188 lines
34 KiB
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188 lines
34 KiB
XML
<document id="B23F5717670333D39B198056D24CA566" ID-CLB-Dataset="6005" ID-DOI="10.5852/ejt.2021.755.1397" ID-GBIF-Dataset="8721d89e-82a8-425a-8707-f53293bbef22" ID-ISSN="2118-9773" ID-Zenodo-Dep="4983284" ID-ZooBank="CEB00D62-0890-403C-8C54-968DC52C8290" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1624019007013" checkinUser="felipe" docAuthor="Moritz, Leif & Wesener, Thomas" docDate="2021" docId="03FC2A12FF8DFFA41A4DFAF3FE00FD7D" docLanguage="en" docName="ejt-755_22-46.pdf" docOrigin="European Journal of Taxonomy 755 (1)" docSource="http://dx.doi.org/10.5852/ejt.2021.755.1397" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Electrocambalidae Moritz & Wesener 2021, fam. nov." docType="treatment" docUuid="C3EA19DA-7BED-407F-BD71-6C17D048E129" docUuidSource="ZooBank" docVersion="8" lastPageNumber="26" masterDocId="FFC5526AFF8EFFA01827FF91FFE9FFB3" masterDocTitle="Electrocambalidae fam. nov., a new family of Cambalidea from Cretaceous Burmese amber (Diplopoda, Spirostreptida)" masterLastPageNumber="46" masterPageNumber="22" pageNumber="25" updateTime="1699017624614" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="CA40C633323C6359B93718834D41C7E4">Electrocambalidae fam. nov., a new family of Cambalidea from Cretaceous Burmese amber (Diplopoda, Spirostreptida)</mods:title>
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<mods:roleTerm id="6F25F8AE026EF5C0C9FF6B95827B9D06">Author</mods:roleTerm>
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<mods:namePart id="D766A0BBF6EF352D0235730A5387074C">Moritz, Leif</mods:namePart>
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<mods:affiliation id="B6DFC730BAB5AD41360BC37387054429">Zoological Research Museum Alexander Koenig (ZFMK), Leibniz Institute for Animal Biodiversity, Adenauerallee 160, D- 53113, Bonn, Germany. & Institute of Evolutionary Biology and Ecology, University of Bonn, An der Immenburg 1, 53121 Bonn, Germany.</mods:affiliation>
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<mods:nameIdentifier id="2DA0997197FEF4BAAF6B8D99DE956D37" type="email">moritz.leif@gmail.com</mods:nameIdentifier>
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<mods:namePart id="4EB8FF1B382093FD99AA9784AEF0B0B9">Wesener, Thomas</mods:namePart>
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<mods:affiliation id="5004ED24EB7D9325F84FC38B3D1AF768">Zoological Research Museum Alexander Koenig (ZFMK), Leibniz Institute for Animal Biodiversity, Adenauerallee 160, D- 53113, Bonn, Germany.</mods:affiliation>
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<mods:nameIdentifier id="6E0D6C3C09D8D6DBF41F03D3F350464E" type="email">t.wesener@leibniz-zfmk.de</mods:nameIdentifier>
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<mods:title id="93F059C6A2CF9E90410C82BB55C45484">European Journal of Taxonomy</mods:title>
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<mods:part id="99FA9254C21D01C39F2625AD28223151">
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<mods:date id="7FE5F8791724C30FFF4F1E2992709567">2021</mods:date>
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<mods:number id="87B48D6A6CDCED764447B6A69C56C382">2021-06-17</mods:number>
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<treatment id="03FC2A12FF8DFFA41A4DFAF3FE00FD7D" ID-DOI="http://doi.org/10.5281/zenodo.4985898" ID-GBIF-Taxon="182958549" ID-Zenodo-Dep="4985898" LSID="urn:lsid:zoobank.org:act:C3EA19DA-7BED-407F-BD71-6C17D048E129" httpUri="http://treatment.plazi.org/id/03FC2A12FF8DFFA41A4DFAF3FE00FD7D" lastPageId="4" lastPageNumber="26" pageId="3" pageNumber="25">
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<subSubSection id="C34FC88FFF8DFFA31A4DFAF3FC23FACE" box="[618,970,1378,1405]" pageId="3" pageNumber="25" type="nomenclature">
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<paragraph id="8BEA9B04FF8DFFA31A4DFAF3FC23FACE" blockId="3.[358,1229,1378,1477]" box="[618,970,1378,1405]" pageId="3" pageNumber="25">
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<taxonomicName id="4C55E087FF8DFFA31A4DFAF3FCBDFACF" ID-CoL="674c1a1a-8cf2-4bc4-b2da-5f47da557f00" authority="Moritz & Wesener, 2021" authorityName="Moritz & Wesener" authorityYear="2021" box="[618,852,1378,1404]" family="Electrocambalidae" kingdom="Animalia" order="Spirostreptida" pageId="3" pageNumber="25" rank="family" status="fam. nov.">
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<emphasis id="B9214716FF8DFFA31A4DFAF3FCBDFACF" bold="true" box="[618,852,1378,1404]" pageId="3" pageNumber="25">Electrocambalidae</emphasis>
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<taxonomicNameLabel id="A212FA6DFF8DFFA31B7BFAF2FC23FACE" box="[860,970,1379,1405]" pageId="3" pageNumber="25" rank="family">fam. nov.</taxonomicNameLabel>
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</paragraph>
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<subSubSection id="C34FC88FFF8DFFA31941FA17FCA4FA76" pageId="3" pageNumber="25" type="description">
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urn:lsid:zoobank.org:act:
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<uuid id="FFF3A1D1FF8DFFA31AA1FA17FB24FA13" box="[646,1229,1414,1440]" pageId="3" pageNumber="25">C3EA19DA-7BED-407F-BD71-6C17D048E129</uuid>
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</uri>
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</paragraph>
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<paragraph id="8BEA9B04FF8DFFA31AC0FA3BFCA4FA76" blockId="3.[358,1229,1378,1477]" box="[743,845,1450,1477]" pageId="3" pageNumber="25">
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<figureCitation id="136E8781FF8DFFA31AC0FA3BFCA4FA76" box="[743,845,1450,1477]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStart-4="Fig" captionStartId-0="5.[189,233,1787,1813]" captionStartId-1="6.[189,233,1964,1990]" captionStartId-2="9.[189,233,1822,1848]" captionStartId-3="12.[188,231,1787,1813]" captionStartId-4="14.[189,233,1822,1848]" captionTargetBox-0="[189,1398,632,1749]" captionTargetBox-1="[189,1398,1368,1924]" captionTargetBox-2="[189,1398,397,1772]" captionTargetBox-3="[188,1399,529,1730]" captionTargetBox-4="[195,1398,525,1784]" captionTargetId-0="figure-69@5.[189,1398,632,1749]" captionTargetId-1="figure-371@6.[189,1398,1368,1924]" captionTargetId-2="figure-54@9.[189,1398,397,1784]" captionTargetId-3="figure-117@12.[188,1399,529,1750]" captionTargetId-4="figure-34@14.[189,1398,525,1784]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionTargetPageId-2="9" captionTargetPageId-3="12" captionTargetPageId-4="14" captionText-0="Fig. 1. Electrocambala ornata gen. et sp. nov., holotype, ♂ (ZFMK-MYR7369). A. Habitus, lateral view, photograph. B. Head and anterior body rings, lateral view, photograph. C. Head and anterior body rings, lateral view, volume rendering. D. Head and anterior body rings, lateral view, drawing. E. Tarsus of midbody leg, photograph. F. Midbody ring, photograph. G. Gonopods, frontal view, segmentation. H. Gonopods, lateral view, segmentation. Abbreviations: ag = anterior gonopod; as = accessory spine; at = antennae; cl = claw; co = collum; cx = coxite; L = leg; la = labrum; md = mandible; mz = metazonite; om = ommatidia; pe = penis; pg = posterior gonopod; pz = prozonite; ta = tarsus; te = telopodite." captionText-1="Fig. 2. Electrocambala ornata gen. et sp. nov., other material, photographs. A. ♂ (BuB1962). B. ♀ (BuB1825). C. Paratype, ♀ (ZFMK MYR7370)." captionText-2="Fig. 3. Electrocambala cretacea gen. et sp. nov., holotype, ♀ (ZFMK-MYR7370). A. Amber piece, photograph. B. Habitus, lateral view, photograph. C. Head and anterior body rings, lateral view, photograph. D. Telson and posterior body rings, lateral view, photograph. E. Head and anterior body rings, lateral view, drawing. F. Antenna, drawing. G. Midbody leg, drawing. Abbreviations: 57 = body ring 57; as = apical spine; at = antennae; av = anal valve; cl = claw; co = collum; cx = coxa; il = incisura lateralis; L = leg; la = labrum; md = mandible; om = ommatidia; ta = tarsus; te = telson." captionText-3="Fig. 4. Kachincambala muelleri gen. et sp. nov., holotype, ♂ (ZFMK-MYR10225). A. Habitus, dorsal view, photograph. B. Head and anterior body rings, ventral view, volume rendering. C. Head and anterior body rings, lateral view, volume rendering. D. head, frontal view, volume rendering. E. Head, ventral view, volume rendering. F. Gonopods, frontal view, segmentation. G. Gonopods, lateral view, segmentation. Abbreviations: ag = anterior gonopod; at = antennae; co = collum; cx = coxite; em = eumentum; gn = gnathochilarium; L = leg; la = labrum; ll = lamella limgualis; md = mandible; om = ommatidia; pe = penis; pg = posterior gonopod; pm = promentum; st = gnathochilarial stipes; te = telopodite." captionText-4="Fig. 5. Kachincambala distorta gen. et sp. nov., holotype, ♂ (ZFMK-MYR7368). A. Amber piece, photograph. B. Head and anterior body rings, lateral view, photograph. C. Head and anterior body rings, lateral view, volume rendering. D. Head, lateral view, drawing. E. Tarsus, drawing. F. Gonopods, anterior view, segmentation. G. Gonopods, lateral view, segmentation. Abbreviations: ag = anterior gonopod; at = antennae; cl = claw; co = collum; cx = coxite; L = leg; la = labrum; md = mandible; om = ommatidia; oz = ozopore; pg = posterior gonopod; ta = tarsus; te = telopodite." figureDoi-0="http://doi.org/10.5281/zenodo.4983286" figureDoi-1="http://doi.org/10.5281/zenodo.4983289" figureDoi-2="http://doi.org/10.5281/zenodo.4983293" figureDoi-3="http://doi.org/10.5281/zenodo.4983295" figureDoi-4="http://doi.org/10.5281/zenodo.4983297" httpUri-0="https://zenodo.org/record/4983286/files/figure.png" httpUri-1="https://zenodo.org/record/4983289/files/figure.png" httpUri-2="https://zenodo.org/record/4983293/files/figure.png" httpUri-3="https://zenodo.org/record/4983295/files/figure.png" httpUri-4="https://zenodo.org/record/4983297/files/figure.png" pageId="3" pageNumber="25">Figs 1–5</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C34FC88FFF8DFFA3189AFA7DFE00F985" pageId="3" pageNumber="25" type="type_taxon">
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<paragraph id="8BEA9B04FF8DFFA3189AFA7DFEA3F9B5" blockId="3.[189,330,1516,1542]" box="[189,330,1516,1542]" pageId="3" pageNumber="25">
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<heading id="D0A22C68FF8DFFA3189AFA7DFEA3F9B5" bold="true" box="[189,330,1516,1542]" fontSize="11" level="3" pageId="3" pageNumber="25" reason="3">
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<emphasis id="B9214716FF8DFFA3189AFA7DFEA3F9B5" bold="true" box="[189,330,1516,1542]" pageId="3" pageNumber="25">
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<typeStatus id="54EE25A6FF8DFFA3189AFA7DFF13F9B5" box="[189,250,1516,1542]" pageId="3" pageNumber="25">Type</typeStatus>
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genus
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BEA9B04FF8DFFA3189AF98AFE00F985" blockId="3.[189,489,1563,1590]" box="[189,489,1563,1590]" pageId="3" pageNumber="25">
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<taxonomicName id="4C55E087FF8DFFA3189AF98AFE9EF986" authority="Moritz & Wesener, 2021" authorityName="Moritz & Wesener" authorityYear="2021" box="[189,375,1563,1589]" class="Diplopoda" family="Pseudonannolenidae" genus="Electrocambala" kingdom="Animalia" order="Spirostreptida" pageId="3" pageNumber="25" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B9214716FF8DFFA3189AF98AFE9EF986" box="[189,375,1563,1589]" italics="true" pageId="3" pageNumber="25">Electrocambala</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A212FA6DFF8DFFA31958F98DFE00F985" box="[383,489,1564,1590]" pageId="3" pageNumber="25" rank="genus">gen. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="C34FC88FFF8DFFA4189AF9CCFD5EFE0F" lastPageId="4" lastPageNumber="26" pageId="3" pageNumber="25" type="diagnosis">
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<paragraph id="8BEA9B04FF8DFFA3189AF9CCFEDFF9C4" blockId="3.[189,310,1629,1655]" box="[189,310,1629,1655]" pageId="3" pageNumber="25">
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<heading id="D0A22C68FF8DFFA3189AF9CCFEDFF9C4" bold="true" box="[189,310,1629,1655]" fontSize="11" level="3" pageId="3" pageNumber="25" reason="3">
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<emphasis id="B9214716FF8DFFA3189AF9CCFEDFF9C4" bold="true" box="[189,310,1629,1655]" pageId="3" pageNumber="25">Diagnosis</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BEA9B04FF8DFFA4189AF91DFD5EFE0F" blockId="3.[189,1399,1676,2025]" lastBlockId="4.[189,1399,265,444]" lastPageId="4" lastPageNumber="26" pageId="3" pageNumber="25">
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The new family
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<taxonomicName id="4C55E087FF8DFFA319A7F91DFDB3F915" authority="Moritz & Wesener, 2021" authorityName="Moritz & Wesener" authorityYear="2021" box="[384,602,1676,1702]" family="Electrocambalidae" kingdom="Animalia" order="Spirostreptida" pageId="3" pageNumber="25" rank="family" status="fam. nov.">Electrocambalidae</taxonomicName>
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<taxonomicNameLabel id="A212FA6DFF8DFFA31A45F91DFD39F914" box="[610,720,1676,1703]" pageId="3" pageNumber="25" rank="family">fam. nov.</taxonomicNameLabel>
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differs from the suborder
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<taxonomicName id="4C55E087FF8DFFA31C22F91DFB5EF915" baseAuthorityName="Jeekel" baseAuthorityYear="1985" box="[1029,1207,1676,1702]" class="Diplopoda" kingdom="Animalia" order="Spirobolida" pageId="3" pageNumber="25" phylum="Arthropoda" rank="subOrder" subOrder="Spirostreptidea">Spirostreptidea</taxonomicName>
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by the structure of the gnathochilarium, with a separate eumentum and promentum, as well as by the presence of two pairs of gonopods.
