283 lines
38 KiB
XML
283 lines
38 KiB
XML
<document id="D467DB7CCAABE89605E369C56BE6FCD6" ID-DOI="10.11646/zootaxa.5006.1.8" ID-ISSN="1175-5326" ID-Zenodo-Dep="5159090" ID-ZooBank="564BEB55-3365-44B7-BEB2-CDBABD7B677F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628067358153" checkinUser="plazi" docAuthor="Eskov, Kirill Y. & Marusik, Yuri M." docDate="2021" docId="1304E835125E8B7345E4F989FD617C13" docLanguage="en" docName="zootaxa.5006.1.8.pdf" docOrigin="Zootaxa 5006 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Glebych minutissimus Eskov & Marusik 2021, sp. n." docType="treatment" docVersion="4" lastPageNumber="51" masterDocId="EF3D904D125C8B754573FFF4FFCE7E0A" masterDocTitle="Glebych minutissimus gen. et sp. nov., the smallest cobweb spider (Araneae Theridiidae)" masterLastPageNumber="53" masterPageNumber="45" pageNumber="47" updateTime="1699033998360" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:mods id="BA816C079ED5041962597B7FE5E62FD7" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="FFCF4F1DFEACAB95BF4EE032D5F184D2">
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<mods:title id="3B562FA53CBE209C06303E46C315C641">Glebych minutissimus gen. et sp. nov., the smallest cobweb spider (Araneae Theridiidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="3DB03AA80552B09B6346DD5F4D5EA2C7" type="personal">
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<mods:role id="EA69AFBD12277A099451F3CC5B30B93C">
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<mods:roleTerm id="E184E402AD63AAD672BF78B10E1FD163">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="EAE726AE1392D3FCBDFA982072CA882E">Eskov, Kirill Y.</mods:namePart>
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<mods:affiliation id="C4C3146425DFD6368D60A8771459E2DC">Palaeontological Institute, RAS, Profsoyuznaya 123, Moscow, 117647, Russia.</mods:affiliation>
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</mods:name>
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<mods:name id="4D458CE426C69C335CE04AAD39B3B050" type="personal">
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<mods:role id="FF0AACCD6CF04EDB5A64316A97965C51">
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<mods:roleTerm id="5F30E3751109F0747569E510B00CDCB8">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="4B057B1B9CAA53BDB579BD15ED05ACAB">Marusik, Yuri M.</mods:namePart>
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<mods:affiliation id="0AA4AFF0A9B3344387693151C0F65E5E">Institute for Biological Problems of the North RAS, Portovaya Str. 18, Magadan, 685000, Russia. & Department of Zoology & Entomology, University of the Free State, Bloemfontein 9300, South Africa. & Zoological Museum, Biodiversity Unit, FI- 20014 University of Turku, Finland</mods:affiliation>
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<mods:typeOfResource id="AD04C0C232ABCD9E048B9821BDD4280E">text</mods:typeOfResource>
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<mods:title id="70E25DCFD38AED9C7178A2BE350532F2">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="D00DC5BC2AF623DE5300F39454B6369F">
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<mods:date id="1E2EFE9C7550DAA5F6DB018DA78401B4">2021</mods:date>
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<mods:number id="42B273AF7A18CFED66ECDDB14FAB1586">2021-07-29</mods:number>
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<mods:detail id="9181662BC40E8FF07271D79756EE9F8A" type="volume">
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<mods:number id="FAEB77860F61109D7CE5B14FA0DE53C5">5006</mods:number>
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<mods:detail id="52311BACA31F2E225CE007361CE25076" type="issue">
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<mods:number id="91A66FBA16F66AC15B1FF782D6AB7833">1</mods:number>
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<mods:start id="15851CC4E3C9F7548CF45A5172828117">45</mods:start>
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<mods:classification id="A5BF02E1232332107F118805D6FEAB67">journal article</mods:classification>
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<mods:identifier id="E49C9D24CF2602C184FA11EAE43274C7" type="DOI">10.11646/zootaxa.5006.1.8</mods:identifier>
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<mods:identifier id="AB131D20AF25E1FFF33D50EA8DC807F3" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="4E10F32E9713D3443DA24C6F88C201B4" type="Zenodo-Dep">5159090</mods:identifier>
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<mods:identifier id="71EFAB9447A08BD43493EE6C0509F293" type="ZooBank">564BEB55-3365-44B7-BEB2-CDBABD7B677F</mods:identifier>
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<treatment id="1304E835125E8B7345E4F989FD617C13" ID-DOI="http://doi.org/10.5281/zenodo.5163122" ID-Zenodo-Dep="5163122" LSID="urn:lsid:plazi:treatment:1304E835125E8B7345E4F989FD617C13" httpUri="http://treatment.plazi.org/id/1304E835125E8B7345E4F989FD617C13" lastPageId="6" lastPageNumber="51" pageId="2" pageNumber="47">
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<subSubSection id="D3B70AA8125E8B7745E4F989FE23789D" box="[151,493,1661,1688]" pageId="2" pageNumber="47" type="nomenclature">
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<paragraph id="9B125923125E8B7745E4F989FE23789D" blockId="2.[151,493,1661,1724]" box="[151,493,1661,1688]" pageId="2" pageNumber="47">
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<heading id="C05AEE4F125E8B7745E4F989FE23789D" bold="true" box="[151,493,1661,1688]" fontSize="11" level="1" pageId="2" pageNumber="47" reason="1">
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<emphasis id="A9D98531125E8B7745E4F989FE23789D" bold="true" box="[151,493,1661,1688]" pageId="2" pageNumber="47">
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<taxonomicName id="5CAD22A0125E8B7745E4F989FE6D789D" ID-CoL="99CFH" authority="Eskov & Marusik, 2021" authorityName="Eskov & Marusik" authorityYear="2021" box="[151,419,1661,1688]" class="Arachnida" family="Theridiidae" genus="Glebych" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="47" phylum="Arthropoda" rank="species" species="minutissimus" status="sp. nov.">
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<emphasis id="A9D98531125E8B7745E4F989FE6D789D" bold="true" box="[151,419,1661,1688]" italics="true" pageId="2" pageNumber="47">Glebych minutissimus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="B2EA384A125E8B7744D8F989FE23789D" box="[427,493,1661,1687]" pageId="2" pageNumber="47" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="D3B70AA8125E8B7745E4F955FEFF7946" pageId="2" pageNumber="47" type="description">
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<paragraph id="9B125923125E8B7745E4F955FF3678B6" blockId="2.[151,493,1661,1724]" box="[151,248,1697,1724]" pageId="2" pageNumber="47">
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<figureCitation id="039645A6125E8B7745E4F955FF3678B6" box="[151,248,1697,1724]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStartId-0="2.[151,250,1483,1507]" captionStartId-1="3.[151,250,1757,1781]" captionStartId-2="4.[151,250,1844,1868]" captionStartId-3="5.[151,250,1593,1617]" captionStartId-4="6.[151,250,1633,1657]" captionTargetBox-0="[163,1433,181,1425]" captionTargetBox-1="[151,1436,381,1731]" captionTargetBox-2="[151,1436,181,1821]" captionTargetBox-3="[151,1435,181,1569]" captionTargetBox-4="[151,1435,561,1610]" captionTargetId-0="figure-21@2.[151,1435,181,1459]" captionTargetId-1="figure-116@3.[151,1436,381,1734]" captionTargetId-2="figure-21@4.[151,1436,181,1821]" captionTargetId-3="figure-17@5.[151,1435,181,1569]" captionTargetId-4="figure-203@6.[151,1435,561,1610]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionTargetPageId-4="6" captionText-0="FIGURE 1. General appearance of Glebych minutissumus sp. n. of the male (A–D) and female (E–F). A prosoma, dorsal; B, E body, lateral; C, F body, ventral; D prosoma, dorso-lateral. Scale = 0.2 mm. Abbreviation: Ts tracheal spiracle." captionText-1="FIGURE 2. SEM images of Glebych minutissimus sp. n., male (A–E, G–I) and female (F, J–K). A–B prosoma, lateral and frontal; C eye group; D spinnerets and colulus; E posterior part of abdomen; F–H chelicera; I terminal part of cymbium with removed bulb showing cymbial spoor; J–K epigyne, ventro-lateral and antero-ventral. Scale = 0.1 mm, unless otherwise indicated. Abbreviations: Cl colulus, Cs cymbial spur, Ep epigynal plate, MAP major ampulate gland spigot, Me median plate of epigyne, Oc copulatory opening, Ss strong seta of cymbial spur, Ts tracheal spiracle." captionText-2="FIGURE 3. SEM images of the male palp of Glebych minutissimus sp. n. A–C intact, meso-ventral, ecto-ventral and ecto-dorsal; D–I expanded palp, different views. Scale = 0.1 mm.Abbreviations: Co conductor, Cs cymbial spur, Eb embolic base, Em embolus, Ma median apophysis, Mp anterior process of median apophysis, Sm serrate retrolateral margin, Ss strong seta of Cs, Tp retrolateral pit of tegulum." captionText-3="FIGURE 4. Glebych minutissimus sp. n., male (A, B, D–G) and female (C, H–L). A prosoma, frontal; B tarsus IV; C palp; D leg I; E abdomen; F–G epiandrus (two individuals); H epigyne, ventral; I–J epigyne, antero-ventral; K epigyne and uterus internus, antero-ventral; L abdomen, antero-ventral. Scale = 0.1 mm. Abbreviations: Cd copulatory duct, Ea epiandrus, Gs gland spigot, Me median plate of epigyne, Oc copulatory opening Re receptacle, To tarsal organ." captionText-4="FIGURE 5. Collecting habitats (A–B) and type locality (C–D) of Glebych minutissimus sp. n." figureDoi-0="http://doi.org/10.5281/zenodo.5159092" figureDoi-1="http://doi.org/10.5281/zenodo.5159094" figureDoi-2="http://doi.org/10.5281/zenodo.5159096" figureDoi-3="http://doi.org/10.5281/zenodo.5159098" figureDoi-4="http://doi.org/10.5281/zenodo.5159102" httpUri-0="https://zenodo.org/record/5159092/files/figure.png" httpUri-1="https://zenodo.org/record/5159094/files/figure.png" httpUri-2="https://zenodo.org/record/5159096/files/figure.png" httpUri-3="https://zenodo.org/record/5159098/files/figure.png" httpUri-4="https://zenodo.org/record/5159102/files/figure.png" pageId="2" pageNumber="47">Figs 1–5</figureCitation>
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</paragraph>
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<paragraph id="9B125923125E8B7745E4F91DFEFF7946" blockId="2.[151,1436,1769,2012]" pageId="2" pageNumber="47">
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<emphasis id="A9D98531125E8B7745E4F91DFF2A7909" bold="true" box="[151,228,1769,1795]" pageId="2" pageNumber="47">Types.</emphasis>
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<materialsCitation id="2BC5537E125E8B774598F91DFEFF7946" collectingDate="2017-09-20" collectionCode="ZMMU, MMUM" collectorName="K. Y. Eskov" country="Peru" elevation="2500" latitude="-11.509167" location="Calabaza Village" longLatPrecision="21" longitude="-74.843056" pageId="2" pageNumber="47" specimenCount="30" specimenCount-female="20" specimenCount-male="10" stateProvince="Junin" typeStatus="holotype">
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<typeStatus id="4416E781125E8B774598F91DFE9D7909" box="[235,339,1769,1795]" pageId="2" pageNumber="47" type="holotype">Holotype</typeStatus>
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<specimenCount id="8DAB92AA125E8B774428F91EFEBE790E" box="[347,368,1770,1796]" pageId="2" pageNumber="47" type="male">♂</specimenCount>
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(
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<collectionCode id="FDBCC1E6125E8B77440CF91DFE147909" box="[383,474,1769,1795]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="2" pageNumber="47" type="Museum">ZMMU</collectionCode>
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) and
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<typeStatus id="4416E781125E8B77476BF91EFD4D790E" box="[536,643,1770,1796]" pageId="2" pageNumber="47" type="paratype">paratypes</typeStatus>
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<specimenCount id="8DAB92AA125E8B7747F8F91DFD607909" box="[651,686,1769,1795]" pageId="2" pageNumber="47" type="male">9♂</specimenCount>
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||
<specimenCount id="8DAB92AA125E8B7747C6F91DFD287909" box="[693,742,1769,1795]" pageId="2" pageNumber="47" type="female">20♀</specimenCount>
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(shared between
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<collectionCode id="FDBCC1E6125E8B7746DFF91EFBCD790E" box="[940,1027,1770,1796]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="2" pageNumber="47" type="Museum">ZMMU</collectionCode>
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and
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<collectionCode id="FDBCC1E6125E8B774148F91EFB52790E" box="[1083,1180,1770,1796]" country="United Kingdom" httpUri="http://grbio.org/cool/c9j8-pjt7" name="University of Manchester Museum, Zoology and Entomology" pageId="2" pageNumber="47" type="Museum">MMUM</collectionCode>
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)
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<collectingCountry id="E3BA19B3125E8B7741D9F91EFB3A790E" box="[1194,1268,1770,1796]" name="Peru" pageId="2" pageNumber="47">PERU</collectingCountry>
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,
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<collectingRegion id="596997C1125E8B774073F91DFA567909" box="[1280,1432,1769,1795]" country="Peru" name="Junin" pageId="2" pageNumber="47">Junin Region</collectingRegion>
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,
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<location id="9E720FF8125E8B7745E4F8F9FE98792D" LSID="urn:lsid:plazi:treatment:1304E835125E8B7345E4F989FD617C13:9E720FF8125E8B7745E4F8F9FE98792D" box="[151,342,1805,1831]" country="Peru" latitude="-11.509167" longLatPrecision="21" longitude="-74.843056" name="Calabaza Village" pageId="2" pageNumber="47" stateProvince="Junin">Calabaza Village</location>
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,
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<geoCoordinate id="FE993FE4125E8B77442DF8F9FE12792D" box="[350,476,1805,1834]" degrees="11" direction="south" minutes="30" orientation="latitude" pageId="2" pageNumber="47" precision="15" seconds="33" value="-11.509167">11º30′33′′S</geoCoordinate>
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<geoCoordinate id="FE993FE4125E8B774493F8F9FDA57922" box="[480,619,1805,1834]" degrees="74" direction="west" minutes="50" orientation="longitude" pageId="2" pageNumber="47" precision="15" seconds="35" value="-74.843056">74º50′35′′W</geoCoordinate>
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,
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<quantity id="5C55F4C6125E8B774700F8F9FD067922" box="[627,712,1805,1832]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="2" pageNumber="47" unit="m" value="2500.0">
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<elevation id="1080BE10125E8B774700F8F9FD067922" box="[627,712,1805,1832]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="2" pageNumber="47" unit="m" value="2500.0">2500 m</elevation>
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</quantity>
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, foggy forest, in epiphytic mosses and bromeliads, 19–
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<date id="EF137FE3125E8B774068F8F9FA52792D" box="[1307,1436,1805,1831]" pageId="2" pageNumber="47" value="2017-09-20">
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<collectingDate id="FF57860B125E8B774068F8F9FA52792D" box="[1307,1436,1805,1831]" pageId="2" pageNumber="47" value="2017-09-20">20.09.2017</collectingDate>
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</date>
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(
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<collectorName id="36583CF5125E8B7745D3F8C5FEEB7946" box="[160,293,1841,1868]" pageId="2" pageNumber="47">K.Y. Eskov</collectorName>
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).
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</materialsCitation>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="D3B70AA8125E8B7745B4F8A1FEE7799E" pageId="2" pageNumber="47" type="etymology">
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<paragraph id="9B125923125E8B7745B4F8A1FEE7799E" blockId="2.[151,1436,1769,2012]" pageId="2" pageNumber="47">
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<emphasis id="A9D98531125E8B7745B4F8A1FE807965" bold="true" box="[199,334,1877,1903]" pageId="2" pageNumber="47">Etymology.</emphasis>
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The species name,
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<taxonomicName id="5CAD22A0125E8B774755F8A2FD767965" authorityName="Eskov & Marusik" authorityYear="2021" box="[550,696,1878,1903]" class="Arachnida" family="Theridiidae" genus="Glebych" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="47" phylum="Arthropoda" rank="species" species="minutissimus">
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<emphasis id="A9D98531125E8B774755F8A2FD767965" box="[550,696,1878,1903]" italics="true" pageId="2" pageNumber="47">minutissimus</emphasis>
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</taxonomicName>
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, refers to the minute size of the species (smallest among all known
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<taxonomicName id="5CAD22A0125E8B7745E4F88DFED0799E" authorityName="Sundevall" authorityYear="1833" box="[151,286,1913,1940]" class="Arachnida" family="Theridiidae" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="47" phylum="Arthropoda" rank="family">Theridiidae</taxonomicName>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="D3B70AA8125E8B7345B4F869FD617C13" lastPageId="6" lastPageNumber="51" pageId="2" pageNumber="47" type="description">
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<paragraph id="9B125923125E8B7745B4F869FCE679BD" blockId="2.[151,1436,1769,2012]" box="[199,808,1949,1976]" pageId="2" pageNumber="47">
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<emphasis id="A9D98531125E8B7745B4F869FE9F79BD" bold="true" box="[199,337,1949,1975]" pageId="2" pageNumber="47">Description</emphasis>
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. Total length
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<specimenCount id="8DAB92AA125E8B77449FF86AFDCF79B2" box="[492,513,1950,1976]" pageId="2" pageNumber="47" type="male">♂</specimenCount>
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0.786–8.57,
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<specimenCount id="8DAB92AA125E8B7747E7F86AFD6779B2" box="[660,681,1950,1976]" pageId="2" pageNumber="47" type="female">♀</specimenCount>
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0.84–0.91.
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</paragraph>
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<paragraph id="9B125923125E8B7645B4F835FCC87F6F" blockId="2.[151,1436,1769,2012]" lastBlockId="3.[151,1437,150,357]" lastPageId="3" lastPageNumber="48" pageId="2" pageNumber="47">
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<emphasis id="A9D98531125E8B7745B4F835FEC579D1" bold="true" box="[199,267,1985,2011]" pageId="2" pageNumber="47">Male.</emphasis>
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Measurements: total length 0.79; carapace 0.36 long, 0.37 wide, 0.23 high; sternum 0.23 long, 0.26 wide; clypeus 0.13 high. Abdomen 0.6 long (longest axis), 0.5 wide, 0.44 high. Chelicera 0.18 long. Palp: 0.59 (0.24 (
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<geoCoordinate id="FE993FE4125F8B7640FEFF63FF027EDF" degrees="0.05" direction="east" orientation="longitude" pageId="3" pageNumber="48" precision="555" value="0.05">ø 0.05</geoCoordinate>
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), 0.1, 0.07, 0.18). Legs: I 1.23 (0.39, 0.16, 0.26, 0.21, 0.21); III 0.81 (0.27, 0.12, 0.13, 0.12, 0.17); IV 1.01 (0.3, 0.11, 0.23, 0.16, 0.21). AME 0.06, AME–AME 0.04. Ocular region with 3 setae anteriorly and 4 setae dorsally. Chelicerae with set of 3 promarginal teeth located on kind of ridge, teeth subequal in length. Sternum strongly bulged (
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<figureCitation id="039645A6125F8B764587FED3FEA17F4B" box="[244,367,295,321]" captionStart="FIGURE 1" captionStartId="2.[151,250,1483,1507]" captionTargetBox="[163,1433,181,1425]" captionTargetId="figure-21@2.[151,1435,181,1459]" captionTargetPageId="2" captionText="FIGURE 1. General appearance of Glebych minutissumus sp. n. of the male (A–D) and female (E–F). A prosoma, dorsal; B, E body, lateral; C, F body, ventral; D prosoma, dorso-lateral. Scale = 0.2 mm. Abbreviation: Ts tracheal spiracle." figureDoi="http://doi.org/10.5281/zenodo.5159092" httpUri="https://zenodo.org/record/5159092/files/figure.png" pageId="3" pageNumber="48">Fig. 1B, E</figureCitation>
|
||
). Abdomen uniformly coloured, evenly covered with setae. Epiandrus slightly bulged, darken, some structure visible through integument (
|
||
<figureCitation id="039645A6125F8B76470CFEBFFD367F6F" box="[639,760,331,357]" captionStart="FIGURE 1" captionStartId="2.[151,250,1483,1507]" captionTargetBox="[163,1433,181,1425]" captionTargetId="figure-21@2.[151,1435,181,1459]" captionTargetPageId="2" captionText="FIGURE 1. General appearance of Glebych minutissumus sp. n. of the male (A–D) and female (E–F). A prosoma, dorsal; B, E body, lateral; C, F body, ventral; D prosoma, dorso-lateral. Scale = 0.2 mm. Abbreviation: Ts tracheal spiracle." figureDoi="http://doi.org/10.5281/zenodo.5159092" httpUri="https://zenodo.org/record/5159092/files/figure.png" pageId="3" pageNumber="48">Fig. 1B–C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="CFD209AB125F8B7645E4F929FC4179A3" ID-DOI="http://doi.org/10.5281/zenodo.5159094" ID-Zenodo-Dep="5159094" httpUri="https://zenodo.org/record/5159094/files/figure.png" pageId="3" pageNumber="48" startId="3.[151,250,1757,1781]" targetBox="[151,1436,381,1731]" targetPageId="3">
|
||
<paragraph id="9B125923125F8B7645E4F929FBA978FF" blockId="3.[151,1436,1757,1962]" box="[151,1127,1757,1781]" pageId="3" pageNumber="48">
|
||
<emphasis id="A9D98531125F8B7645E4F929FEDD78FF" bold="true" box="[151,275,1757,1781]" pageId="3" pageNumber="48">FIGURE 2.</emphasis>
|
||
SEM images of
|
||
<taxonomicName id="5CAD22A0125F8B7644CEF929FD5278FF" authority="Eskov & Marusik, 2021" authorityName="Eskov & Marusik" authorityYear="2021" box="[445,668,1757,1781]" class="Arachnida" family="Theridiidae" genus="Glebych" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="48" phylum="Arthropoda" rank="species" species="minutissimus" status="sp. nov.">
|
||
<emphasis id="A9D98531125F8B7644CEF929FD5278FF" box="[445,668,1757,1781]" italics="true" pageId="3" pageNumber="48">Glebych minutissimus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="B2EA384A125F8B7647D0F92AFD1478FF" box="[675,730,1758,1781]" pageId="3" pageNumber="48" rank="species">sp. n.</taxonomicNameLabel>
|
||
, male (A–E, G–I) and female (F, J–K).
|
||
</paragraph>
|
||
<paragraph id="9B125923125F8B7645E4F8F5FE64796B" blockId="3.[151,1436,1757,1962]" pageId="3" pageNumber="48">A–B prosoma, lateral and frontal; C eye group; D spinnerets and colulus; E posterior part of abdomen; F–H chelicera; I terminal part of cymbium with removed bulb showing cymbial spoor; J–K epigyne, ventro-lateral and antero-ventral. Scale = 0.1 mm, unless otherwise indicated.</paragraph>
|
||
<paragraph id="9B125923125F8B7645E4F899FC4179A3" blockId="3.[151,1436,1757,1962]" pageId="3" pageNumber="48">
|
||
Abbreviations:
|
||
<emphasis id="A9D98531125F8B764447F899FE82798F" box="[308,332,1901,1925]" italics="true" pageId="3" pageNumber="48">Cl</emphasis>
|
||
colulus,
|
||
<emphasis id="A9D98531125F8B7644DBF899FE0D798E" box="[424,451,1901,1924]" italics="true" pageId="3" pageNumber="48">Cs</emphasis>
|
||
cymbial spur,
|
||
<emphasis id="A9D98531125F8B764724F89AFDBD798F" box="[599,627,1902,1925]" italics="true" pageId="3" pageNumber="48">Ep</emphasis>
|
||
epigynal plate,
|
||
<emphasis id="A9D98531125F8B764666F899FC87798E" box="[789,841,1901,1924]" italics="true" pageId="3" pageNumber="48">MAP</emphasis>
|
||
major ampulate gland spigot,
|
||
<emphasis id="A9D98531125F8B76410FF89AFB52798F" box="[1148,1180,1902,1925]" italics="true" pageId="3" pageNumber="48">Me</emphasis>
|
||
median plate of epigyne,
|
||
<emphasis id="A9D98531125F8B7645E4F865FF7B79A2" box="[151,181,1937,1960]" italics="true" pageId="3" pageNumber="48">Oc</emphasis>
|
||
copulatory opening,
|
||
<emphasis id="A9D98531125F8B7644FFF865FE6D79A2" box="[396,419,1937,1960]" italics="true" pageId="3" pageNumber="48">Ss</emphasis>
|
||
strong seta of cymbial spur,
|
||
<emphasis id="A9D98531125F8B7647B4F866FD1379A3" box="[711,733,1938,1961]" italics="true" pageId="3" pageNumber="48">Ts</emphasis>
|
||
tracheal spiracle.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="CFD209AB12588B7145E4F8C0FC1E79B3" ID-DOI="http://doi.org/10.5281/zenodo.5159096" ID-Zenodo-Dep="5159096" httpUri="https://zenodo.org/record/5159096/files/figure.png" pageId="4" pageNumber="49" startId="4.[151,250,1844,1868]" targetBox="[151,1436,181,1821]" targetPageId="4">
|
||
<paragraph id="9B12592312588B7145E4F8C0FC4A7946" blockId="4.[151,1437,1844,1977]" box="[151,900,1844,1869]" pageId="4" pageNumber="49">
|
||
<emphasis id="A9D9853112588B7145E4F8C0FEDD7947" bold="true" box="[151,275,1844,1869]" pageId="4" pageNumber="49">FIGURE 3.</emphasis>
|
||
SEM images of the male palp of
|
||
<taxonomicName id="5CAD22A012588B71471BF8C1FC887946" authority="Eskov & Marusik, 2021" authorityName="Eskov & Marusik" authorityYear="2021" box="[616,838,1845,1869]" class="Arachnida" family="Theridiidae" genus="Glebych" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="49" phylum="Arthropoda" rank="species" species="minutissimus" status="sp. nov.">
|
||
<emphasis id="A9D9853112588B71471BF8C1FC887946" box="[616,838,1845,1869]" italics="true" pageId="4" pageNumber="49">Glebych minutissimus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="B2EA384A12588B71463EF8C1FC4A7946" box="[845,900,1845,1868]" pageId="4" pageNumber="49" rank="species">sp. n.</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph id="9B12592312588B7145E4F8ADFC1E79B3" blockId="4.[151,1437,1844,1977]" pageId="4" pageNumber="49">
|
||
A–C intact, meso-ventral, ecto-ventral and ecto-dorsal; D–I expanded palp, different views. Scale = 0.1 mm. Abbreviations:
|
||
<emphasis id="A9D9853112588B71400CF8ADFA53797A" box="[1407,1437,1881,1904]" italics="true" pageId="4" pageNumber="49">Co</emphasis>
|
||
conductor,
|
||
<emphasis id="A9D9853112588B71447AF889FEEA799E" box="[265,292,1917,1940]" italics="true" pageId="4" pageNumber="49">Cs</emphasis>
|
||
cymbial spur,
|
||
<emphasis id="A9D9853112588B7144CEF889FE17799F" box="[445,473,1917,1941]" italics="true" pageId="4" pageNumber="49">Eb</emphasis>
|
||
embolic base,
|
||
<emphasis id="A9D9853112588B714707F889FD58799E" box="[628,662,1917,1940]" italics="true" pageId="4" pageNumber="49">Em</emphasis>
|
||
embolus,
|
||
<emphasis id="A9D9853112588B714672F889FCED799E" box="[769,803,1917,1940]" italics="true" pageId="4" pageNumber="49">Ma</emphasis>
|
||
median apophysis,
|
||
<emphasis id="A9D9853112588B71469CF889FBDF799E" box="[1007,1041,1917,1940]" italics="true" pageId="4" pageNumber="49">Mp</emphasis>
|
||
anterior process of median apophysis,
|
||
<emphasis id="A9D9853112588B7145E4F855FF7879B2" box="[151,182,1953,1976]" italics="true" pageId="4" pageNumber="49">Sm</emphasis>
|
||
serrate retrolateral margin,
|
||
<emphasis id="A9D9853112588B7144A3F855FE2979B2" box="[464,487,1953,1976]" italics="true" pageId="4" pageNumber="49">Ss</emphasis>
|
||
strong seta of
|
||
<emphasis id="A9D9853112588B714708F855FD5879B2" box="[635,662,1953,1976]" italics="true" pageId="4" pageNumber="49">Cs</emphasis>
|
||
,
|
||
<emphasis id="A9D9853112588B7147D0F855FD7079B2" box="[675,702,1953,1976]" italics="true" pageId="4" pageNumber="49">Tp</emphasis>
|
||
retrolateral pit of tegulum.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="CFD209AB12598B7045E4F9CDFE6978E8" ID-DOI="http://doi.org/10.5281/zenodo.5159098" ID-Zenodo-Dep="5159098" httpUri="https://zenodo.org/record/5159098/files/figure.png" pageId="5" pageNumber="50" startId="5.[151,250,1593,1617]" targetBox="[151,1435,181,1569]" targetPageId="5">
|
||
<paragraph id="9B12592312598B7045E4F9CDFC16785B" blockId="5.[151,1437,1593,1762]" box="[151,984,1593,1617]" pageId="5" pageNumber="50">
|
||
<emphasis id="A9D9853112598B7045E4F9CDFEDD785B" bold="true" box="[151,275,1593,1617]" pageId="5" pageNumber="50">FIGURE 4.</emphasis>
|
||
<taxonomicName id="5CAD22A012598B70446AF9CDFE36785B" authority="Eskov & Marusik, 2021" authorityName="Eskov & Marusik" authorityYear="2021" box="[281,504,1593,1617]" class="Arachnida" family="Theridiidae" genus="Glebych" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="50" phylum="Arthropoda" rank="species" species="minutissimus" status="sp. nov.">
|
||
<emphasis id="A9D9853112598B70446AF9CDFE36785B" box="[281,504,1593,1617]" italics="true" pageId="5" pageNumber="50">Glebych minutissimus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="B2EA384A12598B70448CF9CEFDF8785B" box="[511,566,1594,1617]" pageId="5" pageNumber="50" rank="species">sp. n.</taxonomicNameLabel>
|
||
, male (A, B, D–G) and female (C, H–L).
|
||
</paragraph>
|
||
<paragraph id="9B12592312598B7045E4F9A9FB047893" blockId="5.[151,1437,1593,1762]" pageId="5" pageNumber="50">A prosoma, frontal; B tarsus IV; C palp; D leg I; E abdomen; F–G epiandrus (two individuals); H epigyne, ventral; I–J epigyne, antero-ventral; K epigyne and uterus internus, antero-ventral; L abdomen, antero-ventral. Scale = 0.1 mm.</paragraph>
|
||
<paragraph id="9B12592312598B7045E4F951FE6978E8" blockId="5.[151,1437,1593,1762]" pageId="5" pageNumber="50">
|
||
Abbreviations:
|
||
<emphasis id="A9D9853112598B704444F951FE9B78B7" box="[311,341,1701,1725]" italics="true" pageId="5" pageNumber="50">Cd</emphasis>
|
||
copulatory duct,
|
||
<emphasis id="A9D9853112598B70477EF952FDE778B7" box="[525,553,1702,1725]" italics="true" pageId="5" pageNumber="50">Ea</emphasis>
|
||
epiandrus,
|
||
<emphasis id="A9D9853112598B7047D7F952FD0E78B7" box="[676,704,1702,1725]" italics="true" pageId="5" pageNumber="50">Gs</emphasis>
|
||
gland spigot,
|
||
<emphasis id="A9D9853112598B704624F952FCB978B7" box="[855,887,1702,1725]" italics="true" pageId="5" pageNumber="50">Me</emphasis>
|
||
median plate of epigyne,
|
||
<emphasis id="A9D9853112598B7041F8F952FB6778B7" box="[1163,1193,1702,1725]" italics="true" pageId="5" pageNumber="50">Oc</emphasis>
|
||
copulatory opening
|
||
<emphasis id="A9D9853112598B7040F1F952FA5378B7" box="[1410,1437,1702,1725]" italics="true" pageId="5" pageNumber="50">Re</emphasis>
|
||
receptacle,
|
||
<emphasis id="A9D9853112598B704479F93EFEEC78EB" box="[266,290,1738,1761]" italics="true" pageId="5" pageNumber="50">To</emphasis>
|
||
tarsal organ.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="9B12592312598B7045B4F8FBFBD279B3" blockId="5.[151,1437,1807,2014]" pageId="5" pageNumber="50">
|
||
Palp as in
|
||
<figureCitation id="039645A612598B70444FF8FBFE757920" box="[316,443,1807,1834]" captionStart="FIGURE 1" captionStartId="2.[151,250,1483,1507]" captionTargetBox="[163,1433,181,1425]" captionTargetId="figure-21@2.[151,1435,181,1459]" captionTargetPageId="2" captionText="FIGURE 1. General appearance of Glebych minutissumus sp. n. of the male (A–D) and female (E–F). A prosoma, dorsal; B, E body, lateral; C, F body, ventral; D prosoma, dorso-lateral. Scale = 0.2 mm. Abbreviation: Ts tracheal spiracle." figureDoi="http://doi.org/10.5281/zenodo.5159092" httpUri="https://zenodo.org/record/5159092/files/figure.png" pageId="5" pageNumber="50">Figs 1A–D</figureCitation>
|
||
,
|
||
<figureCitation id="039645A612598B7044B4F8E4FDED7920" box="[455,547,1808,1834]" captionStart="FIGURE 2" captionStartId="3.[151,250,1757,1781]" captionTargetBox="[151,1436,381,1731]" captionTargetId="figure-116@3.[151,1436,381,1734]" captionTargetPageId="3" captionText="FIGURE 2. SEM images of Glebych minutissimus sp. n., male (A–E, G–I) and female (F, J–K). A–B prosoma, lateral and frontal; C eye group; D spinnerets and colulus; E posterior part of abdomen; F–H chelicera; I terminal part of cymbium with removed bulb showing cymbial spoor; J–K epigyne, ventro-lateral and antero-ventral. Scale = 0.1 mm, unless otherwise indicated. Abbreviations: Cl colulus, Cs cymbial spur, Ep epigynal plate, MAP major ampulate gland spigot, Me median plate of epigyne, Oc copulatory opening, Ss strong seta of cymbial spur, Ts tracheal spiracle." figureDoi="http://doi.org/10.5281/zenodo.5159094" httpUri="https://zenodo.org/record/5159094/files/figure.png" pageId="5" pageNumber="50">2A–B, I</figureCitation>
|
||
,
|
||
<figureCitation id="039645A612598B704743F8E4FD8E7920" box="[560,576,1808,1834]" captionStart="FIGURE 3" captionStartId="4.[151,250,1844,1868]" captionTargetBox="[151,1436,181,1821]" captionTargetId="figure-21@4.[151,1436,181,1821]" captionTargetPageId="4" captionText="FIGURE 3. SEM images of the male palp of Glebych minutissimus sp. n. A–C intact, meso-ventral, ecto-ventral and ecto-dorsal; D–I expanded palp, different views. Scale = 0.1 mm.Abbreviations: Co conductor, Cs cymbial spur, Eb embolic base, Em embolus, Ma median apophysis, Mp anterior process of median apophysis, Sm serrate retrolateral margin, Ss strong seta of Cs, Tp retrolateral pit of tegulum." figureDoi="http://doi.org/10.5281/zenodo.5159096" httpUri="https://zenodo.org/record/5159096/files/figure.png" pageId="5" pageNumber="50">3</figureCitation>
|
||
). Femur long and thin, ca. 5 times longer than wide; patella elongate, longer than tibia; tibia with 1 trichobothrium; cymbium square shaped, anterior part not modified, cymbial spur (
|
||
<emphasis id="A9D9853112598B704006F8C0FA5D7944" box="[1397,1427,1844,1870]" italics="true" pageId="5" pageNumber="50">Cs</emphasis>
|
||
) sharply pointed, with 1 strong seta (
|
||
<emphasis id="A9D9853112598B704750F8ACFDF27978" box="[547,572,1880,1906]" italics="true" pageId="5" pageNumber="50">Ss</emphasis>
|
||
) on the base, seta longer than spur; tegulum with retrolateral pit (
|
||
<emphasis id="A9D9853112598B70407EF8ACFAE5797B" box="[1293,1323,1880,1905]" italics="true" pageId="5" pageNumber="50">Tp</emphasis>
|
||
); embolic division (=radix) hidden under median apophysis (
|
||
<emphasis id="A9D9853112598B7047ACF888FCCA799F" box="[735,772,1916,1941]" italics="true" pageId="5" pageNumber="50">Ma</emphasis>
|
||
); median apophysis with large square shaped part having serrate retrolateral margin (
|
||
<emphasis id="A9D9853112598B7044B9F854FE2279B0" box="[458,492,1952,1978]" italics="true" pageId="5" pageNumber="50">Sm</emphasis>
|
||
) and anterior process (
|
||
<emphasis id="A9D9853112598B70479DF854FCDD79B3" box="[750,787,1952,1977]" italics="true" pageId="5" pageNumber="50">Mp</emphasis>
|
||
); conductor (
|
||
<emphasis id="A9D9853112598B7046DFF854FC0379B0" box="[940,973,1952,1978]" italics="true" pageId="5" pageNumber="50">Co</emphasis>
|
||
) large.
|
||
</paragraph>
|
||
<paragraph id="9B12592312598B7345B4F830FA8F7EDF" blockId="5.[151,1437,1807,2014]" lastBlockId="6.[151,1437,150,537]" lastPageId="6" lastPageNumber="51" pageId="5" pageNumber="50">
|
||
<emphasis id="A9D9853112598B7045B4F830FED079D4" bold="true" box="[199,286,1988,2014]" pageId="5" pageNumber="50">Female</emphasis>
|
||
. Measurements: total length 0.91; carapace 0.39 long, 0.4 wide, 0.27 high; sternum 0.24, 0.28 wide; abdomen 0.76 long (longest axis), 0.58 wide, 0.48 high. Palp 0.42 (0.16, 0.08, 0.07, 0.11). Legs: I 1.15 (0.4, 0.14, 0.2, 0.19, 0.2), tarsal organ I 0.7; II 0.86 (0.3, 0.14, 0.15, 0.12, 0.15), IV 0.89 (0.33, 0.12, 0.18, 0.11. 01.5).
|
||
</paragraph>
|
||
<paragraph id="9B125923125A8B7345B4FF2BFD527F17" blockId="6.[151,1437,150,537]" pageId="6" pageNumber="51">
|
||
Chelicerae with set of 5 promarginal teeth located on a kind of ridge, mesal tooth (
|
||
<emphasis id="A9D98531125A8B734129FF2BFBB77EF2" box="[1114,1145,223,248]" italics="true" pageId="6" pageNumber="51">Mt</emphasis>
|
||
) much smaller than other. Palp with claw subequal to diameter of tarsus.
|
||
</paragraph>
|
||
<paragraph id="9B125923125A8B7345B4FED3FD677FDB" blockId="6.[151,1437,150,537]" pageId="6" pageNumber="51">
|
||
Epigyne as in
|
||
<figureCitation id="039645A6125A8B734416FED3FE777F4B" box="[357,441,295,321]" captionStart="FIGURE 1" captionStartId="2.[151,250,1483,1507]" captionTargetBox="[163,1433,181,1425]" captionTargetId="figure-21@2.[151,1435,181,1459]" captionTargetPageId="2" captionText="FIGURE 1. General appearance of Glebych minutissumus sp. n. of the male (A–D) and female (E–F). A prosoma, dorsal; B, E body, lateral; C, F body, ventral; D prosoma, dorso-lateral. Scale = 0.2 mm. Abbreviation: Ts tracheal spiracle." figureDoi="http://doi.org/10.5281/zenodo.5159092" httpUri="https://zenodo.org/record/5159092/files/figure.png" pageId="6" pageNumber="51">Figs 1F</figureCitation>
|
||
,
|
||
<figureCitation id="039645A6125A8B7344B6FED3FDCC7F4B" box="[453,514,295,321]" captionStart="FIGURE 2" captionStartId="3.[151,250,1757,1781]" captionTargetBox="[151,1436,381,1731]" captionTargetId="figure-116@3.[151,1436,381,1734]" captionTargetPageId="3" captionText="FIGURE 2. SEM images of Glebych minutissimus sp. n., male (A–E, G–I) and female (F, J–K). A–B prosoma, lateral and frontal; C eye group; D spinnerets and colulus; E posterior part of abdomen; F–H chelicera; I terminal part of cymbium with removed bulb showing cymbial spoor; J–K epigyne, ventro-lateral and antero-ventral. Scale = 0.1 mm, unless otherwise indicated. Abbreviations: Cl colulus, Cs cymbial spur, Ep epigynal plate, MAP major ampulate gland spigot, Me median plate of epigyne, Oc copulatory opening, Ss strong seta of cymbial spur, Ts tracheal spiracle." figureDoi="http://doi.org/10.5281/zenodo.5159094" httpUri="https://zenodo.org/record/5159094/files/figure.png" pageId="6" pageNumber="51">2J–K</figureCitation>
|
||
,
|
||
<figureCitation id="039645A6125A8B73477EFED3FD9C7F4B" box="[525,594,295,321]" captionStart="FIGURE 4" captionStartId="5.[151,250,1593,1617]" captionTargetBox="[151,1435,181,1569]" captionTargetId="figure-17@5.[151,1435,181,1569]" captionTargetPageId="5" captionText="FIGURE 4. Glebych minutissimus sp. n., male (A, B, D–G) and female (C, H–L). A prosoma, frontal; B tarsus IV; C palp; D leg I; E abdomen; F–G epiandrus (two individuals); H epigyne, ventral; I–J epigyne, antero-ventral; K epigyne and uterus internus, antero-ventral; L abdomen, antero-ventral. Scale = 0.1 mm. Abbreviations: Cd copulatory duct, Ea epiandrus, Gs gland spigot, Me median plate of epigyne, Oc copulatory opening Re receptacle, To tarsal organ." figureDoi="http://doi.org/10.5281/zenodo.5159098" httpUri="https://zenodo.org/record/5159098/files/figure.png" pageId="6" pageNumber="51">4H–L</figureCitation>
|
||
; epigynal plate (
|
||
<emphasis id="A9D98531125A8B734679FED3FCE77F4A" box="[778,809,295,320]" italics="true" pageId="6" pageNumber="51">Ep</emphasis>
|
||
) ca. 1.7 times wider than long, with lighter median part, formed by median plate (
|
||
<emphasis id="A9D98531125A8B7344C2FEBFFE1B7F6E" box="[433,469,331,356]" italics="true" pageId="6" pageNumber="51">Me</emphasis>
|
||
); plate and surroundings with long macrosetae, some as long as median plate’s height; copulatory openings heavily sclerotized, their distance as longs as median plate’s height; furrow between epigynal and median plates leading to copulatory openings (
|
||
<emphasis id="A9D98531125A8B7347B9FE67FD257FA7" box="[714,747,403,429]" italics="true" pageId="6" pageNumber="51">Oc</emphasis>
|
||
); receptacles droplet-shaped, as long as median plate’s height, anterior parts spaced by less than one diameter.
|
||
</paragraph>
|
||
<paragraph id="9B125923125A8B7345B4FE2FFD617C13" blockId="6.[151,1437,150,537]" pageId="6" pageNumber="51">
|
||
<emphasis id="A9D98531125A8B7345B4FE2FFEC87FFF" bold="true" box="[199,262,475,501]" pageId="6" pageNumber="51">Note.</emphasis>
|
||
Postgastral area of the female illustrated (
|
||
<figureCitation id="039645A6125A8B7347AFFE2FFC887FFF" box="[732,838,475,501]" captionStart="FIGURE 4" captionStartId="5.[151,250,1593,1617]" captionTargetBox="[151,1435,181,1569]" captionTargetId="figure-17@5.[151,1435,181,1569]" captionTargetPageId="5" captionText="FIGURE 4. Glebych minutissimus sp. n., male (A, B, D–G) and female (C, H–L). A prosoma, frontal; B tarsus IV; C palp; D leg I; E abdomen; F–G epiandrus (two individuals); H epigyne, ventral; I–J epigyne, antero-ventral; K epigyne and uterus internus, antero-ventral; L abdomen, antero-ventral. Scale = 0.1 mm. Abbreviations: Cd copulatory duct, Ea epiandrus, Gs gland spigot, Me median plate of epigyne, Oc copulatory opening Re receptacle, To tarsal organ." figureDoi="http://doi.org/10.5281/zenodo.5159098" httpUri="https://zenodo.org/record/5159098/files/figure.png" pageId="6" pageNumber="51">Figs 4I–J</figureCitation>
|
||
) has 2 fine pits that are most probably artefacts. Such pits are not visible in SEM images (
|
||
<figureCitation id="039645A6125A8B73475BFE0BFD6C7C13" box="[552,674,511,537]" captionStart="FIGURE 4" captionStartId="5.[151,250,1593,1617]" captionTargetBox="[151,1435,181,1569]" captionTargetId="figure-17@5.[151,1435,181,1569]" captionTargetPageId="5" captionText="FIGURE 4. Glebych minutissimus sp. n., male (A, B, D–G) and female (C, H–L). A prosoma, frontal; B tarsus IV; C palp; D leg I; E abdomen; F–G epiandrus (two individuals); H epigyne, ventral; I–J epigyne, antero-ventral; K epigyne and uterus internus, antero-ventral; L abdomen, antero-ventral. Scale = 0.1 mm. Abbreviations: Cd copulatory duct, Ea epiandrus, Gs gland spigot, Me median plate of epigyne, Oc copulatory opening Re receptacle, To tarsal organ." figureDoi="http://doi.org/10.5281/zenodo.5159098" httpUri="https://zenodo.org/record/5159098/files/figure.png" pageId="6" pageNumber="51">Fig. 4K–L</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |