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<document id="3F30F99A1CC4A325E1166D7CB176E097" ID-DOI="10.5281/zenodo.5170203" ID-ISSN="1942-1354" ID-Zenodo-Dep="5170203" ID-ZooBank="AB1A57F0-7CB4-4830-920B-DF219740A596" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628281492031" checkinUser="felipe" docAuthor="Blahnik, Roger J. &amp; Holzenthal, Ralph W." docDate="2017" docId="03F687A7FFBFF84FFF01BB664297F9CF" docLanguage="en" docName="InsectaMundi.0602.1-251.pdf" docOrigin="Insecta Mundi 2017 (602)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Mortoniella (Mortoniella) variabilis Blahnik &amp; Holzenthal 2017, new species" docType="treatment" docVersion="3" lastPageNumber="71" masterDocId="FFCFFFDFFFF8F807FFC1B865400BFFB4" masterDocTitle="Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *" masterLastPageNumber="251" masterPageNumber="1" pageNumber="70" updateTime="1699039969677" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
<mods:mods id="1839EE5BF41B845D0610E2E252056E22" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="AD37B3C65E45DF4AD993BDD7D6FBAF71">
<mods:title id="EF3936DE71D8A8E566AE3672F7B8EF72">Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *</mods:title>
</mods:titleInfo>
<mods:name id="538C13AFA2D8A157BDD6C7CF9935BD2D" type="personal">
<mods:role id="767051DC8E6258EEB6A523E5BA6C397F">
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<mods:namePart id="F9D51A133624EB8A273BC3B25E517630">Blahnik, Roger J.</mods:namePart>
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<mods:namePart id="6CCBB289BC9227A407507C8A45012682">Holzenthal, Ralph W.</mods:namePart>
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<mods:typeOfResource id="61BA1B314D1AE26BCE1E066296B026E4">text</mods:typeOfResource>
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<mods:title id="D516F93B59310D505D2A2DA5F3628229">Insecta Mundi</mods:title>
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<mods:part id="7412A19532923430D83BEDAFD6C47CC1">
<mods:date id="D8882DDD15EED5F083CFDDFA634742DE">2017</mods:date>
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<mods:number id="825A7764220E5BC162FD2FDE5EB3E6D9">2017-12-29</mods:number>
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<mods:detail id="06A482E8CAA49AD9677A78C5F57CB56E" type="volume">
<mods:number id="E68B15E3E1893E371EB978F731A16CA5">2017</mods:number>
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<mods:number id="1624C4FD988AD89230DD0CE0CECA68AA">602</mods:number>
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<mods:start id="BDABD99CCA9E64835C3909EBFF139FC9">1</mods:start>
<mods:end id="70175441D4631577C75ABC3543336A13">251</mods:end>
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<mods:classification id="803F704207B7581AF3AD1CCB8813CD89">journal article</mods:classification>
<mods:identifier id="B6614595A473BF744B3025F442E02595" type="DOI">10.5281/zenodo.5170203</mods:identifier>
<mods:identifier id="A4791D05440857F55A3CBC1C36A27EBD" type="ISSN">1942-1354</mods:identifier>
<mods:identifier id="8C9FC0190D29090C034D4D3C1B731727" type="Zenodo-Dep">5170203</mods:identifier>
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</mods:mods>
<treatment id="03F687A7FFBFF84FFF01BB664297F9CF" LSID="urn:lsid:plazi:treatment:03F687A7FFBFF84FFF01BB664297F9CF" httpUri="http://treatment.plazi.org/id/03F687A7FFBFF84FFF01BB664297F9CF" lastPageId="72" lastPageNumber="71" pageId="71" pageNumber="70">
<subSubSection id="C345653AFFBFF840FF01BB664361FCAF" box="[192,874,771,795]" pageId="71" pageNumber="70" type="nomenclature">
<paragraph id="8BE036B1FFBFF840FF01BB664361FCAF" blockId="71.[192,874,771,827]" box="[192,874,771,795]" pageId="71" pageNumber="70">
<heading id="D0A881DDFFBFF840FF01BB664361FCAF" bold="true" box="[192,874,771,795]" fontSize="10" level="2" pageId="71" pageNumber="70" reason="3">
<emphasis id="B92BEAA3FFBFF840FF01BB664361FCAF" bold="true" box="[192,874,771,795]" pageId="71" pageNumber="70">
<taxonomicName id="4C5F4D32FFBFF840FF01BB6642B8FCAF" authority="Blahnik &amp; Holzenthal, 2017" authorityName="Blahnik &amp; Holzenthal" authorityYear="2017" box="[192,691,771,795]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="71" pageNumber="70" phylum="Tracheophyta" rank="species" species="variabilis" status="sp. nov." subGenus="Mortoniella">
<emphasis id="B92BEAA3FFBFF840FF01BB664168FCAF" bold="true" box="[192,355,771,795]" italics="true" pageId="71" pageNumber="70">Mortoniella</emphasis>
(
<emphasis id="B92BEAA3FFBFF840FEB7BB664212FCAF" bold="true" box="[374,537,771,795]" italics="true" pageId="71" pageNumber="70">Mortoniella</emphasis>
)
<emphasis id="B92BEAA3FFBFF840FDEABB6642B8FCAF" bold="true" box="[555,691,771,795]" italics="true" pageId="71" pageNumber="70">variabilis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A21857D8FFBFF840FD02BB664361FCAF" box="[707,874,771,795]" pageId="71" pageNumber="70" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C345653AFFBFF840FF01BB46411DFC8F" box="[192,278,803,827]" pageId="71" pageNumber="70" type="description">
<paragraph id="8BE036B1FFBFF840FF01BB46411DFC8F" blockId="71.[192,874,771,827]" box="[192,278,803,827]" pageId="71" pageNumber="70">
<figureCitation id="13642A34FFBFF840FF01BB46411DFC8F" box="[192,278,803,827]" captionStart="Figure 55" captionStartId="195.[192,276,1865,1887]" captionTargetBox="[345,1290,260,1744]" captionTargetId="figure-47@195.[345,1291,260,1758]" captionTargetPageId="195" captionText="Figure 55. Mortoniella (Mortoniella) variabilis, new species male genitalia. A—lateral; B—segment IX and tergum X, dorsal; C—phallic ensemble, ventral; D—lateral, specimen from Colombia." figureDoi="http://doi.org/10.5281/zenodo.5170317" httpUri="https://zenodo.org/record/5170317/files/figure.png" pageId="71" pageNumber="70">Fig. 55</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C345653AFFBFF84FFF31BB064297F9CF" lastPageId="72" lastPageNumber="71" pageId="71" pageNumber="70" type="materials_examined">
<paragraph id="8BE036B1FFBFF840FF31BB064148F98F" blockId="71.[192,1441,867,1595]" pageId="71" pageNumber="70">
<taxonomicName id="4C5F4D32FFBFF840FF31BB064207FCCF" authorityName="Blahnik &amp; Holzenthal" authorityYear="2017" box="[240,524,867,891]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="71" pageNumber="70" phylum="Tracheophyta" rank="species" species="variabilis">
<emphasis id="B92BEAA3FFBFF840FF31BB064207FCCF" box="[240,524,867,891]" italics="true" pageId="71" pageNumber="70">Mortoniella variabilis</emphasis>
</taxonomicName>
is very similar to
<taxonomicName id="4C5F4D32FFBFF840FD38BB01438EFCCF" authority="Blahnik &amp; Holzenthal, 2017" authorityName="Blahnik &amp; Holzenthal" authorityYear="2017" box="[761,901,867,891]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="71" pageNumber="70" phylum="Tracheophyta" rank="species" species="gracilis" status="sp. nov.">
<emphasis id="B92BEAA3FFBFF840FD38BB01438EFCCF" box="[761,901,867,891]" italics="true" pageId="71" pageNumber="70">M. gracilis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A21857D8FFBFF840FC54BB0143D0FCCF" box="[917,987,868,891]" pageId="71" pageNumber="70" rank="species">n. sp.</taxonomicNameLabel>
and
<taxonomicName id="4C5F4D32FFBFF840FBDDBB01450BFCCF" authority="(Flint)" baseAuthorityName="Flint" baseAuthorityYear="1981" box="[1052,1280,867,891]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="71" pageNumber="70" phylum="Tracheophyta" rank="species" species="limona">
<emphasis id="B92BEAA3FFBFF840FBDDBB0144AAFCCF" box="[1052,1185,867,891]" italics="true" pageId="71" pageNumber="70">M. limona</emphasis>
(Flint)
</taxonomicName>
. All of these species have elongate basoventral projections from the phallicata, a distinguishing feature of this complex. The shape of these projections is variable in each of these species, but in
<taxonomicName id="4C5F4D32FFBFF840FB39BBC145ABFC0F" authorityName="Blahnik &amp; Holzenthal" authorityYear="2017" box="[1272,1440,931,955]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="71" pageNumber="70" phylum="Tracheophyta" rank="species" species="variabilis">
<emphasis id="B92BEAA3FFBFF840FB39BBC145ABFC0F" box="[1272,1440,931,955]" italics="true" pageId="71" pageNumber="70">M. variabilis</emphasis>
</taxonomicName>
the projections tend to be narrower and longer than in
<taxonomicName id="4C5F4D32FFBFF840FC54BBA1442AFC6F" authorityName="Blahnik &amp; Holzenthal" authorityYear="2017" box="[917,1057,963,987]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="71" pageNumber="70" phylum="Tracheophyta" rank="species" species="gracilis">
<emphasis id="B92BEAA3FFBFF840FC54BBA1442AFC6F" box="[917,1057,963,987]" italics="true" pageId="71" pageNumber="70">M. gracilis</emphasis>
</taxonomicName>
. The most diagnostic feature distinguishing the species in this complex is the shape of the apex of the dorsal phallic spine. Unfortunately, this is somewhat variable, even within the species, and this is especially so for
<taxonomicName id="4C5F4D32FFBFF840FA46BC614137FB8F" authorityName="Blahnik &amp; Holzenthal" authorityYear="2017" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="71" pageNumber="70" phylum="Tracheophyta" rank="species" species="variabilis">
<emphasis id="B92BEAA3FFBFF840FA46BC614137FB8F" italics="true" pageId="71" pageNumber="70">M variabilis</emphasis>
</taxonomicName>
. In general, the ventral projection at the apex of the dorsal phallic spine is less produced in
<taxonomicName id="4C5F4D32FFBFF840FF22BC214181FBEF" authorityName="Blahnik &amp; Holzenthal" authorityYear="2017" box="[227,394,1091,1115]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="71" pageNumber="70" phylum="Tracheophyta" rank="species" species="variabilis">
<emphasis id="B92BEAA3FFBFF840FF22BC214181FBEF" box="[227,394,1091,1115]" italics="true" pageId="71" pageNumber="70">M. variabilis</emphasis>
</taxonomicName>
than in
<taxonomicName id="4C5F4D32FFBFF840FE3ABC214274FBEF" baseAuthorityName="Flint" baseAuthorityYear="1981" box="[507,639,1091,1115]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="71" pageNumber="70" phylum="Tracheophyta" rank="species" species="limona">
<emphasis id="B92BEAA3FFBFF840FE3ABC214274FBEF" box="[507,639,1091,1115]" italics="true" pageId="71" pageNumber="70">M. limona</emphasis>
</taxonomicName>
, but more so than in
<taxonomicName id="4C5F4D32FFBFF840FC58BC21442FFBEF" authorityName="Blahnik &amp; Holzenthal" authorityYear="2017" box="[921,1060,1091,1115]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="71" pageNumber="70" phylum="Tracheophyta" rank="species" species="gracilis">
<emphasis id="B92BEAA3FFBFF840FC58BC21442FFBEF" box="[921,1060,1091,1115]" italics="true" pageId="71" pageNumber="70">M. gracilis</emphasis>
</taxonomicName>
. The ventral projection often (or perhaps usually) has a distinct, upturned, spine-like projection, not present in
<taxonomicName id="4C5F4D32FFBFF840FB1CBC01456BFBCF" baseAuthorityName="Flint" baseAuthorityYear="1981" box="[1245,1376,1123,1147]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="71" pageNumber="70" phylum="Tracheophyta" rank="species" species="limona">
<emphasis id="B92BEAA3FFBFF840FB1CBC01456BFBCF" box="[1245,1376,1123,1147]" italics="true" pageId="71" pageNumber="70">M. limona</emphasis>
</taxonomicName>
. The development of this spine is quite variable in the material examined, as is the length of the paramere appendages (see
<figureCitation id="13642A34FFBFF840FDE1BCC64284FB0F" box="[544,655,1187,1211]" captionStart="Figure 55" captionStartId="195.[192,276,1865,1887]" captionTargetBox="[345,1290,260,1744]" captionTargetId="figure-47@195.[345,1291,260,1758]" captionTargetPageId="195" captionText="Figure 55. Mortoniella (Mortoniella) variabilis, new species male genitalia. A—lateral; B—segment IX and tergum X, dorsal; C—phallic ensemble, ventral; D—lateral, specimen from Colombia." figureDoi="http://doi.org/10.5281/zenodo.5170317" httpUri="https://zenodo.org/record/5170317/files/figure.png" pageId="71" pageNumber="70">Fig. 55D</figureCitation>
for an extreme variant from
<collectingCountry id="F3487621FFBFF840FBCABCC64482FB0F" box="[1035,1161,1187,1211]" name="Colombia" pageId="71" pageNumber="70">Colombia</collectingCountry>
). We are interpreting this as variation within a species, but the issue may be worthy of reinvestigation when more material is available. There was no evident difference in the female genitalia of specimens from
<collectingCountry id="F3487621FFBFF840FF01BD664132FAAF" box="[192,313,1283,1307]" name="Colombia" pageId="71" pageNumber="70">Colombia</collectingCountry>
and
<collectingCountry id="F3487621FFBFF840FEBCBD66420CFAAF" box="[381,519,1283,1307]" name="Venezuela" pageId="71" pageNumber="70">Venezuela</collectingCountry>
.
<taxonomicName id="4C5F4D32FFBFF840FDD4BD664338FAAF" authorityName="Blahnik &amp; Holzenthal" authorityYear="2017" box="[533,819,1283,1307]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="71" pageNumber="70" phylum="Tracheophyta" rank="species" species="variabilis">
<emphasis id="B92BEAA3FFBFF840FDD4BD664338FAAF" box="[533,819,1283,1307]" italics="true" pageId="71" pageNumber="70">Mortoniella variabilis</emphasis>
</taxonomicName>
is probably more likely to be confused with
<taxonomicName id="4C5F4D32FFBFF840FABEBD614111FA8F" baseAuthorityName="Flint" baseAuthorityYear="1981" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="71" pageNumber="70" phylum="Tracheophyta" rank="species" species="limona">
<emphasis id="B92BEAA3FFBFF840FABEBD614111FA8F" italics="true" pageId="71" pageNumber="70">M. limona</emphasis>
</taxonomicName>
than
<taxonomicName id="4C5F4D32FFBFF840FEADBD4141F2FA8F" authorityName="Blahnik &amp; Holzenthal" authorityYear="2017" box="[364,505,1315,1339]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="71" pageNumber="70" phylum="Tracheophyta" rank="species" species="gracilis">
<emphasis id="B92BEAA3FFBFF840FEADBD4141F2FA8F" box="[364,505,1315,1339]" italics="true" pageId="71" pageNumber="70">M. gracilis</emphasis>
</taxonomicName>
, because the ventral projection of the dorsal phallic spine is distinctly produced in both species, but the character state in
<taxonomicName id="4C5F4D32FFBFF840FC47BD21443AFAEF" authorityName="Blahnik &amp; Holzenthal" authorityYear="2017" box="[902,1073,1347,1371]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="71" pageNumber="70" phylum="Tracheophyta" rank="species" species="variabilis">
<emphasis id="B92BEAA3FFBFF840FC47BD21443AFAEF" box="[902,1073,1347,1371]" italics="true" pageId="71" pageNumber="70">M. variabilis</emphasis>
</taxonomicName>
never reaches the extreme condition found in
<taxonomicName id="4C5F4D32FFBFF840FE69BD014223FACF" baseAuthorityName="Flint" baseAuthorityYear="1981" box="[424,552,1379,1403]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="71" pageNumber="70" phylum="Tracheophyta" rank="species" species="limona">
<emphasis id="B92BEAA3FFBFF840FE69BD014223FACF" box="[424,552,1379,1403]" italics="true" pageId="71" pageNumber="70">M. limona</emphasis>
</taxonomicName>
. Also, the apex of the dorsal phallic spine, above the ventral expansion, tends to be narrower and more attenuate in
<taxonomicName id="4C5F4D32FFBFF840FD3FBDE143AFFA2F" authorityName="Blahnik &amp; Holzenthal" authorityYear="2017" box="[766,932,1411,1435]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="71" pageNumber="70" phylum="Tracheophyta" rank="species" species="variabilis">
<emphasis id="B92BEAA3FFBFF840FD3FBDE143AFFA2F" box="[766,932,1411,1435]" italics="true" pageId="71" pageNumber="70">M. variabilis</emphasis>
</taxonomicName>
than in
<taxonomicName id="4C5F4D32FFBFF840FBD2BDE1449CFA2F" baseAuthorityName="Flint" baseAuthorityYear="1981" box="[1043,1175,1411,1435]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="71" pageNumber="70" phylum="Tracheophyta" rank="species" species="limona">
<emphasis id="B92BEAA3FFBFF840FBD2BDE1449CFA2F" box="[1043,1175,1411,1435]" italics="true" pageId="71" pageNumber="70">M. limona</emphasis>
</taxonomicName>
. In most material of
<taxonomicName id="4C5F4D32FFBFF840FF01BDC14162FA0F" authorityName="Blahnik &amp; Holzenthal" authorityYear="2017" box="[192,361,1443,1467]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="71" pageNumber="70" phylum="Tracheophyta" rank="species" species="variabilis">
<emphasis id="B92BEAA3FFBFF840FF01BDC14162FA0F" box="[192,361,1443,1467]" italics="true" pageId="71" pageNumber="70">M. variabilis</emphasis>
</taxonomicName>
, only 1 pair of endophallic spines is evident, but in specimens with the armature fully expanded, a second, very lightly sclerotized pair may also be evident; the second pair may be a consistent feature, not easily observed unless the armature is completely everted. In the other two species, 2 pairs of sclerotized spines are generally evident, even when the armature is not expanded.
</paragraph>
<paragraph id="8BE036B1FFBFF840FF31BE0641B8F94F" blockId="71.[192,1440,1635,1916]" pageId="71" pageNumber="70">
<emphasis id="B92BEAA3FFBFF840FF31BE064133F9CF" box="[240,312,1635,1659]" italics="true" pageId="71" pageNumber="70">Adult</emphasis>
—Length of forewing: male
<quantity id="4CA79B54FFBFF840FD5ABE064326F9CF" box="[667,813,1635,1659]" metricMagnitude="-3" metricUnit="m" metricValue="3.05" metricValueMax="3.4" metricValueMin="2.7" pageId="71" pageNumber="70" unit="mm" value="3.05" valueMax="3.4" valueMin="2.7">2.7-3.4 mm</quantity>
; female 3.0-3.9. Forewing with forks
<collectionCode id="ED4EAE74FFBFF840FAC6BE01451FF9CF" box="[1287,1300,1636,1659]" country="Romania" lsid="urn:lsid:biocol.org:col:14415" name="&amp;quot;Alexandru Ioan Cuza&amp;quot; University" pageId="71" pageNumber="70" type="Herbarium">I</collectionCode>
,
<collectionCode id="ED4EAE74FFBFF840FAE0BE014531F9CF" box="[1313,1338,1636,1659]" pageId="71" pageNumber="70">II</collectionCode>
, and
<collectionCode id="ED4EAE74FFBFF840FABFBE0145ABF9CF" box="[1406,1440,1636,1659]" pageId="71" pageNumber="70">III</collectionCode>
present, hind wing with forks
<collectionCode id="ED4EAE74FFBFF840FD87BEE14257F92F" box="[582,604,1668,1691]" pageId="71" pageNumber="70">II</collectionCode>
and
<collectionCode id="ED4EAE74FFBFF840FD5CBEE142C8F92F" box="[669,707,1668,1691]" pageId="71" pageNumber="70">III</collectionCode>
. Spur formula 0:4:4. Overall color medium brown, apices of tarsal segments and basal segments of antennae pale brown. Tibial spurs darker than legs, contrasting in color. Wing bar at anastamosis indistinct, marked with light brown setae, most evident at arculus.
</paragraph>
<paragraph id="8BE036B1FFBFF84FFF31BF664218FD6F" blockId="71.[192,1440,1635,1916]" lastBlockId="72.[192,1440,194,1179]" lastPageId="72" lastPageNumber="71" pageId="71" pageNumber="70">
<emphasis id="B92BEAA3FFBFF840FF31BF6641ADF8AF" box="[240,422,1795,1819]" italics="true" pageId="71" pageNumber="70">Male genitalia</emphasis>
—Ventral process of segment
<collectionCode id="ED4EAE74FFBFF840FCD8BF614333F8AF" box="[793,824,1796,1819]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="71" pageNumber="70">VI</collectionCode>
laterally compressed, ventrally projecting, large, subtriangular, length about equal to width at base, process not retracted anterobasally. Segment IX nearly evenly rounded anterolaterally, length greatest midlaterally, posterolateral margin weakly rounded, narrowing ventrally; segment deeply mesally excised dorsally and ventrally, forming lateral lobes, separated dorsomesally by less than ½ width of segment. Tergum X relatively short, lateral margins subparallel, apicomesal margin with V-shaped or U-shaped emargination between apicolateral lobes, extending about 1/3 length of tergum; apicolateral lobes tapered, acute apically, short and narrow as viewed laterally; apicomesal lobes visible in lateral view, rounded apically; ventrolateral lobes distinct, projecting. Inferior appendages somewhat variable shape, with short tapering dorsolateral lobes and apicoventral projections very short or absent. Mesal pockets of inferior appendage with apical processes moderately large, posterodorsally curved. Paramere appendage short or moderate in length (shorter than dorsal phallic spine), narrow, nearly uniformly in width, apex acute. Dorsal phallic, as viewed laterally, compressed apically, with prominent rounded apicoventral expansion and (usually) an acute, dorsally-oriented projection from ventral margin; apex of dorsal phallic spine narrowing and attenuate, often slightly posteriorly curved; spine, in dorsal view, nearly uniform in width, apex acute. Phallicata with dorsal margin sclerotized, extending into endophallic membrane, with apical depression to accommodate apicoventral projection of dorsal phallic spine, basoventrally with pair of very prominent curved, apically rounded, sclerotized, projections. Endophallic membrane short, with one pair of prominent sclerotized spines, and second pair of smaller, lightly sclerotized spines (possibly sometimes absent); ventromesal spine absent.
</paragraph>
<paragraph id="8BE036B1FFB0F84FFF31BA8645ABFCAF" blockId="72.[192,1440,194,1179]" pageId="72" pageNumber="71">
<materialsCitation id="3B373CECFFB0F84FFF31BA8645ABFCAF" collectingDate="1994-01-15" collectionCode="UMSP" country="Venezuela" elevation="360" latitude="10.051" location="Rio Negro in Toromo" longLatPrecision="77" longitude="-72.712" municipality="Parque Nacional Perija" pageId="72" pageNumber="71" specimenCode="UMSP000001386" specimenCount="1" stateProvince="Zulia" typeStatus="holotype">
<emphasis id="B92BEAA3FFB0F84FFF31BA8641BCFD4F" bold="true" box="[240,439,739,763]" italics="true" pageId="72" pageNumber="71">
<typeStatus id="54E48813FFB0F84FFF31BA864160FD4F" box="[240,363,739,763]" pageId="72" pageNumber="71" type="holotype">Holotype</typeStatus>
male
</emphasis>
(pinned)—
<emphasis id="B92BEAA3FFB0F84FFD83BA864357FD4F" bold="true" box="[578,860,739,763]" pageId="72" pageNumber="71">
<collectingCountry id="F3487621FFB0F84FFD83BA8642F4FD4F" box="[578,767,739,763]" name="Venezuela" pageId="72" pageNumber="71">VENEZUELA</collectingCountry>
:
<collectingRegion id="499BF853FFB0F84FFCCABA86435DFD4F" box="[779,854,739,763]" country="Venezuela" name="Zulia" pageId="72" pageNumber="71">Zulia</collectingRegion>
:
</emphasis>
<collectingMunicipality id="6B84ACCBFFB0F84FFCA3BA814480FD4F" box="[866,1163,739,763]" pageId="72" pageNumber="71">Parque Nacional Perijá</collectingMunicipality>
,
<location id="8E80606AFFB0F84FFB57BA864597FD4F" LSID="urn:lsid:plazi:treatment:03F687A7FFBFF84FFF01BB664297F9CF:8E80606AFFB0F84FFB57BA864597FD4F" box="[1174,1436,739,764]" country="Venezuela" latitude="10.051" longLatPrecision="77" longitude="-72.712" municipality="Parque Nacional Perija" name="Rio Negro in Toromo" pageId="72" pageNumber="71" stateProvince="Zulia">Río Negro in Toromo</location>
,
<geoCoordinate id="EE6B5076FFB0F84FFF01BB66414BFCAF" box="[192,320,771,795]" degrees="10.051" direction="north" orientation="latitude" pageId="72" pageNumber="71" precision="55" value="10.051">10.051° N</geoCoordinate>
,
<geoCoordinate id="EE6B5076FFB0F84FFE8DBB6641DAFCAF" box="[332,465,771,795]" degrees="72.712" direction="west" orientation="longitude" pageId="72" pageNumber="71" precision="55" value="-72.712">72.712° W</geoCoordinate>
, el
<quantity id="4CA79B54FFB0F84FFE32BB66424FFCAF" box="[499,580,771,795]" metricMagnitude="2" metricUnit="m" metricValue="3.6" pageId="72" pageNumber="71" unit="m" value="360.0">
<elevation id="0072D182FFB0F84FFE32BB66424FFCAF" box="[499,580,771,795]" metricMagnitude="2" metricUnit="m" metricValue="3.6" pageId="72" pageNumber="71" unit="m" value="360.0">360 m</elevation>
</quantity>
,
<date id="FFE11071FFB0F84FFD91BB6642C3FCAF" box="[592,712,771,795]" pageId="72" pageNumber="71" value="1994-01-15">
<collectingDate id="EFA5E999FFB0F84FFD91BB6642C3FCAF" box="[592,712,771,795]" pageId="72" pageNumber="71" value="1994-01-15">15.i.1994</collectingDate>
</date>
, Holzenthal, Cressa, Rincón (
<specimenCode id="DBF99ECAFFB0F84FFB81BB664529FCAF" box="[1088,1314,771,795]" collectionCode="UMSP" country="USA" httpUri="http://grbio.org/cool/3avb-bd3e" name="University of Minnesota Insect Collection" pageId="72" pageNumber="71" type="University or college">UMSP000001386</specimenCode>
) (
<collectionCode id="ED4EAE74FFB0F84FFAFBBB66459FFCAF" box="[1338,1428,771,795]" country="USA" httpUri="http://grbio.org/cool/3avb-bd3e" name="University of Minnesota Insect Collection" pageId="72" pageNumber="71" type="University or college">UMSP</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BE036B1FFB0F84FFF31BB4643F3FBEF" blockId="72.[192,1440,194,1179]" pageId="72" pageNumber="71">
<materialsCitation id="3B373CECFFB0F84FFF31BB464381FCCF" collectingDate="1997-12-12" collectionCode="F, UMSP" country="Colombia" county="Municipio Cienaga" elevation="930" latitude="11.03944" location="Sierra Nevada de Santa Marta" longLatPrecision="1" longitude="-74.03833" municipality="Rio" pageId="72" pageNumber="71" specimenCount="19" specimenCount-female="14" specimenCount-male="5" stateProvince="Magdalena" typeStatus="paratype">
<emphasis id="B92BEAA3FFB0F84FFF31BB4642FDFC8F" bold="true" box="[240,758,803,827]" pageId="72" pageNumber="71">
<emphasis id="B92BEAA3FFB0F84FFF31BB464174FC8E" bold="true" box="[240,383,803,826]" italics="true" pageId="72" pageNumber="71">
<typeStatus id="54E48813FFB0F84FFF31BB464174FC8E" box="[240,383,803,826]" pageId="72" pageNumber="71" type="paratype">Paratypes</typeStatus>
</emphasis>
—
<collectingCountry id="F3487621FFB0F84FFE5BBB46424FFC8F" box="[410,580,803,827]" name="Colombia" pageId="72" pageNumber="71">COLOMBIA</collectingCountry>
:
<collectingRegion id="499BF853FFB0F84FFD90BB4642FBFC8F" box="[593,752,803,827]" country="Colombia" name="Magdalena" pageId="72" pageNumber="71">Magdalena</collectingRegion>
:
</emphasis>
<collectingCounty id="62814E3DFFB0F84FFD3DBB4643E4FC8F" box="[764,1007,803,827]" pageId="72" pageNumber="71">Municipio Ciénaga</collectingCounty>
,
<collectingMunicipality id="6B84ACCBFFB0F84FFC3ABB46442EFC8F" box="[1019,1061,803,827]" pageId="72" pageNumber="71">Río</collectingMunicipality>
<collectingRegion id="499BF853FFB0F84FFBEDBB464491FC8F" box="[1068,1178,803,827]" country="Colombia" name="Cordoba" pageId="72" pageNumber="71">Cordoba</collectingRegion>
,
<quantity id="4CA79B54FFB0F84FFB64BB4644F9FC8F" box="[1189,1266,803,827]" metricMagnitude="4" metricUnit="m" metricValue="2.5" pageId="72" pageNumber="71" unit="km" value="25.0">25 km</quantity>
NS “Estacion Exp. San Lorenzo,”
<location id="8E80606AFFB0F84FFE74BB264333FCEF" LSID="urn:lsid:plazi:treatment:03F687A7FFBFF84FFF01BB664297F9CF:8E80606AFFB0F84FFE74BB264333FCEF" box="[437,824,835,859]" country="Colombia" county="Municipio Cienaga" latitude="11.03944" longLatPrecision="1" longitude="-74.03833" municipality="Rio" name="Sierra Nevada de Santa Marta" pageId="72" pageNumber="71" stateProvince="Magdalena">Sierra Nevada de Santa Marta</location>
,
<geoCoordinate id="EE6B5076FFB0F84FFC82BB2643EBFCEF" box="[835,992,835,859]" degrees="11.03944" direction="north" orientation="latitude" pageId="72" pageNumber="71" precision="1" value="11.03944">11.03944° N</geoCoordinate>
,
<geoCoordinate id="EE6B5076FFB0F84FFC2ABB264486FCEF" box="[1003,1165,835,859]" degrees="74.03833" direction="west" orientation="longitude" pageId="72" pageNumber="71" precision="1" value="-74.03833">74.03833° W</geoCoordinate>
, el
<quantity id="4CA79B54FFB0F84FFB75BB264509FCEF" box="[1204,1282,835,859]" metricMagnitude="2" metricUnit="m" metricValue="9.3" pageId="72" pageNumber="71" unit="m" value="930.0">
<elevation id="0072D182FFB0F84FFB75BB264509FCEF" box="[1204,1282,835,859]" metricMagnitude="2" metricUnit="m" metricValue="9.3" pageId="72" pageNumber="71" unit="m" value="930.0">930 m</elevation>
</quantity>
,
<date id="FFE11071FFB0F84FFACCBB264590FCEF" box="[1293,1435,835,859]" pageId="72" pageNumber="71" value="1997-12-12">
<collectingDate id="EFA5E999FFB0F84FFACCBB264590FCEF" box="[1293,1435,835,859]" pageId="72" pageNumber="71" value="1997-12-12">12.xii.1997</collectingDate>
</date>
,
<collectionCode id="ED4EAE74FFB0F84FFF01BB0140D9FCCF" box="[192,210,868,891]" country="USA" lsid="urn:lsid:biocol.org:col:15707" name="Field Museum of Natural History, Botany Department" pageId="72" pageNumber="71" type="Herbarium">F</collectionCode>
Muñoz-Q et al.–
<specimenCount id="9D59FD38FFB0F84FFE69BB064205FCCF" box="[424,526,867,891]" pageId="72" pageNumber="71" type="male">5 males</specimenCount>
,
<specimenCount id="9D59FD38FFB0F84FFDDBBB0642AFFCCF" box="[538,676,867,891]" pageId="72" pageNumber="71" type="female">14 females</specimenCount>
(pinned) (
<collectionCode id="ED4EAE74FFB0F84FFCE9BB06438AFCCF" box="[808,897,867,891]" country="USA" httpUri="http://grbio.org/cool/3avb-bd3e" name="University of Minnesota Insect Collection" pageId="72" pageNumber="71" type="University or college">UMSP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B373CECFFB0F84FFC57BB064120FC0F" collectingDate="1997-12-09" collectionCode="F, UMSP" country="Colombia" elevation="570" latitude="11.14306" location="Rio Minca en Minca" longLatPrecision="1" longitude="-74.11611" municipality="Municipio de Santa Marta" pageId="72" pageNumber="71" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Magdalena" typeStatus="paratype">
<collectingMunicipality id="6B84ACCBFFB0F84FFC57BB0644E0FCCF" box="[918,1259,867,891]" pageId="72" pageNumber="71">Municipio de Santa Marta</collectingMunicipality>
,
<location id="8E80606AFFB0F84FFB39BB064119FC2F" LSID="urn:lsid:plazi:treatment:03F687A7FFBFF84FFF01BB664297F9CF:8E80606AFFB0F84FFB39BB064119FC2F" country="Colombia" latitude="11.14306" longLatPrecision="1" longitude="-74.11611" municipality="Municipio de Santa Marta" name="Rio Minca en Minca" pageId="72" pageNumber="71" stateProvince="Magdalena">Río Minca en Minca</location>
,
<geoCoordinate id="EE6B5076FFB0F84FFEDCBBE641B0FC2F" box="[285,443,899,923]" degrees="11.14306" direction="north" orientation="latitude" pageId="72" pageNumber="71" precision="1" value="11.14306">11.14306° N</geoCoordinate>
,
<geoCoordinate id="EE6B5076FFB0F84FFE06BBE64261FC2F" box="[455,618,899,923]" degrees="74.11611" direction="west" orientation="longitude" pageId="72" pageNumber="71" precision="1" value="-74.11611">74.11611° W</geoCoordinate>
, el
<quantity id="4CA79B54FFB0F84FFD50BBE642EAFC2F" box="[657,737,899,923]" metricMagnitude="2" metricUnit="m" metricValue="5.7" pageId="72" pageNumber="71" unit="m" value="570.0">
<elevation id="0072D182FFB0F84FFD50BBE642EAFC2F" box="[657,737,899,923]" metricMagnitude="2" metricUnit="m" metricValue="5.7" pageId="72" pageNumber="71" unit="m" value="570.0">570 m</elevation>
</quantity>
,
<date id="FFE11071FFB0F84FFD2DBBE64360FC2F" box="[748,875,899,923]" pageId="72" pageNumber="71" value="1997-12-09">
<collectingDate id="EFA5E999FFB0F84FFD2DBBE64360FC2F" box="[748,875,899,923]" pageId="72" pageNumber="71" value="1997-12-09">9.xii.1997</collectingDate>
</date>
,
<collectionCode id="ED4EAE74FFB0F84FFCB9BBE14386FC2F" box="[888,909,900,923]" country="USA" lsid="urn:lsid:biocol.org:col:15707" name="Field Museum of Natural History, Botany Department" pageId="72" pageNumber="71" type="Herbarium">F</collectionCode>
. Muñoz-Q et al.–
<specimenCount id="9D59FD38FFB0F84FFBA4BBE644B3FC2F" box="[1125,1208,899,923]" pageId="72" pageNumber="71" type="male">1 male</specimenCount>
,
<specimenCount id="9D59FD38FFB0F84FFB05BBE64525FC2F" box="[1220,1326,899,923]" pageId="72" pageNumber="71" type="female">1 female</specimenCount>
(pinned) (
<collectionCode id="ED4EAE74FFB0F84FFF08BBC64129FC0F" box="[201,290,931,955]" country="USA" httpUri="http://grbio.org/cool/3avb-bd3e" name="University of Minnesota Insect Collection" pageId="72" pageNumber="71" type="University or college">UMSP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B373CECFFB0F84FFEFABBC64138FC6F" collectingDate="1983-02-11" collectionCode="W, OS, NMNH" collectorName="Flint" country="Colombia" location="Jr" municipality="Quebrada Blanca" pageId="72" pageNumber="71" specimenCount="1" specimenCount-male="1" stateProvince="Meta" typeStatus="paratype">
<emphasis id="B92BEAA3FFB0F84FFEFABBC64181FC0F" bold="true" box="[315,394,931,955]" pageId="72" pageNumber="71">
<collectingRegion id="499BF853FFB0F84FFEFABBC6418EFC0F" box="[315,389,931,955]" country="Colombia" name="Meta" pageId="72" pageNumber="71">Meta</collectingRegion>
:
</emphasis>
<collectingMunicipality id="6B84ACCBFFB0F84FFE57BBC64276FC0F" box="[406,637,931,955]" pageId="72" pageNumber="71">Quebrada Blanca</collectingMunicipality>
,
<locationDeviation id="9A522F93FFB0F84FFD4FBBC64373FC0F" box="[654,888,931,955]" location="Restrepo" pageId="72" pageNumber="71">
<quantity id="4CA79B54FFB0F84FFD4FBBC642DBFC0F" box="[654,720,931,955]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="72" pageNumber="71" unit="km" value="3.0">3 km</quantity>
<collectionCode id="ED4EAE74FFB0F84FFD1CBBC142FCFC0F" box="[733,759,932,955]" country="Austria" lsid="urn:lsid:biocol.org:col:15588" name="Naturhistorisches Museum Wien" pageId="72" pageNumber="71" type="Herbarium">W</collectionCode>
Restrepo
</locationDeviation>
,
<date id="FFE11071FFB0F84FFC48BBC64401FC0F" box="[905,1034,931,955]" pageId="72" pageNumber="71" value="1983-02-11">
<collectingDate id="EFA5E999FFB0F84FFC48BBC64401FC0F" box="[905,1034,931,955]" pageId="72" pageNumber="71" value="1983-02-11">11.ii.1983</collectingDate>
</date>
,
<collectionCode id="ED4EAE74FFB0F84FFBDDBBC64449FC0F" box="[1052,1090,931,955]" country="USA" name="Oregon State University" pageId="72" pageNumber="71">OS</collectionCode>
<collectorName id="26AA5367FFB0F84FFB8FBBC6449AFC0F" box="[1102,1169,931,955]" pageId="72" pageNumber="71">Flint</collectorName>
,
<location id="8E80606AFFB0F84FFB63BBC644B5FC0F" LSID="urn:lsid:plazi:treatment:03F687A7FFBFF84FFF01BB664297F9CF:8E80606AFFB0F84FFB63BBC644B5FC0F" box="[1186,1214,931,955]" country="Colombia" municipality="Quebrada Blanca" name="Jr" pageId="72" pageNumber="71" stateProvince="Meta">Jr</location>
–
<specimenCount id="9D59FD38FFB0F84FFB10BBC6452DFC0F" box="[1233,1318,931,955]" pageId="72" pageNumber="71" type="male">1 male</specimenCount>
(alcohol) (
<collectionCode id="ED4EAE74FFB0F84FFF09BBA64120FC6F" box="[200,299,963,987]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="72" pageNumber="71" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B373CECFFB0F84FFEFEBBA64121FBAF" collectingDate="1997-03-21" collectionCode="UMSP" country="Venezuela" county="Parque Nacional Sierra Nevada" elevation="1000" latitude="8.4645" location="Holzenthal" longLatPrecision="1" longitude="-70.84867" municipality="Quebrada San Juan in Santa Rosa" pageId="72" pageNumber="71" specimenCount="7" specimenCount-female="5" specimenCount-male="2" stateProvince="Barinas" typeStatus="paratype">
<emphasis id="B92BEAA3FFB0F84FFEFEBBA6428FFC6F" bold="true" box="[319,644,963,987]" pageId="72" pageNumber="71">
<collectingCountry id="F3487621FFB0F84FFEFEBBA641F5FC6F" box="[319,510,963,987]" name="Venezuela" pageId="72" pageNumber="71">VENEZUELA</collectingCountry>
:
<collectingRegion id="499BF853FFB0F84FFDCABBA64275FC6F" box="[523,638,963,987]" country="Venezuela" name="Barinas" pageId="72" pageNumber="71">Barinas</collectingRegion>
:
</emphasis>
<collectingCounty id="62814E3DFFB0F84FFD4ABBA14428FC6F" box="[651,1059,963,987]" pageId="72" pageNumber="71">Parque Nacional Sierra Nevada</collectingCounty>
,
<collectingMunicipality id="6B84ACCBFFB0F84FFBEEBBA6410AFC4F" pageId="72" pageNumber="71">Quebrada San Juan in Santa Rosa</collectingMunicipality>
,
<geoCoordinate id="EE6B5076FFB0F84FFED1BB8641AFFC4F" box="[272,420,995,1019]" degrees="8.46450" direction="north" orientation="latitude" pageId="72" pageNumber="71" precision="1" value="8.4645">8.46450° N</geoCoordinate>
,
<geoCoordinate id="EE6B5076FFB0F84FFE72BB864251FC4F" box="[435,602,995,1019]" degrees="70.84867" direction="west" orientation="longitude" pageId="72" pageNumber="71" precision="1" value="-70.84867">70.84867° W</geoCoordinate>
, el
<quantity id="4CA79B54FFB0F84FFD4BBB8642E6FC4F" box="[650,749,995,1019]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="72" pageNumber="71" unit="m" value="1000.0">
<elevation id="0072D182FFB0F84FFD4BBB8642E6FC4F" box="[650,749,995,1019]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="72" pageNumber="71" unit="m" value="1000.0">1000 m</elevation>
</quantity>
,
<date id="FFE11071FFB0F84FFD3DBB86438CFC4F" box="[764,903,995,1019]" pageId="72" pageNumber="71" value="1997-03-21">
<collectingDate id="EFA5E999FFB0F84FFD3DBB86438CFC4F" box="[764,903,995,1019]" pageId="72" pageNumber="71" value="1997-03-21">21.iii.1997</collectingDate>
</date>
,
<location id="8E80606AFFB0F84FFC57BB86442FFC4F" LSID="urn:lsid:plazi:treatment:03F687A7FFBFF84FFF01BB664297F9CF:8E80606AFFB0F84FFC57BB86442FFC4F" box="[918,1060,995,1019]" country="Venezuela" county="Parque Nacional Sierra Nevada" latitude="8.4645" longLatPrecision="1" longitude="-70.84867" municipality="Quebrada San Juan in Santa Rosa" name="Holzenthal" pageId="72" pageNumber="71" stateProvince="Barinas">Holzenthal</location>
–
<specimenCount id="9D59FD38FFB0F84FFBF4BB864496FC4F" box="[1077,1181,995,1019]" pageId="72" pageNumber="71" type="male">2 males</specimenCount>
,
<specimenCount id="9D59FD38FFB0F84FFB6DBB864523FC4F" box="[1196,1320,995,1019]" pageId="72" pageNumber="71" type="female">5 females</specimenCount>
(alcohol) (
<collectionCode id="ED4EAE74FFB0F84FFF08BC664129FBAF" box="[201,290,1027,1051]" country="USA" httpUri="http://grbio.org/cool/3avb-bd3e" name="University of Minnesota Insect Collection" pageId="72" pageNumber="71" type="University or college">UMSP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B373CECFFB0F84FFEF9BC664590FBAF" box="[312,1435,1027,1051]" collectingDate="1976-02-24" collectionCode="N, CM, OS, NMNH" collectorName="Flint &amp; Jr" country="Venezuela" location="22 km N Barinitas" pageId="72" pageNumber="71" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Barinas" typeStatus="paratype">
<location id="8E80606AFFB0F84FFEF9BC664220FBAF" LSID="urn:lsid:plazi:treatment:03F687A7FFBFF84FFF01BB664297F9CF:8E80606AFFB0F84FFEF9BC664220FBAF" box="[312,555,1027,1051]" country="Venezuela" name="22 km N Barinitas" pageId="72" pageNumber="71" stateProvince="Barinas">
<quantity id="4CA79B54FFB0F84FFEF9BC66418CFBAF" box="[312,391,1027,1051]" metricMagnitude="4" metricUnit="m" metricValue="2.2" pageId="72" pageNumber="71" unit="km" value="22.0">22 km</quantity>
<collectionCode id="ED4EAE74FFB0F84FFE51BC6141ADFBAF" box="[400,422,1028,1051]" country="China" lsid="urn:lsid:biocol.org:col:13092" name="Nanjing University" pageId="72" pageNumber="71" type="Herbarium">N</collectionCode>
Barinitas
</location>
,
<date id="FFE11071FFB0F84FFDF8BC6642B1FBAF" box="[569,698,1027,1051]" pageId="72" pageNumber="71" value="1976-02-24">
<collectingDate id="EFA5E999FFB0F84FFDF8BC6642B1FBAF" box="[569,698,1027,1051]" pageId="72" pageNumber="71" value="1976-02-24">24.ii.1976</collectingDate>
</date>
,
<collectionCode id="ED4EAE74FFB0F84FFD09BC6642FEFBAF" box="[712,757,1027,1051]" country="China" name="Chongqing Museum" pageId="72" pageNumber="71">CM</collectionCode>
&amp;
<collectionCode id="ED4EAE74FFB0F84FFCDCBC664348FBAF" box="[797,835,1027,1051]" country="USA" name="Oregon State University" pageId="72" pageNumber="71">OS</collectionCode>
<collectorName id="26AA5367FFB0F84FFC8DBC664384FBAF" box="[844,911,1027,1051]" pageId="72" pageNumber="71">Flint</collectorName>
,
<collectorName id="26AA5367FFB0F84FFC5DBC6643B3FBAF" box="[924,952,1027,1051]" pageId="72" pageNumber="71">Jr</collectorName>
–
<specimenCount id="9D59FD38FFB0F84FFC0DBC66442AFBAF" box="[972,1057,1027,1051]" pageId="72" pageNumber="71" type="male">1 male</specimenCount>
,
<specimenCount id="9D59FD38FFB0F84FFBEFBC6644A2FBAF" box="[1070,1193,1027,1051]" pageId="72" pageNumber="71" type="female">2 females</specimenCount>
(alcohol) (
<collectionCode id="ED4EAE74FFB0F84FFAF1BC664598FBAF" box="[1328,1427,1027,1051]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="72" pageNumber="71" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B373CECFFB0F84FFF01BC4643FFFBEF" collectingDate="1994-01-15" collectionCode="UMSP, NMNH, MIZA" country="Venezuela" elevation="360" latitude="10.051" location="Rio Negro in Toromo" longLatPrecision="77" longitude="-72.712" municipality="Parque Nacional Perija" pageId="72" pageNumber="71" specimenCount="15" specimenCount-female="6" specimenCount-male="9" stateProvince="Zulia" typeStatus="paratype">
<emphasis id="B92BEAA3FFB0F84FFF01BC464119FB8F" bold="true" box="[192,274,1059,1083]" pageId="72" pageNumber="71">
<collectingRegion id="499BF853FFB0F84FFF01BC464107FB8F" box="[192,268,1059,1083]" country="Venezuela" name="Zulia" pageId="72" pageNumber="71">Zulia</collectingRegion>
:
</emphasis>
same data as Holotype–
<specimenCount id="9D59FD38FFB0F84FFD90BC4642BCFB8F" box="[593,695,1059,1083]" pageId="72" pageNumber="71" type="male">2 males</specimenCount>
,
<specimenCount id="9D59FD38FFB0F84FFD05BC464334FB8F" box="[708,831,1059,1083]" pageId="72" pageNumber="71" type="female">2 females</specimenCount>
(pinned),
<specimenCount id="9D59FD38FFB0F84FFC05BC46442DFB8F" box="[964,1062,1059,1083]" pageId="72" pageNumber="71" type="male">3 males</specimenCount>
(alcohol) (
<collectionCode id="ED4EAE74FFB0F84FFB6FBC46450CFB8F" box="[1198,1287,1059,1083]" country="USA" httpUri="http://grbio.org/cool/3avb-bd3e" name="University of Minnesota Insect Collection" pageId="72" pageNumber="71" type="University or college">UMSP</collectionCode>
),
<specimenCount id="9D59FD38FFB0F84FFADCBC46458FFB8F" box="[1309,1412,1059,1083]" pageId="72" pageNumber="71" type="male">2 males</specimenCount>
,
<specimenCount id="9D59FD38FFB0F84FFA50BC464128FBEF" pageId="72" pageNumber="71" type="female">2 females</specimenCount>
(alcohol) (
<collectionCode id="ED4EAE74FFB0F84FFE6ABC264205FBEF" box="[427,526,1091,1115]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="72" pageNumber="71" type="Museum">NMNH</collectionCode>
),
<specimenCount id="9D59FD38FFB0F84FFDE5BC264280FBEF" box="[548,651,1091,1115]" pageId="72" pageNumber="71" type="male">2 males</specimenCount>
,
<specimenCount id="9D59FD38FFB0F84FFD59BC264318FBEF" box="[664,787,1091,1115]" pageId="72" pageNumber="71" type="female">2 females</specimenCount>
(alcohol) (
<collectionCode id="ED4EAE74FFB0F84FFC5ABC2643E7FBEF" box="[923,1004,1091,1115]" country="Venezuela" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33834" lsid="urn:lsid:biocol.org:col:33834" name="Museo del Instituto de Zoologia Agricola Francisco Fernandez Yepez" pageId="72" pageNumber="71" type="Museum">MIZA</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BE036B1FFB0F84FFF31BC064491FB2F" blockId="72.[192,1440,194,1179]" pageId="72" pageNumber="71">
<emphasis id="B92BEAA3FFB0F84FFF31BC06417DFBCF" box="[240,374,1123,1147]" italics="true" pageId="72" pageNumber="71">Etymology</emphasis>
—This species is named
<taxonomicName id="4C5F4D32FFB0F84FFD74BC014357FBCF" authorityName="Blahnik &amp; Holzenthal" authorityYear="2017" box="[693,860,1123,1147]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="72" pageNumber="71" phylum="Tracheophyta" rank="species" species="variabilis">
<emphasis id="B92BEAA3FFB0F84FFD74BC014357FBCF" box="[693,860,1123,1147]" italics="true" pageId="72" pageNumber="71">M. variabilis</emphasis>
</taxonomicName>
because of the variability in the structure of the apex of the dorsal phallic spine and length of the paramere appendages.
</paragraph>
<paragraph id="8BE036B1FFB0F84FFF01BC8141B5FB4F" blockId="72.[192,446,1251,1275]" box="[192,446,1251,1275]" pageId="72" pageNumber="71">
—
<emphasis id="B92BEAA3FFB0F84FFF1ABC8641B5FB4F" bold="true" box="[219,446,1251,1275]" pageId="72" pageNumber="71">pocita subgroup</emphasis>
</paragraph>
<paragraph id="8BE036B1FFB0F84FFF01BD4642D4FA8F" blockId="72.[192,735,1315,1339]" box="[192,735,1315,1339]" pageId="72" pageNumber="71">
Included species:
<taxonomicName id="4C5F4D32FFB0F84FFE5DBD4642D0FA8F" authority="(Flint)" baseAuthorityName="Flint" baseAuthorityYear="1983" box="[412,731,1315,1339]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="72" pageNumber="71" phylum="Tracheophyta" rank="species" species="pocita">
<emphasis id="B92BEAA3FFB0F84FFE5DBD464274FA8F" box="[412,639,1315,1339]" italics="true" pageId="72" pageNumber="71">Mortoniella pocita</emphasis>
(Flint)
</taxonomicName>
.
</paragraph>
<paragraph id="8BE036B1FFB0F84FFF31BD064297F9CF" blockId="72.[192,1440,1379,1659]" pageId="72" pageNumber="71">
This subgroup was proposed by
<bibRefCitation id="EFCE4B40FFB0F84FFDBEBD06440EFACF" author="Blahnik, R. J. &amp; R. W. Holzenthal" box="[639,1029,1379,1403]" pageId="72" pageNumber="71" pagination="1 - 74" refId="ref106296" refString="Blahnik, R. J., and R. W. Holzenthal. 2011. Revision of the austral South American species of Mortoniella (Trichoptera: Glossosomatidae: Protoptilinae). Zootaxa 2852: 1 - 74." type="journal article" year="2011">Blahnik and Holzenthal (2011)</bibRefCitation>
to contain only
<taxonomicName id="4C5F4D32FFB0F84FFB0ABD014597FACF" authority="(Flint)" baseAuthorityName="Flint" baseAuthorityYear="1983" box="[1227,1436,1379,1403]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="72" pageNumber="71" phylum="Tracheophyta" rank="species" species="pocita">
<emphasis id="B92BEAA3FFB0F84FFB0ABD01454BFACF" box="[1227,1344,1380,1403]" italics="true" pageId="72" pageNumber="71">M. pocita</emphasis>
(Flint)
</taxonomicName>
. The species was originally described from
<collectingCountry id="F3487621FFB0F84FFD3ABDE64375FA2F" box="[763,894,1411,1435]" name="Argentina" pageId="72" pageNumber="71">Argentina</collectingCountry>
and subsequently listed from
<collectingCountry id="F3487621FFB0F84FFADABDE6457DFA2F" box="[1307,1398,1411,1435]" name="Bolivia" pageId="72" pageNumber="71">Bolivia</collectingCountry>
by
<bibRefCitation id="EFCE4B40FFB0F84FFF01BDC6426BFA0F" author="Rueda Martin, P. A. &amp; F. Gibon" box="[192,608,1443,1467]" pageId="72" pageNumber="71" pagination="215 - 225" refId="ref107162" refString="Rueda Martin, P. A., and F. Gibon. 2008. New species of Glossosomatidae from Bolivia with new records from Bolivia and northwestern Argentina (Trichoptera; Glossosomatidae). Annales de limnologie [= International Journal of Limnology] 44 (3): 215 - 225." type="journal article" year="2008">Rueda Martín and Gibon (2008)</bibRefCitation>
. It has a general similarity to several of the species listed below as leroda group species “unplaced to subgroup,” especially in having elongate inferior appendages and a modified apex of the dorsal phallic spine. However, none of these species have elongate dorsal processes on the phallicata, a character found in
<taxonomicName id="4C5F4D32FFB0F84FFC71BE614420F9AF" baseAuthorityName="Flint" baseAuthorityYear="1983" box="[944,1067,1540,1563]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="72" pageNumber="71" phylum="Tracheophyta" rank="species" species="pocita">
<emphasis id="B92BEAA3FFB0F84FFC71BE614420F9AF" box="[944,1067,1540,1563]" italics="true" pageId="72" pageNumber="71">M. pocita</emphasis>
</taxonomicName>
and also used to define the punensis subgroup. Because of our uncertainty about whether this reflects a relationship between these 2 subgroups, and because of its unusual overall combination of characters, we are continuing to list
<taxonomicName id="4C5F4D32FFB0F84FFED5BE014180F9CF" baseAuthorityName="Flint" baseAuthorityYear="1983" box="[276,395,1636,1659]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="72" pageNumber="71" phylum="Tracheophyta" rank="species" species="pocita">
<emphasis id="B92BEAA3FFB0F84FFED5BE014180F9CF" box="[276,395,1636,1659]" italics="true" pageId="72" pageNumber="71">M. pocita</emphasis>
</taxonomicName>
in its own subgroup.
</paragraph>
</subSubSection>
</treatment>
</document>
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