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<mods:title id="61BEFDB8B3BB75AB8A1768545A240AFC">The bodemeyeri and the convexicollis species groups of the genus Geodromicus Redtenbacher, 1857, and additional taxonomic and faunistic data for some species of the Eastern Palaearctic Region (Coleoptera: Staphylinidae: Omaliinae)</mods:title>
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<heading id="D0A1B42977737A6BDD90F9286AE854F7" bold="true" box="[151,731,1734,1761]" fontSize="11" level="1" pageId="6" pageNumber="457" reason="1">
<taxonomicName id="4C5678C677737A6BDD90F9286AE854F7" ID-CoL="3FPGM" authority="Cameron, 1924" authorityName="Cameron" authorityYear="1924" box="[151,731,1734,1761]" class="Insecta" family="Staphylinidae" genus="Geodromicus" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="457" phylum="Arthropoda" rank="species" species="championi">
<emphasis id="B922DF5777737A6BDD90F9286AE854F7" bold="true" box="[151,731,1734,1761]" pageId="6" pageNumber="457">
<emphasis id="B922DF5777737A6BDD90F928690B54F6" bold="true" box="[151,312,1734,1761]" italics="true" pageId="6" pageNumber="457">Geodromicus</emphasis>
(s.str.)
<emphasis id="B922DF5777737A6BDC93F9286A2654F6" bold="true" box="[404,533,1734,1761]" italics="true" pageId="6" pageNumber="457">championi</emphasis>
<bibRefCitation id="EFC77EB477737A6BDF1BF9286AE854F7" author="Cameron, M." box="[540,731,1734,1760]" pageId="6" pageNumber="457" pagination="160 - 198" refId="ref30826" refString="Cameron, M. (1924). New species of Staphylinidae from India. Transactions of the Entomological Society of London, 1924, 160 - 198. https: // doi. org / 10.1111 / j. 1365 - 2311.1924. tb 03357. x" type="journal article" year="1924">Cameron, 1924</bibRefCitation>
</emphasis>
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<paragraph id="8BE9034577737A6BDD90F90569475512" blockId="6.[151,731,1734,1797]" box="[151,372,1770,1797]" pageId="6" pageNumber="457">
(
<figureCitation id="136D1FC077737A6BDD98F90568DC5512" box="[159,239,1771,1797]" captionStart="FIGURES 16" captionStartId="4.[151,264,1903,1927]" captionTargetBox="[169,1413,194,1867]" captionTargetId="figure-31@4.[154,1433,181,1879]" captionTargetPageId="4" captionText="FIGURES 16. Habitus of Geodromicus: 1—G. bodemeyeri (Turkey, Kagizman), 2—G. championi (lectotype), 3—G. iranicus (holotype), 46—G. zwickianus (Fig. 4: Iran, Qohrud Mts.; Fig. 5: Iran, Küh-e Hezâr; Fig. 6: Turkey, Elmali). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7381609" httpUri="https://zenodo.org/record/7381609/files/figure.png" pageId="6" pageNumber="457">Figs. 2</figureCitation>
,
<figureCitation id="136D1FC077737A6BDDFCF90469705512" box="[251,323,1770,1797]" captionStart="FIGURES 1318" captionStartId="8.[151,264,1943,1967]" captionTargetBox="[237,1350,193,1894]" captionTargetId="figure-31@8.[219,1367,181,1919]" captionTargetPageId="8" captionText="FIGURES 1318. Aedeagus of Geodromicus championi (Figs. 1314: Nepal, Karnali; Fig. 15—Pakistan, Chitral Bumburet) and G. zwickianus (Figs. 1617: Turkey, Yüksekova; Fig. 18: Turkey, Elmali): 13, 1516, 18—parameral view, 14, 17—lateral view. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7381614" httpUri="https://zenodo.org/record/7381614/files/figure.png" pageId="6" pageNumber="457">1315</figureCitation>
,
<figureCitation id="136D1FC077737A6BDC48F905695F5512" box="[335,364,1771,1797]" captionStart="FIGURE 27" captionStartId="13.[151,250,1984,2008]" captionTargetBox="[193,1395,1150,1960]" captionTargetId="figure-31@13.[193,1395,1150,1960]" captionTargetPageId="13" captionText="FIGURE 27. Distribution of Geodromicus championi (circles) and G. malcolmi (square) in the Himalayan Region." figureDoi="http://doi.org/10.5281/zenodo.7381624" httpUri="https://zenodo.org/record/7381624/files/figure.png" pageId="6" pageNumber="457">27</figureCitation>
)
</paragraph>
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<paragraph id="8BE9034577737A6BDD90F8C16C115550" blockId="6.[151,1058,1839,1894]" box="[151,1058,1839,1863]" pageId="6" pageNumber="457">
<treatmentCitationGroup id="AB46246B77737A6BDD90F8C16C115550" box="[151,1058,1839,1863]" pageId="6" pageNumber="457">
<treatmentCitation id="0AF7255477737A6BDD90F8C16A5B5550" author="Cameron, M." box="[151,616,1839,1863]" page="171" pageId="6" pageNumber="457" year="1924">
<taxonomicName id="4C5678C677737A6BDD90F8C16A5B5550" ID-CoL="3FPGM" authority="Cameron, 1924: 171" authorityName="Cameron" authorityPageNumber="171" authorityYear="1924" box="[151,616,1839,1863]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="457" phylum="Arthropoda" rank="species" species="championi">
<emphasis id="B922DF5777737A6BDD90F8C169A35550" box="[151,400,1839,1863]" italics="true" pageId="6" pageNumber="457">Geodromicus championi</emphasis>
<bibRefCitation id="EFC77EB477737A6BDC91F8C16A5B5550" author="Cameron, M." box="[406,616,1839,1863]" pageId="6" pageNumber="457" pagination="160 - 198" refId="ref30826" refString="Cameron, M. (1924). New species of Staphylinidae from India. Transactions of the Entomological Society of London, 1924, 160 - 198. https: // doi. org / 10.1111 / j. 1365 - 2311.1924. tb 03357. x" type="journal article" year="1924">Cameron, 1924: 171</bibRefCitation>
</taxonomicName>
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, 174;
<treatmentCitation id="0AF7255477737A6BDFA1F8C16B425550" author="Cameron, M." box="[678,881,1839,1863]" page="163" pageId="6" pageNumber="457" year="1930">
<bibRefCitation id="EFC77EB477737A6BDFA1F8C16B425550" author="Cameron, M." box="[678,881,1839,1863]" pageId="6" pageNumber="457" refId="ref30874" refString="Cameron, M. (1930) The fauna of British India including Ceylon and Burma. Coleoptera. Staphylinidae. Vol. 1. Taylor and Francis, London, xvii + 471 pp." type="book" year="1930">Cameron 1930: 163</bibRefCitation>
</treatmentCitation>
,
<treatmentCitation id="0AF7255477737A6BDE7BF8C16C115550" author="Puthz, V." box="[892,1058,1839,1863]" page="518" pageId="6" pageNumber="457" year="1973">
<bibRefCitation id="EFC77EB477737A6BDE7BF8C16C115550" author="Puthz, V." box="[892,1058,1839,1863]" pageId="6" pageNumber="457" pagination="517 - 519" refId="ref31675" refString="Puthz, V. (1973) A new Geodromicus from Pakistan (Coleoptera, Staphylinidae). Revue suisse de Zoologie, 80 (2), 517 - 519." type="journal article" year="1973">Puthz 1973: 518</bibRefCitation>
</treatmentCitation>
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</paragraph>
<paragraph id="8BE9034577737A6BDD90F8A06AA05571" blockId="6.[151,1058,1839,1894]" box="[151,659,1870,1894]" pageId="6" pageNumber="457">
<treatmentCitationGroup id="AB46246B77737A6BDD90F8A06AA05571" box="[151,659,1870,1894]" pageId="6" pageNumber="457">
<treatmentCitation id="0AF7255477737A6BDD90F8A06A7D5571" author="Coiffait, H." box="[151,590,1870,1894]" page="267" pageId="6" pageNumber="457" year="1977">
<taxonomicName id="4C5678C677737A6BDD90F8A06A7D5571" ID-CoL="3FPJ7" authority="Coiffait, 1977: 267" authorityName="Coiffait" authorityPageNumber="267" authorityYear="1977" box="[151,590,1870,1894]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="457" phylum="Arthropoda" rank="species" species="nepalicus">
<emphasis id="B922DF5777737A6BDD90F8A069B55571" box="[151,390,1870,1894]" italics="true" pageId="6" pageNumber="457">Geodromicus nepalicus</emphasis>
<bibRefCitation id="EFC77EB477737A6BDC8BF8A06A7D5571" author="Coiffait, H." box="[396,590,1870,1894]" pageId="6" pageNumber="457" pagination="243 - 275" refId="ref31094" refString="Coiffait, H. (1977) Staphilinides recoltes au Nepal par le Professeur Franz (2 e partie). Bulletin de la Societe d'Histoire Naturelle de Toulouse, 112 (1976), 243 - 275." type="journal article" year="1977">Coiffait, 1977: 267</bibRefCitation>
</taxonomicName>
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<emphasis id="B922DF5777737A6BDF53F8A06AA05571" bold="true" box="[596,659,1870,1894]" pageId="6" pageNumber="457">
<taxonomicNameLabel id="A211622C77737A6BDF53F8A06AA05571" box="[596,659,1870,1894]" pageId="6" pageNumber="457">syn.n.</taxonomicNameLabel>
</emphasis>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C34C50CE77737A6ADD90F87A6B6250E6" lastPageId="7" lastPageNumber="458" pageId="6" pageNumber="457" type="materials_examined">
<paragraph id="8BE9034577737A6ADD90F87A6CB35372" blockId="6.[151,1439,1940,2039]" lastBlockId="7.[151,1439,150,2043]" lastPageId="7" lastPageNumber="458" pageId="6" pageNumber="457">
<emphasis id="B922DF5777737A6BDD90F87A69F855B9" bold="true" box="[151,459,1940,1966]" pageId="6" pageNumber="457">Type material examined.</emphasis>
<typeStatus id="54EDBDE777737A6BDCD0F87B6A7455B8" box="[471,583,1941,1967]" pageId="6" pageNumber="457" type="lectotype">Lectotype</typeStatus>
(
<emphasis id="B922DF5777737A6BDF59F87A6B2C55B9" bold="true" box="[606,799,1940,1966]" pageId="6" pageNumber="457">here designated</emphasis>
) of
<taxonomicName id="4C5678C677737A6BDE5EF87A6D0755B9" authority="Cameron, 1924" authorityName="Cameron" authorityYear="1924" box="[857,1332,1940,1966]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="457" phylum="Arthropoda" rank="species" species="championi">
<emphasis id="B922DF5777737A6BDE5EF87A6C4255B9" box="[857,1137,1940,1966]" italics="true" pageId="6" pageNumber="457">Geodromicus championi</emphasis>
<bibRefCitation id="EFC77EB477737A6BD979F87A6D0755B9" author="Cameron, M." box="[1150,1332,1940,1966]" pageId="6" pageNumber="457" pagination="160 - 198" refId="ref30826" refString="Cameron, M. (1924). New species of Staphylinidae from India. Transactions of the Entomological Society of London, 1924, 160 - 198. https: // doi. org / 10.1111 / j. 1365 - 2311.1924. tb 03357. x" type="journal article" year="1924">Cameron, 1924</bibRefCitation>
</taxonomicName>
,
<specimenCount id="9D50C8CC77737A6BD842F87B6D6955B8" box="[1349,1370,1941,1967]" pageId="6" pageNumber="457" type="female"></specimenCount>
(
<figureCitation id="136D1FC077737A6BD868F87A689B55C5" captionStart="FIGURES 16" captionStartId="4.[151,264,1903,1927]" captionTargetBox="[169,1413,194,1867]" captionTargetId="figure-31@4.[154,1433,181,1879]" captionTargetPageId="4" captionText="FIGURES 16. Habitus of Geodromicus: 1—G. bodemeyeri (Turkey, Kagizman), 2—G. championi (lectotype), 3—G. iranicus (holotype), 46—G. zwickianus (Fig. 4: Iran, Qohrud Mts.; Fig. 5: Iran, Küh-e Hezâr; Fig. 6: Turkey, Elmali). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7381609" httpUri="https://zenodo.org/record/7381609/files/figure.png" pageId="6" pageNumber="457">Fig. 2</figureCitation>
): SYN- | TYPE &lt;printed label with blue margins&gt;, Type | H.T. &lt;round label with red margin&gt;, Chakrata Dist. | Sanj Khud 6500` &lt;printed&gt;, Dr. Cameron. | 27. V. [19]22. &lt;printed&gt;,
<taxonomicName id="4C5678C677737A6BD91DF8326C8555E1" authorityName="Redtenbacher" authorityYear="1857" box="[1050,1206,2012,2038]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="457" phylum="Arthropoda" rank="genus">Geodromicus</taxonomicName>
| championi |
<collectingRegion id="4992CDA777737A6BD85EF8326DA555E1" box="[1369,1430,2012,2038]" country="Myanmar" name="Chin" pageId="6" pageNumber="457">Cam</collectingRegion>
. &lt;handwritten&gt;, M. Cameron | Bequest. | B.M. 1955-147. &lt;printed&gt;,
<typeStatus id="54EDBDE777727A6ADEB4FF796C6B52A6" box="[947,1112,151,177]" pageId="7" pageNumber="458" type="lectotype">LECTOTYPE</typeStatus>
|
<taxonomicName id="4C5678C677727A6AD969FF796D3752A6" authorityName="Redtenbacher" authorityYear="1857" box="[1134,1284,151,177]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="458" phylum="Arthropoda" rank="genus">
<emphasis id="B922DF5777727A6AD969FF796D3752A6" box="[1134,1284,151,177]" italics="true" pageId="7" pageNumber="458">Geodromicus</emphasis>
</taxonomicName>
|
<taxonomicName id="4C5678C677727A6AD81EFF796DBC52A6" authorityName="Cameron" authorityYear="1924" box="[1305,1423,151,177]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="458" phylum="Arthropoda" rank="species" species="championi">
<emphasis id="B922DF5777727A6AD81EFF796DBC52A6" box="[1305,1423,151,177]" italics="true" pageId="7" pageNumber="458">championi</emphasis>
</taxonomicName>
|
<bibRefCitation id="EFC77EB477727A6ADD90FF55697752C2" author="Cameron, M." box="[151,324,187,213]" pageId="7" pageNumber="458" pagination="160 - 198" refId="ref30826" refString="Cameron, M. (1924). New species of Staphylinidae from India. Transactions of the Entomological Society of London, 1924, 160 - 198. https: // doi. org / 10.1111 / j. 1365 - 2311.1924. tb 03357. x" type="journal article" year="1924">Cameron, 1924</bibRefCitation>
| Shavrin A.V. des. 2016 &lt;red, printed&gt; (
<collectionCode id="ED479B8077727A6ADE12FF556B5C52C2" box="[789,879,187,213]" pageId="7" pageNumber="458">BMNH</collectionCode>
).
<typeStatus id="54EDBDE777727A6ADE86FF556C1A52C2" box="[897,1065,187,213]" pageId="7" pageNumber="458" type="paralectotype">Paralectotypes</typeStatus>
:
<specimenCount id="9D50C8CC77727A6AD934FF556C4152C2" box="[1075,1138,187,213]" pageId="7" pageNumber="458" type="male">2 ♂♂</specimenCount>
(
<specimenCount id="9D50C8CC77727A6AD987FF556CD452C2" box="[1152,1255,187,213]" pageId="7" pageNumber="458" type="male">one male</specimenCount>
dissected, other male damaged),
<specimenCount id="9D50C8CC77727A6ADC50FF3169AA52EE" box="[343,409,223,249]" pageId="7" pageNumber="458" type="female">4 ♀♀</specimenCount>
(
<specimenCount id="9D50C8CC77727A6ADCACFF316A1852EE" box="[427,555,223,249]" pageId="7" pageNumber="458" type="female">one female</specimenCount>
without antennomeres 36): Chakrata Dist. | Sainj Khud 6500` &lt;printed&gt;, SYN- | TYPE &lt;printed label with blue frame&gt;, Dr. Cameron | 27. V. [19]22. &lt;printed&gt;,
<taxonomicName id="4C5678C677727A6AD9B6FEED6DA9530A" authority="Cam." authorityName="Cameron" authorityYear="1924" box="[1201,1434,259,285]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="458" phylum="Arthropoda" rank="species" species="championi">G. championi Cam.</taxonomicName>
&lt;handwritten&gt;, M. Cameron | Bequest. | B.M. 1955-147. &lt;printed&gt; (
<collectionCode id="ED479B8077727A6ADEB2FEC96C3C5356" box="[949,1039,295,321]" pageId="7" pageNumber="458">BMNH</collectionCode>
).
<typeStatus id="54EDBDE777727A6AD924FEC96C8D5356" box="[1059,1214,295,321]" pageId="7" pageNumber="458" type="paralectotype">Paralectotype</typeStatus>
,
<specimenCount id="9D50C8CC77727A6AD9CCFEC96CD15356" box="[1227,1250,295,321]" pageId="7" pageNumber="458" type="female"></specimenCount>
: Chakrata Dist. Sanj Khud 6500, Dr. Cameron 27.V.[19]22.,
<taxonomicName id="4C5678C677727A6ADFB5FEA56C385372" authority="Cam." authorityName="Cameron" authorityYear="1924" box="[690,1035,331,357]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="458" phylum="Arthropoda" rank="species" species="championi">
<emphasis id="B922DF5777727A6ADFB5FEA56BF65372" box="[690,965,331,357]" italics="true" pageId="7" pageNumber="458">Geodromicus championi</emphasis>
Cam.
</taxonomicName>
(
<collectionCode id="ED479B8077727A6AD919FEA56C475372" box="[1054,1140,331,357]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="7" pageNumber="458">NMPC</collectionCode>
).
</paragraph>
<paragraph id="8BE9034577727A6ADDC0FE816BB9500E" blockId="7.[151,1439,150,2043]" pageId="7" pageNumber="458">
<typeStatus id="54EDBDE777727A6ADDC0FE81691C539E" box="[199,303,367,393]" pageId="7" pageNumber="458" type="holotype">Holotype</typeStatus>
of
<taxonomicName id="4C5678C677727A6ADC55FE816ACE539E" authority="Coiffait, 1977" authorityName="Coiffait" authorityYear="1977" box="[338,765,367,393]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="458" phylum="Arthropoda" rank="species" species="nepalicus">
<emphasis id="B922DF5777727A6ADC55FE816A6B539E" box="[338,600,367,393]" italics="true" pageId="7" pageNumber="458">Geodromicus nepalicus</emphasis>
<bibRefCitation id="EFC77EB477727A6ADF59FE816ACE539E" author="Coiffait, H." box="[606,765,367,393]" pageId="7" pageNumber="458" pagination="243 - 275" refId="ref31094" refString="Coiffait, H. (1977) Staphilinides recoltes au Nepal par le Professeur Franz (2 e partie). Bulletin de la Societe d'Histoire Naturelle de Toulouse, 112 (1976), 243 - 275." type="journal article" year="1977">Coiffait, 1977</bibRefCitation>
</taxonomicName>
,
<specimenCount id="9D50C8CC77727A6ADE01FE816B28539E" box="[774,795,367,393]" pageId="7" pageNumber="458" type="male"></specimenCount>
(dissected): Ufer des Tila Khola | b.Jumla, 26.9.-4.10.72 &lt;printed&gt;, Gebiet von Jumla | Westnepal, lg. H. Franz &lt;printed&gt;,
<taxonomicName id="4C5678C677727A6ADE90FE7D6C0053BA" authorityName="Redtenbacher" authorityYear="1857" box="[919,1075,403,429]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="458" phylum="Arthropoda" rank="genus">Geodromicus</taxonomicName>
| spec. | det. H. Franz [printed] &lt;handwritten&gt;, TYPE &lt;red, printed&gt;,
<taxonomicName id="4C5678C677727A6ADF6AFE596B3A53C6" authorityName="Redtenbacher" authorityYear="1857" box="[621,777,439,465]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="458" phylum="Arthropoda" rank="genus">Geodromicus</taxonomicName>
| nepalicus | H. COIFFAIT [printed] det. 19[printed]76 &lt;handwritten&gt; (
<collectionCode id="ED479B8077727A6ADC55FE35699353E2" box="[338,416,475,501]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="7" pageNumber="458">NMW</collectionCode>
).
<typeStatus id="54EDBDE777727A6ADCBDFE356A2C53E2" box="[442,543,475,501]" pageId="7" pageNumber="458" type="paratype">Paratype</typeStatus>
,
<specimenCount id="9D50C8CC77727A6ADF36FE356A5753E2" box="[561,612,475,501]" pageId="7" pageNumber="458" type="female">1 ♀</specimenCount>
: Sinja Khola zw. Chauta u. Neurigad &lt;printed&gt;, Paratype &lt;red&gt;,
<taxonomicName id="4C5678C677727A6ADD9CFE116A56500E" authority="H. Coiffait 1976" authorityName="H. Coiffait" authorityYear="1976" box="[155,613,511,537]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="458" phylum="Arthropoda" rank="species" species="nepalicus">Geodromicus nepalicus H.Coiffait 1976</taxonomicName>
&lt;handwritten&gt; (
<collectionCode id="ED479B8077727A6ADE25FE116B4D500E" box="[802,894,511,537]" pageId="7" pageNumber="458">MHNH</collectionCode>
).
</paragraph>
<paragraph id="8BE9034577727A6ADDC0FDCC6B6250E6" blockId="7.[151,1439,150,2043]" pageId="7" pageNumber="458">
<emphasis id="B922DF5777727A6ADDC0FDCC6A0E502B" bold="true" box="[199,573,546,572]" pageId="7" pageNumber="458">Additional material examined.</emphasis>
<materialsCitation id="3B3E091877727A6ADF4EFDCC69D75076" ID-GBIF-Occurrence="3979522328" collectingDate="1983-05-24" collectionCode="MHNG" collectorName="C. Besuchet &amp; I. Lobl" country="Pakistan" elevation="2275" location="Chitral Bumburet." pageId="7" pageNumber="458" specimenCount="1" specimenCount-male="1">
<emphasis id="B922DF5777727A6ADF4EFDCC6AEC502B" bold="true" box="[585,735,546,572]" pageId="7" pageNumber="458">
<collectingCountry id="F34143D577727A6ADF4EFDCC6AE9502B" box="[585,730,546,572]" name="Pakistan" pageId="7" pageNumber="458">PAKISTAN</collectingCountry>
:
</emphasis>
<specimenCount id="9D50C8CC77727A6ADFEDFDCD6B28502A" box="[746,795,547,573]" pageId="7" pageNumber="458" type="male">1 ♂</specimenCount>
:
<location id="8E89559E77727A6ADE2BFDCD6BCE502A" LSID="urn:lsid:plazi:treatment:03FFB25377737A64DD90F9286C795102:8E89559E77727A6ADE2BFDCD6BCE502A" box="[812,1021,547,573]" country="Pakistan" name="Chitral Bumburet." pageId="7" pageNumber="458">Chitral Bumburet.</location>
<elevation id="007BE47677727A6AD90FFDCC6CD4502A" box="[1032,1255,546,573]" metricMagnitude="3" metricUnit="m" metricValue="2.275" metricValueMax="2.35" metricValueMin="2.2" pageId="7" pageNumber="458" unit="m" value="2275.0" valueMax="2350.0" valueMin="2200.0">
<quantity id="4CAEAEA077727A6AD90FFDCC6C94502A" box="[1032,1191,546,573]" metricMagnitude="3" metricUnit="m" metricValue="2.275" metricValueMax="2.35" metricValueMin="2.2" pageId="7" pageNumber="458" unit="m" value="2275.0" valueMax="2350.0" valueMin="2200.0">22002350 m</quantity>
a.s.l.
</elevation>
<date id="FFE8258577727A6AD9F5FDCC6D47502A" box="[1266,1396,546,573]" pageId="7" pageNumber="458" value="1983-05-24">
<collectingDate id="EFACDC6D77727A6AD9F5FDCC6D47502A" box="[1266,1396,546,573]" pageId="7" pageNumber="458" value="1983-05-24">24.05.1983</collectingDate>
</date>
.
<collectorName id="26A3669377727A6AD884FDCD68CD5076" pageId="7" pageNumber="458">C. Besuchet</collectorName>
&amp;
<collectorName id="26A3669377727A6ADC24FDA969435076" box="[291,368,583,609]" pageId="7" pageNumber="458">I. Löbl</collectorName>
(
<collectionCode id="ED479B8077727A6ADC87FDA969EF5076" box="[384,476,583,609]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="7" pageNumber="458" type="Museum">MHNG</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B3E091877727A6ADCF7FDA86A965092" ID-GBIF-Occurrence="3979522372" collectingDate="1997-08-06" collectionCode="NME" collectorName="H. F. Grill" country="Nepal" elevation="2800" latitude="29.202778" location="Gothichaur" longitude="82.315556" pageId="7" pageNumber="458" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Karnali">
<emphasis id="B922DF5777727A6ADCF7FDA86AFC5077" bold="true" box="[496,719,582,608]" pageId="7" pageNumber="458">
<collectingCountry id="F34143D577727A6ADCF7FDA86A625077" box="[496,593,582,608]" name="Nepal" pageId="7" pageNumber="458">NEPAL</collectingCountry>
:
<collectingRegion id="4992CDA777727A6ADF5AFDA86AF85077" box="[605,715,582,608]" country="Nepal" name="Karnali" pageId="7" pageNumber="458">KARNALI</collectingRegion>
:
</emphasis>
<specimenCount id="9D50C8CC77727A6ADFD0FDA96B305076" box="[727,771,583,609]" pageId="7" pageNumber="458" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D50C8CC77727A6ADE17FDA96B615076" box="[784,850,583,609]" pageId="7" pageNumber="458" type="female">2 ♀♀</specimenCount>
:
<location id="8E89559E77727A6ADE58FDA96BEC5076" LSID="urn:lsid:plazi:treatment:03FFB25377737A64DD90F9286C795102:8E89559E77727A6ADE58FDA96BEC5076" box="[863,991,583,609]" country="Nepal" longitude="82.31556" name="Gothichaur" pageId="7" pageNumber="458" stateProvince="Karnali">Gothichaur</location>
, 29°1210N 82°1856E.
<elevation id="007BE47677727A6AD803FDA96DA45076" box="[1284,1431,583,609]" metricMagnitude="3" metricUnit="m" metricValue="2.8" pageId="7" pageNumber="458" unit="m" value="2800.0">
<quantity id="4CAEAEA077727A6AD803FDA96D695076" box="[1284,1370,583,609]" metricMagnitude="3" metricUnit="m" metricValue="2.8" pageId="7" pageNumber="458" unit="m" value="2800.0">2800 m</quantity>
a.s.l.
</elevation>
, river bank.
<date id="FFE8258577727A6ADC1CFD8569AF5092" box="[283,412,619,645]" pageId="7" pageNumber="458" value="1997-08-06">
<collectingDate id="EFACDC6D77727A6ADC1CFD8569AF5092" box="[283,412,619,645]" pageId="7" pageNumber="458" value="1997-08-06">08.06.1997</collectingDate>
</date>
.
<collectorName id="26A3669377727A6ADCAEFD856A255092" box="[425,534,619,645]" pageId="7" pageNumber="458">H.F. Grill</collectorName>
leg. (
<collectionCode id="ED479B8077727A6ADF5EFD856AAE5092" box="[601,669,619,645]" country="Germany" httpUri="http://grbio.org/cool/87iw-xebj" name="Sammlung des Naturkundemseum Erfurt" pageId="7" pageNumber="458">NME</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B3E091877727A6ADFB4FD85694D50BE" ID-GBIF-Occurrence="3979522377" collectingDate="1995-01-05" collectorName="A. Weigel" country="Nepal" elevation="2500" location="Jharjwala." municipality="Jumla" pageId="7" pageNumber="458" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D50C8CC77727A6ADFB4FD856AD25092" box="[691,737,619,645]" pageId="7" pageNumber="458" type="male">1 ♂</specimenCount>
:
<collectingMunicipality id="6B8D993F77727A6ADFE8FD856B065092" box="[751,821,619,645]" pageId="7" pageNumber="458">Jumla</collectingMunicipality>
,
<locationDeviation id="9A5B1A6777727A6ADE45FD856BC05092" box="[834,1011,619,645]" location="Jumla" pageId="7" pageNumber="458">
<quantity id="4CAEAEA077727A6ADE45FD856BB85092" box="[834,907,619,645]" metricMagnitude="4" metricUnit="m" metricValue="1.2" pageId="7" pageNumber="458" unit="km" value="12.0">12 km</quantity>
E Jumla
</locationDeviation>
,
<location id="8E89559E77727A6AD907FD856C405092" LSID="urn:lsid:plazi:treatment:03FFB25377737A64DD90F9286C795102:8E89559E77727A6AD907FD856C405092" box="[1024,1139,619,645]" country="Nepal" municipality="Jumla" name="Jharjwala." pageId="7" pageNumber="458">Jharjwala.</location>
<elevation id="007BE47677727A6AD97CFD846D3C5092" box="[1147,1295,618,645]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="7" pageNumber="458" unit="m" value="2500.0">
<quantity id="4CAEAEA077727A6AD97CFD846CE15092" box="[1147,1234,618,645]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="7" pageNumber="458" unit="m" value="2500.0">2500 m</quantity>
a.s.l.
</elevation>
<date id="FFE8258577727A6AD810FD846DAA5092" box="[1303,1433,618,645]" pageId="7" pageNumber="458" value="1995-01-05">
<collectingDate id="EFACDC6D77727A6AD810FD846DAA5092" box="[1303,1433,618,645]" pageId="7" pageNumber="458" value="1995-01-05">01.05.1995</collectingDate>
</date>
.
<collectorName id="26A3669377727A6ADD90FD61693B50BE" box="[151,264,655,681]" pageId="7" pageNumber="458">A. Weigel</collectorName>
leg. (cSh)
</materialsCitation>
;
<materialsCitation id="3B3E091877727A6ADC8DFD6168D150DA" ID-GBIF-Occurrence="3979522337" collectingDate="1995-04-29" collectionCode="NME" collectorName="M. Hartmann &amp; A. Weigel" country="Nepal" elevation="2300" location="Jumla" pageId="7" pageNumber="458" specimenCount="2" specimenCount-female="2">
<specimenCount id="9D50C8CC77727A6ADC8DFD6169FF50BE" box="[394,460,655,681]" pageId="7" pageNumber="458" type="female">2 ♀♀</specimenCount>
:
<location id="8E89559E77727A6ADCDFFD616A2D50BE" LSID="urn:lsid:plazi:treatment:03FFB25377737A64DD90F9286C795102:8E89559E77727A6ADCDFFD616A2D50BE" box="[472,542,655,681]" country="Nepal" name="Jumla" pageId="7" pageNumber="458">Jumla</location>
,
<collectorName id="26A3669377727A6ADF2EFD616AE050BE" box="[553,723,655,681]" pageId="7" pageNumber="458">Tila River, NN</collectorName>
.
<elevation id="007BE47677727A6ADFD9FD616B5D50BE" box="[734,878,655,681]" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="7" pageNumber="458" unit="m" value="2300.0">
<quantity id="4CAEAEA077727A6ADFD9FD616B0050BE" box="[734,819,655,681]" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="7" pageNumber="458" unit="m" value="2300.0">2300 m</quantity>
a.s.l.
</elevation>
<date id="FFE8258577727A6ADE72FD606BC550BE" box="[885,1014,654,681]" pageId="7" pageNumber="458" value="1995-04-29">
<collectingDate id="EFACDC6D77727A6ADE72FD606BC550BE" box="[885,1014,654,681]" pageId="7" pageNumber="458" value="1995-04-29">29.04.1995</collectingDate>
</date>
.
<collectorName id="26A3669377727A6AD906FD616CAB50BE" box="[1025,1176,655,681]" pageId="7" pageNumber="458">M. Hartmann</collectorName>
&amp;
<collectorName id="26A3669377727A6AD9BDFD616D1850BE" box="[1210,1323,655,681]" pageId="7" pageNumber="458">A. Weigel</collectorName>
leg. (cSh,
<collectionCode id="ED479B8077727A6ADD90FD5D68E950DA" box="[151,218,691,717]" country="Germany" httpUri="http://grbio.org/cool/87iw-xebj" name="Sammlung des Naturkundemseum Erfurt" pageId="7" pageNumber="458">NME</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B3E091877727A6ADDF7FD5D6A8050DA" ID-GBIF-Occurrence="3979522339" box="[240,691,691,717]" collectingDate="1995-04-29" collectionCode="NME" collectorName="J. Weipert" country="Nepal" elevation="2300" location="Jumla" pageId="7" pageNumber="458" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D50C8CC77727A6ADDF7FD5D692C50DA" box="[240,287,691,717]" pageId="7" pageNumber="458" type="male">1 ♂</specimenCount>
: same data.
<collectorName id="26A3669377727A6ADCB7FD5D6A1050DA" box="[432,547,691,717]" pageId="7" pageNumber="458">J. Weipert</collectorName>
leg. (
<collectionCode id="ED479B8077727A6ADF60FD5D6A9850DA" box="[615,683,691,717]" country="Germany" httpUri="http://grbio.org/cool/87iw-xebj" name="Sammlung des Naturkundemseum Erfurt" pageId="7" pageNumber="458">NME</collectionCode>
)
</materialsCitation>
;
<emphasis id="B922DF5777727A6ADFC6FD5C6B0550DB" bold="true" box="[705,822,690,716]" pageId="7" pageNumber="458">
<collectingRegion id="4992CDA777727A6ADFC6FD5C6B0150DB" box="[705,818,690,716]" country="Nepal" name="Gandaki" pageId="7" pageNumber="458">GANDAKI</collectingRegion>
:
</emphasis>
<materialsCitation id="3B3E091877727A6ADE38FD5D6B7E50E6" ID-GBIF-Occurrence="3979522338" collectingDate="2012-09-22" collectionCode="NME" collectorName="J. Schmidt" country="Nepal" elevation="3000" latitude="28.502222" location="Dhorpatan" longLatPrecision="20" longitude="83.129166" pageId="7" pageNumber="458" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Dhawalagiri">
<specimenCount id="9D50C8CC77727A6ADE38FD5D6B5D50DA" box="[831,878,691,717]" pageId="7" pageNumber="458" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D50C8CC77727A6ADE7BFD5D6B9850DA" box="[892,939,691,717]" pageId="7" pageNumber="458" type="female">1 ♀</specimenCount>
: SW
<collectingRegion id="4992CDA777727A6ADEEAFD5D6C5B50DA" box="[1005,1128,691,717]" country="Nepal" name="Dhawalagiri" pageId="7" pageNumber="458">Dhaulagiri</collectingRegion>
, E
<location id="8E89559E77727A6AD997FD5D6D3A50DA" LSID="urn:lsid:plazi:treatment:03FFB25377737A64DD90F9286C795102:8E89559E77727A6AD997FD5D6D3A50DA" box="[1168,1289,691,717]" country="Nepal" latitude="28.502222" longLatPrecision="20" longitude="83.129166" name="Dhorpatan" pageId="7" pageNumber="458" stateProvince="Dhawalagiri">Dhorpatan</location>
,
<geoCoordinate id="EE62658277727A6AD811FD5A6DAF50D8" box="[1302,1436,692,719]" degrees="28" direction="north" minutes="30" orientation="latitude" pageId="7" pageNumber="458" precision="15" seconds="08" value="28.502222">28°3008N</geoCoordinate>
<geoCoordinate id="EE62658277727A6ADD90FD36692E50E4" box="[151,285,728,755]" degrees="83" direction="east" minutes="07" orientation="longitude" pageId="7" pageNumber="458" precision="15" seconds="45" value="83.129166">83°0745E</geoCoordinate>
.
<elevation id="007BE47677727A6ADC2FFD39698B50E6" box="[296,440,727,753]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="7" pageNumber="458" unit="m" value="3000.0">
<quantity id="4CAEAEA077727A6ADC2FFD39694E50E6" box="[296,381,727,753]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="7" pageNumber="458" unit="m" value="3000.0">3000 m</quantity>
a.s.l.
</elevation>
<date id="FFE8258577727A6ADCB8FD396A7250E6" box="[447,577,727,753]" pageId="7" pageNumber="458" value="2012-09-22">
<collectingDate id="EFACDC6D77727A6ADCB8FD396A7250E6" box="[447,577,727,753]" pageId="7" pageNumber="458" value="2012-09-22">22.09.2012</collectingDate>
</date>
.
<collectorName id="26A3669377727A6ADF4CFD396AF250E6" box="[587,705,727,753]" pageId="7" pageNumber="458">J. Schmidt</collectorName>
leg. (
<collectionCode id="ED479B8077727A6ADE07FD396B7750E6" box="[768,836,727,753]" country="Germany" httpUri="http://grbio.org/cool/87iw-xebj" name="Sammlung des Naturkundemseum Erfurt" pageId="7" pageNumber="458">NME</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34C50CE77727A64DDC0FD1469BB5356" lastPageId="9" lastPageNumber="460" pageId="7" pageNumber="458" type="description">
<paragraph id="8BE9034577727A6ADDC0FD146BC35196" blockId="7.[151,1439,150,2043]" pageId="7" pageNumber="458">
<emphasis id="B922DF5777727A6ADDC0FD1469465103" bold="true" box="[199,373,762,788]" pageId="7" pageNumber="458">Redescription.</emphasis>
Measurements (n=15): HW: 1.061.20; HL: 0.640.80; OL: 0.320.40; TL: 0.100.12; AL (averaged): 3.54; PL: 1.041.40; PWmax: 1.431.80; PWmin: 0.961.25; ESL: 1.912.35; EW: 2.142.55; MTbL (averaged): 0.68; MTrL (averaged): 0.32 (MTrL 14: 0.17; MTrL 5: 0.15); AW: 1.952.45; AedL: 0.821.00; BL: 5.807.20 (
<typeStatus id="54EDBDE777727A6ADC11FC89694C5196" box="[278,383,871,897]" pageId="7" pageNumber="458" type="lectotype">lectotype</typeStatus>
of
<taxonomicName id="4C5678C677727A6ADCA4FC896A0F5196" authorityName="Cameron" authorityYear="1924" box="[419,572,871,897]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="458" phylum="Arthropoda" rank="species" species="championi">
<emphasis id="B922DF5777727A6ADCA4FC8969845196" box="[419,439,871,897]" italics="true" pageId="7" pageNumber="458">G</emphasis>
.
<emphasis id="B922DF5777727A6ADCC1FC896A0F5196" box="[454,572,871,897]" italics="true" pageId="7" pageNumber="458">championi</emphasis>
</taxonomicName>
: 7.20;
<typeStatus id="54EDBDE777727A6ADF8DFC896ADF5196" box="[650,748,871,897]" pageId="7" pageNumber="458" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C5678C677727A6ADE15FC896BAC5196" authorityName="H. Coiffait" authorityYear="1976" box="[786,927,871,897]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="458" phylum="Arthropoda" rank="species" species="nepalicus">
<emphasis id="B922DF5777727A6ADE15FC896B155196" box="[786,806,871,897]" italics="true" pageId="7" pageNumber="458">G</emphasis>
.
<emphasis id="B922DF5777727A6ADE33FC896BAC5196" box="[820,927,871,897]" italics="true" pageId="7" pageNumber="458">nepalicus</emphasis>
</taxonomicName>
: 6.20).
</paragraph>
<paragraph id="8BE9034577727A6ADDC0FC65698C5623" blockId="7.[151,1439,150,2043]" pageId="7" pageNumber="458">
Habitus as in
<figureCitation id="136D1FC077727A6ADC76FC65698851B2" box="[369,443,907,933]" captionStart="FIGURES 16" captionStartId="4.[151,264,1903,1927]" captionTargetBox="[169,1413,194,1867]" captionTargetId="figure-31@4.[154,1433,181,1879]" captionTargetPageId="4" captionText="FIGURES 16. Habitus of Geodromicus: 1—G. bodemeyeri (Turkey, Kagizman), 2—G. championi (lectotype), 3—G. iranicus (holotype), 46—G. zwickianus (Fig. 4: Iran, Qohrud Mts.; Fig. 5: Iran, Küh-e Hezâr; Fig. 6: Turkey, Elmali). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7381609" httpUri="https://zenodo.org/record/7381609/files/figure.png" pageId="7" pageNumber="458">Fig. 2</figureCitation>
. Body reddish-brown to black (some specimens with paler elytra); antennae brown (some specimens with paler antennomeres 12); mouthparts and legs yellow-brown to black. Forebody without microsculpture except some specimens with traces of indistinct transverse microreticulation in apical portion of head, pronotum with indistinct transverse meshes (some specimens) and abdominal tergites with very dense transverse microsculpture.
</paragraph>
<paragraph id="8BE9034577727A6ADDC0FBD369FD5762" blockId="7.[151,1439,150,2043]" pageId="7" pageNumber="458">Head 1.51.6 times as broad as long, frontal portion with strongly convex supra-antennal elevations, with deep and wide median depression, gradually narrowed and extended basad; temples moderately short, distinctly concaved between posterior margin of eye and smooth postocular ridge (indistinct in some specimens); interocellar depression very deep, moderately narrow or wide; anteocellar foveae deep, reaching level of middle or apical third of eyes. Distance between ocelli 1.21.4 times as long as distance between ocellus and posterior margin of eye. Punctation moderately large and dense, sometimes finer and sparser in middle, and larger and deeper on infraorbital portions. Antenna very long, usually exceeding middle of elytra when reclined; antennomere 2 1.3 times as long as basal antennomere, 3 distinctly longer than 2, 49 slightly shorter than 3, 10 slightly shorter than 9, apical antennomere 1.21.3 times as long as 10.</paragraph>
<paragraph id="8BE9034577727A6ADDC0FA916941543F" blockId="7.[151,1439,150,2043]" pageId="7" pageNumber="458">Pronotum slightly transverse, 1.41.5 times as broad as long, from widest anterior third strongly narrowed apicad toward subparallel and moderately long laterobasal margins; middle portion without or with indistinct longitudinal and transverse, moderately deep mediobasal depressions. Punctation very dense, distinctly larger and deeper than that in middle of head, sparser in middle and mediobasal portions; mediobasal portion of some specimens sometimes without punctation.</paragraph>
<paragraph id="8BE9034577727A6ADDC0F9DC6CB95483" blockId="7.[151,1439,150,2043]" pageId="7" pageNumber="458">Elytra slightly or strongly broadened posteriad, 1.61.8 times as long as pronotum. Punctation very dense, about as that on pronotum or distinctly denser, with interspaces between punctures as long as diameter of onetwo nearest punctures, sometimes denser and coarser in parascutellar portion and along suture.</paragraph>
<paragraph id="8BE9034577727A6ADDC9F9736A4A54A0" blockId="7.[151,1439,150,2043]" box="[206,633,1693,1719]" pageId="7" pageNumber="458">Metatibia twice as long as metatarsus.</paragraph>
<paragraph id="8BE9034577727A6ADDC0F92E69B754E8" blockId="7.[151,1439,150,2043]" pageId="7" pageNumber="458">Abdomen slightly narrower than elytra, with two moderately small transverse tomentose spots in middle of abdominal tergite IV.</paragraph>
<paragraph id="8BE9034577727A6ADDC0F8E66B1555ED" blockId="7.[151,1439,150,2043]" pageId="7" pageNumber="458">
Male. Protarsomeres 14 wide. Apical margin of abdominal tergite VIII straight or slightly concave. Apical margin of abdominal sternite VIII widely concave. Aedeagus with small basal part, gradually narrowed apicad and broadened in about middle of wide median lobe; apical third of median lobe strongly narrowed toward apical portion and from apical portion gradually narrowed toward subacute apex; parameres broadened in apical portion, with four moderately long apical setae; median lobe with two very long sclerotized structures in middle; internal sac with narrow, with very long flagellum, from basal portion stretching apicad between parallel median structures (
<figureCitation id="136D1FC077727A6ADD98F80E691B55EC" box="[159,296,2016,2043]" captionStart="FIGURES 1318" captionStartId="8.[151,264,1943,1967]" captionTargetBox="[237,1350,193,1894]" captionTargetId="figure-31@8.[219,1367,181,1919]" captionTargetPageId="8" captionText="FIGURES 1318. Aedeagus of Geodromicus championi (Figs. 1314: Nepal, Karnali; Fig. 15—Pakistan, Chitral Bumburet) and G. zwickianus (Figs. 1617: Turkey, Yüksekova; Fig. 18: Turkey, Elmali): 13, 1516, 18—parameral view, 14, 17—lateral view. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7381614" httpUri="https://zenodo.org/record/7381614/files/figure.png" pageId="7" pageNumber="458">Figs. 13, 15</figureCitation>
). Lateral aspect of the aedeagus as in
<figureCitation id="136D1FC077727A6ADFD7F80E6B1155ED" box="[720,802,2016,2042]" captionStart="FIGURES 1318" captionStartId="8.[151,264,1943,1967]" captionTargetBox="[237,1350,193,1894]" captionTargetId="figure-31@8.[219,1367,181,1919]" captionTargetPageId="8" captionText="FIGURES 1318. Aedeagus of Geodromicus championi (Figs. 1314: Nepal, Karnali; Fig. 15—Pakistan, Chitral Bumburet) and G. zwickianus (Figs. 1617: Turkey, Yüksekova; Fig. 18: Turkey, Elmali): 13, 1516, 18—parameral view, 14, 17—lateral view. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7381614" httpUri="https://zenodo.org/record/7381614/files/figure.png" pageId="7" pageNumber="458">Fig. 14</figureCitation>
.
</paragraph>
<caption id="DF2953CD777D7A65DD90F87969AE55E0" ID-DOI="http://doi.org/10.5281/zenodo.7381614" ID-Zenodo-Dep="7381614" httpUri="https://zenodo.org/record/7381614/files/figure.png" pageId="8" pageNumber="459" startId="8.[151,264,1943,1967]" targetBox="[237,1350,193,1894]" targetPageId="8">
<paragraph id="8BE90345777D7A65DD90F87969AE55E0" blockId="8.[151,1437,1943,2040]" pageId="8" pageNumber="459">
<emphasis id="B922DF57777D7A65DD90F879696655B8" bold="true" box="[151,341,1943,1967]" pageId="8" pageNumber="459">FIGURES 1318.</emphasis>
Aedeagus of
<taxonomicName id="4C5678C6777D7A65DCE4F8766AED55A7" authorityName="Cameron" authorityYear="1924" box="[483,734,1944,1968]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="459" phylum="Arthropoda" rank="species" species="championi">
<emphasis id="B922DF57777D7A65DCE4F8766AED55A7" box="[483,734,1944,1968]" italics="true" pageId="8" pageNumber="459">Geodromicus championi</emphasis>
</taxonomicName>
(Figs. 1314: Nepal, Karnali; Fig. 15—Pakistan, Chitral Bumburet) and
<taxonomicName id="4C5678C6777D7A65DDC5F852696355C3" authorityName="Fagel" authorityYear="1976" box="[194,336,1980,2004]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="459" phylum="Arthropoda" rank="species" species="zwickianus">
<emphasis id="B922DF57777D7A65DDC5F85268E755C4" box="[194,212,1980,2003]" italics="true" pageId="8" pageNumber="459">G</emphasis>
.
<emphasis id="B922DF57777D7A65DDE6F852696355C3" box="[225,336,1980,2004]" italics="true" pageId="8" pageNumber="459">zwickianus</emphasis>
</taxonomicName>
(Figs. 1617: Turkey, Yüksekova; Fig. 18: Turkey, Elmali): 13, 1516, 18—parameral view, 14, 17—lateral view. Scale bars: 0.1 mm.
</paragraph>
</caption>
<paragraph id="8BE90345777C7A64DDC0FF796A1552C2" blockId="9.[151,1437,151,789]" pageId="9" pageNumber="460">Female. Protarsomeres 14 narrow. Apical margin of abdominal tergite VIII straight or rounded. Apical margin of abdominal sternite VIII rounded.</paragraph>
<paragraph id="8BE90345777C7A64DDC0FF3069BB5356" blockId="9.[151,1437,151,789]" pageId="9" pageNumber="460">
<emphasis id="B922DF57777C7A64DDC0FF30699C52EE" bold="true" box="[199,431,222,249]" pageId="9" pageNumber="460">Comparative notes.</emphasis>
<taxonomicName id="4C5678C6777C7A64DCB2FF316B2052EE" authorityName="Cameron" authorityYear="1924" box="[437,787,223,249]" class="Insecta" family="Staphylinidae" genus="Geodromicus" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="championi">
<emphasis id="B922DF57777C7A64DCB2FF316A7852EE" box="[437,587,223,249]" italics="true" pageId="9" pageNumber="460">Geodromicus</emphasis>
(s.str.)
<emphasis id="B922DF57777C7A64DF9AFF316B2052EE" box="[669,787,223,249]" italics="true" pageId="9" pageNumber="460">championi</emphasis>
</taxonomicName>
can be distinguished from other species of the
<taxonomicName id="4C5678C6777C7A64D81DFF316DAE52EE" authorityName="Bernhauer" authorityYear="1902" box="[1306,1437,223,249]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="bodemeyeri">
<emphasis id="B922DF57777C7A64D81DFF316DAE52EE" box="[1306,1437,223,249]" italics="true" pageId="9" pageNumber="460">bodemeyeri</emphasis>
</taxonomicName>
group by the darker coloration of the body, more transverse pronotum, the shape of the apical portion of the aedeagus and longer flagellum.
</paragraph>
</subSubSection>
<subSubSection id="C34C50CE777C7A64DDC0FEA46A1853C6" pageId="9" pageNumber="460" type="distribution">
<paragraph id="8BE90345777C7A64DDC0FEA46AE1539E" blockId="9.[151,1437,151,789]" pageId="9" pageNumber="460">
<emphasis id="B922DF57777C7A64DDC0FEA469525373" bold="true" box="[199,353,330,356]" pageId="9" pageNumber="460">Distribution.</emphasis>
<taxonomicName id="4C5678C6777C7A64DC6FFEA56AFA5372" authorityName="Cameron" authorityYear="1924" box="[360,713,331,357]" class="Insecta" family="Staphylinidae" genus="Geodromicus" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="championi">
<emphasis id="B922DF57777C7A64DC6FFEA569CD5372" box="[360,510,331,357]" italics="true" pageId="9" pageNumber="460">Geodromicus</emphasis>
(s.str.)
<emphasis id="B922DF57777C7A64DF54FEA56AFA5372" box="[595,713,331,357]" italics="true" pageId="9" pageNumber="460">championi</emphasis>
</taxonomicName>
is widely distributed in the Himalayan Region and known from
<collectingCountry id="F34143D5777C7A64DD90FE8168CA539E" box="[151,249,367,393]" name="Pakistan" pageId="9" pageNumber="460">Pakistan</collectingCountry>
,
<collectingCountry id="F34143D5777C7A64DC03FE81690D539E" box="[260,318,367,393]" name="India" pageId="9" pageNumber="460">India</collectingCountry>
(
<collectingRegion id="4992CDA7777C7A64DC4BFE8169EE539E" box="[332,477,367,393]" country="India" name="Uttarakhand" pageId="9" pageNumber="460">Uttarakhand</collectingRegion>
), and
<collectingCountry id="F34143D5777C7A64DF18FE816A51539E" box="[543,610,367,393]" name="Nepal" pageId="9" pageNumber="460">Nepal</collectingCountry>
(
<figureCitation id="136D1FC0777C7A64DF76FE816AF5539E" box="[625,710,367,393]" captionStart="FIGURE 27" captionStartId="13.[151,250,1984,2008]" captionTargetBox="[193,1395,1150,1960]" captionTargetId="figure-31@13.[193,1395,1150,1960]" captionTargetPageId="13" captionText="FIGURE 27. Distribution of Geodromicus championi (circles) and G. malcolmi (square) in the Himalayan Region." figureDoi="http://doi.org/10.5281/zenodo.7381624" httpUri="https://zenodo.org/record/7381624/files/figure.png" pageId="9" pageNumber="460">Fig. 27</figureCitation>
).
</paragraph>
<paragraph id="8BE90345777C7A64DDC0FE7C6A1853C6" blockId="9.[151,1437,151,789]" pageId="9" pageNumber="460">
<emphasis id="B922DF57777C7A64DDC0FE7C697853BB" bold="true" box="[199,331,402,428]" pageId="9" pageNumber="460">Bionomics.</emphasis>
Specimens were collected at elevations from
<quantity id="4CAEAEA0777C7A64DE66FE7D6C2853BA" box="[865,1051,403,429]" metricMagnitude="3" metricUnit="m" metricValue="2.4899999999999998" metricValueMax="3.0" metricValueMin="1.98" pageId="9" pageNumber="460" unit="m" value="2490.0" valueMax="3000.0" valueMin="1980.0">1980 to 3000 m</quantity>
a.s.l. and were taken from under stones and gravel along river banks.
</paragraph>
</subSubSection>
<subSubSection id="C34C50CE777C7A64DDC0FE356C795102" pageId="9" pageNumber="460" type="discussion">
<paragraph id="8BE90345777C7A64DDC0FE3569B35076" blockId="9.[151,1437,151,789]" pageId="9" pageNumber="460">
<emphasis id="B922DF57777C7A64DDC0FE35690853E2" bold="true" box="[199,315,475,501]" pageId="9" pageNumber="460">Remarks.</emphasis>
Based on the unspecified number of
<typeStatus id="54EDBDE7777C7A64DFE0FE356B7F53E2" box="[743,844,475,501]" pageId="9" pageNumber="460" type="syntype">syntypes</typeStatus>
,
<taxonomicName id="4C5678C6777C7A64DE5FFE356C7353E2" authorityName="Cameron" authorityYear="1924" box="[856,1088,475,501]" class="Insecta" family="Staphylinidae" genus="Geodromicus" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="championi">
<emphasis id="B922DF57777C7A64DE5FFE356B5F53E2" box="[856,876,475,501]" italics="true" pageId="9" pageNumber="460">G</emphasis>
. (s.str.)
<emphasis id="B922DF57777C7A64DECDFE356C7353E2" box="[970,1088,475,501]" italics="true" pageId="9" pageNumber="460">championi</emphasis>
</taxonomicName>
was originally described from “Chakrata District; Sanj Khud and
<taxonomicName id="4C5678C6777C7A64DF19FE116AA2500E" authority=", Gad" authorityName="Gad" box="[542,657,511,537]" class="Insecta" family="Hesperiidae" genus="Bina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="460" phylum="Arthropoda" rank="genus">Bina, Gad</taxonomicName>
,
<quantity id="4CAEAEA0777C7A64DF9BFE106B53500E" box="[668,864,510,537]" metricMagnitude="3" metricUnit="m" metricValue="2.2098" metricValueMax="2.4384" metricValueMin="1.9812" pageId="9" pageNumber="460" unit="ft" value="7250.0" valueMax="8000.0" valueMin="6500.0">6500 to 8000 feet</quantity>
above the sea” (
<collectingCountry id="F34143D5777C7A64D910FE116C66500E" box="[1047,1109,511,537]" name="India" pageId="9" pageNumber="460">India</collectingCountry>
,
<collectingRegion id="4992CDA7777C7A64D967FE116CC3500E" box="[1120,1264,511,537]" country="India" name="Uttarakhand" pageId="9" pageNumber="460">Uttarakhand</collectingRegion>
). The specimen with the best preservation and additional label “Type H.T.” was designated as the
<typeStatus id="54EDBDE7777C7A64D95EFDCD6CF3502A" box="[1113,1216,547,573]" pageId="9" pageNumber="460" type="lectotype">lectotype</typeStatus>
in order to fix the identity of the name.
</paragraph>
<paragraph id="8BE90345777C7A64DDC0FD8568C350E6" blockId="9.[151,1437,151,789]" pageId="9" pageNumber="460">
<taxonomicName id="4C5678C6777C7A64DDC0FD856A2B5092" authorityName="H. Coiffait" authorityYear="1976" box="[199,536,619,645]" class="Insecta" family="Staphylinidae" genus="Geodromicus" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="nepalicus">
<emphasis id="B922DF57777C7A64DDC0FD85696E5092" box="[199,349,619,645]" italics="true" pageId="9" pageNumber="460">Geodromicus</emphasis>
(s.str.)
<emphasis id="B922DF57777C7A64DCAAFD856A2B5092" box="[429,536,619,645]" italics="true" pageId="9" pageNumber="460">nepalicus</emphasis>
</taxonomicName>
was described based on the
<typeStatus id="54EDBDE7777C7A64DE4BFD856B9D5092" box="[844,942,619,645]" pageId="9" pageNumber="460" type="holotype">holotype</typeStatus>
from “Rive du Tila Khola, près Jumla,
<collectingCountry id="F34143D5777C7A64D85EFD856DAF5092" box="[1369,1436,619,645]" name="Nepal" pageId="9" pageNumber="460">Népal</collectingCountry>
occidental…”. During the study of the
<typeStatus id="54EDBDE7777C7A64DF5EFD616A8D50BE" box="[601,702,655,681]" pageId="9" pageNumber="460" type="holotype">holotype</typeStatus>
, I have not found sufficient external and internal morphological differences between it and other studied specimens of
<taxonomicName id="4C5678C6777C7A64DE14FD5D6B9C50DA" authorityName="Cameron" authorityYear="1924" box="[787,943,691,717]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="championi">
<emphasis id="B922DF57777C7A64DE14FD5D6B1450DA" box="[787,807,691,717]" italics="true" pageId="9" pageNumber="460">G</emphasis>
.
<emphasis id="B922DF57777C7A64DE3EFD5D6B9C50DA" box="[825,943,691,717]" italics="true" pageId="9" pageNumber="460">championi</emphasis>
</taxonomicName>
. Thus, it was synonymized with the latter species.
</paragraph>
<paragraph id="8BE90345777C7A64DDC0FD156C795102" blockId="9.[151,1437,151,789]" box="[199,1098,763,789]" pageId="9" pageNumber="460">
<taxonomicName id="4C5678C6777C7A64DDC0FD156A145102" authorityName="Cameron" authorityYear="1924" box="[199,551,763,789]" class="Insecta" family="Staphylinidae" genus="Geodromicus" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="championi">
<emphasis id="B922DF57777C7A64DDC0FD15696E5102" box="[199,349,763,789]" italics="true" pageId="9" pageNumber="460">Geodromicus</emphasis>
(s.str.)
<emphasis id="B922DF57777C7A64DCB6FD156A145102" box="[433,551,763,789]" italics="true" pageId="9" pageNumber="460">championi</emphasis>
</taxonomicName>
is here recorded from
<collectingCountry id="F34143D5777C7A64DE22FD156BB75102" box="[805,900,763,789]" name="Pakistan" pageId="9" pageNumber="460">Pakistan</collectingCountry>
for the first time.
</paragraph>
</subSubSection>
</treatment>
</document>