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<taxonomicName id="4C55E087FF8DFFA319BAF945FD9EF95D" authority="Moritz & Wesener, 2021" authorityName="Moritz & Wesener" authorityYear="2021" box="[413,631,1748,1774]" family="Electrocambalidae" kingdom="Animalia" order="Spirostreptida" pageId="3" pageNumber="25" rank="family" status="fam. nov.">Electrocambalidae</taxonomicName>
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<taxonomicNameLabel id="A212FA6DFF8DFFA31A59F945FD05F95D" box="[638,748,1748,1774]" pageId="3" pageNumber="25" rank="family">fam. nov.</taxonomicNameLabel>
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differs from the cambalidean families
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<taxonomicName id="4C55E087FF8DFFA31C96F945FAACF95D" authorityName="Chamberlain & Hoffman" authorityYear="1950" box="[1201,1349,1748,1774]" class="Diplopoda" family="Choctellidae" kingdom="Animalia" order="Spirostreptida" pageId="3" pageNumber="25" phylum="Arthropoda" rank="family">Choctellidae</taxonomicName>
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and
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<taxonomicName id="4C55E087FF8DFFA3189AF969FE44F8A1" authorityName="Silvestri" authorityYear="1895" box="[189,429,1784,1810]" class="Diplopoda" family="Pseudonannolenidae" kingdom="Animalia" order="Spirostreptida" pageId="3" pageNumber="25" phylum="Arthropoda" rank="family">Pseudonannolenidae</taxonomicName>
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by the presence of well-developed posterior gonopods (
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<figureCitation id="136E8781FF8DFFA31C16F969FB47F8A1" box="[1073,1198,1784,1810]" captionStart="Fig" captionStartId="5.[189,233,1787,1813]" captionTargetBox="[189,1398,632,1749]" captionTargetId="figure-69@5.[189,1398,632,1749]" captionTargetPageId="5" captionText="Fig. 1. Electrocambala ornata gen. et sp. nov., holotype, ♂ (ZFMK-MYR7369). A. Habitus, lateral view, photograph. B. Head and anterior body rings, lateral view, photograph. C. Head and anterior body rings, lateral view, volume rendering. D. Head and anterior body rings, lateral view, drawing. E. Tarsus of midbody leg, photograph. F. Midbody ring, photograph. G. Gonopods, frontal view, segmentation. H. Gonopods, lateral view, segmentation. Abbreviations: ag = anterior gonopod; as = accessory spine; at = antennae; cl = claw; co = collum; cx = coxite; L = leg; la = labrum; md = mandible; mz = metazonite; om = ommatidia; pe = penis; pg = posterior gonopod; pz = prozonite; ta = tarsus; te = telopodite." figureDoi="http://doi.org/10.5281/zenodo.4983286" httpUri="https://zenodo.org/record/4983286/files/figure.png" pageId="3" pageNumber="25">Fig. 1B, G</figureCitation>
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). The new family differs from the
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<taxonomicName id="4C55E087FF8DFFA31954F88AFDECF886" authorityName="Chamberlain & Hoffman" authorityYear="1950" box="[371,517,1819,1845]" class="Diplopoda" family="Choctellidae" kingdom="Animalia" order="Spirostreptida" pageId="3" pageNumber="25" phylum="Arthropoda" rank="family">Choctellidae</taxonomicName>
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and
|
||
<taxonomicName id="4C55E087FF8DFFA31A1CF88AFD0FF886" authorityName="Verhoeff" authorityYear="1924" box="[571,742,1819,1845]" class="Diplopoda" family="Iulomorphidae" kingdom="Animalia" order="Spirostreptida" pageId="3" pageNumber="25" phylum="Arthropoda" rank="family">Iulomorphidae</taxonomicName>
|
||
by having distinctly arched metazonites from body ring 6 onwards, and long body rings with a length/diameter ratio> 0.7 (
|
||
<figureCitation id="136E8781FF8DFFA31B87F8AEFBF5F8EA" box="[928,1052,1855,1881]" captionStart="Fig" captionStartId="5.[189,233,1787,1813]" captionTargetBox="[189,1398,632,1749]" captionTargetId="figure-69@5.[189,1398,632,1749]" captionTargetPageId="5" captionText="Fig. 1. Electrocambala ornata gen. et sp. nov., holotype, ♂ (ZFMK-MYR7369). A. Habitus, lateral view, photograph. B. Head and anterior body rings, lateral view, photograph. C. Head and anterior body rings, lateral view, volume rendering. D. Head and anterior body rings, lateral view, drawing. E. Tarsus of midbody leg, photograph. F. Midbody ring, photograph. G. Gonopods, frontal view, segmentation. H. Gonopods, lateral view, segmentation. Abbreviations: ag = anterior gonopod; as = accessory spine; at = antennae; cl = claw; co = collum; cx = coxite; L = leg; la = labrum; md = mandible; mz = metazonite; om = ommatidia; pe = penis; pg = posterior gonopod; pz = prozonite; ta = tarsus; te = telopodite." figureDoi="http://doi.org/10.5281/zenodo.4983286" httpUri="https://zenodo.org/record/4983286/files/figure.png" pageId="3" pageNumber="25">Fig. 1B–C</figureCitation>
|
||
).
|
||
<taxonomicName id="4C55E087FF8DFFA31C08F8AEFAEFF8EA" authority="Moritz & Wesener, 2021" authorityName="Moritz & Wesener" authorityYear="2021" box="[1071,1286,1855,1881]" family="Electrocambalidae" kingdom="Animalia" order="Spirostreptida" pageId="3" pageNumber="25" rank="family" status="fam. nov.">Electrocambalidae</taxonomicName>
|
||
<taxonomicNameLabel id="A212FA6DFF8DFFA31D2BF8AEFA9EF8E9" box="[1292,1399,1855,1882]" pageId="3" pageNumber="25" rank="family">fam. nov.</taxonomicNameLabel>
|
||
differs from the
|
||
<taxonomicName id="4C55E087FF8DFFA3195CF8F2FDC1F8CE" authorityName="Verhoeff" authorityYear="1924" box="[379,552,1891,1917]" class="Diplopoda" family="Iulomorphidae" kingdom="Animalia" order="Spirostreptida" pageId="3" pageNumber="25" phylum="Arthropoda" rank="family">Iulomorphidae</taxonomicName>
|
||
by the presence of a relatively large first leg pair in the males (
|
||
<figureCitation id="136E8781FF8DFFA31D28F8F2FA99F8CE" box="[1295,1392,1891,1917]" captionStart="Fig" captionStartId="5.[189,233,1787,1813]" captionTargetBox="[189,1398,632,1749]" captionTargetId="figure-69@5.[189,1398,632,1749]" captionTargetPageId="5" captionText="Fig. 1. Electrocambala ornata gen. et sp. nov., holotype, ♂ (ZFMK-MYR7369). A. Habitus, lateral view, photograph. B. Head and anterior body rings, lateral view, photograph. C. Head and anterior body rings, lateral view, volume rendering. D. Head and anterior body rings, lateral view, drawing. E. Tarsus of midbody leg, photograph. F. Midbody ring, photograph. G. Gonopods, frontal view, segmentation. H. Gonopods, lateral view, segmentation. Abbreviations: ag = anterior gonopod; as = accessory spine; at = antennae; cl = claw; co = collum; cx = coxite; L = leg; la = labrum; md = mandible; mz = metazonite; om = ommatidia; pe = penis; pg = posterior gonopod; pz = prozonite; ta = tarsus; te = telopodite." figureDoi="http://doi.org/10.5281/zenodo.4983286" httpUri="https://zenodo.org/record/4983286/files/figure.png" pageId="3" pageNumber="25">Figs 1D</figureCitation>
|
||
,
|
||
<figureCitation id="136E8781FF8DFFA3189AF816FF0BF812" box="[189,226,1927,1953]" captionStart="Fig" captionStartId="12.[188,231,1787,1813]" captionTargetBox="[188,1399,529,1730]" captionTargetId="figure-117@12.[188,1399,529,1750]" captionTargetPageId="12" captionText="Fig. 4. Kachincambala muelleri gen. et sp. nov., holotype, ♂ (ZFMK-MYR10225). A. Habitus, dorsal view, photograph. B. Head and anterior body rings, ventral view, volume rendering. C. Head and anterior body rings, lateral view, volume rendering. D. head, frontal view, volume rendering. E. Head, ventral view, volume rendering. F. Gonopods, frontal view, segmentation. G. Gonopods, lateral view, segmentation. Abbreviations: ag = anterior gonopod; at = antennae; co = collum; cx = coxite; em = eumentum; gn = gnathochilarium; L = leg; la = labrum; ll = lamella limgualis; md = mandible; om = ommatidia; pe = penis; pg = posterior gonopod; pm = promentum; st = gnathochilarial stipes; te = telopodite." figureDoi="http://doi.org/10.5281/zenodo.4983295" httpUri="https://zenodo.org/record/4983295/files/figure.png" pageId="3" pageNumber="25">4E</figureCitation>
|
||
). The new family differs from the families
|
||
<taxonomicName id="4C55E087FF8DFFA31AC8F816FC69F812" authorityName="Bollman" authorityYear="1893" box="[751,896,1927,1953]" class="Diplopoda" family="Cambalidae" kingdom="Animalia" order="Spirostreptida" pageId="3" pageNumber="25" phylum="Arthropoda" rank="family">Cambalidae</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C55E087FF8DFFA31BAAF816FBACF812" authorityName="Cook" authorityYear="1895" box="[909,1093,1927,1953]" class="Diplopoda" family="Cambalopsidae" kingdom="Animalia" order="Spirostreptida" pageId="3" pageNumber="25" phylum="Arthropoda" rank="family">Cambalopsidae</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C55E087FF8DFFA31CA5F816FA9EF812" authorityName="Silvestri" authorityYear="1895" box="[1154,1399,1927,1953]" class="Diplopoda" family="Pseudonannolenidae" kingdom="Animalia" order="Spirostreptida" pageId="3" pageNumber="25" phylum="Arthropoda" rank="family">Pseudonannolenidae</taxonomicName>
|
||
(in which ozopores start from body ring 5) by the ozopores starting from body ring 6 (
|
||
<figureCitation id="136E8781FF8DFFA31CF5F83BFADDF876" box="[1234,1332,1962,1989]" captionStart="Fig" captionStartId="5.[189,233,1787,1813]" captionTargetBox="[189,1398,632,1749]" captionTargetId="figure-69@5.[189,1398,632,1749]" captionTargetPageId="5" captionText="Fig. 1. Electrocambala ornata gen. et sp. nov., holotype, ♂ (ZFMK-MYR7369). A. Habitus, lateral view, photograph. B. Head and anterior body rings, lateral view, photograph. C. Head and anterior body rings, lateral view, volume rendering. D. Head and anterior body rings, lateral view, drawing. E. Tarsus of midbody leg, photograph. F. Midbody ring, photograph. G. Gonopods, frontal view, segmentation. H. Gonopods, lateral view, segmentation. Abbreviations: ag = anterior gonopod; as = accessory spine; at = antennae; cl = claw; co = collum; cx = coxite; L = leg; la = labrum; md = mandible; mz = metazonite; om = ommatidia; pe = penis; pg = posterior gonopod; pz = prozonite; ta = tarsus; te = telopodite." figureDoi="http://doi.org/10.5281/zenodo.4983286" httpUri="https://zenodo.org/record/4983286/files/figure.png" pageId="3" pageNumber="25">Figs 1B</figureCitation>
|
||
,
|
||
<figureCitation id="136E8781FF8DFFA31D64F83BFA80F876" box="[1347,1385,1962,1989]" captionStart="Fig" captionStartId="14.[189,233,1822,1848]" captionTargetBox="[195,1398,525,1784]" captionTargetId="figure-34@14.[189,1398,525,1784]" captionTargetPageId="14" captionText="Fig. 5. Kachincambala distorta gen. et sp. nov., holotype, ♂ (ZFMK-MYR7368). A. Amber piece, photograph. B. Head and anterior body rings, lateral view, photograph. C. Head and anterior body rings, lateral view, volume rendering. D. Head, lateral view, drawing. E. Tarsus, drawing. F. Gonopods, anterior view, segmentation. G. Gonopods, lateral view, segmentation. Abbreviations: ag = anterior gonopod; at = antennae; cl = claw; co = collum; cx = coxite; L = leg; la = labrum; md = mandible; om = ommatidia; oz = ozopore; pg = posterior gonopod; ta = tarsus; te = telopodite." figureDoi="http://doi.org/10.5281/zenodo.4983297" httpUri="https://zenodo.org/record/4983297/files/figure.png" pageId="3" pageNumber="25">5C</figureCitation>
|
||
).
|
||
<taxonomicName id="4C55E087FF8DFFA3189AF85FFE7FF85B" authority="Moritz & Wesener, 2021" authorityName="Moritz & Wesener" authorityYear="2021" box="[189,406,1998,2024]" family="Electrocambalidae" kingdom="Animalia" order="Spirostreptida" pageId="3" pageNumber="25" rank="family" status="fam. nov.">Electrocambalidae</taxonomicName>
|
||
<taxonomicNameLabel id="A212FA6DFF8DFFA319BBF85FFDE1F85A" box="[412,520,1998,2025]" pageId="3" pageNumber="25" rank="family">fam. nov.</taxonomicNameLabel>
|
||
differs from the
|
||
<taxonomicName id="4C55E087FF8DFFA31AE1F85FFCBBF85B" authorityName="Bollman" authorityYear="1893" box="[710,850,1998,2024]" class="Diplopoda" family="Cambalidae" kingdom="Animalia" order="Spirostreptida" pageId="3" pageNumber="25" phylum="Arthropoda" rank="family">Cambalidae</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C55E087FF8DFFA31BA0F85FFBD2F85B" authorityName="Cook" authorityYear="1895" box="[903,1083,1998,2024]" class="Diplopoda" family="Cambalopsidae" kingdom="Animalia" order="Spirostreptida" pageId="3" pageNumber="25" phylum="Arthropoda" rank="family">Cambalopsidae</taxonomicName>
|
||
by the absence of tubercles on the metazonites.
|
||
<taxonomicName id="4C55E087FF8AFFA4198BFE98FD6FFE90" authority="Moritz & Wesener, 2021" authorityName="Moritz & Wesener" authorityYear="2021" box="[428,646,265,291]" family="Electrocambalidae" kingdom="Animalia" order="Spirostreptida" pageId="4" pageNumber="26" rank="family" status="fam. nov.">Electrocambalidae</taxonomicName>
|
||
<taxonomicNameLabel id="A212FA6DFF8AFFA41AA8FE98FD17FE90" box="[655,766,265,291]" pageId="4" pageNumber="26" rank="family">fam. nov.</taxonomicNameLabel>
|
||
differs from other species of the order
|
||
<taxonomicName id="4C55E087FF8AFFA41CF5FE98FA9EFE90" authorityName="Brandt" authorityYear="1833" box="[1234,1399,265,291]" class="Diplopoda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirostreptida" pageId="4" pageNumber="26" phylum="Arthropoda" rank="order">Spirostreptida</taxonomicName>
|
||
in the combination of the following characters: (1) leg pair 3 moved anteriad, resulting in a seemingly legless body ring
|
||
<quantity id="4CAD36E1FF8AFFA419A1FEC4FE58FEDC" box="[390,433,341,367]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="4" pageNumber="26" unit="in" value="3.0">3 in</quantity>
|
||
addition to the legless body ring 4 (
|
||
<figureCitation id="136E8781FF8AFFA41B61FEC4FC74FEDC" box="[838,925,341,367]" captionStart="Fig" captionStartId="5.[189,233,1787,1813]" captionTargetBox="[189,1398,632,1749]" captionTargetId="figure-69@5.[189,1398,632,1749]" captionTargetPageId="5" captionText="Fig. 1. Electrocambala ornata gen. et sp. nov., holotype, ♂ (ZFMK-MYR7369). A. Habitus, lateral view, photograph. B. Head and anterior body rings, lateral view, photograph. C. Head and anterior body rings, lateral view, volume rendering. D. Head and anterior body rings, lateral view, drawing. E. Tarsus of midbody leg, photograph. F. Midbody ring, photograph. G. Gonopods, frontal view, segmentation. H. Gonopods, lateral view, segmentation. Abbreviations: ag = anterior gonopod; as = accessory spine; at = antennae; cl = claw; co = collum; cx = coxite; L = leg; la = labrum; md = mandible; mz = metazonite; om = ommatidia; pe = penis; pg = posterior gonopod; pz = prozonite; ta = tarsus; te = telopodite." figureDoi="http://doi.org/10.5281/zenodo.4983286" httpUri="https://zenodo.org/record/4983286/files/figure.png" pageId="4" pageNumber="26">Fig. 1B</figureCitation>
|
||
), (2) presence of setation on the posterior margin of the metazonites, and (3) pilosity on the head is not restricted to the labrum, but extends further up frontally on the head (
|
||
<figureCitation id="136E8781FF8AFFA419C2FE33FDACFE0F" box="[485,581,418,444]" captionStart="Fig" captionStartId="5.[189,233,1787,1813]" captionTargetBox="[189,1398,632,1749]" captionTargetId="figure-69@5.[189,1398,632,1749]" captionTargetPageId="5" captionText="Fig. 1. Electrocambala ornata gen. et sp. nov., holotype, ♂ (ZFMK-MYR7369). A. Habitus, lateral view, photograph. B. Head and anterior body rings, lateral view, photograph. C. Head and anterior body rings, lateral view, volume rendering. D. Head and anterior body rings, lateral view, drawing. E. Tarsus of midbody leg, photograph. F. Midbody ring, photograph. G. Gonopods, frontal view, segmentation. H. Gonopods, lateral view, segmentation. Abbreviations: ag = anterior gonopod; as = accessory spine; at = antennae; cl = claw; co = collum; cx = coxite; L = leg; la = labrum; md = mandible; mz = metazonite; om = ommatidia; pe = penis; pg = posterior gonopod; pz = prozonite; ta = tarsus; te = telopodite." figureDoi="http://doi.org/10.5281/zenodo.4983286" httpUri="https://zenodo.org/record/4983286/files/figure.png" pageId="4" pageNumber="26">Figs 1D</figureCitation>
|
||
,
|
||
<figureCitation id="136E8781FF8AFFA41A74FE33FD9FFE0F" box="[595,630,418,444]" captionStart="Fig" captionStartId="9.[189,233,1822,1848]" captionTargetBox="[189,1398,397,1772]" captionTargetId="figure-54@9.[189,1398,397,1784]" captionTargetPageId="9" captionText="Fig. 3. Electrocambala cretacea gen. et sp. nov., holotype, ♀ (ZFMK-MYR7370). A. Amber piece, photograph. B. Habitus, lateral view, photograph. C. Head and anterior body rings, lateral view, photograph. D. Telson and posterior body rings, lateral view, photograph. E. Head and anterior body rings, lateral view, drawing. F. Antenna, drawing. G. Midbody leg, drawing. Abbreviations: 57 = body ring 57; as = apical spine; at = antennae; av = anal valve; cl = claw; co = collum; cx = coxa; il = incisura lateralis; L = leg; la = labrum; md = mandible; om = ommatidia; ta = tarsus; te = telson." figureDoi="http://doi.org/10.5281/zenodo.4983293" httpUri="https://zenodo.org/record/4983293/files/figure.png" pageId="4" pageNumber="26">3E</figureCitation>
|
||
,
|
||
<figureCitation id="136E8781FF8AFFA41AA5FE30FD43FE0F" box="[642,682,417,444]" captionStart="Fig" captionStartId="14.[189,233,1822,1848]" captionTargetBox="[195,1398,525,1784]" captionTargetId="figure-34@14.[189,1398,525,1784]" captionTargetPageId="14" captionText="Fig. 5. Kachincambala distorta gen. et sp. nov., holotype, ♂ (ZFMK-MYR7368). A. Amber piece, photograph. B. Head and anterior body rings, lateral view, photograph. C. Head and anterior body rings, lateral view, volume rendering. D. Head, lateral view, drawing. E. Tarsus, drawing. F. Gonopods, anterior view, segmentation. G. Gonopods, lateral view, segmentation. Abbreviations: ag = anterior gonopod; at = antennae; cl = claw; co = collum; cx = coxite; L = leg; la = labrum; md = mandible; om = ommatidia; oz = ozopore; pg = posterior gonopod; ta = tarsus; te = telopodite." figureDoi="http://doi.org/10.5281/zenodo.4983297" httpUri="https://zenodo.org/record/4983297/files/figure.png" pageId="4" pageNumber="26">5D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34FC88FFF8AFFA4189AFE77FDB2FDEB" pageId="4" pageNumber="26" type="etymology">
|
||
<paragraph id="8BEA9B04FF8AFFA4189AFE77FEADFDB3" blockId="4.[189,324,486,512]" box="[189,324,486,512]" pageId="4" pageNumber="26">
|
||
<emphasis id="B9214716FF8AFFA4189AFE77FEADFDB3" bold="true" box="[189,324,486,512]" pageId="4" pageNumber="26">Etymology</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BEA9B04FF8AFFA4189AFD89FDB2FDEB" blockId="4.[189,1398,536,600]" pageId="4" pageNumber="26">
|
||
The family name is derived from the Latin word ʻ
|
||
<emphasis id="B9214716FF8AFFA41ADBFD89FC8AFD81" box="[764,867,536,562]" italics="true" pageId="4" pageNumber="26">electrum</emphasis>
|
||
ʼ meaning ʻamberʼ and the taxon
|
||
<taxonomicName id="4C55E087FF8AFFA41CC4FD89FA9BFD81" authorityName="Cook" authorityYear="1895" box="[1251,1394,536,562]" class="Diplopoda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirostreptida" pageId="4" pageNumber="41" phylum="Arthropoda" rank="subOrder" subOrder="Cambalidea">Cambalidea</taxonomicName>
|
||
, to which the studied fossils belong.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34FC88FFF8AFFA4189AFD13FE00FD7D" pageId="4" pageNumber="26" type="multiple">
|
||
<paragraph id="8BEA9B04FF8AFFA4189AFD13FE3EFD2F" blockId="4.[189,471,642,668]" box="[189,471,642,668]" pageId="4" pageNumber="26">
|
||
<emphasis id="B9214716FF8AFFA4189AFD13FE3EFD2F" bold="true" box="[189,471,642,668]" pageId="4" pageNumber="26">Other genera included</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BEA9B04FF8AFFA4189AFD25FE00FD7D" blockId="4.[189,489,692,718]" box="[189,489,692,718]" pageId="4" pageNumber="26">
|
||
<taxonomicName id="4C55E087FF8AFFA4189AFD25FE9EFD7D" authority="Moritz & Wesener, 2021" authorityName="Moritz & Wesener" authorityYear="2021" box="[189,375,692,718]" class="Diplopoda" family="Pseudonannolenidae" genus="Kachincambala" kingdom="Animalia" order="Spirostreptida" pageId="4" pageNumber="26" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B9214716FF8AFFA4189AFD25FE9EFD7D" box="[189,375,692,718]" italics="true" pageId="4" pageNumber="26">Kachincambala</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A212FA6DFF8AFFA41959FD25FE00FD7D" box="[382,489,692,718]" pageId="4" pageNumber="26" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |