dorsal view), proximally with distinct finger-shaped intercalated lobe; lateral wall consisting of 2 triangular lobes divergent anteriorly, upper lobe medially ridged, with tooth; lower postorbital margin sinuous, distally with broad, rounded lobe; distinct tubercle right behind ventral orbital hiatus. Pterygostomian region moderately inflated, armed with 3 strong spines directed ventrally, second spine largest.
</paragraph>
<paragraph id="C3323693422AA52238D8A3EC2B09FBC2" blockId="16.[707,1236,215,1799]" pageId="16" pageNumber="51">
Basal antennal article (
<figureCitation id="5BB62A16422AA52239DCA3EC2C39FD39" box="[985,1069,735,759]" captionStart="Fig" captionStartId="15.[163,196,1566,1588]" captionTargetBox="[140,1235,215,1532]" captionTargetId="figure-63@15.[138,1237,213,1533]" captionTargetPageId="15" captionText="Fig. 7.ɹNaxioides sahulensis sp. nov., ♀ (holotype, NSMT-Cr 30722; CB 10.6mm excluding lateral spines×PCL 13.8mm excluding intestinal spine) from sta. 32. A: Dorsal habitus. BD: Cephalothorx in dorsal (B), ventral (C) and lateral (D) views." figureDoi="http://doi.org/10.5281/zenodo.13824297" httpUri="https://zenodo.org/record/13824297/files/figure.png" pageId="16" pageNumber="51">Fig. 7C</figureCitation>
) narrow, distolateral angle produced anterolaterally into elongated sharp spine, visible in dorsal view; lateral margin generally straight; strong triangular tooth near mid-length, directed anterolaterally, as long as distolateral spine. Antennal articles slender, penultimate article two-thirds of ultimate article in length; flagellum reaching tip of pseudorostrum.
</paragraph>
<paragraph id="C3323693422AA52238D8A5242BCBFB31" blockId="16.[707,1236,215,1799]" pageId="16" pageNumber="51">Thoracic sternites 1, 2 unarmed, sternite 3 with small spine mesially, sternite 4 occupied with largest spine, sternite 5 with second largest spine mesially, small spine laterally, sternites 6 with 2 tubercles, mesial one larger than lateral one, sternite 7 with low tubercle. Sterno-pleonal cavity not sharply defined.</paragraph>
<paragraph id="C3323693422AA52238D8A4392B9CFA01" blockId="16.[707,1236,215,1799]" pageId="16" pageNumber="51">
Pleon (
<figureCitation id="5BB62A16422AA522392FA4392B6AFAEC" box="[810,894,1290,1314]" captionStart="Fig" captionStartId="15.[163,196,1566,1588]" captionTargetBox="[140,1235,215,1532]" captionTargetId="figure-63@15.[138,1237,213,1533]" captionTargetPageId="15" captionText="Fig. 7.ɹNaxioides sahulensis sp. nov., ♀ (holotype, NSMT-Cr 30722; CB 10.6mm excluding lateral spines×PCL 13.8mm excluding intestinal spine) from sta. 32. A: Dorsal habitus. BD: Cephalothorx in dorsal (B), ventral (C) and lateral (D) views." figureDoi="http://doi.org/10.5281/zenodo.13824297" httpUri="https://zenodo.org/record/13824297/files/figure.png" pageId="16" pageNumber="51">Fig. 7C</figureCitation>
) somites demarcated, but functionally fused in somites 36, telson; somites 16 each armed with large medial spine; somites 36 narrowly ridged medially, somites 15 laterally armed with 15 spines, somite 5 with hard microscopic setae.
</paragraph>
<paragraph id="C3323693422AA52338D8A4E928DAFF21" blockId="16.[707,1236,215,1799]" lastBlockId="17.[140,670,215,1765]" lastPageId="17" lastPageNumber="52" pageId="16" pageNumber="51">
Ambulatory legs (P25) (
<figureCitation id="5BB62A16422AA5223E09A4E92C7DFA3C" box="[1036,1129,1498,1522]" captionStart="Fig" captionStartId="15.[163,196,1566,1588]" captionTargetBox="[140,1235,215,1532]" captionTargetId="figure-63@15.[138,1237,213,1533]" captionTargetPageId="15" captionText="Fig. 7.ɹNaxioides sahulensis sp. nov., ♀ (holotype, NSMT-Cr 30722; CB 10.6mm excluding lateral spines×PCL 13.8mm excluding intestinal spine) from sta. 32. A: Dorsal habitus. BD: Cephalothorx in dorsal (B), ventral (C) and lateral (D) views." figureDoi="http://doi.org/10.5281/zenodo.13824297" httpUri="https://zenodo.org/record/13824297/files/figure.png" pageId="16" pageNumber="51">Fig. 7A</figureCitation>
) slender, much longer than
<collectionCode id="A59CAE56422AA522398EA4CE2BABF9DB" box="[907,959,1533,1557]" pageId="16" pageNumber="51">PCL</collectionCode>
, length decreasing posteriorly. Merus cylindrical, extensor surface terminated with strong spine projecting laterally. Carpus slightly compressed laterally, extensor surface depressed. Propodus cylindrical, 1.2, 1.0, and 0.8 as long as merus in P2, P3, P45, respectively. Dactylus slightly curved distally, 1.1, 0.9. 0.8 as long as propodus in P2, P3, P45, respectively.
</paragraph>
</subSubSection>
<subSubSection id="8B976518422BA5233AA2A1C92A4FFEDC" box="[167,603,250,274]" pageId="17" pageNumber="52" type="etymology">
<paragraph id="C3323693422BA5233AA2A1C92A4FFEDC" blockId="17.[140,670,215,1765]" box="[167,603,250,274]" pageId="17" pageNumber="52">
<emphasis id="F1F9EA81422BA5233AA2A1C9290DFEDC" box="[167,281,250,274]" italics="true" pageId="17" pageNumber="52">Etymology</emphasis>
. Named after the Sahul Shelf.
</paragraph>
</subSubSection>
<subSubSection id="8B976518422BA5233AA2A02E2919F9B2" pageId="17" pageNumber="52" type="discussion">
<paragraph id="C3323693422BA5233AA2A02E2919F9B2" blockId="17.[140,670,215,1765]" pageId="17" pageNumber="52">
<emphasis id="F1F9EA81422BA5233AA2A02E291FFEFB" box="[167,267,285,309]" italics="true" pageId="17" pageNumber="52">Remarks.</emphasis>
Among nine congeneric species of the genus
<taxonomicName id="048D4D10422BA5233B05A00C297FFE99" authorityName="A. Milne-Edwards" authorityYear="1865" box="[256,363,319,343]" class="Malacostraca" family="Epialtidae" genus="Naxioides" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="F1F9EA81422BA5233B05A00C297FFE99" box="[256,363,319,343]" italics="true" pageId="17" pageNumber="52">Naxioides</emphasis>
</taxonomicName>
, the new species described herein,
<taxonomicName id="048D4D10422BA5233AD8A0502978FEB4" authorityName="Manikandan &amp; Megalaa &amp; Valliappan &amp; Nandini &amp; Rani &amp; Dakshinamurthi &amp; Nagappan" authorityYear="2022" box="[221,364,354,378]" class="Malacostraca" family="Epialtidae" genus="Naxioides" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="52" phylum="Arthropoda" rank="species" species="sahulensis">
<emphasis id="F1F9EA81422BA5233AD8A0502978FEB4" box="[221,364,354,378]" italics="true" pageId="17" pageNumber="52">N. sahulensis</emphasis>
</taxonomicName>
, is morphologicially closest to
<taxonomicName id="048D4D10422BA5233AAFA0B629E9FE53" authority="(Haswell, 1880)" baseAuthorityName="Haswell" baseAuthorityYear="1880" box="[170,509,389,413]" class="Malacostraca" family="Epialtidae" genus="Naxioides" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="52" phylum="Arthropoda" rank="species" species="tenuirostris">
<emphasis id="F1F9EA81422BA5233AAFA0B62951FE52" box="[170,325,389,412]" italics="true" pageId="17" pageNumber="52">N. tenuirostris</emphasis>
(Haswell, 1880)
</taxonomicName>
(type locality: Darnley Island, Torres Strait), with which
<taxonomicName id="048D4D10422BA5233858A09B2984FE2C" authority="Ortmann, 1894" authorityName="Ortmann" authorityYear="1894" class="Malacostraca" family="Majidae" genus="Naxia" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="52" phylum="Arthropoda" rank="species" species="cerastes">
<emphasis id="F1F9EA81422BA5233858A09B28F7FE2C" italics="true" pageId="17" pageNumber="52">Naxia cerastes</emphasis>
<bibRefCitation id="A71C4B62422BA5233AE9A0F92984FE2C" author="Ortmann, A." box="[236,400,458,482]" pageId="17" pageNumber="52" pagination="1 - 80" refId="ref29947" refString="Ortmann, A. 1894. Crustaceen. In: Semon, R., Zoologische Forshungsreisen in Australien und dem Malayischen Archipel 5: 1 - 80, pls. 1 - 3." type="journal article" year="1894">Ortmann, 1894</bibRefCitation>
</taxonomicName>
(type locality: Thursday Island, Torres Strait) was synonymized by
<bibRefCitation id="A71C4B62422BA5233853A0DE2979FDE9" author="Griffin, D. J. G. &amp; H. A. Tranter" pageId="17" pageNumber="52" pagination="1 - 335" refId="ref26736" refString="Griffin, D. J. G. and H. A. Tranter 1986. The Decapoda Brachyura of the Siboga Expedition. Part VIII: Maji- dae. Siboga-Expeditie, Monographie 39 (C 4): 1 - 335, pls. 1 - 22." type="journal article" year="1986">Griffin and Tranter (1986)</bibRefCitation>
, known from western to northeastern
<collectingCountry id="BB9A7603422BA5233B13A301296CFD84" box="[278,376,562,586]" name="Australia" pageId="17" pageNumber="52">Australia</collectingCountry>
including the Arafura Sea. Although the
<typeStatus id="1C368831422BA5233B24A366296AFDA3" box="[289,382,597,621]" pageId="17" pageNumber="52" type="holotype">holotype</typeStatus>
female of
<taxonomicName id="048D4D10422BA5233BF1A3662A97FDA3" authorityName="Manikandan &amp; Megalaa &amp; Valliappan &amp; Nandini &amp; Rani &amp; Dakshinamurthi &amp; Nagappan" authorityYear="2022" box="[500,643,597,621]" class="Malacostraca" family="Epialtidae" genus="Naxioides" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="52" phylum="Arthropoda" rank="species" species="sahulensis">
<emphasis id="F1F9EA81422BA5233BF1A3662A97FDA3" box="[500,643,597,621]" italics="true" pageId="17" pageNumber="52">N. sahulensis</emphasis>
</taxonomicName>
is not fully mature, with the left P2, right P4 and both chelipeds missing, two species can be distinguished by the following characteristics that are stable in juveniles through adults of both sexes of
<taxonomicName id="048D4D10422BA5233AFDA230298CFCD4" baseAuthorityName="Haswell" baseAuthorityYear="1880" box="[248,408,771,794]" class="Malacostraca" family="Epialtidae" genus="Naxioides" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="52" phylum="Arthropoda" rank="species" species="tenuirostris">
<emphasis id="F1F9EA81422BA5233AFDA230298CFCD4" box="[248,408,771,794]" italics="true" pageId="17" pageNumber="52">N. tenuirostris</emphasis>
</taxonomicName>
determined from the specimens examined here (and the detailed account and figures of
<bibRefCitation id="A71C4B62422BA5233B8FA2742A3DFC91" author="Griffin, D. J. G." box="[394,553,839,863]" pageId="17" pageNumber="52" pagination="1 - 16" refId="ref26466" refString="Griffin, D. J. G. 1966 a. The genus Chlorinoides (Crustacea, Brachyura, Majidae), 1. A redescription of C. tenu- irostris Haswell and the status of the genus Acanthoph- rys A. Milne Edwards. Records of the Australian Museum 27: 1 - 16." type="journal article" year="1966">Griffin (1966a</bibRefCitation>
: figs. 29) and Griffrin and Tranter (1986): 1) the preorbital spine projects anterodorsally and is subparallel only to the anteriorly inclined anterior gastric spine (versus projecting vertically or inclined posteriorly and subparallel to the upright gastric and cardiac spines in
<taxonomicName id="048D4D10422BA5233B72A52B2A05FBE1" baseAuthorityName="Haswell" baseAuthorityYear="1880" box="[375,529,1048,1071]" class="Malacostraca" family="Epialtidae" genus="Naxioides" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="52" phylum="Arthropoda" rank="species" species="tenuirostris">
<emphasis id="F1F9EA81422BA5233B72A52B2A05FBE1" box="[375,529,1048,1071]" italics="true" pageId="17" pageNumber="52">N. tenuirostris</emphasis>
</taxonomicName>
); 2) the protogastric and hepatic spines of both sides are directed anterolaterally (usually directed laterally in
<taxonomicName id="048D4D10422BA5233AAFA5B32951FB59" baseAuthorityName="Haswell" baseAuthorityYear="1880" box="[170,325,1152,1175]" class="Malacostraca" family="Epialtidae" genus="Naxioides" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="52" phylum="Arthropoda" rank="species" species="tenuirostris">
<emphasis id="F1F9EA81422BA5233AAFA5B32951FB59" box="[170,325,1152,1175]" italics="true" pageId="17" pageNumber="52">N. tenuirostris</emphasis>
</taxonomicName>
); 3) the protogastric spines are similar to the hepatic spines (distinctly longer than the hepatic spines in
<taxonomicName id="048D4D10422BA5233BAFA5F62A51FB12" baseAuthorityName="Haswell" baseAuthorityYear="1880" box="[426,581,1221,1244]" class="Malacostraca" family="Epialtidae" genus="Naxioides" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="52" phylum="Arthropoda" rank="species" species="tenuirostris">
<emphasis id="F1F9EA81422BA5233BAFA5F62A51FB12" box="[426,581,1221,1244]" italics="true" pageId="17" pageNumber="52">N. tenuirostris</emphasis>
</taxonomicName>
); 4) the intestinal spine is long and distinct from the posterior carapace margin (versus proximally fused with additional short projection on the posterior carapace margin in
<taxonomicName id="048D4D10422BA5233B62A4632A10FAA9" baseAuthorityName="Haswell" baseAuthorityYear="1880" box="[359,516,1360,1383]" class="Malacostraca" family="Epialtidae" genus="Naxioides" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="52" phylum="Arthropoda" rank="species" species="tenuirostris">
<emphasis id="F1F9EA81422BA5233B62A4632A10FAA9" box="[359,516,1360,1383]" italics="true" pageId="17" pageNumber="52">N. tenuirostris</emphasis>
</taxonomicName>
, although the intestinal spine itself may also be long and slender); and 5) the P2 dactylus is as long as its propodus and much longer than its merus (shorter than both propodus and merus in
<taxonomicName id="048D4D10422BA5233BF1A4E82A99FA3C" baseAuthorityName="Haswell" baseAuthorityYear="1880" box="[500,653,1499,1522]" class="Malacostraca" family="Epialtidae" genus="Naxioides" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="52" phylum="Arthropoda" rank="species" species="tenuirostris">
<emphasis id="F1F9EA81422BA5233BF1A4E82A99FA3C" box="[500,653,1499,1522]" italics="true" pageId="17" pageNumber="52">N. tenuirostris</emphasis>
</taxonomicName>
). Also, it is remarkable that the pleon is armed with many strong spines in
<taxonomicName id="048D4D10422BA5233BCCA7132A41F9F9" box="[457,597,1567,1591]" class="Malacostraca" family="Majidae" genus="Naxia" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="52" phylum="Arthropoda" rank="species" species="sahuelsns">
<emphasis id="F1F9EA81422BA5233BCCA7132A41F9F9" box="[457,597,1567,1591]" italics="true" pageId="17" pageNumber="52">N. sahuelsns</emphasis>
</taxonomicName>
, compared to the smooth or nodular surface in
<taxonomicName id="048D4D10422BA5233880A7702912F9B2" baseAuthorityName="Haswell" baseAuthorityYear="1880" class="Malacostraca" family="Epialtidae" genus="Naxioides" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="52" phylum="Arthropoda" rank="species" species="tenuirostris">
<emphasis id="F1F9EA81422BA5233880A7702912F9B2" italics="true" pageId="17" pageNumber="52">N. tenuirostris</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="8B976518422BA5233AA2A7B429CCF92B" pageId="17" pageNumber="52" type="distribution">
<paragraph id="C3323693422BA5233AA2A7B429CCF92B" blockId="17.[140,670,215,1765]" pageId="17" pageNumber="52">
<emphasis id="F1F9EA81422BA5233AA2A7B4293DF951" box="[167,297,1671,1695]" italics="true" pageId="17" pageNumber="52">Distribution</emphasis>
. Known only from the Sahul Shelf, the
<typeStatus id="1C368831422BA5233B06A7992925F90C" box="[259,305,1706,1730]" pageId="17" pageNumber="52">type</typeStatus>
locality in the Timor Sea, E of
<collectingCountry id="BB9A7603422BA5233A89A7FE28FAF92B" box="[140,238,1741,1765]" name="Ashmore and Cartier Islands" pageId="17" pageNumber="52">Ashmore</collectingCountry>
Reef,
<quantity id="04759B76422BA5233B31A7FE299BF92A" box="[308,399,1741,1765]" metricMagnitude="1" metricUnit="m" metricValue="7.6" metricValueMax="7.8" metricValueMin="7.4" pageId="17" pageNumber="52" unit="m" value="76.0" valueMax="78.0" valueMin="74.0">7478 m</quantity>
depth.
</paragraph>
</subSubSection>
</treatment>
</document>