224 lines
31 KiB
XML
224 lines
31 KiB
XML
<document id="C8F771DD3326EF8EDD668BBA2EFFE666" ID-DOI="10.11646/zootaxa.5450.1.1" ID-ISSN="1175-5334" ID-Zenodo-Dep="11232500" ID-ZooBank="1B5D192F-1D5B-4460-9133-9AEAE9C920BF" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1716281048833" checkinUser="plazi" docAuthor="Johnson, Robert G." docDate="2024" docId="03FD8227FFBFE341FF78FBAEFC3F82EC" docLanguage="en" docName="zootaxa.5450.1.1.pdf" docOrigin="Zootaxa 5450 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5450.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.19:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="19" docTitle="Stoloharpes calceolus Johnson 2024, n. sp." docType="treatment" docVersion="2" lastPageNumber="98" masterDocId="FFC4FA5FFFDFE320FFEFF87CFFB58771" masterDocTitle="Devonian Harpetidae from the central and eastern Anti-Atlas, Morocco" masterLastPageNumber="185" masterPageNumber="1" pageNumber="97" updateTime="1716282690877" updateUser="ExternalLinkService">
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<mods:title id="0ABBF0EF07A5C95C25FA4A1FE8D528B5">Devonian Harpetidae from the central and eastern Anti-Atlas, Morocco</mods:title>
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<mods:namePart id="7235E8ABCCA6FC6577A035305C49BB51">Johnson, Robert G.</mods:namePart>
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<mods:title id="7CEB59CFE96158CDA79B1CA980C9787D">Zootaxa</mods:title>
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<mods:part id="4F456BC4AE947BA2B4D5D597E7348CE3">
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<mods:date id="D09DDE879B08A54831B34CEAD98DE870">2024</mods:date>
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<mods:number id="F1E28DCC3FC7D27BE212C680F3BEC47B">2024-05-15</mods:number>
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<mods:number id="5E251ED2F4F41112021A68BC3F29FC51">5450</mods:number>
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<mods:url id="B65AF072DB5D5C6F869B33962D6239D5">http://dx.doi.org/10.11646/zootaxa.5450.1.1</mods:url>
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<mods:classification id="4F45D2C4BCA9149F08153E68B2D19CBE">journal article</mods:classification>
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<mods:identifier id="ABE3B202D689311609AD05AF39415700" type="DOI">10.11646/zootaxa.5450.1.1</mods:identifier>
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<mods:identifier id="EF880022A6AE9B4330751422FBF8BE5E" type="ISSN">1175-5334</mods:identifier>
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<mods:identifier id="7E864A1B147EB0C1B1E08361CFF08668" type="Zenodo-Dep">11232500</mods:identifier>
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<treatment id="03FD8227FFBFE341FF78FBAEFC3F82EC" ID-DOI="http://doi.org/10.5281/zenodo.11232658" ID-Zenodo-Dep="11232658" LSID="urn:lsid:plazi:treatment:03FD8227FFBFE341FF78FBAEFC3F82EC" httpUri="http://treatment.plazi.org/id/03FD8227FFBFE341FF78FBAEFC3F82EC" lastPageId="97" lastPageNumber="98" pageId="96" pageNumber="97">
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<subSubSection id="C34E60BAFFBFE340FF78FBAEFE5D849C" box="[151,488,978,1005]" pageId="96" pageNumber="97" type="nomenclature">
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<paragraph id="8BEB3331FFBFE340FF78FBAEFE5D849C" blockId="96.[151,488,978,1041]" box="[151,488,978,1005]" pageId="96" pageNumber="97">
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<heading id="D0A3845DFFBFE340FF78FBAEFE5D849C" bold="true" box="[151,488,978,1005]" fontSize="11" level="2" pageId="96" pageNumber="97" reason="4">
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||
<emphasis id="B920EF23FFBFE340FF78FBAEFE5D849C" bold="true" box="[151,488,978,1005]" pageId="96" pageNumber="97">
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<taxonomicName id="4C5448B2FFBFE340FF78FBAEFE2B849C" authority="Johnson, 2024" authorityName="Johnson" authorityYear="2024" box="[151,414,978,1005]" class="Trilobita" family="Harpetidae" genus="Stoloharpes" kingdom="Animalia" order="Harpetida" pageId="96" pageNumber="97" phylum="Arthropoda" rank="species" species="calceolus" status="sp. nov.">
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<emphasis id="B920EF23FFBFE340FF78FBAEFE2B849C" bold="true" box="[151,414,978,1005]" italics="true" pageId="96" pageNumber="97">Stoloharpes calceolus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2135258FFBFE340FE49FBAFFE5D849C" box="[422,488,979,1005]" pageId="96" pageNumber="97" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C34E60BAFFBFE340FF78FB8BFE9D8360" box="[151,296,1015,1041]" pageId="96" pageNumber="97" type="description">
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<paragraph id="8BEB3331FFBFE340FF78FB8BFE9D8360" blockId="96.[151,488,978,1041]" box="[151,296,1015,1041]" pageId="96" pageNumber="97">
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<figureCitation id="136F2FB4FFBFE340FF78FB8BFE9D8360" box="[151,296,1015,1041]" captionStart="PLATE 49" captionStartId="155.[151,234,1681,1705]" captionTargetBox="[236,1351,181,1658]" captionTargetId="figure-15@155.[236,1351,181,1659]" captionTargetPageId="155" captionText="PLATE 49. A–F Stoloharpes aff. rissaniensis, El Otfal formation, upper Eifelian, Ksar Hannabou (Fig. 2B, Map 6, site 14), dorsal exoskeleton, thorax disarticulated and pygidium missing, NHMUK It 29380. A, dorsal B, lateral and C, anterior view. D, dorsal and E, lateral view of eye lobe (uncoated). F, dorsal view of brim perforations. G–K Stoloharpes calceolus n. sp., “Akantharges Couche”, Eifelian, Ksar Lbor (Fig. 2B, Map 3, site 4), dorsal exoskeleton pygidium enrolled, holotype, NHMUK It 29363. G, dorsal, H, anterior and I, lateral view. J, dorsal view of brim perforations and K, lateral view of pleurae. L–P Stoloharpes aff. calceolus, “Akantharges Couche”, Eifelian, Ksar Lbor (Fig. 2B, Map 3, site 5), dorsal exoskeleton, thorax disarticulated pygidium enrolled, holotype, NHMUK It 29381. L, dorsal, M, anterior, N, lateral and O, posterior view. P, dorsal view of brim perforations. Scale bars shown represent 1 cm unless otherwise indicated." figureDoi="http://doi.org/10.5281/zenodo.11232687" httpUri="https://zenodo.org/record/11232687/files/figure.png" pageId="96" pageNumber="97">Plate 49G–K</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C34E60BAFFBFE340FF78FC42FB7C827C" pageId="96" pageNumber="97" type="diagnosis">
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<paragraph id="8BEB3331FFBFE340FF78FC42FB7C827C" blockId="96.[151,1437,1086,2013]" pageId="96" pageNumber="97">
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<emphasis id="B920EF23FFBFE340FF78FC42FEA78329" bold="true" box="[151,274,1086,1112]" pageId="96" pageNumber="97">Diagnosis.</emphasis>
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Cephalon mildly pyriform; glabella sloping gently anteriorly and narrow, width (tr.) just over 50% of length (sag.) of glabella anterior to S0. Axial furrows nearly straight and weakly converging anteriorly. Preaxial furrows also converging and initially deep, becoming shallower anteriorly as anterior boss deflates to disappear close to girder. Occipital ring with prominent occipital node. Alar furrow well-defined and eyes elongated (exsag.). Genal roll sloping anteriorly at same angle as anterior of glabella, just under
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<superScript id="7C219E79FFBFE340FC1AFCB2FBAC83AD" attach="none" box="[1013,1049,1230,1257]" fontSize="6" pageId="96" pageNumber="97">250</superScript>
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. Laterally, slope steepens strongly to around
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<superScript id="7C219E79FFBFE340FEE6FC8EFE988271" attach="none" box="[265,301,1266,1293]" fontSize="6" pageId="96" pageNumber="97">550</superScript>
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. Brim strongly convex, moderately broad (sag.) and steeply sloping at around
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<superScript id="7C219E79FFBFE340FB71FC8EFB778271" attach="none" box="[1182,1218,1266,1293]" fontSize="6" pageId="96" pageNumber="97">350</superScript>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C34E60BAFFBFE340FF28FD6BFC208240" box="[199,917,1303,1329]" pageId="96" pageNumber="97" type="etymology">
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<paragraph id="8BEB3331FFBFE340FF28FD6BFC208240" blockId="96.[151,1437,1086,2013]" box="[199,917,1303,1329]" pageId="96" pageNumber="97">
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<emphasis id="B920EF23FFBFE340FF28FD6BFEFB8240" bold="true" box="[199,334,1303,1329]" pageId="96" pageNumber="97">Etymology.</emphasis>
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Named after the species’ slipper-like lateral profile.
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</paragraph>
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</subSubSection>
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<subSubSection id="C34E60BAFFBFE340FF28FD46FED98294" pageId="96" pageNumber="97" type="materials_examined">
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<paragraph id="8BEB3331FFBFE340FF28FD46FBB08208" blockId="96.[151,1437,1086,2013]" pageId="96" pageNumber="97">
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<emphasis id="B920EF23FFBFE340FF28FD46FE418224" bold="true" box="[199,500,1338,1365]" pageId="96" pageNumber="97">Material and occurrence</emphasis>
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.
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<materialsCitation id="3B3C396CFFBFE340FDEBFD47FBB08208" collectionCode="NHMUK" pageId="96" pageNumber="97" specimenCount="1" typeStatus="holotype">
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<typeStatus id="54EF8D93FFBFE340FDEBFD47FDC48224" box="[516,625,1339,1365]" pageId="96" pageNumber="97" type="holotype">Holotype</typeStatus>
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:
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<collectionCode id="ED45ABF4FFBFE340FD92FD47FD528224" box="[637,743,1339,1365]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="96" pageNumber="97" type="Museum">NHMUK</collectionCode>
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It 29363,
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<figureCitation id="136F2FB4FFBFE340FC8DFD47FC6A8224" box="[866,991,1339,1365]" captionStart="PLATE 49" captionStartId="155.[151,234,1681,1705]" captionTargetBox="[236,1351,181,1658]" captionTargetId="figure-15@155.[236,1351,181,1659]" captionTargetPageId="155" captionText="PLATE 49. A–F Stoloharpes aff. rissaniensis, El Otfal formation, upper Eifelian, Ksar Hannabou (Fig. 2B, Map 6, site 14), dorsal exoskeleton, thorax disarticulated and pygidium missing, NHMUK It 29380. A, dorsal B, lateral and C, anterior view. D, dorsal and E, lateral view of eye lobe (uncoated). F, dorsal view of brim perforations. G–K Stoloharpes calceolus n. sp., “Akantharges Couche”, Eifelian, Ksar Lbor (Fig. 2B, Map 3, site 4), dorsal exoskeleton pygidium enrolled, holotype, NHMUK It 29363. G, dorsal, H, anterior and I, lateral view. J, dorsal view of brim perforations and K, lateral view of pleurae. L–P Stoloharpes aff. calceolus, “Akantharges Couche”, Eifelian, Ksar Lbor (Fig. 2B, Map 3, site 5), dorsal exoskeleton, thorax disarticulated pygidium enrolled, holotype, NHMUK It 29381. L, dorsal, M, anterior, N, lateral and O, posterior view. P, dorsal view of brim perforations. Scale bars shown represent 1 cm unless otherwise indicated." figureDoi="http://doi.org/10.5281/zenodo.11232687" httpUri="https://zenodo.org/record/11232687/files/figure.png" pageId="96" pageNumber="97">Pl. 49G–K</figureCitation>
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, from
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<emphasis id="B920EF23FFBFE340FBC3FD46FB8F8225" bold="true" box="[1068,1082,1338,1364]" pageId="96" pageNumber="97">“</emphasis>
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<taxonomicName id="4C5448B2FFBFE340FBD5FD47FA908224" authority="Couche" authorityName="Couche" box="[1082,1317,1339,1365]" class="Trilobita" family="Lichidae" genus="Basseiarges" kingdom="Animalia" order="Lichida" pageId="96" pageNumber="97" phylum="Arthropoda" rank="genus">
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<emphasis id="B920EF23FFBFE340FBD5FD47FB758225" box="[1082,1216,1339,1364]" italics="true" pageId="96" pageNumber="97">Basseiarges</emphasis>
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Couche
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</taxonomicName>
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”, Eifelian, Ksar Lbor (
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<figureCitation id="136F2FB4FFBFE340FEF6FD23FEDC8208" box="[281,361,1375,1401]" captionStart="FIGURE 2" captionStartId="7.[152,255,1943,1967]" captionText="FIGURE 2A. Map locations in the Souss–Massa region" pageId="96" pageNumber="97">Fig.2B</figureCitation>
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, Map 3, site 3), dorsal exoskeleton with pygidium enrolled.
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</materialsCitation>
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</paragraph>
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<paragraph id="8BEB3331FFBFE340FF28FDFFFED98294" blockId="96.[151,1437,1086,2013]" pageId="96" pageNumber="97">
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Other material:
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<taxonomicName id="4C5448B2FFBFE340FE93FDFFFD7082EC" authority=", NHMUK It" authorityName="NHMUK It" box="[380,709,1411,1437]" class="Trilobita" family="Harpetidae" genus="Stoloharpes" isUncertain="true" kingdom="Animalia" order="Harpetida" pageId="96" pageNumber="97" phylum="Arthropoda" rank="species" species="calceolus">
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<emphasis id="B920EF23FFBFE340FE93FDFFFE2482EC" box="[380,401,1411,1437]" italics="true" pageId="96" pageNumber="97">S.</emphasis>
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aff.
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<emphasis id="B920EF23FFBFE340FE27FDFFFD8782EC" box="[456,562,1411,1437]" italics="true" pageId="96" pageNumber="97">calceolus</emphasis>
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, NHMUK It
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</taxonomicName>
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299381,
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<figureCitation id="136F2FB4FFBFE340FCDDFDFFFC1182EC" box="[818,932,1411,1437]" captionStart="PLATE 49" captionStartId="155.[151,234,1681,1705]" captionTargetBox="[236,1351,181,1658]" captionTargetId="figure-15@155.[236,1351,181,1659]" captionTargetPageId="155" captionText="PLATE 49. A–F Stoloharpes aff. rissaniensis, El Otfal formation, upper Eifelian, Ksar Hannabou (Fig. 2B, Map 6, site 14), dorsal exoskeleton, thorax disarticulated and pygidium missing, NHMUK It 29380. A, dorsal B, lateral and C, anterior view. D, dorsal and E, lateral view of eye lobe (uncoated). F, dorsal view of brim perforations. G–K Stoloharpes calceolus n. sp., “Akantharges Couche”, Eifelian, Ksar Lbor (Fig. 2B, Map 3, site 4), dorsal exoskeleton pygidium enrolled, holotype, NHMUK It 29363. G, dorsal, H, anterior and I, lateral view. J, dorsal view of brim perforations and K, lateral view of pleurae. L–P Stoloharpes aff. calceolus, “Akantharges Couche”, Eifelian, Ksar Lbor (Fig. 2B, Map 3, site 5), dorsal exoskeleton, thorax disarticulated pygidium enrolled, holotype, NHMUK It 29381. L, dorsal, M, anterior, N, lateral and O, posterior view. P, dorsal view of brim perforations. Scale bars shown represent 1 cm unless otherwise indicated." figureDoi="http://doi.org/10.5281/zenodo.11232687" httpUri="https://zenodo.org/record/11232687/files/figure.png" pageId="96" pageNumber="97">Pl. 49L–P</figureCitation>
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, from
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<emphasis id="B920EF23FFBFE340FC1FFDFEFC4B82ED" bold="true" box="[1008,1022,1410,1436]" pageId="96" pageNumber="97">“</emphasis>
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<taxonomicName id="4C5448B2FFBFE340FC11FDFFFB5D82EC" authority="Couche" authorityName="Couche" box="[1022,1256,1411,1437]" class="Trilobita" family="Lichidae" genus="Basseiarges" kingdom="Animalia" order="Lichida" pageId="96" pageNumber="97" phylum="Arthropoda" rank="genus">
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<emphasis id="B920EF23FFBFE340FC11FDFFFB3182ED" box="[1022,1156,1411,1436]" italics="true" pageId="96" pageNumber="97">Basseiarges</emphasis>
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Couche
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</taxonomicName>
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”, Eifelian, Ksar Lbor,
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<quantity id="4CAC9ED4FFBFE340FF35FDDBFE9482B0" box="[218,289,1447,1473]" metricMagnitude="4" metricUnit="m" metricValue="2.8" pageId="96" pageNumber="97" unit="km" value="28.0">28 km</quantity>
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E of Tinghir, at the base of Jbel Tisdafine (
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<figureCitation id="136F2FB4FFBFE340FCEAFDDBFCE882B0" box="[773,861,1447,1473]" captionStart="FIGURE 2" captionStartId="7.[152,255,1943,1967]" captionText="FIGURE 2A. Map locations in the Souss–Massa region" pageId="96" pageNumber="97">Fig. 2B</figureCitation>
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, Map 3, site 4), dorsal exoskeleton of cephalon and partial thorax only.
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</paragraph>
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</subSubSection>
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<subSubSection id="C34E60BAFFBFE341FF28FD92FE7B8464" lastPageId="97" lastPageNumber="98" pageId="96" pageNumber="97" type="description">
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<paragraph id="8BEB3331FFBFE340FF28FD92FBFB8058" blockId="96.[151,1437,1086,2013]" pageId="96" pageNumber="97">
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<emphasis id="B920EF23FFBFE340FF28FD92FEEC8179" bold="true" box="[199,345,1518,1544]" pageId="96" pageNumber="97">Description.</emphasis>
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Cephalon weakly pyriform, narrow, length (sag.) 85% of width (tr.) and widest at mid-point between eye lobes and alae. Genicranium very narrow, width (tr.) being same as length (sag.), widest at posterior border. Glabella strongly carinate, sloping very gently anteriorly and narrow, width (tr.) at S1 being about half sagittal length of glabella anterior to S0. S1 rises obliquely from axial furrow, before fading halfway up glabella. Other lateral glabellar impressions are missing and L1 inflated. Axial furrows straight and converging weakly anteriorly. Occipital ring slightly broadened medially and with prominent occipital node, base of which reaches down into occipital furrow. Top of occipital node only just lower than crest of glabella anterior to S0. Alae nearly as broad (tr.) as long (exsag.), anterolaterally directed and not quite reaching halfway across genal area. Alar furrows are well-defined. Posterior border short (tr.), around 35% of occipital ring width (tr.).
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</paragraph>
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<paragraph id="8BEB3331FFBFE340FF28FF4FFD4780AC" blockId="96.[151,1437,1086,2013]" pageId="96" pageNumber="97">
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Eye lobes large (exsag.), and no eye ridge. Anterior boss narrow, inflated, strongly connected to glabella but not reaching down to girder. Preaxial furrows slightly convergent anteriorly. Genal roll wide and sloping very gently at just under
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<superScript id="7C219E79FFBFE340FEC3FF06FEE580F9" attach="left" box="[300,336,1914,1941]" fontSize="6" pageId="96" pageNumber="97">250</superScript>
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anteriorly, steepening to about
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<superScript id="7C219E79FFBFE340FD58FF06FD6E80F9" attach="left" box="[695,731,1914,1941]" fontSize="6" pageId="96" pageNumber="97">550</superScript>
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posterolaterally. In lateral view, inner fringe narrows strongly posterolaterally. Fine perforations, relatively sparsely distributed, increase in size on extensions to genal roll on prolongations. Girder narrow and lacking girder kink.
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</paragraph>
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<paragraph id="8BEB3331FFBEE341FF28F8EBFAFA8788" blockId="97.[151,1437,150,1437]" pageId="97" pageNumber="98">
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Brim convex in profile, steeply-sloping at around
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<superScript id="7C219E79FFBEE341FCE2F8EBFC8487D5" attach="left" box="[781,817,150,177]" fontSize="6" pageId="97" pageNumber="98">340</superScript>
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and moderately broad (sag.), being just over 30% of cephalic length (sag.). Brim width ratio 0.70. Brim crossed by light caeca, with a high density of perforations in spaces in between (34 per sq. mm). Diameter of standardised brim perforations averaging just under 95 μm.
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</paragraph>
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||
<paragraph id="8BEB3331FFBEE341FF28F97FFE0386F8" blockId="97.[151,1437,150,1437]" pageId="97" pageNumber="98">Length of prolongations (exsag.) 88% of cephalic length (sag.). External rim narrow, dorsal surface convex. In dorsal view, external rim flares almost imperceptibly outwards as it crosses onto prolongation and then, slanting gently adaxially, continues in a straight line posteriorly for about two-thirds of length (exsag.) of prolongation, where it curves adaxially.</paragraph>
|
||
<paragraph id="8BEB3331FFBEE341FF28F9EFFC1485D8" blockId="97.[151,1437,150,1437]" pageId="97" pageNumber="98">
|
||
The internal rim, leaving posterior border slanting abaxially at an angle of around
|
||
<superScript id="7C219E79FFBEE341FB9FF9EFFB2186D1" attach="left" box="[1136,1172,402,429]" fontSize="6" pageId="97" pageNumber="98">400</superScript>
|
||
to axis, curves steadily adaxially until it meets external rim, where it is again at an angle of around
|
||
<superScript id="7C219E79FFBEE341FC0BF9CBFBBD86B5" attach="left" box="[996,1032,438,465]" fontSize="6" pageId="97" pageNumber="98">400</superScript>
|
||
to axis. In lateral view, external rim curves up distally to meet internal rim at an angle of around
|
||
<superScript id="7C219E79FFBEE341FCD1F9A7FCD78699" attach="none" box="[830,866,474,501]" fontSize="6" pageId="97" pageNumber="98">600</superScript>
|
||
. Internal rim straight in profile, slopes downwards gently from posterior border. Girder, on crossing onto prolongation, slopes sharply upwards, before curving posteriorly to join internal rim about one fifth of its way down prolongation (exsag.). Tubercles on dorsal surface of internal rim. Brim on prolongations quickly steepens posteriorly, to an angle to horizontal of around 80°. Perforations and caeca on brim of prolongations are same as on rest of brim, except for line of slightly bigger perforations which runs just below internal rim. Genal spine short and slanting adaxially.
|
||
</paragraph>
|
||
<paragraph id="8BEB3331FFBEE341FF28FACFFBCE8580" blockId="97.[151,1437,150,1437]" pageId="97" pageNumber="98">
|
||
Thorax has at least 24 segments. Interpleural furrows moderately deep. Distally, pleurae turn down and curve sharply forward, terminating with rounded end (
|
||
<figureCitation id="136F2FB4FFBEE341FD5CFAABFCBB8580" box="[691,782,727,753]" captionStart="PLATE 49" captionStartId="155.[151,234,1681,1705]" captionTargetBox="[236,1351,181,1658]" captionTargetId="figure-15@155.[236,1351,181,1659]" captionTargetPageId="155" captionText="PLATE 49. A–F Stoloharpes aff. rissaniensis, El Otfal formation, upper Eifelian, Ksar Hannabou (Fig. 2B, Map 6, site 14), dorsal exoskeleton, thorax disarticulated and pygidium missing, NHMUK It 29380. A, dorsal B, lateral and C, anterior view. D, dorsal and E, lateral view of eye lobe (uncoated). F, dorsal view of brim perforations. G–K Stoloharpes calceolus n. sp., “Akantharges Couche”, Eifelian, Ksar Lbor (Fig. 2B, Map 3, site 4), dorsal exoskeleton pygidium enrolled, holotype, NHMUK It 29363. G, dorsal, H, anterior and I, lateral view. J, dorsal view of brim perforations and K, lateral view of pleurae. L–P Stoloharpes aff. calceolus, “Akantharges Couche”, Eifelian, Ksar Lbor (Fig. 2B, Map 3, site 5), dorsal exoskeleton, thorax disarticulated pygidium enrolled, holotype, NHMUK It 29381. L, dorsal, M, anterior, N, lateral and O, posterior view. P, dorsal view of brim perforations. Scale bars shown represent 1 cm unless otherwise indicated." figureDoi="http://doi.org/10.5281/zenodo.11232687" httpUri="https://zenodo.org/record/11232687/files/figure.png" pageId="97" pageNumber="98">Pl. 49K</figureCitation>
|
||
). Pygidium enrolled and hidden.
|
||
</paragraph>
|
||
<paragraph id="8BEB3331FFBEE341FF28FA87FE7B8464" blockId="97.[151,1437,150,1437]" box="[199,462,763,789]" pageId="97" pageNumber="98">Hypostome not known.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34E60BAFFBEE341FF28FB63FC3F82EC" pageId="97" pageNumber="98" type="discussion">
|
||
<paragraph id="8BEB3331FFBEE341FF28FB63FAB08344" blockId="97.[151,1437,150,1437]" pageId="97" pageNumber="98">
|
||
<emphasis id="B920EF23FFBEE341FF28FB63FE8E8448" bold="true" box="[199,315,799,825]" pageId="97" pageNumber="98">Remarks.</emphasis>
|
||
<taxonomicName id="4C5448B2FFBEE341FEABFB63FD898448" box="[324,572,799,825]" class="Trilobita" family="Harpetidae" genus="Stoloharpes" kingdom="Animalia" order="Harpetida" pageId="97" pageNumber="98" phylum="Arthropoda" rank="species" species="calceolus">
|
||
<emphasis id="B920EF23FFBEE341FEABFB63FD898448" box="[324,572,799,825]" italics="true" pageId="97" pageNumber="98">Stoloharpes calceolus</emphasis>
|
||
</taxonomicName>
|
||
is similar to
|
||
<taxonomicName id="4C5448B2FFBEE341FD3BFB63FC128449" authorityName="Johnson" authorityYear="2024" box="[724,935,799,825]" class="Trilobita" family="Harpetidae" genus="Stoloharpes" isUncertain="true" kingdom="Animalia" order="Harpetida" pageId="97" pageNumber="98" phylum="Arthropoda" rank="species" species="capricornus">
|
||
<emphasis id="B920EF23FFBEE341FD3BFB63FD578448" box="[724,738,799,825]" italics="true" pageId="97" pageNumber="98">S</emphasis>
|
||
. aff.
|
||
<emphasis id="B920EF23FFBEE341FCCFFB63FC128449" box="[800,935,799,824]" italics="true" pageId="97" pageNumber="98">capricornus</emphasis>
|
||
</taxonomicName>
|
||
but differs in having: a cephalon that is not so strongly pyriform, and widest at the mid-point between the eye lobes and the alae not at the eye lobes; the genicranium widest at the eye lobes and not at the mid-point between the eye lobes and the alae; the flanks of the glabella more tectiform than rounded; the muscle insertion furrow at S1 slanting posteriorly, reaching two thirds of the way up the flank of the glabella, whereas the S1 furrow on
|
||
<taxonomicName id="4C5448B2FFBEE341FC8CFBD3FB8084B9" authorityName="Johnson" authorityYear="2024" box="[867,1077,943,969]" class="Trilobita" family="Harpetidae" genus="Stoloharpes" isUncertain="true" kingdom="Animalia" order="Harpetida" pageId="97" pageNumber="98" phylum="Arthropoda" rank="species" species="capricornus">
|
||
<emphasis id="B920EF23FFBEE341FC8CFBD3FCC484B8" box="[867,881,943,969]" italics="true" pageId="97" pageNumber="98">S</emphasis>
|
||
. aff.
|
||
<emphasis id="B920EF23FFBEE341FC41FBD3FB8084B9" box="[942,1077,943,968]" italics="true" pageId="97" pageNumber="98">capricornus</emphasis>
|
||
</taxonomicName>
|
||
is effaced; an anterior boss and light caeca reaching across brim, while the anterior boss of
|
||
<taxonomicName id="4C5448B2FFBEE341FCDAFBAFFBB0849D" authorityName="Johnson" authorityYear="2024" box="[821,1029,979,1005]" class="Trilobita" family="Harpetidae" genus="Stoloharpes" isUncertain="true" kingdom="Animalia" order="Harpetida" pageId="97" pageNumber="98" phylum="Arthropoda" rank="species" species="capricornus">
|
||
<emphasis id="B920EF23FFBEE341FCDAFBAFFCF6849C" box="[821,835,979,1005]" italics="true" pageId="97" pageNumber="98">S</emphasis>
|
||
. aff.
|
||
<emphasis id="B920EF23FFBEE341FC91FBAFFBB0849D" box="[894,1029,979,1004]" italics="true" pageId="97" pageNumber="98">capricornus</emphasis>
|
||
</taxonomicName>
|
||
is effaced and the brim lacks caeca; prolongations that are about 10% shorter in relation to cephalic length (sag.) than those of
|
||
<taxonomicName id="4C5448B2FFBEE341FB44FB8BFAD18360" box="[1195,1380,1015,1041]" class="Trilobita" family="Harpetidae" genus="Stoloharpes" isUncertain="true" kingdom="Animalia" order="Harpetida" pageId="97" pageNumber="98" phylum="Arthropoda" rank="species" species="calceolus">
|
||
<emphasis id="B920EF23FFBEE341FB44FB8BFB0C8360" box="[1195,1209,1015,1041]" italics="true" pageId="97" pageNumber="98">S</emphasis>
|
||
. aff.
|
||
<emphasis id="B920EF23FFBEE341FB17FB8BFAD18360" box="[1272,1380,1015,1041]" italics="true" pageId="97" pageNumber="98">calceolus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B920EF23FFBEE341FA8CFB8BFADF8360" box="[1379,1386,1015,1041]" italics="true" pageId="97" pageNumber="98">;</emphasis>
|
||
and shorter genal spines that are set at an oblique angle to the exterior rim rather than following its curve.
|
||
</paragraph>
|
||
<paragraph id="8BEB3331FFBEE341FF28FC43FAA3827C" blockId="97.[151,1437,150,1437]" pageId="97" pageNumber="98">
|
||
Other material, NHMUK It 299381 (
|
||
<figureCitation id="136F2FB4FFBEE341FD88FC43FD698328" box="[615,732,1087,1113]" captionStart="PLATE 49" captionStartId="155.[151,234,1681,1705]" captionTargetBox="[236,1351,181,1658]" captionTargetId="figure-15@155.[236,1351,181,1659]" captionTargetPageId="155" captionText="PLATE 49. A–F Stoloharpes aff. rissaniensis, El Otfal formation, upper Eifelian, Ksar Hannabou (Fig. 2B, Map 6, site 14), dorsal exoskeleton, thorax disarticulated and pygidium missing, NHMUK It 29380. A, dorsal B, lateral and C, anterior view. D, dorsal and E, lateral view of eye lobe (uncoated). F, dorsal view of brim perforations. G–K Stoloharpes calceolus n. sp., “Akantharges Couche”, Eifelian, Ksar Lbor (Fig. 2B, Map 3, site 4), dorsal exoskeleton pygidium enrolled, holotype, NHMUK It 29363. G, dorsal, H, anterior and I, lateral view. J, dorsal view of brim perforations and K, lateral view of pleurae. L–P Stoloharpes aff. calceolus, “Akantharges Couche”, Eifelian, Ksar Lbor (Fig. 2B, Map 3, site 5), dorsal exoskeleton, thorax disarticulated pygidium enrolled, holotype, NHMUK It 29381. L, dorsal, M, anterior, N, lateral and O, posterior view. P, dorsal view of brim perforations. Scale bars shown represent 1 cm unless otherwise indicated." figureDoi="http://doi.org/10.5281/zenodo.11232687" httpUri="https://zenodo.org/record/11232687/files/figure.png" pageId="97" pageNumber="98">Pl. 49L–P</figureCitation>
|
||
), is designated as
|
||
<taxonomicName id="4C5448B2FFBEE341FC46FC43FBE88328" box="[937,1117,1087,1113]" class="Trilobita" family="Harpetidae" genus="Stoloharpes" isUncertain="true" kingdom="Animalia" order="Harpetida" pageId="97" pageNumber="98" phylum="Arthropoda" rank="species" species="calceolus">
|
||
<emphasis id="B920EF23FFBEE341FC46FC43FC0B8328" box="[937,958,1087,1113]" italics="true" pageId="97" pageNumber="98">S.</emphasis>
|
||
aff.
|
||
<emphasis id="B920EF23FFBEE341FC1CFC43FBE88328" box="[1011,1117,1087,1113]" italics="true" pageId="97" pageNumber="98">calceolus</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="4C5448B2FFBEE341FB1DFC43FACC8328" box="[1266,1401,1087,1113]" class="Trilobita" family="Harpetidae" genus="Stoloharpes" kingdom="Animalia" order="Harpetida" pageId="97" pageNumber="98" phylum="Arthropoda" rank="species" species="calceolus">
|
||
<emphasis id="B920EF23FFBEE341FB1DFC43FACC8328" box="[1266,1401,1087,1113]" italics="true" pageId="97" pageNumber="98">S. calceolus</emphasis>
|
||
</taxonomicName>
|
||
by having: the widest point of the cephalon at the level of the eyes rather than halfway between the eye lobes and alae; a wider glabella and a L1 that is not inflated; a shallower and wider (sag.) occipital furrow that is medially straight; alae that are laterally rather than anterolaterally directed; a poorly defined eye ridge rather than none; preaxial furrows that do not converge anteriorly; an anterior boss that is less inflated and not joined to the glabella; a higher brim width ratio and a marginal band that is near vertical rather than sloping down under the cephalon.
|
||
</paragraph>
|
||
<paragraph id="8BEB3331FFBEE341FF28FD6BFC3F82EC" blockId="97.[151,1437,150,1437]" pageId="97" pageNumber="98">
|
||
The
|
||
<typeStatus id="54EF8D93FFBEE341FF15FD6BFEE98240" box="[250,348,1303,1329]" pageId="97" pageNumber="98" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C5448B2FFBEE341FE6EFD6BFDA58240" box="[385,528,1303,1329]" class="Trilobita" family="Harpetidae" genus="Stoloharpes" kingdom="Animalia" order="Harpetida" pageId="97" pageNumber="98" phylum="Arthropoda" rank="species" species="calceolus">
|
||
<emphasis id="B920EF23FFBEE341FE6EFD6BFDA58240" box="[385,528,1303,1329]" italics="true" pageId="97" pageNumber="98">M. calceolus</emphasis>
|
||
</taxonomicName>
|
||
is well preserved complete cephalon and thorax of a size that would indicate that they belonged to a well-developed holaspid. The cephalon is symmetrical and without any apparent pathological defects and has significant differences with
|
||
<taxonomicName id="4C5448B2FFBEE341FD7FFD23FCF28208" box="[656,839,1375,1401]" class="Trilobita" family="Harpetidae" genus="Stoloharpes" isUncertain="true" kingdom="Animalia" order="Harpetida" pageId="97" pageNumber="98" phylum="Arthropoda" rank="species" species="calceolus">
|
||
<emphasis id="B920EF23FFBEE341FD7FFD23FD2B8208" box="[656,670,1375,1401]" italics="true" pageId="97" pageNumber="98">S</emphasis>
|
||
. aff.
|
||
<emphasis id="B920EF23FFBEE341FD32FD23FCF28208" box="[733,839,1375,1401]" italics="true" pageId="97" pageNumber="98">calceolus</emphasis>
|
||
</taxonomicName>
|
||
, the species it is most similar too. On this basis the species has been erected although without the support of
|
||
<typeStatus id="54EF8D93FFBEE341FCF8FDFFFC3382EC" box="[791,902,1411,1437]" pageId="97" pageNumber="98" type="paratype">paratypes</typeStatus>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |