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<document id="B324A745BB1035D51DFE9E191C43FF2E" ID-DOI="10.5281/zenodo.5170203" ID-ISSN="1942-1354" ID-Zenodo-Dep="5170203" ID-ZooBank="AB1A57F0-7CB4-4830-920B-DF219740A596" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628281492031" checkinUser="felipe" docAuthor="Blahnik, Roger J. &amp; Holzenthal, Ralph W." docDate="2017" docId="03F687A7FF95F869FF01BC2645ABFC4F" docLanguage="en" docName="InsectaMundi.0602.1-251.pdf" docOrigin="Insecta Mundi 2017 (602)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Mortoniella (Nanotrichia) licina Blahnik &amp; Holzenthal 2017, new species" docType="treatment" docVersion="4" lastPageNumber="109" masterDocId="FFCFFFDFFFF8F807FFC1B865400BFFB4" masterDocTitle="Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *" masterLastPageNumber="251" masterPageNumber="1" pageNumber="108" updateTime="1699039969677" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
<mods:mods id="A14D27478B9BCA9D5B11564F7646FDED" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="57F72C8F88FA0584A01BCB0913C5E1B6">
<mods:title id="B85CB4F3CA864655780B949CAD47B833">Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *</mods:title>
</mods:titleInfo>
<mods:name id="B3F03A4CEAEC64CDBF8293C62C7E6093" type="personal">
<mods:role id="63DD91AA586D40959C4867FCC95FA0D7">
<mods:roleTerm id="11314733843F99B9081DAD50DAADB09A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="DFD041085B1FBABA777CEE4F0BC90199">Blahnik, Roger J.</mods:namePart>
</mods:name>
<mods:name id="4570B999DA74CD7BD050A06466AF2256" type="personal">
<mods:role id="952C703CE69BCCACB7B3F3BAC668F399">
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<mods:namePart id="55515304979DB5B0ED36276CA05A1EE2">Holzenthal, Ralph W.</mods:namePart>
</mods:name>
<mods:typeOfResource id="B59F02C7C6B77A430F63C3A05C0E3FC7">text</mods:typeOfResource>
<mods:relatedItem id="89247C4E2AFC0B7E695E625A508DB879" type="host">
<mods:titleInfo id="7AF956FDF24E0EF19115AA119702734B">
<mods:title id="A8E56384EEE020A5BD5E40D07B7667F9">Insecta Mundi</mods:title>
</mods:titleInfo>
<mods:part id="BF13F2582F807CBD5ED16A0A899B9D8E">
<mods:date id="91B8076F2ACDFB23BE008FEEA0D6FDA6">2017</mods:date>
<mods:detail id="1E5B8255455251071B7EED613AFE1814" type="pubDate">
<mods:number id="8DE15FCADB7D7450315950084693220F">2017-12-29</mods:number>
</mods:detail>
<mods:detail id="2791E07541EBAD4480F2B3F5925AF05B" type="volume">
<mods:number id="24B18E6F5F905B058B694CE8CDA6682C">2017</mods:number>
</mods:detail>
<mods:detail id="FB6FD1A09E95AFA7EDED6B408DBDE69D" type="issue">
<mods:number id="07310134F216B1045EB98CF69E1C6B08">602</mods:number>
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<mods:start id="AE73F55F0FE15CB065E81841972EDFE7">1</mods:start>
<mods:end id="CD674F1D78053AB3A040A039385FF562">251</mods:end>
</mods:extent>
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<mods:classification id="80A50DC724B9E7FC4B28D871F06AC5F4">journal article</mods:classification>
<mods:identifier id="F6B5AEC9CD62302FA2080EA6DFD30723" type="DOI">10.5281/zenodo.5170203</mods:identifier>
<mods:identifier id="91CBEC951D5625A792B89297F8EF1C74" type="ISSN">1942-1354</mods:identifier>
<mods:identifier id="17D647FBCF4236246261933145770002" type="Zenodo-Dep">5170203</mods:identifier>
<mods:identifier id="553901ACBB39D5B54D4B3BE761C1C87B" type="ZooBank">AB1A57F0-7CB4-4830-920B-DF219740A596</mods:identifier>
</mods:mods>
<treatment id="03F687A7FF95F869FF01BC2645ABFC4F" ID-DOI="http://doi.org/10.5281/zenodo.5186347" ID-Zenodo-Dep="5186347" LSID="urn:lsid:plazi:treatment:03F687A7FF95F869FF01BC2645ABFC4F" httpUri="http://treatment.plazi.org/id/03F687A7FF95F869FF01BC2645ABFC4F" lastPageId="110" lastPageNumber="109" pageId="109" pageNumber="108">
<subSubSection id="C345653AFF95F86AFF01BC264337FBEF" box="[192,828,1091,1115]" pageId="109" pageNumber="108" type="nomenclature">
<paragraph id="8BE036B1FF95F86AFF01BC264337FBEF" blockId="109.[192,828,1091,1147]" box="[192,828,1091,1115]" pageId="109" pageNumber="108">
<heading id="D0A881DDFF95F86AFF01BC264337FBEF" bold="true" box="[192,828,1091,1115]" fontSize="10" level="2" pageId="109" pageNumber="108" reason="3">
<taxonomicName id="4C5F4D32FF95F86AFF01BC26428EFBEF" authority="Blahnik &amp; Holzenthal, 2017" authorityName="Blahnik &amp; Holzenthal" authorityYear="2017" box="[192,645,1091,1115]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="109" pageNumber="108" phylum="Tracheophyta" rank="species" species="licina" status="sp. nov." subGenus="Nanotrichia">
<emphasis id="B92BEAA3FF95F86AFF01BC26428EFBEF" bold="true" box="[192,645,1091,1115]" pageId="109" pageNumber="108">
<emphasis id="B92BEAA3FF95F86AFF01BC264168FBEF" bold="true" box="[192,355,1091,1115]" italics="true" pageId="109" pageNumber="108">Mortoniella</emphasis>
(
<emphasis id="B92BEAA3FF95F86AFEB7BC26422AFBEF" bold="true" box="[374,545,1091,1115]" italics="true" pageId="109" pageNumber="108">Nanotrichia</emphasis>
)
<emphasis id="B92BEAA3FF95F86AFDF2BC26428EFBEF" bold="true" box="[563,645,1091,1115]" italics="true" pageId="109" pageNumber="108">licina</emphasis>
</emphasis>
</taxonomicName>
,
<emphasis id="B92BEAA3FF95F86AFD54BC264337FBEF" bold="true" box="[661,828,1091,1115]" pageId="109" pageNumber="108">
<taxonomicNameLabel id="A21857D8FF95F86AFD54BC264337FBEF" box="[661,828,1091,1115]" pageId="109" pageNumber="108" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C345653AFF95F869FF01BC064127FDCF" lastPageId="110" lastPageNumber="109" pageId="109" pageNumber="108" type="description">
<paragraph id="8BE036B1FF95F86AFF01BC064159FBCF" blockId="109.[192,828,1091,1147]" box="[192,338,1123,1147]" pageId="109" pageNumber="108">
<figureCitation id="13642A34FF95F86AFF01BC064113FBCF" box="[192,280,1123,1147]" captionStart="Figure 93" captionStartId="233.[192,276,1836,1858]" captionTargetBox="[354,1293,266,1737]" captionTargetId="figure-56@233.[354,1294,266,1737]" captionTargetPageId="233" captionText="Figure 93. Mortoniella (Nanotrichia) licina, new species, male genitalia. A—lateral; B—segment IX and tergum X, dorsal; C—phallic ensemble, ventral; D—dorsal phallic spine and phallobase, dorsal; E—ventral process of segment VI, lateral." figureDoi="http://doi.org/10.5281/zenodo.5170401" httpUri="https://zenodo.org/record/5170401/files/figure.png" pageId="109" pageNumber="108">Fig. 93</figureCitation>
,
<figureCitation id="13642A34FF95F86AFEE4BC064159FBCF" box="[293,338,1123,1147]" captionStart="Figures 121-125" captionStartId="248.[192,287,1807,1829]" captionTargetBox="[302,1326,269,1692]" captionTargetId="figure-75@248.[302,1327,269,1692]" captionTargetPageId="248" captionText="Figures 121-125. Mortoniella (Nanotrichia), velasquezi group species, female genitalia.121—Mortoniella eduardoi (Rueda Martín and Gibon): A—lateral; B—dorsal. 122—Mortoniella licina, n. sp., dorsal. 123—Mortoniella venezuelensis, n. sp., dorsal. 124—Mortoniella spangleri, n. sp. A—lateral; B—dorsal. 125—Mortoniella coheni, n. sp., dorsal." figureDoi="http://doi.org/10.5281/zenodo.5170423" httpUri="https://zenodo.org/record/5170423/files/figure.png" pageId="109" pageNumber="108">122</figureCitation>
</paragraph>
<paragraph id="8BE036B1FF95F86AFF31BCC644E4FA4F" blockId="109.[192,1440,1187,1531]" pageId="109" pageNumber="108">
<taxonomicName id="4C5F4D32FF95F86AFF31BCC641DCFB0F" authorityName="Blahnik &amp; Holzenthal" authorityYear="2017" box="[240,471,1187,1211]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="109" pageNumber="108" phylum="Tracheophyta" rank="species" species="licina">
<emphasis id="B92BEAA3FF95F86AFF31BCC641DCFB0F" box="[240,471,1187,1211]" italics="true" pageId="109" pageNumber="108">Mortoniella licina</emphasis>
</taxonomicName>
is very closely related to
<taxonomicName id="4C5F4D32FF95F86AFCF2BCC14533FB0F" authority="(Rueda Martin and Gibon)" baseAuthorityName="Rueda Martin and Gibon" baseAuthorityYear="2008" box="[819,1336,1187,1211]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="109" pageNumber="108" phylum="Tracheophyta" rank="species" species="eduardoi">
<emphasis id="B92BEAA3FF95F86AFCF2BCC143DAFB0F" box="[819,977,1187,1211]" italics="true" pageId="109" pageNumber="108">M. eduardoi</emphasis>
(Rueda Martín and Gibon)
</taxonomicName>
and
<taxonomicName id="4C5F4D32FF95F86AFABEBCC1416DFB6F" authority="Blahnik &amp; Holzenthal, 2017" authorityName="Blahnik &amp; Holzenthal" authorityYear="2017" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="109" pageNumber="108" phylum="Tracheophyta" rank="species" species="venezuelensis" status="sp. nov.">
<emphasis id="B92BEAA3FF95F86AFABEBCC1416DFB6F" italics="true" pageId="109" pageNumber="108">M. venezuelensis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A21857D8FF95F86AFEB5BCA141BDFB6F" box="[372,438,1220,1243]" pageId="109" pageNumber="108" rank="species">n. sp.</taxonomicNameLabel>
All of these species have the apical spine-like projections from the mesal pockets of the inferior appendages very elongate and fused to the ventral part of the inferior appendages to form elongate, projecting lobes. All of the species also have an endophallic membrane that is ornamented with a dorsomesal projection with minute spines, membranous lateral lobes with minute spines, and paired ventral sclerites, each with a somewhat forked apex.
<taxonomicName id="4C5F4D32FF95F86AFC6CBD264486FAEF" authorityName="Blahnik &amp; Holzenthal" authorityYear="2017" box="[941,1165,1347,1371]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="109" pageNumber="108" phylum="Tracheophyta" rank="species" species="licina">
<emphasis id="B92BEAA3FF95F86AFC6CBD264486FAEF" box="[941,1165,1347,1371]" italics="true" pageId="109" pageNumber="108">Mortoniella licina</emphasis>
</taxonomicName>
differs from the other two species in lacking a field of upright scale-like setae on the hind wings of males. It most resembles
<taxonomicName id="4C5F4D32FF95F86AFF01BDE1419AFA2F" authorityName="Blahnik &amp; Holzenthal" authorityYear="2017" box="[192,401,1411,1435]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="109" pageNumber="108" phylum="Tracheophyta" rank="species" species="venezuelensis">
<emphasis id="B92BEAA3FF95F86AFF01BDE1419AFA2F" box="[192,401,1411,1435]" italics="true" pageId="109" pageNumber="108">M. venezuelensis</emphasis>
</taxonomicName>
in that the paramere appendages are uniform in width and do not have modified apices. Those in
<taxonomicName id="4C5F4D32FF95F86AFE4DBDC141F7FA0F" authorityName="Blahnik &amp; Holzenthal" authorityYear="2017" box="[396,508,1443,1467]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="109" pageNumber="108" phylum="Tracheophyta" rank="species" species="licina">
<emphasis id="B92BEAA3FF95F86AFE4DBDC141F7FA0F" box="[396,508,1443,1467]" italics="true" pageId="109" pageNumber="108">M. licina</emphasis>
</taxonomicName>
are usually distinctly curved apically, rather than being straight. A subtle, but distinctive character is that the ventral lobes of the inferior appendages and attached spine-like projections of the mesal pockets, are longer and more distinctly upturned in
<taxonomicName id="4C5F4D32FF95F86AFBB9BD8144E3FA4F" authorityName="Blahnik &amp; Holzenthal" authorityYear="2017" box="[1144,1256,1507,1531]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="109" pageNumber="108" phylum="Tracheophyta" rank="species" species="licina">
<emphasis id="B92BEAA3FF95F86AFBB9BD8144E3FA4F" box="[1144,1256,1507,1531]" italics="true" pageId="109" pageNumber="108">M. licina</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BE036B1FF95F86AFF31BE464590F92F" blockId="109.[192,1441,1571,1916]" pageId="109" pageNumber="108">
<emphasis id="B92BEAA3FF95F86AFF31BE46413DF98F" box="[240,310,1571,1595]" italics="true" pageId="109" pageNumber="108">Adult</emphasis>
—Length of forewing: male
<quantity id="4CA79B54FF95F86AFD48BE46431DF98F" box="[649,790,1571,1595]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" metricValueMax="3.1" metricValueMin="2.7" pageId="109" pageNumber="108" unit="mm" value="2.9" valueMax="3.1" valueMin="2.7">2.7-3.1 mm</quantity>
; female
<quantity id="4CA79B54FF95F86AFCB8BE46440DF98F" box="[889,1030,1571,1595]" metricMagnitude="-3" metricUnit="m" metricValue="3.15" metricValueMax="3.5" metricValueMin="2.8" pageId="109" pageNumber="108" unit="mm" value="3.15" valueMax="3.5" valueMin="2.8">2.8-3.5 mm</quantity>
. Forewing with forks I, II, and III present, hind wing with fork II only. Spur formula 0:3:4. Overall color (in alcohol) medium brown. Forewing with narrow whitish wing bar at anastomosis. Males with scale-like setae paralleling veins in fore- and hind wings, hind wing without additional field of darkened, semi-erect, scale-like setae.
</paragraph>
<paragraph id="8BE036B1FF95F869FF31BEC64127FDCF" blockId="109.[192,1441,1571,1916]" lastBlockId="110.[192,1441,195,1019]" lastPageId="110" lastPageNumber="109" pageId="109" pageNumber="108">
<emphasis id="B92BEAA3FF95F86AFF31BEC641AAF90F" box="[240,417,1699,1723]" italics="true" pageId="109" pageNumber="108">Male genitalia</emphasis>
—Ventral process of segment VI posteriorly projecting, narrow basally, length about 3½ times width at base. Segment IX with anterolateral margin rounded and distinctly produced in ventral ½, posterolateral margin nearly straight; segment deeply mesally excised dorsally and ventrally, forming lateral lobes, separated dorsomesally by more than ½ width of segment. Tergum X, as viewed laterally, relatively short, narrowed and slightly ventrally curved apically, basomesally with pair of setose sclerites; as viewed dorsally, short and wide, apicomesally with deep U-shaped emargination, extending more than ½ length of tergum; ventrolateral lobes slightly produced basally, obsolete apically. Inferior appendage with setose, digitate dorsolateral projections, fused apicoventrally to spine-like projections of mesal pockets; composite apical structure forming prominent and very elongate, curved, ventral lobes. Mesal pockets of inferior appendage very large, with elongate, thick, strongly posterodorsally curved, spine-like, apicoventral projections. Paramere appendage elongate, narrow, nearly uniform in width, apex acute, usually distinctly curved in apical part. Phallobase very short, with elongate ventral rod-like projections, each strongly flared apically. Dorsal phallic spine, as viewed laterally, somewhat widened basally, strongly reflexed in about apical 1/3, apex rounded; as viewed dorsally, with distinct dorsolateral projections, widest in basal ½, tapering apically, reflexed apex rounded. Phallicata short, continuous with endophallic membrane apically, laterally with rounded sclerotized projections, surrounding mesal pockets of inferior appendage. Endophallic membrane continuous with phallicata, dorsomesally with slightly raised projection with numerous minute spines, laterally with projecting membranous lobes with minute spines, apically with narrow projecting lobe, ventrally with pair of sclerotized projections, each with v-shaped apical notch (modified phallotremal spines?).
</paragraph>
</subSubSection>
<subSubSection id="C345653AFF96F869FF31BAE645ABFC4F" pageId="110" pageNumber="109" type="materials_examined">
<paragraph id="8BE036B1FF96F869FF31BAE642A3FD0F" blockId="110.[192,1441,195,1019]" pageId="110" pageNumber="109">
<materialsCitation id="3B373CECFF96F869FF31BAE642A3FD0F" collectingDate="1978-11-29" collectionCode="N, NMNH" collectorName="JJ Anderson" country="Ecuador" location="Yanzaza" pageId="110" pageNumber="109" specimenCode="UMSP000096933" specimenCount="1" stateProvince="Zamora-Chinchipe" typeStatus="holotype">
<emphasis id="B92BEAA3FF96F869FF31BAE641BEFD2F" bold="true" box="[240,437,643,667]" italics="true" pageId="110" pageNumber="109">
<typeStatus id="54E48813FF96F869FF31BAE64162FD2F" box="[240,361,643,667]" pageId="110" pageNumber="109" type="holotype">Holotype</typeStatus>
male
</emphasis>
(alcohol)—
<emphasis id="B92BEAA3FF96F869FD88BAE643F7FD2F" bold="true" box="[585,1020,643,667]" pageId="110" pageNumber="109">
<collectingCountry id="F3487621FF96F869FD88BAE642E9FD2F" box="[585,738,643,667]" name="Ecuador" pageId="110" pageNumber="109">ECUADOR</collectingCountry>
:
<collectingRegion id="499BF853FF96F869FD2EBAE643FDFD2F" box="[751,1014,643,667]" country="Ecuador" name="Zamora-Chinchipe" pageId="110" pageNumber="109">Zamora-Chinchipe</collectingRegion>
:
</emphasis>
<location id="8E80606AFF96F869FBC2BAE14466FD2F" LSID="urn:lsid:plazi:treatment:03F687A7FF95F869FF01BC2645ABFC4F:8E80606AFF96F869FBC2BAE14466FD2F" box="[1027,1133,644,667]" country="Ecuador" name="Yanzaza" pageId="110" pageNumber="109" stateProvince="Zamora-Chinchipe">Yanzaza</location>
(
<quantity id="4CA79B54FF96F869FBBFBAE644B6FD2F" box="[1150,1213,643,667]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="110" pageNumber="109" unit="km" value="5.0">5 km</quantity>
<collectionCode id="ED4EAE74FF96F869FB04BAE644D6FD2F" box="[1221,1245,643,667]" country="China" lsid="urn:lsid:biocol.org:col:13092" name="Nanjing University" pageId="110" pageNumber="109" type="Herbarium">N</collectionCode>
),
<date id="FFE11071FF96F869FB32BAE6457EFD2F" box="[1267,1397,643,667]" pageId="110" pageNumber="109" value="1978-11-29">
<collectingDate id="EFA5E999FF96F869FB32BAE6457EFD2F" box="[1267,1397,643,667]" pageId="110" pageNumber="109" value="1978-11-29">29.xi.1978</collectingDate>
</date>
,
<collectorName id="26AA5367FF96F869FA43BAE1413CFD0F" pageId="110" pageNumber="109">JJ Anderson</collectorName>
(
<specimenCode id="DBF99ECAFF96F869FE86BAC64228FD0F" box="[327,547,675,699]" collectionCode="UMSP" country="USA" httpUri="http://grbio.org/cool/3avb-bd3e" name="University of Minnesota Insect Collection" pageId="110" pageNumber="109" type="University or college">UMSP000096933</specimenCode>
) (
<collectionCode id="ED4EAE74FF96F869FDFABAC64297FD0F" box="[571,668,675,699]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="110" pageNumber="109" type="Museum">NMNH</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BE036B1FF96F869FF31BAA64553FC0F" blockId="110.[192,1441,195,1019]" pageId="110" pageNumber="109">
<materialsCitation id="3B373CECFF96F869FF31BAA6413BFD4F" collectingDate="1976-07-08" collectionCode="J, NMNH" collectorName="Cohen" country="Ecuador" location="Puerto Nuevo" pageId="110" pageNumber="109" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Napo" typeStatus="paratype">
<emphasis id="B92BEAA3FF96F869FF31BAA64284FD6F" bold="true" box="[240,655,707,731]" pageId="110" pageNumber="109">
<emphasis id="B92BEAA3FF96F869FF31BAA64170FD6E" bold="true" box="[240,379,707,730]" italics="true" pageId="110" pageNumber="109">
<typeStatus id="54E48813FF96F869FF31BAA64170FD6E" box="[240,379,707,730]" pageId="110" pageNumber="109" type="paratype">Paratypes</typeStatus>
</emphasis>
—
<collectingCountry id="F3487621FF96F869FE57BAA64224FD6F" box="[406,559,707,731]" name="Ecuador" pageId="110" pageNumber="109">ECUADOR</collectingCountry>
:
<collectingRegion id="499BF853FF96F869FDFFBAA64282FD6F" box="[574,649,707,731]" country="Ecuador" name="Napo" pageId="110" pageNumber="109">Napo</collectingRegion>
:
</emphasis>
<location id="8E80606AFF96F869FD58BAA14342FD6F" LSID="urn:lsid:plazi:treatment:03F687A7FF95F869FF01BC2645ABFC4F:8E80606AFF96F869FD58BAA14342FD6F" box="[665,841,708,731]" country="Ecuador" name="Puerto Nuevo" pageId="110" pageNumber="109" stateProvince="Napo">Puerto Nuevo</location>
,
<date id="FFE11071FF96F869FC96BAA643D8FD6F" box="[855,979,707,731]" pageId="110" pageNumber="109" value="1976-07-08">
<collectingDate id="EFA5E999FF96F869FC96BAA643D8FD6F" box="[855,979,707,731]" pageId="110" pageNumber="109" value="1976-07-08">8.vii.1976</collectingDate>
</date>
,
<collectionCode id="ED4EAE74FF96F869FC23BAA143FAFD6F" box="[994,1009,708,731]" country="South Africa" lsid="urn:lsid:biocol.org:col:14647" name="University of the Witwatersrand" pageId="110" pageNumber="109" type="Herbarium">J</collectionCode>
<collectorName id="26AA5367FF96F869FC3ABAA64445FD6F" box="[1019,1102,707,731]" pageId="110" pageNumber="109">Cohen</collectorName>
–
<specimenCount id="9D59FD38FF96F869FB9EBAA644B9FD6F" box="[1119,1202,707,731]" pageId="110" pageNumber="109" type="male">1 male</specimenCount>
,
<specimenCount id="9D59FD38FF96F869FB00BAA64527FD6F" box="[1217,1324,707,731]" pageId="110" pageNumber="109" type="female">1 female</specimenCount>
(alcohol) (
<collectionCode id="ED4EAE74FF96F869FF09BA864123FD4F" box="[200,296,739,763]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="110" pageNumber="109" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B373CECFF96F869FEFEBA864547FD4F" box="[319,1356,739,764]" collectingDate="1977-05-21" collectionCode="DR, NMNH" collectorName="PJ Spangler &amp; Givens" country="Ecuador" location="Puyo" pageId="110" pageNumber="109" specimenCount="1" specimenCount-male="1" stateProvince="Pastaza" typeStatus="paratype">
<emphasis id="B92BEAA3FF96F869FEFEBA8641B8FD4F" bold="true" box="[319,435,739,763]" pageId="110" pageNumber="109">
<collectingRegion id="499BF853FF96F869FEFEBA8641A5FD4F" box="[319,430,739,763]" country="Ecuador" name="Pastaza" pageId="110" pageNumber="109">Pastaza</collectingRegion>
:
</emphasis>
<location id="8E80606AFF96F869FE7CBA8141F5FD48" LSID="urn:lsid:plazi:treatment:03F687A7FF95F869FF01BC2645ABFC4F:8E80606AFF96F869FE7CBA8141F5FD48" box="[445,510,740,764]" country="Ecuador" name="Puyo" pageId="110" pageNumber="109" stateProvince="Pastaza">Puyo</location>
,
<date id="FFE11071FF96F869FDCCBA86428DFD4F" box="[525,646,739,763]" pageId="110" pageNumber="109" value="1977-05-21">
<collectingDate id="EFA5E999FF96F869FDCCBA86428DFD4F" box="[525,646,739,763]" pageId="110" pageNumber="109" value="1977-05-21">21.v.1977</collectingDate>
</date>
,
<collectorName id="26AA5367FF96F869FD54BA81433BFD4F" box="[661,816,739,763]" pageId="110" pageNumber="109">PJ Spangler</collectorName>
and
<collectionCode id="ED4EAE74FF96F869FCB3BA814391FD4F" box="[882,922,740,763]" country="Germany" lsid="urn:lsid:biocol.org:col:15173" name="Technische Universität Dresden" pageId="110" pageNumber="109" type="Herbarium">DR</collectionCode>
<collectorName id="26AA5367FF96F869FC65BA8643F5FD4F" box="[932,1022,739,763]" pageId="110" pageNumber="109">Givens</collectorName>
–
<specimenCount id="9D59FD38FF96F869FBCEBA864454FD4F" box="[1039,1119,739,763]" pageId="110" pageNumber="109" type="male">1 male</specimenCount>
(alcohol) (
<collectionCode id="ED4EAE74FF96F869FB25BA86454FFD4F" box="[1252,1348,739,763]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="110" pageNumber="109" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B373CECFF96F869FA9BBA81436AFCAF" collectingDate="1976-01-30" collectionCode="NMNH" collectorName="Spangler" country="Ecuador" location="Puyo" pageId="110" pageNumber="109" specimenCount="9" specimenCount-male="9" stateProvince="Pastaza" typeStatus="paratype">
<location id="8E80606AFF96F869FA9BBA814590FD48" LSID="urn:lsid:plazi:treatment:03F687A7FF95F869FF01BC2645ABFC4F:8E80606AFF96F869FA9BBA814590FD48" box="[1370,1435,740,764]" country="Ecuador" name="Puyo" pageId="110" pageNumber="109" stateProvince="Pastaza">Puyo</location>
,
<date id="FFE11071FF96F869FF01BB66413FFCAF" box="[192,308,771,795]" pageId="110" pageNumber="109" value="1976-01-30">
<collectingDate id="EFA5E999FF96F869FF01BB66413FFCAF" box="[192,308,771,795]" pageId="110" pageNumber="109" value="1976-01-30">30.i.1976</collectingDate>
</date>
,
<collectorName id="26AA5367FF96F869FE85BB6641BFFCAF" box="[324,436,771,795]" pageId="110" pageNumber="109">Spangler</collectorName>
et al.–
<specimenCount id="9D59FD38FF96F869FDCEBB664279FCAF" box="[527,626,771,795]" pageId="110" pageNumber="109" type="male">9 males</specimenCount>
(alcohol) (
<collectionCode id="ED4EAE74FF96F869FD38BB664352FCAF" box="[761,857,771,795]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="110" pageNumber="109" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B373CECFF96F869FCB0BB6141ABFC8F" collectingDate="1976-05-18" collectionCode="J, NMNH" collectorName="Cohen" country="Ecuador" elevation="500" location="Tewaeno" pageId="110" pageNumber="109" specimenCount="1" specimenCount-male="1" stateProvince="Pastaza" typeStatus="paratype">
<location id="8E80606AFF96F869FCB0BB6143E8FCAF" LSID="urn:lsid:plazi:treatment:03F687A7FF95F869FF01BC2645ABFC4F:8E80606AFF96F869FCB0BB6143E8FCAF" box="[881,995,772,795]" country="Ecuador" name="Tewaeno" pageId="110" pageNumber="109" stateProvince="Pastaza">Tewaeno</location>
,
<quantity id="4CA79B54FF96F869FC30BB664432FCAF" box="[1009,1081,771,795]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="110" pageNumber="109" unit="m" value="500.0">
<elevation id="0072D182FF96F869FC30BB664432FCAF" box="[1009,1081,771,795]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="110" pageNumber="109" unit="m" value="500.0">500m</elevation>
</quantity>
,
<date id="FFE11071FF96F869FB89BB6644C9FCAF" box="[1096,1218,771,795]" pageId="110" pageNumber="109" value="1976-05-18">
<collectingDate id="EFA5E999FF96F869FB89BB6644C9FCAF" box="[1096,1218,771,795]" pageId="110" pageNumber="109" value="1976-05-18">18.v.1976</collectingDate>
</date>
,
<collectionCode id="ED4EAE74FF96F869FB13BB6144EAFCAF" box="[1234,1249,772,795]" country="South Africa" lsid="urn:lsid:biocol.org:col:14647" name="University of the Witwatersrand" pageId="110" pageNumber="109" type="Herbarium">J</collectionCode>
<collectorName id="26AA5367FF96F869FB2DBB664534FCAF" box="[1260,1343,771,795]" pageId="110" pageNumber="109">Cohen</collectorName>
–
<specimenCount id="9D59FD38FF96F869FA8EBB6645ABFCAF" box="[1359,1440,771,795]" pageId="110" pageNumber="109" type="male">1 male</specimenCount>
(alcohol) (
<collectionCode id="ED4EAE74FF96F869FEF9BB464193FC8F" box="[312,408,803,827]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="110" pageNumber="109" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B373CECFF96F869FE6ABB464203FCEF" collectingDate="1976-05-22" collectionCode="NE, J, NMNH" collectorName="Cohen" country="Ecuador" elevation="400" latitude="-1.31667" location="Tigueno" longLatPrecision="1" longitude="-77.46667" municipality="Tzapino" pageId="110" pageNumber="109" specimenCount="1" specimenCount-male="1" stateProvince="Pastaza" typeStatus="paratype">
<collectingMunicipality id="6B84ACCBFF96F869FE6ABB464219FC8F" box="[427,530,803,827]" pageId="110" pageNumber="109">Tzapino</collectingMunicipality>
,
<quantity id="4CA79B54FF96F869FDDCBB464261FC8F" box="[541,618,803,827]" metricMagnitude="4" metricUnit="m" metricValue="3.2" pageId="110" pageNumber="109" unit="km" value="32.0">32 km</quantity>
<collectionCode id="ED4EAE74FF96F869FDB1BB414292FC8F" box="[624,665,804,827]" country="Australia" lsid="urn:lsid:biocol.org:col:14575" name="University of New England" pageId="110" pageNumber="109" type="Herbarium">NE</collectionCode>
<location id="8E80606AFF96F869FD61BB464303FC8F" LSID="urn:lsid:plazi:treatment:03F687A7FF95F869FF01BC2645ABFC4F:8E80606AFF96F869FD61BB464303FC8F" box="[672,776,803,827]" country="Ecuador" latitude="-1.31667" longLatPrecision="1" longitude="-77.46667" municipality="Tzapino" name="Tigueno" pageId="110" pageNumber="109" stateProvince="Pastaza">Tigueno</location>
,
<geoCoordinate id="EE6B5076FF96F869FCD3BB464393FC8F" box="[786,920,803,827]" degrees="1.31667" direction="south" orientation="latitude" pageId="110" pageNumber="109" precision="1" value="-1.31667">1.31667° S</geoCoordinate>
,
<geoCoordinate id="EE6B5076FF96F869FC65BB464448FC8F" box="[932,1091,803,827]" degrees="77.46667" direction="west" orientation="longitude" pageId="110" pageNumber="109" precision="1" value="-77.46667">77.46667° W</geoCoordinate>
,
<quantity id="4CA79B54FF96F869FB8FBB464490FC8F" box="[1102,1179,803,827]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="110" pageNumber="109" unit="m" value="400.0">
<elevation id="0072D182FF96F869FB8FBB464490FC8F" box="[1102,1179,803,827]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="110" pageNumber="109" unit="m" value="400.0">400 m</elevation>
</quantity>
,
<date id="FFE11071FF96F869FB66BB46452AFC8F" box="[1191,1313,803,827]" pageId="110" pageNumber="109" value="1976-05-22">
<collectingDate id="EFA5E999FF96F869FB66BB46452AFC8F" box="[1191,1313,803,827]" pageId="110" pageNumber="109" value="1976-05-22">22.v.1976</collectingDate>
</date>
,
<collectionCode id="ED4EAE74FF96F869FAECBB414537FC8F" box="[1325,1340,804,827]" country="South Africa" lsid="urn:lsid:biocol.org:col:14647" name="University of the Witwatersrand" pageId="110" pageNumber="109" type="Herbarium">J</collectionCode>
<collectorName id="26AA5367FF96F869FA83BB46459AFC8F" box="[1346,1425,803,827]" pageId="110" pageNumber="109">Cohen</collectorName>
–
<specimenCount id="9D59FD38FF96F869FF01BB26411CFCEF" box="[192,279,835,859]" pageId="110" pageNumber="109" type="male">1 male</specimenCount>
(alcohol) (
<collectionCode id="ED4EAE74FF96F869FE61BB26420BFCEF" box="[416,512,835,859]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="110" pageNumber="109" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B373CECFF96F869FDD9BB264203FCCF" collectingDate="1978-12-04" collectionCode="NMNH" collectorName="JJ Anderson" country="Ecuador" location="Zamora" pageId="110" pageNumber="109" specimenCount="512" specimenCount-female="280" specimenCount-male="232" stateProvince="Zamora-Chinchipe" typeStatus="paratype">
<emphasis id="B92BEAA3FF96F869FDD9BB26432EFCEF" bold="true" box="[536,805,835,859]" pageId="110" pageNumber="109">
<collectingRegion id="499BF853FF96F869FDD9BB264314FCEF" box="[536,799,835,859]" country="Ecuador" name="Zamora-Chinchipe" pageId="110" pageNumber="109">Zamora-Chinchipe</collectingRegion>
:
</emphasis>
<location id="8E80606AFF96F869FCF1BB214398FCEF" LSID="urn:lsid:plazi:treatment:03F687A7FF95F869FF01BC2645ABFC4F:8E80606AFF96F869FCF1BB214398FCEF" box="[816,915,836,859]" country="Ecuador" name="Zamora" pageId="110" pageNumber="109" stateProvince="Zamora-Chinchipe">Zamora</location>
,
<date id="FFE11071FF96F869FC63BB264414FCEF" box="[930,1055,835,859]" pageId="110" pageNumber="109" value="1978-12-04">
<collectingDate id="EFA5E999FF96F869FC63BB264414FCEF" box="[930,1055,835,859]" pageId="110" pageNumber="109" value="1978-12-04">4.xii.1978</collectingDate>
</date>
,
<collectorName id="26AA5367FF96F869FBEEBB2144C4FCEF" box="[1071,1231,835,859]" pageId="110" pageNumber="109">JJ Anderson</collectorName>
–
<specimenCount id="9D59FD38FF96F869FB1EBB264568FCEF" box="[1247,1379,835,859]" pageId="110" pageNumber="109" type="male">232 males</specimenCount>
,
<specimenCount id="9D59FD38FF96F869FAB2BB26412BFCCF" pageId="110" pageNumber="109" type="female">280 females</specimenCount>
(alcohol) (
<collectionCode id="ED4EAE74FF96F869FE61BB06420BFCCF" box="[416,512,867,891]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="110" pageNumber="109" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B373CECFF96F869FDD2BB06422FFC2F" collectingDate="1990-09-20" collectionCode="N, OS, UMSP" collectorName="Flint &amp; Jr" country="Ecuador" elevation="880" location="Rio Chicana" pageId="110" pageNumber="109" specimenCount="43" specimenCount-female="32" specimenCount-male="11" stateProvince="Zamora-Chinchipe" typeStatus="paratype">
<location id="8E80606AFF96F869FDD2BB0642A5FCCF" LSID="urn:lsid:plazi:treatment:03F687A7FF95F869FF01BC2645ABFC4F:8E80606AFF96F869FDD2BB0642A5FCCF" box="[531,686,867,891]" country="Ecuador" name="Rio Chicana" pageId="110" pageNumber="109" stateProvince="Zamora-Chinchipe">Río Chicaña</location>
,
<locationDeviation id="9A522F93FF96F869FD78BB064398FCCF" box="[697,915,867,891]" location="Yanzatza" pageId="110" pageNumber="109">
<quantity id="4CA79B54FF96F869FD78BB0642FCFCCF" box="[697,759,867,891]" metricMagnitude="3" metricUnit="m" metricValue="9.0" pageId="110" pageNumber="109" unit="km" value="9.0">9 km</quantity>
<collectionCode id="ED4EAE74FF96F869FD3FBB01431FFCCF" box="[766,788,868,891]" country="China" lsid="urn:lsid:biocol.org:col:13092" name="Nanjing University" pageId="110" pageNumber="109" type="Herbarium">N</collectionCode>
Yanzatza
</locationDeviation>
,
<quantity id="4CA79B54FF96F869FC5FBB0643E7FCCF" box="[926,1004,867,891]" metricMagnitude="2" metricUnit="m" metricValue="8.8" pageId="110" pageNumber="109" unit="m" value="880.0">
<elevation id="0072D182FF96F869FC5FBB0643E7FCCF" box="[926,1004,867,891]" metricMagnitude="2" metricUnit="m" metricValue="8.8" pageId="110" pageNumber="109" unit="m" value="880.0">880 m</elevation>
</quantity>
,
<date id="FFE11071FF96F869FC38BB064470FCCF" box="[1017,1147,867,891]" pageId="110" pageNumber="109" value="1990-09-20">
<collectingDate id="EFA5E999FF96F869FC38BB064470FCCF" box="[1017,1147,867,891]" pageId="110" pageNumber="109" value="1990-09-20">20.ix.1990</collectingDate>
</date>
,
<collectionCode id="ED4EAE74FF96F869FB46BB0644A6FCCF" box="[1159,1197,867,891]" country="USA" name="Oregon State University" pageId="110" pageNumber="109">OS</collectionCode>
<collectorName id="26AA5367FF96F869FB75BB0644FEFCCF" box="[1204,1269,867,891]" pageId="110" pageNumber="109">Flint</collectorName>
,
<collectorName id="26AA5367FF96F869FAC1BB064511FCCF" box="[1280,1306,867,891]" pageId="110" pageNumber="109">Jr</collectorName>
–
<specimenCount id="9D59FD38FF96F869FAEDBB064590FCCF" box="[1324,1435,867,891]" pageId="110" pageNumber="109" type="male">11 males</specimenCount>
,
<specimenCount id="9D59FD38FF96F869FF01BBE6414EFC2F" box="[192,325,899,923]" pageId="110" pageNumber="109" type="female">32 females</specimenCount>
(alcohol) (
<collectionCode id="ED4EAE74FF96F869FE07BBE64217FC2F" box="[454,540,899,923]" country="USA" httpUri="http://grbio.org/cool/3avb-bd3e" name="University of Minnesota Insect Collection" pageId="110" pageNumber="109" type="University or college">UMSP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B373CECFF96F869FDF0BBE1455EFC2F" box="[561,1365,899,923]" collectingDate="1978-11-29" collectionCode="NMNH" collectorName="JJ Anderson" country="Ecuador" location="Yanzaza" pageId="110" pageNumber="109" specimenCount="123" specimenCount-female="110" specimenCount-male="13" stateProvince="Zamora-Chinchipe" typeStatus="paratype">
same data as Holotype–
<specimenCount id="9D59FD38FF96F869FC9FBBE643C7FC2F" box="[862,972,899,923]" pageId="110" pageNumber="109" type="male">13 males</specimenCount>
,
<specimenCount id="9D59FD38FF96F869FC19BBE64466FC2F" box="[984,1133,899,923]" pageId="110" pageNumber="109" type="female">110 females</specimenCount>
(alcohol) (
<collectionCode id="ED4EAE74FF96F869FB2CBBE64546FC2F" box="[1261,1357,899,923]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="110" pageNumber="109" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B373CECFF96F869FAA3BBE64558FC0F" collectingDate="1990-09-21" collectionCode="E, OS, NMNH" collectorName="Flint &amp; Jr" country="Ecuador" elevation="980" location="6 km E Zumbi" pageId="110" pageNumber="109" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Zamora-Chinchipe" typeStatus="paratype">
<location id="8E80606AFF96F869FAA3BBE64125FC0F" LSID="urn:lsid:plazi:treatment:03F687A7FF95F869FF01BC2645ABFC4F:8E80606AFF96F869FAA3BBE64125FC0F" country="Ecuador" name="6 km E Zumbi" pageId="110" pageNumber="109" stateProvince="Zamora-Chinchipe">
<quantity id="4CA79B54FF96F869FAA3BBE645ABFC2F" box="[1378,1440,899,923]" metricMagnitude="3" metricUnit="m" metricValue="6.0" pageId="110" pageNumber="109" unit="km" value="6.0">6 km</quantity>
<collectionCode id="ED4EAE74FF96F869FF01BBC140D8FC0F" box="[192,211,932,955]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="110" pageNumber="109" type="Herbarium">E</collectionCode>
Zumbi
</location>
,
<quantity id="4CA79B54FF96F869FEFABBC64182FC0F" box="[315,393,931,955]" metricMagnitude="2" metricUnit="m" metricValue="9.8" pageId="110" pageNumber="109" unit="m" value="980.0">
<elevation id="0072D182FF96F869FEFABBC64182FC0F" box="[315,393,931,955]" metricMagnitude="2" metricUnit="m" metricValue="9.8" pageId="110" pageNumber="109" unit="m" value="980.0">980 m</elevation>
</quantity>
,
<date id="FFE11071FF96F869FE57BBC64212FC0F" box="[406,537,931,955]" pageId="110" pageNumber="109" value="1990-09-21">
<collectingDate id="EFA5E999FF96F869FE57BBC64212FC0F" box="[406,537,931,955]" pageId="110" pageNumber="109" value="1990-09-21">21.ix.1990</collectingDate>
</date>
,
<collectionCode id="ED4EAE74FF96F869FDE7BBC64247FC0F" box="[550,588,931,955]" country="USA" name="Oregon State University" pageId="110" pageNumber="109">OS</collectionCode>
<collectorName id="26AA5367FF96F869FD92BBC6429FFC0F" box="[595,660,931,955]" pageId="110" pageNumber="109">Flint</collectorName>
,
<collectorName id="26AA5367FF96F869FD61BBC642B0FC0F" box="[672,699,931,955]" pageId="110" pageNumber="109">Jr</collectorName>
–
<specimenCount id="9D59FD38FF96F869FD0FBBC64315FC0F" box="[718,798,931,955]" pageId="110" pageNumber="109" type="male">1 male</specimenCount>
(pinned),
<specimenCount id="9D59FD38FF96F869FC5CBBC643F8FC0F" box="[925,1011,931,955]" pageId="110" pageNumber="109" type="male">1 male</specimenCount>
,
<specimenCount id="9D59FD38FF96F869FC3EBBC64462FC0F" box="[1023,1129,931,955]" pageId="110" pageNumber="109" type="female">1 female</specimenCount>
(alcohol) (
<collectionCode id="ED4EAE74FF96F869FB2ABBC64547FC0F" box="[1259,1356,931,955]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="110" pageNumber="109" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BE036B1FF96F869FF31BBA645ABFC4F" blockId="110.[192,1441,195,1019]" pageId="110" pageNumber="109">
<emphasis id="B92BEAA3FF96F869FF31BBA64179FC6F" box="[240,370,963,987]" italics="true" pageId="110" pageNumber="109">Etymology</emphasis>
—This species is named
<taxonomicName id="4C5F4D32FF96F869FD60BBA1431BFC6F" authorityName="Blahnik &amp; Holzenthal" authorityYear="2017" box="[673,784,963,987]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="110" pageNumber="109" phylum="Tracheophyta" rank="species" species="licina">
<emphasis id="B92BEAA3FF96F869FD60BBA1431BFC6F" box="[673,784,963,987]" italics="true" pageId="110" pageNumber="109">M. licina</emphasis>
</taxonomicName>
from the Latin word
<emphasis id="B92BEAA3FF96F869FBD9BBA64467FC6F" box="[1048,1132,963,987]" italics="true" pageId="110" pageNumber="109">licinus</emphasis>
, meaning bent or turned upward, in reference to the strongly upturned ventral lobes of the inferior appendages in this species.
</paragraph>
</subSubSection>
</treatment>
</document>
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