136 lines
16 KiB
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136 lines
16 KiB
XML
<document id="E3EB05929273019A71F9C5FE8B39176D" ID-DOI="10.5281/zenodo.5170203" ID-ISSN="1942-1354" ID-Zenodo-Dep="5170203" ID-ZooBank="AB1A57F0-7CB4-4830-920B-DF219740A596" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628281492031" checkinUser="felipe" docAuthor="Blahnik, Roger J. & Holzenthal, Ralph W." docDate="2017" docId="03F687A7FFA3F85BFF01BDC643B5FC6F" docLanguage="en" docName="InsectaMundi.0602.1-251.pdf" docOrigin="Insecta Mundi 2017 (602)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Mortoniella (Mortoniella) santiaga Sykora. Unfortunately 1999" docType="treatment" docVersion="3" lastPageNumber="91" masterDocId="FFCFFFDFFFF8F807FFC1B865400BFFB4" masterDocTitle="Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *" masterLastPageNumber="251" masterPageNumber="1" pageNumber="90" updateTime="1699039969677" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
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<mods:titleInfo id="57334BDA42A64961CA43B421F1374A50">
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<mods:title id="F0C11F704B524E9FE8A9119E1209FF4B">Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *</mods:title>
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<mods:namePart id="DA9892C9E9F9C6DD4A281CAB869D6D05">Blahnik, Roger J.</mods:namePart>
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<mods:namePart id="43D800E35A05764B7FC9E47E5165AC65">Holzenthal, Ralph W.</mods:namePart>
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<mods:typeOfResource id="E0016B3FC4A00373E398C65A894BE756">text</mods:typeOfResource>
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<mods:title id="2698C8D23C245976E633562A1AA00966">Insecta Mundi</mods:title>
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</mods:titleInfo>
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<mods:part id="93CBE67B5CBE382E97BC09C0FA50A889">
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<mods:date id="0313294E4BB9175A051AFF9987C14392">2017</mods:date>
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<mods:detail id="3337C18762F931A9353D8FB94DFAD3B7" type="pubDate">
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<mods:number id="9234FB2D01CBDDB3DC861A94A5DCB30D">2017-12-29</mods:number>
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<mods:detail id="AEEC82602A1EB834597433600F155B82" type="volume">
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<mods:number id="C640710CF6E6561DB3A63B830210B2A5">2017</mods:number>
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<mods:detail id="8C3E97F9428F5560C0ADBF44E54E1415" type="issue">
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<mods:number id="E191F3911595EB78D540061CE5F2D8D6">602</mods:number>
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<mods:classification id="DE6DDB23389981AA2428013995FB692E">journal article</mods:classification>
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<mods:identifier id="06D4351AB51C1B278B2A31A9BD677C86" type="DOI">10.5281/zenodo.5170203</mods:identifier>
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<mods:identifier id="297A61AAA18CC835739650205D2BFAC4" type="ISSN">1942-1354</mods:identifier>
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<mods:identifier id="046C7481F4617E7F176C0027498DF39B" type="Zenodo-Dep">5170203</mods:identifier>
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<mods:identifier id="51A5941194C661EF093BEA20D74EF0DB" type="ZooBank">AB1A57F0-7CB4-4830-920B-DF219740A596</mods:identifier>
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<treatment id="03F687A7FFA3F85BFF01BDC643B5FC6F" LSID="urn:lsid:plazi:treatment:03F687A7FFA3F85BFF01BDC643B5FC6F" httpUri="http://treatment.plazi.org/id/03F687A7FFA3F85BFF01BDC643B5FC6F" lastPageId="92" lastPageNumber="91" pageId="91" pageNumber="90">
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<subSubSection id="C345653AFFA3F85CFF01BDC64354FA0F" box="[192,863,1443,1467]" pageId="91" pageNumber="90" type="nomenclature">
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<paragraph id="8BE036B1FFA3F85CFF01BDC64354FA0F" blockId="91.[192,1440,1443,1564]" box="[192,863,1443,1467]" pageId="91" pageNumber="90">
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<heading id="D0A881DDFFA3F85CFF01BDC64354FA0F" bold="true" box="[192,863,1443,1467]" fontSize="10" level="2" pageId="91" pageNumber="90" reason="3">
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<taxonomicName id="4C5F4D32FFA3F85CFF01BDC64354FA0F" authority="Sykora, 1999" authorityName="Sykora. Unfortunately" authorityYear="1999" box="[192,863,1443,1467]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="91" pageNumber="90" phylum="Tracheophyta" rank="species" species="santiaga" subGenus="Mortoniella">
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<emphasis id="B92BEAA3FFA3F85CFF01BDC64354FA0F" bold="true" box="[192,863,1443,1467]" pageId="91" pageNumber="90">
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<emphasis id="B92BEAA3FFA3F85CFF01BDC64168FA0F" bold="true" box="[192,355,1443,1467]" italics="true" pageId="91" pageNumber="90">Mortoniella</emphasis>
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(
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<emphasis id="B92BEAA3FFA3F85CFEB7BDC64212FA0F" bold="true" box="[374,537,1443,1467]" italics="true" pageId="91" pageNumber="90">Mortoniella</emphasis>
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)
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<emphasis id="B92BEAA3FFA3F85CFDEABDC642ADFA0F" bold="true" box="[555,678,1443,1467]" italics="true" pageId="91" pageNumber="90">santiaga</emphasis>
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<bibRefCitation id="EFCE4B40FFA3F85CFD6FBDC64354FA0F" author="Sykora, J." box="[686,863,1443,1467]" pageId="91" pageNumber="90" pagination="377 - 387" refId="ref107309" refString="Sykora, J. 1999. Genus Mortoniella and its distribution in South America (Trichoptera, Glossosomatidae, Protoptilinae). p. 377 - 387. In: H. Malicky and P. Chantaramongkol (eds.). Proceedings of the 9 th International Symposium on Trichoptera. Faculty of Science, Chiang Mai University; Chiang Mai, Thailand. 479 p." type="book chapter" year="1999">Sykora, 1999</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C345653AFFA3F85CFF0BBD864558F88F" pageId="91" pageNumber="90" type="reference_group">
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<paragraph id="8BE036B1FFA3F85CFF0BBD86426CF9A8" blockId="91.[192,1440,1443,1564]" pageId="91" pageNumber="90">
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<treatmentCitationGroup id="AB4F119FFFA3F85CFF0BBD86426CF9A8" pageId="91" pageNumber="90">
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<treatmentCitation id="0AFE10A0FFA3F85CFF0BBD8642B9FA4F" author="Sykora, J." box="[202,690,1507,1531]" page="383" pageId="91" pageNumber="90" year="1999">
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<taxonomicName id="4C5F4D32FFA3F85CFF0BBD8642B9FA4F" authority="Sykora 1999: 383" authorityName="Sykora. Unfortunately" authorityPageNumber="383" authorityYear="1999" box="[202,690,1507,1531]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="91" pageNumber="90" phylum="Tracheophyta" rank="species" species="santiaga">
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<emphasis id="B92BEAA3FFA3F85CFF0BBD8641C4FA4F" box="[202,463,1507,1531]" italics="true" pageId="91" pageNumber="90">Mortoniella santiaga</emphasis>
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<bibRefCitation id="EFCE4B40FFA3F85CFE19BD8642B9FA4F" author="Sykora, J." box="[472,690,1507,1531]" pageId="91" pageNumber="90" pagination="377 - 387" refId="ref107309" refString="Sykora, J. 1999. Genus Mortoniella and its distribution in South America (Trichoptera, Glossosomatidae, Protoptilinae). p. 377 - 387. In: H. Malicky and P. Chantaramongkol (eds.). Proceedings of the 9 th International Symposium on Trichoptera. Faculty of Science, Chiang Mai University; Chiang Mai, Thailand. 479 p." type="book chapter" year="1999">Sykora 1999: 383</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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[member of flinti subgroup];
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<treatmentCitation id="0AFE10A0FFA3F85CFBE4BD8640F0F9AF" author="Blahnik, R. J. & R. W. Holzenthal" page="70" pageId="91" pageNumber="90" year="2008">
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<bibRefCitation id="EFCE4B40FFA3F85CFBE4BD8640F0F9AF" author="Blahnik, R. J. & R. W. Holzenthal" pageId="91" pageNumber="90" pagination="1 - 72" refId="ref106255" refString="Blahnik, R. J., and R. W. Holzenthal. 2008. Revision of the Mexican and Central American species of Mortoniella (Trichoptera: Glossosomatidae: Protoptilinae). Zootaxa 1711: 1 - 72." type="journal article" year="2008">Blahnik and Holzenthal 2008: 70</bibRefCitation>
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</treatmentCitation>
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[member of bilineata group].
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8BE036B1FFA3F85CFF31BE264558F88F" blockId="91.[192,1440,1603,1852]" pageId="91" pageNumber="90">
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Unfortunately, the
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<typeStatus id="54E48813FFA3F85CFE26BE26425AF9EF" box="[487,593,1603,1627]" pageId="91" pageNumber="90" type="holotype">holotype</typeStatus>
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of the species could not be located and thus was not available for direct comparison to
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<taxonomicName id="4C5F4D32FFA3F85CFE0EBE014273F9CF" authority="Blahnik & Holzenthal, 2017" authorityName="Blahnik & Holzenthal" authorityYear="2017" box="[463,632,1636,1659]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="91" pageNumber="90" phylum="Tracheophyta" rank="species" species="acutiterga" status="sp. nov.">
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<emphasis id="B92BEAA3FFA3F85CFE0EBE014273F9CF" box="[463,632,1636,1659]" italics="true" pageId="91" pageNumber="90">M. acutiterga</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A21857D8FFA3F85CFD4BBE0142DBF9CF" box="[650,720,1636,1659]" pageId="91" pageNumber="90" rank="species">n. sp.</taxonomicNameLabel>
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Apparent differences, based on the original description, include paramere appendages with several projections at midlength and a tergum X with flattened ventrolateral processes. This compares to a simple, narrow paramere appendage, without projections, and a tergum X with only narrow, almost spine-like, ventrolateral projections in
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<taxonomicName id="4C5F4D32FFA3F85CFB01BEA14563F96F" authorityName="Blahnik & Holzenthal" authorityYear="2017" box="[1216,1384,1732,1755]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="91" pageNumber="90" phylum="Tracheophyta" rank="species" species="acutiterga">
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<emphasis id="B92BEAA3FFA3F85CFB01BEA14563F96F" box="[1216,1384,1732,1755]" italics="true" pageId="91" pageNumber="90">M. acutiterga</emphasis>
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</taxonomicName>
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. We take these differences to indicate that the two are different species, although the overall similarities are otherwise extensive. The expanded, almost laciniate, apicolateral margin of the dorsal phallic spine, and arched, spine-like apices of tergum X are diagnostic characters for the species pair.
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</paragraph>
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</subSubSection>
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<subSubSection id="C345653AFFA3F85BFF31BF0643B5FC6F" lastPageId="92" lastPageNumber="91" pageId="91" pageNumber="90" type="distribution">
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<paragraph id="8BE036B1FFA3F85CFF31BF064218F8CF" blockId="91.[240,531,1891,1915]" box="[240,531,1891,1915]" pageId="91" pageNumber="90">
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<emphasis id="B92BEAA3FFA3F85CFF31BF064183F8CF" box="[240,392,1891,1915]" italics="true" pageId="91" pageNumber="90">Distribution</emphasis>
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—
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<collectingCountry id="F3487621FFA3F85CFE69BF064204F8CF" box="[424,527,1891,1915]" name="Ecuador" pageId="91" pageNumber="90">Ecuador</collectingCountry>
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.
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</paragraph>
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<paragraph id="8BE036B1FFA4F85BFF01B8A141C7FF6F" blockId="92.[192,460,195,219]" box="[192,460,195,219]" pageId="92" pageNumber="91">
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—
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<emphasis id="B92BEAA3FFA4F85BFF1AB8A641C7FF6F" bold="true" box="[219,460,195,219]" pageId="92" pageNumber="91">tridens subgroup</emphasis>
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</paragraph>
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<paragraph id="8BE036B1FFA4F85BFF01B96643EAFEAF" blockId="92.[192,993,259,283]" box="[192,993,259,283]" pageId="92" pageNumber="91">
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Included species:
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<taxonomicName id="4C5F4D32FFA4F85BFE5AB9664280FEAF" authority="Blahnik & Holzenthal, 2017" authorityName="Blahnik & Holzenthal" authorityYear="2017" box="[411,651,259,283]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="92" pageNumber="91" phylum="Tracheophyta" rank="species" species="tridens" status="sp. nov.">
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<emphasis id="B92BEAA3FFA4F85BFE5AB9664280FEAF" box="[411,651,259,283]" italics="true" pageId="92" pageNumber="91">Mortoniella tridens</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A21857D8FFA4F85BFD5BB96142D4FEAF" box="[666,735,260,283]" pageId="92" pageNumber="91" rank="species">n. sp.</taxonomicNameLabel>
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,
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<taxonomicName id="4C5F4D32FFA4F85BFD2DB9614384FEAF" authority="Blahnik & Holzenthal, 2017" authorityName="Blahnik & Holzenthal" authorityYear="2017" box="[748,911,260,283]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="92" pageNumber="91" phylum="Tracheophyta" rank="species" species="triramosa" status="sp. nov.">
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<emphasis id="B92BEAA3FFA4F85BFD2DB9614384FEAF" box="[748,911,260,283]" italics="true" pageId="92" pageNumber="91">M. triramosa</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A21857D8FFA4F85BFC5FB96143EAFEAF" box="[926,993,260,283]" pageId="92" pageNumber="91" rank="species">n. sp.</taxonomicNameLabel>
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</paragraph>
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<paragraph id="8BE036B1FFA4F85BFF31B92643B5FC6F" blockId="92.[192,1441,323,987]" pageId="92" pageNumber="91">
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The two species included here constitute a closely related species pair and are unusual in that the dorsal phallic spine is apparently 3-partite, deeply divided to form a mesal and 2 lateral projections. A similar character state seems to be indicated in the illustration of
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<taxonomicName id="4C5F4D32FFA4F85BFBEBB9E14568FE2F" authority="(Jacquemart)" baseAuthorityName="Jacquemart" baseAuthorityYear="1963" box="[1066,1379,387,411]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="92" pageNumber="91" phylum="Tracheophyta" rank="species" species="armata">
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<emphasis id="B92BEAA3FFA4F85BFBEBB9E144BBFE2F" box="[1066,1200,388,411]" italics="true" pageId="92" pageNumber="91">M. armata</emphasis>
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(Jacquemart)
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</taxonomicName>
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. The overall illustration of that species is so inadequate that it is difficult to make direct comparisons, but it does not seem to be closely related and we have speculatively placed it in the punensis subgroup (see for more extensive explanation). An unusual feature of the tridens subgroup is that the basal segment of the parameres is flattened and fused to the dorsal margin of the phallobase, appearing as a projecting, flattened lobe. This is connected to the lateral paramere appendage by an elongate membrane, such that the base of the paramere appendage emerges ventrally to the phallic ensemble, with the apical part strongly dorsally curved. A plausible explanation for the tripartite dorsal phallic spine is that the original paramere appendage structure was doubled, as in members of the argentinica subgroup, and the dorsolateral pair, along with the basal structure of the paramere, became fused with the dorsal phallic spine, producing the tripartite assemblage. On this basis, we speculate that the two groups may be related, with
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<taxonomicName id="4C5F4D32FFA4F85BFDBDBA8142FFFD4F" authority="Blahnik & Holzenthal, 2017" authorityName="Blahnik & Holzenthal" authorityYear="2017" box="[636,756,740,763]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="92" pageNumber="91" phylum="Tracheophyta" rank="species" species="esrossi" status="sp. nov.">
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<emphasis id="B92BEAA3FFA4F85BFDBDBA8142FFFD4F" box="[636,756,740,763]" italics="true" pageId="92" pageNumber="91">M. esrossi</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A21857D8FFA4F85BFCC3BA814349FD4F" box="[770,834,740,763]" pageId="92" pageNumber="91" rank="species">n. sp.</taxonomicNameLabel>
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(in the esrossi subgroup) as a possible connecting species (which see for a more complete explanation). Like most members of the argentinica subgroup, the ventral process of the species in the tridens subgroup is large, subtriangular, and ventrally projected, thus resembling species in the leroda group. It seems likely that both of these subgroups are related to that lineage. Unlike species in the argentinica subgroup, the inferior appendages of species in the tridens subgroup are very strongly fused to the phallicata, forming a short setose ventromesal projection, and lack narrow, reflexed dorsolateral lobes. Additionally, the spine-like projections from the mesal pockets of the inferior appendages are not nearly so elongate.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |