261 lines
46 KiB
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261 lines
46 KiB
XML
<document ID-DOI="http://doi.org/10.5281/zenodo.4328172" ID-GBIF-Dataset="faefcf9b-3673-4f50-b6a8-8a5d0cbd01de" ID-GBIF-Taxon="170718955" ID-Zenodo-Dep="4328172" approvalRequired="16" approvalRequired_for_document="1" approvalRequired_for_textStreams="15" checkinTime="1607453490458" checkinUser="plazi" docAuthor="Spangler, Paul J. & Steiner, Warren E." docDate="2005" docId="03C5879CFFCBFF95FF02FF3BBC26FDDF" docLanguage="en" docName="systematicEntomology.30.3.339-357.pdf" docOrigin="Systematic Entomology 30 (3)" docSource="http://dx.doi.org/10.1111/j.1365-3113.2005.00288.x" docTitle="Meru Spangler & Steiner 2005, gen.n." docType="treatment" docVersion="5" lastPageId="8" lastPageNumber="347" masterDocId="FFFCFFE4FFCAFF9DFFA3FFA0B963FFCF" masterDocTitle="A new aquatic beetle family, Meruidae, from Venezuela (Coleoptera: Adephaga)" masterLastPageNumber="357" masterPageNumber="339" pageId="1" pageNumber="340" updateTime="1643586909968" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A new aquatic beetle family, Meruidae, from Venezuela (Coleoptera: Adephaga)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Spangler, Paul J.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Steiner, Warren E.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Systematic Entomology</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2005</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2005-03-07</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>30</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>3</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>339</mods:start>
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<mods:end>357</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.1111/j.1365-3113.2005.00288.x</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.1111/j.1365-3113.2005.00288.x</mods:identifier>
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<mods:identifier type="GBIF-Dataset">faefcf9b-3673-4f50-b6a8-8a5d0cbd01de</mods:identifier>
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<mods:identifier type="Zenodo-Dep">4311852</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.4328172" ID-GBIF-Taxon="170718955" ID-Zenodo-Dep="4328172" LSID="urn:lsid:plazi:treatment:03C5879CFFCBFF95FF02FF3BBC26FDDF" httpUri="http://treatment.plazi.org/id/03C5879CFFCBFF95FF02FF3BBC26FDDF" lastPageId="8" lastPageNumber="347" pageId="1" pageNumber="340">
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<subSubSection box="[161,296,155,177]" pageId="1" pageNumber="340" type="nomenclature">
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<paragraph blockId="1.[161,296,155,177]" box="[161,296,155,177]" pageId="1" pageNumber="340">
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<heading bold="true" box="[161,296,155,177]" fontSize="9" level="1" pageId="1" pageNumber="340" reason="2">
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<emphasis bold="true" box="[161,296,155,177]" pageId="1" pageNumber="340">
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<taxonomicName authority="Spangler & Steiner, 2005" authorityName="Spangler & Steiner" authorityYear="2005" box="[161,218,155,177]" class="Insecta" family="Meruidae" genus="Meru" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="340" phylum="Arthropoda" rank="genus" status="gen.n.">
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<emphasis bold="true" box="[161,218,155,177]" italics="true" pageId="1" pageNumber="340">Meru</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[226,296,155,177]" pageId="1" pageNumber="340" rank="genus">gen.n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection box="[185,538,214,236]" pageId="1" pageNumber="340" type="type_taxon">
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<paragraph blockId="1.[185,538,214,236]" box="[185,538,214,236]" pageId="1" pageNumber="340">
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<emphasis box="[185,479,214,235]" italics="true" pageId="1" pageNumber="340">
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<typeStatus box="[185,235,214,235]" pageId="1" pageNumber="340">Type</typeStatus>
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species.
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<taxonomicName authority="Spangler & Steiner, 2005" authorityName="Spangler & Steiner" authorityYear="2005" box="[330,479,214,235]" class="Insecta" family="Meruidae" genus="Meru" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="340" phylum="Arthropoda" rank="species" species="phyllisae" status="sp.n.">Meru phyllisae</taxonomicName>
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</emphasis>
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,
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<emphasis bold="true" box="[493,538,214,236]" pageId="1" pageNumber="340">
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<taxonomicNameLabel box="[493,538,214,236]" pageId="1" pageNumber="340" rank="species">sp.n.</taxonomicNameLabel>
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</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="8" lastPageNumber="347" pageId="1" pageNumber="340" type="description">
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<paragraph blockId="1.[161,767,273,441]" lastBlockId="1.[830,1436,156,441]" pageId="1" pageNumber="340">
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<emphasis box="[185,307,273,294]" italics="true" pageId="1" pageNumber="340">Description.</emphasis>
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Size very small, length 0.85–0.9 mm; width 0.38–0.41 mm. Body form generally ovate, convex dorsally, less so ventrally (
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<figureCitation box="[343,399,331,353]" captionStart="Fig" captionStartId="1.[161,195,1816,1836]" captionTargetBox="[161,1435,490,1787]" captionTargetId="figure-141@1.[161,1435,490,1788]" captionTargetPageId="1" captionText="Fig. 1. Meru phyllisae, scanning electron micrograph, lateral habitus." figureDoi="http://doi.org/10.5281/zenodo.4311856" httpUri="https://zenodo.org/record/4311856/files/figure.png" pageId="1" pageNumber="340">Fig. 1</figureCitation>
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), widest at basal quarter of elytra; prothorax broadly attached to mesothorax and elytral base (
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<figureCitation box="[168,225,390,412]" captionStart="Fig" captionStartId="2.[151,185,1603,1623]" captionTargetBox="[152,1426,156,1574]" captionTargetId="figure-70@2.[152,1426,156,1574]" captionTargetPageId="2" captionText="Fig. 2. Meru phyllisae. Scanning electron micrographs. A, Habitus, dorsal view; B, habitus, ventral view; C, thoracic sterna; D, head, pronotum and bases of elytra; E, head and prosternal area; F, metasternum and abdomen; G, right elytron; H, pronotal and elytral bases; I, pronotum, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4311858" httpUri="https://zenodo.org/record/4311858/files/figure.png" pageId="1" pageNumber="340">Fig. 2</figureCitation>
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A–I). Colour generally dark brown to yellowish brown. Most surfaces of body bearing flat, wrinkled ‘setae’ (possibly membranous papillae; see comments below) arising from the anterior side of large, shallow punctures.
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</paragraph>
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<paragraph blockId="1.[830,1436,156,441]" lastBlockId="4.[821,1427,1291,1839]" lastPageId="4" lastPageNumber="343" pageId="1" pageNumber="340">
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Head (
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<figureCitation box="[928,1115,214,236]" captionStart="Fig" captionStartId="2.[151,185,1603,1623]" captionTargetBox="[152,1426,156,1574]" captionTargetId="figure-70@2.[152,1426,156,1574]" captionTargetPageId="2" captionText="Fig. 2. Meru phyllisae. Scanning electron micrographs. A, Habitus, dorsal view; B, habitus, ventral view; C, thoracic sterna; D, head, pronotum and bases of elytra; E, head and prosternal area; F, metasternum and abdomen; G, right elytron; H, pronotal and elytral bases; I, pronotum, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4311858" httpUri="https://zenodo.org/record/4311858/files/figure.png" pageId="1" pageNumber="340">Figs 2A, B, D, E</figureCitation>
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,
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<figureCitation box="[1131,1196,214,236]" captionStart="Fig" captionStartId="3.[161,195,1600,1620]" captionTargetBox="[161,1435,156,1571]" captionTargetId="figure-74@3.[161,1435,155,1571]" captionTargetPageId="3" captionText="Fig. 3. Meru phyllisae. A, Head, lateral view; B, head, anterior view; C, mouthparts, ventral view; D, labial palpal bases; E, head, anterior view, detail of mouth; F, right maxillary and labial palpi, ventral view; G, right maxillary palpus, apex; H, left maxillary palpus, apex; I, right labial palpus, apex." figureDoi="http://doi.org/10.5281/zenodo.4311860" httpUri="https://zenodo.org/record/4311860/files/figure.png" pageId="1" pageNumber="340">3A, B</figureCitation>
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) prognathous, slightly longer than wide, with retracted neck region; eyes lateral, broadly elliptic, prominent; ommatidia numbering five by nine across axes of eye. Frons smooth, slightly convex, with reticulate microsculpture; laterally and basally with scattered shallow punctures and fine setae; frontoclypeal suture obscure; labroclypeal suture nearly straight. Labrum trapezoidal, with scalelike microsculpture; front edge with a shallow median notch covered by opposing, overlapping, spatulate setae (
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<figureCitation box="[334,407,1763,1785]" captionStart="Fig" captionStartId="3.[161,195,1600,1620]" captionTargetBox="[161,1435,156,1571]" captionTargetId="figure-74@3.[161,1435,155,1571]" captionTargetPageId="3" captionText="Fig. 3. Meru phyllisae. A, Head, lateral view; B, head, anterior view; C, mouthparts, ventral view; D, labial palpal bases; E, head, anterior view, detail of mouth; F, right maxillary and labial palpi, ventral view; G, right maxillary palpus, apex; H, left maxillary palpus, apex; I, right labial palpus, apex." figureDoi="http://doi.org/10.5281/zenodo.4311860" httpUri="https://zenodo.org/record/4311860/files/figure.png" pageId="2" pageNumber="341">Fig. 3E</figureCitation>
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) and with three tapered setae originating from small pits lateral to base of spatulate seta; a pair of small, flat, toothlike setae imbedded in the anterior margin of the notch. Antenna (
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<figureCitation box="[1231,1290,1734,1756]" captionStart="Fig" captionStartId="4.[151,185,1131,1151]" captionTargetBox="[151,1426,156,1102]" captionTargetId="figure-347@4.[150,1427,155,1103]" captionTargetPageId="4" captionText="Fig. 4. Meru phyllisae, antennae. A, Left, dorsal view; B, right, segments 1–7, ventral view; C, right, segments 7–11, ventral view, showing sensorial patches on segments 7, 9 and 11; D, right, segments 6–8, ventral view, showing sensorial patch on segment 7; E, left, lateral view, apical (eleventh) segment with sensorial patches; F, right, ventral view, apical (eleventh) segment with sensorial patches." figureDoi="http://doi.org/10.5281/zenodo.4311862" httpUri="https://zenodo.org/record/4311862/files/figure.png" pageId="2" pageNumber="341">Fig. 4</figureCitation>
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A–F1) longer than head, filiform, glabrous, eleven-segmented; scapus very short and nearly hidden; pedicellus almost globular, urn-shaped, bearing a few slender setae; antennomere III as long as pedicellus but more narrow, nearly twice as long as wide, subcylindrical, thickest towards apex; antennomeres IV–X of similar form, but
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<collectionCode box="[430,450,1789,1811]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="3" pageNumber="342" type="Museum">V</collectionCode>
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,
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<collectionCode box="[460,496,1789,1811]" pageId="3" pageNumber="342">VII</collectionCode>
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and
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<collectionCode box="[547,575,1789,1811]" pageId="3" pageNumber="342">IX</collectionCode>
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longer and thicker than the adjacent ones; antennomeres
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<collectionCode box="[551,587,1819,1841]" pageId="3" pageNumber="342">VII</collectionCode>
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and
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<collectionCode box="[638,666,1819,1841]" pageId="3" pageNumber="342">IX</collectionCode>
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bearing a platelike sensorial patch ventrally; antennomere XI longer than the preceding two combined, thickest at middle and obliquely tapering to a narrow apex, with two platelike sensorial patches near apex on ventral side. Mandibles (
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<figureCitation box="[159,233,1291,1313]" captionStart="Fig" captionStartId="7.[161,195,1563,1583]" captionTargetBox="[161,766,157,1532]" captionTargetId="figure-198@7.[161,766,157,1535]" captionTargetPageId="7" captionText="Fig. 7. Meru phyllisae. A, Mandibles, dorsal view; B, maxilla, left, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4311870" httpUri="https://zenodo.org/record/4311870/files/figure.png" pageId="4" pageNumber="343">Fig. 7A</figureCitation>
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) nearly twice as long as wide, with a single, blunt, falcate, apical incisor and median terebral tooth; brushes of trichiae, molar teeth, and retinaculum absent. Left mandible slightly longer than right, terebral tooth broadly angular; inferior terebral ridge with one or two minute teeth. Right mandible with terebral tooth truncate and closer to apex than that of left mandible; basal face with several small wrinkles. Maxilla (
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<figureCitation box="[291,363,1495,1517]" captionStart="Fig" captionStartId="7.[161,195,1563,1583]" captionTargetBox="[161,766,157,1532]" captionTargetId="figure-198@7.[161,766,157,1535]" captionTargetPageId="7" captionText="Fig. 7. Meru phyllisae. A, Mandibles, dorsal view; B, maxilla, left, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4311870" httpUri="https://zenodo.org/record/4311870/files/figure.png" pageId="4" pageNumber="343">Fig. 7B</figureCitation>
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) with cardo small, wedge-shaped; stipes and lacinia fused; lacinia stout, elongate, glabrous, with a curved, robust apical spine and seven setae on inner side; galea simple, two-segmented, shorter than lacinia, with apical segment elongate, cylindrical, smoothly rounded at apex; palpus four-segmented; apical segment slightly longer than combined length of segments 1–3, with sensory structures on the distal part (
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<figureCitation box="[266,321,1700,1722]" captionStart="Fig" captionStartId="3.[161,195,1600,1620]" captionTargetBox="[161,1435,156,1571]" captionTargetId="figure-74@3.[161,1435,155,1571]" captionTargetPageId="3" captionText="Fig. 3. Meru phyllisae. A, Head, lateral view; B, head, anterior view; C, mouthparts, ventral view; D, labial palpal bases; E, head, anterior view, detail of mouth; F, right maxillary and labial palpi, ventral view; G, right maxillary palpus, apex; H, left maxillary palpus, apex; I, right labial palpus, apex." figureDoi="http://doi.org/10.5281/zenodo.4311860" httpUri="https://zenodo.org/record/4311860/files/figure.png" pageId="4" pageNumber="343">Fig. 3</figureCitation>
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F–H). Submentum broad, with distinct tentorial pit at basolateral corner (
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<figureCitation box="[481,558,1729,1751]" captionStart="Fig" captionStartId="2.[151,185,1603,1623]" captionTargetBox="[152,1426,156,1574]" captionTargetId="figure-70@2.[152,1426,156,1574]" captionTargetPageId="2" captionText="Fig. 2. Meru phyllisae. Scanning electron micrographs. A, Habitus, dorsal view; B, habitus, ventral view; C, thoracic sterna; D, head, pronotum and bases of elytra; E, head and prosternal area; F, metasternum and abdomen; G, right elytron; H, pronotal and elytral bases; I, pronotum, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4311858" httpUri="https://zenodo.org/record/4311858/files/figure.png" pageId="4" pageNumber="343">Figs 2E</figureCitation>
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,
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<figureCitation box="[572,603,1729,1751]" captionStart="Fig" captionStartId="3.[161,195,1600,1620]" captionTargetBox="[161,1435,156,1571]" captionTargetId="figure-74@3.[161,1435,155,1571]" captionTargetPageId="3" captionText="Fig. 3. Meru phyllisae. A, Head, lateral view; B, head, anterior view; C, mouthparts, ventral view; D, labial palpal bases; E, head, anterior view, detail of mouth; F, right maxillary and labial palpi, ventral view; G, right maxillary palpus, apex; H, left maxillary palpus, apex; I, right labial palpus, apex." figureDoi="http://doi.org/10.5281/zenodo.4311860" httpUri="https://zenodo.org/record/4311860/files/figure.png" pageId="4" pageNumber="343">3C</figureCitation>
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); mentum flat, smooth, slightly more than half as wide as head, with two long setae on each side at approximately middle; prementum recessed, inserted between anterolateral rounded lobes of mentum; base of palpifer with a single small seta; ligula moderately extended and rounded apicomedially, with twelve to fourteen apical setae (
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<figureCitation box="[945,1049,1349,1371]" captionStart="Fig" captionStartId="3.[161,195,1600,1620]" captionTargetBox="[161,1435,156,1571]" captionTargetId="figure-74@3.[161,1435,155,1571]" captionTargetPageId="3" captionText="Fig. 3. Meru phyllisae. A, Head, lateral view; B, head, anterior view; C, mouthparts, ventral view; D, labial palpal bases; E, head, anterior view, detail of mouth; F, right maxillary and labial palpi, ventral view; G, right maxillary palpus, apex; H, left maxillary palpus, apex; I, right labial palpus, apex." figureDoi="http://doi.org/10.5281/zenodo.4311860" httpUri="https://zenodo.org/record/4311860/files/figure.png" pageId="4" pageNumber="343">Fig. 3C, D</figureCitation>
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); labial palpus three-segmented; apical segment swollen and slightly longer than combined length of segments 1 and 2, with apical sensory structures (
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<figureCitation box="[1299,1419,1408,1430]" captionStart="Fig" captionStartId="3.[161,195,1600,1620]" captionTargetBox="[161,1435,156,1571]" captionTargetId="figure-74@3.[161,1435,155,1571]" captionTargetPageId="3" captionText="Fig. 3. Meru phyllisae. A, Head, lateral view; B, head, anterior view; C, mouthparts, ventral view; D, labial palpal bases; E, head, anterior view, detail of mouth; F, right maxillary and labial palpi, ventral view; G, right maxillary palpus, apex; H, left maxillary palpus, apex; I, right labial palpus, apex." figureDoi="http://doi.org/10.5281/zenodo.4311860" httpUri="https://zenodo.org/record/4311860/files/figure.png" pageId="4" pageNumber="343">Fig. 3C, F, I</figureCitation>
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) similar to those of maxillary palpus.
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</paragraph>
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<caption ID-DOI="http://doi.org/10.5281/zenodo.4311856" ID-Zenodo-Dep="4311856" box="[161,799,1816,1836]" httpUri="https://zenodo.org/record/4311856/files/figure.png" pageId="1" pageNumber="340" startId="1.[161,195,1816,1836]" targetBox="[161,1435,490,1787]" targetPageId="1">
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<paragraph blockId="1.[161,799,1816,1836]" box="[161,799,1816,1836]" pageId="1" pageNumber="340">
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<emphasis bold="true" box="[161,220,1816,1836]" pageId="1" pageNumber="340">Fig. 1.</emphasis>
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<taxonomicName authorityName="Spangler & Steiner" authorityYear="2005" box="[231,364,1817,1836]" class="Insecta" family="Meruidae" genus="Meru" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="340" phylum="Arthropoda" rank="species" species="phyllisae">
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<emphasis box="[231,364,1817,1836]" italics="true" pageId="1" pageNumber="340">Meru phyllisae</emphasis>
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</taxonomicName>
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, scanning electron micrograph, lateral habitus.
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</paragraph>
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</caption>
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<caption ID-DOI="http://doi.org/10.5281/zenodo.4311858" ID-Zenodo-Dep="4311858" httpUri="https://zenodo.org/record/4311858/files/figure.png" pageId="2" pageNumber="341" startId="2.[151,185,1603,1623]" targetBox="[152,1426,156,1574]" targetPageId="2">
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<paragraph blockId="2.[151,1427,1603,1676]" pageId="2" pageNumber="341">
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<emphasis bold="true" box="[151,210,1603,1623]" pageId="2" pageNumber="341">Fig. 2.</emphasis>
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<taxonomicName authorityName="Spangler & Steiner" authorityYear="2005" box="[221,357,1604,1623]" class="Insecta" family="Meruidae" genus="Meru" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="341" phylum="Arthropoda" rank="species" species="phyllisae">
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<emphasis box="[221,357,1604,1623]" italics="true" pageId="2" pageNumber="341">Meru phyllisae</emphasis>
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</taxonomicName>
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. Scanning electron micrographs. A, Habitus, dorsal view; B, habitus, ventral view; C, thoracic sterna; D, head, pronotum and bases of elytra; E, head and prosternal area; F, metasternum and abdomen; G, right elytron; H, pronotal and elytral bases; I, pronotum, lateral view.
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</paragraph>
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</caption>
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<caption ID-DOI="http://doi.org/10.5281/zenodo.4311860" ID-Zenodo-Dep="4311860" httpUri="https://zenodo.org/record/4311860/files/figure.png" pageId="3" pageNumber="342" startId="3.[161,195,1600,1620]" targetBox="[161,1435,156,1571]" targetPageId="3">
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<paragraph blockId="3.[161,1437,1600,1673]" pageId="3" pageNumber="342">
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<emphasis bold="true" box="[161,220,1600,1620]" pageId="3" pageNumber="342">Fig. 3.</emphasis>
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<taxonomicName authorityName="Spangler & Steiner" authorityYear="2005" box="[231,364,1601,1620]" class="Insecta" family="Meruidae" genus="Meru" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="342" phylum="Arthropoda" rank="species" species="phyllisae">
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<emphasis box="[231,364,1601,1620]" italics="true" pageId="3" pageNumber="342">Meru phyllisae</emphasis>
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</taxonomicName>
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. A, Head, lateral view; B, head, anterior view; C, mouthparts, ventral view; D, labial palpal bases; E, head, anterior view, detail of mouth; F, right maxillary and labial palpi, ventral view; G, right maxillary palpus, apex; H, left maxillary palpus, apex; I, right labial palpus, apex.
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</paragraph>
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</caption>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.4311862" ID-Zenodo-Dep="4311862" httpUri="https://zenodo.org/record/4311862/files/figure.png" pageId="4" pageNumber="343" startId="4.[151,185,1131,1151]" targetBox="[151,1426,156,1102]" targetPageId="4">
|
||
<paragraph blockId="4.[151,1427,1131,1204]" pageId="4" pageNumber="343">
|
||
<emphasis bold="true" box="[151,210,1131,1151]" pageId="4" pageNumber="343">Fig. 4.</emphasis>
|
||
<taxonomicName authorityName="Spangler & Steiner" authorityYear="2005" box="[221,354,1132,1151]" class="Insecta" family="Meruidae" genus="Meru" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="343" phylum="Arthropoda" rank="species" species="phyllisae">
|
||
<emphasis box="[221,354,1132,1151]" italics="true" pageId="4" pageNumber="343">Meru phyllisae</emphasis>
|
||
</taxonomicName>
|
||
, antennae. A, Left, dorsal view; B, right, segments 1–7, ventral view; C, right, segments 7–11, ventral view, showing sensorial patches on segments 7, 9 and 11; D, right, segments 6–8, ventral view, showing sensorial patch on segment 7; E, left, lateral view, apical (eleventh) segment with sensorial patches; F, right, ventral view, apical (eleventh) segment with sensorial patches.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="4.[821,1427,1291,1839]" lastBlockId="5.[830,1437,156,762]" lastPageId="5" lastPageNumber="344" pageId="4" pageNumber="343">
|
||
Prothorax (
|
||
<figureCitation box="[962,1094,1466,1488]" captionStart="Fig" captionStartId="2.[151,185,1603,1623]" captionTargetBox="[152,1426,156,1574]" captionTargetId="figure-70@2.[152,1426,156,1574]" captionTargetPageId="2" captionText="Fig. 2. Meru phyllisae. Scanning electron micrographs. A, Habitus, dorsal view; B, habitus, ventral view; C, thoracic sterna; D, head, pronotum and bases of elytra; E, head and prosternal area; F, metasternum and abdomen; G, right elytron; H, pronotal and elytral bases; I, pronotum, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4311858" httpUri="https://zenodo.org/record/4311858/files/figure.png" pageId="4" pageNumber="343">Figs 2D, E, I</figureCitation>
|
||
,
|
||
<figureCitation box="[1105,1136,1466,1488]" captionStart="Fig" captionStartId="7.[830,864,1360,1380]" captionTargetBox="[834,1434,158,1330]" captionTargetId="figure-215@7.[831,1436,156,1331]" captionTargetPageId="7" captionText="Fig. 8. Meru phyllisae, sterna. A, Outline of prothorax; B, hind body." figureDoi="http://doi.org/10.5281/zenodo.4311868" httpUri="https://zenodo.org/record/4311868/files/figure.png" pageId="4" pageNumber="343">8A</figureCitation>
|
||
) about four-fifths as wide as elytra; pronotum widest at base, slightly wider than long, lateral margin sinuate, not beaded; anterior margin straight, anterior corners narrowly rounded; posterior corners angled; posterior margin nearly straight between corner and middle, tightly fitted against base of elytron and forming an angle of about 125 with other side at middle; disk evenly convex, surface with four shallow depressions on each side; surface coarsely, densely punctate; each puncture with a long, robust, flattened and wrinkled seta, pleated at base, arising from anterior edge of puncture and recumbent across it; length of seta about twice the diameter of a puncture; scattered, hairlike setae also present, arising from smooth areas between the large punctures. Thoracic sternites with a similar combination of setae and punctures, but larger and more sparsely placed laterally. Prosternum (
|
||
<figureCitation box="[399,478,243,265]" captionStart="Fig" captionStartId="2.[151,185,1603,1623]" captionTargetBox="[152,1426,156,1574]" captionTargetId="figure-70@2.[152,1426,156,1574]" captionTargetPageId="2" captionText="Fig. 2. Meru phyllisae. Scanning electron micrographs. A, Habitus, dorsal view; B, habitus, ventral view; C, thoracic sterna; D, head, pronotum and bases of elytra; E, head and prosternal area; F, metasternum and abdomen; G, right elytron; H, pronotal and elytral bases; I, pronotum, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4311858" httpUri="https://zenodo.org/record/4311858/files/figure.png" pageId="5" pageNumber="344">Figs 2E</figureCitation>
|
||
,
|
||
<figureCitation box="[493,524,243,265]" captionStart="Fig" captionStartId="7.[830,864,1360,1380]" captionTargetBox="[834,1434,158,1330]" captionTargetId="figure-215@7.[831,1436,156,1331]" captionTargetPageId="7" captionText="Fig. 8. Meru phyllisae, sterna. A, Outline of prothorax; B, hind body." figureDoi="http://doi.org/10.5281/zenodo.4311868" httpUri="https://zenodo.org/record/4311868/files/figure.png" pageId="5" pageNumber="344">8A</figureCitation>
|
||
) with notopleural and pleurosternal sutures visible; propleuron large, extending behind coxa; procoxal cavity closed internally. Cervical sclerite absent. Anterior margin of prosternum evenly arcuate with a fine fringe of microsetae along edge and on each side with one large, anteriorly directed, club-shaped seta, with apex flattened and toothed, brushlike (
|
||
<figureCitation box="[679,756,419,441]" captionStart="Fig" captionStartId="2.[151,185,1603,1623]" captionTargetBox="[152,1426,156,1574]" captionTargetId="figure-70@2.[152,1426,156,1574]" captionTargetPageId="2" captionText="Fig. 2. Meru phyllisae. Scanning electron micrographs. A, Habitus, dorsal view; B, habitus, ventral view; C, thoracic sterna; D, head, pronotum and bases of elytra; E, head and prosternal area; F, metasternum and abdomen; G, right elytron; H, pronotal and elytral bases; I, pronotum, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4311858" httpUri="https://zenodo.org/record/4311858/files/figure.png" pageId="5" pageNumber="344">Fig. 2E</figureCitation>
|
||
). Prosternal process very narrow between procoxae but wider and truncate behind them (
|
||
<figureCitation box="[527,605,477,499]" captionStart="Fig" captionStartId="7.[830,864,1360,1380]" captionTargetBox="[834,1434,158,1330]" captionTargetId="figure-215@7.[831,1436,156,1331]" captionTargetPageId="7" captionText="Fig. 8. Meru phyllisae, sterna. A, Outline of prothorax; B, hind body." figureDoi="http://doi.org/10.5281/zenodo.4311868" httpUri="https://zenodo.org/record/4311868/files/figure.png" pageId="5" pageNumber="344">Fig. 8A</figureCitation>
|
||
). Mesosternite, metasternite and lateral part of metacoxae fused, with sutures not discernible laterally (
|
||
<figureCitation box="[535,610,536,558]" captionStart="Fig" captionStartId="7.[830,864,1360,1380]" captionTargetBox="[834,1434,158,1330]" captionTargetId="figure-215@7.[831,1436,156,1331]" captionTargetPageId="7" captionText="Fig. 8. Meru phyllisae, sterna. A, Outline of prothorax; B, hind body." figureDoi="http://doi.org/10.5281/zenodo.4311868" httpUri="https://zenodo.org/record/4311868/files/figure.png" pageId="5" pageNumber="344">Fig. 8B</figureCitation>
|
||
); mesosternite with pentagonal depression partly covered by (articulated with) prosternal process. Transverse ridge of metaventrite absent. Metacoxal plates small, not covering base of trochanter, with lateral margins indistinct anteriorly; median lamina of metacoxae not distinctly separated from lateral part; posterior margin straight, with a small median emargination and deep incisions for trochantinal articulations (
|
||
<figureCitation box="[837,950,156,178]" captionStart="Fig" captionStartId="2.[151,185,1603,1623]" captionTargetBox="[152,1426,156,1574]" captionTargetId="figure-70@2.[152,1426,156,1574]" captionTargetPageId="2" captionText="Fig. 2. Meru phyllisae. Scanning electron micrographs. A, Habitus, dorsal view; B, habitus, ventral view; C, thoracic sterna; D, head, pronotum and bases of elytra; E, head and prosternal area; F, metasternum and abdomen; G, right elytron; H, pronotal and elytral bases; I, pronotum, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4311858" httpUri="https://zenodo.org/record/4311858/files/figure.png" pageId="5" pageNumber="344">Figs 2C, F</figureCitation>
|
||
,
|
||
<figureCitation box="[964,995,156,178]" captionStart="Fig" captionStartId="7.[830,864,1360,1380]" captionTargetBox="[834,1434,158,1330]" captionTargetId="figure-215@7.[831,1436,156,1331]" captionTargetPageId="7" captionText="Fig. 8. Meru phyllisae, sterna. A, Outline of prothorax; B, hind body." figureDoi="http://doi.org/10.5281/zenodo.4311868" httpUri="https://zenodo.org/record/4311868/files/figure.png" pageId="5" pageNumber="344">8B</figureCitation>
|
||
). Metendosternal process slender, median ridge (metacoxal septum) attached to the metasternite for two-thirds of its length, ending with a slenderarmed, Y-shaped metafurca anteriorly (
|
||
<figureCitation box="[1252,1321,243,265]" captionStart="Fig" captionStartId="8.[151,185,1112,1132]" captionTargetBox="[151,756,157,1084]" captionTargetId="figure-654@8.[151,756,157,1084]" captionTargetPageId="8" captionText="Fig. 9. Meru phyllisae, metendosternite, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4311872" httpUri="https://zenodo.org/record/4311872/files/figure.png" pageId="5" pageNumber="344">Figs 9</figureCitation>
|
||
,
|
||
<figureCitation box="[1336,1414,243,265]" captionStart="Fig" captionStartId="9.[161,195,1831,1851]" captionTargetBox="[230,1366,156,1803]" captionTargetId="figure-2@9.[230,1366,156,1803]" captionTargetPageId="9" captionText="Fig. 10. Meru phyllisae, metendosternite, scanning electron micrographs. A, Oblique dorsolateral view; B, dorsolateral view." figureDoi="http://doi.org/10.5281/zenodo.4311874" httpUri="https://zenodo.org/record/4311874/files/figure.png" pageId="5" pageNumber="344">10A, B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="5.[830,1437,156,762]" lastBlockId="7.[830,1437,1409,1840]" lastPageId="7" lastPageNumber="346" pageId="5" pageNumber="344">
|
||
Scutellum not visible. Elytron convex, twice as wide as long, dorsally with setae and punctures like those on pronotum (
|
||
<figureCitation box="[912,1018,331,353]" captionStart="Fig" captionStartId="5.[161,195,1789,1809]" captionTargetBox="[161,1436,820,1761]" captionTargetId="figure-352@5.[160,1437,819,1761]" captionTargetPageId="5" captionText="Fig. 5. Meru phyllisae. A, Left elytron, lateral view; B, left elytron, detail of margin; C, left elytron, inner surface; D, left elytron, wing-folding patch on inner surface; E, abdominal sternites, detail of posterior margins of 1–3; F, abdominal sternite 3, detail of posterior margin." figureDoi="http://doi.org/10.5281/zenodo.4311864" httpUri="https://zenodo.org/record/4311864/files/figure.png" pageId="5" pageNumber="344">Fig. 5A, B</figureCitation>
|
||
) in addition to rows of large, deep, strial punctures that do not bear setae; sutural stria absent and sutural edge not beaded; lateral margin arcuate in lateral view (
|
||
<figureCitation box="[891,970,419,441]" captionStart="Fig" captionStartId="5.[161,195,1789,1809]" captionTargetBox="[161,1436,820,1761]" captionTargetId="figure-352@5.[160,1437,819,1761]" captionTargetPageId="5" captionText="Fig. 5. Meru phyllisae. A, Left elytron, lateral view; B, left elytron, detail of margin; C, left elytron, inner surface; D, left elytron, wing-folding patch on inner surface; E, abdominal sternites, detail of posterior margins of 1–3; F, abdominal sternite 3, detail of posterior margin." figureDoi="http://doi.org/10.5281/zenodo.4311864" httpUri="https://zenodo.org/record/4311864/files/figure.png" pageId="5" pageNumber="344">Fig. 5A</figureCitation>
|
||
); epipleuron indistinct; inner surface smooth, laterally with a large, locking pit at about midlength and a round patch of granular surface (presumably for wing folding) anterior to it (
|
||
<figureCitation box="[1071,1182,506,528]" captionStart="Fig" captionStartId="5.[161,195,1789,1809]" captionTargetBox="[161,1436,820,1761]" captionTargetId="figure-352@5.[160,1437,819,1761]" captionTargetPageId="5" captionText="Fig. 5. Meru phyllisae. A, Left elytron, lateral view; B, left elytron, detail of margin; C, left elytron, inner surface; D, left elytron, wing-folding patch on inner surface; E, abdominal sternites, detail of posterior margins of 1–3; F, abdominal sternite 3, detail of posterior margin." figureDoi="http://doi.org/10.5281/zenodo.4311864" httpUri="https://zenodo.org/record/4311864/files/figure.png" pageId="5" pageNumber="344">Fig. 5C, D</figureCitation>
|
||
). Hind wings dimorphic (
|
||
<figureCitation box="[838,959,536,558]" captionStart="Fig" captionStartId="10.[151,185,696,716]" captionTargetBox="[167,742,156,668]" captionTargetId="figure-2@10.[167,742,156,668]" captionTargetPageId="10" captionText="Fig. 11. Meru phyllisae, hind wing. A, Brachypterous condition; B, macropterous condition." figureDoi="http://doi.org/10.5281/zenodo.4311876" httpUri="https://zenodo.org/record/4311876/files/figure.png" pageId="5" pageNumber="344">Fig. 11A, B</figureCitation>
|
||
), either fully developed with venation moderately developed and distal half folded (
|
||
<figureCitation box="[1286,1377,565,587]" captionStart="Fig" captionStartId="10.[151,185,696,716]" captionTargetBox="[167,742,156,668]" captionTargetId="figure-2@10.[167,742,156,668]" captionTargetPageId="10" captionText="Fig. 11. Meru phyllisae, hind wing. A, Brachypterous condition; B, macropterous condition." figureDoi="http://doi.org/10.5281/zenodo.4311876" httpUri="https://zenodo.org/record/4311876/files/figure.png" pageId="5" pageNumber="344">Figs 11B</figureCitation>
|
||
,
|
||
<figureCitation box="[1398,1424,565,587]" captionStart="Fig" captionStartId="10.[151,185,1224,1244]" captionTargetBox="[167,742,769,1196]" captionTargetId="figure-29@10.[167,742,767,1196]" captionTargetPageId="10" captionText="Fig. 12. Meru phyllisae, hind wing, macropterous condition, in folded position." figureDoi="http://doi.org/10.5281/zenodo.4311878" httpUri="https://zenodo.org/record/4311878/files/figure.png" pageId="5" pageNumber="344">12</figureCitation>
|
||
), or brachypterous, as long as elytron, with venation very reduced and distal half appearing shrunken (
|
||
<figureCitation box="[1334,1424,623,645]" captionStart="Fig" captionStartId="10.[151,185,696,716]" captionTargetBox="[167,742,156,668]" captionTargetId="figure-2@10.[167,742,156,668]" captionTargetPageId="10" captionText="Fig. 11. Meru phyllisae, hind wing. A, Brachypterous condition; B, macropterous condition." figureDoi="http://doi.org/10.5281/zenodo.4311876" httpUri="https://zenodo.org/record/4311876/files/figure.png" pageId="5" pageNumber="344">Fig. 11A</figureCitation>
|
||
); margins with a fringe of long setae; subcubital binding patch absent; oblongum cell not discernible. Procoxa and mesocoxa globular; trochanters smooth, unmodified. Metatrochanter large, about twice the size of pro- and mesotrochanter, globular, swollen posteriorly; junction with metafemur oblique and sinuate. Femora smooth, club-shaped, widest at about apical third, with scattered sparse hairlike setae, some brush-tipped, and a row of five to seven thick, branched setae on dorsal surface. Metafemur very slender basally, attached dorsolaterally to trochanter; distal two-thirds swollen. Tibiae simple, straight, gradually thickening from base to near apex, with sparse hairlike setae, irregularly distributed, becoming more stout towards apex of tibia; each tibia bearing a pair of toothed apical spurs, both trident on pro- and mesotibia, one trident and one laterally pectinate on metatibia (
|
||
<figureCitation box="[571,628,1731,1753]" captionStart="Fig" captionStartId="6.[151,185,1763,1783]" captionTargetBox="[151,1426,310,1734]" captionTargetId="figure-64@6.[150,1427,310,1734]" captionTargetPageId="6" captionText="Fig. 6. Meru phyllisae. A, Front and mid legs, left, anterior view; B, front and mid legs, right, posterior view; C, D, mesotibia, right, apex; E, hind tarsus, left, ventral view; F, hind tarsus, left, posterior view; G, claws, right mesotarsal, lateral view; H, claws, left metatarsal, oblique view; I, claws, right metatarsal, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4311866" httpUri="https://zenodo.org/record/4311866/files/figure.png" pageId="7" pageNumber="346">Fig. 6</figureCitation>
|
||
A–F). Tarsal formula 5-5-5; last tarsomeres longer than combined lengths of the preceding two; tarsomeres 1 and 2 each with four stout, ventral setae, 3 and 4 with two setae, and 5 with one seta; hind tarsus with basal tarsomere longest; length equals combined length of tarsomeres 2–4. Tarsal claws large, pectinate, with four to five teeth (
|
||
<figureCitation box="[1324,1382,1467,1489]" captionStart="Fig" captionStartId="6.[151,185,1763,1783]" captionTargetBox="[151,1426,310,1734]" captionTargetId="figure-64@6.[150,1427,310,1734]" captionTargetPageId="6" captionText="Fig. 6. Meru phyllisae. A, Front and mid legs, left, anterior view; B, front and mid legs, right, posterior view; C, D, mesotibia, right, apex; E, hind tarsus, left, ventral view; F, hind tarsus, left, posterior view; G, claws, right mesotarsal, lateral view; H, claws, left metatarsal, oblique view; I, claws, right metatarsal, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4311866" httpUri="https://zenodo.org/record/4311866/files/figure.png" pageId="7" pageNumber="346">Fig. 6</figureCitation>
|
||
G–I). Empodium small, smooth, padlike.
|
||
</paragraph>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.4311864" ID-Zenodo-Dep="4311864" httpUri="https://zenodo.org/record/4311864/files/figure.png" pageId="5" pageNumber="344" startId="5.[161,195,1789,1809]" targetBox="[161,1436,820,1761]" targetPageId="5">
|
||
<paragraph blockId="5.[161,1437,1789,1836]" pageId="5" pageNumber="344">
|
||
<emphasis bold="true" box="[161,220,1789,1809]" pageId="5" pageNumber="344">Fig. 5.</emphasis>
|
||
<taxonomicName authorityName="Spangler & Steiner" authorityYear="2005" box="[231,362,1790,1809]" class="Insecta" family="Meruidae" genus="Meru" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="344" phylum="Arthropoda" rank="species" species="phyllisae">
|
||
<emphasis box="[231,362,1790,1809]" italics="true" pageId="5" pageNumber="344">Meru phyllisae</emphasis>
|
||
</taxonomicName>
|
||
. A, Left elytron, lateral view; B, left elytron, detail of margin; C, left elytron, inner surface; D, left elytron, wing-folding patch on inner surface; E, abdominal sternites, detail of posterior margins of 1–3; F, abdominal sternite 3, detail of posterior margin.
|
||
</paragraph>
|
||
</caption>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.4311866" ID-Zenodo-Dep="4311866" httpUri="https://zenodo.org/record/4311866/files/figure.png" pageId="6" pageNumber="345" startId="6.[151,185,1763,1783]" targetBox="[151,1426,310,1734]" targetPageId="6">
|
||
<paragraph blockId="6.[151,1427,1763,1836]" pageId="6" pageNumber="345">
|
||
<emphasis bold="true" box="[151,210,1763,1783]" pageId="6" pageNumber="345">Fig. 6.</emphasis>
|
||
<taxonomicName authorityName="Spangler & Steiner" authorityYear="2005" box="[221,354,1764,1783]" class="Insecta" family="Meruidae" genus="Meru" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="345" phylum="Arthropoda" rank="species" species="phyllisae">
|
||
<emphasis box="[221,354,1764,1783]" italics="true" pageId="6" pageNumber="345">Meru phyllisae</emphasis>
|
||
</taxonomicName>
|
||
. A, Front and mid legs, left, anterior view; B, front and mid legs, right, posterior view; C, D, mesotibia, right, apex; E, hind tarsus, left, ventral view; F, hind tarsus, left, posterior view; G, claws, right mesotarsal, lateral view; H, claws, left metatarsal, oblique view; I, claws, right metatarsal, ventral view.
|
||
</paragraph>
|
||
</caption>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.4311868" ID-Zenodo-Dep="4311868" box="[830,1437,1360,1380]" httpUri="https://zenodo.org/record/4311868/files/figure.png" pageId="7" pageNumber="346" startId="7.[830,864,1360,1380]" targetBox="[834,1434,158,1330]" targetPageId="7">
|
||
<paragraph blockId="7.[830,1437,1360,1380]" box="[830,1437,1360,1380]" pageId="7" pageNumber="346">
|
||
<emphasis bold="true" box="[830,890,1360,1380]" pageId="7" pageNumber="346">Fig. 8.</emphasis>
|
||
<taxonomicName authorityName="Spangler & Steiner" authorityYear="2005" box="[900,1027,1361,1380]" class="Insecta" family="Meruidae" genus="Meru" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="346" phylum="Arthropoda" rank="species" species="phyllisae">
|
||
<emphasis box="[900,1027,1361,1380]" italics="true" pageId="7" pageNumber="346">Meru phyllisae</emphasis>
|
||
</taxonomicName>
|
||
, sterna. A, Outline of prothorax; B, hind body.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="7.[830,1437,1409,1840]" pageId="7" pageNumber="346">
|
||
Abdomen with six visible sternites (morphological sterna II–VII); visible sternites 1, 2 and 3 completely fused, with sutures indistinct (positions marked by row of setae); first visible sternite deeply and broadly divided medially for reception of metacoxal processes (
|
||
<figureCitation box="[1179,1248,1642,1664]" captionStart="Fig" captionStartId="10.[151,185,1816,1836]" captionTargetBox="[167,742,1322,1770]" captionTargetId="figure-18@10.[167,742,1305,1788]" captionTargetPageId="10" captionText="Fig. 13. Meru phyllisae, visible abdominal sterna." figureDoi="http://doi.org/10.5281/zenodo.4311882" httpUri="https://zenodo.org/record/4311882/files/figure.png" pageId="7" pageNumber="346">Fig. 13</figureCitation>
|
||
), internally with a heavily sclerotized, transverse septum along posterior margin; posterior margins of sterna 1–5 bearing a comblike row of robust, flattened and grooved setae and forming a fringe along with fewer hairlike setae (
|
||
<figureCitation box="[1230,1307,1759,1781]" captionStart="Fig" captionStartId="2.[151,185,1603,1623]" captionTargetBox="[152,1426,156,1574]" captionTargetId="figure-70@2.[152,1426,156,1574]" captionTargetPageId="2" captionText="Fig. 2. Meru phyllisae. Scanning electron micrographs. A, Habitus, dorsal view; B, habitus, ventral view; C, thoracic sterna; D, head, pronotum and bases of elytra; E, head and prosternal area; F, metasternum and abdomen; G, right elytron; H, pronotal and elytral bases; I, pronotum, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4311858" httpUri="https://zenodo.org/record/4311858/files/figure.png" pageId="7" pageNumber="346">Figs 2F</figureCitation>
|
||
,
|
||
<figureCitation box="[1320,1381,1759,1781]" captionStart="Fig" captionStartId="5.[161,195,1789,1809]" captionTargetBox="[161,1436,820,1761]" captionTargetId="figure-352@5.[160,1437,819,1761]" captionTargetPageId="5" captionText="Fig. 5. Meru phyllisae. A, Left elytron, lateral view; B, left elytron, detail of margin; C, left elytron, inner surface; D, left elytron, wing-folding patch on inner surface; E, abdominal sternites, detail of posterior margins of 1–3; F, abdominal sternite 3, detail of posterior margin." figureDoi="http://doi.org/10.5281/zenodo.4311864" httpUri="https://zenodo.org/record/4311864/files/figure.png" pageId="7" pageNumber="346">5E, F</figureCitation>
|
||
,
|
||
<figureCitation box="[1395,1425,1759,1781]" captionStart="Fig" captionStartId="7.[830,864,1360,1380]" captionTargetBox="[834,1434,158,1330]" captionTargetId="figure-215@7.[831,1436,156,1331]" captionTargetPageId="7" captionText="Fig. 8. Meru phyllisae, sterna. A, Outline of prothorax; B, hind body." figureDoi="http://doi.org/10.5281/zenodo.4311868" httpUri="https://zenodo.org/record/4311868/files/figure.png" pageId="7" pageNumber="346">8B</figureCitation>
|
||
); sternites 1 and 2 with punctures bearing robust wrinkled setae scattered over surface; the flat pleated setae on sternites 3–5
|
||
</paragraph>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.4311870" ID-Zenodo-Dep="4311870" httpUri="https://zenodo.org/record/4311870/files/figure.png" pageId="7" pageNumber="346" startId="7.[161,195,1563,1583]" targetBox="[161,766,157,1532]" targetPageId="7">
|
||
<paragraph blockId="7.[161,768,1563,1609]" pageId="7" pageNumber="346">
|
||
<emphasis bold="true" box="[161,220,1563,1583]" pageId="7" pageNumber="346">Fig. 7.</emphasis>
|
||
<taxonomicName authorityName="Spangler & Steiner" authorityYear="2005" box="[231,363,1564,1583]" class="Insecta" family="Meruidae" genus="Meru" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="346" phylum="Arthropoda" rank="species" species="phyllisae">
|
||
<emphasis box="[231,363,1564,1583]" italics="true" pageId="7" pageNumber="346">Meru phyllisae</emphasis>
|
||
</taxonomicName>
|
||
. A, Mandibles, dorsal view; B, maxilla, left, ventral view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="8.[821,1427,156,441]" pageId="8" pageNumber="347">
|
||
<emphasis box="[845,1202,156,177]" italics="true" pageId="8" pageNumber="347">Wrinkled flat setae on the body.</emphasis>
|
||
These appear to be membranous papillae, very different from the normal, fine setae among them. It is possible that these structures may have a respiratory function, but determining their connection to the tracheal system was beyond the scope of this study. On dry beetles examined under the scanning electron microscope, broken examples of the papillae have been seen and they are hollow, but details of the basal attachment have not been clearly viewed; no opening to the cuticle was observed.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="8" pageNumber="347" type="etymology">
|
||
<paragraph blockId="8.[821,1426,477,528]" pageId="8" pageNumber="347">
|
||
<emphasis box="[845,961,478,499]" italics="true" pageId="8" pageNumber="347">Etymology.</emphasis>
|
||
‘Meru’ means ‘waterfall’ in the language of the Pemon people of the tepui region of
|
||
<collectingCountry box="[1237,1344,506,528]" name="Venezuela" pageId="8" pageNumber="347">Venezuela</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |