266 lines
33 KiB
XML
266 lines
33 KiB
XML
<document id="E8BFB00647E00EE637AF0821C89A3400" ID-DOI="10.5281/zenodo.176760" ID-GBIF-Dataset="5b1b2b40-7287-41d1-97b2-c70a1a41022c" ID-ISSN="1175-5326" ID-Zenodo-Dep="176760" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459838974701" checkinUser="plazi" docAuthor="Evenhuis, Neal L." docDate="2007" docId="03E378594705FFE5CAFDAF34FE42FE76" docLanguage="en" docName="zt01474p062.pdf" docOrigin="Zootaxa 1474" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Empidideicus rhinoclypeatus Evenhuis, n.sp." docType="treatment" docVersion="9" lastPageNumber="61" masterDocId="FFDA00214704FFE3CA6AA904FFE7FF84" masterDocTitle="A remarkable new species of Empidideicus (Diptera: Mythicomyiidae) from Madagascar" masterLastPageNumber="62" masterPageNumber="55" pageNumber="56" updateTime="1698224054689" updateUser="plazi">
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<mods:titleInfo id="D441B1D98F542A63788F48539C2D4710">
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<mods:title id="19D308C280D00CDDED0303A558C860F2">A remarkable new species of Empidideicus (Diptera: Mythicomyiidae) from Madagascar</mods:title>
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<mods:name id="9AC09A7D5B0F26E91ECE13C60EFFB92D" type="personal">
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<mods:namePart id="1DB5009924AEA61190146A9A83CFE0EF">Evenhuis, Neal L.</mods:namePart>
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<mods:title id="B5A247A368C405C259A890FE8B295DA4">Zootaxa</mods:title>
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<mods:part id="02E60D55A4B00A03ACD021602AF5D135">
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<mods:date id="B5AD5181C49CC6DF47CB3DA39192720F">2007</mods:date>
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<mods:number id="E97AE04EED554D97B6AC31A073B124E9">1474</mods:number>
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<mods:identifier id="8F1D3F16E3EC77EF074A0C647992D99F" type="DOI">10.5281/zenodo.176760</mods:identifier>
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<mods:identifier id="7286DC636FAFF0A4DF0398141FD224BB" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="673CD9CE873D7AE83AC049C50968B84B" type="Zenodo-Dep">176760</mods:identifier>
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<treatment id="03E378594705FFE5CAFDAF34FE42FE76" ID-DOI="http://doi.org/10.5281/zenodo.6248709" ID-GBIF-Taxon="119359773" ID-Zenodo-Dep="6248709" LSID="urn:lsid:plazi:treatment:03E378594705FFE5CAFDAF34FE42FE76" httpUri="http://treatment.plazi.org/id/03E378594705FFE5CAFDAF34FE42FE76" lastPageId="6" lastPageNumber="61" pageId="1" pageNumber="56">
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<subSubSection id="C3509AC44705FFE2CAFDAF34FD5DF9CD" box="[151,698,1583,1610]" pageId="1" pageNumber="56" type="nomenclature">
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<paragraph id="8BF5C94F4705FFE2CAFDAF34FD5DF9CD" blockId="1.[151,698,1583,1610]" box="[151,698,1583,1610]" pageId="1" pageNumber="56">
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<heading id="D0BD7E234705FFE2CAFDAF34FD5DF9CD" bold="true" box="[151,698,1583,1610]" fontSize="11" level="1" pageId="1" pageNumber="56" reason="1">
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<emphasis id="B93E155D4705FFE2CAFDAF34FD5DF9CD" bold="true" box="[151,698,1583,1610]" pageId="1" pageNumber="56">
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<taxonomicName id="4C4AB2CC4705FFE2CAFDAF34FD95F9CD" authority="Evenhuis" authorityName="Evenhuis" box="[151,626,1583,1610]" class="Insecta" family="Bombyliidae" genus="Empidideicus" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="56" phylum="Arthropoda" rank="species" species="rhinoclypeatus" status="sp. nov.">
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<emphasis id="B93E155D4705FFE2CAFDAF34FE11F9CE" bold="true" box="[151,502,1584,1610]" italics="true" pageId="1" pageNumber="56">Empidideicus rhinoclypeatus</emphasis>
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Evenhuis
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</taxonomicName>
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,
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<taxonomicNameLabel id="A20DA8264705FFE2C815AF2BFD5DF9CD" box="[639,698,1583,1609]" pageId="1" pageNumber="56" rank="species">n.sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3509AC44705FFE1CAFDAF7EFC8DFE86" lastPageId="2" lastPageNumber="57" pageId="1" pageNumber="56" type="diagnosis">
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<paragraph id="8BF5C94F4705FFE2CAFDAF7EFEF4F86A" blockId="1.[151,1436,1658,1764]" lastBlockId="1.[151,1437,1830,2030]" pageId="1" pageNumber="56">
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<emphasis id="B93E155D4705FFE2CAFDAF7EFEF7F910" bold="true" box="[151,272,1658,1684]" pageId="1" pageNumber="56">Diagnosis</emphasis>
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: This new species belongs in the group of species with a closed cell dm (
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<taxonomicName id="4C4AB2CC4705FFE2CE04AF7FFAFEF910" box="[1134,1305,1659,1684]" class="Insecta" family="Mythicomyiidae" genus="Anomaloptilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="56" phylum="Arthropoda" rank="genus">
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<emphasis id="B93E155D4705FFE2CE04AF7FFAFEF910" box="[1134,1305,1659,1684]" italics="true" pageId="1" pageNumber="56">Anomaloptilus</emphasis>
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</taxonomicName>
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olim). It is easily separated from all
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<taxonomicName id="4C4AB2CC4705FFE2CBD7AFA7FDBAF938" box="[445,605,1699,1724]" class="Insecta" family="Bombyliidae" genus="Empidideicus" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="56" phylum="Arthropoda" rank="genus">
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<emphasis id="B93E155D4705FFE2CBD7AFA7FDBAF938" box="[445,605,1699,1724]" italics="true" pageId="1" pageNumber="56">Empidideicus</emphasis>
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</taxonomicName>
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by the elongate clypeus, extending below the oral margin and one-half the length of the labrum. The female spermatheca resembles
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<taxonomicName id="4C4AB2CC4705FFE2C909AFCFFBC0F960" authority="Frey" authorityName="Frey" box="[867,1063,1738,1764]" class="Insecta" family="Bombyliidae" genus="Empidideicus" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="56" phylum="Arthropoda" rank="species" species="insularis">
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<emphasis id="B93E155D4705FFE2C909AFCFFC0DF960" box="[867,1002,1739,1764]" italics="true" pageId="1" pageNumber="56">E. insularis</emphasis>
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Frey
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</taxonomicName>
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(see
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<bibRefCitation id="EFDBB4BE4705FFE2CE0FAFCEFACEF960" author="Greathead" box="[1125,1321,1738,1764]" pageId="1" pageNumber="61" refString="Greathead, D. J. (1986) The genus Empidideicus Becker (Diptera: Bombyliidae) in the Cape Verde Islands. Entomologist's Monthly Magazine, 122, 85 - 88." type="journal article" year="1986">Greathead, 1986</bibRefCitation>
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, fig. 3) in 1.
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<emphasis id="B93E155D4705FFE2CAA7AE22FED3F8B9" box="[205,308,1830,1853]" italics="true" pageId="1" pageNumber="56">Crytoides</emphasis>
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Engel and
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<taxonomicName id="4C4AB2CC4705FFE2CBC5AE22FD75F8B9" authority="Hesse" authorityName="Hesse" box="[431,658,1830,1853]" class="Insecta" family="Mythicomyiidae" genus="Anomaloptilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="56" phylum="Arthropoda" rank="genus">
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<emphasis id="B93E155D4705FFE2CBC5AE22FDACF8B9" box="[431,587,1830,1853]" italics="true" pageId="1" pageNumber="56">Anomaloptilus</emphasis>
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Hesse
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</taxonomicName>
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were treated as junior synonyms of
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<taxonomicName id="4C4AB2CC4705FFE2CE72AE22FB4EF8B9" box="[1048,1193,1830,1853]" class="Insecta" family="Bombyliidae" genus="Empidideicus" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="56" phylum="Arthropoda" rank="genus">
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<emphasis id="B93E155D4705FFE2CE72AE22FB4EF8B9" box="[1048,1193,1830,1853]" italics="true" pageId="1" pageNumber="56">Empidideicus</emphasis>
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</taxonomicName>
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in
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<bibRefCitation id="EFDBB4BE4705FFE2CEA6AE22FEF9F8DB" author="Greathead" pageId="1" pageNumber="61" refString="Greathead, D. J. & Evenhuis, N. L. (2001 a) Bombylioidea (Diptera: Bombyliidae; Mythicomyiidae) from the island of Sokotra. Zootaxa, 14, 1 - 11." type="journal article" year="2001" yearSuffix="a">Greathead & Evenhuis (2001a)</bibRefCitation>
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based on characters previously used to distinguish the three that were shown to be variable. Further study has shown that there are additional non-variable characters that appear to confidently distinguish these genus-groups. However, until revisionary work on the genus is completed, it is prudent to leave these as species groupings and to not formally resurrect them from synonymy. [Only undescribed species keying to
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<taxonomicName id="4C4AB2CC4705FFE2C9B9AEB5FBDCF84C" box="[979,1083,1969,1992]" class="Insecta" family="Raphitomidae" genus="Cyrtoides" higherTaxonomySource="CoL" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="56" phylum="Arthropoda" rank="genus">
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<emphasis id="B93E155D4705FFE2C9B9AEB5FBDCF84C" box="[979,1083,1969,1992]" italics="true" pageId="1" pageNumber="56">Cyrtoides</emphasis>
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</taxonomicName>
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olim are known from the regions in this key.]
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</paragraph>
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<paragraph id="8BF5C94F4706FFE1CAFDA99CFC8DFE86" blockId="2.[151,1437,152,618]" pageId="2" pageNumber="57">
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having basal striations and a hole at the apex or the reservoir but
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<taxonomicName id="4C4AB2CC4706FFE1C9E1A99CFBF6FF35" box="[907,1041,152,177]" class="Insecta" family="Bombyliidae" genus="Empidideicus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="57" phylum="Arthropoda" rank="species" species="insularis">
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<emphasis id="B93E155D4706FFE1C9E1A99CFBF6FF35" box="[907,1041,152,177]" italics="true" pageId="2" pageNumber="57">E. insularis</emphasis>
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</taxonomicName>
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has cell dm open apically (closed in
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<taxonomicName id="4C4AB2CC4706FFE1CADCA9C4FE63FF5D" box="[182,388,192,217]" class="Insecta" family="Bombyliidae" genus="Empidideicus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="57" phylum="Arthropoda" rank="species" species="rhinoclypeatus">
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<emphasis id="B93E155D4706FFE1CADCA9C4FE63FF5D" box="[182,388,192,217]" italics="true" pageId="2" pageNumber="57">E. rhinoclypeatus</emphasis>
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</taxonomicName>
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). This one of the largest known species in the genus (most are less than 1.0 mm in body length) and is thus easily spotted in collections due to its size.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3509AC44706FFE5CAACA80BFC4EFEFE" lastPageId="6" lastPageNumber="61" pageId="2" pageNumber="57" type="description">
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<paragraph id="8BF5C94F4706FFE1CAACA80BFD8BFDEE" blockId="2.[151,1437,152,618]" pageId="2" pageNumber="57">
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<emphasis id="B93E155D4706FFE1CAACA80BFEB0FEAD" bold="true" box="[198,343,271,297]" pageId="2" pageNumber="57">Description</emphasis>
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:
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<emphasis id="B93E155D4706FFE1CB03A80BFE4FFEAD" bold="true" box="[361,424,271,297]" pageId="2" pageNumber="57">Male</emphasis>
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. Length:
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<quantity id="4CB264AA4706FFE1C875A814FD28FEAE" box="[543,719,272,298]" metricMagnitude="-3" metricUnit="m" metricValue="1.915" metricValueMax="2.38" metricValueMin="1.45" pageId="2" pageNumber="57" unit="mm" value="1.915" valueMax="2.38" valueMin="1.45">1.45–2.38 mm</quantity>
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. Wing:
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<quantity id="4CB264AA4706FFE1C95AA814FC07FEAE" box="[816,992,272,298]" metricMagnitude="-3" metricUnit="m" metricValue="1.975" metricValueMax="2.45" metricValueMin="1.5" pageId="2" pageNumber="57" unit="mm" value="1.975" valueMax="2.45" valueMin="1.5">1.50–2.45 mm</quantity>
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.
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<emphasis id="B93E155D4706FFE1C985A814FBC9FEAD" box="[1007,1070,272,297]" italics="true" pageId="2" pageNumber="57">Head</emphasis>
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(Figs. 1–3): longer than high; eyes dichoptic, separated at vertex by 1.5 times distance between lateral ocelli; occiput and postgena black; front slightly depressed medially, white with subrectangular brown area immediately below ocellar tubercle; face white, tip of oral margin tan to brown; antennae dark brown, set in deep pocket; scape short, subtrapezoidal; pedicel subconical, wider than long; first flagellomere lanceolate, length 3.5 times width; second flagellomere one-half length of first flagellomere, with minute transparent apical style; mentum black; clypeus dark brown, elongate, extending well beyond tip of oral margin, length subequal to head height; proboscis dark brown to black, length 2.5 times head length; labrum sclerotized, stiff, pointed apically, length subequal to clypeus; palpus not evident.
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</paragraph>
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<paragraph id="8BF5C94F4706FFE1CAFDAF5AFEFFF9F1" blockId="2.[151,280,1630,1654]" box="[151,280,1630,1654]" pageId="2" pageNumber="57">
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<emphasis id="B93E155D4706FFE1CAFDAF5AFEF5F9F2" bold="true" box="[151,274,1630,1654]" pageId="2" pageNumber="57">FIGURE 1</emphasis>
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.
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</paragraph>
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<paragraph id="8BF5C94F4706FFE1CAADAFADFBD1F85F" blockId="2.[151,1436,1705,2011]" pageId="2" pageNumber="57">
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<emphasis id="B93E155D4706FFE1CAADAFADFEFBF946" box="[199,284,1705,1730]" italics="true" pageId="2" pageNumber="57">Thorax</emphasis>
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(
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<figureCitation id="1371D5CA4706FFE1CB47AFADFE79F947" box="[301,414,1705,1731]" captionStart="FIGURES 4 – 7" captionStartId="4.[151,269,1618,1642]" captionTargetBox="[156,1429,194,1592]" captionTargetId="figure@4.[156,1431,194,1594]" captionTargetPageId="4" captionText="FIGURES 4 – 7. Empidideicus rhinoclypeatus thoraces, showing differences in mesonotal patterning. 4. Typical male form. 5. Male form with markings. 6. Typical female form. 7. Female form with reduced admedian vittae." httpUri="https://zenodo.org/record/176762/files/figure.png" pageId="2" pageNumber="57">Figs. 4–7</figureCitation>
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,
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<figureCitation id="1371D5CA4706FFE1CBC1AFADFE0DF947" box="[427,490,1705,1731]" captionStart="FIGURES 8 – 10" captionStartId="5.[153,272,1805,1829]" captionTargetBox="[156,1430,434,1780]" captionTargetId="figure@5.[156,1431,434,1781]" captionTargetPageId="5" captionText="FIGURES 8 – 10. Empidideicus rhinoclypeatus. 8. Wing. 9. Male abdomen showing typical pattern. 10. Female abdomen showing typical pattern." httpUri="https://zenodo.org/record/176763/files/figure.png" pageId="2" pageNumber="57">9–10</figureCitation>
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). Mesonotum shiny black with yellow markings in following areas: prescutellar area extending anteriorly for a short distance as traces of admedian vittae; dorsolaterally from humeral callus to transverse suture; post alar callus as anterolateral extension of prescutellar mark; scutellum yellow; mesonotal disc and dorsum of scutellum with short fine brown hairs, small patch of fine minute brown hairs on dorsolateral corner of anepisternum and upper half of propleuron, otherwise, pleura bare; pleura dark brown with pale yellow to white spots on following: posterodorsal corner of anepisternum and anepimeron, laterally on propleuron, upper half of meron; coxae and basal three-fourths of femora and tarsal segments 4 and 5 brown, remainder of legs yellow; halter stem grayish brown, knob white.
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</paragraph>
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<caption id="DF3599C74707FFE0CAFDAC47FDEBFAF9" httpUri="https://zenodo.org/record/176761/files/figure.png" pageId="3" pageNumber="58" targetBox="[370,1212,198,1294]" targetPageId="3">
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<paragraph id="8BF5C94F4707FFE0CAFDAC47FDEBFAF9" blockId="3.[151,1436,1347,1405]" pageId="3" pageNumber="58">
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<emphasis id="B93E155D4707FFE0CAFDAC47FED9FADF" bold="true" box="[151,318,1347,1371]" pageId="3" pageNumber="58">FIGURES 2–3</emphasis>
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. Heads of
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<taxonomicName id="4C4AB2CC4707FFE0CBDCAC47FDA0FADE" box="[438,583,1347,1370]" class="Insecta" family="Bombyliidae" genus="Empidideicus" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="58" phylum="Arthropoda" rank="genus">
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<emphasis id="B93E155D4707FFE0CBDCAC47FDA0FADE" box="[438,583,1347,1370]" italics="true" pageId="3" pageNumber="58">Empidideicus</emphasis>
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</taxonomicName>
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, showing differences in size and sclerotization of clypeus (arrow).
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<emphasis id="B93E155D4707FFE0CF4EAC47FAD6FADF" bold="true" box="[1316,1329,1347,1371]" pageId="3" pageNumber="58">2</emphasis>
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.
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<taxonomicName id="4C4AB2CC4707FFE0CF2AAC47FECEFAF9" authority="Hesse." authorityName="Hesse." class="Insecta" family="Bombyliidae" genus="Empidideicus" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="58" phylum="Arthropoda" rank="species" species="celluliferus">
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<emphasis id="B93E155D4707FFE0CF2AAC47FF3CFAF9" italics="true" pageId="3" pageNumber="58">E. celluliferus</emphasis>
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Hesse.
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</taxonomicName>
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<emphasis id="B93E155D4707FFE0CB44AC61FEDCFAF9" bold="true" box="[302,315,1381,1405]" pageId="3" pageNumber="58">3</emphasis>
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.
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<taxonomicName id="4C4AB2CC4707FFE0CB23AC62FDE2FAF9" box="[329,517,1382,1405]" class="Insecta" family="Bombyliidae" genus="Empidideicus" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="58" phylum="Arthropoda" rank="species" species="rhinoclypeatus">
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<emphasis id="B93E155D4707FFE0CB23AC62FDE2FAF9" box="[329,517,1382,1405]" italics="true" pageId="3" pageNumber="58">E. rhinoclypeatus</emphasis>
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</taxonomicName>
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.
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</paragraph>
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</caption>
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<paragraph id="8BF5C94F4707FFE0CAADACBAFAEDF9FC" blockId="3.[151,1437,1470,2016]" pageId="3" pageNumber="58">
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<emphasis id="B93E155D4707FFE0CAADACBAFEE5FA53" box="[199,258,1470,1495]" italics="true" pageId="3" pageNumber="58">Wing</emphasis>
|
||
(
|
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<figureCitation id="1371D5CA4707FFE0CB7FACBAFEBAFA5C" box="[277,349,1470,1496]" captionStart="FIGURES 8 – 10" captionStartId="5.[153,272,1805,1829]" captionTargetBox="[156,1430,434,1780]" captionTargetId="figure@5.[156,1431,434,1781]" captionTargetPageId="5" captionText="FIGURES 8 – 10. Empidideicus rhinoclypeatus. 8. Wing. 9. Male abdomen showing typical pattern. 10. Female abdomen showing typical pattern." httpUri="https://zenodo.org/record/176763/files/figure.png" pageId="3" pageNumber="58">Fig. 8</figureCitation>
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). Elongate, length 3.5 times width; subhyaline with pale brownish tinge; veins brown, well sclerotized except vein separating basal cells and A1; costa ends slightly beyond end of R4+5; vein Sc incomplete; Rs evanescent at connection with R1; R4+5 slightly bowed; vein M1 slightly curved at wing margin; M2 straight; cell dm closed by apical crossvein; A1 effaced, slightly sinuous to wing margin; fringe of hair on posterior margin of wing well shortest at wing tip becoming longer and more sparse toward base.
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</paragraph>
|
||
<paragraph id="8BF5C94F4707FFE0CAADAF82FD78F89C" blockId="3.[151,1437,1470,2016]" pageId="3" pageNumber="58">
|
||
<emphasis id="B93E155D4707FFE0CAADAF82FED1F91B" box="[199,310,1670,1695]" italics="true" pageId="3" pageNumber="58">Abdomen</emphasis>
|
||
(
|
||
<figureCitation id="1371D5CA4707FFE0CB2CAF82FE69F924" box="[326,398,1670,1696]" captionStart="FIGURES 8 – 10" captionStartId="5.[153,272,1805,1829]" captionTargetBox="[156,1430,434,1780]" captionTargetId="figure@5.[156,1431,434,1781]" captionTargetPageId="5" captionText="FIGURES 8 – 10. Empidideicus rhinoclypeatus. 8. Wing. 9. Male abdomen showing typical pattern. 10. Female abdomen showing typical pattern." httpUri="https://zenodo.org/record/176763/files/figure.png" pageId="3" pageNumber="58">Fig. 9</figureCitation>
|
||
). Dorsum black, concolorous with central mesonotal color; tergite I white laterally; posterior margins of tergites II-VII and lateral portions white; when viewed laterally abdomen with lateral black band consisting of black color on extreme lateral tips of tergites and sternites; remainder of sternites white; tergites with sparsely scattered minute hairs.
|
||
</paragraph>
|
||
<paragraph id="8BF5C94F4707FFE0CAADAE22FD83F864" blockId="3.[151,1437,1470,2016]" pageId="3" pageNumber="58">
|
||
<emphasis id="B93E155D4707FFE0CAADAE22FED1F8BB" box="[199,310,1830,1855]" italics="true" pageId="3" pageNumber="58">Genitalia</emphasis>
|
||
(
|
||
<figureCitation id="1371D5CA4707FFE0CB2FAE22FE7CF8C4" box="[325,411,1830,1856]" captionStart="FIGURES 11 – 12" captionStartId="6.[151,269,1195,1219]" captionTargetBox="[171,1426,571,1168]" captionTargetId="figure@6.[156,1443,554,1185]" captionTargetPageId="6" captionText="FIGURES 11 – 12. Empidideicus rhinoclypeatus, genitalia. 11. Male genitalia; A. lateral view. B. Ventral view. 12. Female genitalia with detail of spermathecal reservoir." httpUri="https://zenodo.org/record/176764/files/figure.png" pageId="3" pageNumber="58">Fig. 11</figureCitation>
|
||
: A, B). Hypopygium shiny brown. Gonocoxa subtriangular in lateral view with rounded subrectangular gonostylus; gonocoxal apodemes extending below gonocoxa, flared apically; aedeagal bulb large, rounded; aedeagal apodeme subequal in length to gonocoxal apodeme, peanut-shaped with short lanceolate lateral rami; epiphallus slightly longer than gonocoxa, with hooked apex; epandrium subrectangular with large pointed dorsolateral process.
|
||
</paragraph>
|
||
<caption id="DF3599C74700FFE7CAFDAF56FB18F90F" httpUri="https://zenodo.org/record/176762/files/figure.png" pageId="4" pageNumber="59" targetBox="[156,1429,194,1592]" targetPageId="4">
|
||
<paragraph id="8BF5C94F4700FFE7CAFDAF56FB18F90F" blockId="4.[151,1436,1618,1676]" pageId="4" pageNumber="59">
|
||
<emphasis id="B93E155D4700FFE7CAFDAF56FED8F9EE" bold="true" box="[151,319,1618,1642]" pageId="4" pageNumber="59">FIGURES 4–7</emphasis>
|
||
.
|
||
<taxonomicName id="4C4AB2CC4700FFE7CB25AF56FD6FF9ED" box="[335,648,1618,1641]" class="Insecta" family="Bombyliidae" genus="Empidideicus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="59" phylum="Arthropoda" rank="species" species="rhinoclypeatus">
|
||
<emphasis id="B93E155D4700FFE7CB25AF56FD6FF9ED" box="[335,648,1618,1641]" italics="true" pageId="4" pageNumber="59">Empidideicus rhinoclypeatus</emphasis>
|
||
</taxonomicName>
|
||
thoraces, showing differences in mesonotal patterning.
|
||
<emphasis id="B93E155D4700FFE7CE9BAF56FB19F9EE" bold="true" box="[1265,1278,1618,1642]" pageId="4" pageNumber="59">4</emphasis>
|
||
. Typical male form.
|
||
<emphasis id="B93E155D4700FFE7CAB2AF70FF02F908" bold="true" box="[216,229,1652,1676]" pageId="4" pageNumber="59">5</emphasis>
|
||
. Male form with markings.
|
||
<emphasis id="B93E155D4700FFE7C87BAF70FDF9F908" bold="true" box="[529,542,1652,1676]" pageId="4" pageNumber="59">6</emphasis>
|
||
. Typical female form.
|
||
<emphasis id="B93E155D4700FFE7C978AF70FCF8F908" bold="true" box="[786,799,1652,1676]" pageId="4" pageNumber="59">7</emphasis>
|
||
. Female form with reduced admedian vittae.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BF5C94F4700FFE7CAADAFE3FED7F84D" blockId="4.[151,1437,1767,1993]" pageId="4" pageNumber="59">
|
||
<emphasis id="B93E155D4700FFE7CAADAFE3FEC5F885" bold="true" box="[199,290,1767,1793]" pageId="4" pageNumber="59">Female</emphasis>
|
||
: As in male except dark coloration of thorax and abdomen brown. Abdominal dark coloration interrupted medially by longitudinal white stripe (
|
||
<figureCitation id="1371D5CA4700FFE7C89AAE0BFCACF8AD" box="[752,843,1807,1833]" captionStart="FIGURES 8 – 10" captionStartId="5.[153,272,1805,1829]" captionTargetBox="[156,1430,434,1780]" captionTargetId="figure@5.[156,1431,434,1781]" captionTargetPageId="5" captionText="FIGURES 8 – 10. Empidideicus rhinoclypeatus. 8. Wing. 9. Male abdomen showing typical pattern. 10. Female abdomen showing typical pattern." httpUri="https://zenodo.org/record/176763/files/figure.png" pageId="4" pageNumber="59">Fig. 10</figureCitation>
|
||
). Genitalia (
|
||
<figureCitation id="1371D5CA4700FFE7C98AAE0BFBDCF8AD" box="[992,1083,1807,1833]" captionStart="FIGURES 11 – 12" captionStartId="6.[151,269,1195,1219]" captionTargetBox="[171,1426,571,1168]" captionTargetId="figure@6.[156,1443,554,1185]" captionTargetPageId="6" captionText="FIGURES 11 – 12. Empidideicus rhinoclypeatus, genitalia. 11. Male genitalia; A. lateral view. B. Ventral view. 12. Female genitalia with detail of spermathecal reservoir." httpUri="https://zenodo.org/record/176764/files/figure.png" pageId="4" pageNumber="59">Fig. 12</figureCitation>
|
||
): spermatheca spheroid-ellipsoid, sclerotized brown, longer than wide, with striations on basal half, large hole apically; apical spermathecal duct ca. 1.5 x length of spermatheca, thin, transparent; sperm pump 3–4 times length of apical duct, transparent, not sclerotized apically or basally, slightly flared and rounded apically; basal duct confluent with sperm pump.
|
||
</paragraph>
|
||
<paragraph id="8BF5C94F4701FFE6CAADA993FC73FEFE" blockId="5.[151,1436,151,378]" pageId="5" pageNumber="60">
|
||
<emphasis id="B93E155D4701FFE6CAADA993FEE9FF35" bold="true" box="[199,270,151,177]" pageId="5" pageNumber="60">
|
||
<typeStatus id="54F177ED4701FFE6CAADA993FEE9FF35" box="[199,270,151,177]" pageId="5" pageNumber="60">Types</typeStatus>
|
||
</emphasis>
|
||
.
|
||
<typeStatus id="54F177ED4701FFE6CB4BA99CFE69FF36" box="[289,398,152,178]" pageId="5" pageNumber="60" type="holotype">Holotype</typeStatus>
|
||
ɗ and 43ɗ, 22Ψ
|
||
<typeStatus id="54F177ED4701FFE6C803A99CFD3EFF36" box="[617,729,152,178]" pageId="5" pageNumber="60" type="paratype">paratypes</typeStatus>
|
||
from
|
||
<emphasis id="B93E155D4701FFE6C94CA993FC1AFF35" bold="true" box="[806,1021,151,177]" pageId="5" pageNumber="60">
|
||
<collectingCountry id="F35D89DF4701FFE6C94CA993FC1AFF35" box="[806,1021,151,177]" name="Madagascar" pageId="5" pageNumber="60">MADAGASCAR</collectingCountry>
|
||
</emphasis>
|
||
: Tulear Province: Cape Ste Marie Special Reserve,
|
||
<quantity id="4CB264AA4701FFE6CB08A9C4FE47FF5E" box="[354,416,192,218]" metricMagnitude="1" metricUnit="m" metricValue="3.7" pageId="5" pageNumber="60" unit="m" value="37.0">37 m</quantity>
|
||
,
|
||
<quantity id="4CB264AA4701FFE6CBC6A9C4FE10FF5E" box="[428,503,192,218]" metricMagnitude="4" metricUnit="m" metricValue="7.4" pageId="5" pageNumber="60" unit="km" value="74.0">74 km</quantity>
|
||
S. of Tsihombe,
|
||
<geoCoordinate id="EE7EAF884701FFE6C8D5A9C4FCBFFF5E" box="[703,856,192,218]" direction="south" orientation="latitude" pageId="5" pageNumber="60" precision="1" value="-25.587267">25°35.236' S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE7EAF884701FFE6C90FA9C4FC0DFF5E" box="[869,1002,192,218]" direction="east" orientation="longitude" pageId="5" pageNumber="60" precision="9" value="45.163">45°09.78'E</geoCoordinate>
|
||
,
|
||
<date id="FFF4EF8F4701FFE6C99CA9C4FB4EFF5E" box="[1014,1193,192,218]" pageId="5" pageNumber="60" value="2003-02-05" valueMax="2003-02-15" valueMin="2003-02-05">5–15 Feb 2003</date>
|
||
, M. Irwin, F. Parker, Malaise trap, spiny bush.
|
||
<emphasis id="B93E155D4701FFE6CBA9A9ECFD66FE85" box="[451,641,232,257]" italics="true" pageId="5" pageNumber="60">
|
||
Other
|
||
<typeStatus id="54F177ED4701FFE6C864A9ECFD66FE85" box="[526,641,232,257]" pageId="5" pageNumber="60" type="paratype">paratypes</typeStatus>
|
||
</emphasis>
|
||
:
|
||
<emphasis id="B93E155D4701FFE6C8FBA9E3FC8FFE85" bold="true" box="[657,872,231,257]" pageId="5" pageNumber="60">
|
||
<collectingCountry id="F35D89DF4701FFE6C8FBA9E3FC8FFE85" box="[657,872,231,257]" name="Madagascar" pageId="5" pageNumber="60">MADAGASCAR</collectingCountry>
|
||
</emphasis>
|
||
: Tulear Province: 2ɗ, 3Ψ, Mikea Forest, NW of Manombo,
|
||
<date id="FFF4EF8F4701FFE6CB74A814FE16FEAE" box="[286,497,272,298]" pageId="5" pageNumber="60" value="2002-04-18" valueMax="2002-04-29" valueMin="2002-04-18">18–29 April 2002</date>
|
||
, M.E. Irwin, R. Harin’Hala, Malaise trap, deciduous dry forest; 3ɗ, Andohahela National Park, Parcel II,
|
||
<date id="FFF4EF8F4701FFE6CBAAA83CFD6FFED6" box="[448,648,312,338]" pageId="5" pageNumber="60" value="2002-12-16" valueMax="2002-12-17" valueMin="2002-12-16">16–17 Dec 2002</date>
|
||
, M.E. Irwin, F.D. Parker, R. Harin’Hala, Malaise trap, transitional forest.
|
||
<typeStatus id="54F177ED4701FFE6CA82A864FEB2FEFE" box="[232,341,352,378]" pageId="5" pageNumber="60" type="holotype">Holotype</typeStatus>
|
||
in CAS.
|
||
<typeStatus id="54F177ED4701FFE6CBA8A864FDD3FEFE" box="[450,564,352,378]" pageId="5" pageNumber="60" type="paratype">Paratypes</typeStatus>
|
||
in CAS, MNHN, and BPBM.
|
||
</paragraph>
|
||
<caption id="DF3599C74701FFE6CAF3AE09FE7EF8C3" httpUri="https://zenodo.org/record/176763/files/figure.png" pageId="5" pageNumber="60" targetBox="[156,1430,434,1780]" targetPageId="5">
|
||
<paragraph id="8BF5C94F4701FFE6CAF3AE09FE7EF8C3" blockId="5.[151,1435,1805,1863]" pageId="5" pageNumber="60">
|
||
<emphasis id="B93E155D4701FFE6CAF3AE09FEADF8A1" bold="true" box="[153,330,1805,1829]" pageId="5" pageNumber="60">FIGURES 8–10</emphasis>
|
||
.
|
||
<taxonomicName id="4C4AB2CC4701FFE6CB3DAE0AFD6BF8A1" box="[343,652,1806,1829]" class="Insecta" family="Bombyliidae" genus="Empidideicus" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="60" phylum="Arthropoda" rank="species" species="rhinoclypeatus">
|
||
<emphasis id="B93E155D4701FFE6CB3DAE0AFD6BF8A1" box="[343,652,1806,1829]" italics="true" pageId="5" pageNumber="60">Empidideicus rhinoclypeatus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="B93E155D4701FFE6C8F3AE09FD41F8A1" bold="true" box="[665,678,1805,1829]" pageId="5" pageNumber="60">8</emphasis>
|
||
. Wing.
|
||
<emphasis id="B93E155D4701FFE6C890AE09FCE0F8A1" bold="true" box="[762,775,1805,1829]" pageId="5" pageNumber="60">9</emphasis>
|
||
. Male abdomen showing typical pattern.
|
||
<emphasis id="B93E155D4701FFE6CED4AE09FB3FF8A1" bold="true" box="[1214,1240,1805,1829]" pageId="5" pageNumber="60">10</emphasis>
|
||
. Female abdomen showing typical pattern.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BF5C94F4701FFE5CAADAEA6FC4EFEFE" blockId="5.[151,1436,1954,2021]" lastBlockId="6.[151,1437,152,498]" lastPageId="6" lastPageNumber="61" pageId="5" pageNumber="60">
|
||
<emphasis id="B93E155D4701FFE6CAADAEA6FEDAF838" bold="true" box="[199,317,1954,1980]" pageId="5" pageNumber="60">Variation</emphasis>
|
||
. The sexual dimorphism exhibited in this species is most evident in the general brown coloration of the thorax and abdomen in females (black in males) and the female abdomen having the transverse dark bands on the tergites interrupted medially (continuous in males). In addition, there is some variation among specimens in the pattern of the mesonotum. In males the typical form (
|
||
<figureCitation id="1371D5CA4702FFE5CE56A9C4FB63FF5E" box="[1084,1156,192,218]" captionStart="FIGURES 4 – 7" captionStartId="4.[151,269,1618,1642]" captionTargetBox="[156,1429,194,1592]" captionTargetId="figure@4.[156,1431,194,1594]" captionTargetPageId="4" captionText="FIGURES 4 – 7. Empidideicus rhinoclypeatus thoraces, showing differences in mesonotal patterning. 4. Typical male form. 5. Male form with markings. 6. Typical female form. 7. Female form with reduced admedian vittae." httpUri="https://zenodo.org/record/176762/files/figure.png" pageId="6" pageNumber="61">Fig. 4</figureCitation>
|
||
) is black with has little evidence of admedian vittae or prescutellar yellow marking, this can vary to clearly marked prescutellar yellow marking (
|
||
<figureCitation id="1371D5CA4702FFE5CB2BA814FE6DFEAE" box="[321,394,272,298]" captionStart="FIGURES 4 – 7" captionStartId="4.[151,269,1618,1642]" captionTargetBox="[156,1429,194,1592]" captionTargetId="figure@4.[156,1431,194,1594]" captionTargetPageId="4" captionText="FIGURES 4 – 7. Empidideicus rhinoclypeatus thoraces, showing differences in mesonotal patterning. 4. Typical male form. 5. Male form with markings. 6. Typical female form. 7. Female form with reduced admedian vittae." httpUri="https://zenodo.org/record/176762/files/figure.png" pageId="6" pageNumber="61">Fig. 5</figureCitation>
|
||
) and some specimens showing traces of admedian vittae. In females, the typical form (
|
||
<figureCitation id="1371D5CA4702FFE5CAF5A83CFF01FED6" box="[159,230,312,338]" captionStart="FIGURES 4 – 7" captionStartId="4.[151,269,1618,1642]" captionTargetBox="[156,1429,194,1592]" captionTargetId="figure@4.[156,1431,194,1594]" captionTargetPageId="4" captionText="FIGURES 4 – 7. Empidideicus rhinoclypeatus thoraces, showing differences in mesonotal patterning. 4. Typical male form. 5. Male form with markings. 6. Typical female form. 7. Female form with reduced admedian vittae." httpUri="https://zenodo.org/record/176762/files/figure.png" pageId="6" pageNumber="61">Fig. 6</figureCitation>
|
||
) has clearly marked admedian vittae as well as distinct yellow prescutellar marking. This can vary to the extreme form of admedian vittae considerably reduced (
|
||
<figureCitation id="1371D5CA4702FFE5C939A864FC7CFEFE" box="[851,923,352,378]" captionStart="FIGURES 4 – 7" captionStartId="4.[151,269,1618,1642]" captionTargetBox="[156,1429,194,1592]" captionTargetId="figure@4.[156,1431,194,1594]" captionTargetPageId="4" captionText="FIGURES 4 – 7. Empidideicus rhinoclypeatus thoraces, showing differences in mesonotal patterning. 4. Typical male form. 5. Male form with markings. 6. Typical female form. 7. Female form with reduced admedian vittae." httpUri="https://zenodo.org/record/176762/files/figure.png" pageId="6" pageNumber="61">Fig. 7</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3509AC44702FFE5CAADA883FE42FE76" pageId="6" pageNumber="61" type="etymology">
|
||
<paragraph id="8BF5C94F4702FFE5CAADA883FE42FE76" blockId="6.[151,1437,152,498]" pageId="6" pageNumber="61">
|
||
<emphasis id="B93E155D4702FFE5CAADA883FEABFE25" bold="true" box="[199,332,391,417]" pageId="6" pageNumber="61">Etymology</emphasis>
|
||
. The specific epithet derives from Greek, r hi nos = “nose” + cl ypeat us = “provide with a shield”; referring to the elongate clypeus of this new species, which is reminiscent of the nose guards of ancient Greek helmets.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |